376 lines
51 KiB
XML
376 lines
51 KiB
XML
<document id="1BAAC764BC88CC098192FE1A679F0557" ID-DOI="10.11646/zootaxa.1859.1.4" ID-ISSN="11755334" ID-Zenodo-Dep="5133117" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1627155571059" checkinUser="felipe" docAuthor="Miller, Kelly B. & Wheeler, Quentin D." docDate="2008" docId="03DD87C9A94CFFFCAEB5F948FD74FE3C" docLanguage="en" docName="zt01859p068.pdf" docOrigin="Zootaxa 1859 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1859.1.4" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Agaporomorphus colberti Miller & Wheeler 2008, sp. n." docType="treatment" docVersion="4" lastPageNumber="67" masterDocId="FFE4FFB1A94DFFF8AE22FF85FF99FF9E" masterDocTitle="A new species of Agaporomorphus Zimmermann from Venezuela, and a review of the A. knischi species group (Coleoptera: Dytiscidae: Copelatinae)" masterLastPageNumber="68" masterPageNumber="63" pageNumber="64" updateTime="1699021298510" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="B03D91E2EBB306C36A347700D6858DB5">A new species of Agaporomorphus Zimmermann from Venezuela, and a review of the A. knischi species group (Coleoptera: Dytiscidae: Copelatinae)</mods:title>
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<mods:name id="4478B8651BDBFBDF4DCA9461D2566DC3" type="personal">
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<mods:roleTerm id="45B6E8FF9D74BBDA6433B4037C422826">Author</mods:roleTerm>
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<mods:namePart id="8EC5144989336F865EE6E88BA3C7A32D">Miller, Kelly B.</mods:namePart>
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<mods:affiliation id="7AAB1889BD86CE7837CBE0C78A38007F">Department of Biology and Museum of Southwestern Biology, University of New Mexico, 167 Castetter Hall, MSC 03 2020, Albuquerque, NM 87131 - 0001, USA. E-mail: kbmiller @ unm. edu</mods:affiliation>
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<mods:roleTerm id="8AEADDBEA518D11E5CE0949FC223C757">Author</mods:roleTerm>
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<mods:namePart id="6964258A7EAE48D85190482523712E6E">Wheeler, Quentin D.</mods:namePart>
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<mods:affiliation id="9B41A7028676F8B66AE462EE85904D3C">School of Life Sciences, Arizona State University, PO Box 876505, Tempe, AZ 85287 - 6505, USA. E-mail: Quentin. Wheeler @ asu. edu Corresponding author</mods:affiliation>
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<mods:typeOfResource id="34B4C450685DD70A3DBD8CBE2C3F633B">text</mods:typeOfResource>
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<mods:title id="508380FF668D398BBC70A0DDB561DE63">Zootaxa</mods:title>
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<mods:part id="A9B21887A14392D5547519BDDE36B190">
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<mods:date id="BB4F80862DA847FBAFEE83E1F611B94B">2008</mods:date>
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<mods:number id="B4D8DC449ECCCFA000EA38996D7CBCF3">2008-08-29</mods:number>
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<mods:number id="3FDFD8A327A4EABFDE71DF63FB31C7D1">1859</mods:number>
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<mods:start id="3EBF81E8A6DEEB7BB27176D82BFAAEE1">63</mods:start>
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<mods:url id="4BF41F7E26345884CD2FDFADF41F1452">https://biotaxa.org/Zootaxa/article/view/zootaxa.1859.1.4</mods:url>
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<mods:classification id="9EB18914DCBD966804341C76E450856F">journal article</mods:classification>
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<mods:identifier id="BBD1B6B300018D752C91687529398D9D" type="DOI">10.11646/zootaxa.1859.1.4</mods:identifier>
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<subSubSection id="C36E6554A94CFFF9AEB5F948FD8FF979" box="[151,534,1741,1767]" pageId="1" pageNumber="64" type="nomenclature">
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<paragraph id="8BCB36DFA94CFFF9AEB5F948FD8FF979" blockId="1.[151,534,1741,1802]" box="[151,534,1741,1767]" pageId="1" pageNumber="64">
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<heading id="D08381B3A94CFFF9AEB5F948FD8FF979" bold="true" box="[151,534,1741,1767]" fontSize="11" level="1" pageId="1" pageNumber="64" reason="1">
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<emphasis id="B900EACDA94CFFF9AEB5F948FD8FF979" bold="true" box="[151,534,1741,1767]" pageId="1" pageNumber="64">
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<taxonomicName id="4C744D5CA94CFFF9AEB5F948FE54F979" ID-CoL="65MFV" authority="Miller & Wheeler, 2008" authorityName="Miller & Wheeler" authorityYear="2008" box="[151,461,1741,1767]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="species" species="colberti" status="sp. nov.">
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<emphasis id="B900EACDA94CFFF9AEB5F948FE54F979" bold="true" box="[151,461,1741,1767]" italics="true" pageId="1" pageNumber="64">Agaporomorphus colberti</emphasis>
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<taxonomicNameLabel id="A23357B6A94CFFF9AFF6F948FD8FF979" box="[468,534,1741,1767]" pageId="1" pageNumber="64" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C36E6554A94CFFF9AEB5F975FEEEF894" box="[151,375,1776,1802]" pageId="1" pageNumber="64" type="description">
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<paragraph id="8BCB36DFA94CFFF9AEB5F975FEEEF894" blockId="1.[151,534,1741,1802]" box="[151,375,1776,1802]" pageId="1" pageNumber="64">
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(
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<figureCitation id="134F2A5AA94CFFF9AEBDF975FE90F894" box="[159,265,1776,1802]" captionStart="FIGURES" captionStartId="2.[151,203,1333,1359]" captionTargetBox="[543,1054,647,1303]" captionTargetId="figure-459@2.[529,1059,637,1306]" captionTargetPageId="2" captionText="FIGURES 1–6. Agaporomorphus species, morphological details. 1, 5) A. colberti; 2) A. silvaticus; 3) A. tambopatensis; 4, 6) A. knischi. 1) dorsal habitus; 2–4) male left antenna, ventral aspect; 5, 6) visible abdominal sternites, ventral aspect. Scale bars; a = 1mm for Figs 1,5,6; b = 0.25mm for Figs 2–4." figureDoi="http://doi.org/10.5281/zenodo.5133121" httpUri="https://zenodo.org/record/5133121/files/figure.png" pageId="1" pageNumber="64">Figs 1, 6</figureCitation>
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, 7–9, 21)
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</paragraph>
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</subSubSection>
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<subSubSection id="C36E6554A94CFFF9AEB5F8BAFE8FF81C" pageId="1" pageNumber="64" type="materials_examined">
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<paragraph id="8BCB36DFA94CFFF9AEB5F8BAFE8FF81C" blockId="1.[151,1436,1855,2002]" pageId="1" pageNumber="64">
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<materialsCitation id="3B1C3C82A94CFFF9AEB5F8BAFE8BF81C" country="Venezuela" county="Departamento" latitude="0.9166667" location="Rio Baria" longLatPrecision="1308" longitude="-66.166664" municipality="Rio Negro" pageId="1" pageNumber="64" specimenCount="1" stateProvince="Amazonas" typeStatus="holotype">
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<emphasis id="B900EACDA94CFFF9AEB5F8BAFEDCF8C7" bold="true" box="[151,325,1855,1881]" pageId="1" pageNumber="64">
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<typeStatus id="54CF887DA94CFFF9AEB5F8BAFF4DF8C7" box="[151,212,1855,1881]" pageId="1" pageNumber="64">Type</typeStatus>
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locality.
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</emphasis>
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<collectingCountry id="F363764FA94CFFF9AF74F8C5FE48F8C4" box="[342,465,1856,1882]" name="Venezuela" pageId="1" pageNumber="64">Venezuela</collectingCountry>
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,
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<collectingCounty id="62AA4E53A94CFFF9AFC4F8C5FD15F8C4" box="[486,652,1856,1882]" pageId="1" pageNumber="64">Departamento</collectingCounty>
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<collectingRegion id="49B0F83DA94CFFF9ACBFF8C5FC82F8C4" box="[669,795,1856,1882]" country="Venezuela" name="Amazonas" pageId="1" pageNumber="64">Amazonas</collectingRegion>
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,
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<collectingMunicipality id="6BAFACA5A94CFFF9AD12F8C5FC2EF8C4" box="[816,951,1856,1882]" pageId="1" pageNumber="64">Rio Negro</collectingMunicipality>
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,
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<location id="8EAB6004A94CFFF9ADEEF8C5FBD0F8C4" LSID="urn:lsid:plazi:treatment:03DD87C9A94CFFFCAEB5F948FD74FE3C:8EAB6004A94CFFF9ADEEF8C5FBD0F8C4" box="[972,1097,1856,1882]" country="Venezuela" county="Departamento" latitude="0.9166667" longLatPrecision="1308" longitude="-66.166664" municipality="Rio Negro" name="Rio Baria" pageId="1" pageNumber="64" stateProvince="Amazonas">Rio Baria</location>
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,
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<location id="8EAB6004A94CFFF9AA7CF8C5FAB8F8C4" LSID="urn:lsid:plazi:treatment:03DD87C9A94CFFFCAEB5F948FD74FE3C:8EAB6004A94CFFF9AA7CF8C5FAB8F8C4" box="[1118,1313,1856,1882]" country="Venezuela" county="Departamento" latitude="0.9166667" longLatPrecision="1308" longitude="-66.166664" municipality="Rio Negro" name="Dpt Rio Negro" pageId="1" pageNumber="64" stateProvince="Amazonas">
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Dpt
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<location id="8EAB6004A94CFFF9AAB9F8C5FAB8F8C4" LSID="urn:lsid:plazi:treatment:03DD87C9A94CFFFCAEB5F948FD74FE3C:8EAB6004A94CFFF9AAB9F8C5FAB8F8C4" box="[1179,1313,1856,1882]" country="Venezuela" county="Departamento" latitude="0.9166667" longLatPrecision="1308" longitude="-66.166664" municipality="Rio Negro" name="Rio Negro" pageId="1" pageNumber="64" stateProvince="Amazonas">Rio Negro</location>
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</location>
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,
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<geoCoordinate id="EE405018A94CFFF9AB15F8C5FA05F8C4" box="[1335,1436,1856,1882]" degrees="00" direction="north" minutes="55" orientation="latitude" pageId="1" pageNumber="64" precision="925" value="0.9166667">00°55’N</geoCoordinate>
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<geoCoordinate id="EE405018A94CFFF9AEB5F8EDFE8BF81C" box="[151,274,1896,1922]" degrees="066" direction="west" minutes="10" orientation="longitude" pageId="1" pageNumber="64" precision="925" value="-66.166664">066°10’W</geoCoordinate>
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C36E6554A94CFFFAAEE7F80AFE51FDDC" lastPageId="2" lastPageNumber="65" pageId="1" pageNumber="64" type="diagnosis">
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<paragraph id="8BCB36DFA94CFFFAAEE7F80AFE51FDDC" blockId="1.[151,1436,1855,2002]" lastBlockId="2.[151,1436,152,578]" lastPageId="2" lastPageNumber="65" pageId="1" pageNumber="64">
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<emphasis id="B900EACDA94CFFF9AEE7F80AFEDCF837" bold="true" box="[197,325,1935,1961]" pageId="1" pageNumber="64">Diagnosis.</emphasis>
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<taxonomicName id="4C744D5CA94CFFF9AF6DF815FDE1F834" box="[335,632,1936,1962]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="species" species="undetermined">
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<emphasis id="B900EACDA94CFFF9AF6DF815FD82F837" box="[335,539,1936,1961]" italics="true" pageId="1" pageNumber="64">Agaporomorphus</emphasis>
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species
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</taxonomicName>
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are extremely similar in characters such as size, general shape and coloration (
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||
<figureCitation id="134F2A5AA94CFFF9AF3FF83DFEF8F84C" box="[285,353,1976,2002]" captionStart="FIGURES" captionStartId="2.[151,203,1333,1359]" captionTargetBox="[543,1054,647,1303]" captionTargetId="figure-459@2.[529,1059,637,1306]" captionTargetPageId="2" captionText="FIGURES 1–6. Agaporomorphus species, morphological details. 1, 5) A. colberti; 2) A. silvaticus; 3) A. tambopatensis; 4, 6) A. knischi. 1) dorsal habitus; 2–4) male left antenna, ventral aspect; 5, 6) visible abdominal sternites, ventral aspect. Scale bars; a = 1mm for Figs 1,5,6; b = 0.25mm for Figs 2–4." figureDoi="http://doi.org/10.5281/zenodo.5133121" httpUri="https://zenodo.org/record/5133121/files/figure.png" pageId="1" pageNumber="64">Fig. 1</figureCitation>
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). This species can be distinguished from all other known
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<taxonomicName id="4C744D5CA94CFFF9AA21F83DFB56F84F" authorityName="Zimmermann" authorityYear="1921" box="[1027,1231,1976,2001]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="genus">
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<emphasis id="B900EACDA94CFFF9AA21F83DFB56F84F" box="[1027,1231,1976,2001]" italics="true" pageId="1" pageNumber="64">Agaporomorphus</emphasis>
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</taxonomicName>
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except
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<taxonomicName id="4C744D5CA94CFFF9AB0BF83DFA05F84F" authorityName="Zimmermann" authorityYear="1921" box="[1321,1436,1976,2001]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="64" phylum="Arthropoda" rank="species" species="knischi">
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<emphasis id="B900EACDA94CFFF9AB0BF83DFA05F84F" box="[1321,1436,1976,2001]" italics="true" pageId="1" pageNumber="64">A. knischi</emphasis>
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</taxonomicName>
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by the following character combination in males: 1) antennomeres V and VI broadly expanded with VI more expanded than V and with a distinct emargination on the posteroventral surface (
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<figureCitation id="134F2A5AA94FFFFAAA64FF45FB14FF44" box="[1094,1165,192,218]" captionStart="FIGURES" captionStartId="2.[151,203,1333,1359]" captionTargetBox="[543,1054,647,1303]" captionTargetId="figure-459@2.[529,1059,637,1306]" captionTargetPageId="2" captionText="FIGURES 1–6. Agaporomorphus species, morphological details. 1, 5) A. colberti; 2) A. silvaticus; 3) A. tambopatensis; 4, 6) A. knischi. 1) dorsal habitus; 2–4) male left antenna, ventral aspect; 5, 6) visible abdominal sternites, ventral aspect. Scale bars; a = 1mm for Figs 1,5,6; b = 0.25mm for Figs 2–4." figureDoi="http://doi.org/10.5281/zenodo.5133121" httpUri="https://zenodo.org/record/5133121/files/figure.png" pageId="2" pageNumber="65">Fig. 4</figureCitation>
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), 2) abdominal sternite V with an apicomedial triangular process (
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<figureCitation id="134F2A5AA94FFFFAACAEFF6DFD6FFE9C" box="[652,758,232,258]" captionStart="FIGURES" captionStartId="2.[151,203,1333,1359]" captionTargetBox="[543,1054,647,1303]" captionTargetId="figure-459@2.[529,1059,637,1306]" captionTargetPageId="2" captionText="FIGURES 1–6. Agaporomorphus species, morphological details. 1, 5) A. colberti; 2) A. silvaticus; 3) A. tambopatensis; 4, 6) A. knischi. 1) dorsal habitus; 2–4) male left antenna, ventral aspect; 5, 6) visible abdominal sternites, ventral aspect. Scale bars; a = 1mm for Figs 1,5,6; b = 0.25mm for Figs 2–4." figureDoi="http://doi.org/10.5281/zenodo.5133121" httpUri="https://zenodo.org/record/5133121/files/figure.png" pageId="2" pageNumber="65">Figs 5, 6</figureCitation>
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), and 3) abdominal sternite VI with a broad medial fovea (
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<figureCitation id="134F2A5AA94FFFFAAEBDFE95FE94FEB4" box="[159,269,272,298]" captionStart="FIGURES" captionStartId="2.[151,203,1333,1359]" captionTargetBox="[543,1054,647,1303]" captionTargetId="figure-459@2.[529,1059,637,1306]" captionTargetPageId="2" captionText="FIGURES 1–6. Agaporomorphus species, morphological details. 1, 5) A. colberti; 2) A. silvaticus; 3) A. tambopatensis; 4, 6) A. knischi. 1) dorsal habitus; 2–4) male left antenna, ventral aspect; 5, 6) visible abdominal sternites, ventral aspect. Scale bars; a = 1mm for Figs 1,5,6; b = 0.25mm for Figs 2–4." figureDoi="http://doi.org/10.5281/zenodo.5133121" httpUri="https://zenodo.org/record/5133121/files/figure.png" pageId="2" pageNumber="65">Figs 5, 6</figureCitation>
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). This new species can be clearly placed within a group of species (the
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<taxonomicName id="4C744D5CA94FFFFAAA53FE95FB7FFEB7" authorityName="Zimmermann" authorityYear="1921" box="[1137,1254,272,297]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="knischi">
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<emphasis id="B900EACDA94FFFFAAA53FE95FB7FFEB7" box="[1137,1254,272,297]" italics="true" pageId="2" pageNumber="65">A. knischi</emphasis>
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</taxonomicName>
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species group) characterized by distinctly modified male genitalia with a short series of fine setae on each side of the ventral surface (Figs 7, 10, 13, 17). Within this group,
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||
<taxonomicName id="4C744D5CA94FFFFAACF2FEE5FCD7FEE7" authority="Miller & Wheeler, 2008" authorityName="Miller & Wheeler" authorityYear="2008" box="[720,846,352,377]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="colberti" status="sp. nov.">
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||
<emphasis id="B900EACDA94FFFFAACF2FEE5FCD7FEE7" box="[720,846,352,377]" italics="true" pageId="2" pageNumber="65">A. colberti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EACDA94FFFFAAD7AFEDAFC05FEE7" bold="true" box="[856,924,351,377]" pageId="2" pageNumber="65">
|
||
<taxonomicNameLabel id="A23357B6A94FFFFAAD7AFEDAFC05FEE7" box="[856,924,351,377]" pageId="2" pageNumber="65" rank="species">sp. n.</taxonomicNameLabel>
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||
</emphasis>
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is most similar to
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<taxonomicName id="4C744D5CA94FFFFAAA5DFEE5FB6CFEE7" authorityName="Zimmermann" authorityYear="1921" box="[1151,1269,352,377]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="knischi">
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<emphasis id="B900EACDA94FFFFAAA5DFEE5FB6CFEE7" box="[1151,1269,352,377]" italics="true" pageId="2" pageNumber="65">A. knischi</emphasis>
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||
</taxonomicName>
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in having the character combination indicated above, but differs from that species in the shape of the male median lobe of the aedeagus (Figs 7, 8) which is broader in ventral and lateral aspects and has a prominently broad and dorsally angulate ventral region near the base that is narrower in
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<taxonomicName id="4C744D5CA94FFFFAAD4EFE5DFC79FE6F" authorityName="Zimmermann" authorityYear="1921" box="[876,992,472,497]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="knischi">
|
||
<emphasis id="B900EACDA94FFFFAAD4EFE5DFC79FE6F" box="[876,992,472,497]" italics="true" pageId="2" pageNumber="65">A. knischi</emphasis>
|
||
</taxonomicName>
|
||
(Figs 17, 18), among numerous other shape features. Also, the posteromedial spine on abdominal sternite V is smaller in
|
||
<taxonomicName id="4C744D5CA94FFFFAAA51FD85FB69FD87" authority="Miller & Wheeler, 2008" authorityName="Miller & Wheeler" authorityYear="2008" box="[1139,1264,512,537]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="colberti" status="sp. nov.">
|
||
<emphasis id="B900EACDA94FFFFAAA51FD85FB69FD87" box="[1139,1264,512,537]" italics="true" pageId="2" pageNumber="65">A. colberti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EACDA94FFFFAAADAFE7AFAA2FD87" bold="true" box="[1272,1339,511,537]" pageId="2" pageNumber="65">
|
||
<taxonomicNameLabel id="A23357B6A94FFFFAAADAFE7AFAA2FD87" box="[1272,1339,511,537]" pageId="2" pageNumber="65" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="134F2A5AA94FFFFAAB6EFD85FA0AFD84" box="[1356,1427,512,538]" captionStart="FIGURES" captionStartId="2.[151,203,1333,1359]" captionTargetBox="[543,1054,647,1303]" captionTargetId="figure-459@2.[529,1059,637,1306]" captionTargetPageId="2" captionText="FIGURES 1–6. Agaporomorphus species, morphological details. 1, 5) A. colberti; 2) A. silvaticus; 3) A. tambopatensis; 4, 6) A. knischi. 1) dorsal habitus; 2–4) male left antenna, ventral aspect; 5, 6) visible abdominal sternites, ventral aspect. Scale bars; a = 1mm for Figs 1,5,6; b = 0.25mm for Figs 2–4." figureDoi="http://doi.org/10.5281/zenodo.5133121" httpUri="https://zenodo.org/record/5133121/files/figure.png" pageId="2" pageNumber="65">Fig. 5</figureCitation>
|
||
) than in
|
||
<taxonomicName id="4C744D5CA94FFFFAAECDFDADFEFBFDDF" authorityName="Zimmermann" authorityYear="1921" box="[239,354,552,577]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="knischi">
|
||
<emphasis id="B900EACDA94FFFFAAECDFDADFEFBFDDF" box="[239,354,552,577]" italics="true" pageId="2" pageNumber="65">A. knischi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="134F2A5AA94FFFFAAF50FDADFE23FDDC" box="[370,442,552,578]" captionStart="FIGURES" captionStartId="2.[151,203,1333,1359]" captionTargetBox="[543,1054,647,1303]" captionTargetId="figure-459@2.[529,1059,637,1306]" captionTargetPageId="2" captionText="FIGURES 1–6. Agaporomorphus species, morphological details. 1, 5) A. colberti; 2) A. silvaticus; 3) A. tambopatensis; 4, 6) A. knischi. 1) dorsal habitus; 2–4) male left antenna, ventral aspect; 5, 6) visible abdominal sternites, ventral aspect. Scale bars; a = 1mm for Figs 1,5,6; b = 0.25mm for Figs 2–4." figureDoi="http://doi.org/10.5281/zenodo.5133121" httpUri="https://zenodo.org/record/5133121/files/figure.png" pageId="2" pageNumber="65">Fig. 6</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF0B6657A94FFFFAAEB5FAB0FCBEFA07" ID-DOI="http://doi.org/10.5281/zenodo.5133121" ID-Zenodo-Dep="5133121" httpUri="https://zenodo.org/record/5133121/files/figure.png" pageId="2" pageNumber="65" startId="2.[151,203,1333,1359]" targetBox="[543,1054,647,1303]" targetPageId="2">
|
||
<paragraph id="8BCB36DFA94FFFFAAEB5FAB0FCBEFA07" blockId="2.[151,1437,1333,1433]" pageId="2" pageNumber="65">
|
||
<emphasis id="B900EACDA94FFFFAAEB5FAB0FEDFFAD0" bold="true" box="[151,326,1333,1359]" pageId="2" pageNumber="65">FIGURES 1–6.</emphasis>
|
||
<taxonomicName id="4C744D5CA94FFFFAAF6FFAB2FDC7FAD0" box="[333,606,1335,1358]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B900EACDA94FFFFAAF6FFAB2FD9FFAD0" box="[333,518,1335,1358]" italics="true" pageId="2" pageNumber="65">Agaporomorphus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, morphological details. 1, 5)
|
||
<taxonomicName id="4C744D5CA94FFFFAADB6FAB2FB9CFAD0" authorityName="Miller & Wheeler" authorityYear="2008" box="[916,1029,1335,1358]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="colberti">
|
||
<emphasis id="B900EACDA94FFFFAADB6FAB2FB9CFAD0" box="[916,1029,1335,1358]" italics="true" pageId="2" pageNumber="65">A. colberti</emphasis>
|
||
</taxonomicName>
|
||
; 2)
|
||
<taxonomicName id="4C744D5CA94FFFFAAA12FAB2FB2BFAD0" authorityName="Miller" authorityYear="2005" box="[1072,1202,1335,1358]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="silvaticus">
|
||
<emphasis id="B900EACDA94FFFFAAA12FAB2FB2BFAD0" box="[1072,1202,1335,1358]" italics="true" pageId="2" pageNumber="65">A. silvaticus</emphasis>
|
||
</taxonomicName>
|
||
; 3)
|
||
<taxonomicName id="4C744D5CA94FFFFAAAFFFAB2FA0CFAD0" authorityName="Miller" authorityYear="2005" box="[1245,1429,1335,1358]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="tambopatensis">
|
||
<emphasis id="B900EACDA94FFFFAAAFFFAB2FA0CFAD0" box="[1245,1429,1335,1358]" italics="true" pageId="2" pageNumber="65">A. tambopatensis</emphasis>
|
||
</taxonomicName>
|
||
; 4, 6)
|
||
<taxonomicName id="4C744D5CA94FFFFAAEECFAD8FEA1FAEA" authorityName="Zimmermann" authorityYear="1921" box="[206,312,1373,1396]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="65" phylum="Arthropoda" rank="species" species="knischi">
|
||
<emphasis id="B900EACDA94FFFFAAEECFAD8FEA1FAEA" box="[206,312,1373,1396]" italics="true" pageId="2" pageNumber="65">A. knischi</emphasis>
|
||
</taxonomicName>
|
||
. 1) dorsal habitus; 2–4) male left antenna, ventral aspect; 5, 6) visible abdominal sternites, ventral aspect. Scale bars; a = 1mm for Figs 1,5,6; b = 0.25mm for Figs 2–4.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C36E6554A94FFFFBAEE7FA55FBDFF91E" lastPageId="3" lastPageNumber="66" pageId="2" pageNumber="65" type="description">
|
||
<paragraph id="8BCB36DFA94FFFFAAEE7FA55FA99F98C" blockId="2.[151,1437,1488,2034]" pageId="2" pageNumber="65">
|
||
<emphasis id="B900EACDA94FFFFAAEE7FA55FEC7FA74" bold="true" box="[197,350,1488,1514]" pageId="2" pageNumber="65">Description.</emphasis>
|
||
<emphasis id="B900EACDA94FFFFAAF46FA55FD94FA77" box="[356,525,1488,1513]" italics="true" pageId="2" pageNumber="65">Measurements</emphasis>
|
||
. TL = 2.93–3.18mm, GW = 1.39–1.57mm, PW = 1.24–1.38mm, HW = 0.78– 0.87mm, EW = 0.49–0.58mm, TL/GW = 1.97–2.18, HW/EW = 1.50–1.64, FW/FL = 2.59–3.22.
|
||
</paragraph>
|
||
<paragraph id="8BCB36DFA94FFFFAAEE4F9A5FE1CF914" blockId="2.[151,1437,1488,2034]" pageId="2" pageNumber="65">
|
||
<emphasis id="B900EACDA94FFFFAAEE4F9A5FEDEF9A7" box="[198,327,1568,1593]" italics="true" pageId="2" pageNumber="65">Coloration</emphasis>
|
||
(
|
||
<figureCitation id="134F2A5AA94FFFFAAF7EF9A5FE3CF9A4" box="[348,421,1568,1594]" captionStart="FIGURES" captionStartId="2.[151,203,1333,1359]" captionTargetBox="[543,1054,647,1303]" captionTargetId="figure-459@2.[529,1059,637,1306]" captionTargetPageId="2" captionText="FIGURES 1–6. Agaporomorphus species, morphological details. 1, 5) A. colberti; 2) A. silvaticus; 3) A. tambopatensis; 4, 6) A. knischi. 1) dorsal habitus; 2–4) male left antenna, ventral aspect; 5, 6) visible abdominal sternites, ventral aspect. Scale bars; a = 1mm for Figs 1,5,6; b = 0.25mm for Figs 2–4." figureDoi="http://doi.org/10.5281/zenodo.5133121" httpUri="https://zenodo.org/record/5133121/files/figure.png" pageId="2" pageNumber="65">Fig. 1</figureCitation>
|
||
). Red-brown on all dorsal surfaces, broadly light yellow along anterior margin of elytron and lateral margins of pronotum. All ventral surfaces and appendages yellow-brown except abdominal sternites red-yellow.
|
||
</paragraph>
|
||
<paragraph id="8BCB36DFA94FFFFAAEE4F91DFD05F8E4" blockId="2.[151,1437,1488,2034]" pageId="2" pageNumber="65">
|
||
<emphasis id="B900EACDA94FFFFAAEE4F91DFE42F92F" box="[198,475,1688,1713]" italics="true" pageId="2" pageNumber="65">Sculpture and structure</emphasis>
|
||
. Pronotum with microsculpture consisting of fine cells, with few very fine punctures interspersed; lateral pronotal bead fine, absent in anterior one-fifth. Prosternum medially strongly carinate, carina extending onto prosternal process; prosternal process medially with a distinct, sharp longitudinal carina extending to apex, laterally with strongly beaded margins, apex pointed. Elytron covered with extremely fine, evenly spaced, short striae, striae more punctiform laterally and apically. Metafemur moderately broad, length about 3 × greatest width.
|
||
</paragraph>
|
||
<paragraph id="8BCB36DFA94FFFFBAEE4F80DFAF8FF44" blockId="2.[151,1437,1488,2034]" lastBlockId="3.[151,1437,152,778]" lastPageId="3" lastPageNumber="66" pageId="2" pageNumber="65">
|
||
<emphasis id="B900EACDA94FFFFAAEE4F80DFEEBF83F" box="[198,370,1928,1953]" italics="true" pageId="2" pageNumber="65">Male genitalia</emphasis>
|
||
; Median lobe in lateral aspect robust, with broad expansion dorsad, apex elongate, sublinear, with distinct dorsally-directed lobe on right side medially and very broad, angular region subbasally, with linear series of fine setae on each side of dorsal midline (Fig. 7); in ventral aspect very robust, broad, with complicated folding and structures (Fig. 8). Lateral lobe broad basally, strongly narrowed medially, apical half very slender, apicomedially with large membranous lobes bearing extensive series of long setae (Figs 9).
|
||
</paragraph>
|
||
<paragraph id="8BCB36DFA94EFFFBAEE4FF6DFD2AFECC" blockId="3.[151,1437,152,778]" pageId="3" pageNumber="66">
|
||
<emphasis id="B900EACDA94EFFFBAEE4FF6DFE16FE9F" box="[198,399,232,257]" italics="true" pageId="3" pageNumber="66">Female genitalia</emphasis>
|
||
; Spermatheca long, loosely coiled, slender, tapered to spermathecal duct; spermathecal duct relatively short; fertilization duct long, coiled; gonocoxa slender, lateral margin broadly convex, apex expanded; laterotergite very slender, elongate.
|
||
</paragraph>
|
||
<paragraph id="8BCB36DFA94EFFFBAEE4FEE5FD69FC94" blockId="3.[151,1437,152,778]" pageId="3" pageNumber="66">
|
||
<emphasis id="B900EACDA94EFFFBAEE4FEE5FE30FEE7" box="[198,425,352,377]" italics="true" pageId="3" pageNumber="66">Sexual dimorphism</emphasis>
|
||
. Male protarsal claws unmodified; male pro- and mesotarsal claws about half length of mesotarsomere V; male without apical lobe on mesotarsomere V; male protarsomeres I and II broadened, protarsomere I with two large adhesive setae, protarsomere II without adhesive setae; male mesotarsomeres I and II slightly broadened, mesotarsomere I with one large, medial adhesive seta and two large, apical adhesive setae, mesotarsomere II with two smaller, apical adhesive setae; female pro- and mesotarsomeres unmodified. Male with fine but distinct triangular, posteriorly-directed prominence medially along posterior margin of visible abdominal sternite V (
|
||
<figureCitation id="134F2A5AA94EFFFBAFEFFDD5FD8AFDF4" box="[461,531,592,618]" captionStart="FIGURES" captionStartId="2.[151,203,1333,1359]" captionTargetBox="[543,1054,647,1303]" captionTargetId="figure-459@2.[529,1059,637,1306]" captionTargetPageId="2" captionText="FIGURES 1–6. Agaporomorphus species, morphological details. 1, 5) A. colberti; 2) A. silvaticus; 3) A. tambopatensis; 4, 6) A. knischi. 1) dorsal habitus; 2–4) male left antenna, ventral aspect; 5, 6) visible abdominal sternites, ventral aspect. Scale bars; a = 1mm for Figs 1,5,6; b = 0.25mm for Figs 2–4." figureDoi="http://doi.org/10.5281/zenodo.5133121" httpUri="https://zenodo.org/record/5133121/files/figure.png" pageId="3" pageNumber="66">Fig. 5</figureCitation>
|
||
); female without prominence. Male with broad, elongate depression medially on abdominal sternite VI (
|
||
<figureCitation id="134F2A5AA94EFFFBAFE9FDFDFD8BFD0C" box="[459,530,632,658]" captionStart="FIGURES" captionStartId="2.[151,203,1333,1359]" captionTargetBox="[543,1054,647,1303]" captionTargetId="figure-459@2.[529,1059,637,1306]" captionTargetPageId="2" captionText="FIGURES 1–6. Agaporomorphus species, morphological details. 1, 5) A. colberti; 2) A. silvaticus; 3) A. tambopatensis; 4, 6) A. knischi. 1) dorsal habitus; 2–4) male left antenna, ventral aspect; 5, 6) visible abdominal sternites, ventral aspect. Scale bars; a = 1mm for Figs 1,5,6; b = 0.25mm for Figs 2–4." figureDoi="http://doi.org/10.5281/zenodo.5133121" httpUri="https://zenodo.org/record/5133121/files/figure.png" pageId="3" pageNumber="66">Fig. 5</figureCitation>
|
||
); female without depression. Male with distinct parallel series of rugulosities on each side of midline on abdominal sternite III (
|
||
<figureCitation id="134F2A5AA94EFFFBACC5FD25FCB7FD24" box="[743,814,672,698]" captionStart="FIGURES" captionStartId="2.[151,203,1333,1359]" captionTargetBox="[543,1054,647,1303]" captionTargetId="figure-459@2.[529,1059,637,1306]" captionTargetPageId="2" captionText="FIGURES 1–6. Agaporomorphus species, morphological details. 1, 5) A. colberti; 2) A. silvaticus; 3) A. tambopatensis; 4, 6) A. knischi. 1) dorsal habitus; 2–4) male left antenna, ventral aspect; 5, 6) visible abdominal sternites, ventral aspect. Scale bars; a = 1mm for Figs 1,5,6; b = 0.25mm for Figs 2–4." figureDoi="http://doi.org/10.5281/zenodo.5133121" httpUri="https://zenodo.org/record/5133121/files/figure.png" pageId="3" pageNumber="66">Fig. 5</figureCitation>
|
||
); female without rugulosities. Male antennomeres V and VI modified, V broadly triangular, VI broad with large posterior emargination (as in
|
||
<figureCitation id="134F2A5AA94EFFFBAA82FD4DFB7FFD7C" box="[1184,1254,712,738]" captionStart="FIGURES" captionStartId="2.[151,203,1333,1359]" captionTargetBox="[543,1054,647,1303]" captionTargetId="figure-459@2.[529,1059,637,1306]" captionTargetPageId="2" captionText="FIGURES 1–6. Agaporomorphus species, morphological details. 1, 5) A. colberti; 2) A. silvaticus; 3) A. tambopatensis; 4, 6) A. knischi. 1) dorsal habitus; 2–4) male left antenna, ventral aspect; 5, 6) visible abdominal sternites, ventral aspect. Scale bars; a = 1mm for Figs 1,5,6; b = 0.25mm for Figs 2–4." figureDoi="http://doi.org/10.5281/zenodo.5133121" httpUri="https://zenodo.org/record/5133121/files/figure.png" pageId="3" pageNumber="66">Fig. 4</figureCitation>
|
||
); female antennomeres unmodified. Male metafemur unmodified.
|
||
</paragraph>
|
||
<paragraph id="8BCB36DFA94EFFFBAEB5F9A1FBDFF91E" blockId="3.[151,1436,1572,1664]" pageId="3" pageNumber="66">
|
||
<emphasis id="B900EACDA94EFFFBAEB5F9A1FEC9F9A2" bold="true" box="[151,336,1572,1596]" pageId="3" pageNumber="66">FIGURES 7–20.</emphasis>
|
||
<taxonomicName id="4C744D5CA94EFFFBAF75F9A1FDF1F9A5" box="[343,616,1572,1595]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B900EACDA94EFFFBAF75F9A1FD89F9A5" box="[343,528,1572,1595]" italics="true" pageId="3" pageNumber="66">Agaporomorphus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
, male gentitalia. 7–9)
|
||
<taxonomicName id="4C744D5CA94EFFFBAD7AF9A1FC53F9A5" authorityName="Miller & Wheeler" authorityYear="2008" box="[856,970,1572,1595]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="colberti">
|
||
<emphasis id="B900EACDA94EFFFBAD7AF9A1FC53F9A5" box="[856,970,1572,1595]" italics="true" pageId="3" pageNumber="66">A. colberti</emphasis>
|
||
</taxonomicName>
|
||
; 10–12)
|
||
<taxonomicName id="4C744D5CA94EFFFBAA09F9A1FB37F9A5" authorityName="Miller" authorityYear="2005" box="[1067,1198,1572,1595]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="silvaticus">
|
||
<emphasis id="B900EACDA94EFFFBAA09F9A1FB37F9A5" box="[1067,1198,1572,1595]" italics="true" pageId="3" pageNumber="66">A. silvaticus</emphasis>
|
||
</taxonomicName>
|
||
; 13–16)
|
||
<taxonomicName id="4C744D5CA94EFFFBAB32F9A1FF55F9C0" authorityName="Miller" authorityYear="2005" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="tambopatensis">
|
||
<emphasis id="B900EACDA94EFFFBAB32F9A1FF55F9C0" italics="true" pageId="3" pageNumber="66">A. tambopatensis</emphasis>
|
||
</taxonomicName>
|
||
; 17–20)
|
||
<taxonomicName id="4C744D5CA94EFFFBAF0EF9C2FE0FF9C0" authorityName="Zimmermann" authorityYear="1921" box="[300,406,1607,1630]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="knischi">
|
||
<emphasis id="B900EACDA94EFFFBAF0EF9C2FE0FF9C0" box="[300,406,1607,1630]" italics="true" pageId="3" pageNumber="66">A. knischi</emphasis>
|
||
</taxonomicName>
|
||
. 7, 10, 13, 17) median lobe, right lateral aspect; 8, 11, 14, 18) median lobe, ventral aspect; 9, 12, 15, 19) right lateral lobe, right lateral aspect. 16, 20) left lateral lobe, right lateral aspect.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6554A94EFFFBAEE7F934FDE4F87A" pageId="3" pageNumber="66" type="etymology">
|
||
<paragraph id="8BCB36DFA94EFFFBAEE7F934FB28F952" blockId="3.[151,1436,1713,2020]" box="[197,1201,1713,1740]" pageId="3" pageNumber="66">
|
||
<emphasis id="B900EACDA94EFFFBAEE7F934FECBF955" bold="true" box="[197,338,1713,1739]" pageId="3" pageNumber="66">Etymology.</emphasis>
|
||
This species is named to honor comedian and author, Stephen T. Colbert.
|
||
</paragraph>
|
||
<paragraph id="8BCB36DFA94EFFFBAEE7F95CFDE4F87A" blockId="3.[151,1436,1713,2020]" pageId="3" pageNumber="66">
|
||
<emphasis id="B900EACDA94EFFFBAEE7F95CFD83F96D" bold="true" box="[197,538,1753,1779]" pageId="3" pageNumber="66">Phylogenetic relationships.</emphasis>
|
||
The phylogenetic analysis resulted in a single parsimony tree (
|
||
<figureCitation id="134F2A5AA94EFFFBAB18F95FFA0FF96A" box="[1338,1430,1754,1780]" captionStart="FIGURE 21" captionStartId="4.[151,254,804,828]" captionTargetBox="[530,1057,479,776]" captionTargetId="figure-555@4.[529,1058,474,780]" captionTargetPageId="4" captionText="FIGURE 21. Single most parsimonious cladogram of Agaporomorphus species (length = 13, CI = 92, RI = 94). Numbers above hatch marks refer to characters numbers. Numbers below hatch marks refer to derived character state transformations." figureDoi="http://doi.org/10.5281/zenodo.5133123" httpUri="https://zenodo.org/record/5133123/files/figure.png" pageId="3" pageNumber="66">Fig. 21</figureCitation>
|
||
, length=13, CI=92, RI=94). This topology is identical to the one found by
|
||
<bibRefCitation id="EFE54B2EA94EFFFBADD0F887FB09F882" author="Miller, K. B." box="[1010,1168,1794,1820]" pageId="3" pageNumber="66" pagination="49 - 59" refId="ref3331" refString="Miller, K. B. (2005) Two New Species of Agaporomorphus Zimmermann (Coleoptera: Dytiscidae) from Peru. Zootaxa, 1059, 49 - 59." type="journal article" year="2005">Miller (2005)</bibRefCitation>
|
||
with the addition of
|
||
<taxonomicName id="4C744D5CA94EFFFBABA1F887FF6BF8DD" authority="Miller & Wheeler, 2008" authorityName="Miller & Wheeler" authorityYear="2008" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="colberti" status="sp. nov.">
|
||
<emphasis id="B900EACDA94EFFFBABA1F887FF6BF8DD" italics="true" pageId="3" pageNumber="66">A. colberti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EACDA94EFFFBAEDBF8ACFEA3F8DD" bold="true" box="[249,314,1833,1859]" pageId="3" pageNumber="66">
|
||
<taxonomicNameLabel id="A23357B6A94EFFFBAEDBF8ACFEA3F8DD" box="[249,314,1833,1859]" pageId="3" pageNumber="66" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
as the unambiguous sister to
|
||
<taxonomicName id="4C744D5CA94EFFFBACB2F8AFFC9AF8DD" authorityName="Zimmermann" authorityYear="1921" box="[656,771,1834,1859]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="knischi">
|
||
<emphasis id="B900EACDA94EFFFBACB2F8AFFC9AF8DD" box="[656,771,1834,1859]" italics="true" pageId="3" pageNumber="66">A. knischi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="134F2A5AA94EFFFBAD30F8AFFCFEF8DA" box="[786,871,1834,1860]" captionStart="FIGURE 21" captionStartId="4.[151,254,804,828]" captionTargetBox="[530,1057,479,776]" captionTargetId="figure-555@4.[529,1058,474,780]" captionTargetPageId="4" captionText="FIGURE 21. Single most parsimonious cladogram of Agaporomorphus species (length = 13, CI = 92, RI = 94). Numbers above hatch marks refer to characters numbers. Numbers below hatch marks refer to derived character state transformations." figureDoi="http://doi.org/10.5281/zenodo.5133123" httpUri="https://zenodo.org/record/5133123/files/figure.png" pageId="3" pageNumber="66">Fig. 21</figureCitation>
|
||
) based on the synapomorphy of modified antennomere V in addition to VI (e.g.
|
||
<figureCitation id="134F2A5AA94EFFFBAC0CF8D7FDE1F8F2" box="[558,632,1874,1900]" captionStart="FIGURES" captionStartId="2.[151,203,1333,1359]" captionTargetBox="[543,1054,647,1303]" captionTargetId="figure-459@2.[529,1059,637,1306]" captionTargetPageId="2" captionText="FIGURES 1–6. Agaporomorphus species, morphological details. 1, 5) A. colberti; 2) A. silvaticus; 3) A. tambopatensis; 4, 6) A. knischi. 1) dorsal habitus; 2–4) male left antenna, ventral aspect; 5, 6) visible abdominal sternites, ventral aspect. Scale bars; a = 1mm for Figs 1,5,6; b = 0.25mm for Figs 2–4." figureDoi="http://doi.org/10.5281/zenodo.5133121" httpUri="https://zenodo.org/record/5133121/files/figure.png" pageId="3" pageNumber="66">Fig. 4</figureCitation>
|
||
,
|
||
<taxonomicName id="4C744D5CA94EFFFBACAAF8D7FCCCF8F5" authorityName="Miller" authorityYear="2005" box="[648,853,1874,1899]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="tambopatensis">
|
||
<emphasis id="B900EACDA94EFFFBACAAF8D7FCCCF8F5" box="[648,853,1874,1899]" italics="true" pageId="3" pageNumber="66">A. tambopatensis</emphasis>
|
||
</taxonomicName>
|
||
also has antennomere VI modified, though to a lesser extent,
|
||
<figureCitation id="134F2A5AA94EFFFBAF19F8FFFE1DF80A" box="[315,388,1914,1940]" captionStart="FIGURES" captionStartId="2.[151,203,1333,1359]" captionTargetBox="[543,1054,647,1303]" captionTargetId="figure-459@2.[529,1059,637,1306]" captionTargetPageId="2" captionText="FIGURES 1–6. Agaporomorphus species, morphological details. 1, 5) A. colberti; 2) A. silvaticus; 3) A. tambopatensis; 4, 6) A. knischi. 1) dorsal habitus; 2–4) male left antenna, ventral aspect; 5, 6) visible abdominal sternites, ventral aspect. Scale bars; a = 1mm for Figs 1,5,6; b = 0.25mm for Figs 2–4." figureDoi="http://doi.org/10.5281/zenodo.5133121" httpUri="https://zenodo.org/record/5133121/files/figure.png" pageId="3" pageNumber="66">Fig. 3</figureCitation>
|
||
). Although not otherwise coded in this analysis because of challenges in homologizing complex shape features,
|
||
<taxonomicName id="4C744D5CA94EFFFBAF94F827FDA8F825" authority="Miller & Wheeler, 2008" authorityName="Miller & Wheeler" authorityYear="2008" box="[438,561,1954,1979]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="colberti" status="sp. nov.">
|
||
<emphasis id="B900EACDA94EFFFBAF94F827FDA8F825" box="[438,561,1954,1979]" italics="true" pageId="3" pageNumber="66">A. colberti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EACDA94EFFFBAC1AF824FDE3F825" bold="true" box="[568,634,1953,1979]" pageId="3" pageNumber="66">
|
||
<taxonomicNameLabel id="A23357B6A94EFFFBAC1AF824FDE3F825" box="[568,634,1953,1979]" pageId="3" pageNumber="66" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C744D5CA94EFFFBAC93F827FCBDF825" authorityName="Zimmermann" authorityYear="1921" box="[689,804,1954,1979]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="66" phylum="Arthropoda" rank="species" species="knischi">
|
||
<emphasis id="B900EACDA94EFFFBAC93F827FCBDF825" box="[689,804,1954,1979]" italics="true" pageId="3" pageNumber="66">A. knischi</emphasis>
|
||
</taxonomicName>
|
||
exhibit considerable general similarity in the shape of the male median lobes (Figs 7, 8, 15, 18).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6554A949FFFCAEE7FF12FAECFF44" pageId="4" pageNumber="67" type="distribution">
|
||
<paragraph id="8BCB36DFA949FFFCAEE7FF12FAECFF44" blockId="4.[151,1436,151,418]" pageId="4" pageNumber="67">
|
||
<emphasis id="B900EACDA949FFFCAEE7FF12FEFFFF2F" bold="true" box="[197,358,151,177]" pageId="4" pageNumber="67">Distribution.</emphasis>
|
||
<taxonomicName id="4C744D5CA949FFFCAF4CFF1DFD07FF2F" authority="Miller & Wheeler, 2008" authorityName="Miller & Wheeler" authorityYear="2008" box="[366,670,152,177]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="colberti" status="sp. nov.">
|
||
<emphasis id="B900EACDA949FFFCAF4CFF1DFD07FF2F" box="[366,670,152,177]" italics="true" pageId="4" pageNumber="67">Agaporomorphus colberti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B900EACDA949FFFCAC84FF12FD73FF2F" bold="true" box="[678,746,151,177]" pageId="4" pageNumber="67">
|
||
<taxonomicNameLabel id="A23357B6A949FFFCAC84FF12FD73FF2F" box="[678,746,151,177]" pageId="4" pageNumber="67" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is currently known only from the
|
||
<typeStatus id="54CF887DA949FFFCAAA1FF1DFB2CFF2C" box="[1155,1205,152,178]" pageId="4" pageNumber="67">type</typeStatus>
|
||
locality in extreme southern
|
||
<collectingCountry id="F363764FA949FFFCAF21FF45FEE2FF44" box="[259,379,192,218]" name="Venezuela" pageId="4" pageNumber="67">Venezuela</collectingCountry>
|
||
near the border with
|
||
<collectingCountry id="F363764FA949FFFCAC54FF45FD58FF44" box="[630,705,192,218]" name="Brazil" pageId="4" pageNumber="67">Brazil</collectingCountry>
|
||
. This is the only
|
||
<taxonomicName id="4C744D5CA949FFFCADA8FF45FBCFFF47" authorityName="Zimmermann" authorityYear="1921" box="[906,1110,192,217]" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B900EACDA949FFFCADA8FF45FBCFFF47" box="[906,1110,192,217]" italics="true" pageId="4" pageNumber="67">Agaporomorphus</emphasis>
|
||
</taxonomicName>
|
||
known from
|
||
<collectingCountry id="F363764FA949FFFCAAD7FF45FAE8FF44" box="[1269,1393,192,218]" name="Venezuela" pageId="4" pageNumber="67">Venezuela</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6554A949FFFCAEE7FF62FD74FE3C" pageId="4" pageNumber="67" type="materials_examined">
|
||
<paragraph id="8BCB36DFA949FFFCAEE7FF62FD74FE3C" blockId="4.[151,1436,151,418]" pageId="4" pageNumber="67">
|
||
<emphasis id="B900EACDA949FFFCAEE7FF62FE22FE9F" bold="true" box="[197,443,231,257]" pageId="4" pageNumber="67">Material examined.</emphasis>
|
||
<materialsCitation id="3B1C3C82A949FFFCAFE7FF6DFB9CFECC" collectionCode="MIZA" collectorName="L. J. Joly A. Chacon" country="Venezuela" county="Dpt Rio Negro" elevation="140" latitude="0.9166667" location="Rio Baria" longLatPrecision="1308" longitude="-66.166664" municipality="Rio Negro" pageId="4" pageNumber="67" specimenCode="II-8" specimenCount="141" specimenCount-male="1" stateProvince="Amazonas" typeStatus="holotype">
|
||
<typeStatus id="54CF887DA949FFFCAFE7FF6DFDFDFE9C" box="[453,612,232,258]" pageId="4" pageNumber="67" type="holotype">HOLOTYPE</typeStatus>
|
||
:
|
||
<specimenCount id="9D72FD56A949FFFCAC51FF6CFD1EFF65" box="[627,647,233,251]" pageId="4" pageNumber="67" type="male">♂</specimenCount>
|
||
in
|
||
<collectionCode id="ED65AE1AA949FFFCAC8CFF6DFD60FE9C" box="[686,761,232,258]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="4" pageNumber="67" type="Museum">MIZA</collectionCode>
|
||
labeled, “
|
||
<collectingCountry id="F363764FA949FFFCAD53FF6DFC75FE9C" box="[881,1004,232,258]" name="Venezuela" pageId="4" pageNumber="67">Venezuela</collectingCountry>
|
||
T-F
|
||
<collectingRegion id="49B0F83DA949FFFCAA0BFF6DFB3DFE9C" box="[1065,1188,232,258]" country="Venezuela" name="Amazonas" pageId="4" pageNumber="67">Amazonas</collectingRegion>
|
||
<collectingCounty id="62AA4E53A949FFFCAA8FFF6DFAC6FE9C" box="[1197,1375,232,258]" pageId="4" pageNumber="67">
|
||
Dpt
|
||
<collectingMunicipality id="6BAFACA5A949FFFCAAC1FF6DFAC6FE9C" box="[1251,1375,232,258]" pageId="4" pageNumber="67">Rio Negro</collectingMunicipality>
|
||
</collectingCounty>
|
||
/
|
||
<location id="8EAB6004A949FFFCAB53FF6DFF4EFEB4" LSID="urn:lsid:plazi:treatment:03DD87C9A94CFFFCAEB5F948FD74FE3C:8EAB6004A949FFFCAB53FF6DFF4EFEB4" country="Venezuela" county="Dpt Rio Negro" latitude="0.9166667" longLatPrecision="1308" longitude="-66.166664" municipality="Rio Negro" name="Rio Baria" pageId="4" pageNumber="67" stateProvince="Amazonas">Rio Baria</location>
|
||
<specimenCount id="9D72FD56A949FFFCAEFDFE95FEB3FEB4" box="[223,298,272,298]" pageId="4" pageNumber="67" type="generic">
|
||
<quantity id="4C8C9B3AA949FFFCAEFDFE95FEBCFEB4" box="[223,293,272,298]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="67" unit="m" value="140.0">
|
||
<elevation id="0059D1ECA949FFFCAEFDFE95FEBCFEB4" box="[223,293,272,298]" metricMagnitude="2" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="67" unit="m" value="140.0">140m</elevation>
|
||
</quantity>
|
||
.
|
||
</specimenCount>
|
||
<geoCoordinate id="EE405018A949FFFCAF11FE95FE10FEB4" box="[307,393,272,298]" degrees="0" direction="north" minutes="55" orientation="latitude" pageId="4" pageNumber="67" precision="925" value="0.9166667">0°55’N</geoCoordinate>
|
||
<geoCoordinate id="EE405018A949FFFCAFB0FE95FE65FEB4" box="[402,508,272,298]" degrees="66" direction="west" minutes="10" orientation="longitude" pageId="4" pageNumber="67" precision="925" value="-66.166664">66°10’W</geoCoordinate>
|
||
/
|
||
<collectorName id="26815309A949FFFCAC2DFE95FD72FEB4" box="[527,747,272,298]" pageId="4" pageNumber="67">L.J.Joly A.Chacon</collectorName>
|
||
4-11-
|
||
<specimenCode id="DBD29EA4A949FFFCAD12FE95FCC7FEB4" box="[816,862,272,298]" collectionCode="II-" pageId="4" pageNumber="67">II-8</specimenCode>
|
||
[date handwritten]/
|
||
<typeStatus id="54CF887DA949FFFCAA70FE95FB74FEB4" box="[1106,1261,272,298]" pageId="4" pageNumber="67" type="holotype">HOLOTYPE</typeStatus>
|
||
:
|
||
<taxonomicName id="4C744D5CA949FFFCAADFFE95FDFFFECC" authority="Miller and Wheeler, 2008" authorityName="Miller and Wheeler" authorityYear="2008" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="colberti">
|
||
<emphasis id="B900EACDA949FFFCAADFFE95FEA8FECF" italics="true" pageId="4" pageNumber="67">Agaporomorphus colberti</emphasis>
|
||
Miller and Wheeler, 2008
|
||
</taxonomicName>
|
||
[red label with black line border].”
|
||
</materialsCitation>
|
||
<materialsCitation id="3B1C3C82A949FFFCAA2FFEBDFD74FE3C" collectionCode="MIZA, USNM, MSBA" country="Venezuela" pageId="4" pageNumber="67" specimenCount="3" stateProvince="Amazonas" typeStatus="paratype">
|
||
<typeStatus id="54CF887DA949FFFCAA2FFEBDFB21FECC" box="[1037,1208,312,338]" pageId="4" pageNumber="67" type="paratype">PARATYPES</typeStatus>
|
||
, 10 total, in
|
||
<collectionCode id="ED65AE1AA949FFFCAB6BFEBDFA0EFECC" box="[1353,1431,312,338]" country="Venezuela" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33834" lsid="urn:lsid:biocol.org:col:33834" name="Museo del Instituto de Zoologia Agricola Francisco Fernandez Yepez" pageId="4" pageNumber="67" type="Museum">MIZA</collectionCode>
|
||
,
|
||
<collectionCode id="ED65AE1AA949FFFCAEB5FEE5FF75FEE4" box="[151,236,352,378]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" lsid="urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="67" type="Museum">USNM</collectionCode>
|
||
and
|
||
<collectionCode id="ED65AE1AA949FFFCAF0AFEE5FEE2FEE4" box="[296,379,352,378]" pageId="4" pageNumber="67">MSBA</collectionCode>
|
||
labeled same as
|
||
<typeStatus id="54CF887DA949FFFCAC68FEE5FD29FEE4" box="[586,688,352,378]" pageId="4" pageNumber="67" type="holotype">holotype</typeStatus>
|
||
except “…/
|
||
<typeStatus id="54CF887DA949FFFCAD69FEE5FC78FEE4" box="[843,993,352,378]" pageId="4" pageNumber="67" type="paratype">PARATYPE</typeStatus>
|
||
:
|
||
<taxonomicName id="4C744D5CA949FFFCADCFFEE5FEDBFE3C" authority="Miller and Wheeler, 2008" authorityName="Miller and Wheeler" authorityYear="2008" class="Insecta" family="Dytiscidae" genus="Agaporomorphus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="67" phylum="Arthropoda" rank="species" species="coberti">
|
||
<emphasis id="B900EACDA949FFFCADCFFEE5FA8FFEE7" box="[1005,1302,352,377]" italics="true" pageId="4" pageNumber="67">Agaporomorphus coberti</emphasis>
|
||
Miller and Wheeler, 2008
|
||
</taxonomicName>
|
||
[blue label with black line border].”
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |