treatments-xml/data/6B/00/87/6B0087EEFFD1FFA4FE94FF497627FD50.xml

<document id="CB0869FF448242F907CD6A04C9659E79" ID-DOI="10.11606/1807-0205/2020.60.special-issue.24" ID-ISSN="1807-0205" ID-Zenodo-Dep="4985575" ID-ZooBank="7863784D-C29C-47AC-B52A-317234626D39" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1624029768623" checkinUser="carolina" docAuthor="Vaz, Stephanie, Silveira, Luiz Felipe Lima da &amp; Rosa, Simone Policena" docDate="2020" docId="6B0087EEFFD1FFA4FE94FF497627FD50" docLanguage="en" docName="PapAvZool.60.special-issue.e202060.24.pdf" docOrigin="Papéis Avulsos de Zoologia (Pap. Avulsos Zool., S. Paulo) 60 (24)" docSource="http://dx.doi.org/10.11606/1807-0205/2020.60.special-issue.24" docStyle="DocumentStyle:6844D5473582683DD13A9A75B001759F.4:PapAvZool.2018-.journal_article" docStyleId="6844D5473582683DD13A9A75B001759F" docStyleName="PapAvZool.2018-.journal_article" docStyleVersion="4" docTitle="Psilocladus costae Vaz &amp; Silveira &amp; Rosa 2020, sp. nov." docType="treatment" docVersion="3" lastPageNumber="12" masterDocId="9739FF96FFD3FFA9FFDEFF937719FFA3" masterDocTitle="Morphology and life cycle of a new species of Psilocladus Blanchard, 1846 (Coleoptera, Lampyridae, Psilocladinae), the first known bromeliad-inhabiting firefly" masterLastPageNumber="15" masterPageNumber="1" pageNumber="3" updateTime="1698969400221" updateUser="plazi" zenodo-license-document="CC-BY-NC-4.0" zenodo-license-figures="CC-BY-NC-4.0">
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<mods:title id="1566D8696B51D481D3A9407B66C22BFC">Morphology and life cycle of a new species of Psilocladus Blanchard, 1846 (Coleoptera, Lampyridae, Psilocladinae), the first known bromeliad-inhabiting firefly</mods:title>
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<mods:namePart id="10F6A58A766E85B9AD4AC09795A09644">Vaz, Stephanie</mods:namePart>
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<mods:namePart id="8C3BDEA0C5295162897B044F88EF9A57">Silveira, Luiz Felipe Lima da</mods:namePart>
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<mods:namePart id="4B31BF4D079725C2740E599DA9DF4B4D">Rosa, Simone Policena</mods:namePart>
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<mods:title id="D8126B3C3324D674EF8DD67FDA0745B2">Papéis Avulsos de Zoologia</mods:title>
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<mods:date id="4CE17828C4BDD6241D2C4684986EFEA2">2020</mods:date>
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<mods:title id="71E07ADFBD326BF60266AFD17DA2263F">Pap. Avulsos Zool., S. Paulo</mods:title>
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<mods:number id="07F111302AAFA43C2116A9352AC24184">60</mods:number>
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<paragraph id="E31636F8FFD1FFABFE94FF497563FF50" blockId="2.[170,795,218,595]" box="[330,634,218,243]" pageId="2" pageNumber="3">
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<taxonomicName id="24A94D7BFFD1FFABFE94FF497502FF50" ID-CoL="8NC64" authority="Vaz &amp; Silveira &amp; Rosa, 2020" authorityName="Vaz &amp; Silveira &amp; Rosa" authorityYear="2020" box="[330,539,218,243]" class="Insecta" family="Lampyridae" genus="Psilocladus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="costae" status="sp. nov.">
<emphasis id="D1DDEAEAFFD1FFABFE94FF497502FF50" bold="true" box="[330,539,218,243]" italics="true" pageId="2" pageNumber="3">Psilocladus costae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAEE5791FFD1FFABFDFEFF497563FF50" box="[544,634,218,243]" pageId="2" pageNumber="3" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="ABB36573FFD1FFABFE7FFF69753AFEB0" box="[417,547,250,275]" pageId="2" pageNumber="3" type="description">
<paragraph id="E31636F8FFD1FFABFE7FFF69753AFEB0" blockId="2.[170,795,218,595]" box="[417,547,250,275]" pageId="2" pageNumber="3">
<emphasis id="D1DDEAEAFFD1FFABFE7FFF69753AFEB0" bold="true" box="[417,547,250,275]" pageId="2" pageNumber="3">
(
<figureCitation id="7B922A7DFFD1FFABFE76FF697502FEB0" box="[424,539,250,275]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStart-10="Figure 13" captionStart-11="Figure 14" captionStart-2="Figure 5" captionStart-3="Figure 6" captionStart-4="Figure 7" captionStart-5="Figure 8" captionStart-6="Figure 9" captionStart-7="Figure 10" captionStart-8="Figure 11" captionStart-9="Figure 12" captionStartId-0="3.[94,153,2106,2131]" captionStartId-1="4.[170,229,2074,2099]" captionStartId-10="12.[170,229,1198,1223]" captionStartId-11="12.[170,229,2106,2131]" captionStartId-2="5.[94,153,2074,2099]" captionStartId-3="6.[172,231,2074,2099]" captionStartId-4="7.[94,153,914,939]" captionStartId-5="7.[94,153,2106,2131]" captionStartId-6="8.[170,229,2074,2099]" captionStartId-7="9.[94,153,2106,2131]" captionStartId-8="10.[170,229,2074,2099]" captionStartId-9="11.[94,153,2074,2099]" captionTargetBox-0="[96,1414,1301,2076]" captionTargetBox-1="[175,1490,721,2029]" captionTargetBox-10="[174,1491,210,1163]" captionTargetBox-11="[170,1493,1245,2076]" captionTargetBox-2="[166,1348,217,2025]" captionTargetBox-3="[176,1493,913,2040]" captionTargetBox-4="[97,1416,212,872]" captionTargetBox-5="[177,1339,1008,2071]" captionTargetBox-6="[183,1491,205,2030]" captionTargetBox-7="[100,1413,1083,2074]" captionTargetBox-8="[176,1489,864,2043]" captionTargetBox-9="[258,1262,1269,2034]" captionTargetId-0="figure-558@3.[94,1417,1301,2076]" captionTargetId-1="figure-225@4.[170,1493,721,2044]" captionTargetId-10="figure-1@12.[170,1493,204,1168]" captionTargetId-11="figure-22@12.[170,1493,1245,2076]" captionTargetId-2="figure-1@5.[151,1360,204,2044]" captionTargetId-3="figure-361@6.[170,1493,910,2044]" captionTargetId-4="figure-21@7.[94,1417,204,884]" captionTargetId-5="figure-1@7.[170,1342,999,2079]" captionTargetId-6="figure-1@8.[170,1493,204,2044]" captionTargetId-7="figure-434@9.[94,1417,1075,2077]" captionTargetId-8="figure-330@10.[170,1493,854,2045]" captionTargetId-9="figure-570@11.[246,1266,1269,2044]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-10="12" captionTargetPageId-11="12" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="7" captionTargetPageId-6="8" captionTargetPageId-7="9" captionTargetPageId-8="10" captionTargetPageId-9="11" captionText-0="Figure 3. Psilocladus costae sp.nov.Adult male habitus.(A) Dorsal;(B) lateral;(C) ventral.Scale bar:= 2 mm." captionText-1="Figure 4. Psilocladus costae sp. nov. Head morphology of adult male. (A-F) Head; (A) dorsal; (B) ventral;(C) lateral; (D) frontal; (E) occipital; (F) posterior;(G) left antenna (dorsal).Scale bar:= 1 mm." captionText-10="Figure 13. Psilocladus costae sp.nov.Pupal habitus.(A) Dorsal;(B) lateral;(C) ventral.Scale bar:= 2 mm." captionText-11="Figure 14. Psilocladus costae sp.nov.Immature bioluminescence.(A) Larva (dorsal);(B) pupa (ventral).Length:larva = 18 mm;pupa = 7 mm." captionText-2="Figure 5. Psilocladus costae sp.nov.Thorax morphology of adult male.(A-E) Prothorax:(A) dorsal;(B) ventral;(C) frontal;(D) posterior;(E)lateral;(F-I) pterothorax, (F) dorsal;(G) lateral;(H) dorsoventral.Scale bar:= 2 mm." captionText-3="Figure 6.Psilocladus costae sp.nov.Thoracic appendages of adult male. (A-C) Left legs,ventral (anterior, middle and posterior,respectively);(D-F) elytron (dorsal, lateral and ventral);(G) left wing.Scale bar:2 mm." captionText-4="Figure 7. Psilocladuscostae sp.nov.Terminal segments and aedeagus of male.(A) SternumVIII;(B) pygidium;(C) syntergite;(D) sternumIX;(E-G) aedeagus (dorsal, lateral and ventral).Scale bar:= 1 mm." captionText-5="Figure 8. Psilocladus costae sp.nov.Adult female habitus.(A) Dorsal;(B) ventral.Scale bar:= 5 mm." captionText-6="Figure 9. Psilocladus costae sp. nov. Adult female morphology. (A-E) Head; (A) dorsal; (B) ventral; (C) frontal; (D) occipital; (E) lateral; (F) left antenna (dorsal); (G) sternumVIII;(H) pygidium;(I, J) ovipositor (dorsal,ventral),arrow indicates stylus.Scale bars:A-E = 1 mm;F-I = 2 mm." captionText-7="Figure 10. Psilocladus costae sp.nov.Mature larval habitus.(A) Dorsal;(B) lateral;(C) ventral.Scale bar:= 2 mm." captionText-8="Figure 11. Psilocladus costae sp. nov. Larval morphology. (A-C) Head (dorsal, lateral, ventral); (D) right anterior, middle and posterior legs (from top to bottom, ventral);(E) left laterotergitesVI-VII;(F) abdominal tergaVI-IX.Scale bars:A-C,E, F = 1 mm;D = 2 mm." captionText-9="Figure 12.Psilocladus costae sp.nov.Larval headmorphology.(A) Head,dorsal;(B)maxillolabial complex,ventral;(C) epipharynx;(D) hypopharynx;(E) leftantenna (ventral,dorsal);(F) mandible (ventral,dorsal).Scale bars:A,B = 1 mm;C,D = 0.25 mm;E,F = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4985583" figureDoi-1="http://doi.org/10.5281/zenodo.4985585" figureDoi-10="http://doi.org/10.5281/zenodo.4985605" figureDoi-11="http://doi.org/10.5281/zenodo.4985609" figureDoi-2="http://doi.org/10.5281/zenodo.4985587" figureDoi-3="http://doi.org/10.5281/zenodo.4985589" figureDoi-4="http://doi.org/10.5281/zenodo.4985591" figureDoi-5="http://doi.org/10.5281/zenodo.4985593" figureDoi-6="http://doi.org/10.5281/zenodo.4985595" figureDoi-7="http://doi.org/10.5281/zenodo.4985597" figureDoi-8="http://doi.org/10.5281/zenodo.4985599" figureDoi-9="http://doi.org/10.5281/zenodo.4985603" httpUri-0="https://zenodo.org/record/4985583/files/figure.png" httpUri-1="https://zenodo.org/record/4985585/files/figure.png" httpUri-10="https://zenodo.org/record/4985605/files/figure.png" httpUri-11="https://zenodo.org/record/4985609/files/figure.png" httpUri-2="https://zenodo.org/record/4985587/files/figure.png" httpUri-3="https://zenodo.org/record/4985589/files/figure.png" httpUri-4="https://zenodo.org/record/4985591/files/figure.png" httpUri-5="https://zenodo.org/record/4985593/files/figure.png" httpUri-6="https://zenodo.org/record/4985595/files/figure.png" httpUri-7="https://zenodo.org/record/4985597/files/figure.png" httpUri-8="https://zenodo.org/record/4985599/files/figure.png" httpUri-9="https://zenodo.org/record/4985603/files/figure.png" pageId="2" pageNumber="3">Figs. 3-14</figureCitation>
)
</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="ABB36573FFD1FFA3FF74FEA97328FD70" lastPageId="10" lastPageNumber="11" pageId="2" pageNumber="3" type="materials_examined">
<paragraph id="E31636F8FFD1FFABFF74FEA97414FDF0" blockId="2.[170,795,218,595]" pageId="2" pageNumber="3">
<emphasis id="D1DDEAEAFFD1FFABFF74FEA9764FFEF0" bold="true" box="[170,342,314,339]" pageId="2" pageNumber="3">Type material:</emphasis>
<materialsCitation id="53C13CA5FFD1FFABFE81FEA9755EFE30" collectingDate="2017-07-02" collectionCode="CLEI" collectorName="S. Rosa &amp; Ladenthin" country="Brazil" location="Campos do Jordao" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo" typeStatus="holotype">
<typeStatus id="3C12885AFFD1FFABFE81FEA976C1FEF0" box="[351,472,314,339]" pageId="2" pageNumber="3" type="holotype">HOLOTYPE</typeStatus>
<specimenCount id="F5AFFD71FFD1FFABFE3EFEA476EAFEF1" box="[480,499,311,338]" pageId="2" pageNumber="3" type="male">♂</specimenCount>
(
<collectionCode id="85B8AE3DFFD1FFABFDDFFEA9752EFEF0" box="[513,567,314,339]" pageId="2" pageNumber="3">CLEI</collectionCode>
): 
<collectingCountry id="9BBE7668FFD1FFABFD96FEA97583FEF0" box="[584,666,314,339]" name="Brazil" pageId="2" pageNumber="3">BRAZIL</collectingCountry>
. 
<collectingRegion id="216DF81AFFD1FFABFD7BFEA9740EFEF0" box="[677,791,314,339]" country="Brazil" name="Sao Paulo" pageId="2" pageNumber="3">São Paulo</collectingRegion>
: 
<location id="E6766023FFD1FFABFF74FEC9769AFED0" LSID="urn:lsid:plazi:treatment:6B0087EEFFD1FFA4FE94FF497627FD50:E6766023FFD1FFABFF74FEC9769AFED0" box="[170,387,346,371]" country="Brazil" name="Campos do Jordao" pageId="2" pageNumber="3" stateProvince="Sao Paulo">Campos do Jordão</location>
, 22°40′51.3″, 45°35′30.6″, 
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, 
<date id="97171038FFD1FFABFF74FEE9763CFE30" box="[170,293,378,403]" pageId="2" pageNumber="3" value="2017-07-02">
<collectingDate id="8753E9D0FFD1FFABFF74FEE9763CFE30" box="[170,293,378,403]" pageId="2" pageNumber="3" value="2017-07-02">02.VII.2017</collectingDate>
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, 
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&amp; 
<collectorName id="4E5C532EFFD1FFABFE7AFEE9750CFE30" box="[420,533,378,403]" pageId="2" pageNumber="3">Ladenthin</collectorName>
leg.
</materialsCitation>
<materialsCitation id="53C13CA5FFD1FFABFD8EFEE97641FE50" collectingDate="2016-12-23" collectingDateMax="2017-12-04" collectingDateMin="2016-12-23" collectionCode="CLEI" collectorName="S. Rosa &amp; Ladenthin" country="Brazil" location="Same" pageId="2" pageNumber="3" specimenCount="3" specimenCount-male="3" stateProvince="Sao Paulo" typeStatus="paratype">
<typeStatus id="3C12885AFFD1FFABFD8EFEE975C8FE30" box="[592,721,378,403]" pageId="2" pageNumber="3" type="paratype">PARATYPES</typeStatus>
. 
<location id="E6766023FFD1FFABFD02FEE97403FE30" LSID="urn:lsid:plazi:treatment:6B0087EEFFD1FFA4FE94FF497627FD50:E6766023FFD1FFABFD02FEE97403FE30" box="[732,794,378,403]" country="Brazil" name="Same" pageId="2" pageNumber="3" stateProvince="Sao Paulo">Same</location>
locality of 
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, 
<date id="97171038FFD1FFABFE7EFE097505FE10" box="[416,540,410,435]" pageId="2" pageNumber="3" value="2016-12-23">
<collectingDate id="8753E9D0FFD1FFABFE7EFE097505FE10" box="[416,540,410,435]" pageId="2" pageNumber="3" value="2016-12-23">23.XII.2016</collectingDate>
</date>
, 
<collectorName id="4E5C532EFFD1FFABFDF5FE097599FE10" box="[555,640,410,435]" pageId="2" pageNumber="3">S. Rosa</collectorName>
&amp; 
<collectorName id="4E5C532EFFD1FFABFD77FE097403FE10" box="[681,794,410,435]" pageId="2" pageNumber="3">Ladenthin</collectorName>
(
<specimenCount id="F5AFFD71FFD1FFABFF6FFE2977F8FE71" box="[177,225,439,467]" pageId="2" pageNumber="3" type="male">3 ♂</specimenCount>
, 
<collectionCode id="85B8AE3DFFD1FFABFF2BFE297630FE70" box="[245,297,442,467]" pageId="2" pageNumber="3">CLEI</collectionCode>
); (collected as larva on 
<date id="97171038FFD1FFABFD8DFE2975D7FE70" box="[595,718,442,467]" pageId="2" pageNumber="3" value="2017-07-02">
<collectingDate id="8753E9D0FFD1FFABFD8DFE2975D7FE70" box="[595,718,442,467]" pageId="2" pageNumber="3" value="2017-07-02">02.VII.2017</collectingDate>
</date>
, adult on 
<date id="97171038FFD1FFABFF0BFE49764BFE50" box="[213,338,474,499]" pageId="2" pageNumber="3" value="2017-12-04">
<collectingDate id="8753E9D0FFD1FFABFF0BFE49764BFE50" box="[213,338,474,499]" pageId="2" pageNumber="3" value="2017-12-04">04.XII.2017</collectingDate>
</date>
)
</materialsCitation>
, 
<materialsCitation id="53C13CA5FFD1FFABFEB6FE49740EFE50" box="[360,791,471,499]" collectionCode="CLEI" collectorName="S. Rosa &amp; Ladenthin" country="Brazil" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo" typeStatus="paratype">
<collectorName id="4E5C532EFFD1FFABFEB6FE4976A4FE50" box="[360,445,474,499]" pageId="2" pageNumber="3">S. Rosa</collectorName>
&amp; 
<collectorName id="4E5C532EFFD1FFABFE39FE497541FE50" box="[487,600,474,499]" pageId="2" pageNumber="3">Ladenthin</collectorName>
leg (
<specimenCount id="F5AFFD71FFD1FFABFD41FE4975D2FE51" box="[671,715,471,499]" pageId="2" pageNumber="3" type="male">1 ♂</specimenCount>
, 
<collectionCode id="85B8AE3DFFD1FFABFD03FE497408FE50" box="[733,785,474,499]" pageId="2" pageNumber="3">CLEI</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="53C13CA5FFD1FFABFF74FE6975D0FDB0" box="[170,713,503,531]" collectingDate="2017-12-02" collectionCode="CLEI" collectorName="Rosa &amp; Ladenthin" country="Brazil" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo" typeStatus="paratype">
<date id="97171038FFD1FFABFF74FE69763FFDB0" box="[170,294,506,531]" pageId="2" pageNumber="3" value="2017-12-02">
<collectingDate id="8753E9D0FFD1FFABFF74FE69763FFDB0" box="[170,294,506,531]" pageId="2" pageNumber="3" value="2017-12-02">02.XII.2017</collectingDate>
</date>
, 
<collectorName id="4E5C532EFFD1FFABFEECFE697670FDB0" box="[306,361,506,531]" pageId="2" pageNumber="3">Rosa</collectorName>
, S &amp; 
<collectorName id="4E5C532EFFD1FFABFE79FE697501FDB0" box="[423,536,506,531]" pageId="2" pageNumber="3">Ladenthin</collectorName>
leg (
<specimenCount id="F5AFFD71FFD1FFABFD86FE697599FDB1" box="[600,640,503,531]" pageId="2" pageNumber="3" type="male">1 ♂</specimenCount>
, 
<collectionCode id="85B8AE3DFFD1FFABFD51FE6975DAFDB0" box="[655,707,506,531]" pageId="2" pageNumber="3">CLEI</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="53C13CA5FFD1FFABFD0BFE697574FD90" collectingDate="2017-12-08" collectingDateMax="2017-12-10" collectingDateMin="2017-12-08" collectionCode="MZSP" collectorName="S. Rosa &amp; Ladenthin" country="Brazil" pageId="2" pageNumber="3" specimenCount="3" specimenCount-male="3" stateProvince="Sao Paulo" typeStatus="paratype">
<date id="97171038FFD1FFABFD0BFE69761CFD90" pageId="2" pageNumber="3" value="2017-12-08" valueMax="2017-12-10" valueMin="2017-12-08">
<collectingDate id="8753E9D0FFD1FFABFD0BFE69761CFD90" pageId="2" pageNumber="3" value="2017-12-08" valueMax="2017-12-10" valueMin="2017-12-08">08-10. XII.2017</collectingDate>
</date>
, 
<collectorName id="4E5C532EFFD1FFABFED3FD897643FD90" box="[269,346,538,563]" pageId="2" pageNumber="3">S. Rosa</collectorName>
&amp; 
<collectorName id="4E5C532EFFD1FFABFEABFD8976FFFD90" box="[373,486,538,563]" pageId="2" pageNumber="3">Ladenthin</collectorName>
(
<specimenCount id="F5AFFD71FFD1FFABFE2CFD897500FD91" box="[498,537,535,563]" pageId="2" pageNumber="3" type="male">3 ♂</specimenCount>
, 
<collectionCode id="85B8AE3DFFD1FFABFDFDFD89757EFD90" box="[547,615,538,563]" country="Brazil" httpUri="http://grbio.org/cool/9yp6-zxp9" name="Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="3" type="Museum">MZSP</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="53C13CA5FFD1FFABFDABFD897666FDF0" collectingDate="2018-01-18" collectingDateMax="2018-01-20" collectingDateMin="2018-01-18" collectionCode="CLEI" collectorName="S. Rosa" country="Brazil" pageId="2" pageNumber="3" specimenCount="1" specimenCount-male="1" stateProvince="Sao Paulo" typeStatus="paratype">
<date id="97171038FFD1FFABFDABFD8975E6FD90" box="[629,767,538,563]" pageId="2" pageNumber="3" value="2018-01-18" valueMax="2018-01-20" valueMin="2018-01-18">
<collectingDate id="8753E9D0FFD1FFABFDABFD8975E6FD90" box="[629,767,538,563]" pageId="2" pageNumber="3" value="2018-01-18" valueMax="2018-01-20" valueMin="2018-01-18">18-20.I.2018</collectingDate>
</date>
, 
<collectorName id="4E5C532EFFD1FFABFCD9FD8977C6FDF0" pageId="2" pageNumber="3">S. Rosa</collectorName>
leg (
<specimenCount id="F5AFFD71FFD1FFABFEC8FDA97623FDF1" box="[278,314,567,595]" pageId="2" pageNumber="3" type="male">1 ♂</specimenCount>
, 
<collectionCode id="85B8AE3DFFD1FFABFE9BFDA97660FDF0" box="[325,377,570,595]" pageId="2" pageNumber="3">CLEI</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="53C13CA5FFD1FFABFE59FDA97413FDF0" box="[391,778,567,595]" collectingDate="2018-02-08" collectingDateMax="2018-02-13" collectingDateMin="2018-02-08" collectionCode="CLEI" collectorName="S. Rosa" country="Brazil" pageId="2" pageNumber="3" specimenCount="1" specimenCount-female="1" stateProvince="Sao Paulo" typeStatus="paratype">
<date id="97171038FFD1FFABFE59FDA9750EFDF0" box="[391,535,570,595]" pageId="2" pageNumber="3" value="2018-02-08" valueMax="2018-02-13" valueMin="2018-02-08">
<collectingDate id="8753E9D0FFD1FFABFE59FDA9750EFDF0" box="[391,535,570,595]" pageId="2" pageNumber="3" value="2018-02-08" valueMax="2018-02-13" valueMin="2018-02-08">08-13.II.2018</collectingDate>
</date>
, 
<collectorName id="4E5C532EFFD1FFABFDFEFDA97575FDF0" box="[544,620,570,595]" pageId="2" pageNumber="3">S. Rosa</collectorName>
leg (
<specimenCount id="F5AFFD71FFD1FFABFD7BFDA975DCFDF1" box="[677,709,567,595]" pageId="2" pageNumber="3" type="female">1 ♀</specimenCount>
, 
<collectionCode id="85B8AE3DFFD1FFABFD11FDA9741AFDF0" box="[719,771,570,595]" pageId="2" pageNumber="3">CLEI</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="E31636F8FFD1FFABFF74FDE9754FFB90" blockId="2.[170,795,634,1075]" pageId="2" pageNumber="3">
<emphasis id="D1DDEAEAFFD1FFABFF74FDE9750AFD30" bold="true" box="[170,531,634,659]" pageId="2" pageNumber="3">Additional material examined:</emphasis>
<materialsCitation id="53C13CA5FFD1FFABFDC7FDE975AFFD10" collectingDate="1998-12" collectionCode="MNHN" collectorName="Gounelle, E." country="Brazil" location="Vale do Rio Pardo" pageId="2" pageNumber="3" specimenCount="1" specimenCount-female="1" stateProvince="Sao Paulo">
<collectingCountry id="9BBE7668FFD1FFABFDC7FDE97572FD30" box="[537,619,634,659]" name="Brazil" pageId="2" pageNumber="3">BRAZIL</collectingCountry>
. 
<collectingRegion id="216DF81AFFD1FFABFDAAFDE975FDFD30" box="[628,740,634,659]" country="Brazil" name="Sao Paulo" pageId="2" pageNumber="3">São Paulo</collectingRegion>
: 
<location id="E6766023FFD1FFABFD35FDE97623FD10" LSID="urn:lsid:plazi:treatment:6B0087EEFFD1FFA4FE94FF497627FD50:E6766023FFD1FFABFD35FDE97623FD10" country="Brazil" name="Vale do Rio Pardo" pageId="2" pageNumber="3" stateProvince="Sao Paulo">Vale do Rio Pardo</location>
, 
<date id="97171038FFD1FFABFE9CFD097685FD10" box="[322,412,666,691]" pageId="2" pageNumber="3" value="1998-12">
<collectingDate id="8753E9D0FFD1FFABFE9CFD097685FD10" box="[322,412,666,691]" pageId="2" pageNumber="3" value="1998-12">XII.1998</collectingDate>
</date>
, 
<collectorName id="4E5C532EFFD1FFABFE7BFD097531FD10" box="[421,552,666,691]" pageId="2" pageNumber="3">Gounelle, E.</collectorName>
(
<specimenCount id="F5AFFD71FFD1FFABFDE9FD09754EFD11" box="[567,599,663,691]" pageId="2" pageNumber="3" type="female">1 ♀</specimenCount>
, 
<collectionCode id="85B8AE3DFFD1FFABFDBCFD0975A9FD10" box="[610,688,666,691]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="3">MNHN</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="53C13CA5FFD1FFABFD61FD09766AFCB0" collectingDate="2018-07-05" collectingDateMax="2018-12-17" collectingDateMin="2018-07-05" collectionCode="CLEI" collectorName="S. Rosa" country="Brazil" elevation="1464" location="Campos do Jordao" pageId="2" pageNumber="3" specimenCount="1" specimenCount-pupa="1">
<location id="E6766023FFD1FFABFD61FD09763AFD70" LSID="urn:lsid:plazi:treatment:6B0087EEFFD1FFA4FE94FF497627FD50:E6766023FFD1FFABFD61FD09763AFD70" country="Brazil" name="Campos do Jordao" pageId="2" pageNumber="3">Campos do Jordão</location>
, 22°40′51.3″, 45°35′30.6″, 
<quantity id="24519B1DFFD1FFABFD8AFD2975AAFD70" box="[596,691,698,723]" metricMagnitude="3" metricUnit="m" metricValue="1.464" pageId="2" pageNumber="3" unit="m" value="1464.0">
<elevation id="6884D1CBFFD1FFABFD8AFD2975AAFD70" box="[596,691,698,723]" metricMagnitude="3" metricUnit="m" metricValue="1.464" pageId="2" pageNumber="3" unit="m" value="1464.0">1,464 m</elevation>
</quantity>
(collect- ed as larva on 
<date id="97171038FFD1FFABFE8EFD4976D5FD50" box="[336,460,730,755]" pageId="2" pageNumber="3" value="2018-07-05">
<collectingDate id="8753E9D0FFD1FFABFE8EFD4976D5FD50" box="[336,460,730,755]" pageId="2" pageNumber="3" value="2018-07-05">05.VII.2018</collectingDate>
</date>
, pupa on 
<date id="97171038FFD1FFABFDE3FD4975A3FD50" box="[573,698,730,755]" pageId="2" pageNumber="3" value="2018-12-17">
<collectingDate id="8753E9D0FFD1FFABFDE3FD4975A3FD50" box="[573,698,730,755]" pageId="2" pageNumber="3" value="2018-12-17">17.XII.2018</collectingDate>
</date>
), 
<collectorName id="4E5C532EFFD1FFABFD15FD497403FD50" box="[715,794,730,755]" pageId="2" pageNumber="3">S. Rosa</collectorName>
leg. (
<specimenCount id="F5AFFD71FFD1FFABFF3CFD697636FCB0" box="[226,303,762,787]" pageId="2" pageNumber="3" type="pupa">1 pupa</specimenCount>
, 
<collectionCode id="85B8AE3DFFD1FFABFEE7FD697674FCB0" box="[313,365,762,787]" pageId="2" pageNumber="3">CLEI</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="53C13CA5FFD1FFABFEA2FD69740EFCB0" box="[380,791,762,787]" collectingDate="2017-05-27" collectingDateMax="2018-12-17" collectingDateMin="2018-07-05" collectionCode="CLEI" collectorName="S. Rosa" country="Brazil" elevation="1464" isTailEnumeration="true" location="Campos do Jordao" pageId="2" pageNumber="3" specimenCount="2" specimenCount-larva="2">
<date id="97171038FFD1FFABFEA2FD6976F0FCB0" box="[380,489,762,787]" pageId="2" pageNumber="3" value="2017-05-27">
<collectingDate id="8753E9D0FFD1FFABFEA2FD6976F0FCB0" box="[380,489,762,787]" pageId="2" pageNumber="3" value="2017-05-27">27.V.2017</collectingDate>
</date>
, 
<collectorName id="4E5C532EFFD1FFABFE2CFD697526FCB0" box="[498,575,762,787]" pageId="2" pageNumber="3">S. Rosa</collectorName>
leg. (
<specimenCount id="F5AFFD71FFD1FFABFDA7FD6975CDFCB0" box="[633,724,762,787]" pageId="2" pageNumber="3" type="larva">2 larvae</specimenCount>
, 
<collectionCode id="85B8AE3DFFD1FFABFD03FD697408FCB0" box="[733,785,762,787]" pageId="2" pageNumber="3">CLEI</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="53C13CA5FFD1FFABFF74FC89755EFC90" box="[170,583,794,819]" collectingDate="2017-11-09" collectionCode="UNIFEI" collectorName="S Rosa" country="Brazil" pageId="2" pageNumber="3" specimenCount="1" specimenCount-larva="1">
<date id="97171038FFD1FFABFF74FC897639FC90" box="[170,288,794,819]" pageId="2" pageNumber="3" value="2017-11-09">
<collectingDate id="8753E9D0FFD1FFABFF74FC897639FC90" box="[170,288,794,819]" pageId="2" pageNumber="3" value="2017-11-09">09.XI.2017</collectingDate>
</date>
, 
<collectorName id="4E5C532EFFD1FFABFEF8FC897675FC90" box="[294,364,794,819]" pageId="2" pageNumber="3">S Rosa</collectorName>
leg.(
<specimenCount id="F5AFFD71FFD1FFABFE7AFC8976F5FC90" box="[420,492,794,819]" pageId="2" pageNumber="3" type="larva">1 larva</specimenCount>
, 
<collectionCode id="85B8AE3DFFD1FFABFE2CFC897559FC90" box="[498,576,794,819]" pageId="2" pageNumber="3">UNIFEI</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="53C13CA5FFD1FFABFD93FC89769CFCF0" collectingDate="2017-12-04" collectionCode="UNIFEI" collectorName="S. Rosa" country="Brazil" pageId="2" pageNumber="3" specimenCount="1" specimenCount-larva="1">
<date id="97171038FFD1FFABFD93FC8975D1FC90" box="[589,712,794,819]" pageId="2" pageNumber="3" value="2017-12-04">
<collectingDate id="8753E9D0FFD1FFABFD93FC8975D1FC90" box="[589,712,794,819]" pageId="2" pageNumber="3" value="2017-12-04">04.XII.2017</collectingDate>
</date>
, 
<collectorName id="4E5C532EFFD1FFABFD11FC897403FC90" box="[719,794,794,819]" pageId="2" pageNumber="3">S.Rosa</collectorName>
leg.(
<specimenCount id="F5AFFD71FFD1FFABFF3EFCA97630FCF0" box="[224,297,826,851]" pageId="2" pageNumber="3" type="larva">1 larva</specimenCount>
, 
<collectionCode id="85B8AE3DFFD1FFABFEEEFCA97667FCF0" box="[304,382,826,851]" pageId="2" pageNumber="3">UNIFEI</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="53C13CA5FFD1FFABFE52FCA9740EFCF0" box="[396,791,826,851]" collectingDate="2018-03-22" collectionCode="CLEI" collectorName="S. Rosa" country="Brazil" pageId="2" pageNumber="3" specimenCount="1" specimenCount-larva="1">
<date id="97171038FFD1FFABFE52FCA976E6FCF0" box="[396,511,826,851]" pageId="2" pageNumber="3" value="2018-03-22">
<collectingDate id="8753E9D0FFD1FFABFE52FCA976E6FCF0" box="[396,511,826,851]" pageId="2" pageNumber="3" value="2018-03-22">22.III.2018</collectingDate>
</date>
, 
<collectorName id="4E5C532EFFD1FFABFDD8FCA9754BFCF0" box="[518,594,826,851]" pageId="2" pageNumber="3">S.Rosa</collectorName>
leg. (
<specimenCount id="F5AFFD71FFD1FFABFD53FCA975CFFCF0" box="[653,726,826,851]" pageId="2" pageNumber="3" type="larva">1 larva</specimenCount>
, 
<collectionCode id="85B8AE3DFFD1FFABFD03FCA97408FCF0" box="[733,785,826,851]" pageId="2" pageNumber="3">CLEI</collectionCode>
)
</materialsCitation>
; 
<date id="97171038FFD1FFABFF74FCC97604FCD0" box="[170,285,858,883]" pageId="2" pageNumber="3" value="2018-03-25">25.III.2018</date>
, S Rosa leg. (
<specimenCount id="F5AFFD71FFD1FFABFE69FCC9751DFCD0" box="[439,516,858,883]" pageId="2" pageNumber="3" type="larva">1 larva</specimenCount>
, 1 larval exuvia, UNIFEI); 
<materialsCitation id="53C13CA5FFD1FFABFF74FCE9756CFC30" box="[170,629,890,915]" collectingDate="2018-05-05" collectionCode="UNIFEI" collectorName="S. Rosa" country="Brazil" pageId="2" pageNumber="3" specimenCount="1" specimenCount-larva="1">
<date id="97171038FFD1FFABFF74FCE97601FC30" box="[170,280,890,915]" pageId="2" pageNumber="3" value="2018-05-05">
<collectingDate id="8753E9D0FFD1FFABFF74FCE97601FC30" box="[170,280,890,915]" pageId="2" pageNumber="3" value="2018-05-05">05.V.2018</collectingDate>
</date>
, 
<collectorName id="4E5C532EFFD1FFABFEF8FCE97660FC30" box="[294,377,890,915]" pageId="2" pageNumber="3">S. Rosa</collectorName>
leg. (
<specimenCount id="F5AFFD71FFD1FFABFE1CFCE9750BFC30" box="[450,530,890,915]" pageId="2" pageNumber="3" type="larva">1 larva</specimenCount>
, 
<collectionCode id="85B8AE3DFFD1FFABFDFFFCE97576FC30" box="[545,623,890,915]" pageId="2" pageNumber="3">UNIFEI</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="53C13CA5FFD1FFABFD5DFCE976D3FC10" collectingDate="2018-06-24" collectionCode="UNIFEI" collectorName="S. Rosa" country="Brazil" pageId="2" pageNumber="3" specimenCount="1" specimenCount-larva="1">
<date id="97171038FFD1FFABFD5DFCE975E1FC30" box="[643,760,890,915]" pageId="2" pageNumber="3" value="2018-06-24">
<collectingDate id="8753E9D0FFD1FFABFD5DFCE975E1FC30" box="[643,760,890,915]" pageId="2" pageNumber="3" value="2018-06-24">24.VI.2018</collectingDate>
</date>
, 
<collectorName id="4E5C532EFFD1FFABFCD9FCE977C6FC10" pageId="2" pageNumber="3">S. Rosa</collectorName>
leg. (
<specimenCount id="F5AFFD71FFD1FFABFEC1FC097672FC10" box="[287,363,922,947]" pageId="2" pageNumber="3" type="larva">1 larva</specimenCount>
, 
<collectionCode id="85B8AE3DFFD1FFABFEA8FC0976DDFC10" box="[374,452,922,947]" pageId="2" pageNumber="3">UNIFEI</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="53C13CA5FFD1FFABFE0AFC09762EFC70" collectingDate="2018-06-30" collectionCode="UNIFEI" collectorName="S. Rosa" country="Brazil" pageId="2" pageNumber="3" specimenCount="2" specimenCount-larva="2">
<date id="97171038FFD1FFABFE0AFC097550FC10" box="[468,585,922,947]" pageId="2" pageNumber="3" value="2018-06-30">
<collectingDate id="8753E9D0FFD1FFABFE0AFC097550FC10" box="[468,585,922,947]" pageId="2" pageNumber="3" value="2018-06-30">30.VI.2018</collectingDate>
</date>
, 
<collectorName id="4E5C532EFFD1FFABFD8DFC0975BAFC10" box="[595,675,922,947]" pageId="2" pageNumber="3">S. Rosa</collectorName>
leg. (
<specimenCount id="F5AFFD71FFD1FFABFD3EFC0977CAFC70" pageId="2" pageNumber="3" type="larva">2 larvae</specimenCount>
, 
<collectionCode id="85B8AE3DFFD1FFABFF3DFC297628FC70" box="[227,305,954,979]" pageId="2" pageNumber="3">UNIFEI</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="53C13CA5FFD1FFABFE98FC29740EFC70" box="[326,791,954,979]" collectingDate="2018-07-16" collectionCode="CLEI" collectorName="S. Rosa" country="Brazil" pageId="2" pageNumber="3" specimenCount="2" specimenCount-larva="2">
<date id="97171038FFD1FFABFE98FC2976DBFC70" box="[326,450,954,979]" pageId="2" pageNumber="3" value="2018-07-16">
<collectingDate id="8753E9D0FFD1FFABFE98FC2976DBFC70" box="[326,450,954,979]" pageId="2" pageNumber="3" value="2018-07-16">16.VII.2018</collectingDate>
</date>
, 
<collectorName id="4E5C532EFFD1FFABFE0FFC29753CFC70" box="[465,549,954,979]" pageId="2" pageNumber="3">S. Rosa</collectorName>
leg. (
<specimenCount id="F5AFFD71FFD1FFABFDB3FC2975D7FC70" box="[621,718,954,979]" pageId="2" pageNumber="3" type="larva">2 larvae</specimenCount>
, 
<collectionCode id="85B8AE3DFFD1FFABFD03FC297408FC70" box="[733,785,954,979]" pageId="2" pageNumber="3">CLEI</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="53C13CA5FFD1FFABFF74FC497551FC50" box="[170,584,986,1011]" collectingDate="2018-08-27" collectionCode="CLEI" collectorName="S. Rosa" country="Brazil" pageId="2" pageNumber="3" specimenCount="1" specimenCount-larva="1">
<date id="97171038FFD1FFABFF74FC497635FC50" box="[170,300,986,1011]" pageId="2" pageNumber="3" value="2018-08-27">
<collectingDate id="8753E9D0FFD1FFABFF74FC497635FC50" box="[170,300,986,1011]" pageId="2" pageNumber="3" value="2018-08-27">27.VIII.2018</collectingDate>
</date>
, 
<collectorName id="4E5C532EFFD1FFABFEEAFC497698FC50" box="[308,385,986,1011]" pageId="2" pageNumber="3">S. Rosa</collectorName>
leg. (
<specimenCount id="F5AFFD71FFD1FFABFE63FC49751FFC50" box="[445,518,986,1011]" pageId="2" pageNumber="3" type="larva">1 larva</specimenCount>
, 
<collectionCode id="85B8AE3DFFD1FFABFDD0FC49755BFC50" box="[526,578,986,1011]" pageId="2" pageNumber="3">CLEI</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="53C13CA5FFD1FFABFD8EFC497661FBB0" collectingDate="2018-09-17" collectionCode="CLEI" collectorName="S. Rosa" country="Brazil" pageId="2" pageNumber="3" specimenCount="2" specimenCount-larva="2">
<date id="97171038FFD1FFABFD8EFC4975DCFC50" box="[592,709,986,1011]" pageId="2" pageNumber="3" value="2018-09-17">
<collectingDate id="8753E9D0FFD1FFABFD8EFC4975DCFC50" box="[592,709,986,1011]" pageId="2" pageNumber="3" value="2018-09-17">17.IX.2018</collectingDate>
</date>
, 
<collectorName id="4E5C532EFFD1FFABFD13FC497403FC50" box="[717,794,986,1011]" pageId="2" pageNumber="3">S. Rosa</collectorName>
leg.(
<specimenCount id="F5AFFD71FFD1FFABFF03FC69762FFBB0" box="[221,310,1018,1043]" pageId="2" pageNumber="3" type="larva">2 larvae</specimenCount>
, 
<collectionCode id="85B8AE3DFFD1FFABFEE0FC69766BFBB0" box="[318,370,1018,1043]" pageId="2" pageNumber="3">CLEI</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="53C13CA5FFD1FFABFEA1FC69740EFBB0" box="[383,791,1018,1043]" collectingDate="2019-03-08" collectionCode="CLEI" collectorName="S. Rosa" country="Brazil" pageId="2" pageNumber="3" specimenCount="2" specimenCount-larva="2">
<date id="97171038FFD1FFABFEA1FC6976E8FBB0" box="[383,497,1018,1043]" pageId="2" pageNumber="3" value="2019-03-08">
<collectingDate id="8753E9D0FFD1FFABFEA1FC6976E8FBB0" box="[383,497,1018,1043]" pageId="2" pageNumber="3" value="2019-03-08">08.III.2019</collectingDate>
</date>
, 
<collectorName id="4E5C532EFFD1FFABFE27FC69755CFBB0" box="[505,581,1018,1043]" pageId="2" pageNumber="3">S.Rosa</collectorName>
leg. (
<specimenCount id="F5AFFD71FFD1FFABFDA3FC6975CFFBB0" box="[637,726,1018,1043]" pageId="2" pageNumber="3" type="larva">2 larvae</specimenCount>
, 
<collectionCode id="85B8AE3DFFD1FFABFD03FC697408FBB0" box="[733,785,1018,1043]" pageId="2" pageNumber="3">CLEI</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="53C13CA5FFD1FFABFF74FB89754BFB90" box="[170,594,1050,1075]" collectingDate="2019-10-27" collectionCode="UNIFEI" collectorName="S. Rosa" country="Brazil" pageId="2" pageNumber="3" specimenCount="1" specimenCount-larva="1">
<date id="97171038FFD1FFABFF74FB897600FB90" box="[170,281,1050,1075]" pageId="2" pageNumber="3" value="2019-10-27">
<collectingDate id="8753E9D0FFD1FFABFF74FB897600FB90" box="[170,281,1050,1075]" pageId="2" pageNumber="3" value="2019-10-27">27.X.2019</collectingDate>
</date>
, 
<collectorName id="4E5C532EFFD1FFABFEFCFB897676FB90" box="[290,367,1050,1075]" pageId="2" pageNumber="3">S. Rosa</collectorName>
leg. (
<specimenCount id="F5AFFD71FFD1FFABFE75FB8976ECFB90" box="[427,501,1050,1075]" pageId="2" pageNumber="3" type="larva">1 larva</specimenCount>
, 
<collectionCode id="85B8AE3DFFD1FFABFE20FB897555FB90" box="[510,588,1050,1075]" pageId="2" pageNumber="3">UNIFEI</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="E31636F8FFD1FFABFF74FBC977F5FBD0" blockId="2.[170,236,1114,1139]" box="[170,236,1114,1139]" pageId="2" pageNumber="3">
<emphasis id="D1DDEAEAFFD1FFABFF74FBC977F5FBD0" bold="true" box="[170,236,1114,1139]" pageId="2" pageNumber="3">Adult</emphasis>
</paragraph>
<paragraph id="E31636F8FFD1FFABFCBBFF497232FE90" blockId="2.[869,1493,218,307]" pageId="2" pageNumber="3">jected toward phallobase; parameres symmetric, apically acute; phallobase symmetric, with posterior margin rounded and lateral margins emarginate.</paragraph>
<paragraph id="E31636F8FFD1FFABFCBBFEC97358FE10" blockId="2.[869,1493,346,435]" pageId="2" pageNumber="3">
<emphasis id="D1DDEAEAFFD1FFABFCBBFEC974DBFED0" bold="true" box="[869,962,346,371]" pageId="2" pageNumber="3">Female:</emphasis>
Sternum VII with light organ almost as wide as sternum width, with anterior and posterior margins slightly emarginate.
</paragraph>
<paragraph id="E31636F8FFD1FFABFCBBFE497638FA30" blockId="2.[869,1494,474,723]" lastBlockId="2.[170,795,1178,1427]" pageId="2" pageNumber="3">
<emphasis id="D1DDEAEAFFD1FFABFCBBFE497317FE50" bold="true" box="[869,1038,474,499]" pageId="2" pageNumber="3">Color pattern:</emphasis>
Frons, antennae and mouthparts dark brown (
<figureCitation id="7B922A7DFFD1FFABFC62FE69731AFDB0" box="[956,1027,506,531]" captionStart="Figure 3" captionStartId="3.[94,153,2106,2131]" captionTargetBox="[96,1414,1301,2076]" captionTargetId="figure-558@3.[94,1417,1301,2076]" captionTargetPageId="3" captionText="Figure 3. Psilocladus costae sp.nov.Adult male habitus.(A) Dorsal;(B) lateral;(C) ventral.Scale bar:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985583" httpUri="https://zenodo.org/record/4985583/files/figure.png" pageId="2" pageNumber="3">Figs. 3</figureCitation>
B-C, 4A-D); pronotum orange (
<figureCitation id="7B922A7DFFD1FFABFA85FE6972BAFDB0" box="[1371,1443,506,531]" captionStart="Figure 5" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[166,1348,217,2025]" captionTargetId="figure-1@5.[151,1360,204,2044]" captionTargetPageId="5" captionText="Figure 5. Psilocladus costae sp.nov.Thorax morphology of adult male.(A-E) Prothorax:(A) dorsal;(B) ventral;(C) frontal;(D) posterior;(E)lateral;(F-I) pterothorax, (F) dorsal;(G) lateral;(H) dorsoventral.Scale bar:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985587" httpUri="https://zenodo.org/record/4985587/files/figure.png" pageId="2" pageNumber="3">Figs. 5</figureCitation>
A-E); scutellum light brown (
<figureCitation id="7B922A7DFFD1FFABFBB2FD8973A3FD90" box="[1132,1210,538,563]" captionStart="Figure 5" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[166,1348,217,2025]" captionTargetId="figure-1@5.[151,1360,204,2044]" captionTargetPageId="5" captionText="Figure 5. Psilocladus costae sp.nov.Thorax morphology of adult male.(A-E) Prothorax:(A) dorsal;(B) ventral;(C) frontal;(D) posterior;(E)lateral;(F-I) pterothorax, (F) dorsal;(G) lateral;(H) dorsoventral.Scale bar:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985587" httpUri="https://zenodo.org/record/4985587/files/figure.png" pageId="2" pageNumber="3">Fig. 5F</figureCitation>
); elytra black with lateral margins light brown (
<figureCitation id="7B922A7DFFD1FFABFB8FFDA9738EFDF0" box="[1105,1175,570,595]" captionStart="Figure 6" captionStartId="6.[172,231,2074,2099]" captionTargetBox="[176,1493,913,2040]" captionTargetId="figure-361@6.[170,1493,910,2044]" captionTargetPageId="6" captionText="Figure 6.Psilocladus costae sp.nov.Thoracic appendages of adult male. (A-C) Left legs,ventral (anterior, middle and posterior,respectively);(D-F) elytron (dorsal, lateral and ventral);(G) left wing.Scale bar:2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985589" httpUri="https://zenodo.org/record/4985589/files/figure.png" pageId="2" pageNumber="3">Figs. 6</figureCitation>
D-F); pro and mesocoxae entirely orange; trochanter orange; femur, tibia and tarsus dark brown (
<figureCitation id="7B922A7DFFD1FFABFC2DFDE97323FD30" box="[1011,1082,634,659]" captionStart="Figure 6" captionStartId="6.[172,231,2074,2099]" captionTargetBox="[176,1493,913,2040]" captionTargetId="figure-361@6.[170,1493,910,2044]" captionTargetPageId="6" captionText="Figure 6.Psilocladus costae sp.nov.Thoracic appendages of adult male. (A-C) Left legs,ventral (anterior, middle and posterior,respectively);(D-F) elytron (dorsal, lateral and ventral);(G) left wing.Scale bar:2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985589" httpUri="https://zenodo.org/record/4985589/files/figure.png" pageId="2" pageNumber="3">Figs. 6</figureCitation>
A-C); sterna II-V dark brown (
<figureCitation id="7B922A7DFFD1FFABFAA2FDE972D5FD30" box="[1404,1484,634,659]" captionStart="Figure 3" captionStartId="3.[94,153,2106,2131]" captionTargetBox="[96,1414,1301,2076]" captionTargetId="figure-558@3.[94,1417,1301,2076]" captionTargetPageId="3" captionText="Figure 3. Psilocladus costae sp.nov.Adult male habitus.(A) Dorsal;(B) lateral;(C) ventral.Scale bar:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985583" httpUri="https://zenodo.org/record/4985583/files/figure.png" pageId="2" pageNumber="3">Fig. 3C</figureCitation>
), sterna VI-VII with median region translucent (
<figureCitation id="7B922A7DFFD1FFABFA94FD09728FFD10" box="[1354,1430,666,691]" captionStart="Figure 3" captionStartId="3.[94,153,2106,2131]" captionTargetBox="[96,1414,1301,2076]" captionTargetId="figure-558@3.[94,1417,1301,2076]" captionTargetPageId="3" captionText="Figure 3. Psilocladus costae sp.nov.Adult male habitus.(A) Dorsal;(B) lateral;(C) ventral.Scale bar:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985583" httpUri="https://zenodo.org/record/4985583/files/figure.png" pageId="2" pageNumber="3">Fig. 3C</figureCitation>
), sternum VIII brown (
<figureCitation id="7B922A7DFFD1FFABFBCFFD297343FD70" box="[1041,1114,698,723]" captionStart="Figure 7" captionStartId="7.[94,153,914,939]" captionTargetBox="[97,1416,212,872]" captionTargetId="figure-21@7.[94,1417,204,884]" captionTargetPageId="7" captionText="Figure 7. Psilocladuscostae sp.nov.Terminal segments and aedeagus of male.(A) SternumVIII;(B) pygidium;(C) syntergite;(D) sternumIX;(E-G) aedeagus (dorsal, lateral and ventral).Scale bar:= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4985591" httpUri="https://zenodo.org/record/4985591/files/figure.png" pageId="2" pageNumber="3">Fig.7A</figureCitation>
) and pygidium dark brown (
<figureCitation id="7B922A7DFFD1FFABFA5CFD2972D2FD70" box="[1410,1483,698,723]" captionStart="Figure 7" captionStartId="7.[94,153,914,939]" captionTargetBox="[97,1416,212,872]" captionTargetId="figure-21@7.[94,1417,204,884]" captionTargetPageId="7" captionText="Figure 7. Psilocladuscostae sp.nov.Terminal segments and aedeagus of male.(A) SternumVIII;(B) pygidium;(C) syntergite;(D) sternumIX;(E-G) aedeagus (dorsal, lateral and ventral).Scale bar:= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4985591" httpUri="https://zenodo.org/record/4985591/files/figure.png" pageId="2" pageNumber="3">Fig.7B</figureCitation>
). 
<emphasis id="D1DDEAEAFFD1FFABFF74FB0976B9FB10" bold="true" box="[170,416,1178,1203]" pageId="2" pageNumber="3">
Diagnosis (
<figureCitation id="7B922A7DFFD1FFABFEF2FB09768BFB10" box="[300,402,1178,1203]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStart-2="Figure 5" captionStart-3="Figure 6" captionStart-4="Figure 7" captionStart-5="Figure 8" captionStart-6="Figure 9" captionStartId-0="3.[94,153,2106,2131]" captionStartId-1="4.[170,229,2074,2099]" captionStartId-2="5.[94,153,2074,2099]" captionStartId-3="6.[172,231,2074,2099]" captionStartId-4="7.[94,153,914,939]" captionStartId-5="7.[94,153,2106,2131]" captionStartId-6="8.[170,229,2074,2099]" captionTargetBox-0="[96,1414,1301,2076]" captionTargetBox-1="[175,1490,721,2029]" captionTargetBox-2="[166,1348,217,2025]" captionTargetBox-3="[176,1493,913,2040]" captionTargetBox-4="[97,1416,212,872]" captionTargetBox-5="[177,1339,1008,2071]" captionTargetBox-6="[183,1491,205,2030]" captionTargetId-0="figure-558@3.[94,1417,1301,2076]" captionTargetId-1="figure-225@4.[170,1493,721,2044]" captionTargetId-2="figure-1@5.[151,1360,204,2044]" captionTargetId-3="figure-361@6.[170,1493,910,2044]" captionTargetId-4="figure-21@7.[94,1417,204,884]" captionTargetId-5="figure-1@7.[170,1342,999,2079]" captionTargetId-6="figure-1@8.[170,1493,204,2044]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionTargetPageId-2="5" captionTargetPageId-3="6" captionTargetPageId-4="7" captionTargetPageId-5="7" captionTargetPageId-6="8" captionText-0="Figure 3. Psilocladus costae sp.nov.Adult male habitus.(A) Dorsal;(B) lateral;(C) ventral.Scale bar:= 2 mm." captionText-1="Figure 4. Psilocladus costae sp. nov. Head morphology of adult male. (A-F) Head; (A) dorsal; (B) ventral;(C) lateral; (D) frontal; (E) occipital; (F) posterior;(G) left antenna (dorsal).Scale bar:= 1 mm." captionText-2="Figure 5. Psilocladus costae sp.nov.Thorax morphology of adult male.(A-E) Prothorax:(A) dorsal;(B) ventral;(C) frontal;(D) posterior;(E)lateral;(F-I) pterothorax, (F) dorsal;(G) lateral;(H) dorsoventral.Scale bar:= 2 mm." captionText-3="Figure 6.Psilocladus costae sp.nov.Thoracic appendages of adult male. (A-C) Left legs,ventral (anterior, middle and posterior,respectively);(D-F) elytron (dorsal, lateral and ventral);(G) left wing.Scale bar:2 mm." captionText-4="Figure 7. Psilocladuscostae sp.nov.Terminal segments and aedeagus of male.(A) SternumVIII;(B) pygidium;(C) syntergite;(D) sternumIX;(E-G) aedeagus (dorsal, lateral and ventral).Scale bar:= 1 mm." captionText-5="Figure 8. Psilocladus costae sp.nov.Adult female habitus.(A) Dorsal;(B) ventral.Scale bar:= 5 mm." captionText-6="Figure 9. Psilocladus costae sp. nov. Adult female morphology. (A-E) Head; (A) dorsal; (B) ventral; (C) frontal; (D) occipital; (E) lateral; (F) left antenna (dorsal); (G) sternumVIII;(H) pygidium;(I, J) ovipositor (dorsal,ventral),arrow indicates stylus.Scale bars:A-E = 1 mm;F-I = 2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4985583" figureDoi-1="http://doi.org/10.5281/zenodo.4985585" figureDoi-2="http://doi.org/10.5281/zenodo.4985587" figureDoi-3="http://doi.org/10.5281/zenodo.4985589" figureDoi-4="http://doi.org/10.5281/zenodo.4985591" figureDoi-5="http://doi.org/10.5281/zenodo.4985593" figureDoi-6="http://doi.org/10.5281/zenodo.4985595" httpUri-0="https://zenodo.org/record/4985583/files/figure.png" httpUri-1="https://zenodo.org/record/4985585/files/figure.png" httpUri-2="https://zenodo.org/record/4985587/files/figure.png" httpUri-3="https://zenodo.org/record/4985589/files/figure.png" httpUri-4="https://zenodo.org/record/4985591/files/figure.png" httpUri-5="https://zenodo.org/record/4985593/files/figure.png" httpUri-6="https://zenodo.org/record/4985595/files/figure.png" pageId="2" pageNumber="3">Figs. 3-9</figureCitation>
):
</emphasis>
Antennae with 11 antennomeres, biflabellate, covered by dense, upright bristles, males with flabellae twice longer than antennomere body length, basally attached and decreasing in length towards antennal apex. Antennal sockets large, as wide as ⅓ frontal width. Apical maxillary and labial palpomeres securiform. Pronotum semilunar, outlined by deep punctures.
</paragraph>
<paragraph id="E31636F8FFD1FFAAFF74FA2976CBFCD0" blockId="2.[170,794,1466,1715]" lastBlockId="3.[94,719,218,883]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
<emphasis id="D1DDEAEAFFD1FFABFF74FA2977F2FA70" bold="true" box="[170,235,1466,1491]" pageId="2" pageNumber="3">Male:</emphasis>
Sternum VI with light organ as long as ¾ sternum length and as wide as ½ sternum width. Sternum VII with light organ as long as ⅔ sternum length and as wide as sternum width. Sternum VIII with posterior margin slightly emarginate; syntergite robust, slightly longer than sternum IX with anterior margin emarginate; phallus with dorsal and ventral plates; dorsal plate basally fused to parameres, slightly smaller than parameres, struts pro- 
<emphasis id="D1DDEAEAFFD0FFAAFF80FF49762EFF50" bold="true" box="[94,311,218,243]" pageId="3" pageNumber="4">
Head (
<figureCitation id="7B922A7DFFD0FFAAFF64FF497636FF50" box="[186,303,218,243]" captionStart-0="Figure 3" captionStart-1="Figure 4" captionStartId-0="3.[94,153,2106,2131]" captionStartId-1="4.[170,229,2074,2099]" captionTargetBox-0="[96,1414,1301,2076]" captionTargetBox-1="[175,1490,721,2029]" captionTargetId-0="figure-558@3.[94,1417,1301,2076]" captionTargetId-1="figure-225@4.[170,1493,721,2044]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Figure 3. Psilocladus costae sp.nov.Adult male habitus.(A) Dorsal;(B) lateral;(C) ventral.Scale bar:= 2 mm." captionText-1="Figure 4. Psilocladus costae sp. nov. Head morphology of adult male. (A-F) Head; (A) dorsal; (B) ventral;(C) lateral; (D) frontal; (E) occipital; (F) posterior;(G) left antenna (dorsal).Scale bar:= 1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4985583" figureDoi-1="http://doi.org/10.5281/zenodo.4985585" httpUri-0="https://zenodo.org/record/4985583/files/figure.png" httpUri-1="https://zenodo.org/record/4985585/files/figure.png" pageId="3" pageNumber="4">Figs. 3-4</figureCitation>
)
</emphasis>
entirely covered by pronotum (
<figureCitation id="7B922A7DFFD0FFAAFFBAFF6977B7FEB0" box="[100,174,250,275]" captionStart="Figure 3" captionStartId="3.[94,153,2106,2131]" captionTargetBox="[96,1414,1301,2076]" captionTargetId="figure-558@3.[94,1417,1301,2076]" captionTargetPageId="3" captionText="Figure 3. Psilocladus costae sp.nov.Adult male habitus.(A) Dorsal;(B) lateral;(C) ventral.Scale bar:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985583" httpUri="https://zenodo.org/record/4985583/files/figure.png" pageId="3" pageNumber="4">Figs. 3</figureCitation>
A-C); almost 2× wider than tall (
<figureCitation id="7B922A7DFFD0FFAAFDFCFF69756CFEB0" box="[546,629,250,275]" captionStart="Figure 4" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[175,1490,721,2029]" captionTargetId="figure-225@4.[170,1493,721,2044]" captionTargetPageId="4" captionText="Figure 4. Psilocladus costae sp. nov. Head morphology of adult male. (A-F) Head; (A) dorsal; (B) ventral;(C) lateral; (D) frontal; (E) occipital; (F) posterior;(G) left antenna (dorsal).Scale bar:= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4985585" httpUri="https://zenodo.org/record/4985585/files/figure.png" pageId="3" pageNumber="4">Fig. 4D</figureCitation>
); lateral margins slightly convergent posteriad (
<figureCitation id="7B922A7DFFD0FFAAFDF6FE8975DDFE90" box="[552,708,282,307]" captionStart="Figure 4" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[175,1490,721,2029]" captionTargetId="figure-225@4.[170,1493,721,2044]" captionTargetPageId="4" captionText="Figure 4. Psilocladus costae sp. nov. Head morphology of adult male. (A-F) Head; (A) dorsal; (B) ventral;(C) lateral; (D) frontal; (E) occipital; (F) posterior;(G) left antenna (dorsal).Scale bar:= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4985585" httpUri="https://zenodo.org/record/4985585/files/figure.png" pageId="3" pageNumber="4">Figs. 4B, E, F</figureCitation>
). Frons slightly emarginate (
<figureCitation id="7B922A7DFFD0FFAAFE59FEA976CEFEF0" box="[391,471,314,339]" captionStart="Figure 4" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[175,1490,721,2029]" captionTargetId="figure-225@4.[170,1493,721,2044]" captionTargetPageId="4" captionText="Figure 4. Psilocladus costae sp. nov. Head morphology of adult male. (A-F) Head; (A) dorsal; (B) ventral;(C) lateral; (D) frontal; (E) occipital; (F) posterior;(G) left antenna (dorsal).Scale bar:= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4985585" httpUri="https://zenodo.org/record/4985585/files/figure.png" pageId="3" pageNumber="4">Fig. 4D</figureCitation>
). Antennal sockets reniform, ½ as wide as frons (
<figureCitation id="7B922A7DFFD0FFAAFE50FEC97539FED0" box="[398,544,346,371]" captionStart="Figure 4" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[175,1490,721,2029]" captionTargetId="figure-225@4.[170,1493,721,2044]" captionTargetPageId="4" captionText="Figure 4. Psilocladus costae sp. nov. Head morphology of adult male. (A-F) Head; (A) dorsal; (B) ventral;(C) lateral; (D) frontal; (E) occipital; (F) posterior;(G) left antenna (dorsal).Scale bar:= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4985585" httpUri="https://zenodo.org/record/4985585/files/figure.png" pageId="3" pageNumber="4">Figs. 4A, C, D</figureCitation>
); vertex convex (
<figureCitation id="7B922A7DFFD0FFAAFFBAFEE977B6FE30" box="[100,175,378,403]" captionStart="Figure 4" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[175,1490,721,2029]" captionTargetId="figure-225@4.[170,1493,721,2044]" captionTargetPageId="4" captionText="Figure 4. Psilocladus costae sp. nov. Head morphology of adult male. (A-F) Head; (A) dorsal; (B) ventral;(C) lateral; (D) frontal; (E) occipital; (F) posterior;(G) left antenna (dorsal).Scale bar:= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4985585" httpUri="https://zenodo.org/record/4985585/files/figure.png" pageId="3" pageNumber="4">Figs. 4</figureCitation>
E-F). Antennae with 11 antennomeres, covered by dense, upright bristles, scape robust apically, pedicel almost as long as wide, antennomeres III-X biflabellate, cylindrical, decreasing in length, apical antennomere as long as scape (
<figureCitation id="7B922A7DFFD0FFAAFEFDFE69766DFDB0" box="[291,372,506,531]" captionStart="Figure 4" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[175,1490,721,2029]" captionTargetId="figure-225@4.[170,1493,721,2044]" captionTargetPageId="4" captionText="Figure 4. Psilocladus costae sp. nov. Head morphology of adult male. (A-F) Head; (A) dorsal; (B) ventral;(C) lateral; (D) frontal; (E) occipital; (F) posterior;(G) left antenna (dorsal).Scale bar:= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4985585" httpUri="https://zenodo.org/record/4985585/files/figure.png" pageId="3" pageNumber="4">Fig. 4G</figureCitation>
). Frontoclypeus slightly curved (
<figureCitation id="7B922A7DFFD0FFAAFFBAFD8977F0FD90" box="[100,233,538,563]" captionStart="Figure 4" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[175,1490,721,2029]" captionTargetId="figure-225@4.[170,1493,721,2044]" captionTargetPageId="4" captionText="Figure 4. Psilocladus costae sp. nov. Head morphology of adult male. (A-F) Head; (A) dorsal; (B) ventral;(C) lateral; (D) frontal; (E) occipital; (F) posterior;(G) left antenna (dorsal).Scale bar:= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4985585" httpUri="https://zenodo.org/record/4985585/files/figure.png" pageId="3" pageNumber="4">Figs. 4A, D</figureCitation>
). Labrum connected to frontoclypeus by membrane (
<figureCitation id="7B922A7DFFD0FFAAFF39FDA97621FDF0" box="[231,312,570,595]" captionStart="Figure 4" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[175,1490,721,2029]" captionTargetId="figure-225@4.[170,1493,721,2044]" captionTargetPageId="4" captionText="Figure 4. Psilocladus costae sp. nov. Head morphology of adult male. (A-F) Head; (A) dorsal; (B) ventral;(C) lateral; (D) frontal; (E) occipital; (F) posterior;(G) left antenna (dorsal).Scale bar:= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4985585" httpUri="https://zenodo.org/record/4985585/files/figure.png" pageId="3" pageNumber="4">Fig. 4D</figureCitation>
). Mandibles stout basally, acute apically, with a basal fringe of bristles (
<figureCitation id="7B922A7DFFD0FFAAFE37FDC97578FDD0" box="[489,609,602,627]" captionStart="Figure 4" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[175,1490,721,2029]" captionTargetId="figure-225@4.[170,1493,721,2044]" captionTargetPageId="4" captionText="Figure 4. Psilocladus costae sp. nov. Head morphology of adult male. (A-F) Head; (A) dorsal; (B) ventral;(C) lateral; (D) frontal; (E) occipital; (F) posterior;(G) left antenna (dorsal).Scale bar:= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4985585" httpUri="https://zenodo.org/record/4985585/files/figure.png" pageId="3" pageNumber="4">Figs. 4A, D</figureCitation>
). Maxillae with cardo well-sclerotized; stipites rounded.Palpi 4-segmented: I short, trapezoidal with angles acute in relation to II, II rectangular 2× longer than III, III rounded, IV securiform, 1.5× longer than II and 2× wider than II narrowing towards apex. Labium with palpi 3-segmented, palpomeres I and II rectangular, slightly enlarged towards apex;III triangular and elongated,2× longer than II. Gular sutures almost indistinct (
<figureCitation id="7B922A7DFFD0FFAAFEA4FCC976DEFCD0" box="[378,455,858,883]" captionStart="Figure 4" captionStartId="4.[170,229,2074,2099]" captionTargetBox="[175,1490,721,2029]" captionTargetId="figure-225@4.[170,1493,721,2044]" captionTargetPageId="4" captionText="Figure 4. Psilocladus costae sp. nov. Head morphology of adult male. (A-F) Head; (A) dorsal; (B) ventral;(C) lateral; (D) frontal; (E) occipital; (F) posterior;(G) left antenna (dorsal).Scale bar:= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4985585" httpUri="https://zenodo.org/record/4985585/files/figure.png" pageId="3" pageNumber="4">Fig. 4B</figureCitation>
).
</paragraph>
<caption id="B7D66670FFD1FFABFCBBF9C9737AF910" ID-DOI="http://doi.org/10.5281/zenodo.4985577" ID-Zenodo-Dep="4985577" httpUri="https://zenodo.org/record/4985577/files/figure.png" pageId="2" pageNumber="3" startId="2.[869,928,1626,1651]" targetBox="[869,1493,746,1596]" targetPageId="2">
<paragraph id="E31636F8FFD1FFABFCBBF9C9737AF910" blockId="2.[869,1494,1626,1715]" pageId="2" pageNumber="3">
Figure 1. 
<taxonomicName id="24A94D7BFFD1FFABFC65F9C97350F9D0" authority="Vaz &amp; Silveira &amp; Rosa, 2020" authorityName="Vaz &amp; Silveira &amp; Rosa" authorityYear="2020" box="[955,1097,1626,1651]" class="Insecta" family="Lampyridae" genus="Psilocladus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="costae" status="sp. nov.">
<emphasis id="D1DDEAEAFFD1FFABFC65F9C97350F9D0" bold="true" box="[955,1097,1626,1651]" italics="true" pageId="2" pageNumber="3">Psilocladus costae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAEE5791FFD1FFABFB93F9C97395F9D0" box="[1101,1164,1626,1651]" pageId="2" pageNumber="3" rank="species">sp.nov.</taxonomicNameLabel>
Habitat at Campos do Jordão,São Paulo. (A) Legal reserve along deforested area; (B, C) canopy bromeliads; (D) fallen bromeliads 
<taxonomicName id="24A94D7BFFD1FFABFC1BF9097347F910" authorityName="Wawra" authorityYear="1862" box="[965,1118,1690,1715]" class="Liliopsida" family="Bromeliaceae" genus="Vriesea" kingdom="Plantae" order="Poales" pageId="2" pageNumber="3" phylum="Tracheophyta" rank="species" species="bituminosa">Vriesea bituminosa</taxonomicName>
.
</paragraph>
</caption>
<caption id="B7D66670FFD1FFABFF74F7897677F7F0" ID-DOI="http://doi.org/10.5281/zenodo.4985581" ID-Zenodo-Dep="4985581" httpUri="https://zenodo.org/record/4985581/files/figure.png" pageId="2" pageNumber="3" startId="2.[170,229,2074,2099]" targetBox="[170,1493,1742,2044]" targetPageId="2">
<paragraph id="E31636F8FFD1FFABFF74F7897677F7F0" blockId="2.[170,1494,2074,2131]" pageId="2" pageNumber="3">
Figure 2. 
<taxonomicName id="24A94D7BFFD1FFABFEDEF7897697F790" authority="Vaz &amp; Silveira &amp; Rosa, 2020" authorityName="Vaz &amp; Silveira &amp; Rosa" authorityYear="2020" box="[256,398,2074,2099]" class="Insecta" family="Lampyridae" genus="Psilocladus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="costae" status="sp. nov.">
<emphasis id="D1DDEAEAFFD1FFABFEDEF7897697F790" bold="true" box="[256,398,2074,2099]" italics="true" pageId="2" pageNumber="3">Psilocladus costae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAEE5791FFD1FFABFE4DF78976C8F790" box="[403,465,2074,2099]" pageId="2" pageNumber="3" rank="species">sp.nov.</taxonomicNameLabel>
Live larvae.(A, B) Larva on bromeliad leaf blades (dorsal);(C) larva on a detached innermost leaf sheath of bromeliad rosette. Length of larva:20 mm.
</paragraph>
</caption>
<paragraph id="E31636F8FFD0FFAAFF80FC09738CFC70" blockId="3.[94,719,922,1267]" lastBlockId="3.[794,1418,218,979]" pageId="3" pageNumber="4">
<emphasis id="D1DDEAEAFFD0FFAAFF80FC097620FC10" bold="true" box="[94,313,922,947]" pageId="3" pageNumber="4">
Thorax (
<figureCitation id="7B922A7DFFD0FFAAFF1CFC097632FC10" box="[194,299,922,947]" captionStart-0="Figure 5" captionStart-1="Figure 6" captionStartId-0="5.[94,153,2074,2099]" captionStartId-1="6.[172,231,2074,2099]" captionTargetBox-0="[166,1348,217,2025]" captionTargetBox-1="[176,1493,913,2040]" captionTargetId-0="figure-1@5.[151,1360,204,2044]" captionTargetId-1="figure-361@6.[170,1493,910,2044]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Figure 5. Psilocladus costae sp.nov.Thorax morphology of adult male.(A-E) Prothorax:(A) dorsal;(B) ventral;(C) frontal;(D) posterior;(E)lateral;(F-I) pterothorax, (F) dorsal;(G) lateral;(H) dorsoventral.Scale bar:= 2 mm." captionText-1="Figure 6.Psilocladus costae sp.nov.Thoracic appendages of adult male. (A-C) Left legs,ventral (anterior, middle and posterior,respectively);(D-F) elytron (dorsal, lateral and ventral);(G) left wing.Scale bar:2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4985587" figureDoi-1="http://doi.org/10.5281/zenodo.4985589" httpUri-0="https://zenodo.org/record/4985587/files/figure.png" httpUri-1="https://zenodo.org/record/4985589/files/figure.png" pageId="3" pageNumber="4">Figs. 5-6</figureCitation>
).
</emphasis>
Pronotum semilunar, anterior margin rounded, lateral margins divergent, posterior margin with central region slightly projected posteriad and medially emarginate, posterior angles rounded, regularly punctured, punctures small and bristled (
<figureCitation id="7B922A7DFFD0FFAAFD90FB89758FFB90" box="[590,662,1050,1075]" captionStart="Figure 5" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[166,1348,217,2025]" captionTargetId="figure-1@5.[151,1360,204,2044]" captionTargetPageId="5" captionText="Figure 5. Psilocladus costae sp.nov.Thorax morphology of adult male.(A-E) Prothorax:(A) dorsal;(B) ventral;(C) frontal;(D) posterior;(E)lateral;(F-I) pterothorax, (F) dorsal;(G) lateral;(H) dorsoventral.Scale bar:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985587" httpUri="https://zenodo.org/record/4985587/files/figure.png" pageId="3" pageNumber="4">Figs. 5</figureCitation>
A-D). Hypomeron 2× longer than tall (
<figureCitation id="7B922A7DFFD0FFAAFE3AFBA9752FFBF0" box="[484,566,1082,1107]" captionStart="Figure 5" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[166,1348,217,2025]" captionTargetId="figure-1@5.[151,1360,204,2044]" captionTargetPageId="5" captionText="Figure 5. Psilocladus costae sp.nov.Thorax morphology of adult male.(A-E) Prothorax:(A) dorsal;(B) ventral;(C) frontal;(D) posterior;(E)lateral;(F-I) pterothorax, (F) dorsal;(G) lateral;(H) dorsoventral.Scale bar:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985587" httpUri="https://zenodo.org/record/4985587/files/figure.png" pageId="3" pageNumber="4">Fig. 5E</figureCitation>
). Prosternum slightly constricted medially (
<figureCitation id="7B922A7DFFD0FFAAFE7BFBC976EBFBD0" box="[421,498,1114,1139]" captionStart="Figure 5" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[166,1348,217,2025]" captionTargetId="figure-1@5.[151,1360,204,2044]" captionTargetPageId="5" captionText="Figure 5. Psilocladus costae sp.nov.Thorax morphology of adult male.(A-E) Prothorax:(A) dorsal;(B) ventral;(C) frontal;(D) posterior;(E)lateral;(F-I) pterothorax, (F) dorsal;(G) lateral;(H) dorsoventral.Scale bar:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985587" httpUri="https://zenodo.org/record/4985587/files/figure.png" pageId="3" pageNumber="4">Fig. 5B</figureCitation>
). Elytra ellipsoid, 4× longer than wide at greatest width, secondary pubescence absent (
<figureCitation id="7B922A7DFFD0FFAAFED7FB09764CFB10" box="[265,341,1178,1203]" captionStart="Figure 6" captionStartId="6.[172,231,2074,2099]" captionTargetBox="[176,1493,913,2040]" captionTargetId="figure-361@6.[170,1493,910,2044]" captionTargetPageId="6" captionText="Figure 6.Psilocladus costae sp.nov.Thoracic appendages of adult male. (A-C) Left legs,ventral (anterior, middle and posterior,respectively);(D-F) elytron (dorsal, lateral and ventral);(G) left wing.Scale bar:2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985589" httpUri="https://zenodo.org/record/4985589/files/figure.png" pageId="3" pageNumber="4">Figs. 6</figureCitation>
D-F). Hind wing well-developed, posterior margin sinuose, 2× longer than wide, r4 almost 3× longer than r3, radial cell 3× wider than long, almost reaching anterior margin; CuA2 cross vein absent; MP3 + 4 Y-shaped and basally connected with Cu; RP + MP1 + 2 of three fourths r4 length, almost reaching distal margin; AA3 parallel with CuP + AA3; CuA truncate in the origin (
<figureCitation id="7B922A7DFFD0FFAAFC70FEC974E7FED0" box="[942,1022,346,371]" captionStart="Figure 6" captionStartId="6.[172,231,2074,2099]" captionTargetBox="[176,1493,913,2040]" captionTargetId="figure-361@6.[170,1493,910,2044]" captionTargetPageId="6" captionText="Figure 6.Psilocladus costae sp.nov.Thoracic appendages of adult male. (A-C) Left legs,ventral (anterior, middle and posterior,respectively);(D-F) elytron (dorsal, lateral and ventral);(G) left wing.Scale bar:2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985589" httpUri="https://zenodo.org/record/4985589/files/figure.png" pageId="3" pageNumber="4">Fig. 6G</figureCitation>
). Alinotum slightly wider than long, lateral margins slightly convergent posteriad, posterior margin straight; prescutum extending slightly less than half metascutum length; rounded area of scutum weakly sclerotized, scutum-prescutal plates sclerotized, extending ridges almost up to posterior margin; metascutellum glabrous. Mesosternum weakly sclerotized, acute medially, attached to metasternum by a suture, almost as wide as mesosternum. Mesepimeron attached to metasternum by membrane. Mesosternum/mesanepisternum suture inconspicuous. Mesanepisternum/mesepimeron suture conspicuous. Metasternum oblique and strongly depressed by mesocoxae, anterior medial keel prominent up to anterior third, discrimen distinct, lateral margins divergent posteriad up to lateral-most part of metacoxae, then convergent posteriad, posterior margin bisinuose (
<figureCitation id="7B922A7DFFD0FFAAFC4AFCC974F9FCD0" box="[916,992,858,883]" captionStart="Figure 5" captionStartId="5.[94,153,2074,2099]" captionTargetBox="[166,1348,217,2025]" captionTargetId="figure-1@5.[151,1360,204,2044]" captionTargetPageId="5" captionText="Figure 5. Psilocladus costae sp.nov.Thorax morphology of adult male.(A-E) Prothorax:(A) dorsal;(B) ventral;(C) frontal;(D) posterior;(E)lateral;(F-I) pterothorax, (F) dorsal;(G) lateral;(H) dorsoventral.Scale bar:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985587" httpUri="https://zenodo.org/record/4985587/files/figure.png" pageId="3" pageNumber="4">Figs. 5</figureCitation>
F-I). Femur slightly shorter than tibia (
<figureCitation id="7B922A7DFFD0FFAAFCFEFCE9747FFC30" box="[800,870,890,915]" captionStart="Figure 6" captionStartId="6.[172,231,2074,2099]" captionTargetBox="[176,1493,913,2040]" captionTargetId="figure-361@6.[170,1493,910,2044]" captionTargetPageId="6" captionText="Figure 6.Psilocladus costae sp.nov.Thoracic appendages of adult male. (A-C) Left legs,ventral (anterior, middle and posterior,respectively);(D-F) elytron (dorsal, lateral and ventral);(G) left wing.Scale bar:2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985589" httpUri="https://zenodo.org/record/4985589/files/figure.png" pageId="3" pageNumber="4">Figs. 6</figureCitation>
A-C). Tarsomere I 2× longer than II, II slightly longer than III, III subequal in length to IV, IV bilobed, lobes reaching half V length (
<figureCitation id="7B922A7DFFD0FFAAFBC6FC297347FC70" box="[1048,1118,954,979]" captionStart="Figure 6" captionStartId="6.[172,231,2074,2099]" captionTargetBox="[176,1493,913,2040]" captionTargetId="figure-361@6.[170,1493,910,2044]" captionTargetPageId="6" captionText="Figure 6.Psilocladus costae sp.nov.Thoracic appendages of adult male. (A-C) Left legs,ventral (anterior, middle and posterior,respectively);(D-F) elytron (dorsal, lateral and ventral);(G) left wing.Scale bar:2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985589" httpUri="https://zenodo.org/record/4985589/files/figure.png" pageId="3" pageNumber="4">Figs. 6</figureCitation>
A-C).
</paragraph>
<paragraph id="E31636F8FFD0FFADFCC4FC6976F0FD90" blockId="3.[794,1418,1018,1267]" lastBlockId="4.[170,795,218,563]" lastPageId="4" lastPageNumber="5" pageId="3" pageNumber="4">
<emphasis id="D1DDEAEAFFD0FFAAFCC4FC6974F6FBB0" bold="true" box="[794,1007,1018,1043]" pageId="3" pageNumber="4">
Abdomen (
<figureCitation id="7B922A7DFFD0FFAAFC40FC6974F8FBB0" box="[926,993,1018,1043]" captionStart="Figure 7" captionStartId="7.[94,153,914,939]" captionTargetBox="[97,1416,212,872]" captionTargetId="figure-21@7.[94,1417,204,884]" captionTargetPageId="7" captionText="Figure 7. Psilocladuscostae sp.nov.Terminal segments and aedeagus of male.(A) SternumVIII;(B) pygidium;(C) syntergite;(D) sternumIX;(E-G) aedeagus (dorsal, lateral and ventral).Scale bar:= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4985591" httpUri="https://zenodo.org/record/4985591/files/figure.png" pageId="3" pageNumber="4">Fig. 7</figureCitation>
):
</emphasis>
Seven visible sterna (II-VIII) (
<figureCitation id="7B922A7DFFD0FFAAFAF1FC697299FBB0" box="[1327,1408,1018,1043]" captionStart="Figure 3" captionStartId="3.[94,153,2106,2131]" captionTargetBox="[96,1414,1301,2076]" captionTargetId="figure-558@3.[94,1417,1301,2076]" captionTargetPageId="3" captionText="Figure 3. Psilocladus costae sp.nov.Adult male habitus.(A) Dorsal;(B) lateral;(C) ventral.Scale bar:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985583" httpUri="https://zenodo.org/record/4985583/files/figure.png" pageId="3" pageNumber="4">Fig. 3C</figureCitation>
), lateral margins convergent posteriad,spiracles dorsal (i.e. dorsally reflexed margins of sternite) (
<figureCitation id="7B922A7DFFD0FFAAFB17FBA97203FBF0" box="[1225,1306,1082,1107]" captionStart="Figure 3" captionStartId="3.[94,153,2106,2131]" captionTargetBox="[96,1414,1301,2076]" captionTargetId="figure-558@3.[94,1417,1301,2076]" captionTargetPageId="3" captionText="Figure 3. Psilocladus costae sp.nov.Adult male habitus.(A) Dorsal;(B) lateral;(C) ventral.Scale bar:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985583" httpUri="https://zenodo.org/record/4985583/files/figure.png" pageId="3" pageNumber="4">Fig. 3B</figureCitation>
); sternum VIII 3× wider than long, posterior margin emarginate (
<figureCitation id="7B922A7DFFD0FFAAFCFEFBE97477FB30" box="[800,878,1146,1171]" captionStart="Figure 7" captionStartId="7.[94,153,914,939]" captionTargetBox="[97,1416,212,872]" captionTargetId="figure-21@7.[94,1417,204,884]" captionTargetPageId="7" captionText="Figure 7. Psilocladuscostae sp.nov.Terminal segments and aedeagus of male.(A) SternumVIII;(B) pygidium;(C) syntergite;(D) sternumIX;(E-G) aedeagus (dorsal, lateral and ventral).Scale bar:= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4985591" httpUri="https://zenodo.org/record/4985591/files/figure.png" pageId="3" pageNumber="4">Fig. 7A</figureCitation>
); pygidium sclerotized, anterior margin indented, lateral margins rounded and posterior margin with two parassagital sinuosities, lateral angles poorly developed (
<figureCitation id="7B922A7DFFD0FFAAFCFEFB497469FB50" box="[800,880,1242,1267]" captionStart="Figure 7" captionStartId="7.[94,153,914,939]" captionTargetBox="[97,1416,212,872]" captionTargetId="figure-21@7.[94,1417,204,884]" captionTargetPageId="7" captionText="Figure 7. Psilocladuscostae sp.nov.Terminal segments and aedeagus of male.(A) SternumVIII;(B) pygidium;(C) syntergite;(D) sternumIX;(E-G) aedeagus (dorsal, lateral and ventral).Scale bar:= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4985591" httpUri="https://zenodo.org/record/4985591/files/figure.png" pageId="3" pageNumber="4">Fig. 7B</figureCitation>
); syntergite subtriangular, subdivided and with anterior margin emarginate (
<figureCitation id="7B922A7DFFD7FFADFE26FF497552FF50" box="[504,587,218,243]" captionStart="Figure 7" captionStartId="7.[94,153,914,939]" captionTargetBox="[97,1416,212,872]" captionTargetId="figure-21@7.[94,1417,204,884]" captionTargetPageId="7" captionText="Figure 7. Psilocladuscostae sp.nov.Terminal segments and aedeagus of male.(A) SternumVIII;(B) pygidium;(C) syntergite;(D) sternumIX;(E-G) aedeagus (dorsal, lateral and ventral).Scale bar:= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4985591" httpUri="https://zenodo.org/record/4985591/files/figure.png" pageId="4" pageNumber="5">Fig. 7C</figureCitation>
); sternum IX with posterior margin convergent posteriad, posterior margin rounded, with a spine in central region of the anterior margin (
<figureCitation id="7B922A7DFFD7FFADFEECFEA9769AFEF0" box="[306,387,314,339]" captionStart="Figure 7" captionStartId="7.[94,153,914,939]" captionTargetBox="[97,1416,212,872]" captionTargetId="figure-21@7.[94,1417,204,884]" captionTargetPageId="7" captionText="Figure 7. Psilocladuscostae sp.nov.Terminal segments and aedeagus of male.(A) SternumVIII;(B) pygidium;(C) syntergite;(D) sternumIX;(E-G) aedeagus (dorsal, lateral and ventral).Scale bar:= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4985591" httpUri="https://zenodo.org/record/4985591/files/figure.png" pageId="4" pageNumber="5">Fig. 7D</figureCitation>
). 
<emphasis id="D1DDEAEAFFD7FFADFE4DFEA97582FEF0" bold="true" box="[403,667,314,339]" pageId="4" pageNumber="5">
Aedeagus (
<figureCitation id="7B922A7DFFD7FFADFDCBFEA97578FEF0" box="[533,609,314,339]" captionStart="Figure 7" captionStartId="7.[94,153,914,939]" captionTargetBox="[97,1416,212,872]" captionTargetId="figure-21@7.[94,1417,204,884]" captionTargetPageId="7" captionText="Figure 7. Psilocladuscostae sp.nov.Terminal segments and aedeagus of male.(A) SternumVIII;(B) pygidium;(C) syntergite;(D) sternumIX;(E-G) aedeagus (dorsal, lateral and ventral).Scale bar:= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4985591" httpUri="https://zenodo.org/record/4985591/files/figure.png" pageId="4" pageNumber="5">Figs. 7</figureCitation>
E-G):
</emphasis>
Phallobase symmetrical, with a distinct longitudinal line (
<figureCitation id="7B922A7DFFD7FFADFD40FEC97416FED0" box="[670,783,346,371]" captionStart="Figure 7" captionStartId="7.[94,153,914,939]" captionTargetBox="[97,1416,212,872]" captionTargetId="figure-21@7.[94,1417,204,884]" captionTargetPageId="7" captionText="Figure 7. Psilocladuscostae sp.nov.Terminal segments and aedeagus of male.(A) SternumVIII;(B) pygidium;(C) syntergite;(D) sternumIX;(E-G) aedeagus (dorsal, lateral and ventral).Scale bar:= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4985591" httpUri="https://zenodo.org/record/4985591/files/figure.png" pageId="4" pageNumber="5">Figs. 7E, G</figureCitation>
); parameres symmetrical, connected basally by a thin bridge, apically acute, apical ½ membranous, internal margin sinuose (
<figureCitation id="7B922A7DFFD7FFADFEB6FE2976ABFE70" box="[360,434,442,467]" captionStart="Figure 7" captionStartId="7.[94,153,914,939]" captionTargetBox="[97,1416,212,872]" captionTargetId="figure-21@7.[94,1417,204,884]" captionTargetPageId="7" captionText="Figure 7. Psilocladuscostae sp.nov.Terminal segments and aedeagus of male.(A) SternumVIII;(B) pygidium;(C) syntergite;(D) sternumIX;(E-G) aedeagus (dorsal, lateral and ventral).Scale bar:= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4985591" httpUri="https://zenodo.org/record/4985591/files/figure.png" pageId="4" pageNumber="5">Figs. 7</figureCitation>
E-G); phallus with struts almost straight, extending basally slightly beyond parameres; ventral plate coriaceous, dorsal plate flat, apically acute, strongly sclerotized (
<figureCitation id="7B922A7DFFD7FFADFE51FD8976C7FD90" box="[399,478,538,563]" captionStart="Figure 7" captionStartId="7.[94,153,914,939]" captionTargetBox="[97,1416,212,872]" captionTargetId="figure-21@7.[94,1417,204,884]" captionTargetPageId="7" captionText="Figure 7. Psilocladuscostae sp.nov.Terminal segments and aedeagus of male.(A) SternumVIII;(B) pygidium;(C) syntergite;(D) sternumIX;(E-G) aedeagus (dorsal, lateral and ventral).Scale bar:= 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4985591" httpUri="https://zenodo.org/record/4985591/files/figure.png" pageId="4" pageNumber="5">Fig. 7G</figureCitation>
).
</paragraph>
<caption id="B7D66670FFD0FFAAFF80F7A974C0F7F0" ID-DOI="http://doi.org/10.5281/zenodo.4985583" ID-Zenodo-Dep="4985583" box="[94,985,2106,2131]" httpUri="https://zenodo.org/record/4985583/files/figure.png" pageId="3" pageNumber="4" startId="3.[94,153,2106,2131]" targetBox="[96,1414,1301,2076]" targetPageId="3">
<paragraph id="E31636F8FFD0FFAAFF80F7A974C0F7F0" blockId="3.[94,985,2106,2131]" box="[94,985,2106,2131]" pageId="3" pageNumber="4">
Figure 3. 
<taxonomicName id="24A94D7BFFD0FFAAFF6AF7A9765BF7F0" authority="Vaz &amp; Silveira &amp; Rosa, 2020" authorityName="Vaz &amp; Silveira &amp; Rosa" authorityYear="2020" box="[180,322,2106,2131]" class="Insecta" family="Lampyridae" genus="Psilocladus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="costae" status="sp. nov.">
<emphasis id="D1DDEAEAFFD0FFAAFF6AF7A9765BF7F0" bold="true" box="[180,322,2106,2131]" italics="true" pageId="3" pageNumber="4">Psilocladus costae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAEE5791FFD0FFAAFE98F7A9769DF7F0" box="[326,388,2106,2131]" pageId="3" pageNumber="4" rank="species">sp.nov.</taxonomicNameLabel>
Adult male habitus.(A) Dorsal;(B) lateral;(C) ventral.Scale bar:= 2 mm.
</paragraph>
</caption>
<paragraph id="E31636F8FFD7FFADFF74FDC9738FFDB0" blockId="4.[170,794,602,691]" lastBlockId="4.[869,1493,218,531]" pageId="4" pageNumber="5">
<emphasis id="D1DDEAEAFFD7FFADFF74FDC97698FDD0" bold="true" box="[170,385,602,627]" pageId="4" pageNumber="5">
Female (
<figureCitation id="7B922A7DFFD7FFADFED2FDC97640FDD0" box="[268,345,602,627]" captionStart="Figure 8" captionStartId="7.[94,153,2106,2131]" captionTargetBox="[177,1339,1008,2071]" captionTargetId="figure-1@7.[170,1342,999,2079]" captionTargetPageId="7" captionText="Figure 8. Psilocladus costae sp.nov.Adult female habitus.(A) Dorsal;(B) ventral.Scale bar:= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4985593" httpUri="https://zenodo.org/record/4985593/files/figure.png" pageId="4" pageNumber="5">Figs. 8</figureCitation>
, 
<figureCitation id="7B922A7DFFD7FFADFEBDFDC9766AFDD0" box="[355,371,602,627]" captionStart="Figure 9" captionStartId="8.[170,229,2074,2099]" captionTargetBox="[183,1491,205,2030]" captionTargetId="figure-1@8.[170,1493,204,2044]" captionTargetPageId="8" captionText="Figure 9. Psilocladus costae sp. nov. Adult female morphology. (A-E) Head; (A) dorsal; (B) ventral; (C) frontal; (D) occipital; (E) lateral; (F) left antenna (dorsal); (G) sternumVIII;(H) pygidium;(I, J) ovipositor (dorsal,ventral),arrow indicates stylus.Scale bars:A-E = 1 mm;F-I = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985595" httpUri="https://zenodo.org/record/4985595/files/figure.png" pageId="4" pageNumber="5">9</figureCitation>
):
</emphasis>
Eye almost ⅔ as wide as frons in frontal view (
<figureCitation id="7B922A7DFFD7FFADFECBFDE97669FD30" box="[277,368,634,659]" captionStart="Figure 8" captionStartId="7.[94,153,2106,2131]" captionTargetBox="[177,1339,1008,2071]" captionTargetId="figure-1@7.[170,1342,999,2079]" captionTargetPageId="7" captionText="Figure 8. Psilocladus costae sp.nov.Adult female habitus.(A) Dorsal;(B) ventral.Scale bar:= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4985593" httpUri="https://zenodo.org/record/4985593/files/figure.png" pageId="4" pageNumber="5">Figs. 8B</figureCitation>
, 
<figureCitation id="7B922A7DFFD7FFADFEA3FDE97693FD30" box="[381,394,634,659]" captionStart="Figure 9" captionStartId="8.[170,229,2074,2099]" captionTargetBox="[183,1491,205,2030]" captionTargetId="figure-1@8.[170,1493,204,2044]" captionTargetPageId="8" captionText="Figure 9. Psilocladus costae sp. nov. Adult female morphology. (A-E) Head; (A) dorsal; (B) ventral; (C) frontal; (D) occipital; (E) lateral; (F) left antenna (dorsal); (G) sternumVIII;(H) pygidium;(I, J) ovipositor (dorsal,ventral),arrow indicates stylus.Scale bars:A-E = 1 mm;F-I = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985595" httpUri="https://zenodo.org/record/4985595/files/figure.png" pageId="4" pageNumber="5">9</figureCitation>
A-E); antennae with short flabellae which are slightly longer than core antennomere (
<figureCitation id="7B922A7DFFD7FFADFD18FD097416FD10" box="[710,783,666,691]" captionStart="Figure 9" captionStartId="8.[170,229,2074,2099]" captionTargetBox="[183,1491,205,2030]" captionTargetId="figure-1@8.[170,1493,204,2044]" captionTargetPageId="8" captionText="Figure 9. Psilocladus costae sp. nov. Adult female morphology. (A-E) Head; (A) dorsal; (B) ventral; (C) frontal; (D) occipital; (E) lateral; (F) left antenna (dorsal); (G) sternumVIII;(H) pygidium;(I, J) ovipositor (dorsal,ventral),arrow indicates stylus.Scale bars:A-E = 1 mm;F-I = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985595" httpUri="https://zenodo.org/record/4985595/files/figure.png" pageId="4" pageNumber="5">Fig.9F</figureCitation>
); pronotum 2× wider than long (
<figureCitation id="7B922A7DFFD7FFADFB1DFF49720AFF50" box="[1219,1299,218,243]" captionStart="Figure 8" captionStartId="7.[94,153,2106,2131]" captionTargetBox="[177,1339,1008,2071]" captionTargetId="figure-1@7.[170,1342,999,2079]" captionTargetPageId="7" captionText="Figure 8. Psilocladus costae sp.nov.Adult female habitus.(A) Dorsal;(B) ventral.Scale bar:= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4985593" httpUri="https://zenodo.org/record/4985593/files/figure.png" pageId="4" pageNumber="5">Fig. 8A</figureCitation>
); sternum VI with subrectangular light organ (
<figureCitation id="7B922A7DFFD7FFADFB79FF6973EFFEB0" box="[1191,1270,250,275]" captionStart="Figure 8" captionStartId="7.[94,153,2106,2131]" captionTargetBox="[177,1339,1008,2071]" captionTargetId="figure-1@7.[170,1342,999,2079]" captionTargetPageId="7" captionText="Figure 8. Psilocladus costae sp.nov.Adult female habitus.(A) Dorsal;(B) ventral.Scale bar:= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4985593" httpUri="https://zenodo.org/record/4985593/files/figure.png" pageId="4" pageNumber="5">Fig. 8B</figureCitation>
) almost as wide as sternum width; sternum VII lacking light organ (
<figureCitation id="7B922A7DFFD7FFADFAA3FE8972D2FE90" box="[1405,1483,282,307]" captionStart="Figure 8" captionStartId="7.[94,153,2106,2131]" captionTargetBox="[177,1339,1008,2071]" captionTargetId="figure-1@7.[170,1342,999,2079]" captionTargetPageId="7" captionText="Figure 8. Psilocladus costae sp.nov.Adult female habitus.(A) Dorsal;(B) ventral.Scale bar:= 5 mm." figureDoi="http://doi.org/10.5281/zenodo.4985593" httpUri="https://zenodo.org/record/4985593/files/figure.png" pageId="4" pageNumber="5">Fig. 8B</figureCitation>
); sternum VIII with lateral margins converging posteriad and posterior margin with central region emarginate (
<figureCitation id="7B922A7DFFD7FFADFCB2FEE974A4FE30" box="[876,957,378,403]" captionStart="Figure 9" captionStartId="8.[170,229,2074,2099]" captionTargetBox="[183,1491,205,2030]" captionTargetId="figure-1@8.[170,1493,204,2044]" captionTargetPageId="8" captionText="Figure 9. Psilocladus costae sp. nov. Adult female morphology. (A-E) Head; (A) dorsal; (B) ventral; (C) frontal; (D) occipital; (E) lateral; (F) left antenna (dorsal); (G) sternumVIII;(H) pygidium;(I, J) ovipositor (dorsal,ventral),arrow indicates stylus.Scale bars:A-E = 1 mm;F-I = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985595" httpUri="https://zenodo.org/record/4985595/files/figure.png" pageId="4" pageNumber="5">Fig. 9G</figureCitation>
); pygidium with lateral margins convergent and posterior margin rounded (
<figureCitation id="7B922A7DFFD7FFADFB45FE0973F4FE10" box="[1179,1261,410,435]" captionStart="Figure 9" captionStartId="8.[170,229,2074,2099]" captionTargetBox="[183,1491,205,2030]" captionTargetId="figure-1@8.[170,1493,204,2044]" captionTargetPageId="8" captionText="Figure 9. Psilocladus costae sp. nov. Adult female morphology. (A-E) Head; (A) dorsal; (B) ventral; (C) frontal; (D) occipital; (E) lateral; (F) left antenna (dorsal); (G) sternumVIII;(H) pygidium;(I, J) ovipositor (dorsal,ventral),arrow indicates stylus.Scale bars:A-E = 1 mm;F-I = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985595" httpUri="https://zenodo.org/record/4985595/files/figure.png" pageId="4" pageNumber="5">Fig. 9H</figureCitation>
). 
<emphasis id="D1DDEAEAFFD7FFADFADFFE09726DFE10" bold="true" box="[1281,1396,410,435]" pageId="4" pageNumber="5">Genitalia:</emphasis>
ovipositor with baculus sclerotized, coxites membranous, styli (pointed) apically sclerotized and bristled outwards at the lateral edges (
<figureCitation id="7B922A7DFFD7FFADFBF7FE697369FDB0" box="[1065,1136,506,531]" captionStart="Figure 9" captionStartId="8.[170,229,2074,2099]" captionTargetBox="[183,1491,205,2030]" captionTargetId="figure-1@8.[170,1493,204,2044]" captionTargetPageId="8" captionText="Figure 9. Psilocladus costae sp. nov. Adult female morphology. (A-E) Head; (A) dorsal; (B) ventral; (C) frontal; (D) occipital; (E) lateral; (F) left antenna (dorsal); (G) sternumVIII;(H) pygidium;(I, J) ovipositor (dorsal,ventral),arrow indicates stylus.Scale bars:A-E = 1 mm;F-I = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985595" httpUri="https://zenodo.org/record/4985595/files/figure.png" pageId="4" pageNumber="5">Figs. 9</figureCitation>
I-J).
</paragraph>
<paragraph id="E31636F8FFD7FFADFCBBFDA97335FDF0" blockId="4.[869,1068,570,595]" box="[869,1068,570,595]" pageId="4" pageNumber="5">
<emphasis id="D1DDEAEAFFD7FFADFCBBFDA97335FDF0" bold="true" box="[869,1068,570,595]" pageId="4" pageNumber="5">Immature stages</emphasis>
</paragraph>
<paragraph id="E31636F8FFD7FFADFCBBFDE972CCFD10" blockId="4.[869,1493,634,691]" pageId="4" pageNumber="5">
<emphasis id="D1DDEAEAFFD7FFADFCBBFDE97298FD30" bold="true" box="[869,1409,634,659]" pageId="4" pageNumber="5">
Mature larva (
<figureCitation id="7B922A7DFFD7FFADFBECFDE973D3FD30" box="[1074,1226,634,659]" captionStart-0="Figure 10" captionStart-1="Figure 11" captionStart-2="Figure 12" captionStartId-0="9.[94,153,2106,2131]" captionStartId-1="10.[170,229,2074,2099]" captionStartId-2="11.[94,153,2074,2099]" captionTargetBox-0="[100,1413,1083,2074]" captionTargetBox-1="[176,1489,864,2043]" captionTargetBox-2="[258,1262,1269,2034]" captionTargetId-0="figure-434@9.[94,1417,1075,2077]" captionTargetId-1="figure-330@10.[170,1493,854,2045]" captionTargetId-2="figure-570@11.[246,1266,1269,2044]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionTargetPageId-2="11" captionText-0="Figure 10. Psilocladus costae sp.nov.Mature larval habitus.(A) Dorsal;(B) lateral;(C) ventral.Scale bar:= 2 mm." captionText-1="Figure 11. Psilocladus costae sp. nov. Larval morphology. (A-C) Head (dorsal, lateral, ventral); (D) right anterior, middle and posterior legs (from top to bottom, ventral);(E) left laterotergitesVI-VII;(F) abdominal tergaVI-IX.Scale bars:A-C,E, F = 1 mm;D = 2 mm." captionText-2="Figure 12.Psilocladus costae sp.nov.Larval headmorphology.(A) Head,dorsal;(B)maxillolabial complex,ventral;(C) epipharynx;(D) hypopharynx;(E) leftantenna (ventral,dorsal);(F) mandible (ventral,dorsal).Scale bars:A,B = 1 mm;C,D = 0.25 mm;E,F = 0.5 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4985597" figureDoi-1="http://doi.org/10.5281/zenodo.4985599" figureDoi-2="http://doi.org/10.5281/zenodo.4985603" httpUri-0="https://zenodo.org/record/4985597/files/figure.png" httpUri-1="https://zenodo.org/record/4985599/files/figure.png" httpUri-2="https://zenodo.org/record/4985603/files/figure.png" pageId="4" pageNumber="5">Figs. 10-12</figureCitation>
). Description:
</emphasis>
Body (
<figureCitation id="7B922A7DFFD7FFADFCB5FD0974A6FD10" box="[875,959,666,691]" captionStart="Figure 10" captionStartId="9.[94,153,2106,2131]" captionTargetBox="[100,1413,1083,2074]" captionTargetId="figure-434@9.[94,1417,1075,2077]" captionTargetPageId="9" captionText="Figure 10. Psilocladus costae sp.nov.Mature larval habitus.(A) Dorsal;(B) lateral;(C) ventral.Scale bar:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985597" httpUri="https://zenodo.org/record/4985597/files/figure.png" pageId="4" pageNumber="5">Figs. 10</figureCitation>
A-C) 
<quantity id="24519B1DFFD7FFADFC26FD097373FD10" box="[1016,1130,666,691]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="4" pageNumber="5" unit="mm" value="18.5" valueMax="22.0" valueMin="15.0">15-22 mm</quantity>
long, oblong (pronotum semio-
</paragraph>
<caption id="B7D66670FFD7FFADFF74F78976CEF7F0" ID-DOI="http://doi.org/10.5281/zenodo.4985585" ID-Zenodo-Dep="4985585" httpUri="https://zenodo.org/record/4985585/files/figure.png" pageId="4" pageNumber="5" startId="4.[170,229,2074,2099]" targetBox="[175,1490,721,2029]" targetPageId="4">
<paragraph id="E31636F8FFD7FFADFF74F78976CEF7F0" blockId="4.[170,1493,2074,2131]" pageId="4" pageNumber="5">
Figure 4. 
<taxonomicName id="24A94D7BFFD7FFADFEDFF7897689F790" authority="Vaz &amp; Silveira &amp; Rosa, 2020" authorityName="Vaz &amp; Silveira &amp; Rosa" authorityYear="2020" box="[257,400,2074,2099]" class="Insecta" family="Lampyridae" genus="Psilocladus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="costae" status="sp. nov.">
<emphasis id="D1DDEAEAFFD7FFADFEDFF7897689F790" bold="true" box="[257,400,2074,2099]" italics="true" pageId="4" pageNumber="5">Psilocladus costae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAEE5791FFD7FFADFE4BF78976CDF790" box="[405,468,2074,2099]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
Head morphology of adult male. (A-F) Head; (A) dorsal; (B) ventral;(C) lateral; (D) frontal; (E) occipital; (F) posterior;(G) left antenna (dorsal).Scale bar:= 1 mm.
</paragraph>
</caption>
<caption id="B7D66670FFD6FFACFF80F7897520F7F0" ID-DOI="http://doi.org/10.5281/zenodo.4985587" ID-Zenodo-Dep="4985587" httpUri="https://zenodo.org/record/4985587/files/figure.png" pageId="5" pageNumber="6" startId="5.[94,153,2074,2099]" targetBox="[166,1348,217,2025]" targetPageId="5">
<paragraph id="E31636F8FFD6FFACFF80F7897520F7F0" blockId="5.[94,1419,2074,2131]" pageId="5" pageNumber="6">
Figure 5. 
<taxonomicName id="24A94D7BFFD6FFACFF6DF7897658F790" authority="Vaz &amp; Silveira &amp; Rosa, 2020" authorityName="Vaz &amp; Silveira &amp; Rosa" authorityYear="2020" box="[179,321,2074,2099]" class="Insecta" family="Lampyridae" genus="Psilocladus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="costae" status="sp. nov.">
<emphasis id="D1DDEAEAFFD6FFACFF6DF7897658F790" bold="true" box="[179,321,2074,2099]" italics="true" pageId="5" pageNumber="6">Psilocladus costae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAEE5791FFD6FFACFE9BF789769AF790" box="[325,387,2074,2099]" pageId="5" pageNumber="6" rank="species">sp.nov.</taxonomicNameLabel>
Thorax morphology of adult male.(A-E) Prothorax:(A) dorsal;(B) ventral;(C) frontal;(D) posterior;(E)lateral;(F-I) pterothorax, (F) dorsal;(G) lateral;(H) dorsoventral.Scale bar:= 2 mm.
</paragraph>
</caption>
<paragraph id="E31636F8FFD5FFAFFF74FF497645FE50" blockId="6.[170,794,218,499]" pageId="6" pageNumber="7">
val, widest at metathorax and gradually decreasing in width posteriorly from abdominal segment III), dorsoventrally flattened. Head dark brown, antennae, palpi and mandibles lighter (
<figureCitation id="7B922A7DFFD5FFAFFE62FEA97508FEF0" box="[444,529,314,339]" captionStart="Figure 11" captionStartId="10.[170,229,2074,2099]" captionTargetBox="[176,1489,864,2043]" captionTargetId="figure-330@10.[170,1493,854,2045]" captionTargetPageId="10" captionText="Figure 11. Psilocladus costae sp. nov. Larval morphology. (A-C) Head (dorsal, lateral, ventral); (D) right anterior, middle and posterior legs (from top to bottom, ventral);(E) left laterotergitesVI-VII;(F) abdominal tergaVI-IX.Scale bars:A-C,E, F = 1 mm;D = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985599" httpUri="https://zenodo.org/record/4985599/files/figure.png" pageId="6" pageNumber="7">Figs. 11</figureCitation>
A-C); dorsal surface of body brown, with yellow spot on anterior ⅔ of lateral margins of thorax and abdominal terga I-VI (
<figureCitation id="7B922A7DFFD5FFAFFD54FEE975FCFE30" box="[650,741,378,403]" captionStart="Figure 10" captionStartId="9.[94,153,2106,2131]" captionTargetBox="[100,1413,1083,2074]" captionTargetId="figure-434@9.[94,1417,1075,2077]" captionTargetPageId="9" captionText="Figure 10. Psilocladus costae sp.nov.Mature larval habitus.(A) Dorsal;(B) lateral;(C) ventral.Scale bar:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985597" httpUri="https://zenodo.org/record/4985597/files/figure.png" pageId="6" pageNumber="7">Fig. 10A</figureCitation>
); terga VII-VIII with lateral vitreous spots (
<figureCitation id="7B922A7DFFD5FFAFFD9EFE097580FE10" box="[576,665,410,435]" captionStart="Figure 11" captionStartId="10.[170,229,2074,2099]" captionTargetBox="[176,1489,864,2043]" captionTargetId="figure-330@10.[170,1493,854,2045]" captionTargetPageId="10" captionText="Figure 11. Psilocladus costae sp. nov. Larval morphology. (A-C) Head (dorsal, lateral, ventral); (D) right anterior, middle and posterior legs (from top to bottom, ventral);(E) left laterotergitesVI-VII;(F) abdominal tergaVI-IX.Scale bars:A-C,E, F = 1 mm;D = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985599" httpUri="https://zenodo.org/record/4985599/files/figure.png" pageId="6" pageNumber="7">Fig. 11F</figureCitation>
). Tegument densely covered with short, very fine, grayish pubescence (
<figureCitation id="7B922A7DFFD5FFAFFF29FE497648FE50" box="[247,337,474,499]" captionStart="Figure 10" captionStartId="9.[94,153,2106,2131]" captionTargetBox="[100,1413,1083,2074]" captionTargetId="figure-434@9.[94,1417,1075,2077]" captionTargetPageId="9" captionText="Figure 10. Psilocladus costae sp.nov.Mature larval habitus.(A) Dorsal;(B) lateral;(C) ventral.Scale bar:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985597" httpUri="https://zenodo.org/record/4985597/files/figure.png" pageId="6" pageNumber="7">Fig. 10B</figureCitation>
).
</paragraph>
<paragraph id="E31636F8FFD5FFA0FF74FD89763FFBB0" blockId="6.[170,794,538,883]" lastBlockId="9.[94,719,218,1043]" lastPageId="9" lastPageNumber="10" pageId="6" pageNumber="7">
<emphasis id="D1DDEAEAFFD5FFAFFF74FD89750DFD90" bold="true" box="[170,532,538,563]" pageId="6" pageNumber="7">
Head (
<figureCitation id="7B922A7DFFD5FFAFFF2AFD897654FD90" box="[244,333,538,563]" captionStart="Figure 10" captionStartId="9.[94,153,2106,2131]" captionTargetBox="[100,1413,1083,2074]" captionTargetId="figure-434@9.[94,1417,1075,2077]" captionTargetPageId="9" captionText="Figure 10. Psilocladus costae sp.nov.Mature larval habitus.(A) Dorsal;(B) lateral;(C) ventral.Scale bar:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985597" httpUri="https://zenodo.org/record/4985597/files/figure.png" pageId="6" pageNumber="7">Figs. 10</figureCitation>
B-C, 11A-C, 12A):
</emphasis>
As long as wide, almost entirely retractable into prothorax (
<figureCitation id="7B922A7DFFD5FFAFFDE1FDA9758EFDF0" box="[575,663,570,595]" captionStart="Figure 10" captionStartId="9.[94,153,2106,2131]" captionTargetBox="[100,1413,1083,2074]" captionTargetId="figure-434@9.[94,1417,1075,2077]" captionTargetPageId="9" captionText="Figure 10. Psilocladus costae sp.nov.Mature larval habitus.(A) Dorsal;(B) lateral;(C) ventral.Scale bar:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985597" httpUri="https://zenodo.org/record/4985597/files/figure.png" pageId="6" pageNumber="7">Figs. 10</figureCitation>
B-C); dorsal surface with long, fine and sparse setae on anterior and lateral regions, one stemma with convex lens laterally at base of antennifer; antennifer membranous and glabrous,as long as basal antennomere (
<figureCitation id="7B922A7DFFD5FFAFFD9FFD2975A4FD70" box="[577,701,698,723]" captionStart="Figure 12" captionStartId="11.[94,153,2074,2099]" captionTargetBox="[258,1262,1269,2034]" captionTargetId="figure-570@11.[246,1266,1269,2044]" captionTargetPageId="11" captionText="Figure 12.Psilocladus costae sp.nov.Larval headmorphology.(A) Head,dorsal;(B)maxillolabial complex,ventral;(C) epipharynx;(D) hypopharynx;(E) leftantenna (ventral,dorsal);(F) mandible (ventral,dorsal).Scale bars:A,B = 1 mm;C,D = 0.25 mm;E,F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4985603" httpUri="https://zenodo.org/record/4985603/files/figure.png" pageId="6" pageNumber="7">Figs.12A,E</figureCitation>
). Frontal arms V-shaped, well impressed at posterior half, weakly impressed at anterior half, almost reaching base of antennifer (
<figureCitation id="7B922A7DFFD5FFAFFF37FC89765FFC90" box="[233,326,794,819]" captionStart="Figure 12" captionStartId="11.[94,153,2074,2099]" captionTargetBox="[258,1262,1269,2034]" captionTargetId="figure-570@11.[246,1266,1269,2044]" captionTargetPageId="11" captionText="Figure 12.Psilocladus costae sp.nov.Larval headmorphology.(A) Head,dorsal;(B)maxillolabial complex,ventral;(C) epipharynx;(D) hypopharynx;(E) leftantenna (ventral,dorsal);(F) mandible (ventral,dorsal).Scale bars:A,B = 1 mm;C,D = 0.25 mm;E,F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4985603" httpUri="https://zenodo.org/record/4985603/files/figure.png" pageId="6" pageNumber="7">Fig. 12A</figureCitation>
); epicranial stem very short; clypeolabrum fused to frons, partly membranous, lateral part darkly sclerotized and bilobate, median part translucent, with dark fusiform plate at middle; fusiform plate with anteri- or part fused to cranium, forming acute tooth; posterior part fused to epipharynx and visible through translucent cuticle (
<figureCitation id="7B922A7DFFD5FFAFFC61FEA97330FEF0" box="[959,1065,314,339]" captionStart="Figure 11" captionStartId="10.[170,229,2074,2099]" captionTargetBox="[176,1489,864,2043]" captionTargetId="figure-330@10.[170,1493,854,2045]" captionTargetPageId="10" captionText="Figure 11. Psilocladus costae sp. nov. Larval morphology. (A-C) Head (dorsal, lateral, ventral); (D) right anterior, middle and posterior legs (from top to bottom, ventral);(E) left laterotergitesVI-VII;(F) abdominal tergaVI-IX.Scale bars:A-C,E, F = 1 mm;D = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985599" httpUri="https://zenodo.org/record/4985599/files/figure.png" pageId="6" pageNumber="7">Figs. 11A</figureCitation>
, 
<figureCitation id="7B922A7DFFD5FFAFFBE9FEA97364FEF0" box="[1079,1149,314,339]" captionStart="Figure 12" captionStartId="11.[94,153,2074,2099]" captionTargetBox="[258,1262,1269,2034]" captionTargetId="figure-570@11.[246,1266,1269,2044]" captionTargetPageId="11" captionText="Figure 12.Psilocladus costae sp.nov.Larval headmorphology.(A) Head,dorsal;(B)maxillolabial complex,ventral;(C) epipharynx;(D) hypopharynx;(E) leftantenna (ventral,dorsal);(F) mandible (ventral,dorsal).Scale bars:A,B = 1 mm;C,D = 0.25 mm;E,F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4985603" httpUri="https://zenodo.org/record/4985603/files/figure.png" pageId="6" pageNumber="7">12C,D</figureCitation>
). Antennae (
<figureCitation id="7B922A7DFFD5FFAFFAD3FEA97281FEF0" box="[1293,1432,314,339]" captionStart="Figure 11" captionStartId="10.[170,229,2074,2099]" captionTargetBox="[176,1489,864,2043]" captionTargetId="figure-330@10.[170,1493,854,2045]" captionTargetPageId="10" captionText="Figure 11. Psilocladus costae sp. nov. Larval morphology. (A-C) Head (dorsal, lateral, ventral); (D) right anterior, middle and posterior legs (from top to bottom, ventral);(E) left laterotergitesVI-VII;(F) abdominal tergaVI-IX.Scale bars:A-C,E, F = 1 mm;D = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985599" httpUri="https://zenodo.org/record/4985599/files/figure.png" pageId="6" pageNumber="7">Figs. 11A, C</figureCitation>
, 
<figureCitation id="7B922A7DFFD5FFAFFA7BFEA972D6FEF0" box="[1445,1487,314,339]" captionStart="Figure 12" captionStartId="11.[94,153,2074,2099]" captionTargetBox="[258,1262,1269,2034]" captionTargetId="figure-570@11.[246,1266,1269,2044]" captionTargetPageId="11" captionText="Figure 12.Psilocladus costae sp.nov.Larval headmorphology.(A) Head,dorsal;(B)maxillolabial complex,ventral;(C) epipharynx;(D) hypopharynx;(E) leftantenna (ventral,dorsal);(F) mandible (ventral,dorsal).Scale bars:A,B = 1 mm;C,D = 0.25 mm;E,F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4985603" httpUri="https://zenodo.org/record/4985603/files/figure.png" pageId="6" pageNumber="7">12E</figureCitation>
) elongate, with three antennomeres, antennomere I partially sclerotized, sparsely setose, setae on membranous region very short; antennomere II 1.1× as long as I, fully sclerotized, sparsely setose, laterally flattened, apex ventrally with elliptical, flattened sensorium; antennomere III attached dorsally to antennomere II, digitiform, 0.2× as long as antennomere II, with few fine setae subapically, one very short spiniform projection at apex. Epipharynx (
<figureCitation id="7B922A7DFFD5FFAFFCB2FDC974D0FDD0" box="[876,969,602,627]" captionStart="Figure 12" captionStartId="11.[94,153,2074,2099]" captionTargetBox="[258,1262,1269,2034]" captionTargetId="figure-570@11.[246,1266,1269,2044]" captionTargetPageId="11" captionText="Figure 12.Psilocladus costae sp.nov.Larval headmorphology.(A) Head,dorsal;(B)maxillolabial complex,ventral;(C) epipharynx;(D) hypopharynx;(E) leftantenna (ventral,dorsal);(F) mandible (ventral,dorsal).Scale bars:A,B = 1 mm;C,D = 0.25 mm;E,F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4985603" httpUri="https://zenodo.org/record/4985603/files/figure.png" pageId="6" pageNumber="7">Fig. 12C</figureCitation>
) with cross-shaped sclerite and two triangular striate plates; plates with anterior margin densely covered with fine setae and two orifices at lateral margins; hypopharynx (
<figureCitation id="7B922A7DFFD5FFAFFBD3FD297376FD70" box="[1037,1135,698,723]" captionStart="Figure 12" captionStartId="11.[94,153,2074,2099]" captionTargetBox="[258,1262,1269,2034]" captionTargetId="figure-570@11.[246,1266,1269,2044]" captionTargetPageId="11" captionText="Figure 12.Psilocladus costae sp.nov.Larval headmorphology.(A) Head,dorsal;(B)maxillolabial complex,ventral;(C) epipharynx;(D) hypopharynx;(E) leftantenna (ventral,dorsal);(F) mandible (ventral,dorsal).Scale bars:A,B = 1 mm;C,D = 0.25 mm;E,F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4985603" httpUri="https://zenodo.org/record/4985603/files/figure.png" pageId="6" pageNumber="7">Fig. 12D</figureCitation>
) with anterior lobe triangular, densely setose; posterior part sclerotized, glabrous, with elongate concave projection. Mandibles (
<figureCitation id="7B922A7DFFD5FFAFFAAAFD6972D6FCB0" box="[1396,1487,762,787]" captionStart="Figure 12" captionStartId="11.[94,153,2074,2099]" captionTargetBox="[258,1262,1269,2034]" captionTargetId="figure-570@11.[246,1266,1269,2044]" captionTargetPageId="11" captionText="Figure 12.Psilocladus costae sp.nov.Larval headmorphology.(A) Head,dorsal;(B)maxillolabial complex,ventral;(C) epipharynx;(D) hypopharynx;(E) leftantenna (ventral,dorsal);(F) mandible (ventral,dorsal).Scale bars:A,B = 1 mm;C,D = 0.25 mm;E,F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4985603" httpUri="https://zenodo.org/record/4985603/files/figure.png" pageId="6" pageNumber="7">Fig. 12F</figureCitation>
) symmetrical, falcate, with a channel opening near apex, with ventral tuft of fine setae near channel opening; lateral margin at base covered with fine long setae; mesal margin at base with very short setae; row of sparse setae along channel dorsally; retinaculum well developed, forming a large and acute tooth; mesal membranous extension densely setose. Maxillolabial complex (
<figureCitation id="7B922A7DFFDAFFA0FDB3FEA975D1FEF0" box="[621,712,314,339]" captionStart="Figure 12" captionStartId="11.[94,153,2074,2099]" captionTargetBox="[258,1262,1269,2034]" captionTargetId="figure-570@11.[246,1266,1269,2044]" captionTargetPageId="11" captionText="Figure 12.Psilocladus costae sp.nov.Larval headmorphology.(A) Head,dorsal;(B)maxillolabial complex,ventral;(C) epipharynx;(D) hypopharynx;(E) leftantenna (ventral,dorsal);(F) mandible (ventral,dorsal).Scale bars:A,B = 1 mm;C,D = 0.25 mm;E,F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4985603" httpUri="https://zenodo.org/record/4985603/files/figure.png" pageId="9" pageNumber="10">Fig. 12B</figureCitation>
) separated from ventral head capsule by broad membrane; maxillae with cardo narrow, elongate, 0.5× as long as stripes, subparallel-sided; stipites elongate, with short membranous area on anterior margin, few fine setae irregularly distributed and a stout seta near base of galea; palpus 4-segmented, tapering toward apex with sparse fine setae and a stout seta on anterolateral margin of palpomere I; galea 2-articulated: basal palpomere triangular, as wide as long; apical palpomere digitiform, 5× longer than wide, with one stout long seta apically and few shorter setae subapically; lacinia consisting of densely pubescent structure connected to dorsomesal stipital edge; labium (
<figureCitation id="7B922A7DFFDAFFA0FE5FFD4976C6FD50" box="[385,479,730,755]" captionStart="Figure 12" captionStartId="11.[94,153,2074,2099]" captionTargetBox="[258,1262,1269,2034]" captionTargetId="figure-570@11.[246,1266,1269,2044]" captionTargetPageId="11" captionText="Figure 12.Psilocladus costae sp.nov.Larval headmorphology.(A) Head,dorsal;(B)maxillolabial complex,ventral;(C) epipharynx;(D) hypopharynx;(E) leftantenna (ventral,dorsal);(F) mandible (ventral,dorsal).Scale bars:A,B = 1 mm;C,D = 0.25 mm;E,F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4985603" httpUri="https://zenodo.org/record/4985603/files/figure.png" pageId="9" pageNumber="10">Fig. 12B</figureCitation>
): prementum densely covered with moderately long setae, emarginated between palpi; palpus 2-articulated, apical palpomere as long as the basal one, strongly tapered apicad, with slot sensorium on outer side; mentum with anterior ⅓ membranous; posterior ⅔ sclerotized with pair of long setae at midlength; submentum and gula membranous. Post-occipital region as long as head, glabrous, with 3 elongate weakly sclerotized areas contiguous to anterior prothoracic collar.
</paragraph>
<caption id="B7D66670FFD5FFAFFF72F789755FF7F0" ID-DOI="http://doi.org/10.5281/zenodo.4985589" ID-Zenodo-Dep="4985589" httpUri="https://zenodo.org/record/4985589/files/figure.png" pageId="6" pageNumber="7" startId="6.[172,231,2074,2099]" targetBox="[176,1493,913,2040]" targetPageId="6">
<paragraph id="E31636F8FFD5FFAFFF72F789755FF7F0" blockId="6.[170,1494,2074,2131]" pageId="6" pageNumber="7">
Figure 6. 
<taxonomicName id="24A94D7BFFD5FFAFFEDEF7897697F790" authority="Vaz &amp; Silveira &amp; Rosa, 2020" authorityName="Vaz &amp; Silveira &amp; Rosa" authorityYear="2020" box="[256,398,2074,2099]" class="Insecta" family="Lampyridae" genus="Psilocladus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="costae" status="sp. nov.">
<emphasis id="D1DDEAEAFFD5FFAFFEDEF7897697F790" bold="true" box="[256,398,2074,2099]" italics="true" pageId="6" pageNumber="7">Psilocladus costae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAEE5791FFD5FFAFFE4DF78976C8F790" box="[403,465,2074,2099]" pageId="6" pageNumber="7" rank="species">sp.nov.</taxonomicNameLabel>
Thoracic appendages of adult male. (A-C) Left legs,ventral (anterior, middle and posterior,respectively);(D-F) elytron (dorsal, lateral and ventral);(G) left wing.Scale bar:2 mm.
</paragraph>
</caption>
<caption id="B7D66670FFD4FFAEFF80FC0176B9FC68" ID-DOI="http://doi.org/10.5281/zenodo.4985591" ID-Zenodo-Dep="4985591" httpUri="https://zenodo.org/record/4985591/files/figure.png" pageId="7" pageNumber="8" startId="7.[94,153,914,939]" targetBox="[97,1416,212,872]" targetPageId="7">
<paragraph id="E31636F8FFD4FFAEFF80FC0176B9FC68" blockId="7.[94,1419,914,971]" pageId="7" pageNumber="8">
Figure 7. 
<taxonomicName id="24A94D7BFFD4FFAEFF6DFC017659FC08" authority="Vaz &amp; Silveira &amp; Rosa, 2020" authorityName="Vaz &amp; Silveira &amp; Rosa" authorityYear="2020" box="[179,320,914,939]" class="Insecta" family="Lampyridae" genus="Psilocladus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="costae" status="sp. nov.">
<emphasis id="D1DDEAEAFFD4FFAEFF6DFC017659FC08" bold="true" box="[179,320,914,939]" italics="true" pageId="7" pageNumber="8">Psilocladus costae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAEE5791FFD4FFAEFE9AFC01769BFC08" box="[324,386,914,939]" pageId="7" pageNumber="8" rank="species">sp.nov.</taxonomicNameLabel>
Terminal segments and aedeagus of male.(A) SternumVIII;(B) pygidium;(C) syntergite;(D) sternum IX;(E-G) aedeagus (dorsal, lateral and ventral).Scale bar:= 1 mm.
</paragraph>
</caption>
<caption id="B7D66670FFD4FFAEFF80F7A97489F7F0" ID-DOI="http://doi.org/10.5281/zenodo.4985593" ID-Zenodo-Dep="4985593" box="[94,912,2106,2131]" httpUri="https://zenodo.org/record/4985593/files/figure.png" pageId="7" pageNumber="8" startId="7.[94,153,2106,2131]" targetBox="[177,1339,1008,2071]" targetPageId="7">
<paragraph id="E31636F8FFD4FFAEFF80F7A97489F7F0" blockId="7.[94,912,2106,2131]" box="[94,912,2106,2131]" pageId="7" pageNumber="8">
Figure 8. 
<taxonomicName id="24A94D7BFFD4FFAEFF6AF7A9765BF7F0" authority="Vaz &amp; Silveira &amp; Rosa, 2020" authorityName="Vaz &amp; Silveira &amp; Rosa" authorityYear="2020" box="[180,322,2106,2131]" class="Insecta" family="Lampyridae" genus="Psilocladus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="costae" status="sp. nov.">
<emphasis id="D1DDEAEAFFD4FFAEFF6AF7A9765BF7F0" bold="true" box="[180,322,2106,2131]" italics="true" pageId="7" pageNumber="8">Psilocladus costae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAEE5791FFD4FFAEFE98F7A9769DF7F0" box="[326,388,2106,2131]" pageId="7" pageNumber="8" rank="species">sp.nov.</taxonomicNameLabel>
Adult female habitus.(A) Dorsal;(B) ventral.Scale bar:= 5 mm.
</paragraph>
</caption>
<caption id="B7D66670FFDBFFA1FF74F789738CF7F0" ID-DOI="http://doi.org/10.5281/zenodo.4985595" ID-Zenodo-Dep="4985595" httpUri="https://zenodo.org/record/4985595/files/figure.png" pageId="8" pageNumber="9" startId="8.[170,229,2074,2099]" targetBox="[183,1491,205,2030]" targetPageId="8">
<paragraph id="E31636F8FFDBFFA1FF74F789738CF7F0" blockId="8.[170,1494,2074,2131]" pageId="8" pageNumber="9">
Figure 9. 
<taxonomicName id="24A94D7BFFDBFFA1FEDDF789768BF790" authority="Vaz &amp; Silveira &amp; Rosa, 2020" authorityName="Vaz &amp; Silveira &amp; Rosa" authorityYear="2020" box="[259,402,2074,2099]" class="Insecta" family="Lampyridae" genus="Psilocladus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="costae" status="sp. nov.">
<emphasis id="D1DDEAEAFFDBFFA1FEDDF789768BF790" bold="true" box="[259,402,2074,2099]" italics="true" pageId="8" pageNumber="9">Psilocladus costae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAEE5791FFDBFFA1FE46F78976C1F790" box="[408,472,2074,2099]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
Adult female morphology. (A-E) Head; (A) dorsal; (B) ventral; (C) frontal; (D) occipital; (E) lateral; (F) left antenna (dorsal); (G) sternum VIII;(H) pygidium;(I, J) ovipositor (dorsal,ventral),arrow indicates stylus.Scale bars:A-E = 1 mm;F-I = 2 mm.
</paragraph>
</caption>
<paragraph id="E31636F8FFDAFFA0FCC4FF49722DFCF0" blockId="9.[794,1419,218,851]" pageId="9" pageNumber="10">
<emphasis id="D1DDEAEAFFDAFFA0FCC4FF4974CDFF50" bold="true" box="[794,980,218,243]" pageId="9" pageNumber="10">
Thorax(
<figureCitation id="7B922A7DFFDAFFA0FCA6FF4974DEFF50" box="[888,967,218,243]" captionStart="Figure 10" captionStartId="9.[94,153,2106,2131]" captionTargetBox="[100,1413,1083,2074]" captionTargetId="figure-434@9.[94,1417,1075,2077]" captionTargetPageId="9" captionText="Figure 10. Psilocladus costae sp.nov.Mature larval habitus.(A) Dorsal;(B) lateral;(C) ventral.Scale bar:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985597" httpUri="https://zenodo.org/record/4985597/files/figure.png" pageId="9" pageNumber="10">Fig. 10</figureCitation>
):
</emphasis>
tip of posterior angles of terga with long stout seta; pronotum semielliptical 1.3-1.6× wider than long, with posterior margin slightly emarginate, prosternum weakly sclerotized with short endocarinate rod between procoxae; anterior prothoracic collar membranous dorsally,strongly sclerotized on anterior ⅔ ventrally; sclerotized region finely punctate/setose. Mesonotum as long as metanotum, both transverse with anterior angles rounded, posterior angles straight; mesonotum 3-3.3× wider than long; metanotum 3.3-3.6× wider than long; mesepisternum with a functional biforous spiracle on anterior corner; ventral surface of meso- and metathorax membranous medially, with sclerotized regions on pleurae laterally to coxae. Legs (
<figureCitation id="7B922A7DFFDAFFA0FB59FDE973FFFD30" box="[1159,1254,634,659]" captionStart="Figure 11" captionStartId="10.[170,229,2074,2099]" captionTargetBox="[176,1489,864,2043]" captionTargetId="figure-330@10.[170,1493,854,2045]" captionTargetPageId="10" captionText="Figure 11. Psilocladus costae sp. nov. Larval morphology. (A-C) Head (dorsal, lateral, ventral); (D) right anterior, middle and posterior legs (from top to bottom, ventral);(E) left laterotergitesVI-VII;(F) abdominal tergaVI-IX.Scale bars:A-C,E, F = 1 mm;D = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985599" httpUri="https://zenodo.org/record/4985599/files/figure.png" pageId="9" pageNumber="10">Fig. 11D</figureCitation>
): coxa conical, strongly projected ventrad, partly strongly sclerotized, with fine long setae; trochanter with a few short stout setae; femur with sparse, evenly distributed spiniform setae and few short stout setae, and one long stout seta ventrally; tibiotarsus densely covered with spiniform setae; pretarsus with one seta on each side at base.
</paragraph>
<paragraph id="E31636F8FFDAFFA3FCC4FCE9753AFC90" blockId="9.[794,1418,890,1043]" lastBlockId="10.[170,795,218,819]" lastPageId="10" lastPageNumber="11" pageId="9" pageNumber="10">
<emphasis id="D1DDEAEAFFDAFFA0FCC4FCE97341FC30" bold="true" box="[794,1112,890,915]" pageId="9" pageNumber="10">
Abdomen (
<figureCitation id="7B922A7DFFDAFFA0FC42FCE974E2FC30" box="[924,1019,890,915]" captionStart="Figure 10" captionStartId="9.[94,153,2106,2131]" captionTargetBox="[100,1413,1083,2074]" captionTargetId="figure-434@9.[94,1417,1075,2077]" captionTargetPageId="9" captionText="Figure 10. Psilocladus costae sp.nov.Mature larval habitus.(A) Dorsal;(B) lateral;(C) ventral.Scale bar:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985597" httpUri="https://zenodo.org/record/4985597/files/figure.png" pageId="9" pageNumber="10">Figs. 10</figureCitation>
, 
<figureCitation id="7B922A7DFFDAFFA0FBD8FCE97306FC30" box="[1030,1055,890,915]" captionStart="Figure 11" captionStartId="10.[170,229,2074,2099]" captionTargetBox="[176,1489,864,2043]" captionTargetId="figure-330@10.[170,1493,854,2045]" captionTargetPageId="10" captionText="Figure 11. Psilocladus costae sp. nov. Larval morphology. (A-C) Head (dorsal, lateral, ventral); (D) right anterior, middle and posterior legs (from top to bottom, ventral);(E) left laterotergitesVI-VII;(F) abdominal tergaVI-IX.Scale bars:A-C,E, F = 1 mm;D = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985599" httpUri="https://zenodo.org/record/4985599/files/figure.png" pageId="9" pageNumber="10">11</figureCitation>
E-F):
</emphasis>
Gradually narrowed posteriorly from segment III; terga I-VIII with well impressed sagittal ecdisial line, anterior angles rounded, posterior angles acute and strongly projected posteriad with stout long seta at tip; terga I-VI transversal, subequal in length; tergum I about 4.3× wider than long, VI 3.7× wider than long; tergum VIII 1.5× wider than VII, with posterior margin broadly emarginate; tergum IX as long as wide, lateral margins subparallel, posterior angles weakly projected with tuft of foliaceus setae at tip, posterior margin slightly rounded. Sternites I-IX sclerotized, with two pairs of parasagittal very short stout setae (at midlength and posterior margin) and long stout seta at posterior angles; sternites VI-VIII with additional pair of stout seta near lateral margins; stout setae increasing in length from sternites I-IX; laterotergites I-VIII 0.6-0.8× as large as respective sternites, sclerotized, posterior corners projected, with 2 prominences; anterior prominence with biforous spiracle, posterior prominence with a stout long seta; sternite VIII with pair of light organ on anterior corners. Segment X (
<figureCitation id="7B922A7DFFD9FFA3FF10FD297633FD70" box="[206,298,698,723]" captionStart="Figure 10" captionStartId="9.[94,153,2106,2131]" captionTargetBox="[100,1413,1083,2074]" captionTargetId="figure-434@9.[94,1417,1075,2077]" captionTargetPageId="9" captionText="Figure 10. Psilocladus costae sp.nov.Mature larval habitus.(A) Dorsal;(B) lateral;(C) ventral.Scale bar:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985597" httpUri="https://zenodo.org/record/4985597/files/figure.png" pageId="10" pageNumber="11">Figs. 10</figureCitation>
B-C) attached under segment IX, tergum broadly conate or fused to sternum IX; sternum glabrous, most part sclerotized, except by subapical membranous transverse area; pygopod present.
</paragraph>
<caption id="B7D66670FFDAFFA0FF80F7A974EEF7F0" ID-DOI="http://doi.org/10.5281/zenodo.4985597" ID-Zenodo-Dep="4985597" box="[94,1015,2106,2131]" httpUri="https://zenodo.org/record/4985597/files/figure.png" pageId="9" pageNumber="10" startId="9.[94,153,2106,2131]" targetBox="[100,1413,1083,2074]" targetPageId="9">
<paragraph id="E31636F8FFDAFFA0FF80F7A974EEF7F0" blockId="9.[94,1015,2106,2131]" box="[94,1015,2106,2131]" pageId="9" pageNumber="10">
Figure 10. 
<taxonomicName id="24A94D7BFFDAFFA0FF61F7A97654F7F0" authority="Vaz &amp; Silveira &amp; Rosa, 2020" authorityName="Vaz &amp; Silveira &amp; Rosa" authorityYear="2020" box="[191,333,2106,2131]" class="Insecta" family="Lampyridae" genus="Psilocladus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="species" species="costae" status="sp. nov.">
<emphasis id="D1DDEAEAFFDAFFA0FF61F7A97654F7F0" bold="true" box="[191,333,2106,2131]" italics="true" pageId="9" pageNumber="10">Psilocladus costae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAEE5791FFDAFFA0FE8FF7A97696F7F0" box="[337,399,2106,2131]" pageId="9" pageNumber="10" rank="species">sp.nov.</taxonomicNameLabel>
Mature larval habitus.(A) Dorsal;(B) lateral;(C) ventral.Scale bar:= 2 mm.
</paragraph>
</caption>
<paragraph id="E31636F8FFD9FFA3FCBBFF497328FD70" blockId="10.[869,1494,218,723]" pageId="10" pageNumber="11">
<emphasis id="D1DDEAEAFFD9FFA3FCBBFF497308FF50" bold="true" box="[869,1041,218,243]" pageId="10" pageNumber="11">
Pupa (
<figureCitation id="7B922A7DFFD9FFA3FC6FFF49731DFF50" box="[945,1028,218,243]" captionStart="Figure 13" captionStartId="12.[170,229,1198,1223]" captionTargetBox="[174,1491,210,1163]" captionTargetId="figure-1@12.[170,1493,204,1168]" captionTargetPageId="12" captionText="Figure 13. Psilocladus costae sp.nov.Pupal habitus.(A) Dorsal;(B) lateral;(C) ventral.Scale bar:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985605" httpUri="https://zenodo.org/record/4985605/files/figure.png" pageId="10" pageNumber="11">Fig. 13</figureCitation>
):
</emphasis>
Total length 
<emphasis id="D1DDEAEAFFD9FFA3FB78FF4973DCFF50" box="[1190,1221,218,243]" italics="true" pageId="10" pageNumber="11">ca.</emphasis>
7.0 mm, adecticous, exarate, white. Pronotum semicircular, anterior and lateral margins densely setose, setae long and stout. Dorsal surface with short setae. Posterior margin with posteri- or angles rounded, projected posteriad, median region straight. Mesothorax with a pair of reniform spiracles on posterior half of pleurite. Metanotum 1.1× as long as mesonotum, with a pair of long setae on posterior third. Abdominal terga I-VIII transverse, with posterior angles acute, strongly projected posteriad; lateral margins densely setose; setae long stout; tergite IX subquadrate with posterior angles weakly projected, with tuft of stout setae.Pair of light organ on sterna VI and VII (
<figureCitation id="7B922A7DFFD9FFA3FA91FDC972B0FDD0" box="[1359,1449,602,627]" captionStart="Figure 14" captionStartId="12.[170,229,2106,2131]" captionTargetBox="[170,1493,1245,2076]" captionTargetId="figure-22@12.[170,1493,1245,2076]" captionTargetPageId="12" captionText="Figure 14. Psilocladus costae sp.nov.Immature bioluminescence.(A) Larva (dorsal);(B) pupa (ventral).Length:larva = 18 mm;pupa = 7 mm." figureDoi="http://doi.org/10.5281/zenodo.4985609" httpUri="https://zenodo.org/record/4985609/files/figure.png" pageId="10" pageNumber="11">Fig. 14B</figureCitation>
), inconspicuous in alcohol-preserved specimen. Abdominal spiracles located ventrally on posterior angles of pleurae on segments I-VIII.
</paragraph>
</subSubSection>
<subSubSection id="ABB36573FFD9FFA2FCBBFD697299FED0" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="discussion">
<paragraph id="E31636F8FFD9FFA2FCBBFD697299FED0" blockId="10.[869,1493,762,819]" lastBlockId="11.[794,1418,218,371]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<emphasis id="D1DDEAEAFFD9FFA3FCBBFD6974CAFCB0" bold="true" box="[869,979,762,787]" pageId="10" pageNumber="11">Remarks:</emphasis>
<taxonomicName id="24A94D7BFFD9FFA3FC37FD6973A1FCB0" authority="Vaz &amp; Silveira &amp; Rosa, 2020" authorityName="Vaz &amp; Silveira &amp; Rosa" authorityYear="2020" box="[1001,1208,762,787]" class="Insecta" family="Lampyridae" genus="Psilocladus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="costae" status="sp. nov.">
<emphasis id="D1DDEAEAFFD9FFA3FC37FD6973A1FCB0" box="[1001,1208,762,787]" italics="true" pageId="10" pageNumber="11">Psilocladus costae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAEE5791FFD9FFA3FB11FD69722AFCB0" box="[1231,1331,762,787]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
is the first 
<taxonomicName id="24A94D7BFFD9FFA3FCBBFC8974EFFC90" box="[869,1014,794,819]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="subFamily" subFamily="Psilocladinae">Psilocladinae</taxonomicName>
species to have their immature stages de- scribed. The larva of this species is more similar to those of 
<taxonomicName id="24A94D7BFFD8FFA2FFA7FF6977EDFEB0" authorityName="Lacordaire" authorityYear="1857" box="[121,244,250,275]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subFamily" subFamily="Photurinae">Photurinae</taxonomicName>
species: 
<taxonomicName id="24A94D7BFFD8FFA2FE8DFF6975D7FEB0" authority="Curtis (Rosa, 2007)" authorityName="Curtis (Rosa" authorityYear="2007" box="[339,718,250,275]" class="Insecta" family="Lampyridae" genus="Photuris" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="fulvipes">
<emphasis id="D1DDEAEAFFD8FFA2FE8DFF697518FEB0" box="[339,513,250,275]" italics="true" pageId="11" pageNumber="12">Photuris fulvipes</emphasis>
Curtis (
<bibRefCitation id="87384B09FFD8FFA2FD8EFF6975D0FEB0" author="Rosa, S. P." box="[592,713,250,275]" pageId="11" pageNumber="12" pagination="125 - 130" refId="ref8750" refString="Rosa, S. P. 2007. Description of Photuris fulvipes (Blanchard) immatures (Coleoptera, Lampyridae, Photurinae) and bionomic aspects under laboratory conditions. Revista Brasileira de Entomologia, 51 (2): 125 - 130. DOI" type="journal article" year="2007">Rosa,2007</bibRefCitation>
)
</taxonomicName>
and an unidentified species of 
<taxonomicName id="24A94D7BFFD8FFA2FE68FE897630FEF0" authority="Motschulsky (Costa et al., 1988)" class="Insecta" family="Lampyridae" genus="Bicellonycha" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="D1DDEAEAFFD8FFA2FE68FE897525FE90" box="[438,572,282,307]" italics="true" pageId="11" pageNumber="12">Bicellonycha</emphasis>
Motschulsky (Costa 
<emphasis id="D1DDEAEAFFD8FFA2FF74FEA977FAFEF0" box="[170,227,314,339]" italics="true" pageId="11" pageNumber="12">et al.,</emphasis>
1988)
</taxonomicName>
. Those species share the oblong body (with semioval pronotum, widest at metathorax and gradually decreasing in width from abdominal segment III) (
<figureCitation id="7B922A7DFFD8FFA2FF5BFE0977FBFE10" box="[133,226,410,435]" captionStart="Figure 10" captionStartId="9.[94,153,2106,2131]" captionTargetBox="[100,1413,1083,2074]" captionTargetId="figure-434@9.[94,1417,1075,2077]" captionTargetPageId="9" captionText="Figure 10. Psilocladus costae sp.nov.Mature larval habitus.(A) Dorsal;(B) lateral;(C) ventral.Scale bar:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985597" httpUri="https://zenodo.org/record/4985597/files/figure.png" pageId="11" pageNumber="12">Fig. 10A</figureCitation>
), head with frontal arms well impressed posteriorly and divergent anteriad (V- or U-shaped), clypeolabrum with a median tooth darkly sclerotized, sensorium on second antennomere flattened (
<figureCitation id="7B922A7DFFD8FFA2FDD0FE69756DFDB0" box="[526,628,506,531]" captionStart="Figure 11" captionStartId="10.[170,229,2074,2099]" captionTargetBox="[176,1489,864,2043]" captionTargetId="figure-330@10.[170,1493,854,2045]" captionTargetPageId="10" captionText="Figure 11. Psilocladus costae sp. nov. Larval morphology. (A-C) Head (dorsal, lateral, ventral); (D) right anterior, middle and posterior legs (from top to bottom, ventral);(E) left laterotergitesVI-VII;(F) abdominal tergaVI-IX.Scale bars:A-C,E, F = 1 mm;D = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985599" httpUri="https://zenodo.org/record/4985599/files/figure.png" pageId="11" pageNumber="12">Figs. 11A</figureCitation>
, 
<figureCitation id="7B922A7DFFD8FFA2FDA3FE6975DDFDB0" box="[637,708,506,531]" captionStart="Figure 12" captionStartId="11.[94,153,2074,2099]" captionTargetBox="[258,1262,1269,2034]" captionTargetId="figure-570@11.[246,1266,1269,2044]" captionTargetPageId="11" captionText="Figure 12.Psilocladus costae sp.nov.Larval headmorphology.(A) Head,dorsal;(B)maxillolabial complex,ventral;(C) epipharynx;(D) hypopharynx;(E) leftantenna (ventral,dorsal);(F) mandible (ventral,dorsal).Scale bars:A,B = 1 mm;C,D = 0.25 mm;E,F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4985603" httpUri="https://zenodo.org/record/4985603/files/figure.png" pageId="11" pageNumber="12">12A, E</figureCitation>
), pretarsus with one basal seta on each side, and presence of localized stout setae on thorax and abdomen (
<figureCitation id="7B922A7DFFD8FFA2FDB3FDA975DDFDF0" box="[621,708,570,595]" captionStart="Figure 11" captionStartId="10.[170,229,2074,2099]" captionTargetBox="[176,1489,864,2043]" captionTargetId="figure-330@10.[170,1493,854,2045]" captionTargetPageId="10" captionText="Figure 11. Psilocladus costae sp. nov. Larval morphology. (A-C) Head (dorsal, lateral, ventral); (D) right anterior, middle and posterior legs (from top to bottom, ventral);(E) left laterotergitesVI-VII;(F) abdominal tergaVI-IX.Scale bars:A-C,E, F = 1 mm;D = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985599" httpUri="https://zenodo.org/record/4985599/files/figure.png" pageId="11" pageNumber="12">Fig.11E</figureCitation>
). They differ in the density and distributional pattern of the stout setae on thoracic and abdominal segments. Stout setae are sparser in 
<taxonomicName id="24A94D7BFFD8FFA2FE5EFD097558FD10" authority="Vaz &amp; Silveira &amp; Rosa, 2020" authorityName="Vaz &amp; Silveira &amp; Rosa" authorityYear="2020" box="[384,577,666,691]" class="Insecta" family="Lampyridae" genus="Psilocladus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="costae" status="sp. nov.">
<emphasis id="D1DDEAEAFFD8FFA2FE5EFD097558FD10" box="[384,577,666,691]" italics="true" pageId="11" pageNumber="12">Psilocladus costae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAEE5791FFD8FFA2FD96FD097584FD10" box="[584,669,666,691]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
and in 
<emphasis id="D1DDEAEAFFD8FFA2FFA0FD297651FD70" box="[126,328,698,723]" italics="true" pageId="11" pageNumber="12">
<taxonomicName id="24A94D7BFFD8FFA2FFA0FD29765CFD70" box="[126,325,698,723]" class="Insecta" family="Lampyridae" genus="Photuris" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="femoralis">Photuris femoralis</taxonomicName>
,
</emphasis>
but in 
<taxonomicName id="24A94D7BFFD8FFA2FE7AFD29751BFD70" authority="Vaz &amp; Silveira &amp; Rosa, 2020" authorityName="Vaz &amp; Silveira &amp; Rosa" authorityYear="2020" box="[420,514,698,723]" class="Insecta" family="Lampyridae" genus="Psilocladus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="costae" status="sp. nov.">
<emphasis id="D1DDEAEAFFD8FFA2FE7AFD29751BFD70" box="[420,514,698,723]" italics="true" pageId="11" pageNumber="12">P. costae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAEE5791FFD8FFA2FDD3FD29757FFD70" box="[525,614,698,723]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
they are shorter and absent on median part of thoracic and abdominal terga. Larvae of 
<taxonomicName id="24A94D7BFFD8FFA2FEAEFD6976D3FCB0" authority="Vaz &amp; Silveira &amp; Rosa, 2020" authorityName="Vaz &amp; Silveira &amp; Rosa" authorityYear="2020" box="[368,458,762,787]" class="Insecta" family="Lampyridae" genus="Psilocladus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="costae" status="sp. nov.">
<emphasis id="D1DDEAEAFFD8FFA2FEAEFD6976D3FCB0" box="[368,458,762,787]" italics="true" pageId="11" pageNumber="12">P.costae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAEE5791FFD8FFA2FE0EFD69753AFCB0" box="[464,547,762,787]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
also differ from both 
<taxonomicName id="24A94D7BFFD8FFA2FF43FC897601FC90" box="[157,280,794,819]" class="Insecta" family="Lampyridae" genus="Photuris" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="femoralis">
<emphasis id="D1DDEAEAFFD8FFA2FF43FC897601FC90" box="[157,280,794,819]" italics="true" pageId="11" pageNumber="12">P. femoralis</emphasis>
</taxonomicName>
and 
<taxonomicName id="24A94D7BFFD8FFA2FE8BFC897510FC90" box="[341,521,794,819]" class="Insecta" family="Lampyridae" genus="Bycellonycha" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="D1DDEAEAFFD8FFA2FE8BFC8976F8FC90" box="[341,481,794,819]" italics="true" pageId="11" pageNumber="12">Bycellonycha</emphasis>
sp.
</taxonomicName>
larvae by having mandible sparsely setose and abruptly curved on apical half (
<figureCitation id="7B922A7DFFD8FFA2FF4DFCC977F5FCD0" box="[147,236,858,883]" captionStart="Figure 12" captionStartId="11.[94,153,2074,2099]" captionTargetBox="[258,1262,1269,2034]" captionTargetId="figure-570@11.[246,1266,1269,2044]" captionTargetPageId="11" captionText="Figure 12.Psilocladus costae sp.nov.Larval headmorphology.(A) Head,dorsal;(B)maxillolabial complex,ventral;(C) epipharynx;(D) hypopharynx;(E) leftantenna (ventral,dorsal);(F) mandible (ventral,dorsal).Scale bars:A,B = 1 mm;C,D = 0.25 mm;E,F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4985603" httpUri="https://zenodo.org/record/4985603/files/figure.png" pageId="11" pageNumber="12">Fig. 12F</figureCitation>
), hypopharynx with single anterior lobe and posterior part elongate (
<figureCitation id="7B922A7DFFD8FFA2FEB6FCE976DCFC30" box="[360,453,890,915]" captionStart="Figure 12" captionStartId="11.[94,153,2074,2099]" captionTargetBox="[258,1262,1269,2034]" captionTargetId="figure-570@11.[246,1266,1269,2044]" captionTargetPageId="11" captionText="Figure 12.Psilocladus costae sp.nov.Larval headmorphology.(A) Head,dorsal;(B)maxillolabial complex,ventral;(C) epipharynx;(D) hypopharynx;(E) leftantenna (ventral,dorsal);(F) mandible (ventral,dorsal).Scale bars:A,B = 1 mm;C,D = 0.25 mm;E,F = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4985603" httpUri="https://zenodo.org/record/4985603/files/figure.png" pageId="11" pageNumber="12">Fig. 12D</figureCitation>
), vitreous spots on terga VII and VIII, and tergite IX somewhat quadrate (
<figureCitation id="7B922A7DFFD8FFA2FDB1FC0975D1FC10" box="[623,712,922,947]" captionStart="Figure 11" captionStartId="10.[170,229,2074,2099]" captionTargetBox="[176,1489,864,2043]" captionTargetId="figure-330@10.[170,1493,854,2045]" captionTargetPageId="10" captionText="Figure 11. Psilocladus costae sp. nov. Larval morphology. (A-C) Head (dorsal, lateral, ventral); (D) right anterior, middle and posterior legs (from top to bottom, ventral);(E) left laterotergitesVI-VII;(F) abdominal tergaVI-IX.Scale bars:A-C,E, F = 1 mm;D = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985599" httpUri="https://zenodo.org/record/4985599/files/figure.png" pageId="11" pageNumber="12">Fig. 11F</figureCitation>
) (in 
<taxonomicName id="24A94D7BFFD8FFA2FF58FC297618FC70" authorityName="Lacordaire" authorityYear="1857" box="[134,257,954,979]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subFamily" subFamily="Photurinae">Photurinae</taxonomicName>
larvae, mandible is densely setose and gradually curved, the hypopharynx is bilobed with posterior part short, vitreous spots are absent, and the tergite IX is semioval). The pupa of 
<taxonomicName id="24A94D7BFFD8FFA2FE1BFB89753BFB90" authority="Vaz &amp; Silveira &amp; Rosa, 2020" authorityName="Vaz &amp; Silveira &amp; Rosa" authorityYear="2020" box="[453,546,1050,1075]" class="Insecta" family="Lampyridae" genus="Psilocladus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="costae" status="sp. nov.">
<emphasis id="D1DDEAEAFFD8FFA2FE1BFB89753BFB90" box="[453,546,1050,1075]" italics="true" pageId="11" pageNumber="12">P. costae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAEE5791FFD8FFA2FDF4FB897598FB90" box="[554,641,1050,1075]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
is also more similar to the pupae of the photurine species than to pupae of other Neotropical species belonging to the genera 
<taxonomicName id="24A94D7BFFD8FFA2FF6BFBE97699FB30" authority="Laporte" authorityName="Laporte" box="[181,384,1146,1171]" class="Insecta" family="Lampyridae" genus="Aspisoma" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="D1DDEAEAFFD8FFA2FF6BFBE97607FB30" box="[181,286,1146,1171]" italics="true" pageId="11" pageNumber="12">Aspisoma</emphasis>
Laporte
</taxonomicName>
, 
<taxonomicName id="24A94D7BFFD8FFA2FE52FBE9753EFB30" authority="Laporte" authorityName="Laporte" box="[396,551,1146,1171]" class="Insecta" family="Lampyridae" genus="Lucio" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="D1DDEAEAFFD8FFA2FE52FBE976DCFB30" box="[396,453,1146,1171]" italics="true" pageId="11" pageNumber="12">Lucio</emphasis>
Laporte
</taxonomicName>
, 
<taxonomicName id="24A94D7BFFD8FFA2FDEDFBE977F0FB10" authority="Motschulsky" authorityName="Motschulsky" class="Insecta" family="Lampyridae" genus="Cratomorphus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="D1DDEAEAFFD8FFA2FDEDFBE975D7FB30" box="[563,718,1146,1171]" italics="true" pageId="11" pageNumber="12">Cratomorphus</emphasis>
Motschulsky
</taxonomicName>
and 
<taxonomicName id="24A94D7BFFD8FFA2FEF5FB09750AFB10" authority="Olivier." authorityName="Olivier." box="[299,531,1178,1203]" class="Insecta" family="Lampyridae" genus="Pyractonema" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="genus">
<emphasis id="D1DDEAEAFFD8FFA2FEF5FB0976A3FB10" box="[299,442,1178,1203]" italics="true" pageId="11" pageNumber="12">Pyractonema</emphasis>
Olivier.
</taxonomicName>
<taxonomicName id="24A94D7BFFD8FFA2FDC3FB097581FB10" authorityName="Lacordaire" authorityYear="1857" box="[541,664,1178,1203]" class="Insecta" family="Lampyridae" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="subFamily" subFamily="Photurinae">Photurinae</taxonomicName>
and 
<taxonomicName id="24A94D7BFFD8FFA2FF80FB2977A4FB70" authority="Vaz &amp; Silveira &amp; Rosa, 2020" authorityName="Vaz &amp; Silveira &amp; Rosa" authorityYear="2020" box="[94,189,1210,1235]" class="Insecta" family="Lampyridae" genus="Psilocladus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="costae" status="sp. nov.">
<emphasis id="D1DDEAEAFFD8FFA2FF80FB2977A4FB70" box="[94,189,1210,1235]" italics="true" pageId="11" pageNumber="12">P. costae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAEE5791FFD8FFA2FF16FB297638FB70" box="[200,289,1210,1235]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
pupae share the presence of dense, long and stout setae on pronotal margins and posteri- or angles of abdominal terga (
<figureCitation id="7B922A7DFFD8FFA2FBB4FF6973AEFEB0" box="[1130,1207,250,275]" captionStart="Figure 13" captionStartId="12.[170,229,1198,1223]" captionTargetBox="[174,1491,210,1163]" captionTargetId="figure-1@12.[170,1493,204,1168]" captionTargetPageId="12" captionText="Figure 13. Psilocladus costae sp.nov.Pupal habitus.(A) Dorsal;(B) lateral;(C) ventral.Scale bar:= 2 mm." figureDoi="http://doi.org/10.5281/zenodo.4985605" httpUri="https://zenodo.org/record/4985605/files/figure.png" pageId="11" pageNumber="12">Fig. 13</figureCitation>
). 
<taxonomicName id="24A94D7BFFD8FFA2FB16FF697290FEB0" authority="Vaz &amp; Silveira &amp; Rosa, 2020" authorityName="Vaz &amp; Silveira &amp; Rosa" authorityYear="2020" box="[1224,1417,250,275]" class="Insecta" family="Lampyridae" genus="Psilocladus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="costae" status="sp. nov.">
<emphasis id="D1DDEAEAFFD8FFA2FB16FF697290FEB0" box="[1224,1417,250,275]" italics="true" pageId="11" pageNumber="12">Psilocladus costae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAEE5791FFD8FFA2FCC4FE897477FE90" box="[794,878,282,307]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
pupa differs by having photophores on abdominal segments VI and VII (
<figureCitation id="7B922A7DFFD8FFA2FBFDFEA97364FEF0" box="[1059,1149,314,339]" captionStart="Figure 14" captionStartId="12.[170,229,2106,2131]" captionTargetBox="[170,1493,1245,2076]" captionTargetId="figure-22@12.[170,1493,1245,2076]" captionTargetPageId="12" captionText="Figure 14. Psilocladus costae sp.nov.Immature bioluminescence.(A) Larva (dorsal);(B) pupa (ventral).Length:larva = 18 mm;pupa = 7 mm." figureDoi="http://doi.org/10.5281/zenodo.4985609" httpUri="https://zenodo.org/record/4985609/files/figure.png" pageId="11" pageNumber="12">Fig. 14B</figureCitation>
), whereas photurine pupae have only one pair of photophores on segment VIII.
</paragraph>
</subSubSection>
<caption id="B7D66670FFD9FFA3FF74F78974ECF7F0" ID-DOI="http://doi.org/10.5281/zenodo.4985599" ID-Zenodo-Dep="4985599" httpUri="https://zenodo.org/record/4985599/files/figure.png" pageId="10" pageNumber="11" startId="10.[170,229,2074,2099]" targetBox="[176,1489,864,2043]" targetPageId="10">
<paragraph id="E31636F8FFD9FFA3FF74F78974ECF7F0" blockId="10.[170,1494,2074,2131]" pageId="10" pageNumber="11">
Figure 11. 
<taxonomicName id="24A94D7BFFD9FFA3FED3F7897684F790" authority="Vaz &amp; Silveira &amp; Rosa, 2020" authorityName="Vaz &amp; Silveira &amp; Rosa" authorityYear="2020" box="[269,413,2074,2099]" class="Insecta" family="Lampyridae" genus="Psilocladus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="costae" status="sp. nov.">
<emphasis id="D1DDEAEAFFD9FFA3FED3F7897684F790" bold="true" box="[269,413,2074,2099]" italics="true" pageId="10" pageNumber="11">Psilocladus costae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAEE5791FFD9FFA3FE7DF78976FBF790" box="[419,482,2074,2099]" pageId="10" pageNumber="11" rank="species">sp. nov.</taxonomicNameLabel>
Larval morphology. (A-C) Head (dorsal, lateral, ventral); (D) right anterior, middle and posterior legs (from top to bottom, ventral);(E) left laterotergites VI-VII;(F) abdominal terga VI-IX.Scale bars:A-C,E, F = 1 mm;D = 2 mm.
</paragraph>
</caption>
<subSubSection id="ABB36573FFD8FFA4FCC4FE097627FD50" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" type="etymology">
<paragraph id="E31636F8FFD8FFA2FCC4FE09749BFDB0" blockId="11.[794,1418,410,531]" pageId="11" pageNumber="12">
<emphasis id="D1DDEAEAFFD8FFA2FCC4FE0974B8FE10" bold="true" box="[794,929,410,435]" pageId="11" pageNumber="12">Etymology:</emphasis>
The specific epithet is in honor to Professor Dr. Cleide Costa who dedicated many studies to immature beetles, especially Elateroidea. Singular genitive, feminine.
</paragraph>
<paragraph id="E31636F8FFD8FFA4FCC4FDA97627FD50" blockId="11.[794,1418,570,1235]" lastBlockId="13.[94,719,218,755]" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12">
<emphasis id="D1DDEAEAFFD8FFA2FCC4FDA973B6FDF0" bold="true" box="[794,1199,570,595]" pageId="11" pageNumber="12">Field and laboratory observations:</emphasis>
We collected 
<specimenCount id="F5AFFD71FFD8FFA2FA9BFDA97458FDD0" pageId="11" pageNumber="12" type="larva">49 larvae</specimenCount>
inside fallen epiphytic bromeliads. Few larvae were collected outside of bromeliads during the night when they were localized by their very bright bioluminescence (
<figureCitation id="7B922A7DFFD8FFA2FCFEFD297498FD70" box="[800,897,698,723]" captionStart="Figure 14" captionStartId="12.[170,229,2106,2131]" captionTargetBox="[170,1493,1245,2076]" captionTargetId="figure-22@12.[170,1493,1245,2076]" captionTargetPageId="12" captionText="Figure 14. Psilocladus costae sp.nov.Immature bioluminescence.(A) Larva (dorsal);(B) pupa (ventral).Length:larva = 18 mm;pupa = 7 mm." figureDoi="http://doi.org/10.5281/zenodo.4985609" httpUri="https://zenodo.org/record/4985609/files/figure.png" pageId="11" pageNumber="12">Fig. 14A</figureCitation>
). Those larvae were identified by comparison with the bromeliad larvae. Of the 61 fallen bromeliads investigated, 26 had larvae of 
<taxonomicName id="24A94D7BFFD8FFA2FBB6FD697230FCB0" authority="Vaz &amp; Silveira &amp; Rosa, 2020" authorityName="Vaz &amp; Silveira &amp; Rosa" authorityYear="2020" box="[1128,1321,762,787]" class="Insecta" family="Lampyridae" genus="Psilocladus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="costae" status="sp. nov.">
<emphasis id="D1DDEAEAFFD8FFA2FBB6FD697230FCB0" box="[1128,1321,762,787]" italics="true" pageId="11" pageNumber="12">Psilocladus costae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="CAEE5791FFD8FFA2FAEEFD69729EFCB0" box="[1328,1415,762,787]" pageId="11" pageNumber="12" rank="species">sp. nov.</taxonomicNameLabel>
, the majority of which found in the innermost leaves of the bromeliad rosette. The number of larvae inside each bromeliad ranged from one to five, but in most of them (n = 16) we found only 
<specimenCount id="F5AFFD71FFD8FFA2FBF2FCE97381FC30" box="[1068,1176,890,915]" pageId="11" pageNumber="12" type="larva">one larva</specimenCount>
. They were found inside bromeliads throughout the year (
<specimenCount id="F5AFFD71FFD8FFA2FB02FC097251FC10" box="[1244,1352,922,947]" pageId="11" pageNumber="12" type="larva">22 larvae</specimenCount>
from April to August and 
<specimenCount id="F5AFFD71FFD8FFA2FC2BFC297340FC70" box="[1013,1113,954,979]" pageId="11" pageNumber="12" type="larva">21 larvae</specimenCount>
from September to March), whilst active glowing larvae outside of bromeliads were found in the beginning of the rainy season (
<specimenCount id="F5AFFD71FFD8FFA2FAC3FC697290FBB0" box="[1309,1417,1018,1043]" pageId="11" pageNumber="12" type="larva">six larvae</specimenCount>
in October and November). At night larvae were found by their intense glow near bromeliad tanks, above leaf blades (
<figureCitation id="7B922A7DFFD8FFA2FCAFFBC974D7FBD0" box="[881,974,1114,1139]" captionStart="Figure 14" captionStartId="12.[170,229,2106,2131]" captionTargetBox="[170,1493,1245,2076]" captionTargetId="figure-22@12.[170,1493,1245,2076]" captionTargetPageId="12" captionText="Figure 14. Psilocladus costae sp.nov.Immature bioluminescence.(A) Larva (dorsal);(B) pupa (ventral).Length:larva = 18 mm;pupa = 7 mm." figureDoi="http://doi.org/10.5281/zenodo.4985609" httpUri="https://zenodo.org/record/4985609/files/figure.png" pageId="11" pageNumber="12">Fig. 14A</figureCitation>
) and on trunks and branches near bromeliads fixed on trees. Larvae emit green light, easily seen from above through the vitreous spots on tergum VIII when the larvae are stretched, as well as through vitreous spots on tergum VII, that covers the VIII when larvae shrink. In the lab, larvae preyed on termites and scirtid larvae. Mortality was very high and only two firefly larvae molted into pupa and adult, therefore we could not determine the number of larval instars. Larval length ranged from 
<quantity id="24519B1DFFDEFFA4FF2BFEE97662FE30" box="[245,379,378,403]" metricMagnitude="-2" metricUnit="m" metricValue="1.55" metricValueMax="2.2" metricValueMin="0.9" pageId="13" pageNumber="14" unit="mm" value="15.5" valueMax="22.0" valueMin="9.0">9 to 22 mm</quantity>
and the larvae which molted into pupa and adult were 
<emphasis id="D1DDEAEAFFDEFFA4FE5AFE0976BAFE10" box="[388,419,410,435]" italics="true" pageId="13" pageNumber="14">ca.</emphasis>
<quantity id="24519B1DFFDEFFA4FE74FE0976E0FE10" box="[426,505,410,435]" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="14" unit="mm" value="15.0">15 mm</quantity>
long, therefore we infer that mature larva may be 
<quantity id="24519B1DFFDEFFA4FE6CFE29753AFE70" box="[434,547,442,467]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="13" pageNumber="14" unit="mm" value="18.5" valueMax="22.0" valueMin="15.0">15-22 mm</quantity>
long. A few larvae possibly died due to acari infestation, but the causes of overall mortality were not known. Adults were observed on wet and dark nights in December and January. They flew at about 10 meters height whilst glowing green continuously, rarely below 5 meters, yet 
<specimenCount id="F5AFFD71FFDEFFA4FDB4FDC97782FD30" pageId="13" pageNumber="14" type="generic">10 specimens</specimenCount>
could be collected on the grass and on leaf blades of bromeliads fixed lower on the trees. The flying adults concentrated in areas with higher density of canopy bromeliads (
<figureCitation id="7B922A7DFFDEFFA4FF1FFD49762AFD50" box="[193,307,730,755]" captionStart="Figure 1" captionStartId="2.[869,928,1626,1651]" captionTargetBox="[869,1493,746,1596]" captionTargetId="figure-512@2.[869,1493,746,1596]" captionTargetPageId="2" captionText="Figure 1. Psilocladus costae sp.nov.Habitat at Campos do Jordão,São Paulo. (A) Legal reserve along deforested area; (B, C) canopy bromeliads; (D) fallen bromeliads Vrieseabituminosa." figureDoi="http://doi.org/10.5281/zenodo.4985577" httpUri="https://zenodo.org/record/4985577/files/figure.png" pageId="13" pageNumber="14">Figs. 1B, C</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>