208 lines
34 KiB
XML
208 lines
34 KiB
XML
<document id="CCA5BB624533EA19F90A1A911CB0E070" ID-CLB-Dataset="2931" ID-DOI="10.11646/zootaxa.5079.1.1" ID-GBIF-Dataset="010695ef-a142-42ad-81d7-2e7ed1f7e89e" ID-ISSN="1175-5326" ID-Zenodo-Dep="5766402" ID-ZooBank="2B118B81-7353-4A2C-A892-5466DFC83230" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638952199435" checkinUser="plazi" docAuthor="Takano, Kohei Takenaka, Gao, Jian-Jun, Hu, Yao-Guang, Li, Nan-Nan, Yafuso, Masako, Suwito, Awit, Repin, Rimi, Pungga, Runi Anak Sylvester, Meleng, Paulus Ak, Kaliang, Clement Het, Chong, Lucy & Toda, Masanori J." docDate="2021" docId="03D75C2D45762E23FF526197B225B631" docLanguage="en" docName="zootaxa.5079.1.1.pdf" docOrigin="Zootaxa 5079 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Colocasiomyia cristata de Meijere 1914" docType="treatment" docVersion="7" lastPageNumber="29" masterDocId="FFEE2455456A2E3FFFC56005B34CB33C" masterDocTitle="Phylogeny, taxonomy and flower-breeding ecology of the Colocasiomyia cristata species group (Diptera: Drosophilidae), with descriptions of ten new species" masterLastPageNumber="70" masterPageNumber="1" pageNumber="29" updateTime="1699472124804" updateUser="ExternalLinkService">
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<mods:title id="37B23277F0096CD97E8F07160BCDD65F">Phylogeny, taxonomy and flower-breeding ecology of the Colocasiomyia cristata species group (Diptera: Drosophilidae), with descriptions of ten new species</mods:title>
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<mods:namePart id="70DF4F4191648CC5A705A9D310E96CDB">Takano, Kohei Takenaka</mods:namePart>
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<mods:namePart id="7EFF5212F88CCE320D6C11C8977373AC">Gao, Jian-Jun</mods:namePart>
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<mods:namePart id="8BD4993EC86AF15F3EC4AF7D832BAC38">Hu, Yao-Guang</mods:namePart>
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<mods:namePart id="EDE8DAE91CC37AE5439B8CEB681FAA40">Li, Nan-Nan</mods:namePart>
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<mods:namePart id="D7C7102800412560C17090350FD9762B">Yafuso, Masako</mods:namePart>
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<mods:namePart id="D043F512538D701F7510A3627F931C4B">Suwito, Awit</mods:namePart>
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<mods:namePart id="73B99DE5A55CCAC885FD4117FF69A92D">Repin, Rimi</mods:namePart>
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<mods:namePart id="FE50A1147B83DE0432CEE653A28A12E0">Pungga, Runi Anak Sylvester</mods:namePart>
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<mods:namePart id="1F3D566B8D08712B5691E926D06C8F3F">Meleng, Paulus Ak</mods:namePart>
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<mods:namePart id="9501AB256D346CEB472D346C4EFDD73C">Kaliang, Clement Het</mods:namePart>
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<mods:namePart id="12B00B0527A77AF28CC568884DD2D728">Chong, Lucy</mods:namePart>
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<mods:namePart id="EE33DE704D69C29BF217E11EF2604C00">Toda, Masanori J.</mods:namePart>
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<mods:title id="17821D8AF2F842966DE31FCAE527CDB5">Zootaxa</mods:title>
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<mods:date id="9999DF92507C04F602CC33A917267CEC">2021</mods:date>
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<mods:number id="D8F3784E1F61479265481BDF5FE6C678">2021-12-08</mods:number>
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<mods:number id="1C1CE39D34D0E7896756A8815A6E390C">5079</mods:number>
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<mods:classification id="11BCDB8A97162C3F808C976C86BA27D4">journal article</mods:classification>
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<mods:identifier id="C09F7B3C549BCDFAB5E8D304BB6640F4" type="DOI">10.11646/zootaxa.5079.1.1</mods:identifier>
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<treatment id="03D75C2D45762E23FF526197B225B631" ID-DOI="http://doi.org/10.5281/zenodo.5786159" ID-GBIF-Taxon="191651669" ID-Zenodo-Dep="5786159" LSID="urn:lsid:plazi:treatment:03D75C2D45762E23FF526197B225B631" httpUri="http://treatment.plazi.org/id/03D75C2D45762E23FF526197B225B631" lastPageNumber="29" pageId="28" pageNumber="29">
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<subSubSection id="C364BEB045762E23FF526197B1A0B291" box="[151,748,402,429]" pageId="28" pageNumber="29" type="nomenclature">
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<paragraph id="8BC1ED3B45762E23FF526197B1A0B291" blockId="28.[151,748,402,429]" box="[151,748,402,429]" pageId="28" pageNumber="29">
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<emphasis id="B90A312945762E23FF526197B1A0B291" bold="true" box="[151,748,402,429]" pageId="28" pageNumber="29">
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2.2.
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<taxonomicName id="4C7E96B845762E23FF0E6197B291B291" ID-CoL="5ZMS3" authorityName="de Meijere" authorityYear="1914" box="[203,477,402,429]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="cristata">
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<emphasis id="B90A312945762E23FF0E6197B291B291" bold="true" box="[203,477,402,429]" italics="true" pageId="28" pageNumber="29">Colocasiomyia cristata</emphasis>
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</taxonomicName>
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species complex, new
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</emphasis>
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</paragraph>
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</subSubSection>
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<subSubSection id="C364BEB045762E23FF5261DFB003B7F9" pageId="28" pageNumber="29" type="diagnosis">
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<paragraph id="8BC1ED3B45762E23FF5261DFB070B15D" blockId="28.[151,1437,474,1293]" pageId="28" pageNumber="29">
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<emphasis id="B90A312945762E23FF5261DFB246B2C8" bold="true" box="[151,266,474,500]" pageId="28" pageNumber="29">Diagnosis</emphasis>
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. Mid tibia with only 1 large, apical seta (ch.15-1;
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<figureCitation id="1345F1BE45762E23FC8B61DEB0E5B2C9" box="[846,937,475,501]" captionStart="FIGURE 7" captionStartId="12.[151,250,903,927]" captionTargetBox="[151,1435,181,878]" captionTargetId="figure-22@12.[151,1435,181,879]" captionTargetPageId="12" captionText="FIGURE 7. Characters 13‒15 on the fore (A, B, distal portion of tibia and tarsus) and mid (C, D, ditto) legs of Colocasiomyia colocasiae (Duda) (A, C) and C. cristata de Meijere (B, D). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766416" httpUri="https://zenodo.org/record/5766416/files/figure.png" pageId="28" pageNumber="29">Fig. 7D</figureCitation>
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). Male abdominal sternite VI with a pair of processes bearing prominent seta on apex and stout setae on submedial to distal portion (ch.26,27-1;
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<figureCitation id="1345F1BE45762E23FAD661FAB6DDB125" box="[1299,1425,511,537]" captionStart="FIGURE 9" captionStartId="13.[151,250,1780,1804]" captionTargetBox="[151,1435,181,1757]" captionTargetId="figure-18@13.[151,1435,181,1757]" captionTargetPageId="13" captionText="FIGURE 9. Male abdominal sternite VIs of Colocasiomyia colocasiae (Duda) (B in anteroventral view, C in lateral view), C. stamenicola (Carson & Okada) (D in anterior view) and all species of the C. cristata subgroup (A in ventral view, E‒R in anterior view): A, C. ecornuta sp. nov.; E, C. grandis sp. nov.; F, C. vieti sp. nov.; G, C. cristata de Meijere; H, C. matthewsi sp. nov.; I, C. sulawesiana Okada & Yafuso; J, C. kinabaluana sp. nov.; K, C. kotana sp. nov.; L, C. sabahana sp. nov.; M, C. sarawakana sp. nov.; N, C. pistilicola (Carson & Okada); O, C. xenalocasiae (Okada); P, C. diconica (Toda & Okada); Q, C. sumatrana sp. nov.; and R, C. leucocasiae sp. nov. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766420" httpUri="https://zenodo.org/record/5766420/files/figure.png" pageId="28" pageNumber="29">Fig. 9G–K</figureCitation>
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). Lateral pieces of subepandrial sclerite broad plates somewhat sclerotized at least on ventral margin (ch.37-4;
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<figureCitation id="1345F1BE45762E23FAB56226B380B15D" captionStart="FIGURE 10" captionStartId="14.[151,250,1695,1719]" captionTargetBox="[151,1435,885,1666]" captionTargetId="figure-299@14.[151,1435,885,1672]" captionTargetPageId="14" captionText="FIGURE 10. Characters 30‒39 on the periphallic organs of Colocasiomyia colocasiae (Duda) (A, lateral view; B, anterior view), C. cristata de Meijere (C, lateral view; D, ventral view; E, surstylus) and C. xenalocasiae (Okada) (F, oblique lateral view; G, ventral view; H, surstylus). Abbreviations: cerc, cercus; epand, epandrium; sur, surstylus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766422" httpUri="https://zenodo.org/record/5766422/files/figure.png" pageId="28" pageNumber="29">Fig. 10D</figureCitation>
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). Phallal sheath thicker than wide (ch.43-0;
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<figureCitation id="1345F1BE45762E23FD7D6242B062B15D" box="[696,814,583,609]" captionStart="FIGURE 11" captionStartId="16.[151,250,1315,1339]" captionTargetBox="[151,1435,237,1292]" captionTargetId="figure-43@16.[151,1435,237,1292]" captionTargetPageId="16" captionText="FIGURE 11. Characters 40‒47 on the phallic organs of Colocasiomyia colocasiae (Duda) (A, lateral view; B, ventral view; C, hypandrium and gonocoxites, oblique lateral view), C. xenalocasiae (Okada) (D, lateral view; E, ventral view), C. sabahana sp. nov. (F, lateral view; G, ventral view) and C. cristata de Meijere (H, lateral view; I, ventral view).Abbreviations: aed, aedeagus; goncx, gonocoxite; hypd, hypandrium; phap, phallapodeme; ph sh, phallal sheath. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766424" httpUri="https://zenodo.org/record/5766424/files/figure.png" pageId="28" pageNumber="29">Fig. 11H,I</figureCitation>
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).
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</paragraph>
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<paragraph id="8BC1ED3B45762E23FF02626FB003B7F9" blockId="28.[151,1437,474,1293]" pageId="28" pageNumber="29">
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<emphasis id="B90A312945762E23FF02626FB2E0B1B9" bold="true" box="[199,428,618,645]" pageId="28" pageNumber="29">Shared characters.</emphasis>
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Anterior reclinate orbital seta small, situated behind proclinate orbital seta; additional, interfrontal setulae on fronto-orbital plate present (ch.2-0;
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<figureCitation id="1345F1BE45762E23FCE1628AB034B195" box="[804,888,655,681]" captionStart="FIGURE 3" captionStartId="9.[151,250,1541,1565]" captionTargetBox="[151,1435,181,1517]" captionTargetId="figure-18@9.[151,1435,181,1517]" captionTargetPageId="9" captionText="FIGURE 3. Characters 1‒4 on the head of Colocasiomyia colocasiae (Duda) (A‒D) and C. cristata de Meijere (E‒H). A, E, Caudal view; B, F, dorsal view; C, G, frontal view; D, H, right eye (frontal view). Abbreviations: ar, arista; a rc orb s, anterior reclinate orbital seta; i ocp s, inner occipital setae; pc orb s, proclinate orbital seta; spc s, supracervical setae. Character states are indicated in the form of ‘character No.-state code’. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766408" httpUri="https://zenodo.org/record/5766408/files/figure.png" pageId="28" pageNumber="29">Fig. 3F</figureCitation>
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). Eye with stout, dense interfacetal setulae (ch.3- 1;
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<figureCitation id="1345F1BE45762E23FF7762B6B246B1F1" box="[178,266,691,717]" captionStart="FIGURE 3" captionStartId="9.[151,250,1541,1565]" captionTargetBox="[151,1435,181,1517]" captionTargetId="figure-18@9.[151,1435,181,1517]" captionTargetPageId="9" captionText="FIGURE 3. Characters 1‒4 on the head of Colocasiomyia colocasiae (Duda) (A‒D) and C. cristata de Meijere (E‒H). A, E, Caudal view; B, F, dorsal view; C, G, frontal view; D, H, right eye (frontal view). Abbreviations: ar, arista; a rc orb s, anterior reclinate orbital seta; i ocp s, inner occipital setae; pc orb s, proclinate orbital seta; spc s, supracervical setae. Character states are indicated in the form of ‘character No.-state code’. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766408" httpUri="https://zenodo.org/record/5766408/files/figure.png" pageId="28" pageNumber="29">Fig. 3H</figureCitation>
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). Distance between antennal sockets narrower than half of socket width (ch.4-1;
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<figureCitation id="1345F1BE45762E23FB4562B6B79BB1F1" box="[1152,1239,691,717]" captionStart="FIGURE 3" captionStartId="9.[151,250,1541,1565]" captionTargetBox="[151,1435,181,1517]" captionTargetId="figure-18@9.[151,1435,181,1517]" captionTargetPageId="9" captionText="FIGURE 3. Characters 1‒4 on the head of Colocasiomyia colocasiae (Duda) (A‒D) and C. cristata de Meijere (E‒H). A, E, Caudal view; B, F, dorsal view; C, G, frontal view; D, H, right eye (frontal view). Abbreviations: ar, arista; a rc orb s, anterior reclinate orbital seta; i ocp s, inner occipital setae; pc orb s, proclinate orbital seta; spc s, supracervical setae. Character states are indicated in the form of ‘character No.-state code’. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766408" httpUri="https://zenodo.org/record/5766408/files/figure.png" pageId="28" pageNumber="29">Fig. 3G</figureCitation>
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). Cibarial, medial sensilla in parallel rows narrower than sensilla campaniformia (ch.5-2;
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<figureCitation id="1345F1BE45762E23FC0E62D2B769B1CD" box="[971,1061,727,753]" captionStart="FIGURE 4" captionStartId="10.[151,250,798,822]" captionTargetBox="[151,1435,181,773]" captionTargetId="figure-22@10.[151,1435,181,774]" captionTargetPageId="10" captionText="FIGURE 4. Characters 5 and 6 on the cibarium (dorsal view) of Colocasiomyia xenalocasiae (Okada) (A), C. cristata de Meijere (B) and C. sarawakana sp. nov. (C). Abbreviations: a s, anterior sensilla; m s, medial sensilla; p s, posterior sensillum; s c, sensilla campaniformia. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766410" httpUri="https://zenodo.org/record/5766410/files/figure.png" pageId="28" pageNumber="29">Fig. 4B</figureCitation>
|
||
). Neither additional dorsocentral setae on presutural area nor prescutellar acrostichal setae present; acrostichal setulae in 4 rows (ch.8-1, 9-0;
|
||
<figureCitation id="1345F1BE45762E23FAB562FEB3F0B005" captionStart="FIGURE 5" captionStartId="10.[151,250,1874,1898]" captionTargetBox="[151,1435,920,1850]" captionTargetId="figure-75@10.[151,1435,919,1851]" captionTargetPageId="10" captionText="FIGURE 5. Characters 7‒11 on the thorax (A‒D, dorsal view; E‒G, lateral view) of Colocasiomyia colocasiae (Duda) (A, E), C. grandis sp. nov. (B), C. cristata de Meijere (C, F), C. diconica (Toda & Okada) (D) and C. ecornuta sp. nov. (G). Abbreviations: a dc s, anterior dorsocentral setae; ap sctl s, apical scutellar seta; b sctl s, basal scutellar seta; p kepst s, posterior katepisternal seta; pprn s, postpronotal seta; trn sut, transverse suture. Scale bars: 0.5 mm in A‒D, 0.1 mm in E‒G." figureDoi="http://doi.org/10.5281/zenodo.5766412" httpUri="https://zenodo.org/record/5766412/files/figure.png" pageId="28" pageNumber="29">Fig. 5C</figureCitation>
|
||
). Katepisternal setae small, shorter than postpronotal setae (ch.10-1;
|
||
<figureCitation id="1345F1BE45762E23FC75631AB74FB005" box="[944,1027,798,825]" captionStart="FIGURE 5" captionStartId="10.[151,250,1874,1898]" captionTargetBox="[151,1435,920,1850]" captionTargetId="figure-75@10.[151,1435,919,1851]" captionTargetPageId="10" captionText="FIGURE 5. Characters 7‒11 on the thorax (A‒D, dorsal view; E‒G, lateral view) of Colocasiomyia colocasiae (Duda) (A, E), C. grandis sp. nov. (B), C. cristata de Meijere (C, F), C. diconica (Toda & Okada) (D) and C. ecornuta sp. nov. (G). Abbreviations: a dc s, anterior dorsocentral setae; ap sctl s, apical scutellar seta; b sctl s, basal scutellar seta; p kepst s, posterior katepisternal seta; pprn s, postpronotal seta; trn sut, transverse suture. Scale bars: 0.5 mm in A‒D, 0.1 mm in E‒G." figureDoi="http://doi.org/10.5281/zenodo.5766412" httpUri="https://zenodo.org/record/5766412/files/figure.png" pageId="28" pageNumber="29">Fig. 5F</figureCitation>
|
||
). Basal scutellar setae long; posterior tip beyond half of apical scutellar setae (ch.11-1;
|
||
<figureCitation id="1345F1BE45762E23FD016346B051B061" box="[708,797,834,861]" captionStart="FIGURE 5" captionStartId="10.[151,250,1874,1898]" captionTargetBox="[151,1435,920,1850]" captionTargetId="figure-75@10.[151,1435,919,1851]" captionTargetPageId="10" captionText="FIGURE 5. Characters 7‒11 on the thorax (A‒D, dorsal view; E‒G, lateral view) of Colocasiomyia colocasiae (Duda) (A, E), C. grandis sp. nov. (B), C. cristata de Meijere (C, F), C. diconica (Toda & Okada) (D) and C. ecornuta sp. nov. (G). Abbreviations: a dc s, anterior dorsocentral setae; ap sctl s, apical scutellar seta; b sctl s, basal scutellar seta; p kepst s, posterior katepisternal seta; pprn s, postpronotal seta; trn sut, transverse suture. Scale bars: 0.5 mm in A‒D, 0.1 mm in E‒G." figureDoi="http://doi.org/10.5281/zenodo.5766412" httpUri="https://zenodo.org/record/5766412/files/figure.png" pageId="28" pageNumber="29">Fig. 5C</figureCitation>
|
||
). Apical scutellar setae nearer to each other than to basal scutellar seta (
|
||
<figureCitation id="1345F1BE45762E23FEFF6362B2D8B0BD" box="[314,404,870,897]" captionStart="FIGURE 5" captionStartId="10.[151,250,1874,1898]" captionTargetBox="[151,1435,920,1850]" captionTargetId="figure-75@10.[151,1435,919,1851]" captionTargetPageId="10" captionText="FIGURE 5. Characters 7‒11 on the thorax (A‒D, dorsal view; E‒G, lateral view) of Colocasiomyia colocasiae (Duda) (A, E), C. grandis sp. nov. (B), C. cristata de Meijere (C, F), C. diconica (Toda & Okada) (D) and C. ecornuta sp. nov. (G). Abbreviations: a dc s, anterior dorsocentral setae; ap sctl s, apical scutellar seta; b sctl s, basal scutellar seta; p kepst s, posterior katepisternal seta; pprn s, postpronotal seta; trn sut, transverse suture. Scale bars: 0.5 mm in A‒D, 0.1 mm in E‒G." figureDoi="http://doi.org/10.5281/zenodo.5766412" httpUri="https://zenodo.org/record/5766412/files/figure.png" pageId="28" pageNumber="29">Fig. 5C</figureCitation>
|
||
). Costal setae in middle row all weak, trichoid (ch.12-2;
|
||
<figureCitation id="1345F1BE45762E23FBD96362B738B0BE" box="[1052,1140,871,898]" captionStart="FIGURE 6" captionStartId="11.[151,250,1951,1975]" captionTargetBox="[286,1299,181,1927]" captionTargetId="figure-18@11.[286,1300,181,1927]" captionTargetPageId="11" captionText="FIGURE 6. Character 12 of costal setae on the wing (anterodistal portion) of Colocasiomyia xenalocasiae (Okada) (A), C. colocasiae (Duda) (B) and C. cristata de Meijere (C). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766414" httpUri="https://zenodo.org/record/5766414/files/figure.png" pageId="28" pageNumber="29">Fig. 6C</figureCitation>
|
||
). Patch covered with only minute pubescence absent on anterodorsal portion of fore tarsomere I (ch.13-1;
|
||
<figureCitation id="1345F1BE45762E23FBD8638EB73AB099" box="[1053,1142,907,933]" captionStart="FIGURE 7" captionStartId="12.[151,250,903,927]" captionTargetBox="[151,1435,181,878]" captionTargetId="figure-22@12.[151,1435,181,879]" captionTargetPageId="12" captionText="FIGURE 7. Characters 13‒15 on the fore (A, B, distal portion of tibia and tarsus) and mid (C, D, ditto) legs of Colocasiomyia colocasiae (Duda) (A, C) and C. cristata de Meijere (B, D). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766416" httpUri="https://zenodo.org/record/5766416/files/figure.png" pageId="28" pageNumber="29">Fig. 7B</figureCitation>
|
||
). Male abdominal sternite V medially concaved on posterior margin. Female abdominal sternites III and IV longer than wide (ch.19,20-1). Male abdominal sternite VI not pubescent at least partly (ch.23-0;
|
||
<figureCitation id="1345F1BE45762E23FCBB63D6B0B7B0D1" box="[894,1019,979,1005]" captionStart="FIGURE 9" captionStartId="13.[151,250,1780,1804]" captionTargetBox="[151,1435,181,1757]" captionTargetId="figure-18@13.[151,1435,181,1757]" captionTargetPageId="13" captionText="FIGURE 9. Male abdominal sternite VIs of Colocasiomyia colocasiae (Duda) (B in anteroventral view, C in lateral view), C. stamenicola (Carson & Okada) (D in anterior view) and all species of the C. cristata subgroup (A in ventral view, E‒R in anterior view): A, C. ecornuta sp. nov.; E, C. grandis sp. nov.; F, C. vieti sp. nov.; G, C. cristata de Meijere; H, C. matthewsi sp. nov.; I, C. sulawesiana Okada & Yafuso; J, C. kinabaluana sp. nov.; K, C. kotana sp. nov.; L, C. sabahana sp. nov.; M, C. sarawakana sp. nov.; N, C. pistilicola (Carson & Okada); O, C. xenalocasiae (Okada); P, C. diconica (Toda & Okada); Q, C. sumatrana sp. nov.; and R, C. leucocasiae sp. nov. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766420" httpUri="https://zenodo.org/record/5766420/files/figure.png" pageId="28" pageNumber="29">Fig. 9G–K</figureCitation>
|
||
). Epandrial phragma small (narrower than epandrial anteroventral corner) expansion on antero-subapical margin (ch.33-2;
|
||
<figureCitation id="1345F1BE45762E23FB8B63F2B7F8B72D" box="[1102,1204,1015,1041]" captionStart="FIGURE 10" captionStartId="14.[151,250,1695,1719]" captionTargetBox="[151,1435,885,1666]" captionTargetId="figure-299@14.[151,1435,885,1672]" captionTargetPageId="14" captionText="FIGURE 10. Characters 30‒39 on the periphallic organs of Colocasiomyia colocasiae (Duda) (A, lateral view; B, anterior view), C. cristata de Meijere (C, lateral view; D, ventral view; E, surstylus) and C. xenalocasiae (Okada) (F, oblique lateral view; G, ventral view; H, surstylus). Abbreviations: cerc, cercus; epand, epandrium; sur, surstylus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766422" httpUri="https://zenodo.org/record/5766422/files/figure.png" pageId="28" pageNumber="29">Fig. 10C</figureCitation>
|
||
). Surstylus vestigial, very narrow process, apically with 1 or 2 tiny, trichoid seta(e) (ch.34,35-1;
|
||
<figureCitation id="1345F1BE45762E23FC1E641EB773B709" box="[987,1087,1051,1077]" captionStart="FIGURE 10" captionStartId="14.[151,250,1695,1719]" captionTargetBox="[151,1435,885,1666]" captionTargetId="figure-299@14.[151,1435,885,1672]" captionTargetPageId="14" captionText="FIGURE 10. Characters 30‒39 on the periphallic organs of Colocasiomyia colocasiae (Duda) (A, lateral view; B, anterior view), C. cristata de Meijere (C, lateral view; D, ventral view; E, surstylus) and C. xenalocasiae (Okada) (F, oblique lateral view; G, ventral view; H, surstylus). Abbreviations: cerc, cercus; epand, epandrium; sur, surstylus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766422" httpUri="https://zenodo.org/record/5766422/files/figure.png" pageId="28" pageNumber="29">Fig. 10E</figureCitation>
|
||
). Median piece of subepandrial sclerite broad, somewhat quadrate plate (ch.36-0). Cercal, ventral/apical portion more or less differentiated from upper portion (ch.38-0;
|
||
<figureCitation id="1345F1BE45762E23FE766466B153B741" box="[435,543,1123,1149]" captionStart="FIGURE 10" captionStartId="14.[151,250,1695,1719]" captionTargetBox="[151,1435,885,1666]" captionTargetId="figure-299@14.[151,1435,885,1672]" captionTargetPageId="14" captionText="FIGURE 10. Characters 30‒39 on the periphallic organs of Colocasiomyia colocasiae (Duda) (A, lateral view; B, anterior view), C. cristata de Meijere (C, lateral view; D, ventral view; E, surstylus) and C. xenalocasiae (Okada) (F, oblique lateral view; G, ventral view; H, surstylus). Abbreviations: cerc, cercus; epand, epandrium; sur, surstylus. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766422" httpUri="https://zenodo.org/record/5766422/files/figure.png" pageId="28" pageNumber="29">Fig. 10C</figureCitation>
|
||
). Phallal sheath not pubescent, gently curved ventrad, projected anteriad at dorsobasal corner in lateral view, apically not bilobed (ch.40-1, 41-3, 42-0, 44-1;
|
||
<figureCitation id="1345F1BE45762E23FB896482B7F9B79D" box="[1100,1205,1159,1185]" captionStart="FIGURE 11" captionStartId="16.[151,250,1315,1339]" captionTargetBox="[151,1435,237,1292]" captionTargetId="figure-43@16.[151,1435,237,1292]" captionTargetPageId="16" captionText="FIGURE 11. Characters 40‒47 on the phallic organs of Colocasiomyia colocasiae (Duda) (A, lateral view; B, ventral view; C, hypandrium and gonocoxites, oblique lateral view), C. xenalocasiae (Okada) (D, lateral view; E, ventral view), C. sabahana sp. nov. (F, lateral view; G, ventral view) and C. cristata de Meijere (H, lateral view; I, ventral view).Abbreviations: aed, aedeagus; goncx, gonocoxite; hypd, hypandrium; phap, phallapodeme; ph sh, phallal sheath. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766424" httpUri="https://zenodo.org/record/5766424/files/figure.png" pageId="28" pageNumber="29">Fig. 11H</figureCitation>
|
||
). Gonocoxites long, narrow slips (
|
||
<figureCitation id="1345F1BE45762E23FEF564AEB2D4B7F9" box="[304,408,1195,1221]" captionStart="FIGURE 11" captionStartId="16.[151,250,1315,1339]" captionTargetBox="[151,1435,237,1292]" captionTargetId="figure-43@16.[151,1435,237,1292]" captionTargetPageId="16" captionText="FIGURE 11. Characters 40‒47 on the phallic organs of Colocasiomyia colocasiae (Duda) (A, lateral view; B, ventral view; C, hypandrium and gonocoxites, oblique lateral view), C. xenalocasiae (Okada) (D, lateral view; E, ventral view), C. sabahana sp. nov. (F, lateral view; G, ventral view) and C. cristata de Meijere (H, lateral view; I, ventral view).Abbreviations: aed, aedeagus; goncx, gonocoxite; hypd, hypandrium; phap, phallapodeme; ph sh, phallal sheath. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766424" httpUri="https://zenodo.org/record/5766424/files/figure.png" pageId="28" pageNumber="29">Fig. 11H</figureCitation>
|
||
). Epiproct with pubescence (
|
||
<figureCitation id="1345F1BE45762E23FD1964AEB00FB7F9" box="[732,835,1195,1221]" captionStart="FIGURE 12" captionStartId="17.[151,250,762,786]" captionTargetBox="[151,1435,182,734]" captionTargetId="figure-18@17.[151,1435,181,737]" captionTargetPageId="17" captionText="FIGURE 12. Characters 48‒51 on the female terminalia of Colocasiomyia colocasiae (Duda) (A, lateral view; B, hypoproct in ventral view) and C. cristata de Meijere (C, lateral view; D, hypoproct in ventral view). Abbreviations: epiprct, epiproct; hyprct, hypoproct; ov, oviscapt; t-VIII, tergite VIII. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766426" httpUri="https://zenodo.org/record/5766426/files/figure.png" pageId="28" pageNumber="29">Fig. 12C</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364BEB045762E23FF0264CAB225B631" pageId="28" pageNumber="29" type="reference_group">
|
||
<paragraph id="8BC1ED3B45762E23FF0264CAB225B631" blockId="28.[151,1437,474,1293]" pageId="28" pageNumber="29">
|
||
<emphasis id="B90A312945762E23FF0264CAB2DDB7D4" bold="true" box="[199,401,1230,1257]" pageId="28" pageNumber="29">Included species.</emphasis>
|
||
<taxonomicName id="4C7E96B845762E23FE5D64CAB1D1B7D4" ID-CoL="5ZMS3" authorityName="de Meijere" authorityYear="1914" box="[408,669,1231,1257]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="cristata">
|
||
<emphasis id="B90A312945762E23FE5D64CAB1D1B7D4" box="[408,669,1231,1257]" italics="true" pageId="28" pageNumber="29">Colocasiomyia cristata</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7E96B845762E23FD6E64CAB019B7D5" ID-CoL="X6V5" authorityName="Okada & Yafuso" authorityYear="1989" box="[683,853,1231,1257]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="sulawesiana">
|
||
<emphasis id="B90A312945762E23FD6E64CAB019B7D5" box="[683,853,1231,1257]" italics="true" pageId="28" pageNumber="29">C. sulawesiana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7E96B845762E23FCA164CAB75DB7D5" ID-CoL="9HZHM" authority="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda, 2021" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[868,1041,1231,1257]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="kinabaluana" status="sp. nov.">
|
||
<emphasis id="B90A312945762E23FCA164CAB75DB7D5" box="[868,1041,1231,1257]" italics="true" pageId="28" pageNumber="29">C. kinabaluana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90A312945762E23FBD264CAB73DB7D5" bold="true" box="[1047,1137,1231,1257]" pageId="28" pageNumber="29">
|
||
<taxonomicNameLabel id="A2398C5245762E23FBD264CAB73DB7D5" box="[1047,1137,1231,1257]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
,
|
||
<taxonomicName id="4C7E96B845762E23FBBA64CAB7A0B7D5" ID-CoL="9HV2V" authority="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda, 2021" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[1151,1260,1231,1257]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="kotana" status="sp. nov.">
|
||
<emphasis id="B90A312945762E23FBBA64CAB7A0B7D5" box="[1151,1260,1231,1257]" italics="true" pageId="28" pageNumber="29">C. kotana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90A312945762E23FB3664CAB600B7D5" bold="true" box="[1267,1356,1231,1257]" pageId="28" pageNumber="29">
|
||
<taxonomicNameLabel id="A2398C5245762E23FB3664CAB600B7D5" box="[1267,1356,1231,1257]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C7E96B845762E23FA4664CAB245B631" ID-CoL="9HWJN" authority="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda, 2021" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="matthewsi" status="sp. nov.">
|
||
<emphasis id="B90A312945762E23FA4664CAB6DAB7D5" box="[1411,1430,1231,1257]" italics="true" pageId="28" pageNumber="29">C</emphasis>
|
||
.
|
||
<emphasis id="B90A312945762E23FF5264F6B245B631" box="[151,265,1267,1293]" italics="true" pageId="28" pageNumber="29">matthewsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90A312945762E23FED564F6B225B631" bold="true" box="[272,361,1267,1293]" pageId="28" pageNumber="29">
|
||
<taxonomicNameLabel id="A2398C5245762E23FED564F6B225B631" box="[272,361,1267,1293]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |