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<document id="CFCF96B7E7D3E1CEC5A5A34747D7D8B6" ID-DOI="10.11646/zootaxa.4020.3.9" ID-GBIF-Dataset="ff6b61bc-bb45-4436-9d88-8f1c678daf06" ID-ISSN="1175-5326" ID-Zenodo-Dep="238547" ID-ZooBank="A254E350-F0C8-495D-A003-43EAF1512B0F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461271511584" checkinUser="plazi" docAuthor="De Matos-Pita, Susana S. &amp; Ramil, Fran" docDate="2015" docId="002487F22013FFD7FF142B0EFCDEFBBE" docLanguage="en" docName="zt04020p587.pdf" docOrigin="Zootaxa 4020 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ezaxius Matos-Pita &amp; Ramil, 2015, n. gen." docType="treatment" docVersion="7" lastPageNumber="579" masterDocId="FC1DFF8A2014FFDFFF83282EFFCCFFE6" masterDocTitle="Additions to thalassinidean fauna (Crustacea: Decapoda) off Mauritania (NW Africa) with the description of a new genus and a new species" masterLastPageNumber="587" masterPageNumber="571" pageNumber="578" updateTime="1698611913965" updateUser="plazi">
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<mods:title id="77217F4BAAA8D3CE43AFDD30DC0C1DB8">Additions to thalassinidean fauna (Crustacea: Decapoda) off Mauritania (NW Africa) with the description of a new genus and a new species</mods:title>
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<mods:namePart id="FD96CFB41FEB623311E1325BEFCD6B0A">De Matos-Pita, Susana S.</mods:namePart>
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<mods:namePart id="47F6BB4B584C0311734EA8814A4A3897">Ramil, Fran</mods:namePart>
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<treatment id="002487F22013FFD7FF142B0EFCDEFBBE" ID-DOI="http://doi.org/10.5281/zenodo.6108165" ID-GBIF-Taxon="119638868" ID-Zenodo-Dep="6108165" LSID="urn:lsid:plazi:treatment:002487F22013FFD7FF142B0EFCDEFBBE" httpUri="http://treatment.plazi.org/id/002487F22013FFD7FF142B0EFCDEFBBE" lastPageId="8" lastPageNumber="579" pageId="7" pageNumber="578">
<subSubSection id="C097656F2013FFD8FF142B0EFE6BFCDC" box="[151,423,800,826]" pageId="7" pageNumber="578" type="nomenclature">
<paragraph id="883236E42013FFD8FF142B0EFE6BFCDC" blockId="7.[151,423,800,826]" box="[151,423,800,826]" pageId="7" pageNumber="578">
<heading id="D37A81882013FFD8FF142B0EFE6BFCDC" bold="true" box="[151,423,800,826]" fontSize="11" level="1" pageId="7" pageNumber="578" reason="1">
<emphasis id="BAF9EAF62013FFD8FF142B0EFE6BFCDC" bold="true" box="[151,423,800,826]" pageId="7" pageNumber="578">
Genus
<taxonomicName id="4F8D4D672013FFD8FF6D2B0EFE82FCDC" box="[238,334,800,826]" class="Malacostraca" family="Axiidae" genus="Ezaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="BAF9EAF62013FFD8FF6D2B0EFE82FCDC" bold="true" box="[238,334,800,826]" italics="true" pageId="7" pageNumber="578">Ezaxius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1CA578D2013FFD8FED62B0EFE6BFCDC" box="[341,423,800,826]" pageId="7" pageNumber="578" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C097656F2013FFD8FF142B48FB09FAC9" pageId="7" pageNumber="578" type="diagnosis">
<paragraph id="883236E42013FFD8FF142B48FB09FAC9" blockId="7.[151,1437,870,1975]" pageId="7" pageNumber="578">
<emphasis id="BAF9EAF62013FFD8FF142B48FEDDFC99" bold="true" box="[151,273,870,895]" pageId="7" pageNumber="578">Diagnosis.</emphasis>
Carapace faintly pitted. Rostrum triangular with upturned tip, laterally spinose, extending posteriorly as lateral carinae. Gastric region elevated, bearing 5 carinae: median, submedian and short lateral; submedian carinae converging anteriorly to form a horseshoe-like shape. No linea thalassinica. Cervical groove distinct only on dorsal surface, postcervical carina wide and short, weakly visible only over posterior median lobe. Eyes short, unpigmented, one-third rostrum length. Antennal acicle elongate, protruding straight forward. Anterolateral margin broadly rounded, unarmed. Maxilla 2 exopod posterior lobe with one long seta. Maxillipeds 13 with exopod and single epipod; maxilliped 2 with single podobranchia; maxilliped 3 with double podobranchia and single arthrobranchia. Pereiopods 1 and 2 chelate, pereiopods 14 with single epipod and double arthrobranchia, pereiopods 13 with double podobranchia. Pleurobranchia absent. Pleomeres unarmed; first somite with pleura acute; second broad, anteriorly and posteriorly rounded; 35 anteriorly rounded, declining to a rounded angle posteriorly; pleura on somite 6 rounded. Pleopod 1 uniramous, unsegmented; pleopods 25 alike, slender, with appendix interna; pleopod 2 without appendix masculina. Uropod exopod with spinose transverse suture. Telson longer than wide, with posterior margin rounded, without posteromedian tooth. Hermaphrodite.
</paragraph>
</subSubSection>
<subSubSection id="C097656F2013FFD8FF442D14FC28FAB5" box="[199,996,1338,1363]" pageId="7" pageNumber="578" type="materials_examined">
<paragraph id="883236E42013FFD8FF442D14FC28FAB5" blockId="7.[151,1437,870,1975]" box="[199,996,1338,1363]" pageId="7" pageNumber="578">
<emphasis id="BAF9EAF62013FFD8FF442D14FEAEFAB5" bold="true" box="[199,354,1338,1363]" pageId="7" pageNumber="578">
<typeStatus id="573688462013FFD8FF442D14FECEFAB5" box="[199,258,1338,1363]" pageId="7" pageNumber="578">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="4F8D4D672013FFD8FEEA2D12FD89FAB5" box="[361,581,1338,1363]" class="Malacostraca" family="Axiidae" genus="Ezaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="species" species="ferachevali" status="sp. nov.">
<emphasis id="BAF9EAF62013FFD8FEEA2D12FD89FAB5" box="[361,581,1338,1363]" italics="true" pageId="7" pageNumber="578">Ezaxius ferachevali</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1CA578D2013FFD8FDCF2D15FD1AFAB5" box="[588,726,1338,1363]" pageId="7" pageNumber="578" rank="species">new species</taxonomicNameLabel>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="C097656F2013FFD8FF442D70FBFEFA7D" pageId="7" pageNumber="578" type="etymology">
<paragraph id="883236E42013FFD8FF442D70FBFEFA7D" blockId="7.[151,1437,870,1975]" pageId="7" pageNumber="578">
<emphasis id="BAF9EAF62013FFD8FF442D70FE82FA91" bold="true" box="[199,334,1374,1399]" pageId="7" pageNumber="578">Etymology.</emphasis>
Generic name is a combination of the name
<emphasis id="BAF9EAF62013FFD8FCC32D71FCA6FA90" box="[832,874,1375,1398]" italics="true" pageId="7" pageNumber="578">Eza</emphasis>
from the Spanish R/V Vizconde de Eza, on which the Maurit surveys were conducted, and the genus name
<taxonomicName id="4F8D4D672013FFD8FC902DADFC9DFA7C" box="[787,849,1411,1434]" class="Malacostraca" family="Axiidae" genus="Axius" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="genus">
<emphasis id="BAF9EAF62013FFD8FC902DADFC9DFA7C" box="[787,849,1411,1434]" italics="true" pageId="7" pageNumber="578">Axius</emphasis>
</taxonomicName>
. Gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="C097656F2013FFD7FF442D88FCDEFBBE" lastPageId="8" lastPageNumber="579" pageId="7" pageNumber="578" type="discussion">
<paragraph id="883236E42013FFD8FF442D88FEBDF939" blockId="7.[151,1437,870,1975]" pageId="7" pageNumber="578">
<emphasis id="BAF9EAF62013FFD8FF442D88FEF7FA59" bold="true" box="[199,315,1446,1471]" pageId="7" pageNumber="578">Remarks.</emphasis>
<taxonomicName id="4F8D4D672013FFD8FEC52D89FE51FA58" box="[326,413,1447,1470]" class="Malacostraca" family="Axiidae" genus="Ezaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="BAF9EAF62013FFD8FEC52D89FE51FA58" box="[326,413,1447,1470]" italics="true" pageId="7" pageNumber="578">Ezaxius</emphasis>
</taxonomicName>
<emphasis id="BAF9EAF62013FFD8FE2A2D88FE30FA59" bold="true" box="[425,508,1446,1471]" pageId="7" pageNumber="578">
<taxonomicNameLabel id="A1CA578D2013FFD8FE2A2D88FE30FA59" box="[425,508,1446,1471]" pageId="7" pageNumber="578" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
, described here with pereiopod 2 chelate, should be included in the infraorder Axiidea/Callianassidea (see
<tableCitation id="C50F035F2013FFD8FE562DE4FDE4FA05" box="[469,552,1482,1507]" captionStart="TABLE 1" captionStartId="3.[151,239,152,175]" captionText="TABLE 1. The two main recent tendencies in the systematic arrangement of the thalassinidean families." pageId="7" pageNumber="578">Table 1</tableCitation>
). Within this infraorder, the following combination of characters separates the new genus from all except the
<taxonomicName id="4F8D4D672013FFD8FD832DC0FD95F9E1" box="[512,601,1518,1543]" class="Insecta" family="Axiidae" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="family">Axiidae</taxonomicName>
and related families (see
<tableCitation id="C50F035F2013FFD8FC052DC0FC16F9E1" box="[902,986,1518,1543]" captionStart="TABLE 1" captionStartId="3.[151,239,152,175]" captionText="TABLE 1. The two main recent tendencies in the systematic arrangement of the thalassinidean families." pageId="7" pageNumber="578">Table 1</tableCitation>
): the absence of linea thalassinica, the presence of a conspicuous rostrum and the presence of a long seta on maxilla 2 exopod posterior lobe. The presence of a uropodal exopod transverse suture, epipods on pereiopods 14 and an appendix masculina attached mesially on pleopod 2 endopod exclude the new genus from Sakai
<emphasis id="BAF9EAF62013FFD8FB862E72FB85F995" box="[1029,1097,1626,1651]" italics="true" pageId="7" pageNumber="578">et al.</emphasis>
families
<taxonomicName id="4F8D4D672013FFD8FB4C2E74FAADF995" box="[1231,1377,1626,1651]" class="Malacostraca" family="Axiidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="family">Eiconaxiidae</taxonomicName>
and
<taxonomicName id="4F8D4D672013FFD8FF142E50FEAAF971" box="[151,358,1662,1687]" class="Insecta" family="Axiidae" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="578" phylum="Arthropoda" rank="family">Eiconaxiopsididae</taxonomicName>
(both without uropodal suture), Coralaxiidae (lacking epipods on pereiopods 14) and from
<taxonomicName id="4F8D4D672013FFD8FF142E8CFEFFF95D" box="[151,307,1698,1723]" class="Malacostraca" family="Calocarididae" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="family">Calocarididae</taxonomicName>
(in which the appendix masculina is attached terminally). Therefore, we subsume
<taxonomicName id="4F8D4D672013FFD8FB512E8AFAE5F95D" box="[1234,1321,1700,1723]" class="Malacostraca" family="Axiidae" genus="Ezaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="578" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="BAF9EAF62013FFD8FB512E8AFAE5F95D" box="[1234,1321,1700,1723]" italics="true" pageId="7" pageNumber="578">Ezaxius</emphasis>
</taxonomicName>
<emphasis id="BAF9EAF62013FFD8FAB32E8CFAB3F95D" bold="true" box="[1328,1407,1698,1723]" pageId="7" pageNumber="578">
<taxonomicNameLabel id="A1CA578D2013FFD8FAB32E8CFAB3F95D" box="[1328,1407,1698,1723]" pageId="7" pageNumber="578" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
to the family Axiidea.
</paragraph>
<paragraph id="883236E42013FFD8FF442EC4FDA5F851" blockId="7.[151,1437,870,1975]" pageId="7" pageNumber="578">
Since the first attempt to identify valuable characters to define axiid genera by
<bibRefCitation id="EC1C4B152013FFD8FBC82EC4FF62F8C1" author="Kensley" pageId="7" pageNumber="578" refString="Kensley, B. &amp; Gore, R. H. (1981) Coralaxius abelei, new genus and new species (Crustacea: Decapoda: Thalassinidea: Axiidae): a coral-inhabiting shrimp from the Florida Keys and the western Caribbean Sea. Proceedings of the Biological Society of Washington, 93, 1277 - 1294." type="journal article" year="1981">
Kensley &amp; Gore (1981:
<tableCitation id="C50F035F2013FFD8FADD2EC4FF68F8C1" captionStart="TABLE 1" captionStartId="3.[151,239,152,175]" captionText="TABLE 1. The two main recent tendencies in the systematic arrangement of the thalassinidean families." pageId="7" pageNumber="578">Table 1</tableCitation>
)
</bibRefCitation>
, the various authors have grouped the species mainly based on the morphology of the rostrum, length ratio rostrum/eyestalk, carapace carinae, antennal acicle, maxilliped 3, symmetry of the chelipeds, pleonal pleura, telson and uropods, as well as on the presence or absence of maxilla 2 posterior lobe long setae, epipods, exopods, branchiae, pleopods 1 and 2, appendix masculina, appendix interna and/or uropodal exopod transverse suture (e.g.
<bibRefCitation id="EC1C4B152013FFD8FF142FB1FED6F851" author="Poore" box="[151,282,1951,1975]" pageId="7" pageNumber="578" refString="Poore, G. C. B. (1994) A phylogeny of the families of Thalassinidea (Crustacea: Decapoda) with keys to families and genera. Memoirs of Museum Victoria, 54, 79 - 120." type="journal article" year="1994">Poore 1994</bibRefCitation>
;
<bibRefCitation id="EC1C4B152013FFD8FEA52FB1FE1CF851" author="Ngoc-Ho" box="[294,464,1951,1975]" pageId="7" pageNumber="578" refString="Ngoc-Ho, N. (2003) European and Mediterranean Thalassinidea (Crustacea, Decapoda). Zoosystema, 25 (3), 439 - 555." type="journal article" year="2003">Ngoc-Ho 2003</bibRefCitation>
;
<bibRefCitation id="EC1C4B152013FFD8FE5F2FB0FD91F851" author="Sakai" box="[476,605,1950,1975]" pageId="7" pageNumber="578" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). In: Fransen, C. H. J. M. &amp; Vaupel Klein, J. C. von (Eds.), Crustaceana Monographs. Vol. 13. Leiden, Brill, pp. 1 - 520. http: // dx. doi. org / 10.1163 / 9789047424185" type="book" year="2011">Sakai 2011</bibRefCitation>
).
</paragraph>
<paragraph id="883236E4201CFFD7FF4428B6FC17FD86" blockId="8.[151,1437,151,1112]" pageId="8" pageNumber="579">
Some of these characters must be taken with caution because they have been demonstrated to be ambivalent. Thus, although lateral and median carapace carinae is usually constant, the submedian carina can be poorly defined or blurred in different specimens of the same species, such as in
<taxonomicName id="4F8D4D67201CFFD7FCF428F1FBA5FF1E" box="[887,1129,223,248]" class="Malacostraca" family="Axiidae" genus="Calaxius" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="579" phylum="Arthropoda" rank="species" species="pitatucensis">
<emphasis id="BAF9EAF6201CFFD7FCF428F1FBA5FF1E" box="[887,1129,223,248]" italics="true" pageId="8" pageNumber="579">Calaxius pitatucensis</emphasis>
</taxonomicName>
(de
<bibRefCitation id="EC1C4B15201CFFD7FB1F28CEFAD4FF1E" author="Man" box="[1180,1304,223,248]" pageId="8" pageNumber="579" refString="Man, J. G. de (1925) The Decapoda of the Siboga-Expedition. Part VI. The Axiidae collected by the Siboga-Expedition. Siboga Expeditie, 39 a 5, 1 - 127." type="journal article" year="1925">
<collectingCountry id="F09A7674201CFFD7FB1F28CEFB1EFF11" box="[1180,1234,224,247]" name="Isle of Man" pageId="8" pageNumber="579">Man</collectingCountry>
, 1925
</bibRefCitation>
) (see
<bibRefCitation id="EC1C4B15201CFFD7FADD28F1FF1EFEFA" author="Sakai" pageId="8" pageNumber="579" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). In: Fransen, C. H. J. M. &amp; Vaupel Klein, J. C. von (Eds.), Crustaceana Monographs. Vol. 13. Leiden, Brill, pp. 1 - 520. http: // dx. doi. org / 10.1163 / 9789047424185" type="book" year="2011">Sakai 2011</bibRefCitation>
: 104; as
<taxonomicName id="4F8D4D67201CFFD7FEBB292AFDABFEFA" box="[312,615,260,285]" class="Malacostraca" family="Axiidae" genus="Colemanaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="579" phylum="Arthropoda" rank="species" species="pitatucensis">
<emphasis id="BAF9EAF6201CFFD7FEBB292AFDABFEFA" box="[312,615,260,285]" italics="true" pageId="8" pageNumber="579">Colemanaxius pitatucensis</emphasis>
</taxonomicName>
), in
<taxonomicName id="4F8D4D67201CFFD7FD1E292AFBE1FEFA" authority="Nobili, 1904" authorityName="Nobili" authorityYear="1904" box="[669,1069,260,285]" class="Malacostraca" family="Axiidae" genus="Allaxius" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="579" phylum="Arthropoda" rank="species" species="aethiopicus">
<emphasis id="BAF9EAF6201CFFD7FD1E292AFC4FFEFB" box="[669,899,260,285]" italics="true" pageId="8" pageNumber="579">Allaxius aethiopicus</emphasis>
(
<bibRefCitation id="EC1C4B15201CFFD7FC10292AFBE9FEFA" author="Nobili" box="[915,1061,260,285]" pageId="8" pageNumber="579" refString="Nobili, G. (1904) Diagnoses preliminaires de vingt-huit especes nouvelles de Stomatopodes et Decapodes Macroures de la Mer Rouge. Bulletin du Museum d'Histoire naturelle, Paris, 10 (5), 228 - 238." type="journal article" year="1904">Nobili, 1904</bibRefCitation>
)
</taxonomicName>
and in
<taxonomicName id="4F8D4D67201CFFD7FB07292AFE54FEA6" authority="A. Milne-Edwards, 1873" authorityName="A. Milne-Edwards" authorityYear="1873" class="Malacostraca" family="Axiidae" genus="Axiopsis" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="579" phylum="Arthropoda" rank="species" species="serratifrons">
<emphasis id="BAF9EAF6201CFFD7FB07292AFAA3FEFA" box="[1156,1391,260,284]" italics="true" pageId="8" pageNumber="579">Axiopsis serratifrons</emphasis>
(
<bibRefCitation id="EC1C4B15201CFFD7FA03292AFE5CFEA6" author="Milne-Edwards" pageId="8" pageNumber="579" refString="Milne-Edwards, A. (1873) Description de quelques Crustaces Nouveaux ou peu connus provenant du Musee de M. C. Godeffroy. Journal des Museum Godeffroy, 1 (4), 77 - 88 (253 - 264), pls. 12 - 13. http: // dx. doi. org / 10.5962 / bhl. title. 10644" type="journal article" year="1873">A. Milne-Edwards, 1873</bibRefCitation>
)
</taxonomicName>
(see
<bibRefCitation id="EC1C4B15201CFFD7FE582909FDACFEA6" author="Sakai" box="[475,608,295,320]" pageId="8" pageNumber="579" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). In: Fransen, C. H. J. M. &amp; Vaupel Klein, J. C. von (Eds.), Crustaceana Monographs. Vol. 13. Leiden, Brill, pp. 1 - 520. http: // dx. doi. org / 10.1163 / 9789047424185" type="book" year="2011">Sakai 2011</bibRefCitation>
: 41, 56, respectively); moreover, postcervical carina can be posteriorly distinct in larger specimens but indistinct in smaller mature specimens, such as in
<taxonomicName id="4F8D4D67201CFFD7FBFD2962FEF2FE6E" authority="Herbst, 1796" authorityName="Herbst" authorityYear="1796" class="Malacostraca" family="Axiidae" genus="Eutrichocheles" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="579" phylum="Arthropoda" rank="species" species="modestus">
<emphasis id="BAF9EAF6201CFFD7FBFD2962FA50FE83" box="[1150,1436,332,357]" italics="true" pageId="8" pageNumber="579">Eutrichocheles modestus</emphasis>
(
<bibRefCitation id="EC1C4B15201CFFD7FF1D2941FEF9FE6E" author="Herbst" box="[158,309,367,392]" pageId="8" pageNumber="579" refString="Herbst, J. F. W. (1796) Versuch einer Naturgeschichte der Krabben und Krebse nebst einer systematischen Beschreibung ihrer verschiedenen Arten. Vol. 2. Bei Gottlieb August Lange, Berlin und Stralsund, 225 pp., 25 pls. [pls. 22 - 46]" type="book chapter" year="1796">Herbst, 1796</bibRefCitation>
)
</taxonomicName>
(see
<bibRefCitation id="EC1C4B15201CFFD7FE032941FDC8FE6E" author="Sakai" box="[384,516,367,392]" pageId="8" pageNumber="579" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). In: Fransen, C. H. J. M. &amp; Vaupel Klein, J. C. von (Eds.), Crustaceana Monographs. Vol. 13. Leiden, Brill, pp. 1 - 520. http: // dx. doi. org / 10.1163 / 9789047424185" type="book" year="2011">Sakai 2011</bibRefCitation>
: 112). The presence or absence of pleopod 1 can also be confusing, mainly in species descriptions based on only one specimen, because of the absence of this appendage in young specimens of some species, such as
<taxonomicName id="4F8D4D67201CFFD7FE1A2997FDA1FE36" box="[409,621,439,464]" class="Malacostraca" family="Axiidae" genus="Paraxiopsis" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="579" phylum="Arthropoda" rank="species" species="brocki">
<emphasis id="BAF9EAF6201CFFD7FE1A2997FDA1FE36" box="[409,621,439,464]" italics="true" pageId="8" pageNumber="579">Paraxiopsis brocki</emphasis>
</taxonomicName>
(de
<bibRefCitation id="EC1C4B15201CFFD7FD272996FCEEFE36" author="Man" box="[676,802,440,464]" pageId="8" pageNumber="579" refString="Man, J. G. de (1888) Bericht uber die von Herrn Dr. J. Brock im indischen Archipel gesammelten Decapoden und Stomatopoden. Archiv fur Naturgeschichte, 53 (1), 215 - 600, pls. 7 - 22 a." type="journal article" year="1888">
<collectingCountry id="F09A7674201CFFD7FD272996FD16FE29" box="[676,730,440,463]" name="Isle of Man" pageId="8" pageNumber="579">Man</collectingCountry>
, 1888
</bibRefCitation>
), in which pleopod 1 is a segmented flagellum in large ovigerous females but lacking in small females (
<bibRefCitation id="EC1C4B15201CFFD7FD4F29F2FC51FE13" author="Sakai" box="[716,925,476,501]" pageId="8" pageNumber="579" refString="Sakai, K. (1992 b) Notes on some species of Thalassinidea from French Polynesia (Crustacea: Decapoda). Senckenbergiana maritima, 22 (3 / 6), 211 - 216." type="journal article" year="1992" yearSuffix="b">Sakai 1992b: 216</bibRefCitation>
). Variations in the armature of several body structures, including rostrum, gastric submedian carinae, telson, third maxilliped, chelipeds, second pereiopods, uropods and the length of the scaphocerite, were also observed between
<typeStatus id="57368846201CFFD7FC752A0AFB94FDDB" box="[1014,1112,548,573]" pageId="8" pageNumber="579" type="holotype">holotype</typeStatus>
and
<typeStatus id="57368846201CFFD7FB1B2A0BFB34FDDB" box="[1176,1272,549,573]" pageId="8" pageNumber="579" type="paratype">paratype</typeStatus>
specimens of
<emphasis id="BAF9EAF6201CFFD7FF142A67FE48FD86" box="[151,388,583,608]" italics="true" pageId="8" pageNumber="579">
<taxonomicName id="4F8D4D67201CFFD7FF142A67FEE5FD86" box="[151,297,585,608]" class="Malacostraca" family="Axiidae" genus="Formosaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="579" phylum="Arthropoda" rank="genus">Formosaxius</taxonomicName>
dorsum
</emphasis>
<bibRefCitation id="EC1C4B15201CFFD7FE082A69FD7DFD86" author="Komai" box="[395,689,583,608]" pageId="8" pageNumber="579" refString="Komai, T., Lin, F. - J. &amp; Chan, T. - Y. (2010) Five new species of Axiidae (Crustacea: Decapoda: Axiidea) from deep-water off Taiwan, with description of a new genus. Zootaxa, 2352, 1 - 28." type="journal article" year="2010">Komai, Lin &amp; Chan, 2010</bibRefCitation>
by
<bibRefCitation id="EC1C4B15201CFFD7FD582A69FC7EFD86" author="Komai" box="[731,946,583,608]" pageId="8" pageNumber="579" refString="Komai, T., Lin, F. - J. &amp; Chan, T. - Y. (2010) Five new species of Axiidae (Crustacea: Decapoda: Axiidea) from deep-water off Taiwan, with description of a new genus. Zootaxa, 2352, 1 - 28." type="journal article" year="2010">
Komai
<emphasis id="BAF9EAF6201CFFD7FCAD2A67FCAAFD86" box="[814,870,583,608]" italics="true" pageId="8" pageNumber="579">et al.</emphasis>
(2010
</bibRefCitation>
: 3).
</paragraph>
<paragraph id="883236E4201CFFD7FF442A42FCDEFBBE" blockId="8.[151,1437,151,1112]" pageId="8" pageNumber="579">
Our material does not share all the foregoing characters with the 19 axiid genera cited by
<bibRefCitation id="EC1C4B15201CFFD7FB442A43FA91FD62" author="Poore" box="[1223,1373,620,644]" pageId="8" pageNumber="579" refString="Poore, G. C. B. (1994) A phylogeny of the families of Thalassinidea (Crustacea: Decapoda) with keys to families and genera. Memoirs of Museum Victoria, 54, 79 - 120." type="journal article" year="1994">Poore (1994)</bibRefCitation>
, with the updated 21 axiid genera listed by
<bibRefCitation id="EC1C4B15201CFFD7FDC72AA1FC49FD4E" author="Komai" box="[580,901,655,680]" pageId="8" pageNumber="579" refString="Komai, T. &amp; Tachikawa, H. (2007) New genus and species of axiid shrimp (Crustacea, Decapoda, Thalassinidea) from Japan. Bulletin of the National Museum of Nature and Science, series A (Zoology), 33, 113 - 126." type="journal article" year="2007">Komai &amp; Tachikawa (2007)</bibRefCitation>
, with the 44 genera included in the
<taxonomicName id="4F8D4D67201CFFD7FA9C2AA1FAB4FD4E" box="[1311,1400,655,680]" class="Insecta" family="Axiidae" kingdom="Animalia" order="Lepidoptera" pageId="8" pageNumber="579" phylum="Arthropoda" rank="family">Axiidae</taxonomicName>
by
<bibRefCitation id="EC1C4B15201CFFD7FF142A9AFEE4FD2A" author="Sakai" box="[151,296,692,717]" pageId="8" pageNumber="579" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). In: Fransen, C. H. J. M. &amp; Vaupel Klein, J. C. von (Eds.), Crustaceana Monographs. Vol. 13. Leiden, Brill, pp. 1 - 520. http: // dx. doi. org / 10.1163 / 9789047424185" type="book" year="2011">Sakai (2011)</bibRefCitation>
, or with the features of the axiid genus
<taxonomicName id="4F8D4D67201CFFD7FD652A9BFB67FD2A" authority="Komai, Lin &amp; Chan, 2010" authorityName="Komai, Lin &amp; Chan" authorityYear="2010" box="[742,1195,692,717]" class="Malacostraca" family="Axiidae" genus="Formosaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="BAF9EAF6201CFFD7FD652A9BFCB5FD2A" box="[742,889,693,716]" italics="true" pageId="8" pageNumber="579">Formosaxius</emphasis>
<bibRefCitation id="EC1C4B15201CFFD7FCFC2A9AFB67FD2A" author="Komai" box="[895,1195,692,717]" pageId="8" pageNumber="579" refString="Komai, T., Lin, F. - J. &amp; Chan, T. - Y. (2010) Five new species of Axiidae (Crustacea: Decapoda: Axiidea) from deep-water off Taiwan, with description of a new genus. Zootaxa, 2352, 1 - 28." type="journal article" year="2010">Komai, Lin &amp; Chan, 2010</bibRefCitation>
</taxonomicName>
. A triangular rostrum with lateral teeth continuous with lateral carinae, the presence of five gastric anterior carinae, the elongate antennal acicle, pleopods 35 endopods with appendix interna and the presence of a transverse suture on uropodal exopod link
<taxonomicName id="4F8D4D67201CFFD7FF4F2B0FFEEFFCDE" box="[204,291,801,824]" class="Malacostraca" family="Axiidae" genus="Ezaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="579" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="BAF9EAF6201CFFD7FF4F2B0FFEEFFCDE" box="[204,291,801,824]" italics="true" pageId="8" pageNumber="579">Ezaxius</emphasis>
</taxonomicName>
<emphasis id="BAF9EAF6201CFFD7FEAE2B31FEB2FCDE" bold="true" box="[301,382,799,824]" pageId="8" pageNumber="579">
<taxonomicNameLabel id="A1CA578D201CFFD7FEAE2B31FEB2FCDE" box="[301,382,799,824]" pageId="8" pageNumber="579" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
with
<taxonomicName id="4F8D4D67201CFFD7FE402B31FD37FCDE" authority="Sakai, 2011" authorityName="Sakai" authorityYear="2011" box="[451,763,799,824]" class="Malacostraca" family="Axiidae" genus="Albatrossaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="BAF9EAF6201CFFD7FE402B31FDA4FCDE" box="[451,616,799,824]" italics="true" pageId="8" pageNumber="579">Albatrossaxius</emphasis>
<bibRefCitation id="EC1C4B15201CFFD7FDF12B31FD37FCDE" author="Sakai" box="[626,763,799,824]" pageId="8" pageNumber="579" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). In: Fransen, C. H. J. M. &amp; Vaupel Klein, J. C. von (Eds.), Crustaceana Monographs. Vol. 13. Leiden, Brill, pp. 1 - 520. http: // dx. doi. org / 10.1163 / 9789047424185" type="book" year="2011">Sakai, 2011</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4F8D4D67201CFFD7FC8A2B0EFB0BFCDE" authority="Kensley &amp; Simmons, 1988" authorityName="Kensley &amp; Simmons" authorityYear="1988" box="[777,1223,799,824]" class="Malacostraca" family="Axiidae" genus="Axiorygma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="BAF9EAF6201CFFD7FC8A2B0EFC48FCD1" box="[777,900,800,823]" italics="true" pageId="8" pageNumber="579">Axiorygma</emphasis>
<bibRefCitation id="EC1C4B15201CFFD7FC0E2B31FB0BFCDE" author="Kensley" box="[909,1223,799,824]" pageId="8" pageNumber="579" refString="Kensley, B. &amp; Simmons, G. M. (1988) Axiorygma nethertoni, a new genus and species of thalassinidean shrimp from Florida (Decapoda: Axiidae). Journal of Crustacean Biology, 8 (4), 657 - 667. http: // dx. doi. org / 10.2307 / 1548701" type="journal article" year="1988">Kensley &amp; Simmons, 1988</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4F8D4D67201CFFD7FB562B31FF1DFCBA" authority="Sakai, 2011" authorityName="Sakai" authorityYear="2011" class="Malacostraca" family="Axiidae" genus="Balssaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="BAF9EAF6201CFFD7FB562B31FA81FCDE" box="[1237,1357,799,824]" italics="true" pageId="8" pageNumber="579">Balssaxius</emphasis>
<bibRefCitation id="EC1C4B15201CFFD7FAD52B31FF1DFCBA" author="Sakai" pageId="8" pageNumber="579" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). In: Fransen, C. H. J. M. &amp; Vaupel Klein, J. C. von (Eds.), Crustaceana Monographs. Vol. 13. Leiden, Brill, pp. 1 - 520. http: // dx. doi. org / 10.1163 / 9789047424185" type="book" year="2011">Sakai, 2011</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4F8D4D67201CFFD7FF612B6BFDC5FCBA" authority="Sakai, 2011" authorityName="Sakai" authorityYear="2011" box="[226,521,836,861]" class="Malacostraca" family="Axiidae" genus="Guyanacaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="BAF9EAF6201CFFD7FF612B6BFEBEFCBA" box="[226,370,837,860]" italics="true" pageId="8" pageNumber="579">Guyanacaris</emphasis>
<bibRefCitation id="EC1C4B15201CFFD7FEFC2B6AFDC5FCBA" author="Sakai" box="[383,521,836,861]" pageId="8" pageNumber="579" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). In: Fransen, C. H. J. M. &amp; Vaupel Klein, J. C. von (Eds.), Crustaceana Monographs. Vol. 13. Leiden, Brill, pp. 1 - 520. http: // dx. doi. org / 10.1163 / 9789047424185" type="book" year="2011">Sakai, 2011</bibRefCitation>
</taxonomicName>
and
<emphasis id="BAF9EAF6201CFFD7FDC82B6AFD28FCBB" box="[587,740,836,861]" italics="true" pageId="8" pageNumber="579">Leonardaxius</emphasis>
<bibRefCitation id="EC1C4B15201CFFD7FD722B6AFCB2FCBA" author="Sakai" box="[753,894,836,861]" pageId="8" pageNumber="579" refString="Sakai, K. (2011) Axioidea of the world and a reconsideration of the Callianassoidea (Decapoda, Thalassinidea, Callianassida). In: Fransen, C. H. J. M. &amp; Vaupel Klein, J. C. von (Eds.), Crustaceana Monographs. Vol. 13. Leiden, Brill, pp. 1 - 520. http: // dx. doi. org / 10.1163 / 9789047424185" type="book" year="2011">Sakai, 2011</bibRefCitation>
. However, all of them present a conspicuous posteromedian tooth on telson (vs no tooth in
<taxonomicName id="4F8D4D67201CFFD7FD552B47FCE1FC66" box="[726,813,873,896]" class="Malacostraca" family="Axiidae" genus="Ezaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="579" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="BAF9EAF6201CFFD7FD552B47FCE1FC66" box="[726,813,873,896]" italics="true" pageId="8" pageNumber="579">Ezaxius</emphasis>
</taxonomicName>
<emphasis id="BAF9EAF6201CFFD7FCB72B49FC41FC66" bold="true" box="[820,909,871,896]" pageId="8" pageNumber="579">
<taxonomicNameLabel id="A1CA578D201CFFD7FCB72B49FC48FC66" box="[820,900,871,896]" pageId="8" pageNumber="579" rank="genus">n. gen.</taxonomicNameLabel>
)
</emphasis>
and none of them was described as being hermaphrodite. Moreover,
<taxonomicName id="4F8D4D67201CFFD7FE442BA3FDD2FC42" box="[455,542,909,932]" class="Malacostraca" family="Axiidae" genus="Ezaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="579" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="BAF9EAF6201CFFD7FE442BA3FDD2FC42" box="[455,542,909,932]" italics="true" pageId="8" pageNumber="579">Ezaxius</emphasis>
</taxonomicName>
<emphasis id="BAF9EAF6201CFFD7FDA62BA2FDB8FC43" bold="true" box="[549,628,908,933]" pageId="8" pageNumber="579">
<taxonomicNameLabel id="A1CA578D201CFFD7FDA62BA2FDB8FC43" box="[549,628,908,933]" pageId="8" pageNumber="579" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
can be separated from
<taxonomicName id="4F8D4D67201CFFD7FC002BA2FBE4FC43" box="[899,1064,908,933]" class="Malacostraca" family="Axiidae" genus="Albatrossaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="BAF9EAF6201CFFD7FC002BA2FBE4FC43" box="[899,1064,908,933]" italics="true" pageId="8" pageNumber="579">Albatrossaxius</emphasis>
</taxonomicName>
,
<taxonomicName id="4F8D4D67201CFFD7FBBB2BA3FB05FC42" box="[1080,1225,909,932]" class="Malacostraca" family="Axiidae" genus="Guyanacaris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="BAF9EAF6201CFFD7FBBB2BA3FB05FC42" box="[1080,1225,909,932]" italics="true" pageId="8" pageNumber="579">Guyanacaris</emphasis>
</taxonomicName>
and
<emphasis id="BAF9EAF6201CFFD7FA802BA2FA50FC43" box="[1283,1436,908,933]" italics="true" pageId="8" pageNumber="579">Leonardaxius</emphasis>
by the presence of appendix masculina (vs absent in our specimen); from
<taxonomicName id="4F8D4D67201CFFD7FC4F2B81FB88FC2E" box="[972,1092,943,968]" class="Malacostraca" family="Axiidae" genus="Balssaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="BAF9EAF6201CFFD7FC4F2B81FB88FC2E" box="[972,1092,943,968]" italics="true" pageId="8" pageNumber="579">Balssaxius</emphasis>
</taxonomicName>
by the cervical groove distinct along the entire length (vs dorsally but not fully laterally distinct) and from
<taxonomicName id="4F8D4D67201CFFD7FBBF2BFAFB74FC0D" box="[1084,1208,980,1003]" class="Malacostraca" family="Axiidae" genus="Axiorygma" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="579" phylum="Arthropoda" rank="genus">
<emphasis id="BAF9EAF6201CFFD7FBBF2BFAFB74FC0D" box="[1084,1208,980,1003]" italics="true" pageId="8" pageNumber="579">Axiorygma</emphasis>
</taxonomicName>
by the absence of postcervical carina and pleopod 1 (vs presence). Therefore, while generic definitions in this family continue to be unclear or vague (
<bibRefCitation id="EC1C4B15201CFFD7FEE22C32FD42FBD2" author="Kensley" box="[353,654,1052,1077]" pageId="8" pageNumber="579" refString="Kensley, B. &amp; Simmons, G. M. (1988) Axiorygma nethertoni, a new genus and species of thalassinidean shrimp from Florida (Decapoda: Axiidae). Journal of Crustacean Biology, 8 (4), 657 - 667. http: // dx. doi. org / 10.2307 / 1548701" type="journal article" year="1988">Kensley &amp; Simmons 1988</bibRefCitation>
;
<bibRefCitation id="EC1C4B15201CFFD7FD192C33FCD2FBD2" author="Poore" box="[666,798,1052,1076]" pageId="8" pageNumber="579" refString="Poore, G. C. B. (1994) A phylogeny of the families of Thalassinidea (Crustacea: Decapoda) with keys to families and genera. Memoirs of Museum Victoria, 54, 79 - 120." type="journal article" year="1994">Poore 1994</bibRefCitation>
; see above), we describe
<taxonomicName id="4F8D4D67201CFFD7FBBF2C33FB5FFBD2" box="[1084,1171,1053,1076]" class="Malacostraca" family="Axiidae" genus="Ezaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="579" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="BAF9EAF6201CFFD7FBBF2C33FB5FFBD2" box="[1084,1171,1053,1076]" italics="true" pageId="8" pageNumber="579">Ezaxius</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A1CA578D201CFFD7FB192C32FB24FBD3" box="[1178,1256,1052,1077]" pageId="8" pageNumber="579" rank="genus">
<emphasis id="BAF9EAF6201CFFD7FB192C32FB24FBD3" bold="true" box="[1178,1256,1052,1077]" pageId="8" pageNumber="579">n. gen.</emphasis>
</taxonomicNameLabel>
here in order to accommodate the new species
<taxonomicName id="4F8D4D67201CFFD7FE732C6FFD00FBBE" box="[496,716,1087,1112]" class="Malacostraca" family="Axiidae" genus="Ezaxius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="579" phylum="Arthropoda" rank="species" species="ferachevali" status="sp. nov.">
<emphasis id="BAF9EAF6201CFFD7FE732C6FFD00FBBE" box="[496,716,1087,1112]" italics="true" pageId="8" pageNumber="579">Ezaxius ferachevali</emphasis>
</taxonomicName>
<emphasis id="BAF9EAF6201CFFD7FD502C11FCDEFBBE" bold="true" box="[723,786,1087,1112]" pageId="8" pageNumber="579">
<taxonomicNameLabel id="A1CA578D201CFFD7FD502C11FCDEFBBE" box="[723,786,1087,1112]" pageId="8" pageNumber="579" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>