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<document ID-DOI="http://doi.org/10.5281/zenodo.6025478" ID-GBIF-Dataset="a01119e3-5395-4d97-8fa6-bb6742b6722d" ID-GBIF-Taxon="142611027" ID-Zenodo-Dep="6025478" checkinTime="1521998155865" checkinUser="plazi" docAuthor="Diego N. Barbosa, Ricardo Kawada &amp; Celso O. Azevedo" docDate="2017" docId="03DD87B3FF8BFF82FF98B2D829D1FD41" docLanguage="en" docName="322-335 Bethylidae.pdf" docOrigin="Arthropod fauna of the UAE 6" docStyle="DocumentStyle{}" docTitle="Glenosema dura Diego N. Barbosa &amp; Ricardo Kawada &amp; Celso O. Azevedo 2017, sp. nov." docType="treatment" docVersion="5" lastPageId="12" lastPageNumber="334" masterDocId="FFE4FFCBFF81FF8EFFF2B0782A36FFD8" masterDocTitle="Order Hymenoptera, family Bethylidae Genus Glenosema Kieffer" masterLastPageNumber="335" masterPageNumber="322" pageId="10" pageNumber="332" updateTime="1644420597770" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Order Hymenoptera, family Bethylidae Genus Glenosema Kieffer</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Diego N. Barbosa</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ricardo Kawada</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Celso O. Azevedo</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Arthropod fauna of the UAE</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2017</mods:date>
<mods:detail type="volume">
<mods:number>6</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>322</mods:start>
<mods:end>335</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.5281/zenodo.1207201</mods:identifier>
<mods:identifier type="GBIF-Dataset">a01119e3-5395-4d97-8fa6-bb6742b6722d</mods:identifier>
<mods:identifier type="Zenodo-Dep">1207201</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.6025478" ID-GBIF-Taxon="142611027" ID-Zenodo-Dep="6025478" LSID="urn:lsid:plazi:treatment:03DD87B3FF8BFF82FF98B2D829D1FD41" httpUri="http://treatment.plazi.org/id/03DD87B3FF8BFF82FF98B2D829D1FD41" lastPageId="12" lastPageNumber="334" pageId="10" pageNumber="332">
<subSubSection box="[106,705,672,696]" pageId="10" pageNumber="332" type="nomenclature">
<paragraph blockId="10.[106,1112,672,1579]" box="[106,705,672,696]" pageId="10" pageNumber="332">
<taxonomicName authority="Barbosa, Kawada &amp; Azevedo" authorityName="Diego N. Barbosa &amp; Ricardo Kawada &amp; Celso O. Azevedo" authorityYear="2017" box="[106,611,672,696]" class="Insecta" family="Bethylidae" genus="Glenosema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="10" pageNumber="332" phylum="Arthropoda" rank="species" species="dura" status="sp. nov.">
<emphasis bold="true" box="[106,288,672,696]" italics="true" pageId="10" pageNumber="332">Glenosema dura</emphasis>
Barbosa, Kawada &amp; Azevedo
</taxonomicName>
<taxonomicNameLabel box="[618,705,672,696]" pageId="10" pageNumber="332" rank="species">
<emphasis bold="true" box="[618,643,672,696]" pageId="10" pageNumber="332">sp</emphasis>
.
<emphasis bold="true" box="[656,697,672,696]" pageId="10" pageNumber="332">nov</emphasis>
.
</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection box="[974,1111,672,696]" pageId="10" pageNumber="332" type="description">
<paragraph blockId="10.[106,1112,672,1579]" box="[974,1111,672,696]" pageId="10" pageNumber="332">
<figureCitation box="[974,1111,672,696]" captionStart="Plates 1923" captionStartId="11.[136,193,1585,1607]" captionTargetBox="[160,1118,144,1552]" captionTargetId="figure@11.[161,1116,146,1550]" captionText="Plates 1923. Glenosema dura sp. nov., female, holotype. 19: Habitus in lateral view; 20: Mesosoma in dorsal view;21: Head indorsal view; 22:Mesosoma inlateral view; 23:Fore wing in dorsal view." httpUri="https://zenodo.org/record/1207213/files/figure.png" pageId="10" pageNumber="332">Plates 1923</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="332" type="materials_examined">
<paragraph blockId="10.[106,1112,672,1579]" pageId="10" pageNumber="332">
Specimens examined:
<materialsCitation ID-GBIF-Occurrence="1836402782" collectingDate="2000-07" collectionCode="CNCI" collectorName="A. van Harten" country="Yemen" location="Malaise trap," municipality="Lahj" pageId="10" pageNumber="332" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<typeStatus box="[337,430,702,724]" pageId="10" pageNumber="332">Holotype</typeStatus>
: “
<specimenCount box="[457,475,702,723]" pageId="10" pageNumber="332" type="female"></specimenCount>
<collectingCountry box="[488,576,702,723]" name="Yemen" pageId="10" pageNumber="332">YEMEN</collectingCountry>
,
<collectingMunicipality box="[593,640,702,724]" pageId="10" pageNumber="332">Lahj</collectingMunicipality>
,
<collectingDate box="[657,744,702,723]" pageId="10" pageNumber="332" value="2000-07">
<date box="[657,744,702,723]" pageId="10" pageNumber="332" value="2000-07">VII.2000</date>
</collectingDate>
<location LSID="urn:lsid:plazi:treatment:03DD87B3FF8BFF82FF98B2D829D1FD41:8EAB607EFF8BFF84FCFDB2C629A2FD0B" box="[783,916,702,724]" country="Yemen" municipality="Lahj" name="Malaise trap," pageId="10" pageNumber="332">Malaise trap,</location>
<collectorName box="[929,1079,702,723]" pageId="10" pageNumber="332">A. van Harten</collectorName>
&amp; col[lector] (
<collectionCode box="[219,282,730,752]" collectionName="Canada, Ontario, Ottawa, Canadian National Collection of Insects" pageId="10" pageNumber="332">CNCI</collectionCode>
)”.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection lastPageId="12" lastPageNumber="334" pageId="10" pageNumber="332" type="description">
<paragraph blockId="10.[106,1112,672,1579]" box="[106,885,758,782]" pageId="10" pageNumber="332">
Description: Holotype female (
<figureCitation box="[437,529,758,782]" captionStart="Plates 1923" captionStartId="11.[136,193,1585,1607]" captionTargetBox="[160,1118,144,1552]" captionTargetId="figure@11.[161,1116,146,1550]" captionText="Plates 1923. Glenosema dura sp. nov., female, holotype. 19: Habitus in lateral view; 20: Mesosoma in dorsal view;21: Head indorsal view; 22:Mesosoma inlateral view; 23:Fore wing in dorsal view." httpUri="https://zenodo.org/record/1207213/files/figure.png" pageId="10" pageNumber="332">Plate 19</figureCitation>
). Body 3.61 mm. LFW 1.77 mm.
</paragraph>
<paragraph blockId="10.[106,1112,672,1579]" pageId="10" pageNumber="332">
Colours: Head and mesosoma dark castaneous nearly black; metasoma, scape, pedicel, mandible, coxae, trochanters, femora, castaneous; dark castaneous; flagellum, palpi, tegula, tarsi light castaneous; wings clear hyaline with distal spot; wing venation light castaneous Head (
<figureCitation box="[177,270,881,905]" captionStart="Plates 1923" captionStartId="11.[136,193,1585,1607]" captionTargetBox="[160,1118,144,1552]" captionTargetId="figure@11.[161,1116,146,1550]" captionText="Plates 1923. Glenosema dura sp. nov., female, holotype. 19: Habitus in lateral view; 20: Mesosoma in dorsal view;21: Head indorsal view; 22:Mesosoma inlateral view; 23:Fore wing in dorsal view." httpUri="https://zenodo.org/record/1207213/files/figure.png" pageId="10" pageNumber="332">Plate 21</figureCitation>
): Mandible with seven sharp apical teeth, with upper and lower edge of inner margin serrulate, lower corner of right mandible with translucid and very thin subtriangular projection. Clypeus with apical margin rounded, without lateral lobe; median clypeal carina inconspicuous, fused with median lobe, median surface with spine-shaped process. Antennal foramina distance more than antennal foramen diameter. Antenna with insertion as high as clypeal carina, with short and sub-erect pubescence; scape long and curved, with same diameter along its extension; pedicel unusually long, 3.5
<emphasis box="[718,742,1067,1088]" italics="true" pageId="10" pageNumber="332">×</emphasis>
as long as wide, and 3.5
<emphasis box="[1012,1036,1067,1088]" italics="true" pageId="10" pageNumber="332">×</emphasis>
longer than flagellomere I; flagellomeres longer than wide. Eye elliptical, small, placed dorsally. Frons polished, slightly punctured, without setae. First five antennal segments in ratio of about 14:9:4:3:3. LH 1.0
<emphasis box="[396,420,1159,1180]" italics="true" pageId="10" pageNumber="332">×</emphasis>
WH. WF 2.0
<emphasis box="[574,598,1159,1180]" italics="true" pageId="10" pageNumber="332">×</emphasis>
HE. WF 0.66
<emphasis box="[757,781,1159,1180]" italics="true" pageId="10" pageNumber="332">×</emphasis>
WH. OOL 3.6
<emphasis box="[950,974,1159,1180]" italics="true" pageId="10" pageNumber="332">×</emphasis>
WOT. VOL 0.5
<emphasis box="[139,163,1190,1211]" italics="true" pageId="10" pageNumber="332">×</emphasis>
HE. Distance of posterior ocellus to vertex crest 1.5
<emphasis box="[744,768,1190,1211]" italics="true" pageId="10" pageNumber="332">×</emphasis>
DAO. Frontal angle of ocellar triangle right; ocelli very small; ocellar triangle compact. Vertex almost straight. Occipital carina present. Gena polished. Hypostomal carina forming an acute angle. Palpal formula 6:3, segments with circular cross section, equally wide, apices of segments with some short setae. Mesosoma (
<figureCitation box="[238,386,1310,1334]" captionStart="Plates 1923" captionStartId="11.[136,193,1585,1607]" captionTargetBox="[160,1118,144,1552]" captionTargetId="figure@11.[161,1116,146,1550]" captionText="Plates 1923. Glenosema dura sp. nov., female, holotype. 19: Habitus in lateral view; 20: Mesosoma in dorsal view;21: Head indorsal view; 22:Mesosoma inlateral view; 23:Fore wing in dorsal view." httpUri="https://zenodo.org/record/1207213/files/figure.png" pageId="10" pageNumber="332">Plates 20, 22</figureCitation>
): Pronotal disc 1.16
<emphasis box="[607,631,1312,1333]" italics="true" pageId="10" pageNumber="332">×</emphasis>
as long as wide; polished, trapezoidal, side slightly concave; anterior slope of pronotal disc inclined in profile. Mesoscutum coriaceous, as long as mesoscutellum; notaulus absent; parapsidal signum present and conspicuous. Mesoscutellum coriaceous, touching propodeal disc; mesoscutellar sulcus very narrow, arched. Propodeal disc 1.0
<emphasis box="[394,418,1435,1456]" italics="true" pageId="10" pageNumber="332">×</emphasis>
as long as wide; coriaceous, with seven carinae; median carina complete; inner discal carinae incomplete; second pair of discal carina incomplete; outer discal carina complete; sublateral carina absent; lateral carina present and conspicuous; posterior carina complete; spiracle elliptical, placed at lateral carina anteriorly. Declivity of propodeum coriaceous, with median carina. Lateral surface of propodeum strigate.
</paragraph>
<caption httpUri="https://zenodo.org/record/1207213/files/figure.png" pageId="11" pageNumber="333" startId="11.[136,193,1585,1607]" targetBox="[160,1118,144,1552]" targetPageId="11">
<paragraph blockId="11.[136,1142,1585,1634]" pageId="11" pageNumber="333">
Plates 1923.
<taxonomicName authority="Diego N. Barbosa &amp; Ricardo Kawada &amp; Celso O. Azevedo, 2017" authorityName="Diego N. Barbosa &amp; Ricardo Kawada &amp; Celso O. Azevedo" authorityYear="2017" box="[273,433,1585,1606]" class="Insecta" family="Bethylidae" genus="Glenosema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="11" pageNumber="333" phylum="Arthropoda" rank="species" species="dura" status="sp. nov.">
<emphasis box="[273,433,1585,1606]" italics="true" pageId="11" pageNumber="333">Glenosema dura</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[440,519,1585,1606]" pageId="11" pageNumber="333" rank="species">sp. nov.</taxonomicNameLabel>
, female, holotype. 19: Habitus in lateral view; 20: Mesosoma in dorsal view; 21: Head in dorsal view; 22: Mesosoma in lateral view; 23: Fore wing in dorsal view.
</paragraph>
</caption>
<paragraph blockId="12.[106,1112,151,665]" pageId="12" pageNumber="334">Prosternum conspicuous. Mesopleuron with acropleural sulcus continuous with mesopleural suture, without fovea; mesopleural suture with inner margin not striate.</paragraph>
<paragraph blockId="12.[106,1112,151,665]" pageId="12" pageNumber="334">
Wings: Forewing elongate (
<figureCitation box="[422,518,212,236]" captionStart="Plates 1923" captionStartId="11.[136,193,1585,1607]" captionTargetBox="[160,1118,144,1552]" captionTargetId="figure@11.[161,1116,146,1550]" captionText="Plates 1923. Glenosema dura sp. nov., female, holotype. 19: Habitus in lateral view; 20: Mesosoma in dorsal view;21: Head indorsal view; 22:Mesosoma inlateral view; 23:Fore wing in dorsal view." httpUri="https://zenodo.org/record/1207213/files/figure.png" pageId="12" pageNumber="334">Plate 23</figureCitation>
), dorsal face without setae, with three closed cells; median cell almost rectangular; stigma developed and elongate; r-rs &amp; Rs vein medium. Hind wing with costal vein long, dorsal face without setae; with four hamuli.
</paragraph>
<paragraph blockId="12.[106,1112,151,665]" pageId="12" pageNumber="334">Legs: Profemur not dilated. Trochanters clearly strong. Spur of protibia strong. Tarsi elongate. Tarsal claw simple and arched, with small basal tooth.</paragraph>
<paragraph blockId="12.[106,1112,151,665]" pageId="12" pageNumber="334">
Metasoma 2.28
<emphasis box="[272,296,368,389]" italics="true" pageId="12" pageNumber="334">×</emphasis>
longer than wide; coriaceous, tergites III with pair of conspicuous lateral spiracles; sternite II with antero-lateral flap projected to petiole.
</paragraph>
</subSubSection>
<subSubSection pageId="12" pageNumber="334" type="discussion">
<paragraph blockId="12.[106,1112,151,665]" pageId="12" pageNumber="334">
Remarks: This species is similar to the Thai species
<taxonomicName authority="Terayama, 1996" authorityName="Terayama" authorityYear="1996" box="[676,1061,427,451]" class="Insecta" family="Bethylidae" genus="Glenosema" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="334" phylum="Arthropoda" rank="species" species="chiangmaiensis">
<emphasis box="[676,876,427,451]" italics="true" pageId="12" pageNumber="334">G. chiangmaiensis</emphasis>
Terayama, 1996
</taxonomicName>
, and
<taxonomicName box="[106,524,457,482]" pageId="12" pageNumber="334">
<emphasis box="[106,326,457,481]" italics="true" pageId="12" pageNumber="334">G. doiinthanonensis</emphasis>
Terayama, 1996,
</taxonomicName>
by having fully developed wings and spine-shaped clypeal process. However, the spine-shaped clypeal process of the former is much longer than in the two latter. In addition, in
<taxonomicName authority="Diego N. Barbosa &amp; Ricardo Kawada &amp; Celso O. Azevedo, 2017" authorityName="Diego N. Barbosa &amp; Ricardo Kawada &amp; Celso O. Azevedo" authorityYear="2017" box="[458,542,519,543]" class="Insecta" family="Bethylidae" genus="Glenosema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="334" phylum="Arthropoda" rank="species" species="dura" status="sp. nov.">
<emphasis box="[458,542,519,543]" italics="true" pageId="12" pageNumber="334">G. dura</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[551,638,519,542]" pageId="12" pageNumber="334" rank="species">sp. nov.</taxonomicNameLabel>
the pedicel is proportionally longer than in
<taxonomicName box="[106,308,549,573]" class="Insecta" family="Bethylidae" genus="Glenosema" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="334" phylum="Arthropoda" rank="species" species="chiangmaiensis">
<emphasis box="[106,308,549,573]" italics="true" pageId="12" pageNumber="334">G. chiangmaiensis</emphasis>
</taxonomicName>
and
<taxonomicName box="[368,586,549,573]" class="Insecta" family="Bethylidae" genus="Glenosema" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="334" phylum="Arthropoda" rank="species" species="doiinthanonensis">
<emphasis box="[368,586,549,573]" italics="true" pageId="12" pageNumber="334">G. doiinthanonensis</emphasis>
</taxonomicName>
, and the frontal angle ocellar triangle is acute, whereas is right-angled in
<taxonomicName box="[389,587,580,604]" class="Insecta" family="Bethylidae" genus="Glenosema" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="334" phylum="Arthropoda" rank="species" species="chiangmaiensis">
<emphasis box="[389,587,580,604]" italics="true" pageId="12" pageNumber="334">G. chiangmaiensis</emphasis>
</taxonomicName>
and
<taxonomicName box="[639,854,580,604]" class="Insecta" family="Bethylidae" genus="Glenosema" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="334" phylum="Arthropoda" rank="species" species="doiinthanonensis">
<emphasis box="[639,854,580,604]" italics="true" pageId="12" pageNumber="334">G. doiinthanonensis</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection box="[106,490,611,635]" pageId="12" pageNumber="334" type="distribution">
<paragraph blockId="12.[106,1112,151,665]" box="[106,490,611,635]" pageId="12" pageNumber="334">Distribution: United Arab Emirates.</paragraph>
</subSubSection>
<subSubSection box="[106,999,641,665]" pageId="12" pageNumber="334" type="etymology">
<paragraph blockId="12.[106,1112,151,665]" box="[106,999,641,665]" pageId="12" pageNumber="334">
Etymology: The specific epithet
<taxonomicName authority="Latin" authorityName="Latin" box="[456,569,641,665]" class="Insecta" family="Bethylidae" genus="Glenosema" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="12" pageNumber="334" phylum="Arthropoda" rank="species" species="dura">
<emphasis box="[456,507,641,665]" italics="true" pageId="12" pageNumber="334">dura</emphasis>
Latin
</taxonomicName>
refers the spine-shaped clypeal process.
</paragraph>
</subSubSection>
</treatment>
</document>