treatments-xml/data/51/8B/85/518B854C2FC85CBDA694CB3BCA1CBA86.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1147.93824" ID-Pensoft-Pub="1313-2970-1147-1" ID-Pensoft-UUID="FF2B6B317B505F9EA0E1000BDCD16CE7" ID-ZooBank="9BB9881B0076473D8E53155D37CA1F50" ModsDocID="1313-2970-1147-1" checkinTime="1676378625502" checkinUser="pensoft" docAuthor="Pall-Gergely, Barna, Hunyadi, Andras, Vermeulen, Jaap J., Grego, Jozef, Sutcharit, Chirasak, Reischuetz, Alexander, Dumrongrojwattana, Pongrat, Botta-Dukat, Zoltan, Oerstan, Aydin, Fekete, Judit &amp; Jochum, Adrienne" docDate="2023" docId="518B854C2FC85CBDA694CB3BCA1CBA86" docLanguage="en" docName="ZooKeys 1147: 1-177" docOrigin="ZooKeys 1147" docPubDate="2023-02-13" docSource="http://dx.doi.org/10.3897/zookeys.1147.93824" docTitle="Angustopila apokritodon Pall-Gergely &amp; Hunyadi 2023, sp. nov." docType="treatment" docUuid="A9BB647B-6D1B-4FB1-BD26-0644957B9A7D" docUuidSource="ZooBank" docVersion="1" id="FF2B6B317B505F9EA0E1000BDCD16CE7" lastPageNumber="1" masterDocId="FF2B6B317B505F9EA0E1000BDCD16CE7" masterDocTitle="Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)" masterLastPageNumber="177" masterPageNumber="1" pageNumber="1" updateTime="1676378625502" updateUser="pensoft">
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<mods:title>Five times over: 42 new Angustopila species highlight Southeast Asia's rich biodiversity (Gastropoda, Stylommatophora, Hypselostomatidae)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Pall-Gergely, Barna</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6167-7221</mods:nameIdentifier>
<mods:affiliation>Centre for Agricultural Research, Plant Protection Institute, Eoetvoes Lorand Research Network, Herman Otto ut 15, H- 1022 Budapest, Hungary</mods:affiliation>
<mods:nameIdentifier type="email">pallgergely2@gmail.com</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hunyadi, Andras</mods:namePart>
<mods:affiliation>Adria setany 10 G 2 / 5., H- 1148 Budapest, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Vermeulen, Jaap J.</mods:namePart>
<mods:affiliation>JK Art and Science, Lauwerbes 8, 2318 AT Leiden, Netherlands</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grego, Jozef</mods:namePart>
<mods:affiliation>Horna Micina 219, SK- 97401 Banska Bystrica, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Sutcharit, Chirasak</mods:namePart>
<mods:affiliation>Animal Systematic Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Reischuetz, Alexander</mods:namePart>
<mods:affiliation>Puechhaimgasse 52, A- 3580 Horn, Austria</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Dumrongrojwattana, Pongrat</mods:namePart>
<mods:affiliation>Department of Biology, Faculty of Science, Burapha University, 169 Longhardbangsaen Road, Muang District, Chonburi, 20131, Thailand</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Botta-Dukat, Zoltan</mods:namePart>
<mods:affiliation>Centre for Ecological Research, Institute of Ecology and Botany, Alkotmany 2 - 4, H- 2600, Vacratot, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Oerstan, Aydin</mods:namePart>
<mods:affiliation>12501 Milestone Manor Lane, Germantown, Maryland, 20876, USA</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Fekete, Judit</mods:namePart>
<mods:affiliation>University of Pannonia, Centre of Natural Science, Research Group of Limnology, Egyetem u. 10, H- 8200 Veszprem, Hungary &amp; Centre for Ecological Research, Institute of Aquatic Ecology, Department of Tisza Research, 18 / c Bem square, H- 4026 Debrecen, Hungary</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Jochum, Adrienne</mods:namePart>
<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-6624-6412</mods:nameIdentifier>
<mods:affiliation>Naturhistorisches Museum der Burgergemeinde Bern, CH- 3005 Bern, Switzerland &amp; Institute of Ecology and Evolution, University of Bern, CH- 3012 Bern, Switzerland &amp; Senckenberg Forschungsinstitut und Naturmuseum, 60325 Frankfurt am Main, Germany</mods:affiliation>
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<mods:date>2023</mods:date>
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<mods:number>2023-02-13</mods:number>
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<mods:detail type="volume">
<mods:number>1147</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1147.93824</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1147.93824</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:A9BB647B-6D1B-4FB1-BD26-0644957B9A7D" httpUri="http://treatment.plazi.org/id/518B854C2FC85CBDA694CB3BCA1CBA86" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="https://zoobank.org/A9BB647B-6D1B-4FB1-BD26-0644957B9A7D" authority="Páll-Gergely &amp; Hunyadi" authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Angustopila apokritodon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="apokritodon" status="sp. nov.">
Angustopila apokritodon 
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi
</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">
<figureCitation captionStart="Figure 87" captionStartId="F87" captionText="Figure 87. Angustopila apokritodon Pall-Gergely &amp; Hunyadi, sp. nov. A paratype B-I holotype (HNHM 105317). Apertural (A, B), lateral (C), apical (D) and ventral (E) sides of the shell; sculpture of the protoconch with protoconch-teleoconch boundary (F); aperture (G); ventral (H) and frontal (I) surface of the body whorl." figureDoi="10.3897/zookeys.1147.93824.figure87" httpUri="https://binary.pensoft.net/fig/808896" pageId="0" pageNumber="1">Fig. 87</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type material">
<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: Vietnam • 1 empty shell (H: 0.87 mm, D: 0.81 mm); 
<normalizedToken originalValue="Sơn">Sơn</normalizedToken>
La Province, Quỳnh Nhai district, 20 km north from crossing point to Thuận 
<normalizedToken originalValue="Châu">Chau</normalizedToken>
, Chiềng Khoang, cave above the village (locality code: 2020/9); 
<geoCoordinate degrees="21" direction="north" minutes="33.44" orientation="latitude" precision="9" value="21.557333">21°33.44'N</geoCoordinate>
, 
<geoCoordinate degrees="103" direction="east" minutes="40.91" orientation="longitude" precision="9" value="103.68183">103°40.91'E</geoCoordinate>
; 315 m a.s.l.; 7 Feb. 2020; A. Hunyadi leg.; HNHM 105317.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
: Vietnam • 4 shells; same data as for holotype; coll. HA.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
A medium-sized, conical 
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
</taxonomicName>
species with three prominent apertural barriers (long parietal tooth reaching parietal callus, prominent lower palatal tooth, prominent subcolumellar tooth).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">Shell of normal size for the genus, slightly higher than wide; off-white, translucent; conical with regularly growing whorls; body whorl widest in standard apertural view; protoconch consists of 1.5 whorls, with slight spiral striation on the last protoconch whorl; teleoconch with some strong to weak radial striae, occasionally grouped in multiple thread-like lines between stronger radial growth lines and spiral striae (ca. 12-14 on body whorl from standard apertural view) of comparable strength, resulting in a reticulated sculpture; whorls 4.25-4.5, rounded; aperture slightly oblique to shell axis in lateral view; umbilicus wide; aperture broad piriform-subquadrate with straight concave parietal part; peristome expanded, not reflected; parietal callus not detached from penultimate whorl; parietal tooth large and straight, directed towards lower palatal tooth; anterior to large parietal tooth (on the parietal callus) the callus is supported with a whitish triangular thickening facing to the front; palatal tooth situated on lower part of palatal wall (probably homologous with the lower palatal tooth), small, denticle-like, sits close to basal lip; subcolumellar tooth situated at junction of columellar and basal positions, similar in size and alignment to the peristome edge and in line with the palatal tooth.</paragraph>
<caption doi="10.3897/zookeys.1147.93824.figure87" httpUri="https://binary.pensoft.net/fig/808896" pageId="0" pageNumber="1" start="Figure 87" startId="F87">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Figure 87.</emphasis>
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila apokritodon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="apokritodon">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila apokritodon</emphasis>
</taxonomicName>
<normalizedToken originalValue="Páll-Gergely">Pall-Gergely</normalizedToken>
&amp; Hunyadi, sp. nov. 
<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
paratype 
<emphasis bold="true" pageId="0" pageNumber="1">B-I</emphasis>
holotype (HNHM 105317). Apertural (
<emphasis bold="true" pageId="0" pageNumber="1">A, B</emphasis>
), lateral (
<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
), apical (
<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
) and ventral (
<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
) sides of the shell; sculpture of the protoconch with protoconch-teleoconch boundary (
<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
); aperture (
<emphasis bold="true" pageId="0" pageNumber="1">G</emphasis>
); ventral (
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
) and frontal (
<emphasis bold="true" pageId="0" pageNumber="1">I</emphasis>
) surface of the body whorl.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Measurements (in mm).</paragraph>
<paragraph pageId="0" pageNumber="1">
H = 0.87-0.92, D = 0.81-0.86, H/D*100 = 101.2-109.6 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 4), RUD = 30.5-30.7 (
<emphasis italics="true" pageId="0" pageNumber="1">n</emphasis>
= 2).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
No other 
<taxonomicName class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila</emphasis>
</taxonomicName>
species shows similar arrangement of apertural barriers (parietal + lower palatal + subcolumellar), and the thick callus anterior to the parietal tooth has not been observed in any other congeners. Maybe 
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila tetradon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tetradon">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila tetradon</emphasis>
</taxonomicName>
sp. nov. could be considered as being most similar, but it has an additional upper palatal tooth and a lower conical shell shape. Furthermore, 
<taxonomicName lsidName="A. rara" pageId="0" pageNumber="1" rank="species" species="rara">
<emphasis italics="true" pageId="0" pageNumber="1">A. rara</emphasis>
</taxonomicName>
sp. nov. has a narrower umbilicus, a wider subcolumellar tooth, and a more apically situated palatal tooth.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
The specific epithet 
<taxonomicName authorityName="Pall-Gergely &amp; Hunyadi" authorityYear="2023" class="Gastropoda" family="Gastrocoptidae" genus="Angustopila" kingdom="Animalia" lsidName="Angustopila apokritodon" order="Stylommatophora" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="apokritodon">
<emphasis italics="true" pageId="0" pageNumber="1">Angustopila apokritodon</emphasis>
</taxonomicName>
derives from the Greek words 
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="ἀπόκριτος">ἀπόκριτος</normalizedToken>
</emphasis>
(= separated) and 
<emphasis italics="true" pageId="0" pageNumber="1">
<normalizedToken originalValue="ὀδούς">ὀδούς</normalizedToken>
</emphasis>
(= tooth) referring to the large distance between the palatal and subcolumellar teeth.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">
This species is only known from the type locality in 
<normalizedToken originalValue="Sơn">Sơn</normalizedToken>
La Province in Vietnam (Fig. 
<figureCitation captionStart="Figure 27" captionStartId="F27" captionText="Figure 27. Distribution of Angustopila species." figureDoi="10.3897/zookeys.1147.93824.figure27" httpUri="https://binary.pensoft.net/fig/808836" pageId="0" pageNumber="1">27</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
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