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<document id="FA4386F416482DC6A03CE87385E6F205" ID-DOI="10.11646/zootaxa.3947.1.6" ID-GBIF-Dataset="1cec74cd-3ea8-4aa9-980a-c820df158f39" ID-ISSN="1175-5326" ID-Zenodo-Dep="243970" ID-ZooBank="6D036999-335A-4C36-8243-0608938C8DD0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461039611458" checkinUser="plazi" docAuthor="Mengual, Ximo, Kazerani, Farzaneh, Talebi, Ali Asghar &amp; Gilasian, Ebrahim" docDate="2015" docId="03E1513DD440FFB9FF3EF92EEE4EFBEF" docLanguage="en" docName="zt03947p108.pdf" docOrigin="Zootaxa 3947 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pelecocera persiana Kuznetzov" docType="treatment" docVersion="8" lastPageNumber="101" masterDocId="FFD82945D441FFBBFFA9FFA2EB58FF90" masterDocTitle="A revision of the genus Pelecocera Meigen with the description of the male of Pelecocera persiana Kuznetzov from Iran (Diptera: Syrphidae)" masterLastPageNumber="108" masterPageNumber="99" pageNumber="100" updateTime="1698575321531" updateUser="plazi">
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<mods:title id="986281A2B8E1399645C3AA4E01E7A985">A revision of the genus Pelecocera Meigen with the description of the male of Pelecocera persiana Kuznetzov from Iran (Diptera: Syrphidae)</mods:title>
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<mods:namePart id="68F0888704A122AE898AB9DC8236F505">Mengual, Ximo</mods:namePart>
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<mods:namePart id="022F627E6D13FC913D07930B58DE4D01">Kazerani, Farzaneh</mods:namePart>
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<mods:namePart id="DF2B0FBBA8F80BF8C84FF6CFD72A9D39">Talebi, Ali Asghar</mods:namePart>
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<mods:namePart id="80E21681DAA6A406F9622BEEBEE286A3">Gilasian, Ebrahim</mods:namePart>
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<treatment id="03E1513DD440FFB9FF3EF92EEE4EFBEF" ID-DOI="http://doi.org/10.5281/zenodo.6108183" ID-GBIF-Taxon="119609560" ID-Zenodo-Dep="6108183" LSID="urn:lsid:plazi:treatment:03E1513DD440FFB9FF3EF92EEE4EFBEF" httpUri="http://treatment.plazi.org/id/03E1513DD440FFB9FF3EF92EEE4EFBEF" lastPageId="2" lastPageNumber="101" pageId="1" pageNumber="100">
<subSubSection id="C352B3A0D440FFBAFF3EF92EEE4FF854" pageId="1" pageNumber="100" type="nomenclature">
<paragraph id="8BF7E02BD440FFBAFF3EF92EE94FF936" blockId="1.[151,535,1676,1736]" box="[151,535,1676,1702]" pageId="1" pageNumber="100">
<heading id="D0BF5747D440FFBAFF3EF92EE94FF936" bold="true" box="[151,535,1676,1702]" fontSize="11" level="1" pageId="1" pageNumber="100" reason="1">
<taxonomicName id="4C489BA8D440FFBAFF3EF92EE94FF936" ID-CoL="75Z98" authority="Kuznetzov" authorityName="Kuznetzov" box="[151,535,1676,1702]" class="Insecta" family="Syrphidae" genus="Pelecocera" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="persiana">
<emphasis id="B93C3C39D440FFBAFF3EF92EE94FF936" bold="true" box="[151,535,1676,1702]" pageId="1" pageNumber="100">
<emphasis id="B93C3C39D440FFBAFF3EF92EEAD2F936" bold="true" box="[151,394,1676,1702]" italics="true" pageId="1" pageNumber="100">Pelecocera persiana</emphasis>
Kuznetzov
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BF7E02BD440FFBAFF3EF90DEA43F958" blockId="1.[151,535,1676,1736]" box="[151,283,1711,1736]" pageId="1" pageNumber="100">
<figureCitation id="1373FCAED440FFBAFF3EF90DEA4FF958" box="[151,279,1711,1736]" captionStart="FIGURES 1 6. 1" captionStartId="4.[151,264,1924,1947]" captionTargetBox="[243,1348,202,1893]" captionTargetId="figure@4.[236,1350,195,1900]" captionTargetPageId="4" captionText="FIGURES 1 6. 1. Pelecocera persiana, male, dorsal view (scale: 2 mm). 2. Pelecocera latifrons, male, dorsal view (scale: 2 mm). 3. Pelecocera persiana, male, lateral view (scale: 2 mm). 4. Pelecocera persiana, male, frontal view (scale: 1 mm). 5. Pelecocera latifrons, male, lateral view (scale: 2 mm). 6. Pelecocera latifrons, male, frontal view (scale: 1 mm)." httpUri="https://zenodo.org/record/243971/files/figure.png" pageId="1" pageNumber="100">Figs 1, 3, 4</figureCitation>
.
</paragraph>
<paragraph id="8BF7E02BD440FFBAFF3EF955EE4FF854" blockId="1.[151,1437,1783,2024]" pageId="1" pageNumber="100">
<emphasis id="B93C3C39D440FFBAFF3EF955EAC6F880" bold="true" box="[151,414,1783,1808]" pageId="1" pageNumber="100">Differential diagnosis.</emphasis>
Species with almost the entire body densely grey pollinose and the frons broader than eye width in dorsal view.
<taxonomicName id="4C489BA8D440FFBAFE20F8B9E928F8A3" box="[393,624,1819,1844]" class="Insecta" family="Syrphidae" genus="Pelecocera" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="persiana">
<emphasis id="B93C3C39D440FFBAFE20F8B9E928F8A3" box="[393,624,1819,1844]" italics="true" pageId="1" pageNumber="100">Pelecocera persiana</emphasis>
</taxonomicName>
is close to
<taxonomicName id="4C489BA8D440FFBAFD46F8BEE831F8A4" box="[751,873,1819,1844]" class="Insecta" family="Syrphidae" genus="Pelecocera" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="B93C3C39D440FFBAFD46F8BEE831F8A4" box="[751,873,1819,1844]" italics="true" pageId="1" pageNumber="100">P. latifrons</emphasis>
</taxonomicName>
, but differs in the following characters: male with postpronotum yellow (black in
<taxonomicName id="4C489BA8D440FFBAFE5DF8E2E936F8C8" box="[500,622,1855,1880]" class="Insecta" family="Syrphidae" genus="Pelecocera" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="B93C3C39D440FFBAFE5DF8E2E936F8C8" box="[500,622,1855,1880]" italics="true" pageId="1" pageNumber="100">P. latifrons</emphasis>
</taxonomicName>
), male with scutellum black with pale posterior margin (entirely black in
<taxonomicName id="4C489BA8D440FFBAFF3EF8C6EA4DF8EC" box="[151,277,1891,1916]" class="Insecta" family="Syrphidae" genus="Pelecocera" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="B93C3C39D440FFBAFF3EF8C6EA4DF8EC" box="[151,277,1891,1916]" italics="true" pageId="1" pageNumber="100">P. latifrons</emphasis>
</taxonomicName>
); following
<bibRefCitation id="EFD99DDAD440FFBAFE00F8C6E923F8EC" author="Kuznetzov" box="[425,635,1892,1916]" pageId="1" pageNumber="100" refString="Kuznetzov, S. Y. (1989) Hover-flies of the genus Pelecocera Mg. (Diptera, Syrphidae). Latvian Entomology, 32, 80 - 85." type="journal article" year="1989">Kuznetzov (1989)</bibRefCitation>
, female with broader frons and differently shaped basoflagellomere. Male genitalia are very similar in these two species and are a bit larger in
<taxonomicName id="4C489BA8D440FFBAFC77F82AEF05F80F" box="[990,1117,1928,1951]" class="Insecta" family="Syrphidae" genus="Pelecocera" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="persiana">
<emphasis id="B93C3C39D440FFBAFC77F82AEF05F80F" box="[990,1117,1928,1951]" italics="true" pageId="1" pageNumber="100">P. persiana</emphasis>
</taxonomicName>
. However,
<taxonomicName id="4C489BA8D440FFBAFB49F82AEE02F830" box="[1248,1370,1927,1952]" class="Insecta" family="Syrphidae" genus="Pelecocera" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="latifrons">
<emphasis id="B93C3C39D440FFBAFB49F82AEE02F830" box="[1248,1370,1927,1952]" italics="true" pageId="1" pageNumber="100">P. latifrons</emphasis>
</taxonomicName>
has a slightly larger cercus and a dorsal incision on the surstylus that
<taxonomicName id="4C489BA8D440FFBAFCF7F80EE883F853" box="[862,987,1964,1987]" class="Insecta" family="Syrphidae" genus="Pelecocera" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="100" phylum="Arthropoda" rank="species" species="persiana">
<emphasis id="B93C3C39D440FFBAFCF7F80EE883F853" box="[862,987,1964,1987]" italics="true" pageId="1" pageNumber="100">P. persiana</emphasis>
</taxonomicName>
seems to lack (
<figureCitation id="1373FCAED440FFBAFB21F809EE53F854" box="[1160,1291,1963,1988]" captionStart="FIGURES 16 23. 16" captionStartId="6.[151,264,1871,1894]" captionTargetBox="[257,1335,197,1846]" captionTargetId="figure@6.[250,1340,193,1849]" captionTargetPageId="6" captionText="FIGURES 16 23. 16. Pelecocera pergandei, male, lateral view (scale: 2 mm). 17. Pelecocera latifrons, lectotype male, frontal view (scale: 1 mm). 18. Pelecocera pergandei, male, dorsal view (scale: 2 mm). 19. Pelecocera latifrons, lectotype male, dorsal view (scale: 2 mm). 20. Pelecocera latifrons, lectotype male, lateral view (scale: 2 mm). 21. Pelecocera latifrons, lectotype male, labels. 22. Pelecocera latifrons, lectotype male, lateral view male genitalia (scale: 0.5 mm). 23. Pelecocera persiana, male, lateral view male genitalia (scale: 0.5 mm)." httpUri="https://zenodo.org/record/243973/files/figure.png" pageId="1" pageNumber="100">Figs 22, 23</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C352B3A0D440FFB9FF6EF86DEE31FC7F" lastPageId="2" lastPageNumber="101" pageId="1" pageNumber="100" type="description">
<paragraph id="8BF7E02BD440FFB9FF6EF86DEE31FC7F" blockId="1.[151,1437,1783,2024]" lastBlockId="2.[151,1437,151,1151]" lastPageId="2" lastPageNumber="101" pageId="1" pageNumber="100">
<emphasis id="B93C3C39D440FFBAFF6EF86DEA9FF878" bold="true" box="[199,455,1999,2024]" pageId="1" pageNumber="100">Description of male.</emphasis>
Body length: 7.1 mm; wing length: 5.2 mm.
<emphasis id="B93C3C39D440FFBAFBA1F86DEF10F878" bold="true" box="[1032,1096,1999,2024]" pageId="1" pageNumber="100">Head</emphasis>
(
<figureCitation id="1373FCAED440FFBAFBF6F86DEF8CF878" box="[1119,1236,1999,2024]" captionStart="FIGURES 1 6. 1" captionStartId="4.[151,264,1924,1947]" captionTargetBox="[243,1348,202,1893]" captionTargetId="figure@4.[236,1350,195,1900]" captionTargetPageId="4" captionText="FIGURES 1 6. 1. Pelecocera persiana, male, dorsal view (scale: 2 mm). 2. Pelecocera latifrons, male, dorsal view (scale: 2 mm). 3. Pelecocera persiana, male, lateral view (scale: 2 mm). 4. Pelecocera persiana, male, frontal view (scale: 1 mm). 5. Pelecocera latifrons, male, lateral view (scale: 2 mm). 6. Pelecocera latifrons, male, frontal view (scale: 1 mm)." httpUri="https://zenodo.org/record/243971/files/figure.png" pageId="1" pageNumber="100">Figs 3, 4</figureCitation>
). Face produced anteroventrally, triangular in frontal view, with facial tubercle placed ventrally (tip below ventral margin of eye), yellow-brown medially, darker below antennal insertion point and black along oral margin, with long pale pile on ventral part and on paraface; face densely grey pollinose, very concave between antenna and facial tubercle; gena pale (yellow-brown), black posteriorly, densely grey pollinose with long pale pile; lunule dark brown, shining; frons dark brown, with long pale pile, densely grey pollinose except area around lunule and below vertical triangle, with a depression between anterior ocellus and antenna; vertical triangle raised, broad, scalene, pale pilose; antenna dark brown, basoflagellomere somewhat paler ventrally, dark pilose; ratio scape:pedicel:basoflagellomere = 0.8:1.2:1.7; basoflagellomere broadened dorsally and ventrally, broader than long (
<figureCitation id="1373FCAED443FFB9FBECFE36EFD3FE3C" box="[1093,1163,404,429]" captionStart="FIGURES 1 6. 1" captionStartId="4.[151,264,1924,1947]" captionTargetBox="[243,1348,202,1893]" captionTargetId="figure@4.[236,1350,195,1900]" captionTargetPageId="4" captionText="FIGURES 1 6. 1. Pelecocera persiana, male, dorsal view (scale: 2 mm). 2. Pelecocera latifrons, male, dorsal view (scale: 2 mm). 3. Pelecocera persiana, male, lateral view (scale: 2 mm). 4. Pelecocera persiana, male, frontal view (scale: 1 mm). 5. Pelecocera latifrons, male, lateral view (scale: 2 mm). 6. Pelecocera latifrons, male, frontal view (scale: 1 mm)." httpUri="https://zenodo.org/record/243971/files/figure.png" pageId="2" pageNumber="101">Fig. 4</figureCitation>
), with broad arista with three clearly-defined segments; basoflagellomere and arista microtrichose; eye bare, dichoptic, distance between eyes (3.2 mm at lunule level) broader than eye width; occiput black, grey-silver pollinose, yellowish-white pilose.
<emphasis id="B93C3C39D443FFB9FF3EFE5DEBA8FD88" bold="true" box="[151,240,511,536]" pageId="2" pageNumber="101">Thorax</emphasis>
(
<figureCitation id="1373FCAED443FFB9FEABFE5DEA28FD88" box="[258,368,511,536]" captionStart="FIGURES 1 6. 1" captionStartId="4.[151,264,1924,1947]" captionTargetBox="[243,1348,202,1893]" captionTargetId="figure@4.[236,1350,195,1900]" captionTargetPageId="4" captionText="FIGURES 1 6. 1. Pelecocera persiana, male, dorsal view (scale: 2 mm). 2. Pelecocera latifrons, male, dorsal view (scale: 2 mm). 3. Pelecocera persiana, male, lateral view (scale: 2 mm). 4. Pelecocera persiana, male, frontal view (scale: 1 mm). 5. Pelecocera latifrons, male, lateral view (scale: 2 mm). 6. Pelecocera latifrons, male, frontal view (scale: 1 mm)." httpUri="https://zenodo.org/record/243971/files/figure.png" pageId="2" pageNumber="101">Figs 1, 3</figureCitation>
). Scutum black, densely grey pollinose except medially, with long pale pile; postpronotum yellow, pilose; notopleuron black, pale pilose; scutellum black with yellow macula on posterior margin, yellow pilose, densely grey pollinose, with pale subscutellar fringe. Pleuron mostly dark brown to black, grey pollinose, with long pale pile on proepimeron, posterior anepisternum, dorsal and ventral katepisternum, and anterior anepimeron; dorsal and ventral pile patches on katepisternum broadly separated; metaepisternum and metasternum bare; calypter yellow, yellow pilose; plumula yellow; halter yellow, capitulum slightly darker; posterior spiracular fringes yellow.
<emphasis id="B93C3C39D443FFB9FEEAFD75EAD2FD60" bold="true" box="[323,394,727,752]" pageId="2" pageNumber="101">Wing.</emphasis>
Wing membrane hyaline, stigma brownish; entirely microtrichose. Alula present, as broad as costal cell or a bit narrower, microtrichose. Cell r1 open, vein R4+5 straight, and crossvein r-m basal to middle of cell dm.
<emphasis id="B93C3C39D443FFB9FF62FC80EA52FCAB" bold="true" box="[203,266,802,827]" pageId="2" pageNumber="101">Legs.</emphasis>
Coxae black, grey pollinose; trochanters paler, yellowish; femora black, yellow basally and apically; tibiae yellow with a broad black ring; tarsi yellow; mostly yellow pilose.
<emphasis id="B93C3C39D443FFB9FB8BFCE4EFCEFCCF" bold="true" box="[1058,1174,838,863]" pageId="2" pageNumber="101">Abdomen</emphasis>
(
<figureCitation id="1373FCAED443FFB9FB07FCE4EFAEFCCF" box="[1198,1270,838,863]" captionStart="FIGURES 1 6. 1" captionStartId="4.[151,264,1924,1947]" captionTargetBox="[243,1348,202,1893]" captionTargetId="figure@4.[236,1350,195,1900]" captionTargetPageId="4" captionText="FIGURES 1 6. 1. Pelecocera persiana, male, dorsal view (scale: 2 mm). 2. Pelecocera latifrons, male, dorsal view (scale: 2 mm). 3. Pelecocera persiana, male, lateral view (scale: 2 mm). 4. Pelecocera persiana, male, frontal view (scale: 1 mm). 5. Pelecocera latifrons, male, lateral view (scale: 2 mm). 6. Pelecocera latifrons, male, frontal view (scale: 1 mm)." httpUri="https://zenodo.org/record/243971/files/figure.png" pageId="2" pageNumber="101">Fig. 1</figureCitation>
). Broad, flat, unmargined, grey pollinose, with long pale pile. Tergum 1 black; tergum 2 black with two broad, elongated, yellow maculae not reaching lateral margins, except on anterolateral margin; terga 3 and 4 black with two broad, elongate, yellow maculae not reaching lateral margins, but reaching anterior margin of the tergum; sterna pale, yellow pilose. Genitalia as in
<figureCitation id="1373FCAED443FFB9FE96FC74EAD7FC7F" box="[319,399,982,1007]" captionStart="FIGURES 16 23. 16" captionStartId="6.[151,264,1871,1894]" captionTargetBox="[257,1335,197,1846]" captionTargetId="figure@6.[250,1340,193,1849]" captionTargetPageId="6" captionText="FIGURES 16 23. 16. Pelecocera pergandei, male, lateral view (scale: 2 mm). 17. Pelecocera latifrons, lectotype male, frontal view (scale: 1 mm). 18. Pelecocera pergandei, male, dorsal view (scale: 2 mm). 19. Pelecocera latifrons, lectotype male, dorsal view (scale: 2 mm). 20. Pelecocera latifrons, lectotype male, lateral view (scale: 2 mm). 21. Pelecocera latifrons, lectotype male, labels. 22. Pelecocera latifrons, lectotype male, lateral view male genitalia (scale: 0.5 mm). 23. Pelecocera persiana, male, lateral view male genitalia (scale: 0.5 mm)." httpUri="https://zenodo.org/record/243973/files/figure.png" pageId="2" pageNumber="101">Fig. 23</figureCitation>
(right surstylus is broken and left surstylus is missing).
<emphasis id="B93C3C39D443FFB9FBABFC74EF38FC7F" bold="true" box="[1026,1120,982,1007]" pageId="2" pageNumber="101">Female.</emphasis>
See
<bibRefCitation id="EFD99DDAD443FFB9FB3EFC7AEE3DFC7F" author="Kuznetzov" box="[1175,1381,983,1007]" pageId="2" pageNumber="101" refString="Kuznetzov, S. Y. (1989) Hover-flies of the genus Pelecocera Mg. (Diptera, Syrphidae). Latvian Entomology, 32, 80 - 85." type="journal article" year="1989">Kuznetzov (1989)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C352B3A0D443FFB9FF6EFC58E83CFB83" box="[199,868,1018,1043]" pageId="2" pageNumber="101" type="distribution">
<paragraph id="8BF7E02BD443FFB9FF6EFC58E83CFB83" blockId="2.[151,1437,151,1151]" box="[199,868,1018,1043]" pageId="2" pageNumber="101">
<emphasis id="B93C3C39D443FFB9FF6EFC58EA38FB83" bold="true" box="[199,352,1018,1043]" pageId="2" pageNumber="101">Distribution.</emphasis>
<collectingCountry id="F35FA0BBD443FFB9FECEFC59EACCFB82" box="[359,404,1019,1042]" name="Iran" pageId="2" pageNumber="101">Iran</collectingCountry>
(East
<collectingCountry id="F35FA0BBD443FFB9FE72FC58E90EFB83" box="[475,598,1018,1043]" name="Azerbaijan" pageId="2" pageNumber="101">Azerbaijan</collectingCountry>
and Qazvin Provinces).
</paragraph>
</subSubSection>
<subSubSection id="C352B3A0D443FFB9FF6EFBBCE97AFBCA" pageId="2" pageNumber="101" type="materials_examined">
<paragraph id="8BF7E02BD443FFB9FF6EFBBCE97AFBCA" blockId="2.[151,1437,151,1151]" pageId="2" pageNumber="101">
<emphasis id="B93C3C39D443FFB9FF6EFBBCEAE8FBA7" bold="true" box="[199,432,1054,1079]" pageId="2" pageNumber="101">Material examined.</emphasis>
<collectingCountry id="F35FA0BBD443FFB9FE1EFBBDE958FBA7" box="[439,512,1055,1079]" name="Iran" pageId="2" pageNumber="101">IRAN</collectingCountry>
: Qazvin Province, Zereshk road,
<quantity id="4CB04DCED443FFB9FCDFFBBDE895FBA7" box="[886,973,1055,1079]" metricMagnitude="3" metricUnit="m" metricValue="1.9969999999999999" pageId="2" pageNumber="101" unit="m" value="1997.0">1997 m</quantity>
, 36˚25'10.9''N 50˚06'10.3''E,
<date id="FFF6C6EBD443FFB9FABEFBBCEEC0FBA7" box="[1303,1432,1054,1079]" pageId="2" pageNumber="101" value="2011-07-06">06.vii.2011</date>
. Leg.: M. Kheirandish [
<specimenCount id="9D4E2BA2D443FFB9FE3EFBE1EAE5FBCB" box="[407,445,1091,1115]" pageId="2" pageNumber="101" type="generic">1♂</specimenCount>
,
<collectionCode id="ED5978EED443FFB9FE60FBE1E94FFBCA" box="[457,535,1091,1114]" pageId="2" pageNumber="101">TMUI</collectionCode>
].
</paragraph>
</subSubSection>
<subSubSection id="C352B3A0D443FFB9FF6EFBC4EE4EFBEF" box="[199,1302,1126,1151]" pageId="2" pageNumber="101" type="discussion">
<paragraph id="8BF7E02BD443FFB9FF6EFBC4EE4EFBEF" blockId="2.[151,1437,151,1151]" box="[199,1302,1126,1151]" pageId="2" pageNumber="101">
<emphasis id="B93C3C39D443FFB9FF6EFBC4EA63FBEF" bold="true" box="[199,315,1126,1151]" pageId="2" pageNumber="101">Remarks.</emphasis>
This male specimen was collected approximately
<quantity id="4CB04DCED443FFB9FCD9FBC4E89DFBEF" box="[880,965,1126,1151]" metricMagnitude="5" metricUnit="m" metricValue="4.5" pageId="2" pageNumber="101" unit="km" value="450.0">450 km</quantity>
southeast of the type-locality.
</paragraph>
</subSubSection>
</treatment>
</document>