363 lines
47 KiB
XML
363 lines
47 KiB
XML
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<mods:title id="D0B694B964EC6FA203C3918175B0B472">Phylogeny, taxonomy and flower-breeding ecology of the Colocasiomyia cristata species group (Diptera: Drosophilidae), with descriptions of ten new species</mods:title>
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<mods:namePart id="44A518E526AAB37CF656D50BA1A3F162">Takano, Kohei Takenaka</mods:namePart>
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<mods:namePart id="0C086FCEDF0F382D56975E292B47C8D6">Toda, Masanori J.</mods:namePart>
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<mods:date id="D1381A2DAA1AB4FF042C5ACB4C61F100">2021</mods:date>
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<mods:number id="E9935DB7FFCE7BB9C3FF54FBD70225F6">2021-12-08</mods:number>
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<mods:number id="67E726562FBE521865374DABE01EB7F8">5079</mods:number>
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<treatment id="03D75C2D454B2E1BFF526683B62CB1F1" ID-DOI="http://doi.org/10.5281/zenodo.5786177" ID-GBIF-Taxon="191651680" ID-Zenodo-Dep="5786177" LSID="urn:lsid:plazi:treatment:03D75C2D454B2E1BFF526683B62CB1F1" httpUri="http://treatment.plazi.org/id/03D75C2D454B2E1BFF526683B62CB1F1" lastPageId="36" lastPageNumber="37" pageId="33" pageNumber="34">
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<emphasis id="B90A3129454B2E1EFF526683B1D0B59D" bold="true" box="[151,668,1670,1697]" pageId="33" pageNumber="34">
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8)
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<taxonomicName id="4C7E96B8454B2E1EFF726683B17DB59D" ID-CoL="9HWJN" authority="Toda" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[183,561,1670,1697]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="matthewsi" status="sp. nov.">
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<emphasis id="B90A3129454B2E1EFF726683B2A6B59D" bold="true" box="[183,490,1670,1697]" italics="true" pageId="33" pageNumber="34">Colocasiomyia matthewsi</emphasis>
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Toda
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</taxonomicName>
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,
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<taxonomicNameLabel id="A2398C52454B2E1EFDFB6682B1D0B59D" box="[574,668,1671,1697]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</paragraph>
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<subSubSection id="C364BEB0454B2E1EFF5266AEB227B5F9" box="[151,363,1706,1733]" pageId="33" pageNumber="34" type="description">
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<paragraph id="8BC1ED3B454B2E1EFF5266AEB227B5F9" blockId="33.[151,668,1670,1733]" box="[151,363,1706,1733]" pageId="33" pageNumber="34">
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(
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<figureCitation id="1345F1BE454B2E1EFF5A66AEB3B0B5F9" box="[159,252,1707,1733]" captionStart="FIGURE 9" captionStartId="13.[151,250,1780,1804]" captionTargetBox="[151,1435,181,1757]" captionTargetId="figure-18@13.[151,1435,181,1757]" captionTargetPageId="13" captionText="FIGURE 9. Male abdominal sternite VIs of Colocasiomyia colocasiae (Duda) (B in anteroventral view, C in lateral view), C. stamenicola (Carson & Okada) (D in anterior view) and all species of the C. cristata subgroup (A in ventral view, E‒R in anterior view): A, C. ecornuta sp. nov.; E, C. grandis sp. nov.; F, C. vieti sp. nov.; G, C. cristata de Meijere; H, C. matthewsi sp. nov.; I, C. sulawesiana Okada & Yafuso; J, C. kinabaluana sp. nov.; K, C. kotana sp. nov.; L, C. sabahana sp. nov.; M, C. sarawakana sp. nov.; N, C. pistilicola (Carson & Okada); O, C. xenalocasiae (Okada); P, C. diconica (Toda & Okada); Q, C. sumatrana sp. nov.; and R, C. leucocasiae sp. nov. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766420" httpUri="https://zenodo.org/record/5766420/files/figure.png" pageId="33" pageNumber="34">Figs 9H</figureCitation>
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,
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<figureCitation id="1345F1BE454B2E1EFECD66AFB277B5F9" box="[264,315,1706,1733]" captionStart="FIGURE 15" captionStartId="21.[151,250,1936,1960]" captionTargetBox="[151,1435,860,1912]" captionTargetId="figure-242@21.[151,1435,860,1912]" captionTargetPageId="21" captionText="FIGURE 15. Habitus (upper, lateral view; lower, dorsal view) of species of the Colocasiomyia cristata subgroup.A, C. ecornuta sp. nov. ♂; B, C. grandis sp. nov. ♂; C, C. vieti sp. nov. ♂; D, C. cristata de Meijere ♂; E, C. sulawesiana Okada & Yafuso ♂; F, C. kinabaluana sp. nov. ♂; G, C. kotana sp. nov. ♂; and H, C. matthewsi sp. nov. ♂. Scale bars: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766434" httpUri="https://zenodo.org/record/5766434/files/figure.png" pageId="33" pageNumber="34">15H</figureCitation>
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,
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<figureCitation id="1345F1BE454B2E1EFE8366AEB22FB5F9" box="[326,355,1707,1733]" captionStart="FIGURE 23" captionStartId="35.[151,250,1464,1488]" captionTargetBox="[151,1435,183,1439]" captionTargetId="figure-18@35.[151,1435,181,1440]" captionTargetPageId="35" captionText="FIGURE 23. Colocasiomyia matthewsi Toda, sp. nov. (♂ and ♀ paratypes). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766452" httpUri="https://zenodo.org/record/5766452/files/figure.png" pageId="33" pageNumber="34">23</figureCitation>
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)
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</paragraph>
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<subSubSection id="C364BEB0454B2E1EFF5266EDB19AB43C" box="[151,726,1768,1793]" pageId="33" pageNumber="34" type="reference_group">
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<paragraph id="8BC1ED3B454B2E1EFF5266EDB19AB43C" blockId="33.[151,726,1768,1793]" box="[151,726,1768,1793]" pageId="33" pageNumber="34">
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<treatmentCitationGroup id="AB6ECA15454B2E1EFF5266EDB19AB43C" box="[151,726,1768,1793]" pageId="33" pageNumber="34">
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<taxonomicName id="4C7E96B8454B2E1EFF5266EDB19EB43C" authority=": Fartyal et al., 2013: 768" authorityName="Fartyal" authorityPageNumber="768" authorityYear="2013" box="[151,722,1768,1793]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="sp.">
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<emphasis id="B90A3129454B2E1EFF5266EDB263B43C" box="[151,303,1768,1792]" italics="true" pageId="33" pageNumber="34">Colocasiomyia</emphasis>
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sp.
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<treatmentCitation id="0ADFCB2A454B2E1EFE9C66EDB19EB43C" author="Fartyal, R. S. & Gao, J. J. & Toda, M. J. & Hu, Y. G. & Takano, K. T. & Suwito, A. & Katoh, T. & Takigahira, T. & Yin, J. T." box="[345,722,1768,1793]" page="768" pageId="33" pageNumber="34" year="2013">
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aff.
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<emphasis id="B90A3129454B2E1EFE4466ECB282B43C" box="[385,462,1769,1792]" italics="true" pageId="33" pageNumber="34">cristata</emphasis>
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:
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<bibRefCitation id="EFEF90CA454B2E1EFE1966EDB19EB43C" author="Fartyal, R. S. & Gao, J. J. & Toda, M. J. & Hu, Y. G. & Takano, K. T. & Suwito, A. & Katoh, T. & Takigahira, T. & Yin, J. T." box="[476,722,1768,1793]" pageId="33" pageNumber="34" pagination="763 - 782" refId="ref32284" refString="Fartyal, R. S., Gao, J. J., Toda, M. J., Hu, Y. G., Takano, K. T., Suwito, A., Katoh, T., Takigahira, T. & Yin, J. T. (2013) Colocasiomyia (Diptera: Drosophilidae) revised phylogenetically, with a new species group having peculiar lifecycles on monsteroid (Araceae) host plants. Systematic Entomology, 38, 763 - 782. https: // doi. org / 10.1111 / syen. 12027" type="journal article" year="2013">
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Fartyal
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<emphasis id="B90A3129454B2E1EFDEF66ECB11BB43C" box="[554,599,1768,1792]" italics="true" pageId="33" pageNumber="34">et al</emphasis>
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., 2013: 768
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</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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.
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</paragraph>
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<subSubSection id="C364BEB0454B2E1EFF52672DB65CB4CA" pageId="33" pageNumber="34" type="diagnosis">
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<paragraph id="8BC1ED3B454B2E1EFF52672DB65CB4CA" blockId="33.[151,1437,1832,2039]" pageId="33" pageNumber="34">
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<emphasis id="B90A3129454B2E1EFF52672DB25EB47E" bold="true" box="[151,274,1832,1858]" pageId="33" pageNumber="34">Diagnosis.</emphasis>
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A pair of processes on male abdominal sternite VI long (>
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<quantity id="4C8640DE454B2E1EFC4E672DB0BFB47F" box="[907,1011,1832,1859]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="33" pageNumber="34" unit="mm" value="0.05">0.05 mm</quantity>
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), widely separated, strongly divergent, slightly curved inward (
|
||
<figureCitation id="1345F1BE454B2E1EFE5A6749B2B4B45A" box="[415,504,1868,1894]" captionStart="FIGURE 9" captionStartId="13.[151,250,1780,1804]" captionTargetBox="[151,1435,181,1757]" captionTargetId="figure-18@13.[151,1435,181,1757]" captionTargetPageId="13" captionText="FIGURE 9. Male abdominal sternite VIs of Colocasiomyia colocasiae (Duda) (B in anteroventral view, C in lateral view), C. stamenicola (Carson & Okada) (D in anterior view) and all species of the C. cristata subgroup (A in ventral view, E‒R in anterior view): A, C. ecornuta sp. nov.; E, C. grandis sp. nov.; F, C. vieti sp. nov.; G, C. cristata de Meijere; H, C. matthewsi sp. nov.; I, C. sulawesiana Okada & Yafuso; J, C. kinabaluana sp. nov.; K, C. kotana sp. nov.; L, C. sabahana sp. nov.; M, C. sarawakana sp. nov.; N, C. pistilicola (Carson & Okada); O, C. xenalocasiae (Okada); P, C. diconica (Toda & Okada); Q, C. sumatrana sp. nov.; and R, C. leucocasiae sp. nov. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766420" httpUri="https://zenodo.org/record/5766420/files/figure.png" pageId="33" pageNumber="34">Fig. 9H</figureCitation>
|
||
). Epandrium pubescent except for anteroventral elongation and anterior, ventral and posteroventral margins, posteroventrally extended as ventral lobe somewhat pointed and bearing many small setae on apex, with 7–8 setae on lateral portion and 30–31 setae as thick as cercal setae on ventral lobe of each side (
|
||
<figureCitation id="1345F1BE454B2E1EFAAA6791B386B4EE" captionStart="FIGURE 23" captionStartId="35.[151,250,1464,1488]" captionTargetBox="[151,1435,183,1439]" captionTargetId="figure-18@35.[151,1435,181,1440]" captionTargetPageId="35" captionText="FIGURE 23. Colocasiomyia matthewsi Toda, sp. nov. (♂ and ♀ paratypes). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766452" httpUri="https://zenodo.org/record/5766452/files/figure.png" pageId="33" pageNumber="34">Fig. 23C</figureCitation>
|
||
). Phallal sheath apicoventrally narrowed and apically round in lateral view (
|
||
<figureCitation id="1345F1BE454B2E1EFBF367BDB7ECB4EE" box="[1078,1184,1976,2002]" captionStart="FIGURE 23" captionStartId="35.[151,250,1464,1488]" captionTargetBox="[151,1435,183,1439]" captionTargetId="figure-18@35.[151,1435,181,1440]" captionTargetPageId="35" captionText="FIGURE 23. Colocasiomyia matthewsi Toda, sp. nov. (♂ and ♀ paratypes). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766452" httpUri="https://zenodo.org/record/5766452/files/figure.png" pageId="33" pageNumber="34">Fig. 23D</figureCitation>
|
||
). Oviscapt very short, only twice as long as wide, somewhat oval, with 23–24 ovisensilla on nearly entire surface (
|
||
<figureCitation id="1345F1BE454B2E1EFB6567D9B648B4CA" box="[1184,1284,2012,2038]" captionStart="FIGURE 23" captionStartId="35.[151,250,1464,1488]" captionTargetBox="[151,1435,183,1439]" captionTargetId="figure-18@35.[151,1435,181,1440]" captionTargetPageId="35" captionText="FIGURE 23. Colocasiomyia matthewsi Toda, sp. nov. (♂ and ♀ paratypes). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766452" httpUri="https://zenodo.org/record/5766452/files/figure.png" pageId="33" pageNumber="34">Fig. 23F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF01BDB345482E1DFF526734B199B4E5" ID-DOI="http://doi.org/10.5281/zenodo.5766450" ID-Zenodo-Dep="5766450" httpUri="https://zenodo.org/record/5766450/files/figure.png" pageId="34" pageNumber="35" startId="34.[151,250,1841,1865]" targetBox="[274,1313,181,1818]" targetPageId="34">
|
||
<paragraph id="8BC1ED3B45482E1DFF526734B199B4E5" blockId="34.[151,1437,1841,2010]" pageId="34" pageNumber="35">
|
||
<emphasis id="B90A312945482E1DFF526734B26DB476" bold="true" box="[151,289,1841,1866]" pageId="34" pageNumber="35">FIGURE 22.</emphasis>
|
||
<taxonomicName id="4C7E96B845482E1DFEED6737B1F8B476" authority="Toda & Takano" authorityName="Toda & Takano" authorityYear="2021" box="[296,692,1842,1866]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="kotana" status="sp. nov.">
|
||
<emphasis id="B90A312945482E1DFEED6737B140B476" box="[296,524,1842,1866]" italics="true" pageId="34" pageNumber="35">Colocasiomyia kotana</emphasis>
|
||
Toda & Takano
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B90A312945482E1DFD7A6737B05CB476" bold="true" box="[703,784,1842,1866]" pageId="34" pageNumber="35">
|
||
<taxonomicNameLabel id="A2398C5245482E1DFD7A6737B05CB476" box="[703,784,1842,1866]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(♂ and ♀ paratypes and puparium from Kota Kinabalu, Sabah, Malaysia). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view); G, anterior spiracles of puparium (dorsal view); H, puparium (ventral habitus; inset: caudal-most portion); I, mouth hook; J, cephalopharyngeal skeleton. Scale bars: 0.1 mm in A–F and J, 1.0 mm in H.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF01BDB345492E1CFF5265BDB041B52B" ID-DOI="http://doi.org/10.5281/zenodo.5766452" ID-Zenodo-Dep="5766452" httpUri="https://zenodo.org/record/5766452/files/figure.png" pageId="35" pageNumber="36" startId="35.[151,250,1464,1488]" targetBox="[151,1435,183,1439]" targetPageId="35">
|
||
<paragraph id="8BC1ED3B45492E1CFF5265BDB041B52B" blockId="35.[151,1437,1464,1560]" pageId="35" pageNumber="36">
|
||
<emphasis id="B90A312945492E1CFF5265BDB26EB6EC" bold="true" box="[151,290,1464,1488]" pageId="35" pageNumber="36">FIGURE 23.</emphasis>
|
||
<taxonomicName id="4C7E96B845492E1CFEEF65BDB122B6EC" authority="Toda" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[298,622,1464,1488]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="matthewsi" status="sp. nov.">
|
||
<emphasis id="B90A312945492E1CFEEF65BDB17DB6EC" box="[298,561,1464,1488]" italics="true" pageId="35" pageNumber="36">Colocasiomyia matthewsi</emphasis>
|
||
Toda
|
||
</taxonomicName>
|
||
,
|
||
<emphasis id="B90A312945492E1CFDBF65BDB181B6EC" bold="true" box="[634,717,1464,1488]" pageId="35" pageNumber="36">
|
||
<taxonomicNameLabel id="A2398C5245492E1CFDBF65BDB181B6EC" box="[634,717,1464,1488]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(♂ and ♀ paratypes). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view). Scale bars: 0.1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C364BEB045492E1BFF026643B243B3E9" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="description">
|
||
<paragraph id="8BC1ED3B45492E1CFF026643B19FB594" blockId="35.[151,1437,1606,2029]" pageId="35" pageNumber="36">
|
||
<emphasis id="B90A312945492E1CFF026643B21DB55C" bold="true" box="[199,337,1606,1632]" pageId="35" pageNumber="36">Description</emphasis>
|
||
(
|
||
<specimenCount id="9D7826B245492E1CFEA56643B239B55C" box="[352,373,1606,1632]" pageId="35" pageNumber="36" type="male">♂</specimenCount>
|
||
and
|
||
<specimenCount id="9D7826B245492E1CFE6C6642B2F3B55D" box="[425,447,1607,1633]" pageId="35" pageNumber="36" type="female">♀</specimenCount>
|
||
; not repeating characters common to
|
||
<taxonomicName id="4C7E96B845492E1CFC9B6642B745B55C" authority="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda, 2021" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[862,1033,1606,1633]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="kinabaluana" status="sp. nov.">
|
||
<emphasis id="B90A312945492E1CFC9B6642B03DB55D" box="[862,881,1607,1633]" italics="true" pageId="35" pageNumber="36">C</emphasis>
|
||
.
|
||
<emphasis id="B90A312945492E1CFCB86643B745B55C" box="[893,1033,1606,1632]" italics="true" pageId="35" pageNumber="36">kinabaluana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90A312945492E1CFBCA6643B72BB55C" bold="true" box="[1039,1127,1606,1632]" pageId="35" pageNumber="36">
|
||
<taxonomicNameLabel id="A2398C5245492E1CFBCA6643B72BB55C" box="[1039,1127,1606,1632]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
). Head. Supracervical setae 5–8 per side, distally more or less curved, longer than inner occipital setae. Cibarial, medial sensilla 3–4 per side. Supralateral setae outside prementum 3–5 per side.
|
||
</paragraph>
|
||
<paragraph id="8BC1ED3B45492E1CFF0266B7B03EB5F1" blockId="35.[151,1437,1606,2029]" box="[199,882,1714,1741]" pageId="35" pageNumber="36">Thorax. Anterior dorsocentral setae behind transverse suture.</paragraph>
|
||
<paragraph id="8BC1ED3B45492E1CFF0266D3B671B428" blockId="35.[151,1437,1606,2029]" pageId="35" pageNumber="36">
|
||
Abdomen. Male sternite III longer than wide, posteriorly slightly narrowing; IV wider than long (
|
||
<figureCitation id="1345F1BE45492E1CFAE066D3B6DCB5CC" box="[1317,1424,1750,1776]" captionStart="FIGURE 23" captionStartId="35.[151,250,1464,1488]" captionTargetBox="[151,1435,183,1439]" captionTargetId="figure-18@35.[151,1435,181,1440]" captionTargetPageId="35" captionText="FIGURE 23. Colocasiomyia matthewsi Toda, sp. nov. (♂ and ♀ paratypes). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766452" httpUri="https://zenodo.org/record/5766452/files/figure.png" pageId="35" pageNumber="36">Fig. 23A</figureCitation>
|
||
). Female sternite V wider than long; VI wider than long, medially concaved on posterior margin (
|
||
<figureCitation id="1345F1BE45492E1CFB0F66FFB67DB428" box="[1226,1329,1786,1812]" captionStart="FIGURE 23" captionStartId="35.[151,250,1464,1488]" captionTargetBox="[151,1435,183,1439]" captionTargetId="figure-18@35.[151,1435,181,1440]" captionTargetPageId="35" captionText="FIGURE 23. Colocasiomyia matthewsi Toda, sp. nov. (♂ and ♀ paratypes). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766452" httpUri="https://zenodo.org/record/5766452/files/figure.png" pageId="35" pageNumber="36">Fig. 23B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC1ED3B45492E1CFF02671BB158B4BC" blockId="35.[151,1437,1606,2029]" pageId="35" pageNumber="36">
|
||
Male terminalia. Cercus pubescent only on dorsal 2/5, with 56–57 setae, extended below, apically forming broad lobe marginally roundish and fringed with small setae (
|
||
<figureCitation id="1345F1BE45492E1CFC9A6747B084B460" box="[863,968,1858,1884]" captionStart="FIGURE 23" captionStartId="35.[151,250,1464,1488]" captionTargetBox="[151,1435,183,1439]" captionTargetId="figure-18@35.[151,1435,181,1440]" captionTargetPageId="35" captionText="FIGURE 23. Colocasiomyia matthewsi Toda, sp. nov. (♂ and ♀ paratypes). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766452" httpUri="https://zenodo.org/record/5766452/files/figure.png" pageId="35" pageNumber="36">Fig. 23C</figureCitation>
|
||
). Phallapodeme sclerotized plate slightly oblique to phallal axis (
|
||
<figureCitation id="1345F1BE45492E1CFE5A6763B14BB4BC" box="[415,519,1894,1920]" captionStart="FIGURE 23" captionStartId="35.[151,250,1464,1488]" captionTargetBox="[151,1435,183,1439]" captionTargetId="figure-18@35.[151,1435,181,1440]" captionTargetPageId="35" captionText="FIGURE 23. Colocasiomyia matthewsi Toda, sp. nov. (♂ and ♀ paratypes). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766452" httpUri="https://zenodo.org/record/5766452/files/figure.png" pageId="35" pageNumber="36">Fig. 23D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC1ED3B45492E1CFF02678FB737B498" blockId="35.[151,1437,1606,2029]" box="[199,1147,1930,1957]" pageId="35" pageNumber="36">
|
||
Female terminalia. Hypoproct medially with a small patch of pubescence (
|
||
<figureCitation id="1345F1BE45492E1CFBCF678FB722B498" box="[1034,1134,1930,1956]" captionStart="FIGURE 23" captionStartId="35.[151,250,1464,1488]" captionTargetBox="[151,1435,183,1439]" captionTargetId="figure-18@35.[151,1435,181,1440]" captionTargetPageId="35" captionText="FIGURE 23. Colocasiomyia matthewsi Toda, sp. nov. (♂ and ♀ paratypes). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766452" httpUri="https://zenodo.org/record/5766452/files/figure.png" pageId="35" pageNumber="36">Fig. 23F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC1ED3B45492E1BFF0267ABB243B3E9" blockId="35.[151,1437,1606,2029]" lastBlockId="36.[151,1437,150,717]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
|
||
Indices (range of
|
||
<specimenCount id="9D7826B245492E1CFE4A67ABB28CB4F4" box="[399,448,1966,1992]" pageId="35" pageNumber="36" type="male">10♂</specimenCount>
|
||
and
|
||
<specimenCount id="9D7826B245492E1CFE3C67ABB166B4F4" box="[505,554,1966,1992]" pageId="35" pageNumber="36" type="female">10♀</specimenCount>
|
||
<typeStatus id="54C5539945492E1CFDF767AAB1EFB4F5" box="[562,675,1967,1993]" pageId="35" pageNumber="36" type="paratype">paratypes</typeStatus>
|
||
): FW/HW = 0.53–0.58, ch/o = 0.28–0.36, prorb = 0.86–1.14, rcorb = 0.33–0.56, vb = 0.26–0.47, dcl = 0.57–0.74, sctl = 0.74–0.98, sterno = 0.65–1.10, orbito = 0.45–0.82, dcp = 0.97–1.30, sctlp = 1.27–1.82, C = 1.74–2.12, 4c = 0.92–1.08, 4v = 1.42–1.60, 5x = 0.72–1.14, ac = 2.60–3.45, M = 0.28–0.40.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364BEB0454E2E1BFF0260DAB772B27D" pageId="36" pageNumber="37" type="materials_examined">
|
||
<paragraph id="8BC1ED3B454E2E1BFF0260DAB28EB221" blockId="36.[151,1437,150,717]" pageId="36" pageNumber="37">
|
||
<materialsCitation id="3B16E766454E2E1BFF0260DAB28EB221" ID-GBIF-Occurrence="3418896747" collectingDate="2012-12-02" collectionCode="MPMP" country="Philippines" location="Gandara" pageId="36" pageNumber="37" specimenCount="1" specimenCount-male="1" stateProvince="Samar" typeStatus="holotype">
|
||
<emphasis id="B90A3129454E2E1BFF0260DAB277B3C5" bold="true" box="[199,315,223,249]" pageId="36" pageNumber="37">
|
||
<typeStatus id="54C55399454E2E1BFF0260DAB27AB3C5" box="[199,310,223,249]" pageId="36" pageNumber="37" type="holotype">Holotype</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<specimenCount id="9D7826B2454E2E1BFE8760DAB21BB3C5" box="[322,343,223,249]" pageId="36" pageNumber="37" type="male">♂</specimenCount>
|
||
(
|
||
<collectionCode id="ED6F75FE454E2E1BFEA060DAB2F0B3C5" box="[357,444,223,249]" country="Philippines" httpUri="http://grbio.org/cool/i3xh-eq16" name="National Museum of the Philippines" pageId="36" pageNumber="37">MPMP</collectionCode>
|
||
), “
|
||
<location id="8EA1BBE0454E2E1BFE1C60DAB171B3C5" LSID="urn:lsid:plazi:treatment:03D75C2D454B2E1BFF526683B62CB1F1:8EA1BBE0454E2E1BFE1C60DAB171B3C5" box="[473,573,223,249]" country="Philippines" name="Gandara" pageId="36" pageNumber="37" stateProvince="Samar">Gandara</location>
|
||
,
|
||
<collectingRegion id="49BA23D9454E2E1BFD8D60DAB1DDB3C5" box="[584,657,223,249]" country="Philippines" name="Samar" pageId="36" pageNumber="37">Samar</collectingRegion>
|
||
,
|
||
<collectingCountry id="F369ADAB454E2E1BFD5960DAB051B3C5" box="[668,797,223,249]" name="Philippines" pageId="36" pageNumber="37">Philippines</collectingCountry>
|
||
,
|
||
<date id="FFC0CBFB454E2E1BFCED60DAB0D7B3C5" box="[808,923,223,249]" pageId="36" pageNumber="37" value="2012-12-02">
|
||
<collectingDate id="EF843213454E2E1BFCED60DAB0D7B3C5" box="[808,923,223,249]" pageId="36" pageNumber="37" value="2012-12-02">2.xii.2012</collectingDate>
|
||
</date>
|
||
, ex
|
||
<taxonomicName id="4C7E96B8454E2E1BFC0D60DAB2F8B221" authority=", P. J. Matthews, M. Medecilo & J. R. Castillo" authorityName="P. J. Matthews, M. Medecilo & J. R. Castillo" class="Liliopsida" family="Araceae" genus="Alocasia" kingdom="Plantae" order="Alismatales" pageId="36" pageNumber="37" phylum="Tracheophyta" rank="species" species="macrorrhizos">
|
||
<emphasis id="B90A3129454E2E1BFC0D60DAB78AB3C5" box="[968,1222,223,249]" italics="true" pageId="36" pageNumber="37">Alocasia macrorrhizos</emphasis>
|
||
, P.J. Matthews, M. Medecilo & J.R. Castillo
|
||
</taxonomicName>
|
||
”.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BC1ED3B454E2E1BFF026122B772B27D" blockId="36.[151,1437,150,717]" box="[199,1086,295,321]" pageId="36" pageNumber="37">
|
||
<materialsCitation id="3B16E766454E2E1BFF026122B776B27D" ID-GBIF-Occurrence="3418896567" box="[199,1082,295,321]" collectingDate="2012-12-02" collectionCode="MPMP, SEHU" country="Philippines" location="Gandara" pageId="36" pageNumber="37" specimenCount="20" specimenCount-female="10" specimenCount-male="10" stateProvince="Samar" typeStatus="paratype">
|
||
<emphasis id="B90A3129454E2E1BFF026122B20AB27D" bold="true" box="[199,326,295,321]" pageId="36" pageNumber="37">
|
||
<typeStatus id="54C55399454E2E1BFF026122B20DB27D" box="[199,321,295,321]" pageId="36" pageNumber="37" type="paratype">Paratypes</typeStatus>
|
||
.
|
||
</emphasis>
|
||
<collectingCountry id="F369ADAB454E2E1BFE886122B29CB27D" box="[333,464,295,321]" name="Philippines" pageId="36" pageNumber="37">Philippines</collectingCountry>
|
||
:
|
||
<specimenCount id="9D7826B2454E2E1BFE1F6122B141B27D" box="[474,525,295,321]" pageId="36" pageNumber="37" type="male">10♂</specimenCount>
|
||
,
|
||
<specimenCount id="9D7826B2454E2E1BFDDC6122B100B27D" box="[537,588,295,321]" pageId="36" pageNumber="37" type="female">10♀</specimenCount>
|
||
, same data as the holotype (
|
||
<collectionCode id="ED6F75FE454E2E1BFC436122B095B27D" box="[902,985,295,321]" country="Philippines" httpUri="http://grbio.org/cool/i3xh-eq16" name="National Museum of the Philippines" pageId="36" pageNumber="37">MPMP</collectionCode>
|
||
,
|
||
<collectionCode id="ED6F75FE454E2E1BFC216122B77EB27D" box="[996,1074,295,321]" pageId="36" pageNumber="37">SEHU</collectionCode>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364BEB0454E2E1BFF02614FB102B259" box="[199,590,330,357]" pageId="36" pageNumber="37" type="distribution">
|
||
<paragraph id="8BC1ED3B454E2E1BFF02614FB102B259" blockId="36.[151,1437,150,717]" box="[199,590,330,357]" pageId="36" pageNumber="37">
|
||
<emphasis id="B90A3129454E2E1BFF02614FB22DB258" bold="true" box="[199,353,330,356]" pageId="36" pageNumber="37">Distribution.</emphasis>
|
||
<collectingCountry id="F369ADAB454E2E1BFEA2614EB2A9B259" box="[359,485,331,357]" name="Philippines" pageId="36" pageNumber="37">Philippines</collectingCountry>
|
||
(
|
||
<collectingRegion id="49BA23D9454E2E1BFE31614EB10EB259" box="[500,578,331,357]" country="Philippines" name="Samar" pageId="36" pageNumber="37">Samar</collectingRegion>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364BEB0454E2E1BFF02616AB0D4B195" pageId="36" pageNumber="37" type="discussion">
|
||
<paragraph id="8BC1ED3B454E2E1BFF02616AB0D4B195" blockId="36.[151,1437,150,717]" pageId="36" pageNumber="37">
|
||
<emphasis id="B90A3129454E2E1BFF02616AB277B2B5" bold="true" box="[199,315,367,393]" pageId="36" pageNumber="37">Remarks.</emphasis>
|
||
This species was placed as the basal branch sister to the clade (BP = 100) consisting of the remaining four species of the
|
||
<taxonomicName id="4C7E96B8454E2E1BFEB46196B28BB290" box="[369,455,403,428]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="cristata">
|
||
<emphasis id="B90A3129454E2E1BFEB46196B28BB290" box="[369,455,403,428]" italics="true" pageId="36" pageNumber="37">cristata</emphasis>
|
||
</taxonomicName>
|
||
complex (
|
||
<figureCitation id="1345F1BE454E2E1BFD856196B1D8B291" box="[576,660,403,429]" captionStart="FIGURE 13" captionStartId="18.[151,250,1328,1352]" captionTargetBox="[151,1433,181,1302]" captionTargetId="figure-22@18.[151,1436,181,1305]" captionTargetPageId="18" captionText="FIGURE 13. The strict consensus tree resulting from the parsimony analysis (PAUP* v4.0a168) of the data matrix of 25 spp. × 51 morphological characters (Appendix 6) for 20 species of the Colocasiomyia cristata group (as ingroup) and five species of the C. gigantea group (as outgroup). The tree is rooted by outgroup rooting, and synapomorphies (solid circle: nonhomoplastic; open circle: homoplastic) inferred from both ACCTRAN and DELTRAN character optimization are indicated on each internal branch along with support value (BP). Codes of clades (I and II) and subclades (IIa–c) are shown beside the corresponding branches (see the text for more explanation)." figureDoi="http://doi.org/10.5281/zenodo.5766428" httpUri="https://zenodo.org/record/5766428/files/figure.png" pageId="36" pageNumber="37">Fig. 13</figureCitation>
|
||
), lacking the following synapomorphies for the latter clade: anterior dorsocentral setae just beside transverse suture (ch.7-1), epandrium posteroventrally roundish (ch.30-1) and with setae thicker than cercal setae (ch.32-1), phallapodeme less sclerotized thin plate (ch.45-2) directed nearly perpendicular to axis of phallal sheath (ch.46-1), and hypoproct not pubescent (ch.48-1). In addition, within the latter clade,
|
||
<taxonomicName id="4C7E96B8454E2E1BFF256226B2C7B101" box="[224,395,547,573]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="sulawesiana">
|
||
<emphasis id="B90A3129454E2E1BFF256226B2C7B101" box="[224,395,547,573]" italics="true" pageId="36" pageNumber="37">C. sulawesiana</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7E96B8454E2E1BFE5C6226B109B101" authority="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda, 2021" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[409,581,547,573]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="kinabaluana" status="sp. nov.">
|
||
<emphasis id="B90A3129454E2E1BFE5C6226B109B101" box="[409,581,547,573]" italics="true" pageId="36" pageNumber="37">C. kinabaluana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90A3129454E2E1BFD896226B1E9B101" bold="true" box="[588,677,547,573]" pageId="36" pageNumber="37">
|
||
<taxonomicNameLabel id="A2398C52454E2E1BFD896226B1E9B101" box="[588,677,547,573]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C7E96B8454E2E1BFD1E6226B004B101" authority="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda, 2021" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[731,840,547,573]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="kotana" status="sp. nov.">
|
||
<emphasis id="B90A3129454E2E1BFD1E6226B004B101" box="[731,840,547,573]" italics="true" pageId="36" pageNumber="37">C. kotana</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90A3129454E2E1BFC8A6226B0E4B101" bold="true" box="[847,936,547,573]" pageId="36" pageNumber="37">
|
||
<taxonomicNameLabel id="A2398C52454E2E1BFC8A6226B0E4B101" box="[847,936,547,573]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
formed a clade (BP = 75;
|
||
<figureCitation id="1345F1BE454E2E1BFB0B6226B652B101" box="[1230,1310,547,573]" captionStart="FIGURE 13" captionStartId="18.[151,250,1328,1352]" captionTargetBox="[151,1433,181,1302]" captionTargetId="figure-22@18.[151,1436,181,1305]" captionTargetPageId="18" captionText="FIGURE 13. The strict consensus tree resulting from the parsimony analysis (PAUP* v4.0a168) of the data matrix of 25 spp. × 51 morphological characters (Appendix 6) for 20 species of the Colocasiomyia cristata group (as ingroup) and five species of the C. gigantea group (as outgroup). The tree is rooted by outgroup rooting, and synapomorphies (solid circle: nonhomoplastic; open circle: homoplastic) inferred from both ACCTRAN and DELTRAN character optimization are indicated on each internal branch along with support value (BP). Codes of clades (I and II) and subclades (IIa–c) are shown beside the corresponding branches (see the text for more explanation)." figureDoi="http://doi.org/10.5281/zenodo.5766428" httpUri="https://zenodo.org/record/5766428/files/figure.png" pageId="36" pageNumber="37">Fig. 13</figureCitation>
|
||
) supported by an autapomorphy, the short (<
|
||
<quantity id="4C8640DE454E2E1BFDD66243B130B15D" box="[531,636,582,609]" metricMagnitude="-5" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="37" unit="mm" value="0.05">0.05 mm</quantity>
|
||
) processes on the male sternite VI (ch.25-1;
|
||
<figureCitation id="1345F1BE454E2E1BFB446242B7B8B15D" box="[1153,1268,583,609]" captionStart="FIGURE 9" captionStartId="13.[151,250,1780,1804]" captionTargetBox="[151,1435,181,1757]" captionTargetId="figure-18@13.[151,1435,181,1757]" captionTargetPageId="13" captionText="FIGURE 9. Male abdominal sternite VIs of Colocasiomyia colocasiae (Duda) (B in anteroventral view, C in lateral view), C. stamenicola (Carson & Okada) (D in anterior view) and all species of the C. cristata subgroup (A in ventral view, E‒R in anterior view): A, C. ecornuta sp. nov.; E, C. grandis sp. nov.; F, C. vieti sp. nov.; G, C. cristata de Meijere; H, C. matthewsi sp. nov.; I, C. sulawesiana Okada & Yafuso; J, C. kinabaluana sp. nov.; K, C. kotana sp. nov.; L, C. sabahana sp. nov.; M, C. sarawakana sp. nov.; N, C. pistilicola (Carson & Okada); O, C. xenalocasiae (Okada); P, C. diconica (Toda & Okada); Q, C. sumatrana sp. nov.; and R, C. leucocasiae sp. nov. Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766420" httpUri="https://zenodo.org/record/5766420/files/figure.png" pageId="36" pageNumber="37">Fig. 9I–K</figureCitation>
|
||
). Furthermore,
|
||
<taxonomicName id="4C7E96B8454E2E1BFF52626EB266B1B9" authority="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda, 2021" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[151,298,619,645]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="matthewsi" status="sp. nov.">
|
||
<emphasis id="B90A3129454E2E1BFF52626EB266B1B9" box="[151,298,619,645]" italics="true" pageId="36" pageNumber="37">C. matthewsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90A3129454E2E1BFEF4626EB2C6B1B9" bold="true" box="[305,394,619,645]" pageId="36" pageNumber="37">
|
||
<taxonomicNameLabel id="A2398C52454E2E1BFEF4626EB2C6B1B9" box="[305,394,619,645]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
somewhat resembles
|
||
<taxonomicName id="4C7E96B8454E2E1BFD44626EB1B4B1B9" authority="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda, 2021" authorityName="Takano & Gao & Hu & Li & Yafuso & Suwito & Repin & Pungga & Meleng & Kaliang & Chong & Toda" authorityYear="2021" box="[641,760,619,645]" class="Insecta" family="Drosophilidae" genus="Colocasiomyia" kingdom="Animalia" order="Diptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="species" species="grandis" status="sp. nov.">
|
||
<emphasis id="B90A3129454E2E1BFD44626EB1B4B1B9" box="[641,760,619,645]" italics="true" pageId="36" pageNumber="37">C. grandis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B90A3129454E2E1BFD3A626EB014B1B9" bold="true" box="[767,856,619,645]" pageId="36" pageNumber="37">
|
||
<taxonomicNameLabel id="A2398C52454E2E1BFD3A626EB014B1B9" box="[767,856,619,645]" pageId="36" pageNumber="37" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in the morphology of periphallic organs (
|
||
<figureCitation id="1345F1BE454E2E1BFAE9626EB6DBB1B9" box="[1324,1431,619,645]" captionStart="FIGURE 16" captionStartId="24.[151,250,1785,1809]" captionTargetBox="[242,1345,181,1761]" captionTargetId="figure-22@24.[242,1345,181,1761]" captionTargetPageId="24" captionText="FIGURE 16. Colocasiomyia ecornuta Toda & Takano, sp. nov. (♂ and ♀ paratypes and puparium from Soppeng, South Sulawesi, Indonesia). A, Head (frontal view); B, scutum and scutellum (dorsal view); C, D, abdominal sternites in ventral view (C, male; D, female); E, periphallic organs (caudolateral view; inset: cercal ventral lobe); F, phallal sheath (dorsal view); G, H, phallic organs (G, lateral view; H, ventral view); I peri-oviscapt (caudal view); J, oviscapt (ventral view); K, puparium (ventral habitus; inset: caudal-most portion at lower right and ventrolateral portion of segments at the upper right); L, mouth hook; M, cephalopharyngeal skeleton. Abbreviations: i vt s, inner vertical setae; sctl, scutellum. Scale bars: 0.1 mm in A–J and M, 1.0 mm in K." figureDoi="http://doi.org/10.5281/zenodo.5766438" httpUri="https://zenodo.org/record/5766438/files/figure.png" pageId="36" pageNumber="37">Figs 16D</figureCitation>
|
||
,
|
||
<figureCitation id="1345F1BE454E2E1BFF52628AB385B195" box="[151,201,655,681]" captionStart="FIGURE 23" captionStartId="35.[151,250,1464,1488]" captionTargetBox="[151,1435,183,1439]" captionTargetId="figure-18@35.[151,1435,181,1440]" captionTargetPageId="35" captionText="FIGURE 23. Colocasiomyia matthewsi Toda, sp. nov. (♂ and ♀ paratypes). A, Male abdominal sternites (ventral view); B, female abdominal sternites (ventral view); C, periphallic organs (lateral view); D, phallic organs (lateral view); E, ditto (ventral view); F, female terminalia (lateral view). Scale bars: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.5766452" httpUri="https://zenodo.org/record/5766452/files/figure.png" pageId="36" pageNumber="37">23C</figureCitation>
|
||
), but can be distinguished from it by their diagnostic characters.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C364BEB0454E2E1BFF0262B6B62CB1F1" box="[199,1376,691,717]" pageId="36" pageNumber="37" type="etymology">
|
||
<paragraph id="8BC1ED3B454E2E1BFF0262B6B62CB1F1" blockId="36.[151,1437,150,717]" box="[199,1376,691,717]" pageId="36" pageNumber="37">
|
||
<emphasis id="B90A3129454E2E1BFF0262B6B202B1F1" bold="true" box="[199,334,691,717]" pageId="36" pageNumber="37">Etymology.</emphasis>
|
||
Patronym, dedicated to Prof. Peter J. Matthews, one of the three collectors of the specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |