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<document ID-DOI="10.3853/j.2201-4349.69.2017.1653" ID-ISSN="2201-4349" ID-Zenodo-Dep="5237800" ID-ZooBank="68F315FF-3FEB-410E-96EC-5F494510F440" approvalRequired="186" approvalRequired_for_taxonomicNames="6" approvalRequired_for_textStreams="100" approvalRequired_for_treatments="80" checkinTime="1629535407606" checkinUser="felipe" docAuthor="Parnaby, Harry E., Ingleby, Sandy &amp; Divljan, Anja" docDate="2017" docId="03DD87C8FFFD73731B6EFEFDFD889206" docLanguage="en" docName="RecAustMus.69.5.277-420.pdf" docOrigin="Records of the Australian Museum 69 (5)" docSource="http://dx.doi.org/10.3853/j.2201-4349.69.2017.1653" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Canis familiaris var. papuensis Ramsay 1879" docType="treatment" docVersion="3" lastPageNumber="347" masterDocId="FFE4FFB0FFB973351832FFEBFFD5973C" masterDocTitle="Type Specimens of Non-fossil Mammals in the Australian Museum, Sydney" masterLastPageNumber="420" masterPageNumber="277" pageNumber="345" updateTime="1674153641407" updateUser="juliana" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title>Type Specimens of Non-fossil Mammals in the Australian Museum, Sydney</mods:title>
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<mods:namePart>Parnaby, Harry E.</mods:namePart>
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<mods:namePart>Ingleby, Sandy</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Divljan, Anja</mods:namePart>
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<mods:date>2017</mods:date>
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<mods:number>2017-10-06</mods:number>
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<mods:number>69</mods:number>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.5238002" ID-Zenodo-Dep="5238002" LSID="urn:lsid:plazi:treatment:03DD87C8FFFD73731B6EFEFDFD889206" httpUri="http://treatment.plazi.org/id/03DD87C8FFFD73731B6EFEFDFD889206" lastPageId="70" lastPageNumber="347" pageId="68" pageNumber="345">
<subSubSection pageId="68" pageNumber="345" type="nomenclature">
<paragraph blockId="68.[860,1446,278,305]" box="[860,1446,278,305]" pageId="68" pageNumber="345">
<heading box="[860,1446,278,305]" centered="true" fontSize="11" level="2" pageId="68" pageNumber="345" reason="2">
<taxonomicName authority="Ramsay, 1879 b" authorityName="Ramsay" authorityYear="1879" box="[860,1446,278,305]" class="Mammalia" family="Canidae" genus="Canis" kingdom="Animalia" order="Carnivora" pageId="68" pageNumber="345" phylum="Chordata" rank="variety" species="familiaris" variety="papuensis">
<emphasis bold="true" box="[860,1446,278,305]" pageId="68" pageNumber="345">
<emphasis bold="true" box="[860,1059,278,305]" italics="true" pageId="68" pageNumber="345">Canis familiaris</emphasis>
var.
<emphasis bold="true" box="[1122,1245,278,304]" italics="true" pageId="68" pageNumber="345">papuensis</emphasis>
<bibRefCitation author="Ramsay, E" box="[1254,1446,278,305]" pageId="68" pageNumber="345" pagination="241 - 305" refId="ref142705" refString="Ramsay, E. P. 1879 b. Contributions to the zoology of New Guinea. Parts I and II. Proceedings of the Linnean Society of New South Wales (series 1) 3 (3): 241 - 305." type="journal article" year="1879">Ramsay, 1879b</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph blockId="68.[869,1435,325,347]" box="[869,1435,325,347]" pageId="68" pageNumber="345">
<emphasis box="[869,1151,325,346]" italics="true" pageId="68" pageNumber="345">Proc. Linn. Soc. N.S.W. (ser.1)</emphasis>
3(3): 242. (?Jan?
<date box="[1317,1425,325,347]" pageId="68" pageNumber="345" value="1879-04">April 1879</date>
).
</paragraph>
</subSubSection>
<subSubSection pageId="68" pageNumber="345" type="description">
<paragraph blockId="68.[839,1260,368,393]" box="[839,1260,368,393]" pageId="68" pageNumber="345">
<emphasis bold="true" box="[839,1013,368,392]" pageId="68" pageNumber="345">Common name</emphasis>
.?Papuan Village Dog.
</paragraph>
<paragraph blockId="68.[838,1467,413,555]" pageId="68" pageNumber="345">
<emphasis bold="true" box="[839,999,413,438]" pageId="68" pageNumber="345">Current name</emphasis>
.
<taxonomicName authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[1011,1353,413,438]" class="Mammalia" family="Canidae" genus="Canis" kingdom="Animalia" order="Carnivora" pageId="68" pageNumber="345" phylum="Chordata" rank="species" species="familiaris">
<emphasis box="[1011,1182,414,438]" italics="true" pageId="68" pageNumber="345">Canis familiaris</emphasis>
<bibRefCitation author="Linnaeus, C" box="[1187,1353,413,438]" pageId="68" pageNumber="345" refId="ref137371" refString="Linnaeus, C. 1758. Systema Naturae per regna tria naturae, secundum classes, ordines, genera, species, cum characteribus, differentiis, synonymis, locis. Editio decima, reformata. Tome 1. Holmiae: Laurentii Salvii. 824 pp. https: // doi. org / 10.5962 / bhl. title. 542" type="book" year="1758">Linnaeus, 1758</bibRefCitation>
</taxonomicName>
; following rationale of Jackson &amp; Groves (2015: 288) and
<bibRefCitation author="Jackson, S &amp; Groves, P &amp; Fleming, K &amp; Aplin, M &amp; Eldridge, A" pageId="68" pageNumber="345" pagination="201 - 224" refId="ref133800" refString="Jackson, S. M., C. P. Groves, P. J. S. Fleming, K. P. Aplin, M. D. B. Eldridge, A. Gonzalez, and K. M. Helgen. 2017. The Wayward Dog: is the Australian native dog or Dingo a distinct species? Zootaxa: 4317 (2): 201 - 224. https: // doi. org / 10.11646 / zootaxa. 4317.2.1" type="journal article" year="2017">
Jackson
<emphasis italics="true" pageId="68" pageNumber="345">et al.</emphasis>
(2017)
</bibRefCitation>
; also see Dwyer &amp; Minnegal (2016) for a review of the relationship between New Guinean village and feral dog populations.
</paragraph>
</subSubSection>
<subSubSection pageId="68" pageNumber="345" type="materials_examined">
<paragraph blockId="68.[837,1468,576,1099]" pageId="68" pageNumber="345">
<materialsCitation collectingDate="1878-12" collectionCode="AM" collectorName="A Register. No" country="Guinea-Bissau" county="Troughton's" location="Annual" municipality="Alternatively" pageId="68" pageNumber="345" specimenCount="2" typeStatus="holotype">
<emphasis bold="true" box="[839,944,576,600]" pageId="68" pageNumber="345">
<typeStatus box="[839,944,576,600]" pageId="68" pageNumber="345">Holotype</typeStatus>
</emphasis>
. A.3652, designated here. Skull (
<figureCitation box="[1323,1407,576,600]" captionStart="Figure 28" captionStartId="69.[161,224,1928,1950]" captionTargetBox="[269,1273,155,1879]" captionTargetId="figure-9@69.[240,1302,151,1882]" captionTargetPageId="69" captionText="Figure 28. AM A.3652, holotype skull of Canis familiaris var. papuensis Ramsay, 1879b. (Photography by Stuart Humphreys)." figureDoi="http://doi.org/10.5281/zenodo.5237862" httpUri="https://zenodo.org/record/5237862/files/figure.png" pageId="68" pageNumber="345">Fig. 28</figureCitation>
), sex not recorded, purchased from Andrew Goldie, registered
<date box="[839,1013,635,659]" pageId="68" pageNumber="345" value="1878-12">
<collectingDate box="[839,1013,635,659]" pageId="68" pageNumber="345" value="1878-12">December 1878</collectingDate>
</date>
. Collection date and collector not given in the
<collectorName box="[879,1042,664,688]" pageId="68" pageNumber="345">A Register. No</collectorName>
specific locality is entered against this specimen, but an earlier register entry indicates that it was part of a large collection from the “Port Moresby district” purchased from Andrew Goldie. In his original account, Ramsay states that the specimen was obtained on Andrew Goldies second expedition to New
<collectingCountry box="[1227,1308,811,835]" name="Guinea-Bissau" pageId="68" pageNumber="345">Guinea</collectingCountry>
. No associated skeletal elements or skin have yet been located in the collection but it is possible that the
<typeStatus box="[1215,1308,870,894]" pageId="68" pageNumber="345">holotype</typeStatus>
was originally a mounted skeleton, as “1 dog, from New
<collectingCountry box="[1289,1368,899,923]" name="Guinea-Bissau" pageId="68" pageNumber="345">Guinea</collectingCountry>
” is listed amongst mammal specimens articulated during
<quantity box="[1380,1466,928,952]" metricMagnitude="1" metricUnit="m" metricValue="4.77774" pageId="68" pageNumber="345" unit="in" value="1881.0">1881 in</quantity>
the
<collectionCode box="[878,921,958,982]" country="Australia" name="Australian Museum" pageId="68" pageNumber="345" type="Museum">AM</collectionCode>
<location LSID="urn:lsid:plazi:treatment:03DD87C8FFFD73731B6EFEFDFD889206:8EAB6005FFFD73711BADFC55FC3B94EA" box="[927,1006,958,982]" country="Guinea-Bissau" county="Troughton's" municipality="Alternatively" name="Annual" pageId="68" pageNumber="345">Annual</location>
report (
<bibRefCitation author="Ramsay, E" box="[1093,1263,958,982]" pageId="68" pageNumber="345" pagination="17 - 18" refId="ref142921" refString="Ramsay, E. P. 1882 c. Appendix XI, List of specimens cured and mounted and other work done by the taxidermists and articulator. In Report of the Trustees for 1881. Australian Museum, pp. 17 - 18. Sydney, NSW: Thomas Richards, Government Printer." type="book chapter" year="1882">Ramsay, 1882c</bibRefCitation>
).
<collectingMunicipality box="[1280,1422,958,982]" pageId="68" pageNumber="345">Alternatively</collectingMunicipality>
, the mounted specimen may have been a specimen acquired from another source, possibly from Miklouho-Maclay. The skull has the registration number written in what is likely to be
<collectingCounty box="[841,970,1075,1099]" pageId="68" pageNumber="345">Troughtons</collectingCounty>
handwriting.
</materialsCitation>
</paragraph>
<paragraph blockId="68.[837,1467,1123,1264]" pageId="68" pageNumber="345">
<emphasis bold="true" box="[839,960,1123,1147]" pageId="68" pageNumber="345">Condition</emphasis>
. Cranium missing upper right 1st and 2nd premolars (possibly lost while the animal was alive), missing medial side of right auditory bulla, anterior tips of nasal bones eroded, hole in occipital. Left dentary has two holes in ramus, right dentary complete.
</paragraph>
<paragraph blockId="68.[837,1467,1285,1456]" pageId="68" pageNumber="345">
<emphasis bold="true" box="[839,1199,1285,1310]" pageId="68" pageNumber="345">Cranial measurements (mm)</emphasis>
. A.3652: GL, 165.25; ConL,154.36; BasL, 147.39; NasL, 50.78*; NasB, 17.22; UC1C1 (alv.), 30.25; APV, 10.58; PAL, 82.42; UPM (alv.), 16.73; UMR (alv.), 16.32; ZB, 88.73; POC, 24.86; BUL, 19.95; MB, 58.74; DL (condyl.), 123.28; LPM (alv.), 10.14; LMR (alv.), 29.92. [* = estimate, nasal tips broken].
</paragraph>
<paragraph blockId="68.[839,1465,1480,1534]" pageId="68" pageNumber="345">
<emphasis bold="true" box="[839,994,1480,1504]" pageId="68" pageNumber="345">Type locality</emphasis>
.
<collectingRegion box="[1012,1256,1480,1504]" country="Papua New Guinea" name="National Capital" pageId="68" pageNumber="345">Port Moresby district</collectingRegion>
(
<bibRefCitation author="Ramsay, E" box="[1274,1454,1480,1504]" pageId="68" pageNumber="345" pagination="241 - 305" refId="ref142705" refString="Ramsay, E. P. 1879 b. Contributions to the zoology of New Guinea. Parts I and II. Proceedings of the Linnean Society of New South Wales (series 1) 3 (3): 241 - 305." type="journal article" year="1879">Ramsay, 1879b</bibRefCitation>
), Central Province,
<collectingCountry box="[1032,1242,1510,1534]" name="Papua New Guinea" pageId="68" pageNumber="345">Papua New Guinea</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="70" lastPageNumber="347" pageId="68" pageNumber="345" type="discussion">
<paragraph blockId="68.[838,1467,1557,1992]" pageId="68" pageNumber="345">
<emphasis bold="true" box="[839,967,1557,1581]" pageId="68" pageNumber="345">Comments</emphasis>
. Volume 3 issue 3 of the
<emphasis italics="true" pageId="68" pageNumber="345">Proc. Linn. Soc. N.S.W.</emphasis>
is the only issue for which
<bibRefCitation author="Fletcher, J" box="[1197,1362,1587,1611]" pageId="68" pageNumber="345" pagination="533 - 536" refId="ref129679" refString="Fletcher, J. J. 1896. On the dates of publication of the early volumes of the Society's Proceedings. Proceedings of the Linnean Society of New South Wales (series 2) 10 (4): 533 - 536. https: // doi. org / 10.5962 / bhl. part. 24365" type="journal article" year="1896">Fletcher (1896)</bibRefCitation>
could not determine the publication date; it was probably between
<date box="[839,1047,1646,1670]" pageId="68" pageNumber="345" value="1879-01" valueMax="1879-05-31" valueMin="1879-01-00">JanuaryMay 1879</date>
, based on publication dates of conjoint issues.
</paragraph>
<paragraph blockId="68.[838,1467,1557,1992]" lastBlockId="70.[166,794,152,264]" lastPageId="70" lastPageNumber="347" pageId="68" pageNumber="345">
Previous assessments of the status of this taxon have presumably been made without examination of Ramsays original material that has remained unreported in the AM Collection since its description.
<bibRefCitation author="Troughton, E" box="[1182,1371,1792,1816]" pageId="68" pageNumber="345" pagination="93 - 98" refId="ref148095" refString="Troughton, E. [L. G.] 1971. The early history and relationships of the New Guinea Highland dog (Canis hallstromi). Proceedings of the Linnean Society of New South Wales 96 (2): 93 - 98." type="journal article" year="1971">Troughton (1971)</bibRefCitation>
reported that Ramsays specimen had been destroyed but we are confident that the dog skull labelled A.3652 is the specimen reported by Ramsay. Although he did not specify the collection locality, sex, or provide any measurements of the specimen, Ramsay stated that it was obtained from Goldie and his wording implies a single specimen. Only one “
<taxonomicName class="Mammalia" family="Canidae" genus="Canis" kingdom="Animalia" lastPageId="70" lastPageNumber="347" order="Carnivora" pageId="68" pageNumber="345" phylum="Chordata" rank="species" species="papuensis">
<emphasis italics="true" lastPageId="70" lastPageNumber="347" pageId="68" pageNumber="345">Canis papuensis</emphasis>
</taxonomicName>
” is listed in the AM specimen acquisitions for 1878, purchased from Goldie and listed, remarkably, under “Collection of fish specimens from New
<collectingCountry box="[592,669,210,234]" name="Guinea-Bissau" pageId="70" pageNumber="347">Guinea</collectingCountry>
(in spirits)” (
<bibRefCitation author="Ramsay, E" box="[173,342,240,264]" pageId="70" pageNumber="347" pagination="11 - 15" refId="ref142749" refString="Ramsay, E. P. 1879 c. Appendix VIII, Specimens purchased. In Report of the Trustees for the year ending 31 December, 1878. Australian Museum, pp. 11 - 15. Sydney, NSW: Thomas Richards, Government Printer." type="book chapter" year="1879">Ramsay, 1879c</bibRefCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5237862" ID-Zenodo-Dep="5237862" box="[161,1382,1928,1950]" httpUri="https://zenodo.org/record/5237862/files/figure.png" pageId="69" pageNumber="346" startId="69.[161,224,1928,1950]" targetBox="[269,1273,155,1879]" targetPageId="69">
<paragraph blockId="69.[161,1382,1928,1950]" box="[161,1382,1928,1950]" pageId="69" pageNumber="346">
Figure 28. AM A.3652, holotype skull of
<taxonomicName authority="Ramsay, 1879 b" authorityName="Ramsay" authorityYear="1879" box="[560,1016,1928,1950]" class="Mammalia" family="Canidae" genus="Canis" kingdom="Animalia" order="Carnivora" pageId="69" pageNumber="346" phylum="Chordata" rank="variety" species="familiaris" variety="papuensis">
<emphasis box="[560,715,1928,1950]" italics="true" pageId="69" pageNumber="346">Canis familiaris</emphasis>
var.
<emphasis box="[763,859,1929,1950]" italics="true" pageId="69" pageNumber="346">papuensis</emphasis>
<bibRefCitation author="Ramsay, E" box="[865,1016,1928,1950]" pageId="69" pageNumber="346" pagination="241 - 305" refId="ref142705" refString="Ramsay, E. P. 1879 b. Contributions to the zoology of New Guinea. Parts I and II. Proceedings of the Linnean Society of New South Wales (series 1) 3 (3): 241 - 305." type="journal article" year="1879">Ramsay, 1879b</bibRefCitation>
</taxonomicName>
. (Photography by Stuart Humphreys).
</paragraph>
</caption>
<paragraph blockId="70.[164,794,285,720]" pageId="70" pageNumber="347">
Specimen A.3652 was registered in
<date box="[548,717,285,309]" pageId="70" pageNumber="347" value="1878-12">December 1878</date>
as part of a large consignment of specimens in spirit purchased from Goldie; these were entered in the register under the heading “Collection of spirit specimensAndrew Goldie not in the best state” and many were later marked as “destroyed”. Taken along with Ramsays comment that the dog specimen was received from Goldie in a poor state of preservation in spirits, these entries may have influenced
<bibRefCitation author="Troughton, E" box="[525,713,490,514]" pageId="70" pageNumber="347" pagination="93 - 98" refId="ref148095" refString="Troughton, E. [L. G.] 1971. The early history and relationships of the New Guinea Highland dog (Canis hallstromi). Proceedings of the Linnean Society of New South Wales 96 (2): 93 - 98." type="journal article" year="1971">Troughton (1971)</bibRefCitation>
to state that Ramsays
<typeStatus box="[332,426,520,544]" pageId="70" pageNumber="347">holotype</typeStatus>
was no longer in the collection. However, A.3652 was actually marked as being transferred to the Palaeontology Dept. (no date given) where it was found in 2012 during preparation of this paper. Troughton (1957) initially treated
<taxonomicName box="[412,519,638,661]" class="Mammalia" family="Canidae" genus="Canis" kingdom="Animalia" order="Carnivora" pageId="70" pageNumber="347" phylum="Chordata" rank="species" species="papuensis">
<emphasis box="[412,519,638,661]" italics="true" pageId="70" pageNumber="347">papuensis</emphasis>
</taxonomicName>
as a
<emphasis box="[571,722,637,661]" italics="true" pageId="70" pageNumber="347">nomen nudum</emphasis>
but he later equivocated and even commented that the name had been ignored by most authors (
<bibRefCitation author="Troughton, E" box="[498,681,696,720]" pageId="70" pageNumber="347" pagination="93 - 98" refId="ref148095" refString="Troughton, E. [L. G.] 1971. The early history and relationships of the New Guinea Highland dog (Canis hallstromi). Proceedings of the Linnean Society of New South Wales 96 (2): 93 - 98." type="journal article" year="1971">Troughton, 1971</bibRefCitation>
).
</paragraph>
<paragraph blockId="70.[164,794,741,882]" pageId="70" pageNumber="347">
The ratio zygomatic breadth/condylobasal length (87.8/
<quantity metricMagnitude="-1" metricUnit="m" metricValue="1.685" pageId="70" pageNumber="347" unit="mm" value="168.5">168.5 mm</quantity>
) of 0.52 for A.3652 falls below the range 0.640.74 given for
<bibRefCitation author="Koler-Matznick, J &amp; Brisbin, Jr, M" box="[279,602,800,824]" pageId="70" pageNumber="347" pagination="109 - 118" refId="ref134761" refString="Koler-Matznick, J., I. L. Brisbin, Jr, M. Feinstein, and S. Bulmer. 2003. An updated description of the New Guinea singing dog (Canis hallstromi, Troughton 1957). Journal of Zoology (London) 261: 109 - 118. https: // doi. org / 10.1017 / S 0952836903004060" type="journal article" year="2003">
Koler-Matznick
<emphasis box="[462,520,800,824]" italics="true" pageId="70" pageNumber="347">et al.</emphasis>
(2003)
</bibRefCitation>
for New
<collectingCountry box="[716,794,800,824]" name="Guinea-Bissau" pageId="70" pageNumber="347">Guinea</collectingCountry>
Singing Dogs “
<taxonomicName authorityName="Troughton" authorityYear="1957" box="[327,472,829,853]" class="Mammalia" family="Canidae" genus="Canis" kingdom="Animalia" order="Carnivora" pageId="70" pageNumber="347" phylum="Chordata" rank="species" species="hallstromi">
<emphasis box="[327,345,829,852]" italics="true" pageId="70" pageNumber="347">C</emphasis>
.
<emphasis box="[356,472,829,853]" italics="true" pageId="70" pageNumber="347">hallstromi</emphasis>
</taxonomicName>
<emphasis box="[472,479,829,853]" italics="true" pageId="70" pageNumber="347"></emphasis>
and it is also below 0.610.62 given by those authors for Papuan village dogs.
</paragraph>
<paragraph blockId="70.[164,796,904,1337]" pageId="70" pageNumber="347">
Ramsay stated that his knowledge of this taxon, including its small body size and inability to bark, drew from the observations of Morton and Broadbent about Papuan dogs. Although it is not known who collected the
<typeStatus box="[628,719,992,1016]" pageId="70" pageNumber="347">holotype</typeStatus>
during Goldies expedition, it is possible that the specimen came from a moderate to high elevation in the Owen Stanley Range behind Port Moresby, given that collecting occurred quite early over a wide range of altitudes (
<bibRefCitation author="Moore, C" box="[441,697,1109,1133]" pageId="70" pageNumber="347" pagination="39 - 127" refId="ref139377" refString="Moore, C., and S. Mullins. 2012. Andrew Goldie's memoir: 1875 - 1879. Memoirs of the Queensland Museum - Culture 6: 39 - 127." type="journal article" year="2012">Moore &amp; Mullins, 2012</bibRefCitation>
;
<bibRefCitation author="Morton, A" pageId="70" pageNumber="347" pagination="65 - 84" refId="ref139469" refString="Morton, A. 1885 a. Notes of a trip to the Islands of Torres Straits and the south-east coast of New Guinea. Proceedings of the Geographic Society of Australasia, New South Wales and Victorian Branches. 1: 65 - 84, 3 pls." type="journal article" year="1885">Morton, 1885a</bibRefCitation>
). In an account of his work with Goldie,
<bibRefCitation author="Morton, A" box="[633,795,1138,1162]" pageId="70" pageNumber="347" pagination="1 - 2" refId="ref139518" refString="Morton, A. 1885 b. Notes on New Guinea - III. The Mercury (Hobart) XLVI (4715): 4, col. 1 - 2, 31 March." type="journal article" year="1885">Morton (1885b)</bibRefCitation>
indicates in relation to “the New
<collectingCountry box="[492,567,1168,1192]" name="Guinea-Bissau" pageId="70" pageNumber="347">Guinea</collectingCountry>
dogs” that “we did not find them in a wild state”, only in villages. However, Broadbent had a separate collecting camp and might have obtained the
<typeStatus box="[167,257,1256,1280]" pageId="70" pageNumber="347">holotype</typeStatus>
as a feral animal. It is possible that the
<typeStatus box="[644,734,1256,1280]" pageId="70" pageNumber="347">holotype</typeStatus>
was a village dog but that Ramsays description reported observations of the singing calls heard by field workers.
</paragraph>
</subSubSection>
</treatment>
</document>