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<document id="B7E6A0D3E092E9ADD075BB5A2F7B3B90" ID-CLB-Dataset="298461" ID-DOI="10.11646/zootaxa.5463.1.7" ID-GBIF-Dataset="5437a261-1c86-45c2-a113-83c4fcc30ef9" ID-ISSN="1175-5326" ID-Zenodo-Dep="11610306" ID-ZooBank="7C3B419A-C6BB-497F-A43E-A260CCB355B2" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="carolina" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.tables_approvedBy="carolina" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1718181091375" checkinUser="plazi" docAuthor="Provenzano-Rizzi, Francisco, Barriga-Salazar, Ramiro &amp; Stewart, Donald J." docDate="2024" docId="C27487A0FF8EFFF0B6AE644A6DECDCEB" docLanguage="en" docName="zootaxa.5463.1.7.pdf" docOrigin="Zootaxa 5463 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5463.1.7" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Panaqolus orcesi Provenzano-Rizzi &amp; Barriga-Salazar &amp; Stewart 2024, new species" docType="treatment" docVersion="7" lastPageNumber="120" masterDocId="3E4DFFD8FF8CFFF8B639652C6F71DD1F" masterDocTitle="Two new species of the armored catfish genus Panaqolus (Siluriformes, Loricariidae) from the Ecuadorian Amazon" masterLastPageNumber="126" masterPageNumber="112" pageNumber="114" updateTime="1718321359055" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="E3914F82C0BF6B251A4608BF2FC780A6">Two new species of the armored catfish genus Panaqolus (Siluriformes, Loricariidae) from the Ecuadorian Amazon</mods:title>
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<mods:namePart id="8023BE97D57BB818A9412007F4F8070A">Provenzano-Rizzi, Francisco</mods:namePart>
<mods:affiliation id="3BB2A0FA1E4168F065692787668D504C">Centro MBUCV, Instituto de Zoología y Ecología Tropical, Facultad de Ciencias, UCV, Caracas, Venezuela</mods:affiliation>
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<mods:namePart id="BEC528CF2A19AD2372F4121A36D89DB1">Barriga-Salazar, Ramiro</mods:namePart>
<mods:affiliation id="19408977EFA9BB7C08DA69C43E0C5580">Departamento de Biología, Facultad de Ciencias, Escuela Politécnica Nacional, Quito, Ecuador</mods:affiliation>
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</mods:role>
<mods:namePart id="886929CEB5C836D459EFFC383A6DE2D4">Stewart, Donald J.</mods:namePart>
<mods:affiliation id="A97ACA0759344963660E68B09D951464">Department of Environmental Biology, SUNY College of Environmental Science and Forestry, Syracuse, New York, 13210, USA</mods:affiliation>
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<mods:title id="5D4135F26286286B4DAC33A96F50955B">Zootaxa</mods:title>
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<mods:date id="34394F8E46EE957ACA5DF8744A19215E">2024</mods:date>
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<mods:number id="429A5875F8E0784943AA7FF460347507">2024-06-04</mods:number>
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<mods:number id="DE6361C7E3F805739DEBB3A1EDA5B3FA">5463</mods:number>
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<mods:url id="43A277B8373D8A105DCDFE8B19E9CEB0">http://dx.doi.org/10.11646/zootaxa.5463.1.7</mods:url>
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<mods:classification id="CEC13AB9250718F7C48892551CE57C9C">journal article</mods:classification>
<mods:identifier id="EA3F86A79375D1EA9095564303E0BFA4" type="CLB-Dataset">298461</mods:identifier>
<mods:identifier id="FAA31BE574F0A81F97A92935CEFB29B7" type="DOI">10.11646/zootaxa.5463.1.7</mods:identifier>
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<treatment id="C27487A0FF8EFFF0B6AE644A6DECDCEB" ID-DOI="http://doi.org/10.5281/zenodo.11645869" ID-GBIF-Taxon="232750432" ID-Zenodo-Dep="11645869" LSID="urn:lsid:plazi:treatment:C27487A0FF8EFFF0B6AE644A6DECDCEB" httpUri="http://treatment.plazi.org/id/C27487A0FF8EFFF0B6AE644A6DECDCEB" lastPageId="8" lastPageNumber="120" pageId="2" pageNumber="114">
<subSubSection id="02C7653DFF8EFFFAB6AE644A6CA6DCBA" pageId="2" pageNumber="114" type="nomenclature">
<paragraph id="4A6236B6FF8EFFFAB6AE644A6CA6DCBA" blockId="2.[151,983,358,457]" pageId="2" pageNumber="114">
<heading id="112A81DAFF8EFFFAB6AE644A6D74DC9F" bold="true" box="[151,517,358,385]" fontSize="11" level="1" pageId="2" pageNumber="114" reason="1">
<taxonomicName id="8DDD4D35FF8EFFFAB6AE644A6E16DC9E" authority="Provenzano-Rizzi &amp; Barriga-Salazar &amp; Stewart, 2024" authorityName="Provenzano-Rizzi &amp; Barriga-Salazar &amp; Stewart" authorityYear="2024" box="[151,359,358,385]" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="114" phylum="Chordata" rank="species" species="orcesi" status="sp. nov.">Panaqolus orcesi</taxonomicName>
,
<taxonomicNameLabel id="639A57DFFF8EFFFAB74F644B6D74DC9F" box="[374,517,358,385]" pageId="2" pageNumber="114" rank="species">new species</taxonomicNameLabel>
</heading>
<uri id="3E4C3AB4FF8EFFFAB6AE64A66CA6DCBA" box="[151,983,394,421]" pageId="2" pageNumber="114">
urn:lsid:zoobank.org:act:
<uuid id="3E7B0C63FF8EFFFAB79564A66CA6DCBA" box="[428,983,394,421]" pageId="2" pageNumber="114">C6918B6A-5879-45B7-8DC1-AB0AA772A4FB</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="02C7653DFF8EFFFAB6AE64836EB2DCD6" box="[151,451,431,457]" pageId="2" pageNumber="114" type="description">
<paragraph id="4A6236B6FF8EFFFAB6AE64836EB2DCD6" blockId="2.[151,983,358,457]" box="[151,451,431,457]" pageId="2" pageNumber="114">
(
<figureCitation id="D2E62A33FF8EFFFAB6A664836F9CDCD6" box="[159,237,431,457]" captionStart="FIGURE 1" captionStartId="4.[152,255,1961,1985]" captionTargetBox="[202,1386,179,1935]" captionTargetId="figure-17@4.[202,1386,179,1935]" captionTargetPageId="4" captionText="FIGURE 1. Panaqolus orcesi, MEPN 19492, 180.6 mm SL, holotype. Scale bar = 1 cm. Photos by F.P-R." figureDoi="http://doi.org/10.5281/zenodo.11610308" httpUri="https://zenodo.org/record/11610308/files/figure.png" pageId="2" pageNumber="114">Figs. 1</figureCitation>
,
<figureCitation id="D2E62A33FF8EFFFAB6C264836E67DCD6" box="[251,278,431,457]" captionStart="FIGURE 2" captionStartId="5.[152,255,743,767]" captionTargetBox="[164,1423,181,719]" captionTargetId="figure-19@5.[164,1423,181,719]" captionTargetPageId="5" captionText="FIGURE 2. Close up of mouth and buccal disk for: a) Panaqolus orcesi, MEPN 19492, 180.6 mm SL, holotype; and b) P. pantostiktos, MEPN 19495, 163.4 mm SL, holotype. Photos by F.P-R." figureDoi="http://doi.org/10.5281/zenodo.11610310" httpUri="https://zenodo.org/record/11610310/files/figure.png" pageId="2" pageNumber="114">2a</figureCitation>
and
<figureCitation id="D2E62A33FF8EFFFAB77564836E18DCD6" box="[332,361,431,457]" captionStart="FIGURE 3" captionStartId="5.[152,255,1968,1992]" captionTargetBox="[164,1423,964,1945]" captionTargetId="figure-85@5.[164,1423,964,1945]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal view of color patterns in specimens soon after capture for: a) Panaqolus orcesi, with cream-colored dots, and b) Panaqolus pantostiktos, with yellow- or gold-colored dots. Photos by R.B-S." figureDoi="http://doi.org/10.5281/zenodo.11610312" httpUri="https://zenodo.org/record/11610312/files/figure.png" pageId="2" pageNumber="114">3a</figureCitation>
;
<tableCitation id="075F030DFF8EFFFAB74D64836EC9DCD6" box="[372,440,431,457]" captionStart="TABLE 1" captionStartId="2.[152,244,828,852]" captionTargetPageId="2" captionText="TABLE 1. Morphometric data of Panaqolus orcesi, expressed as percent of SL or percent of HL." httpUri="http://table.plazi.org/id/1EA2663EFF8EFFFAB6A166106BDBDE4B" pageId="2" pageNumber="114" tableUuid="1EA2663EFF8EFFFAB6A166106BDBDE4B">Tab. 1</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="02C7653DFF8EFFFAB6AE64DB6D7FDE12" pageId="2" pageNumber="114" type="materials_examined">
<paragraph id="4A6236B6FF8EFFFAB6AE64DB6EAFDF46" blockId="2.[151,1437,503,781]" pageId="2" pageNumber="114">
<materialsCitation id="FAB53CEBFF8EFFFAB6AE64DB6EAFDF46" ID-GBIF-Occurrence="4876552304" collectingDate="2006-06-26" collectionCode="MEPN-I" collectorName="R. Barriga" county="Ecuadorian Amazon" latitude="-0.83416665" location="Napo River basin" longLatPrecision="21" longitude="-75.56612" municipality="River" pageId="2" pageNumber="114" specimenCount="1" stateProvince="Orellana Province" typeStatus="holotype">
<emphasis id="78A9EAA4FF8EFFFAB6AE64DB6E7ADF0E" bold="true" box="[151,267,503,529]" pageId="2" pageNumber="114">
<typeStatus id="95668814FF8EFFFAB6AE64DB6E77DF0E" box="[151,262,503,529]" pageId="2" pageNumber="114" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="2CCCAE73FF8EFFFAB72B64DB6E03DF0E" box="[274,370,503,529]" pageId="2" pageNumber="114">MEPN-I</collectionCode>
<quantity id="8D259B53FF8EFFFAB74064DB6D4FDF0E" box="[377,574,503,529]" metricMagnitude="4" metricUnit="m" metricValue="1.94921806" pageId="2" pageNumber="114" unit="mm" value="1.94921806E7">19492, 180.6 mm</quantity>
SL,
<collectingCounty id="A3034E3AFF8EFFFAB44B64DB6C29DF0E" box="[626,856,503,529]" pageId="2" pageNumber="114">Ecuadorian Amazon</collectingCounty>
,
<collectingRegion id="8819F854FF8EFFFAB55A64DB6B41DF0E" box="[867,1072,503,529]" pageId="2" pageNumber="114">Orellana Province</collectingRegion>
,
<location id="4F02606DFF8EFFFAB20264DB6A70DF0E" LSID="urn:lsid:plazi:treatment:C27487A0FF8EFFF0B6AE644A6DECDCEB:4F02606DFF8EFFFAB20264DB6A70DF0E" box="[1083,1281,503,529]" county="Ecuadorian Amazon" latitude="-0.83416665" longLatPrecision="21" longitude="-75.56612" municipality="River" name="Napo River basin" pageId="2" pageNumber="114" stateProvince="Orellana Province">Napo River basin</location>
,
<location id="4F02606DFF8EFFFAB33264DB6F86DF2A" LSID="urn:lsid:plazi:treatment:C27487A0FF8EFFF0B6AE644A6DECDCEB:4F02606DFF8EFFFAB33264DB6F86DF2A" county="Ecuadorian Amazon" latitude="-0.83416665" longLatPrecision="21" longitude="-75.56612" municipality="River" name="Quebrada La Cascada" pageId="2" pageNumber="114" stateProvince="Orellana Province">Quebrada La Cascada</location>
,
<location id="4F02606DFF8EFFFAB73A67376ED3DF2A" LSID="urn:lsid:plazi:treatment:C27487A0FF8EFFF0B6AE644A6DECDCEB:4F02606DFF8EFFFAB73A67376ED3DF2A" box="[259,418,539,565]" county="Ecuadorian Amazon" latitude="-0.83416665" longLatPrecision="21" longitude="-75.56612" municipality="River" name="Tiputini River" pageId="2" pageNumber="114" stateProvince="Orellana Province">Tiputini River</location>
tributary, approx.
<geoCoordinate id="2FE95071FF8EFFFAB44C67366C72DF2A" box="[629,771,538,565]" degrees="00" direction="south" minutes="50" orientation="latitude" pageId="2" pageNumber="114" precision="15" seconds="03" value="-0.83416665">00°50’03” S</geoCoordinate>
,
<geoCoordinate id="2FE95071FF8EFFFAB52867366CD9DF2A" box="[785,936,538,565]" degrees="75" direction="west" minutes="33" orientation="longitude" pageId="2" pageNumber="114" precision="15" seconds="58" value="-75.56612">75°33’58” W</geoCoordinate>
, 190 meters above sea level (masl),
<date id="3E631076FF8EFFFAB37C67376FA3DF46" pageId="2" pageNumber="114" value="2006-06-26">
<collectingDate id="2E27E99EFF8EFFFAB37C67376FA3DF46" pageId="2" pageNumber="114" value="2006-06-26">26 June 2006</collectingDate>
</date>
,
<collectorName id="E7285360FF8EFFFAB6E467136E24DF46" box="[221,341,575,601]" pageId="2" pageNumber="114">R. Barriga</collectorName>
, RBS06-12.
</materialsCitation>
</paragraph>
<paragraph id="4A6236B6FF8EFFFAB6FE674F6D7FDE12" blockId="2.[151,1437,503,781]" pageId="2" pageNumber="114">
<materialsCitation id="FAB53CEBFF8EFFFAB6FE674F6C91DFDA" ID-GBIF-Occurrence="4876552302" collectingDate="2005-04-20" collectingDateMax="2005-04-22" collectingDateMin="2005-04-20" collectionCode="MEPN-I" collectorName="R. Barriga &amp; T. Caiza &amp; J. Nenhimo" county="Ecuadorian Amazon" elevation="100" latitude="-0.62222224" location="Rio Mengo Garita" longLatPrecision="21" longitude="-76.50694" municipality="River" pageId="2" pageNumber="114" specimenCount="2" stateProvince="Orellana Province, Napo" typeStatus="paratype">
<emphasis id="78A9EAA4FF8EFFFAB6FE674F6E37DF62" bold="true" box="[199,326,611,637]" pageId="2" pageNumber="114">
<typeStatus id="95668814FF8EFFFAB6FE674F6E30DF62" box="[199,321,611,637]" pageId="2" pageNumber="114" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="5CDBFD3FFF8EFFFAB774674F6E8DDF62" box="[333,508,611,637]" count="2" pageId="2" pageNumber="114" type="generic">Two specimens</specimenCount>
from
<collectingCounty id="A3034E3AFF8EFFFAB47B674F6C5BDF62" box="[578,810,611,637]" pageId="2" pageNumber="114">Ecuadorian Amazon</collectingCounty>
,
<collectingRegion id="8819F854FF8EFFFAB50F674F6B3FDF62" box="[822,1102,611,637]" isEnumeration="true" pageId="2" pageNumber="114">Orellana Province, Napo</collectingRegion>
River basin:
<collectionCode id="2CCCAE73FF8EFFFAB2DE674F6A36DF62" box="[1255,1351,611,637]" pageId="2" pageNumber="114">MEPN-I</collectionCode>
19493, 170.9 mm SL,
<location id="4F02606DFF8EFFFAB70267AB6D79DFBE" LSID="urn:lsid:plazi:treatment:C27487A0FF8EFFF0B6AE644A6DECDCEB:4F02606DFF8EFFFAB70267AB6D79DFBE" box="[315,520,647,673]" county="Ecuadorian Amazon" latitude="-0.62222224" longLatPrecision="21" longitude="-76.50694" municipality="River" name="Rio Mengo Garita" pageId="2" pageNumber="114" stateProvince="Orellana Province, Napo">Rio Mengo Garita</location>
, 100 m upstream from bridge on
<location id="4F02606DFF8EFFFAB54D67AB6B76DFBE" LSID="urn:lsid:plazi:treatment:C27487A0FF8EFFF0B6AE644A6DECDCEB:4F02606DFF8EFFFAB54D67AB6B76DFBE" box="[884,1031,647,673]" county="Ecuadorian Amazon" latitude="-0.62222224" longLatPrecision="21" longitude="-76.50694" municipality="River" name="Yasuni River" pageId="2" pageNumber="114" stateProvince="Orellana Province, Napo">Yasuni River</location>
, approx.
<geoCoordinate id="2FE95071FF8EFFFAB25567AA6B86DFBE" box="[1132,1271,646,673]" degrees="00" direction="south" minutes="50" orientation="latitude" pageId="2" pageNumber="114" precision="15" seconds="03" value="-0.83416665">00°50’03” S</geoCoordinate>
,
<geoCoordinate id="2FE95071FF8EFFFAB33A67AB6AE9DFBE" box="[1283,1432,647,673]" degrees="76" direction="west" minutes="20" orientation="longitude" pageId="2" pageNumber="114" precision="15" seconds="42" value="-76.345">76°20’42” W</geoCoordinate>
, 250 masl,
<date id="3E631076FF8EFFFAB73067876EBEDFDA" box="[265,463,682,709]" pageId="2" pageNumber="114" value="2005-04-20" valueMax="2005-04-22" valueMin="2005-04-20">
<collectingDate id="2E27E99EFF8EFFFAB73067876EBEDFDA" box="[265,463,682,709]" pageId="2" pageNumber="114" value="2005-04-20" valueMax="2005-04-22" valueMin="2005-04-20">20–22 April 2005</collectingDate>
</date>
,
<collectorName id="E7285360FF8EFFFAB7E067876D3EDFDA" box="[473,591,683,709]" pageId="2" pageNumber="114">R. Barriga</collectorName>
,
<collectorName id="E7285360FF8EFFFAB46067876DC4DFDA" box="[601,693,683,709]" pageId="2" pageNumber="114">T. Caiza</collectorName>
&amp;
<collectorName id="E7285360FF8EFFFAB4EF67876C28DFDA" box="[726,857,683,709]" pageId="2" pageNumber="114">J. Nenhimo</collectorName>
, RBS05-08.
</materialsCitation>
<materialsCitation id="FAB53CEBFF8EFFFAB5DF67876D7BDE12" ID-GBIF-Occurrence="4876552306" collectingDate="2007-06-17" collectingDateMax="2007-06-17" collectingDateMin="2005-04-20" collectionCode="MEPN-I" collectorName="R. Barriga &amp; J. Silva" county="Ecuadorian Amazon" elevation="236" latitude="-0.62222224" location="Tiputini River" longLatPrecision="21" longitude="-76.50694" municipality="River" pageId="2" pageNumber="114" specimenCount="1" stateProvince="Orellana Province, Napo" typeStatus="paratype">
<collectionCode id="2CCCAE73FF8EFFFAB5DF67876B37DFDA" box="[998,1094,683,709]" pageId="2" pageNumber="114">MEPN-I</collectionCode>
19494, 136.5 mm SL;
<location id="4F02606DFF8EFFFAB37A67876FA6DFF6" LSID="urn:lsid:plazi:treatment:C27487A0FF8EFFF0B6AE644A6DECDCEB:4F02606DFF8EFFFAB37A67876FA6DFF6" county="Ecuadorian Amazon" latitude="-0.62222224" longLatPrecision="21" longitude="-76.50694" municipality="River" name="Tiputini River" pageId="2" pageNumber="114" stateProvince="Orellana Province, Napo">Tiputini River</location>
,
<locationDeviation id="5BD02F94FF8EFFFAB6DB67E36D98DFF6" box="[226,745,719,745]" location="Campamento Politecnica" pageId="2" pageNumber="114">1 km upstream from Campamento Politécnica</locationDeviation>
, approx.
<geoCoordinate id="2FE95071FF8EFFFAB56967E36CACDFF6" box="[848,989,719,745]" degrees="00" direction="south" minutes="37" orientation="latitude" pageId="2" pageNumber="114" precision="15" seconds="20" value="-0.62222224">00°37´20” S</geoCoordinate>
,
<geoCoordinate id="2FE95071FF8EFFFAB5D367E26BF0DFF6" box="[1002,1153,718,745]" degrees="76" direction="west" minutes="30" orientation="longitude" pageId="2" pageNumber="114" precision="15" seconds="25" value="-76.50694">76°30´25” W</geoCoordinate>
, 236 masl,
<date id="3E631076FF8EFFFAB33967E36AE9DFF6" box="[1280,1432,719,745]" pageId="2" pageNumber="114" value="2007-06-17">
<collectingDate id="2E27E99EFF8EFFFAB33967E36AE9DFF6" box="[1280,1432,719,745]" pageId="2" pageNumber="114" value="2007-06-17">17 June 2007</collectingDate>
</date>
,
<collectorName id="E7285360FF8EFFFAB6AE67DF6E7DDE12" box="[151,268,755,781]" pageId="2" pageNumber="114">R. Barriga</collectorName>
&amp;
<collectorName id="E7285360FF8EFFFAB70967DF6EF4DE12" box="[304,389,755,781]" pageId="2" pageNumber="114">J. Silva</collectorName>
, RBS07-15
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="1EA2663EFF8EFFFAB6A166106BDBDE4B" ID-Table-UUID="1EA2663EFF8EFFFAB6A166106BDBDE4B" box="[152,1194,828,853]" httpUri="http://table.plazi.org/id/1EA2663EFF8EFFFAB6A166106BDBDE4B" pageId="2" pageNumber="114" startId="2.[152,244,828,852]" targetBox="[159,1378,870,2008]" targetIsTable="true" targetPageId="2" targetType="table">
<paragraph id="4A6236B6FF8EFFFAB6A166106BDBDE4B" blockId="2.[152,1194,828,853]" box="[152,1194,828,853]" pageId="2" pageNumber="114">
<emphasis id="78A9EAA4FF8EFFFAB6A166106E7EDE4B" bold="true" box="[152,271,828,852]" pageId="2" pageNumber="114">TABLE 1.</emphasis>
Morphometric data of
<taxonomicName id="8DDD4D35FF8EFFFAB43F66106DCEDE4B" authorityName="Provenzano-Rizzi &amp; Barriga-Salazar &amp; Stewart" authorityYear="2024" box="[518,703,828,852]" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="114" phylum="Chordata" rank="species" species="orcesi">
<emphasis id="78A9EAA4FF8EFFFAB43F66106DCEDE4B" box="[518,703,828,852]" italics="true" pageId="2" pageNumber="114">Panaqolus orcesi</emphasis>
</taxonomicName>
, expressed as percent of SL or percent of HL.
</paragraph>
</caption>
<paragraph id="4A6236B6FF8EFFFAB546664A6BFBDAC8" pageId="2" pageNumber="114">
<table id="38DDC416FF8E0007B6A6664A6A13DAC7" box="[159,1378,870,2008]" gridcols="5" gridrows="28" pageId="2" pageNumber="114">
<tr id="F4ED34F4FF8E0007B6A6664A6A13DE64" box="[159,1378,870,891]" gridrow="0" pageId="2" pageNumber="114">
<th id="B73C5D88FF8E0007B6A6664A6A13DE64" box="[159,1378,870,891]" colspan="5" colspanRight="4" gridcol="0" gridrow="0" pageId="2" pageNumber="114">
<taxonomicName id="8DDD4D35FF8EFFFAB546664A6B54DE64" authority="MEPN" authorityName="MEPN" box="[895,1061,870,891]" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="114" phylum="Chordata" rank="species" species="orcesi">
<emphasis id="78A9EAA4FF8EFFFAB546664A6B54DE64" box="[895,1061,870,891]" italics="true" pageId="2" pageNumber="114">Panaqolus orcesi</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="F4ED34F4FF8E0007B6A666A16A13DEBD" box="[159,1378,909,930]" gridrow="1" pageId="2" pageNumber="114" rowspan-0="1" rowspan-4="1">
<td id="B73C5D88FF8E0007B47A66A16DB9DEBD" box="[579,712,909,930]" gridcol="1" gridrow="1" pageId="2" pageNumber="114">MEPN 19492</td>
<td id="B73C5D88FF8E0007B50E66A16CCDDEBD" box="[823,956,909,930]" gridcol="2" gridrow="1" pageId="2" pageNumber="114">MEPN 19493</td>
<td id="B73C5D88FF8E0007B21866A16BD7DEBD" box="[1057,1190,909,930]" gridcol="3" gridrow="1" pageId="2" pageNumber="114">MEPN 19494</td>
</tr>
<tr id="F4ED34F4FF8E0007B6A666986A13DED5" box="[159,1378,948,970]" gridrow="2" pageId="2" pageNumber="114">
<th id="B73C5D88FF8E0007B6A666986EB9DED5" box="[159,456,948,970]" gridcol="0" gridrow="2" pageId="2" pageNumber="114">Character</th>
<td id="B73C5D88FF8E0007B47A66986DB9DED5" box="[579,712,948,970]" gridcol="1" gridrow="2" pageId="2" pageNumber="114">Holotype</td>
<td id="B73C5D88FF8E0007B50E66986CCDDED5" box="[823,956,948,970]" gridcol="2" gridrow="2" pageId="2" pageNumber="114">Paratype</td>
<td id="B73C5D88FF8E0007B21866986BD7DED5" box="[1057,1190,948,970]" gridcol="3" gridrow="2" pageId="2" pageNumber="114">Paratype</td>
<td id="B73C5D88FF8E0007B32866986A13DED5" box="[1297,1378,948,970]" gridcol="4" gridrow="2" pageId="2" pageNumber="114">Mean %</td>
</tr>
<tr id="F4ED34F4FF8E0007B6A666F76A13DEEE" box="[159,1378,987,1009]" gridrow="3" pageId="2" pageNumber="114" rowspan-4="1">
<th id="B73C5D88FF8E0007B6A666F76EB9DEEE" box="[159,456,987,1009]" gridcol="0" gridrow="3" pageId="2" pageNumber="114">SL (mm)</th>
<td id="B73C5D88FF8E0007B47A66F76DB9DEEE" box="[579,712,987,1009]" gridcol="1" gridrow="3" pageId="2" pageNumber="114">180.6</td>
<td id="B73C5D88FF8E0007B50E66F76CCDDEEE" box="[823,956,987,1009]" gridcol="2" gridrow="3" pageId="2" pageNumber="114">170.9</td>
<td id="B73C5D88FF8E0007B21866F76BD7DEEE" box="[1057,1190,987,1009]" gridcol="3" gridrow="3" pageId="2" pageNumber="114">136.5</td>
</tr>
<tr id="F4ED34F4FF8E0007B6A661296A13D904" box="[159,1378,1029,1051]" gridrow="4" pageId="2" pageNumber="114">
<th id="B73C5D88FF8E0007B6A661296A13D904" box="[159,1378,1029,1051]" colspan="5" colspanRight="4" gridcol="0" gridrow="4" pageId="2" pageNumber="114">
<emphasis id="78A9EAA4FF8EFFFAB6A661296E5BD904" bold="true" box="[159,298,1029,1051]" pageId="2" pageNumber="114">Percent of SL</emphasis>
</th>
</tr>
<tr id="F4ED34F4FF8E0007B6A661036A13D95A" box="[159,1378,1071,1093]" gridrow="5" pageId="2" pageNumber="114">
<th id="B73C5D88FF8E0007B6A661036EB9D95A" box="[159,456,1071,1093]" gridcol="0" gridrow="5" pageId="2" pageNumber="114">Head length</th>
<td id="B73C5D88FF8E0007B47A61036DB9D95A" box="[579,712,1071,1093]" gridcol="1" gridrow="5" pageId="2" pageNumber="114">33.2</td>
<td id="B73C5D88FF8E0007B50E61036CCDD95A" box="[823,956,1071,1093]" gridcol="2" gridrow="5" pageId="2" pageNumber="114">33.8</td>
<td id="B73C5D88FF8E0007B21861036BD7D95A" box="[1057,1190,1071,1093]" gridcol="3" gridrow="5" pageId="2" pageNumber="114">34.6</td>
<td id="B73C5D88FF8E0007B32861036A13D95A" box="[1297,1378,1071,1093]" gridcol="4" gridrow="5" pageId="2" pageNumber="114">33.9</td>
</tr>
<tr id="F4ED34F4FF8E0007B6A661746A13D971" box="[159,1378,1112,1134]" gridrow="6" pageId="2" pageNumber="114">
<th id="B73C5D88FF8E0007B6A661746EB9D971" box="[159,456,1112,1134]" gridcol="0" gridrow="6" pageId="2" pageNumber="114">Predorsal length</th>
<td id="B73C5D88FF8E0007B47A61746DB9D971" box="[579,712,1112,1134]" gridcol="1" gridrow="6" pageId="2" pageNumber="114">40.6</td>
<td id="B73C5D88FF8E0007B50E61746CCDD971" box="[823,956,1112,1134]" gridcol="2" gridrow="6" pageId="2" pageNumber="114">41.4</td>
<td id="B73C5D88FF8E0007B21861746BD7D971" box="[1057,1190,1112,1134]" gridcol="3" gridrow="6" pageId="2" pageNumber="114">42.7</td>
<td id="B73C5D88FF8E0007B32861746A13D971" box="[1297,1378,1112,1134]" gridcol="4" gridrow="6" pageId="2" pageNumber="114">41.6</td>
</tr>
<tr id="F4ED34F4FF8E0007B6A661AE6A13D987" box="[159,1378,1154,1176]" gridrow="7" pageId="2" pageNumber="114">
<th id="B73C5D88FF8E0007B6A661AE6EB9D987" box="[159,456,1154,1176]" gridcol="0" gridrow="7" pageId="2" pageNumber="114">Postdorsal length</th>
<td id="B73C5D88FF8E0007B47A61AE6DB9D987" box="[579,712,1154,1176]" gridcol="1" gridrow="7" pageId="2" pageNumber="114">30.9</td>
<td id="B73C5D88FF8E0007B50E61AE6CCDD987" box="[823,956,1154,1176]" gridcol="2" gridrow="7" pageId="2" pageNumber="114">31.6</td>
<td id="B73C5D88FF8E0007B21861AE6BD7D987" box="[1057,1190,1154,1176]" gridcol="3" gridrow="7" pageId="2" pageNumber="114">32.1</td>
<td id="B73C5D88FF8E0007B32861AE6A13D987" box="[1297,1378,1154,1176]" gridcol="4" gridrow="7" pageId="2" pageNumber="114">31.6</td>
</tr>
<tr id="F4ED34F4FF8E0007B6A661806A13D9DD" box="[159,1378,1196,1218]" gridrow="8" pageId="2" pageNumber="114">
<th id="B73C5D88FF8E0007B6A661806EB9D9DD" box="[159,456,1196,1218]" gridcol="0" gridrow="8" pageId="2" pageNumber="114">Interdorsal length</th>
<td id="B73C5D88FF8E0007B47A61806DB9D9DD" box="[579,712,1196,1218]" gridcol="1" gridrow="8" pageId="2" pageNumber="114">12.1</td>
<td id="B73C5D88FF8E0007B50E61806CCDD9DD" box="[823,956,1196,1218]" gridcol="2" gridrow="8" pageId="2" pageNumber="114">12.4</td>
<td id="B73C5D88FF8E0007B21861806BD7D9DD" box="[1057,1190,1196,1218]" gridcol="3" gridrow="8" pageId="2" pageNumber="114">12.7</td>
<td id="B73C5D88FF8E0007B32861806A13D9DD" box="[1297,1378,1196,1218]" gridcol="4" gridrow="8" pageId="2" pageNumber="114">12.4</td>
</tr>
<tr id="F4ED34F4FF8E0007B6A661F96A13D9F4" box="[159,1378,1237,1259]" gridrow="9" pageId="2" pageNumber="114">
<th id="B73C5D88FF8E0007B6A661F96EB9D9F4" box="[159,456,1237,1259]" gridcol="0" gridrow="9" pageId="2" pageNumber="114">Preanal length</th>
<td id="B73C5D88FF8E0007B47A61F96DB9D9F4" box="[579,712,1237,1259]" gridcol="1" gridrow="9" pageId="2" pageNumber="114">69.7</td>
<td id="B73C5D88FF8E0007B50E61F96CCDD9F4" box="[823,956,1237,1259]" gridcol="2" gridrow="9" pageId="2" pageNumber="114">70.2</td>
<td id="B73C5D88FF8E0007B21861F96BD7D9F4" box="[1057,1190,1237,1259]" gridcol="3" gridrow="9" pageId="2" pageNumber="114">70.2</td>
<td id="B73C5D88FF8E0007B32861F96A13D9F4" box="[1297,1378,1237,1259]" gridcol="4" gridrow="9" pageId="2" pageNumber="114">70.0</td>
</tr>
<tr id="F4ED34F4FF8E0007B6A661D36A13D80A" box="[159,1378,1279,1301]" gridrow="10" pageId="2" pageNumber="114">
<th id="B73C5D88FF8E0007B6A661D36EB9D80A" box="[159,456,1279,1301]" gridcol="0" gridrow="10" pageId="2" pageNumber="114">Postanal length</th>
<td id="B73C5D88FF8E0007B47A61D36DB9D80A" box="[579,712,1279,1301]" gridcol="1" gridrow="10" pageId="2" pageNumber="114">28.6</td>
<td id="B73C5D88FF8E0007B50E61D36CCDD80A" box="[823,956,1279,1301]" gridcol="2" gridrow="10" pageId="2" pageNumber="114">27.8</td>
<td id="B73C5D88FF8E0007B21861D36BD7D80A" box="[1057,1190,1279,1301]" gridcol="3" gridrow="10" pageId="2" pageNumber="114">27.4</td>
<td id="B73C5D88FF8E0007B32861D36A13D80A" box="[1297,1378,1279,1301]" gridcol="4" gridrow="10" pageId="2" pageNumber="114">28.0</td>
</tr>
<tr id="F4ED34F4FF8E0007B6A660046A13D821" box="[159,1378,1320,1342]" gridrow="11" pageId="2" pageNumber="114">
<th id="B73C5D88FF8E0007B6A660046EB9D821" box="[159,456,1320,1342]" gridcol="0" gridrow="11" pageId="2" pageNumber="114">Thoracic length</th>
<td id="B73C5D88FF8E0007B47A60046DB9D821" box="[579,712,1320,1342]" gridcol="1" gridrow="11" pageId="2" pageNumber="114">24.2</td>
<td id="B73C5D88FF8E0007B50E60046CCDD821" box="[823,956,1320,1342]" gridcol="2" gridrow="11" pageId="2" pageNumber="114">28.2</td>
<td id="B73C5D88FF8E0007B21860046BD7D821" box="[1057,1190,1320,1342]" gridcol="3" gridrow="11" pageId="2" pageNumber="114">28.6</td>
<td id="B73C5D88FF8E0007B32860046A13D821" box="[1297,1378,1320,1342]" gridcol="4" gridrow="11" pageId="2" pageNumber="114">27.0</td>
</tr>
<tr id="F4ED34F4FF8E0007B6A6607E6A13D877" box="[159,1378,1362,1384]" gridrow="12" pageId="2" pageNumber="114">
<th id="B73C5D88FF8E0007B6A6607E6EB9D877" box="[159,456,1362,1384]" gridcol="0" gridrow="12" pageId="2" pageNumber="114">Abdominal length</th>
<td id="B73C5D88FF8E0007B47A607E6DB9D877" box="[579,712,1362,1384]" gridcol="1" gridrow="12" pageId="2" pageNumber="114">24.1</td>
<td id="B73C5D88FF8E0007B50E607E6CCDD877" box="[823,956,1362,1384]" gridcol="2" gridrow="12" pageId="2" pageNumber="114">25.3</td>
<td id="B73C5D88FF8E0007B218607E6BD7D877" box="[1057,1190,1362,1384]" gridcol="3" gridrow="12" pageId="2" pageNumber="114">23.0</td>
<td id="B73C5D88FF8E0007B328607E6A13D877" box="[1297,1378,1362,1384]" gridcol="4" gridrow="12" pageId="2" pageNumber="114">24.1</td>
</tr>
<tr id="F4ED34F4FF8E0007B6A660576A13D88D" box="[159,1378,1403,1426]" gridrow="13" pageId="2" pageNumber="114">
<th id="B73C5D88FF8E0007B6A660576EB9D88D" box="[159,456,1403,1426]" gridcol="0" gridrow="13" pageId="2" pageNumber="114">Dorsal fin base</th>
<td id="B73C5D88FF8E0007B47A60576DB9D88D" box="[579,712,1403,1426]" gridcol="1" gridrow="13" pageId="2" pageNumber="114">33.1</td>
<td id="B73C5D88FF8E0007B50E60576CCDD88D" box="[823,956,1403,1426]" gridcol="2" gridrow="13" pageId="2" pageNumber="114">33.2</td>
<td id="B73C5D88FF8E0007B21860576BD7D88D" box="[1057,1190,1403,1426]" gridcol="3" gridrow="13" pageId="2" pageNumber="114">35.0</td>
<td id="B73C5D88FF8E0007B32860576A13D88D" box="[1297,1378,1403,1426]" gridcol="4" gridrow="13" pageId="2" pageNumber="114">33.7</td>
</tr>
<tr id="F4ED34F4FF8E0007B6A660896A13D8A4" box="[159,1378,1445,1467]" gridrow="14" pageId="2" pageNumber="114">
<th id="B73C5D88FF8E0007B6A660896EB9D8A4" box="[159,456,1445,1467]" gridcol="0" gridrow="14" pageId="2" pageNumber="114">Dorsal spine length</th>
<td id="B73C5D88FF8E0007B47A60896DB9D8A4" box="[579,712,1445,1467]" gridcol="1" gridrow="14" pageId="2" pageNumber="114">34.0</td>
<td id="B73C5D88FF8E0007B50E60896CCDD8A4" box="[823,956,1445,1467]" gridcol="2" gridrow="14" pageId="2" pageNumber="114">?</td>
<td id="B73C5D88FF8E0007B21860896BD7D8A4" box="[1057,1190,1445,1467]" gridcol="3" gridrow="14" pageId="2" pageNumber="114">34.1</td>
<td id="B73C5D88FF8E0007B32860896A13D8A4" box="[1297,1378,1445,1467]" gridcol="4" gridrow="14" pageId="2" pageNumber="114">34.1</td>
</tr>
<tr id="F4ED34F4FF8E0007B6A660E36A13D8FA" box="[159,1378,1487,1509]" gridrow="15" pageId="2" pageNumber="114">
<th id="B73C5D88FF8E0007B6A660E36EB9D8FA" box="[159,456,1487,1509]" gridcol="0" gridrow="15" pageId="2" pageNumber="114">Pectoral spine length</th>
<td id="B73C5D88FF8E0007B47A60E36DB9D8FA" box="[579,712,1487,1509]" gridcol="1" gridrow="15" pageId="2" pageNumber="114">35.8</td>
<td id="B73C5D88FF8E0007B50E60E36CCDD8FA" box="[823,956,1487,1509]" gridcol="2" gridrow="15" pageId="2" pageNumber="114">37.4</td>
<td id="B73C5D88FF8E0007B21860E36BD7D8FA" box="[1057,1190,1487,1509]" gridcol="3" gridrow="15" pageId="2" pageNumber="114">35.9</td>
<td id="B73C5D88FF8E0007B32860E36A13D8FA" box="[1297,1378,1487,1509]" gridcol="4" gridrow="15" pageId="2" pageNumber="114">36.3</td>
</tr>
<tr id="F4ED34F4FF8E0007B6A660D46A13DB11" box="[159,1378,1528,1550]" gridrow="16" pageId="2" pageNumber="114">
<th id="B73C5D88FF8E0007B6A660D46EB9DB11" box="[159,456,1528,1550]" gridcol="0" gridrow="16" pageId="2" pageNumber="114">Pelvic spine length</th>
<td id="B73C5D88FF8E0007B47A60D46DB9DB11" box="[579,712,1528,1550]" gridcol="1" gridrow="16" pageId="2" pageNumber="114">26.9</td>
<td id="B73C5D88FF8E0007B50E60D46CCDDB11" box="[823,956,1528,1550]" gridcol="2" gridrow="16" pageId="2" pageNumber="114">29.1</td>
<td id="B73C5D88FF8E0007B21860D46BD7DB11" box="[1057,1190,1528,1550]" gridcol="3" gridrow="16" pageId="2" pageNumber="114">27.4</td>
<td id="B73C5D88FF8E0007B32860D46A13DB11" box="[1297,1378,1528,1550]" gridcol="4" gridrow="16" pageId="2" pageNumber="114">27.8</td>
</tr>
<tr id="F4ED34F4FF8E0007B6A6630E6A13DB27" box="[159,1378,1570,1592]" gridrow="17" pageId="2" pageNumber="114">
<th id="B73C5D88FF8E0007B6A6630E6EB9DB27" box="[159,456,1570,1592]" gridcol="0" gridrow="17" pageId="2" pageNumber="114">Cleithral width</th>
<td id="B73C5D88FF8E0007B47A630E6DB9DB27" box="[579,712,1570,1592]" gridcol="1" gridrow="17" pageId="2" pageNumber="114">33.0</td>
<td id="B73C5D88FF8E0007B50E630E6CCDDB27" box="[823,956,1570,1592]" gridcol="2" gridrow="17" pageId="2" pageNumber="114">32.4</td>
<td id="B73C5D88FF8E0007B218630E6BD7DB27" box="[1057,1190,1570,1592]" gridcol="3" gridrow="17" pageId="2" pageNumber="114">33.5</td>
<td id="B73C5D88FF8E0007B328630E6A13DB27" box="[1297,1378,1570,1592]" gridcol="4" gridrow="17" pageId="2" pageNumber="114">33.0</td>
</tr>
<tr id="F4ED34F4FF8E0007B6A663676A13DB7D" box="[159,1378,1611,1634]" gridrow="18" pageId="2" pageNumber="114">
<th id="B73C5D88FF8E0007B6A663676EB9DB7D" box="[159,456,1611,1634]" gridcol="0" gridrow="18" pageId="2" pageNumber="114">Caudal peduncle depth</th>
<td id="B73C5D88FF8E0007B47A63676DB9DB7D" box="[579,712,1611,1634]" gridcol="1" gridrow="18" pageId="2" pageNumber="114">12.6</td>
<td id="B73C5D88FF8E0007B50E63676CCDDB7D" box="[823,956,1611,1634]" gridcol="2" gridrow="18" pageId="2" pageNumber="114">11.8</td>
<td id="B73C5D88FF8E0007B21863676BD7DB7D" box="[1057,1190,1611,1634]" gridcol="3" gridrow="18" pageId="2" pageNumber="114">12.7</td>
<td id="B73C5D88FF8E0007B32863676A13DB7D" box="[1297,1378,1611,1634]" gridcol="4" gridrow="18" pageId="2" pageNumber="114">12.4</td>
</tr>
<tr id="F4ED34F4FF8E0007B6A663596A13DB94" box="[159,1378,1653,1675]" gridrow="19" pageId="2" pageNumber="114">
<th id="B73C5D88FF8E0007B6A663596A13DB94" box="[159,1378,1653,1675]" colspan="5" colspanRight="4" gridcol="0" gridrow="19" pageId="2" pageNumber="114">
<emphasis id="78A9EAA4FF8EFFFAB6A663596E41DB95" bold="true" box="[159,304,1653,1675]" pageId="2" pageNumber="114">Percent of HL</emphasis>
</th>
</tr>
<tr id="F4ED34F4FF8E0007B6A663B36A13DBAA" box="[159,1378,1695,1717]" gridrow="20" pageId="2" pageNumber="114">
<th id="B73C5D88FF8E0007B6A663B36EB9DBAA" box="[159,456,1695,1717]" gridcol="0" gridrow="20" pageId="2" pageNumber="114">Head depth</th>
<td id="B73C5D88FF8E0007B47A63B36DB9DBAA" box="[579,712,1695,1717]" gridcol="1" gridrow="20" pageId="2" pageNumber="114">63.9</td>
<td id="B73C5D88FF8E0007B50E63B36CCDDBAA" box="[823,956,1695,1717]" gridcol="2" gridrow="20" pageId="2" pageNumber="114">67.1</td>
<td id="B73C5D88FF8E0007B21863B36BD7DBAA" box="[1057,1190,1695,1717]" gridcol="3" gridrow="20" pageId="2" pageNumber="114">63.6</td>
<td id="B73C5D88FF8E0007B32863B36A13DBAA" box="[1297,1378,1695,1717]" gridcol="4" gridrow="20" pageId="2" pageNumber="114">64.8</td>
</tr>
<tr id="F4ED34F4FF8E0007B6A663E46A13DBC1" box="[159,1378,1736,1758]" gridrow="21" pageId="2" pageNumber="114">
<th id="B73C5D88FF8E0007B6A663E46EB9DBC1" box="[159,456,1736,1758]" gridcol="0" gridrow="21" pageId="2" pageNumber="114">Snout length</th>
<td id="B73C5D88FF8E0007B47A63E46DB9DBC1" box="[579,712,1736,1758]" gridcol="1" gridrow="21" pageId="2" pageNumber="114">68.8</td>
<td id="B73C5D88FF8E0007B50E63E46CCDDBC1" box="[823,956,1736,1758]" gridcol="2" gridrow="21" pageId="2" pageNumber="114">68.5</td>
<td id="B73C5D88FF8E0007B21863E46BD7DBC1" box="[1057,1190,1736,1758]" gridcol="3" gridrow="21" pageId="2" pageNumber="114">70.3</td>
<td id="B73C5D88FF8E0007B32863E46A13DBC1" box="[1297,1378,1736,1758]" gridcol="4" gridrow="21" pageId="2" pageNumber="114">69.2</td>
</tr>
<tr id="F4ED34F4FF8E0007B6A663DE6A13DA17" box="[159,1378,1778,1800]" gridrow="22" pageId="2" pageNumber="114">
<th id="B73C5D88FF8E0007B6A663DE6EB9DA17" box="[159,456,1778,1800]" gridcol="0" gridrow="22" pageId="2" pageNumber="114">Interorbital width</th>
<td id="B73C5D88FF8E0007B47A63DE6DB9DA17" box="[579,712,1778,1800]" gridcol="1" gridrow="22" pageId="2" pageNumber="114">45.2</td>
<td id="B73C5D88FF8E0007B50E63DE6CCDDA17" box="[823,956,1778,1800]" gridcol="2" gridrow="22" pageId="2" pageNumber="114">45.8</td>
<td id="B73C5D88FF8E0007B21863DE6BD7DA17" box="[1057,1190,1778,1800]" gridcol="3" gridrow="22" pageId="2" pageNumber="114">44.5</td>
<td id="B73C5D88FF8E0007B32863DE6A13DA17" box="[1297,1378,1778,1800]" gridcol="4" gridrow="22" pageId="2" pageNumber="114">45.2</td>
</tr>
<tr id="F4ED34F4FF8E0007B6A662376A13DA2D" box="[159,1378,1819,1842]" gridrow="23" pageId="2" pageNumber="114">
<th id="B73C5D88FF8E0007B6A662376EB9DA2D" box="[159,456,1819,1842]" gridcol="0" gridrow="23" pageId="2" pageNumber="114">Orbital diameter</th>
<td id="B73C5D88FF8E0007B47A62376DB9DA2D" box="[579,712,1819,1842]" gridcol="1" gridrow="23" pageId="2" pageNumber="114">17.3</td>
<td id="B73C5D88FF8E0007B50E62376CCDDA2D" box="[823,956,1819,1842]" gridcol="2" gridrow="23" pageId="2" pageNumber="114">18.3</td>
<td id="B73C5D88FF8E0007B21862376BD7DA2D" box="[1057,1190,1819,1842]" gridcol="3" gridrow="23" pageId="2" pageNumber="114">17.1</td>
<td id="B73C5D88FF8E0007B32862376A13DA2D" box="[1297,1378,1819,1842]" gridcol="4" gridrow="23" pageId="2" pageNumber="114">17.6</td>
</tr>
<tr id="F4ED34F4FF8E0007B6A662696A13DA44" box="[159,1378,1861,1883]" gridrow="24" pageId="2" pageNumber="114">
<th id="B73C5D88FF8E0007B6A662696EB9DA44" box="[159,456,1861,1883]" gridcol="0" gridrow="24" pageId="2" pageNumber="114">Mandibular ramus length</th>
<td id="B73C5D88FF8E0007B47A62696DB9DA44" box="[579,712,1861,1883]" gridcol="1" gridrow="24" pageId="2" pageNumber="114">13.3</td>
<td id="B73C5D88FF8E0007B50E62696CCDDA44" box="[823,956,1861,1883]" gridcol="2" gridrow="24" pageId="2" pageNumber="114">11.4</td>
<td id="B73C5D88FF8E0007B21862696BD7DA44" box="[1057,1190,1861,1883]" gridcol="3" gridrow="24" pageId="2" pageNumber="114">13.2</td>
<td id="B73C5D88FF8E0007B32862696A13DA44" box="[1297,1378,1861,1883]" gridcol="4" gridrow="24" pageId="2" pageNumber="114">12.6</td>
</tr>
<tr id="F4ED34F4FF8E0007B6A662436A13DA9B" box="[159,1378,1903,1924]" gridrow="25" pageId="2" pageNumber="114">
<th id="B73C5D88FF8E0007B6A662436A13DA9B" box="[159,1378,1903,1924]" colspan="5" colspanRight="4" gridcol="0" gridrow="25" pageId="2" pageNumber="114">
<emphasis id="78A9EAA4FF8EFFFAB6A662436E54DA9B" bold="true" box="[159,293,1903,1924]" pageId="2" pageNumber="114">Dot numbers</emphasis>
</th>
</tr>
<tr id="F4ED34F4FF8E0007B6A662B46A13DAB1" box="[159,1378,1944,1966]" gridrow="26" pageId="2" pageNumber="114" rowspan-4="1">
<th id="B73C5D88FF8E0007B6A662B46EB9DAB1" box="[159,456,1944,1966]" gridcol="0" gridrow="26" pageId="2" pageNumber="114">Dots on pectoral fin Left/Right</th>
<td id="B73C5D88FF8E0007B47A62B46DB9DAB1" box="[579,712,1944,1966]" gridcol="1" gridrow="26" pageId="2" pageNumber="114">215/230</td>
<td id="B73C5D88FF8E0007B50E62B46CCDDAB1" box="[823,956,1944,1966]" gridcol="2" gridrow="26" pageId="2" pageNumber="114">129/114</td>
<td id="B73C5D88FF8E0007B21862B46BD7DAB1" box="[1057,1190,1944,1966]" gridcol="3" gridrow="26" pageId="2" pageNumber="114">112/104</td>
</tr>
<tr id="F4ED34F4FF8E0007B6A662EE6A13DAC7" box="[159,1378,1986,2008]" gridrow="27" pageId="2" pageNumber="114" rowspan-4="1">
<th id="B73C5D88FF8E0007B6A662EE6EB9DAC7" box="[159,456,1986,2008]" gridcol="0" gridrow="27" pageId="2" pageNumber="114">Dots on pelvic-fin Left/Right</th>
<td id="B73C5D88FF8E0007B47A62EE6DB9DAC7" box="[579,712,1986,2008]" gridcol="1" gridrow="27" pageId="2" pageNumber="114">161/204</td>
<td id="B73C5D88FF8E0007B50E62EE6CCDDAC7" box="[823,956,1986,2008]" gridcol="2" gridrow="27" pageId="2" pageNumber="114">95/91</td>
<td id="B73C5D88FF8E0007B21862EE6BD7DAC7" box="[1057,1190,1986,2008]" gridcol="3" gridrow="27" pageId="2" pageNumber="114">114/107</td>
</tr>
</table>
</paragraph>
<subSubSection id="02C7653DFF8FFFFBB6FE65BA6E7DDE30" pageId="3" pageNumber="115" type="diagnosis">
<paragraph id="4A6236B6FF8FFFFBB6FE65BA6E7DDE30" blockId="3.[151,1437,150,2021]" pageId="3" pageNumber="115">
<emphasis id="78A9EAA4FF8FFFFBB6FE65BA6E33DDAF" bold="true" box="[199,322,150,176]" pageId="3" pageNumber="115">Diagnosis.</emphasis>
<taxonomicName id="8DDD4D35FF8FFFFBB77065BB6D7DDDAF" box="[329,524,151,177]" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="115" phylum="Chordata" rank="species" species="orcesi">
<emphasis id="78A9EAA4FF8FFFFBB77065BB6D7DDDAF" box="[329,524,151,177]" italics="true" pageId="3" pageNumber="115">Panaqolus orcesi</emphasis>
</taxonomicName>
can be distinguished from all congeners, except
<emphasis id="78A9EAA4FF8FFFFBB20065BB6A39DDAF" box="[1081,1352,151,177]" italics="true" pageId="3" pageNumber="115">
<taxonomicName id="8DDD4D35FF8FFFFBB20065BB6B8CDDAE" baseAuthorityName="Kanazawa" baseAuthorityYear="1958" box="[1081,1277,151,177]" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="115" phylum="Chordata" rank="species" species="albomaculatus">P. albomaculatus</taxonomicName>
,
<taxonomicName id="8DDD4D35FF8FFFFBB33065BB6A39DDAF" authorityName="Cramer &amp; Rapp Py-Daniel" authorityYear="2015" box="[1289,1352,151,176]" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="115" phylum="Chordata" rank="species" species="nix">P. nix</taxonomicName>
</emphasis>
, and
<taxonomicName id="8DDD4D35FF8FFFFBB3B165BB6E6FDDCB" authorityName="Provenzano-Rizzi &amp; Barriga-Salazar &amp; Stewart" authorityYear="2024" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="115" phylum="Chordata" rank="species" species="pantostiktos">
<emphasis id="78A9EAA4FF8FFFFBB3B165BB6E6FDDCB" italics="true" pageId="3" pageNumber="115">P. pantostiktos</emphasis>
</taxonomicName>
by base color of head, body, and fins uniformly grey, brown, or black, with light (cream, yellow or gold) dots
<emphasis id="78A9EAA4FF8FFFFBB6F465F26F9ADDE8" box="[205,235,222,247]" italics="true" pageId="3" pageNumber="115">vs.</emphasis>
head, body, and fins with light or dark bars, bands, stripes, or saddles contrasting with the base color, never with light dots or rounded blotches.
<taxonomicName id="8DDD4D35FF8FFFFBB416642D6D80DC04" box="[559,753,257,283]" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="115" phylum="Chordata" rank="species" species="orcesi">
<emphasis id="78A9EAA4FF8FFFFBB416642D6D80DC04" box="[559,753,257,283]" italics="true" pageId="3" pageNumber="115">Panaqolus orcesi</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="8DDD4D35FF8FFFFBB21A642E6B95DC04" baseAuthorityName="Kanazawa" baseAuthorityYear="1958" box="[1059,1252,257,283]" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="115" phylum="Chordata" rank="species" species="albomaculatus">
<emphasis id="78A9EAA4FF8FFFFBB21A642E6B95DC04" box="[1059,1252,257,283]" italics="true" pageId="3" pageNumber="115">P. albomaculatus</emphasis>
</taxonomicName>
by having more dentary teeth (5–7
<emphasis id="78A9EAA4FF8FFFFBB75164096EF6DC21" box="[360,391,293,318]" italics="true" pageId="3" pageNumber="115">vs.</emphasis>
3–5), having distal tips of the dentaries converging to form an acute angle
<emphasis id="78A9EAA4FF8FFFFBB2FF64096B94DC21" box="[1222,1253,293,318]" italics="true" pageId="3" pageNumber="115">vs.</emphasis>
dentaries nearly parallel, and by reaching larger sizes (maximum standard length
<quantity id="8D259B53FF8FFFFBB55964646CA0DC7C" box="[864,977,328,355]" metricMagnitude="-1" metricUnit="m" metricValue="1.8059999999999998" pageId="3" pageNumber="115" unit="mm" value="180.6">180.6 mm</quantity>
<emphasis id="78A9EAA4FF8FFFFBB5EF64656C85DC7D" box="[982,1012,329,354]" italics="true" pageId="3" pageNumber="115">vs.</emphasis>
<quantity id="8D259B53FF8FFFFBB5C364646B11DC7D" box="[1018,1120,328,355]" metricMagnitude="-2" metricUnit="m" metricValue="9.05" pageId="3" pageNumber="115" unit="mm" value="90.5">90.5 mm</quantity>
).
<taxonomicName id="8DDD4D35FF8FFFFBB24B64646A43DC7D" box="[1138,1330,328,354]" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="115" phylum="Chordata" rank="species" species="orcesi">
<emphasis id="78A9EAA4FF8FFFFBB24B64646A43DC7D" box="[1138,1330,328,354]" italics="true" pageId="3" pageNumber="115">Panaqolus orcesi</emphasis>
</taxonomicName>
is further distinguished from
<taxonomicName id="8DDD4D35FF8FFFFBB74A64406EC3DC9A" authorityName="Cramer &amp; Rapp Py-Daniel" authorityYear="2015" box="[371,434,364,389]" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="115" phylum="Chordata" rank="species" species="nix">
<emphasis id="78A9EAA4FF8FFFFBB74A64406EC3DC9A" box="[371,434,364,389]" italics="true" pageId="3" pageNumber="115">P. nix</emphasis>
</taxonomicName>
by shorter inter-dorsal distance (12.1–12.7% SL
<emphasis id="78A9EAA4FF8FFFFBB5D864406B71DC9A" box="[993,1024,364,389]" italics="true" pageId="3" pageNumber="115">vs.</emphasis>
13.7–19.4% SL), shorter abdominal length (23.0–25.3% SL
<emphasis id="78A9EAA4FF8FFFFBB79364BC6EB9DCB6" box="[426,456,400,425]" italics="true" pageId="3" pageNumber="115">vs.</emphasis>
29.5–34.9% SL), longer dorsal-fin base length (33.1–35.0% SL
<emphasis id="78A9EAA4FF8FFFFBB28A64BC6BA0DCB6" box="[1203,1233,400,425]" italics="true" pageId="3" pageNumber="115">vs.</emphasis>
20.5–28.8% SL), longer dorsal- and pectoral-fin spines (34.0–34.1% SL
<emphasis id="78A9EAA4FF8FFFFBB539649F6C6EDCD3" box="[768,799,435,460]" italics="true" pageId="3" pageNumber="115">vs.</emphasis>
26.2–32.5% SL and 35.8–37.4% SL
<emphasis id="78A9EAA4FF8FFFFBB2F9649F6BAEDCD3" box="[1216,1247,435,460]" italics="true" pageId="3" pageNumber="115">vs.</emphasis>
29.5–34.9% SL, respectively), a deeper caudal peduncle (11.8–12.7% SL
<emphasis id="78A9EAA4FF8FFFFBB52564FB6C4ADCEF" box="[796,827,471,496]" italics="true" pageId="3" pageNumber="115">vs.</emphasis>
9.3–11.4% SL), longer snout length (68.5–70.3% HL
<emphasis id="78A9EAA4FF8FFFFBB6AE64D66FC7DF0C" box="[151,182,506,531]" italics="true" pageId="3" pageNumber="115">vs.</emphasis>
49.7–63.7% HL), larger orbital diameter (17.1–18.3% HL
<emphasis id="78A9EAA4FF8FFFFBB57264D66C18DF0C" box="[843,873,506,531]" italics="true" pageId="3" pageNumber="115">vs.</emphasis>
12.5–16.8% HL), and apparently reaching greater sizes (maximum standard length
<quantity id="8D259B53FF8FFFFBB43467316DF1DF28" box="[525,640,541,567]" metricMagnitude="-1" metricUnit="m" metricValue="1.8059999999999998" pageId="3" pageNumber="115" unit="mm" value="180.6">180.6 mm</quantity>
<emphasis id="78A9EAA4FF8FFFFBB4BE67326DD7DF28" box="[647,678,542,567]" italics="true" pageId="3" pageNumber="115">vs.</emphasis>
<quantity id="8D259B53FF8FFFFBB49767316C55DF28" box="[686,804,541,567]" metricMagnitude="-1" metricUnit="m" metricValue="1.122" pageId="3" pageNumber="115" unit="mm" value="112.2">112.2 mm</quantity>
).
<taxonomicName id="8DDD4D35FF8FFFFBB50E67316C8BDF28" box="[823,1018,541,567]" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="115" phylum="Chordata" rank="species" species="orcesi">
<emphasis id="78A9EAA4FF8FFFFBB50E67316C8BDF28" box="[823,1018,541,567]" italics="true" pageId="3" pageNumber="115">Panaqolus orcesi</emphasis>
</taxonomicName>
is distinguished from
<taxonomicName id="8DDD4D35FF8FFFFBB2C167326AEDDF28" authorityName="Provenzano-Rizzi &amp; Barriga-Salazar &amp; Stewart" authorityYear="2024" box="[1272,1436,541,567]" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="115" phylum="Chordata" rank="species" species="pantostiktos">
<emphasis id="78A9EAA4FF8FFFFBB2C167326AEDDF28" box="[1272,1436,541,567]" italics="true" pageId="3" pageNumber="115">P. pantostiktos</emphasis>
</taxonomicName>
by its wider interorbital distance (44.5–45.8% HL
<emphasis id="78A9EAA4FF8FFFFBB4E3676D6D88DF45" box="[730,761,577,602]" italics="true" pageId="3" pageNumber="115">vs.</emphasis>
40.2–43.7% HL), longer snout length (68.5–70.3% HL
<emphasis id="78A9EAA4FF8FFFFBB347676D6AEDDF45" box="[1406,1436,577,602]" italics="true" pageId="3" pageNumber="115">vs.</emphasis>
62.7–68.9% HL), and longer inter-dorsal distance (12.1–12.7% SL
<emphasis id="78A9EAA4FF8FFFFBB5AB67496CC0DF61" box="[914,945,613,638]" italics="true" pageId="3" pageNumber="115">vs.</emphasis>
10.3–12.0% SL) (
<figureCitation id="D2E62A33FF8FFFFBB2BA67486B83DF61" box="[1155,1266,612,638]" captionStart="FIGURE 4" captionStartId="6.[152,255,1944,1968]" captionTargetBox="[429,1158,189,1912]" captionTargetId="figure-17@6.[411,1176,181,1920]" captionTargetPageId="6" captionText="FIGURE 4. Comparisons of diagnostic relationships between the two new Panaqolus spp. for selected morphometric variables. Probability that allometric relationships for the two species are the same is indicated in each panel based on analysis of covariance." figureDoi="http://doi.org/10.5281/zenodo.11610316" httpUri="https://zenodo.org/record/11610316/files/figure.png" pageId="3" pageNumber="115">Fig. 4a–c</figureCitation>
);
<taxonomicName id="8DDD4D35FF8FFFFBB33E67496A16DF61" box="[1287,1383,613,638]" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="115" phylum="Chordata" rank="species" species="orcesi">
<emphasis id="78A9EAA4FF8FFFFBB33E67496A16DF61" box="[1287,1383,613,638]" italics="true" pageId="3" pageNumber="115">P. orcesi</emphasis>
</taxonomicName>
also appears to have a shorter dorsal-fin spine (34.0–34.1% SL
<emphasis id="78A9EAA4FF8FFFFBB51D67A46C32DFBE" box="[804,835,648,673]" italics="true" pageId="3" pageNumber="115">vs.</emphasis>
35.2–38.4% SL), but due to some broken spines, data were insufficient for a statistical test. Further distinguished from
<taxonomicName id="8DDD4D35FF8FFFFBB5BA67876B56DFDA" authorityName="Provenzano-Rizzi &amp; Barriga-Salazar &amp; Stewart" authorityYear="2024" box="[899,1063,683,709]" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="115" phylum="Chordata" rank="species" species="pantostiktos">
<emphasis id="78A9EAA4FF8FFFFBB5BA67876B56DFDA" box="[899,1063,683,709]" italics="true" pageId="3" pageNumber="115">P. pantostiktos</emphasis>
</taxonomicName>
by having smaller average mid-lateral dot sizes for a given size individual (
<figureCitation id="D2E62A33FF8FFFFBB44267E26DBADFF6" box="[635,715,718,745]" captionStart="FIGURE 5" captionStartId="7.[152,255,1909,1933]" captionTargetBox="[279,1310,207,1859]" captionTargetId="figure-19@7.[253,1335,181,1885]" captionTargetPageId="7" captionText="FIGURE 5. Comparisons of diagnostic differences versus SL in: a) mid-lateral dot sizes (based on averages for 10 of the largest dots below dorsal fin), and b) pectoral-fin dot counts (based on average of counts for both pectoral fins of each specimen). Probability that allometric relationships for the two species are the same is indicated in each panel based on analysis of covariance." figureDoi="http://doi.org/10.5281/zenodo.11610318" httpUri="https://zenodo.org/record/11610318/files/figure.png" pageId="3" pageNumber="115">Fig. 5a</figureCitation>
), and by having higher average number of dots on dorsal surface of pectoral fin (
<figureCitation id="D2E62A33FF8FFFFBB77167DE6EECDE13" box="[328,413,753,780]" captionStart="FIGURE 5" captionStartId="7.[152,255,1909,1933]" captionTargetBox="[279,1310,207,1859]" captionTargetId="figure-19@7.[253,1335,181,1885]" captionTargetPageId="7" captionText="FIGURE 5. Comparisons of diagnostic differences versus SL in: a) mid-lateral dot sizes (based on averages for 10 of the largest dots below dorsal fin), and b) pectoral-fin dot counts (based on average of counts for both pectoral fins of each specimen). Probability that allometric relationships for the two species are the same is indicated in each panel based on analysis of covariance." figureDoi="http://doi.org/10.5281/zenodo.11610318" httpUri="https://zenodo.org/record/11610318/files/figure.png" pageId="3" pageNumber="115">Fig. 5b</figureCitation>
). Relatively smaller dot sizes and higher dot densities in
<taxonomicName id="8DDD4D35FF8FFFFBB21C67DE6BF4DE14" box="[1061,1157,754,779]" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="115" phylum="Chordata" rank="species" species="orcesi">
<emphasis id="78A9EAA4FF8FFFFBB21C67DE6BF4DE14" box="[1061,1157,754,779]" italics="true" pageId="3" pageNumber="115">P. orcesi</emphasis>
</taxonomicName>
are also evident in
<figureCitation id="D2E62A33FF8FFFFBB35C67DE6FB0DE30" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="4.[152,255,1961,1985]" captionStartId-1="5.[152,255,743,767]" captionStartId-2="5.[152,255,1968,1992]" captionTargetBox-0="[202,1386,179,1935]" captionTargetBox-1="[164,1423,181,719]" captionTargetBox-2="[164,1423,964,1945]" captionTargetId-0="figure-17@4.[202,1386,179,1935]" captionTargetId-1="figure-19@5.[164,1423,181,719]" captionTargetId-2="figure-85@5.[164,1423,964,1945]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="5" captionText-0="FIGURE 1. Panaqolus orcesi, MEPN 19492, 180.6 mm SL, holotype. Scale bar = 1 cm. Photos by F.P-R." captionText-1="FIGURE 2. Close up of mouth and buccal disk for: a) Panaqolus orcesi, MEPN 19492, 180.6 mm SL, holotype; and b) P. pantostiktos, MEPN 19495, 163.4 mm SL, holotype. Photos by F.P-R." captionText-2="FIGURE 3. Dorsal view of color patterns in specimens soon after capture for: a) Panaqolus orcesi, with cream-colored dots, and b) Panaqolus pantostiktos, with yellow- or gold-colored dots. Photos by R.B-S." figureDoi-0="http://doi.org/10.5281/zenodo.11610308" figureDoi-1="http://doi.org/10.5281/zenodo.11610310" figureDoi-2="http://doi.org/10.5281/zenodo.11610312" httpUri-0="https://zenodo.org/record/11610308/files/figure.png" httpUri-1="https://zenodo.org/record/11610310/files/figure.png" httpUri-2="https://zenodo.org/record/11610312/files/figure.png" pageId="3" pageNumber="115">Figs. 1–3</figureCitation>
and
<figureCitation id="D2E62A33FF8FFFFBB6C166396E78DE30" box="[248,265,789,815]" captionStart="FIGURE 6" captionStartId="10.[152,255,2004,2028]" captionTargetBox="[164,1423,309,1980]" captionTargetId="figure-64@10.[164,1423,309,1980]" captionTargetPageId="10" captionText="FIGURE 6. Panaqolus pantostiktos, MEPN 19495, 163.4 mm SL, holotype. Scale bars = 1 cm. Photos by F.P-R." figureDoi="http://doi.org/10.5281/zenodo.11610320" httpUri="https://zenodo.org/record/11610320/files/figure.png" pageId="3" pageNumber="115">6</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="02C7653DFF8FFFF0B6FE66146DE1DC03" lastPageId="8" lastPageNumber="120" pageId="3" pageNumber="115" type="description">
<paragraph id="4A6236B6FF8FFFFBB6FE66146EB8D938" blockId="3.[151,1437,150,2021]" pageId="3" pageNumber="115">
<emphasis id="78A9EAA4FF8FFFFBB6FE66146E28DE4D" bold="true" box="[199,345,824,850]" pageId="3" pageNumber="115">Description.</emphasis>
Morphometric data in
<tableCitation id="075F030DFF8FFFFBB46A66146DE6DE4C" box="[595,663,824,851]" captionStart="TABLE 1" captionStartId="2.[152,244,828,852]" captionTargetPageId="2" captionText="TABLE 1. Morphometric data of Panaqolus orcesi, expressed as percent of SL or percent of HL." httpUri="http://table.plazi.org/id/1EA2663EFF8EFFFAB6A166106BDBDE4B" pageId="3" pageNumber="115" tableUuid="1EA2663EFF8EFFFAB6A166106BDBDE4B">Tab. 1</tableCitation>
. Medium-sized loricariid with standard length of measured specimens up to
<quantity id="8D259B53FF8FFFFBB6E166706E3BDE68" box="[216,330,860,887]" metricMagnitude="-1" metricUnit="m" metricValue="1.8059999999999998" pageId="3" pageNumber="115" unit="mm" value="180.6">180.6 mm</quantity>
SL. Body robust, quite depressed anteriorly and compressed posteriorly (
<figureCitation id="D2E62A33FF8FFFFBB2BF66706BB9DE69" box="[1158,1224,860,886]" captionStart="FIGURE 1" captionStartId="4.[152,255,1961,1985]" captionTargetBox="[202,1386,179,1935]" captionTargetId="figure-17@4.[202,1386,179,1935]" captionTargetPageId="4" captionText="FIGURE 1. Panaqolus orcesi, MEPN 19492, 180.6 mm SL, holotype. Scale bar = 1 cm. Photos by F.P-R." figureDoi="http://doi.org/10.5281/zenodo.11610308" httpUri="https://zenodo.org/record/11610308/files/figure.png" pageId="3" pageNumber="115">Fig. 1</figureCitation>
). Dorsal profile of head from snout tip to anterior border of orbit straight and inclined with slope of approximately 45 degrees, then gently convex or straight to dorsal-fin origin and descending straight or gently convex to origin of first procurrent caudal-fin ray. Orbit dorsal margin and posterior tip of supraoccipital slightly elevated. Greatest body depth near dorsal-fin origin. Caudal peduncle compressed, ovoid in cross-section, deep and robust (
<figureCitation id="D2E62A33FF8FFFFBB24666C66BB0D91B" box="[1151,1217,1002,1028]" captionStart="FIGURE 1" captionStartId="4.[152,255,1961,1985]" captionTargetBox="[202,1386,179,1935]" captionTargetId="figure-17@4.[202,1386,179,1935]" captionTargetPageId="4" captionText="FIGURE 1. Panaqolus orcesi, MEPN 19492, 180.6 mm SL, holotype. Scale bar = 1 cm. Photos by F.P-R." figureDoi="http://doi.org/10.5281/zenodo.11610308" httpUri="https://zenodo.org/record/11610308/files/figure.png" pageId="3" pageNumber="115">Fig. 1</figureCitation>
). Ventral profile of head and body flat (
<figureCitation id="D2E62A33FF8FFFFBB74F61216EC6D938" box="[374,439,1037,1063]" captionStart="FIGURE 1" captionStartId="4.[152,255,1961,1985]" captionTargetBox="[202,1386,179,1935]" captionTargetId="figure-17@4.[202,1386,179,1935]" captionTargetPageId="4" captionText="FIGURE 1. Panaqolus orcesi, MEPN 19492, 180.6 mm SL, holotype. Scale bar = 1 cm. Photos by F.P-R." figureDoi="http://doi.org/10.5281/zenodo.11610308" httpUri="https://zenodo.org/record/11610308/files/figure.png" pageId="3" pageNumber="115">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="4A6236B6FF8FFFFBB6FE611D6DFCD98D" blockId="3.[151,1437,150,2021]" pageId="3" pageNumber="115">
In dorsal view, body widest near dorsal-fin origin, narrowing posteriorly. Eye dorsolaterally placed; orbit diameter 17.1–18.3% HL. Interorbital width 44.5–45.8% HL, almost flat or slightly concave near eyes. Nares small and ovoid, slightly longer than wide (
<figureCitation id="D2E62A33FF8FFFFBB40261546D0DD98D" box="[571,636,1144,1170]" captionStart="FIGURE 1" captionStartId="4.[152,255,1961,1985]" captionTargetBox="[202,1386,179,1935]" captionTargetId="figure-17@4.[202,1386,179,1935]" captionTargetPageId="4" captionText="FIGURE 1. Panaqolus orcesi, MEPN 19492, 180.6 mm SL, holotype. Scale bar = 1 cm. Photos by F.P-R." figureDoi="http://doi.org/10.5281/zenodo.11610308" httpUri="https://zenodo.org/record/11610308/files/figure.png" pageId="3" pageNumber="115">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="4A6236B6FF8FFFFBB6FE61B76B6CD83F" blockId="3.[151,1437,150,2021]" pageId="3" pageNumber="115">
Oral disk longitudinally elliptical, maxillary barbels developed and evident; lips papillose with smooth borders; lower lip larger and more developed than upper (
<figureCitation id="D2E62A33FF8FFFFBB4F461936C50D9C6" box="[717,801,1215,1241]" captionStart="FIGURE 2" captionStartId="5.[152,255,743,767]" captionTargetBox="[164,1423,181,719]" captionTargetId="figure-19@5.[164,1423,181,719]" captionTargetPageId="5" captionText="FIGURE 2. Close up of mouth and buccal disk for: a) Panaqolus orcesi, MEPN 19492, 180.6 mm SL, holotype; and b) P. pantostiktos, MEPN 19495, 163.4 mm SL, holotype. Photos by F.P-R." figureDoi="http://doi.org/10.5281/zenodo.11610310" httpUri="https://zenodo.org/record/11610310/files/figure.png" pageId="3" pageNumber="115">Fig. 2a</figureCitation>
). Teeth spoon-shaped and unicuspid or with very small lateral cusp. Premaxillary teeth 4–6 per ramus (mode 5); dentary teeth 5–7 per ramus (mode 6). Premaxillary and dentary tooth rows both meet distally at an angle of approximately 45° (
<figureCitation id="D2E62A33FF8FFFFBB587602A6B61D83F" box="[958,1040,1286,1312]" captionStart="FIGURE 2" captionStartId="5.[152,255,743,767]" captionTargetBox="[164,1423,181,719]" captionTargetId="figure-19@5.[164,1423,181,719]" captionTargetPageId="5" captionText="FIGURE 2. Close up of mouth and buccal disk for: a) Panaqolus orcesi, MEPN 19492, 180.6 mm SL, holotype; and b) P. pantostiktos, MEPN 19495, 163.4 mm SL, holotype. Photos by F.P-R." figureDoi="http://doi.org/10.5281/zenodo.11610310" httpUri="https://zenodo.org/record/11610310/files/figure.png" pageId="3" pageNumber="115">Fig. 2a</figureCitation>
).
</paragraph>
<paragraph id="4A6236B6FF8FFFFBB6FE60056DCFDB24" blockId="3.[151,1437,150,2021]" pageId="3" pageNumber="115">
Head and body covered with plates. Ventral surface of head and belly covered with small and irregularly shaped plates, with small or very small odontodes. In the relatively large specimens available, lateral plates of body have enlarged odontodes, like spines (
<figureCitation id="D2E62A33FF8FFFFBB42C605C6D25D895" box="[533,596,1392,1418]" captionStart="FIGURE 1" captionStartId="4.[152,255,1961,1985]" captionTargetBox="[202,1386,179,1935]" captionTargetId="figure-17@4.[202,1386,179,1935]" captionTargetPageId="4" captionText="FIGURE 1. Panaqolus orcesi, MEPN 19492, 180.6 mm SL, holotype. Scale bar = 1 cm. Photos by F.P-R." figureDoi="http://doi.org/10.5281/zenodo.11610308" httpUri="https://zenodo.org/record/11610308/files/figure.png" pageId="3" pageNumber="115">Fig 1</figureCitation>
). Posterior to anal-fin origin, the developed spines on lateral plates are arranged in five longitudinal rows, which seem to form keels (
<figureCitation id="D2E62A33FF8FFFFBB56B60BF6CFCD8B1" box="[850,909,1427,1454]" captionStart="FIGURE 1" captionStartId="4.[152,255,1961,1985]" captionTargetBox="[202,1386,179,1935]" captionTargetId="figure-17@4.[202,1386,179,1935]" captionTargetPageId="4" captionText="FIGURE 1. Panaqolus orcesi, MEPN 19492, 180.6 mm SL, holotype. Scale bar = 1 cm. Photos by F.P-R." figureDoi="http://doi.org/10.5281/zenodo.11610308" httpUri="https://zenodo.org/record/11610308/files/figure.png" pageId="3" pageNumber="115">Fig 1</figureCitation>
); 24–25 lateral plates. Supraoccipital bordered posteriorly by 2 plates on each side; 9–11 plates along dorsal-fin base; 5–7 plates between dorsal and adipose fins; 3–5 plates between adipose-fin and caudal-fin origin; 3 plates along anal-fin base; 9–12 plates between last anal-fin ray and caudal-fin origin. Hypertrophied cheek odontodes 15–22, increasing in size posteriorly; longest odontodes surpass pectoral-fin origin; odontode tips curved.
</paragraph>
<paragraph id="4A6236B6FF8FFFFBB6FE63696A25DA65" blockId="3.[151,1437,150,2021]" pageId="3" pageNumber="115">Dorsal-fin I,7 with small anterior triangular spinelet; pectoral-fin I,6; pelvic-fin I,5; anal-fin i,4; caudal-fin i,14,i. Dorsal-fin origin anterior to pelvic-fin origin; when adpressed overlaps adipose-fin origin; dorsal-fin spine lock functional; last ray of dorsal-fin without posterior membrane. Pectoral-fin origin anterior to posterior margin of orbit; when adpressed, pectoral spine almost surpasses half of pelvic-fin spine; pectoral-fin spine robust, dorsal surface of spine with developed odontodes, larger at distal tip; membrane between spine and first ray without fleshy extension. Pelvic-fin origin at vertical through base of third dorsal-fin ray; distal tip of pelvic spine slightly curved, when adpressed reaching mid-length of anal-fin base. Anal-fin origin behind the vertical below last dorsal-fin ray. Triangular adipose-fin developed, with well-ossified spine bearing odontodes; adipose-fin membrane with posterior margin concave to nearly vertical. Caudal-fin obliquely truncate; unbranched caudal-fin rays with filaments.</paragraph>
<paragraph id="4A6236B6FF8FFFFBB6FE62A86C77DAFA" blockId="3.[151,1437,150,2021]" pageId="3" pageNumber="115">
<emphasis id="78A9EAA4FF8FFFFBB6FE62A86EFEDA81" bold="true" box="[199,399,1924,1950]" pageId="3" pageNumber="115">Color in alcohol.</emphasis>
Whole body and all fins have base color uniform brown, black or dark grey, with cream or beige rounded small dots, mostly uniform sized on head and body, but some between pectoral and pelvic fins may be more variable in size and irregularly shaped (
<figureCitation id="D2E62A33FF8FFFFBB48A62E76D84DAFA" box="[691,757,1995,2021]" captionStart="FIGURE 1" captionStartId="4.[152,255,1961,1985]" captionTargetBox="[202,1386,179,1935]" captionTargetId="figure-17@4.[202,1386,179,1935]" captionTargetPageId="4" captionText="FIGURE 1. Panaqolus orcesi, MEPN 19492, 180.6 mm SL, holotype. Scale bar = 1 cm. Photos by F.P-R." figureDoi="http://doi.org/10.5281/zenodo.11610308" httpUri="https://zenodo.org/record/11610308/files/figure.png" pageId="3" pageNumber="115">Fig. 1</figureCitation>
).
</paragraph>
<caption id="1EA2663EFF88FFFCB6A162856BA5DADF" ID-DOI="http://doi.org/10.5281/zenodo.11610308" ID-Zenodo-Dep="11610308" box="[152,1236,1961,1985]" httpUri="https://zenodo.org/record/11610308/files/figure.png" pageId="4" pageNumber="116" startId="4.[152,255,1961,1985]" targetBox="[202,1386,179,1935]" targetPageId="4" targetType="figure">
<paragraph id="4A6236B6FF88FFFCB6A162856BA5DADF" blockId="4.[152,1236,1961,1985]" box="[152,1236,1961,1985]" pageId="4" pageNumber="116">
<emphasis id="78A9EAA4FF88FFFCB6A162856E69DADE" bold="true" box="[152,280,1961,1985]" pageId="4" pageNumber="116">FIGURE 1.</emphasis>
<taxonomicName id="8DDD4D35FF88FFFCB72662856D50DADF" authority=", MEPN" authorityName="MEPN" box="[287,545,1961,1985]" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="116" phylum="Chordata" rank="species" species="orcesi">
<emphasis id="78A9EAA4FF88FFFCB72662856EBFDADE" box="[287,462,1961,1985]" italics="true" pageId="4" pageNumber="116">Panaqolus orcesi</emphasis>
, MEPN
</taxonomicName>
19492, 180.6 mm SL, holotype. Scale bar = 1 cm. Photos by F.P-R.
</paragraph>
</caption>
<caption id="1EA2663EFF89FFFDB6A167CB6C11DE3C" ID-DOI="http://doi.org/10.5281/zenodo.11610310" ID-Zenodo-Dep="11610310" httpUri="https://zenodo.org/record/11610310/files/figure.png" pageId="5" pageNumber="117" startId="5.[152,255,743,767]" targetBox="[164,1423,181,719]" targetPageId="5" targetType="figure">
<paragraph id="4A6236B6FF89FFFDB6A167CB6C11DE3C" blockId="5.[151,1437,743,804]" pageId="5" pageNumber="117">
<emphasis id="78A9EAA4FF89FFFDB6A167CB6E6BDFE0" bold="true" box="[152,282,743,767]" pageId="5" pageNumber="117">FIGURE 2.</emphasis>
Close up of mouth and buccal disk for: a)
<taxonomicName id="8DDD4D35FF89FFFDB4E767CB6C95DFE0" authority=", MEPN" authorityName="MEPN" box="[734,996,743,767]" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="117" phylum="Chordata" rank="species" species="orcesi">
<emphasis id="78A9EAA4FF89FFFDB4E767CB6CE1DFE0" box="[734,912,743,767]" italics="true" pageId="5" pageNumber="117">Panaqolus orcesi</emphasis>
, MEPN
</taxonomicName>
19492, 180.6 mm SL, holotype; and b)
<taxonomicName id="8DDD4D35FF89FFFDB3B367C46E15DE3C" authority=", MEPN" authorityName="MEPN" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="117" phylum="Chordata" rank="species" species="pantostiktos">
<emphasis id="78A9EAA4FF89FFFDB3B367C46E60DE3C" italics="true" pageId="5" pageNumber="117">P. pantostiktos</emphasis>
, MEPN
</taxonomicName>
19495, 163.4 mm SL, holotype. Photos by F.P-R.
</paragraph>
</caption>
<paragraph id="4A6236B6FF89FFFDB6FE66416D6EDEB3" blockId="5.[151,1436,877,940]" pageId="5" pageNumber="117">
<emphasis id="78A9EAA4FF89FFFDB6FE66416E25DE98" bold="true" box="[199,340,877,903]" pageId="5" pageNumber="117">Color in life</emphasis>
. In a photo of freshly captured specimen, cream-colored dots on a dark gray background cover the entire fish, including fins (
<figureCitation id="D2E62A33FF89FFFDB7F966BE6D63DEB3" box="[448,530,914,940]" captionStart="FIGURE 3" captionStartId="5.[152,255,1968,1992]" captionTargetBox="[164,1423,964,1945]" captionTargetId="figure-85@5.[164,1423,964,1945]" captionTargetPageId="5" captionText="FIGURE 3. Dorsal view of color patterns in specimens soon after capture for: a) Panaqolus orcesi, with cream-colored dots, and b) Panaqolus pantostiktos, with yellow- or gold-colored dots. Photos by R.B-S." figureDoi="http://doi.org/10.5281/zenodo.11610312" httpUri="https://zenodo.org/record/11610312/files/figure.png" pageId="5" pageNumber="117">Fig. 3a</figureCitation>
).
</paragraph>
<caption id="1EA2663EFF89FFFDB6A1629C6C98DAF2" ID-DOI="http://doi.org/10.5281/zenodo.11610312" ID-Zenodo-Dep="11610312" httpUri="https://zenodo.org/record/11610312/files/figure.png" pageId="5" pageNumber="117" startId="5.[152,255,1968,1992]" targetBox="[164,1423,964,1945]" targetPageId="5" targetType="figure">
<paragraph id="4A6236B6FF89FFFDB6A1629C6C98DAF2" blockId="5.[151,1436,1968,2029]" pageId="5" pageNumber="117">
<emphasis id="78A9EAA4FF89FFFDB6A1629C6E68DAD6" bold="true" box="[152,281,1968,1993]" pageId="5" pageNumber="117">FIGURE 3.</emphasis>
Dorsal view of color patterns in specimens soon after capture for: a)
<taxonomicName id="8DDD4D35FF89FFFDB5D8629D6BE0DAD7" box="[993,1169,1969,1993]" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="117" phylum="Chordata" rank="species" species="orcesi">
<emphasis id="78A9EAA4FF89FFFDB5D8629D6BE0DAD7" box="[993,1169,1969,1993]" italics="true" pageId="5" pageNumber="117">Panaqolus orcesi</emphasis>
</taxonomicName>
, with cream-colored dots, and b)
<taxonomicName id="8DDD4D35FF89FFFDB6E762F96EBBDAF2" box="[222,458,2005,2029]" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="117" phylum="Chordata" rank="species" species="pantostiktos">
<emphasis id="78A9EAA4FF89FFFDB6E762F96EBBDAF2" box="[222,458,2005,2029]" italics="true" pageId="5" pageNumber="117">Panaqolus pantostiktos</emphasis>
</taxonomicName>
, with yellow- or gold-colored dots. Photos by R.B-S.
</paragraph>
</caption>
<caption id="1EA2663EFF8AFFFEB6A162B46E59DAE7" ID-DOI="http://doi.org/10.5281/zenodo.11610316" ID-Zenodo-Dep="11610316" httpUri="https://zenodo.org/record/11610316/files/figure.png" pageId="6" pageNumber="118" startId="6.[152,255,1944,1968]" targetBox="[429,1158,189,1912]" targetPageId="6" targetType="figure">
<paragraph id="4A6236B6FF8AFFFEB6A162B46E59DAE7" blockId="6.[151,1437,1944,2040]" pageId="6" pageNumber="118">
<emphasis id="78A9EAA4FF8AFFFEB6A162B46E6FDAAF" bold="true" box="[152,286,1944,1968]" pageId="6" pageNumber="118">FIGURE 4.</emphasis>
Comparisons of diagnostic relationships between the two new
<taxonomicName id="8DDD4D35FF8AFFFEB5EF62B46B08DAAF" box="[982,1145,1944,1968]" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="6" pageNumber="118" phylum="Chordata" rank="species" species="undetermined">
<emphasis id="78A9EAA4FF8AFFFEB5EF62B46B33DAAF" box="[982,1090,1944,1968]" italics="true" pageId="6" pageNumber="118">Panaqolus</emphasis>
spp.
</taxonomicName>
for selected morphometric variables. Probability that allometric relationships for the two species are the same is indicated in each panel based on analysis of covariance.
</paragraph>
</caption>
<caption id="1EA2663EFF8BFFFFB6A162596E7DDAE6" ID-DOI="http://doi.org/10.5281/zenodo.11610318" ID-Zenodo-Dep="11610318" httpUri="https://zenodo.org/record/11610318/files/figure.png" pageId="7" pageNumber="119" startId="7.[152,255,1909,1933]" targetBox="[279,1310,207,1859]" targetPageId="7" targetType="figure">
<paragraph id="4A6236B6FF8BFFFFB6A162596E7DDAE6" blockId="7.[151,1437,1909,2041]" pageId="7" pageNumber="119">
<emphasis id="78A9EAA4FF8BFFFFB6A162596E66DA92" bold="true" box="[152,279,1909,1933]" pageId="7" pageNumber="119">FIGURE 5.</emphasis>
Comparisons of diagnostic differences versus SL in: a) mid-lateral dot sizes (based on averages for 10 of the largest dots below dorsal fin), and b) pectoral-fin dot counts (based on average of counts for both pectoral fins of each specimen). Probability that allometric relationships for the two species are the same is indicated in each panel based on analysis of covariance.
</paragraph>
</caption>
<paragraph id="4A6236B6FF84FFF0B6FE65BA6DE1DC03" blockId="8.[151,1437,150,500]" pageId="8" pageNumber="120">
<emphasis id="78A9EAA4FF84FFF0B6FE65BA6EC6DDAF" bold="true" box="[199,439,150,176]" pageId="8" pageNumber="120">Sexual dimorphism.</emphasis>
The
<specimenCount id="5CDBFD3FFF84FFF0B7D665BA6DD2DDAF" box="[495,675,150,176]" count="3" pageId="8" pageNumber="120" type="generic">three specimens</specimenCount>
analyzed have well-developed odontodes on the pectoral spines and on the lateral and dorsal plates of the trunk, mostly posteriorly; some odontodes on this area have notable size (
<figureCitation id="D2E62A33FF84FFF0B35665966FD5DDE7" captionStart="FIGURE 1" captionStartId="4.[152,255,1961,1985]" captionTargetBox="[202,1386,179,1935]" captionTargetId="figure-17@4.[202,1386,179,1935]" captionTargetPageId="4" captionText="FIGURE 1. Panaqolus orcesi, MEPN 19492, 180.6 mm SL, holotype. Scale bar = 1 cm. Photos by F.P-R." figureDoi="http://doi.org/10.5281/zenodo.11610308" httpUri="https://zenodo.org/record/11610308/files/figure.png" pageId="8" pageNumber="120">Fig. 1</figureCitation>
). These features have been indicated in various other species of hypostomines as secondary sexual dimorphism for males, but our specimens were not sexed.
</paragraph>
</subSubSection>
<subSubSection id="02C7653DFF84FFF0B6FE640A6BFBDCB3" pageId="8" pageNumber="120" type="distribution">
<paragraph id="4A6236B6FF84FFF0B6FE640A6BFBDCB3" blockId="8.[151,1437,150,500]" pageId="8" pageNumber="120">
<emphasis id="78A9EAA4FF84FFF0B6FE640A6E87DC5F" bold="true" box="[199,502,294,320]" pageId="8" pageNumber="120">Distribution and Habitat.</emphasis>
The known distribution of this species is restricted to the Tiputini and Yasuni rivers, southern tributaries of the
<collectingRegion id="8819F854FF84FFF0B78664676E8DDC7A" box="[447,508,331,357]" country="Ecuador" name="Napo" pageId="8" pageNumber="120">Napo</collectingRegion>
River (
<figureCitation id="D2E62A33FF84FFF0B46964666DE5DC7B" box="[592,660,330,356]" captionStart="FIGURE 7" captionStartId="11.[152,255,1968,1992]" captionTargetBox="[176,1411,633,1945]" captionTargetId="figure-216@11.[176,1411,633,1945]" captionTargetPageId="11" captionText="FIGURE 7. Map of Ecuador and adjacent areas showing capture localities for Panaqolus orcesi red circles and red star (type locality), and for Panaqolus pantostiktos red rhombus (type locality)." figureDoi="http://doi.org/10.5281/zenodo.11610322" httpUri="https://zenodo.org/record/11610322/files/figure.png" pageId="8" pageNumber="120">Fig. 7</figureCitation>
). Capture sites were in relatively small lowland to piedmont streams. Specimens were collected on submerged woody debris at about
<quantity id="8D259B53FF84FFF0B54D64426CC7DC96" box="[884,950,366,393]" metricMagnitude="-1" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="120" unit="m" value="0.8">0.8 m</quantity>
depth. The stream at the
<typeStatus id="95668814FF84FFF0B2E264436A7ADC96" box="[1243,1291,367,393]" pageId="8" pageNumber="120">type</typeStatus>
locality was about
<quantity id="8D259B53FF84FFF0B6E564BE6E76DCB2" box="[220,263,402,429]" metricMagnitude="0" metricUnit="m" metricValue="8.0" pageId="8" pageNumber="120" unit="m" value="8.0">8 m</quantity>
wide with black water (pH 6.1), 23°C, and with sand and mud along the shores.
</paragraph>
</subSubSection>
<subSubSection id="02C7653DFF84FFF0B6FE649A6DECDCEB" pageId="8" pageNumber="120" type="etymology">
<paragraph id="4A6236B6FF84FFF0B6FE649A6DECDCEB" blockId="8.[151,1437,150,500]" pageId="8" pageNumber="120">
<emphasis id="78A9EAA4FF84FFF0B6FE649A6E3FDCCF" bold="true" box="[199,334,438,464]" pageId="8" pageNumber="120">Etymology.</emphasis>
The specific epithet honors Gustavo Orces, in recognition of his extensive contributions to the knowledge vertebrate biodiversity in
<collectingCountry id="32CA7626FF84FFF0B40364F66DE8DCEB" box="[570,665,474,500]" name="Ecuador" pageId="8" pageNumber="120">Ecuador</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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