278 lines
28 KiB
XML
278 lines
28 KiB
XML
<document id="798C29641849353AE57ECE387C7B6C90" ID-DOI="10.5281/zenodo.190015" ID-GBIF-Dataset="a4c97e03-e184-4220-bbc2-ba1f955c580c" ID-ISSN="1175-5326" ID-Zenodo-Dep="190015" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461107024962" checkinUser="plazi" docAuthor="Kohlmann, Bert & Solis, Angel" docDate="2009" docId="03DD8799FFF7852FFF66FB98FBADF84D" docLanguage="en" docName="zt02219p037.pdf" docOrigin="Zootaxa 2219" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Canthidium (Eucanthidium) darwini Kohlmann & Solis, sp. nov." docType="treatment" docVersion="9" lastPageNumber="36" masterDocId="FFE4FFE1FFF4852AFFF1FF8CFFE7FFA6" masterDocTitle="New species of Ateuchus and Canthidium (Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica" masterLastPageNumber="37" masterPageNumber="31" pageNumber="34" updateTime="1698593446454" updateUser="plazi">
|
||
<mods:mods id="7FFBF86656FFAFB25C5B55454971F28A" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="1797E23189CBD0AD004E5DB4ABBD9A37">
|
||
<mods:title id="6877D1D3BC43FA01680C7FB3E2E9B1BF">New species of Ateuchus and Canthidium (Coleoptera: Scarabaeidae: Scarabaeinae) from Costa Rica</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="47E691CE4E35981AF108DA7F027EF480" type="personal">
|
||
<mods:role id="F1C463C9A1B6890C3EB0F06D708D1C7E">
|
||
<mods:roleTerm id="CCD3A8648EDE711273A1C99A78361969">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="5BDA43D4AA241EDD3B68F1FE92D084C2">Kohlmann, Bert</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="10CE9E194FA33E6E4289FDFD037D8B03" type="personal">
|
||
<mods:role id="725F580EB2AAFD67E9E8DE9D4252E9CE">
|
||
<mods:roleTerm id="E7A7108DE667F00DD9C874F47D5BD377">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="88CDDF73F19E7A61980682882731807F">Solis, Angel</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="86D97E40849F3D146C3F4AA723DBF7D1">text</mods:typeOfResource>
|
||
<mods:relatedItem id="18187095D6068A2DABFEF6F5A62F1ECF" type="host">
|
||
<mods:titleInfo id="59C0E0D87FF1B1B7002F17BB38FD728B">
|
||
<mods:title id="42BB4FB3F4A60ABF546D66ABC0A8E3AD">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="96C6A74A46EFF97FE7E749267AFB2EE7">
|
||
<mods:date id="C446BD3B8F9BD78666F86D432464B40A">2009</mods:date>
|
||
<mods:detail id="BA88D0ED2EDD8F1C0BA9431972DC3DFE" type="volume">
|
||
<mods:number id="11521DA58591EF5E82B302090830663C">2219</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="4DE72A6B878F3AB0B9C96017D6613E9F" unit="page">
|
||
<mods:start id="ACF61BBDB58F3584CB301F82B25CB48F">31</mods:start>
|
||
<mods:end id="9C13ED97A25C07AAF1F61D2E7A43B0EB">37</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="F8A82649244A968CD1499896C5FB502E">journal article</mods:classification>
|
||
<mods:identifier id="3D894529B9259B1E60C0044F91C6AEBB" type="DOI">10.5281/zenodo.190015</mods:identifier>
|
||
<mods:identifier id="0393EA18701F9A5D7E9DCC5CB408AD44" type="GBIF-Dataset">a4c97e03-e184-4220-bbc2-ba1f955c580c</mods:identifier>
|
||
<mods:identifier id="9D037560E26C13A2804E25D2389FC1C2" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="1E67A972608DAC3018C02992ED506166" type="Zenodo-Dep">190015</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03DD8799FFF7852FFF66FB98FBADF84D" ID-DOI="http://doi.org/10.5281/zenodo.6216043" ID-GBIF-Taxon="119623533" ID-Zenodo-Dep="6216043" LSID="urn:lsid:plazi:treatment:03DD8799FFF7852FFF66FB98FBADF84D" httpUri="http://treatment.plazi.org/id/03DD8799FFF7852FFF66FB98FBADF84D" lastPageId="5" lastPageNumber="36" pageId="3" pageNumber="34">
|
||
<subSubSection id="C36E6504FFF78529FF66FB98FEFFFBF7" pageId="3" pageNumber="34" type="nomenclature">
|
||
<paragraph id="8BCB368FFFF78529FF66FB98FC48FB8B" blockId="3.[151,943,1043,1105]" box="[151,943,1043,1070]" pageId="3" pageNumber="34">
|
||
<heading id="D08381E3FFF78529FF66FB98FC48FB8B" bold="true" box="[151,943,1043,1070]" fontSize="11" level="1" pageId="3" pageNumber="34" reason="1">
|
||
<emphasis id="B900EA9DFFF78529FF66FB98FC48FB8B" bold="true" box="[151,943,1043,1070]" pageId="3" pageNumber="34">
|
||
<taxonomicName id="4C744D0CFFF78529FF66FB98FCA2FB8B" ID-CoL="699FM" authority="Kohlmann & Solis" authorityName="Kohlmann & Solis" box="[151,837,1043,1070]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="34" phylum="Arthropoda" rank="species" species="darwini" status="sp. nov." subGenus="Eucanthidium">
|
||
<emphasis id="B900EA9DFFF78529FF66FB98FECFFB88" bold="true" box="[151,296,1044,1070]" italics="true" pageId="3" pageNumber="34">Canthidium</emphasis>
|
||
(
|
||
<emphasis id="B900EA9DFFF78529FEC8FB98FE00FB88" bold="true" box="[313,487,1044,1070]" italics="true" pageId="3" pageNumber="34">Eucanthidium</emphasis>
|
||
)
|
||
<emphasis id="B900EA9DFFF78529FE09FB98FDB2FB88" bold="true" box="[504,597,1044,1070]" italics="true" pageId="3" pageNumber="34">darwini</emphasis>
|
||
Kohlmann & Solís
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="A23357E6FFF78529FCA3FB9FFC48FB8B" box="[850,943,1043,1069]" pageId="3" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BCB368FFFF78529FF66FBBBFEFFFBF7" blockId="3.[151,943,1043,1105]" box="[151,280,1079,1105]" pageId="3" pageNumber="34">
|
||
(
|
||
<figureCitation id="134F2A0AFFF78529FF6EFBBBFEE9FBF7" box="[159,270,1079,1105]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="4.[151,255,1988,2012]" captionStartId-1="5.[151,255,1308,1332]" captionTargetBox-1="[226,1360,194,1284]" captionTargetId-1="figure@5.[226,1360,194,1285]" captionTargetPageId-1="5" captionText-0="FIGURE 3. Dorsal view of a male Canthidium darwini." captionText-1="FIGURE 4. Known distribution of the new species in Costa Rica: A. earthorum (․) and C. darwini (˔)." httpUri-1="https://zenodo.org/record/190016/files/figure.png" pageId="3" pageNumber="34">Figs. 3–4</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6504FFF78529FF66FB08FAB6FAB4" pageId="3" pageNumber="34" type="diagnosis">
|
||
<paragraph id="8BCB368FFFF78529FF66FB08FAB6FAB4" blockId="3.[151,1437,1156,2033]" pageId="3" pageNumber="34">
|
||
<emphasis id="B900EA9DFFF78529FF66FB08FEFAFB38" bold="true" box="[151,285,1156,1182]" pageId="3" pageNumber="34">Diagnosis:</emphasis>
|
||
This species is distinguished from other Costa Rican species by the following combination of characters: Frontoclypeal region with three conical tubercles; clypeal indentation V-shaped; pygidium minutely punctate; elytral striae fine, superficially impressed; protibia with two clearly developed lateral teeth, and third weakly developed; head, pronotal, and elytral surface clearly and evidently shagreened.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6504FFF78529FF34FA92FE83F805" pageId="3" pageNumber="34" type="materials_examined">
|
||
<paragraph id="8BCB368FFFF78529FF34FA92FD15FA20" blockId="3.[151,1437,1156,2033]" pageId="3" pageNumber="34">
|
||
<emphasis id="B900EA9DFFF78529FF34FA92FE05FA9E" bold="true" box="[197,482,1310,1336]" pageId="3" pageNumber="34">
|
||
<typeStatus id="54CF882DFFF78529FF34FA92FEDBFA9E" box="[197,316,1310,1336]" pageId="3" pageNumber="34" type="holotype">Holotype</typeStatus>
|
||
description:
|
||
</emphasis>
|
||
Male (
|
||
<figureCitation id="134F2A0AFFF78529FDCDFA93FD51FA9F" box="[572,694,1311,1337]" captionStart-0="FIGURE 3" captionStart-1="FIGURE 4" captionStartId-0="4.[151,255,1988,2012]" captionStartId-1="5.[151,255,1308,1332]" captionTargetBox-1="[226,1360,194,1284]" captionTargetId-1="figure@5.[226,1360,194,1285]" captionTargetPageId-1="5" captionText-0="FIGURE 3. Dorsal view of a male Canthidium darwini." captionText-1="FIGURE 4. Known distribution of the new species in Costa Rica: A. earthorum (․) and C. darwini (˔)." httpUri-1="https://zenodo.org/record/190016/files/figure.png" pageId="3" pageNumber="34">Figs. 3–4</figureCitation>
|
||
). Total length 4.0 mm. Elytral width
|
||
<quantity id="4C8C9B6AFFF78529FB74FA93FB0FFA9F" box="[1157,1256,1311,1337]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="34" unit="mm" value="2.7">2.7 mm</quantity>
|
||
. Body slightly globose, head and pronotal surface green with golden reflections, elytra reddish brown with golden reflections, dorsum shagreened; venter dark brown.
|
||
</paragraph>
|
||
<paragraph id="8BCB368FFFF78529FF34FA1EFBBCFA5C" blockId="3.[151,1437,1156,2033]" pageId="3" pageNumber="34">
|
||
Clypeal margin anteriorly wrinkled and V-shaped, tooth on each side rounded, margin arcuate; dorsal surface of head finely punctate and shagreened; frons with three conical tubercles forming a triangle, area between the tubercles depressed; eye dorsally small (
|
||
<date id="FFCA104FFFF78529FCF5FA6CFCD6FA5C" box="[772,817,1504,1530]" pageId="3" pageNumber="34" value="1915-10">15x</date>
|
||
the interocular distance).
|
||
</paragraph>
|
||
<paragraph id="8BCB368FFFF78529FF37F98BFDB0F9E1" blockId="3.[151,1437,1156,2033]" pageId="3" pageNumber="34">Pronotum finely punctate and uniformly and evidently shagreened, midline not impressed. Proepisternum finely wrinkled proepimeron granular.</paragraph>
|
||
<paragraph id="8BCB368FFFF78529FF37F9D8FE3EF91D" blockId="3.[151,1437,1156,2033]" pageId="3" pageNumber="34">Elytral surface uniformly shagreened; elytral striae broad, dark, smooth, and very slightly impressed; surface dotted with small, polished lunules. Pygidium smooth, shiny, uniformly and minutely punctate, slightly shagreened at base.</paragraph>
|
||
<paragraph id="8BCB368FFFF78529FF37F944FE4FF889" blockId="3.[151,1437,1156,2033]" pageId="3" pageNumber="34">Protibia with two developed external teeth, the third weakly developed; protibiae long and slender (same in female), protibial anterior internal angle elongated as a broad, strong spine; mesotibiae and metatibiae clearly dilated apically.</paragraph>
|
||
<paragraph id="8BCB368FFFF78529FF34F8B0FE83F805" blockId="3.[151,1437,1156,2033]" pageId="3" pageNumber="34">
|
||
<emphasis id="B900EA9DFFF78529FF34F8B0FEA6F8F0" bold="true" box="[197,321,1852,1878]" pageId="3" pageNumber="34">Allotype:</emphasis>
|
||
Female. Total length:
|
||
<quantity id="4C8C9B6AFFF78529FD93F8B0FD20F8F0" box="[610,711,1852,1878]" metricMagnitude="-3" metricUnit="m" metricValue="4.2" pageId="3" pageNumber="34" unit="mm" value="4.2">4.2 mm</quantity>
|
||
. Elytral width:
|
||
<quantity id="4C8C9B6AFFF78529FC7CF8B0FC16F8F0" box="[909,1009,1852,1878]" metricMagnitude="-3" metricUnit="m" metricValue="2.9" pageId="3" pageNumber="34" unit="mm" value="2.9">2.9 mm</quantity>
|
||
. Differs from the
|
||
<typeStatus id="54CF882DFFF78529FB27F8B0FAA3F8F0" box="[1238,1348,1852,1878]" pageId="3" pageNumber="34" type="holotype">holotype</typeStatus>
|
||
by the following characters: protibial anterior internal angle elongated as a fine, slender spine; protibia slightly shorter (
|
||
<quantity id="4C8C9B6AFFF78529FF09F805FEB0F805" box="[248,343,1929,1955]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="34" unit="mm" value="0.8">0.8 mm</quantity>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6504FFF78529FF34F83CFEC2F857" pageId="3" pageNumber="34" type="description">
|
||
<paragraph id="8BCB368FFFF78529FF34F83CFEC2F857" blockId="3.[151,1437,1156,2033]" pageId="3" pageNumber="34">
|
||
<emphasis id="B900EA9DFFF78529FF34F83CFEA5F86C" bold="true" box="[197,322,1968,1994]" pageId="3" pageNumber="34">Variation:</emphasis>
|
||
Total length:
|
||
<quantity id="4C8C9B6AFFF78529FE14F83CFD91F86C" box="[485,630,1968,1994]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.1" metricValueMin="3.9" pageId="3" pageNumber="34" unit="mm" value="4.0" valueMax="4.1" valueMin="3.9">3.9–4.1 mm</quantity>
|
||
. Elytral width:
|
||
<quantity id="4C8C9B6AFFF78529FCD9F83CFC5EF86C" box="[808,953,1968,1994]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" metricValueMax="2.7" metricValueMin="2.5" pageId="3" pageNumber="34" unit="mm" value="2.6" valueMax="2.7" valueMin="2.5">2.5–2.7 mm</quantity>
|
||
. Pronotal reflections can be green, red, or dark brown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6504FFF0852FFF34FF1BFAF8FA01" lastPageId="5" lastPageNumber="36" pageId="4" pageNumber="35" type="materials_examined">
|
||
<paragraph id="8BCB368FFFF0852EFF34FF1BFC4EFD92" blockId="4.[151,1438,151,564]" pageId="4" pageNumber="35">
|
||
<emphasis id="B900EA9DFFF0852EFF34FF1BFE5FFF17" bold="true" box="[197,440,151,177]" pageId="4" pageNumber="35">Material Examined</emphasis>
|
||
(
|
||
<specimenCount id="9D72FD06FFF0852EFE36FF14FD8EFF14" box="[455,617,152,178]" pageId="4" pageNumber="35" type="generic">48 specimens</specimenCount>
|
||
):
|
||
<emphasis id="B900EA9DFFF0852EFD70FF1BFD1CFF17" bold="true" box="[641,763,151,177]" pageId="4" pageNumber="35">
|
||
<typeStatus id="54CF882DFFF0852EFD70FF1BFD12FF17" box="[641,757,151,177]" pageId="4" pageNumber="35" type="holotype">Holotype</typeStatus>
|
||
,
|
||
</emphasis>
|
||
male:
|
||
<collectingCountry id="F363761FFFF0852EFCBBFF14FC1EFF14" box="[842,1017,152,178]" name="Costa Rica" pageId="4" pageNumber="35">COSTA RICA</collectingCountry>
|
||
:
|
||
<emphasis id="B900EA9DFFF0852EFBF5FF14FBA8FF17" box="[1028,1103,152,177]" italics="true" pageId="4" pageNumber="35">Limón</emphasis>
|
||
. P.N. La Amistad. Punto #5.
|
||
<quantity id="4C8C9B6AFFF0852EFF66FF32FED8FF7E" box="[151,319,190,216]" metricMagnitude="3" metricUnit="m" metricValue="1.55" metricValueMax="1.6" metricValueMin="1.5" pageId="4" pageNumber="35" unit="m" value="1550.0" valueMax="1600.0" valueMin="1500.0">1500–1600 m</quantity>
|
||
.
|
||
<date id="FFCA104FFFF0852EFEBBFF32FDF9FF7E" box="[330,542,190,216]" pageId="4" pageNumber="35" value="2007-10-24" valueMax="2007-10-25" valueMin="2007-10-24">24–25 OCT 2007</date>
|
||
. B. Gamboa, M. Monge. Tp. Foso. L N 198990 627455 #92828.
|
||
<emphasis id="B900EA9DFFF0852EFADBFF32FA7BFF7E" bold="true" box="[1322,1436,190,216]" pageId="4" pageNumber="35">Allotype,</emphasis>
|
||
female:
|
||
<emphasis id="B900EA9DFFF0852EFF07FF69FEA2FF58" box="[246,325,229,254]" italics="true" pageId="4" pageNumber="35">ibidem</emphasis>
|
||
.
|
||
<emphasis id="B900EA9DFFF0852EFEA5FF69FE3EFF59" bold="true" box="[340,473,229,255]" pageId="4" pageNumber="35">
|
||
<typeStatus id="54CF882DFFF0852EFEA5FF69FE33FF59" box="[340,468,229,255]" pageId="4" pageNumber="35" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<emphasis id="B900EA9DFFF0852EFE10FF69FDD7FF58" box="[481,560,229,254]" italics="true" pageId="4" pageNumber="35">ibidem</emphasis>
|
||
,
|
||
<specimenCount id="9D72FD06FFF0852EFDCEFF69FD34FF59" box="[575,723,229,255]" pageId="4" pageNumber="35" type="generic">6 specimens</specimenCount>
|
||
;
|
||
<emphasis id="B900EA9DFFF0852EFD2EFF69FCC9FF58" box="[735,814,229,254]" italics="true" pageId="4" pageNumber="35">ibidem</emphasis>
|
||
, Z.P. Río Banano, Campamento base,
|
||
<quantity id="4C8C9B6AFFF0852EFB03FF69FA70FF59" box="[1266,1431,229,255]" metricMagnitude="3" metricUnit="m" metricValue="1.35" metricValueMax="1.4" metricValueMin="1.3" pageId="4" pageNumber="35" unit="m" value="1350.0" valueMax="1400.0" valueMin="1300.0">1300–1400 m</quantity>
|
||
.
|
||
<date id="FFCA104FFFF0852EFF66FE80FE8CFE80" box="[151,363,268,294]" pageId="4" pageNumber="35" value="2007-10-25" valueMax="2007-10-26" valueMin="2007-10-25">25–26 OCT 2007</date>
|
||
. L N 199938 627615 #92827,
|
||
<specimenCount id="9D72FD06FFF0852EFD28FE80FC88FE80" box="[729,879,268,294]" pageId="4" pageNumber="35" type="generic">5 specimens</specimenCount>
|
||
;
|
||
<emphasis id="B900EA9DFFF0852EFC8AFE80FC2CFE83" box="[891,971,268,293]" italics="true" pageId="4" pageNumber="35">ibidem</emphasis>
|
||
, P.I. La Amistad Caribe. Camp. 2: Río Coén, Transecto 1.
|
||
<quantity id="4C8C9B6AFFF0852EFE8DFEBEFDFDFEEA" box="[380,538,306,332]" metricMagnitude="3" metricUnit="m" metricValue="1.75" metricValueMax="1.8" metricValueMin="1.7" pageId="4" pageNumber="35" unit="m" value="1750.0" valueMax="1800.0" valueMin="1700.0">1700–1800m</quantity>
|
||
.
|
||
<date id="FFCA104FFFF0852EFDD4FEBEFD15FEEA" box="[549,754,306,332]" pageId="4" pageNumber="35" value="2007-02-21" valueMax="2007-02-23" valueMin="2007-02-21">21–23 FEB 2007</date>
|
||
. B. Gamboa, M. Moraga. Tp. Foso. L_S_370381_549794 #90734,
|
||
<specimenCount id="9D72FD06FFF0852EFF0CFED5FE78FED5" box="[253,415,345,371]" pageId="4" pageNumber="35" type="generic">12 specimens</specimenCount>
|
||
.
|
||
<emphasis id="B900EA9DFFF0852EFE58FED5FDECFED4" box="[425,523,345,370]" italics="true" pageId="4" pageNumber="35">Cartago</emphasis>
|
||
. La Unión. Z.P. C. Carpintera. Cima.
|
||
<quantity id="4C8C9B6AFFF0852EFC33FED5FBFFFED5" box="[962,1048,345,371]" metricMagnitude="3" metricUnit="m" metricValue="1.8699999999999999" pageId="4" pageNumber="35" unit="m" value="1870.0">1870m</quantity>
|
||
.
|
||
<date id="FFCA104FFFF0852EFBD3FED5FB0EFED5" box="[1058,1257,345,371]" pageId="4" pageNumber="35" value="2008-07-23" valueMax="2008-07-25" valueMin="2008-07-23">23–25 JUL 2008</date>
|
||
. J. Azofeifa, B. Hernández, M. Moraga, M. Zumbado. Tp. Malaise/ Intersección. L_N_207500_538000 #94456,
|
||
<specimenCount id="9D72FD06FFF0852EFA8EFE0CFEF2FE66" pageId="4" pageNumber="35" type="generic">11 specimens</specimenCount>
|
||
;
|
||
<emphasis id="B900EA9DFFF0852EFED0FE2BFE96FE66" box="[289,369,423,448]" italics="true" pageId="4" pageNumber="35">ibidem</emphasis>
|
||
, Z.P. C. Carpintera. Q. Chirraca.
|
||
<quantity id="4C8C9B6AFFF0852EFD0FFE2AFCB4FE66" box="[766,851,422,448]" metricMagnitude="3" metricUnit="m" metricValue="1.75" pageId="4" pageNumber="35" unit="m" value="1750.0">1750m</quantity>
|
||
.
|
||
<date id="FFCA104FFFF0852EFCAEFE2AFBCEFE66" box="[863,1065,422,448]" pageId="4" pageNumber="35" value="2008-07-22" valueMax="2008-07-25" valueMin="2008-07-22">22–25 JUL 2008</date>
|
||
. Azofeifa, Hernández, Moraga, Zumbado. Tp. Malaise/ Intersección. L_N_208250_539000 #94458,
|
||
<specimenCount id="9D72FD06FFF0852EFBE6FE41FB2CFE41" box="[1047,1227,461,487]" pageId="4" pageNumber="35" type="generic">12 specimens</specimenCount>
|
||
;
|
||
<emphasis id="B900EA9DFFF0852EFB28FE41FAD7FE40" box="[1241,1328,461,486]" italics="true" pageId="4" pageNumber="35">ibidem</emphasis>
|
||
, Z.P. C. Carpintera. Fca. Istarú.
|
||
<quantity id="4C8C9B6AFFF0852EFE34FE78FD89FDA8" box="[453,622,500,526]" metricMagnitude="3" metricUnit="m" metricValue="1.85" metricValueMax="1.86" metricValueMin="1.8399999999999999" pageId="4" pageNumber="35" unit="m" value="1850.0" valueMax="1860.0" valueMin="1840.0">1840–1860m</quantity>
|
||
.
|
||
<date id="FFCA104FFFF0852EFD8AFE78FCB4FDA8" box="[635,851,500,526]" pageId="4" pageNumber="35" value="2008-07-22" valueMax="2008-07-25" valueMin="2008-07-22">22–25 JUL 2008</date>
|
||
. Azofeifa, Hernández, Moraga, Zumbado. Tp. Malaise/ Intersección. L_N_207300_539000 #94463,
|
||
<specimenCount id="9D72FD06FFF0852EFCE3FD96FC42FD92" box="[786,933,538,564]" pageId="4" pageNumber="35" type="generic">6 specimens</specimenCount>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF0B6607FFF0852EFF66F848FD17F87A" box="[151,752,1988,2012]" pageId="4" pageNumber="35">
|
||
<paragraph id="8BCB368FFFF0852EFF66F848FD17F87A" blockId="4.[151,752,1988,2012]" box="[151,752,1988,2012]" pageId="4" pageNumber="35">
|
||
<emphasis id="B900EA9DFFF0852EFF66F848FEFEF87A" bold="true" box="[151,281,1988,2012]" pageId="4" pageNumber="35">FIGURE 3.</emphasis>
|
||
Dorsal view of a male
|
||
<taxonomicName id="4C744D0CFFF0852EFDE3F849FD0EF87A" box="[530,745,1989,2012]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="35" phylum="Arthropoda" rank="species" species="darwini">
|
||
<emphasis id="B900EA9DFFF0852EFDE3F849FD0EF87A" box="[530,745,1989,2012]" italics="true" pageId="4" pageNumber="35">Canthidium darwini</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF0B6607FFF1852FFF66FA90FB12FA92" box="[151,1269,1308,1332]" httpUri="https://zenodo.org/record/190016/files/figure.png" pageId="5" pageNumber="36" targetBox="[226,1360,194,1284]" targetPageId="5">
|
||
<paragraph id="8BCB368FFFF1852FFF66FA90FB12FA92" blockId="5.[151,1269,1308,1332]" box="[151,1269,1308,1332]" pageId="5" pageNumber="36">
|
||
<emphasis id="B900EA9DFFF1852FFF66FA90FEFEFA92" bold="true" box="[151,281,1308,1332]" pageId="5" pageNumber="36">FIGURE 4.</emphasis>
|
||
Known distribution of the new species in Costa Rica:
|
||
<taxonomicName id="4C744D0CFFF1852FFCAEFA91FC0AFA92" box="[863,1005,1309,1332]" class="Insecta" family="Scarabaeidae" genus="Ateuchus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="earthorum">
|
||
<emphasis id="B900EA9DFFF1852FFCAEFA91FC0AFA92" box="[863,1005,1309,1332]" italics="true" pageId="5" pageNumber="36">A. earthorum</emphasis>
|
||
</taxonomicName>
|
||
(․) and
|
||
<taxonomicName id="4C744D0CFFF1852FFBB8FA91FB5CFA92" box="[1097,1211,1309,1332]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="darwini">
|
||
<emphasis id="B900EA9DFFF1852FFBB8FA91FB5CFA92" box="[1097,1211,1309,1332]" italics="true" pageId="5" pageNumber="36">C. darwini</emphasis>
|
||
</taxonomicName>
|
||
(˔).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BCB368FFFF1852FFF34FAEAFAF8FA01" blockId="5.[151,1437,1382,2027]" pageId="5" pageNumber="36">
|
||
<emphasis id="B900EA9DFFF1852FFF34FAEAFED7FA26" bold="true" box="[197,304,1382,1408]" pageId="5" pageNumber="36">Habitat:</emphasis>
|
||
The species has been collected in February, July, and October between
|
||
<date id="FFCA104FFFF1852FFB7FFAEAFAD8FA26" box="[1166,1343,1382,1408]" pageId="5" pageNumber="36" value="1300" valueMax="1870">
|
||
1300 and
|
||
<quantity id="4C8C9B6AFFF1852FFAF5FAEAFAD8FA26" box="[1284,1343,1382,1408]" metricMagnitude="3" metricUnit="m" metricValue="1.8699999999999999" pageId="5" pageNumber="36" unit="m" value="1870.0">1870</quantity>
|
||
</date>
|
||
m with Malaise/interception and hog dung-baited traps in cloud forest mixed with
|
||
<taxonomicName id="4C744D0CFFF1852FFBF5FA01FB81FA00" box="[1028,1126,1421,1446]" class="Magnoliopsida" family="Fagaceae" genus="Quercus" kingdom="Plantae" order="Fagales" pageId="5" pageNumber="36" phylum="Tracheophyta" rank="genus">
|
||
<emphasis id="B900EA9DFFF1852FFBF5FA01FB81FA00" box="[1028,1126,1421,1446]" italics="true" pageId="5" pageNumber="36">Quercus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C744D0CFFF1852FFB6EFA01FAFCFA01" box="[1183,1307,1421,1447]" class="Magnoliopsida" family="Lauraceae" kingdom="Plantae" order="Laurales" pageId="5" pageNumber="36" phylum="Tracheophyta" rank="family">Lauraceae</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6504FFF1852FFF34FA3FFBFCF88C" pageId="5" pageNumber="36" type="description">
|
||
<paragraph id="8BCB368FFFF1852FFF34FA3FFF07FA52" blockId="5.[151,1437,1382,2027]" pageId="5" pageNumber="36">
|
||
<emphasis id="B900EA9DFFF1852FFF34FA3FFD9BFA6B" bold="true" box="[197,636,1459,1485]" pageId="5" pageNumber="36">
|
||
Geographical distribution (
|
||
<figureCitation id="134F2A0AFFF1852FFDEEFA3FFD8CFA6B" box="[543,619,1459,1485]" captionStart="FIGURE 4" captionStartId="5.[151,255,1308,1332]" captionTargetBox="[226,1360,194,1284]" captionTargetId="figure@5.[226,1360,194,1285]" captionTargetPageId="5" captionText="FIGURE 4. Known distribution of the new species in Costa Rica: A. earthorum (․) and C. darwini (˔)." httpUri="https://zenodo.org/record/190016/files/figure.png" pageId="5" pageNumber="36">Fig. 4</figureCitation>
|
||
):
|
||
</emphasis>
|
||
Known only from the Caribbean slope of the Talamanca mountain range.
|
||
</paragraph>
|
||
<paragraph id="8BCB368FFFF1852FFF34F98CFD64F9CE" blockId="5.[151,1437,1382,2027]" pageId="5" pageNumber="36">
|
||
<emphasis id="B900EA9DFFF1852FFF34F98CFE01F9BC" bold="true" box="[197,486,1536,1562]" pageId="5" pageNumber="36">Chorological affinities:</emphasis>
|
||
The new species is basically occupying a similar altitude (
|
||
<quantity id="4C8C9B6AFFF1852FFB53F98DFAAFF9BD" box="[1186,1352,1537,1563]" metricMagnitude="3" metricUnit="m" metricValue="1.3" metricValueMax="1.6" metricValueMin="1.0" pageId="5" pageNumber="36" unit="m" value="1300.0" valueMax="1600.0" valueMin="1000.0">1000–1600 m</quantity>
|
||
) in the Caribbean slope of the Talamanca mountain range, as its sister species,
|
||
<taxonomicName id="4C744D0CFFF1852FFC07F9A4FB45F9E7" box="[1014,1186,1576,1601]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="marianelae">
|
||
<emphasis id="B900EA9DFFF1852FFC07F9A4FB45F9E7" box="[1014,1186,1576,1601]" italics="true" pageId="5" pageNumber="36">C. marianelae</emphasis>
|
||
</taxonomicName>
|
||
, does in the Central, Tilarán, and Guanacaste mountain ranges.
|
||
</paragraph>
|
||
<paragraph id="8BCB368FFFF1852FFF34F9F9FBFCF88C" blockId="5.[151,1437,1382,2027]" pageId="5" pageNumber="36">
|
||
<emphasis id="B900EA9DFFF1852FFF34F9F9FDE7F929" bold="true" box="[197,512,1653,1679]" pageId="5" pageNumber="36">Taxonomic relationships:</emphasis>
|
||
This species is hypothesized to be the sister species of
|
||
<taxonomicName id="4C744D0CFFF1852FFB77F9F9FAC9F928" box="[1158,1326,1653,1678]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="marianelae">
|
||
<emphasis id="B900EA9DFFF1852FFB77F9F9FAC9F928" box="[1158,1326,1653,1678]" italics="true" pageId="5" pageNumber="36">C. marianelae</emphasis>
|
||
</taxonomicName>
|
||
based on shared morphological characters. These species can be easily separated because the new species is heavily shagreened on the thorax and elytra; whereas
|
||
<taxonomicName id="4C744D0CFFF1852FFD29F94EFC6DF97D" box="[728,906,1730,1755]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="marianelae">
|
||
<emphasis id="B900EA9DFFF1852FFD29F94EFC6DF97D" box="[728,906,1730,1755]" italics="true" pageId="5" pageNumber="36">C. marianelae</emphasis>
|
||
</taxonomicName>
|
||
is only slightly shagreened on the thorax. Moreover, the new species has evident broad, dark, flat, and superficial elytral striae; whereas,
|
||
<taxonomicName id="4C744D0CFFF1852FFB05F965FA7CF8A4" box="[1268,1435,1769,1794]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="marianelae">
|
||
<emphasis id="B900EA9DFFF1852FFB05F965FA7CF8A4" box="[1268,1435,1769,1794]" italics="true" pageId="5" pageNumber="36">C. marianelae</emphasis>
|
||
</taxonomicName>
|
||
has broad, flat, and superficial elytral striae, that are very difficult to discern.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C36E6504FFF1852FFF34F8BAFBADF84D" pageId="5" pageNumber="36" type="etymology">
|
||
<paragraph id="8BCB368FFFF1852FFF34F8BAFBADF84D" blockId="5.[151,1437,1382,2027]" pageId="5" pageNumber="36">
|
||
<emphasis id="B900EA9DFFF1852FFF34F8BAFEB1F8F6" bold="true" box="[197,342,1846,1872]" pageId="5" pageNumber="36">Etymology:</emphasis>
|
||
“
|
||
<taxonomicName id="4C744D0CFFF1852FFE9BF8BBFE2BF8F6" box="[362,460,1847,1872]" class="Insecta" family="Scarabaeidae" genus="Canthidium" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="36" phylum="Arthropoda" rank="species" species="darwini">
|
||
<emphasis id="B900EA9DFFF1852FFE9BF8BBFE2BF8F6" box="[362,460,1847,1872]" italics="true" pageId="5" pageNumber="36">Darwini</emphasis>
|
||
</taxonomicName>
|
||
”, is the genitive form of Darwin. This species is primarily dedicated to the eminent biologist and evolutionist, Charles Darwin. It is additionally dedicated to the
|
||
<collectingCountry id="F363761FFFF1852FFBC2F8D1FBB9F8D1" box="[1075,1118,1885,1911]" name="United Kingdom" pageId="5" pageNumber="36">UK</collectingCountry>
|
||
Darwin Initiative for their contribution to the preservation of tropical biodiversity and as an acknowledgement for financing the project “Basic Tools for Managing the La Amistad (
|
||
<collectingCountry id="F363761FFFF1852FFD6CF826FCF9F862" box="[669,798,1962,1988]" name="Costa Rica" pageId="5" pageNumber="36">Costa Rica</collectingCountry>
|
||
–
|
||
<collectingCountry id="F363761FFFF1852FFCCBF826FC7EF862" box="[826,921,1962,1988]" name="Panama" pageId="5" pageNumber="36">Panama</collectingCountry>
|
||
) International Park.” This project supported the expeditions that resulted in the collection of the specimens used in this study.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |