treatments-xml/data/B0/71/87/B07187DFA821FFA9FED7F9DB9F33FAC6.xml

<document id="704D9308ECB5B1646E7FF43BCDA1D48B" ID-CLB-Dataset="153629" ID-DOI="10.18590/euscorpius.2016.vol2016.iss220.1" ID-GBIF-Dataset="03196918-10d5-44d2-ab42-d38f88314c7b" ID-ISSN="1536-9307" ID-Zenodo-Dep="7124466" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638249082081" checkinUser="felipe" docAuthor="Kovařík, František, Lowe, Graeme, Ranawana, Kithsiri B., Hoferek, David &amp; Š, V. A." docDate="2016" docId="B07187DFA821FFA9FED7F9DB9F33FAC6" docLanguage="en" docName="Euscorpius.220.1-133.pdf" docOrigin="Euscorpius 220" docSource="https://doi.org/10.18590/euscorpius.2016.vol2016.iss220.1" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Reddyanus Vachon 1972" docType="treatment" docVersion="7" lastPageNumber="56" masterDocId="4C48FFA7A817FF90FFC0FFD79E62FFBD" masterDocTitle="Scorpions of Sri Lanka (Scorpiones Buthidae, Chaerilidae, Scorpionidae) with description of four new species of the genera Charmus Karsch, 1879 and Reddyanus Vachon, 1972, stat n" masterLastPageNumber="133" masterPageNumber="1" pageNumber="53" updateTime="1699471154095" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
<mods:mods id="399CC7FB854D40E80A8A5ABB1856FFC2" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E064CC75251A36C2787891D909E30452">
<mods:title id="5FFBBF638C24FD66959E86A893F4EBF6">Scorpions of Sri Lanka (Scorpiones Buthidae, Chaerilidae, Scorpionidae) with description of four new species of the genera Charmus Karsch, 1879 and Reddyanus Vachon, 1972, stat n</mods:title>
</mods:titleInfo>
<mods:name id="EEE106E571097A4B8337C19E4706BD82" type="personal">
<mods:role id="1B644C90D6A8B48D1097B0834C8867F9">
<mods:roleTerm id="0AB8D1E0FC622CC0B27821A2AE4FE111">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="20401C641993C036A7220F8C531C89EF">Kovařík, František</mods:namePart>
</mods:name>
<mods:name id="8911928BD5074D1741B8C9D39C27BC29" type="personal">
<mods:role id="E5EE231EF184CFF7B8E1DC825BDBD847">
<mods:roleTerm id="94377AD8F8B65ED33F94D547F23A4E66">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E49E9C93C614389CA0D22A94B79BE0B0">Lowe, Graeme</mods:namePart>
</mods:name>
<mods:name id="E499EA0CB3911BC9117D161882A9CEC0" type="personal">
<mods:role id="3DCECC37D223397F6DA1296D02EBB03E">
<mods:roleTerm id="15063392087B665BA0E1C0D6DD32842D">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C06A1A042A789CE28BDB3D55F3A7E1C8">Ranawana, Kithsiri B.</mods:namePart>
</mods:name>
<mods:name id="A2F909291C31812852AD1C95076D46F0" type="personal">
<mods:role id="79956928DB9B5837A3019A5CEE093771">
<mods:roleTerm id="EC120560DC28D1312A358482E8FB3FAA">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4313C59477F1115C268B4FA54A44F46D">Hoferek, David</mods:namePart>
</mods:name>
<mods:name id="B9B7136A5A30AF6CB883481774C5B90E" type="personal">
<mods:role id="0055C554AF2AADD05BBEEFBE73A66BF9">
<mods:roleTerm id="2BCDB14595DA0ED4B90736DFD83F1956">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7B2B3AC3C7E150D6D575430F4A337A81">Š, V. A.</mods:namePart>
</mods:name>
<mods:typeOfResource id="13A34936ACA2FBE57CADE4CA39A55866">text</mods:typeOfResource>
<mods:relatedItem id="111F148E148B6D8BA6C6B15A6FCDDE04" type="host">
<mods:titleInfo id="0FF41F95A9E13A918453F41DCE4787A6">
<mods:title id="A267888EBCAAD1FBCD8D4F5169E2A2A0">Euscorpius</mods:title>
</mods:titleInfo>
<mods:part id="FB1791F26F7B87A6EBACF1F051D4408A">
<mods:date id="7389E6727341D01C95DEBD43C091185C">2016</mods:date>
<mods:detail id="7C4FA48A705942AB857A924127A70ED8" type="pubDate">
<mods:number id="F29CAF46F389E9C5F75143D584620725">2016-12-31</mods:number>
</mods:detail>
<mods:detail id="CD3C1B1535A2852ECA3A589FAF1E2B97" type="volume">
<mods:number id="912ECEC05CF0A4972D43DBD8E37C15F4">220</mods:number>
</mods:detail>
<mods:extent id="047599E1E7B8EC89FA26FA6193F01DBA" unit="page">
<mods:start id="23CB1370C75C6ADD0AE60E2BB7589A39">1</mods:start>
<mods:end id="76478C4753FE9B1613AA07792389157E">133</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="783142B6A8AB9E59CC4BD8B83407446C">
<mods:url id="FC8ECD73285E35DE7F2CA6D9F0671777">https://doi.org/10.18590/euscorpius.2016.vol2016.iss220.1</mods:url>
</mods:location>
<mods:classification id="131B6599E47E420F653B058121ED5EDB">journal article</mods:classification>
<mods:identifier id="36BF507A5D90E3B7914973684C824C55" type="CLB-Dataset">153629</mods:identifier>
<mods:identifier id="46BD2B689F46DFF68F07A0C04B0A93A4" type="DOI">10.18590/euscorpius.2016.vol2016.iss220.1</mods:identifier>
<mods:identifier id="131044CE092D92D4446AC68C63B10A5E" type="GBIF-Dataset">03196918-10d5-44d2-ab42-d38f88314c7b</mods:identifier>
<mods:identifier id="12DB7F342D948FAD162D7B2104C26EEA" type="ISSN">1536-9307</mods:identifier>
<mods:identifier id="31C07400107E09D3CAC2FD52FF146D99" type="Zenodo-Dep">7124466</mods:identifier>
<mods:identifier id="7F89AD1CB8AB431B28E9E505FF2B123B" type="ZooBank">DD0DF45D-F63A-4AA2-8EFF-03CF99E297EF</mods:identifier>
</mods:mods>
<treatment id="B07187DFA821FFA9FED7F9DB9F33FAC6" ID-DOI="http://doi.org/10.5281/zenodo.7124533" ID-GBIF-Taxon="201134857" ID-Zenodo-Dep="7124533" LSID="urn:lsid:plazi:treatment:B07187DFA821FFA9FED7F9DB9F33FAC6" httpUri="http://treatment.plazi.org/id/B07187DFA821FFA9FED7F9DB9F33FAC6" lastPageId="57" lastPageNumber="56" pageId="54" pageNumber="53">
<subSubSection id="70C26542A821FFA6FED7F9DB9CA9F99E" box="[279,715,1547,1572]" pageId="54" pageNumber="53" type="nomenclature">
<paragraph id="386736C9A821FFA6FED7F9DB9CA9F99E" blockId="54.[206,787,1547,1633]" box="[279,715,1547,1572]" pageId="54" pageNumber="53">
Genus 
<taxonomicName id="FFD84D4AA821FFA6FEA2F9DC9C1AF99E" ID-CoL="77FW" authority="Vachon, 1972" authorityName="Vachon" authorityYear="1972" box="[354,632,1547,1571]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA821FFA6FEA2F9DC9FB9F99E" bold="true" box="[354,475,1547,1571]" italics="true" pageId="54" pageNumber="53">Reddyanus</emphasis>
<bibRefCitation id="5C494B38A821FFA6FE22F9DB9C1AF99E" author="Vachon" box="[482,632,1548,1571]" firstAuthor="Vachon" journalOrPublisher="Cahiers Pacifique" pageId="54" pageNumber="53" pagination="169 - 180" part="16" refId="ref45078" refString="VACHON, M. 1972. Remarques sur les scorpions appartenant au genre Isometrus H. et E. (Buthidae) a propos de l`espece Isometrus maculatus (Geer) habitant Ile de Paques. Cahiers Pacifique, 16: 169 - 180." title="Remarques sur les scorpions appartenant au genre Isometrus H. et E. (Buthidae) a propos de l`espece Isometrus maculatus (Geer) habitant Ile de Paques" type="journal article" year="1972">Vachon, 1972</bibRefCitation>
</taxonomicName>
<emphasis id="0AACEADBA821FFA6FDBEF9DB9CC5F999" bold="true" box="[638,679,1548,1572]" pageId="54" pageNumber="53">stat</emphasis>
. 
<emphasis id="0AACEADBA821FFA6FD75F9DB9CA6F999" bold="true" box="[693,708,1548,1572]" pageId="54" pageNumber="53">n</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="70C26542A821FFA6FF0EF9FC9CB2F9DC" pageId="54" pageNumber="53" type="description">
<paragraph id="386736C9A821FFA6FF0EF9FC9CB2F9DC" blockId="54.[206,787,1547,1633]" pageId="54" pageNumber="53">
(
<figureCitation id="A0E32A4CA821FFA6FF16F9FC9F4DF9FF" box="[214,303,1579,1602]" captionStart="Figures 12–15" captionStartId="6.[192,278,1885,1909]" captionTargetBox="[191,1440,167,1856]" captionTargetId="figure-11@6.[192,1439,168,1855]" captionTargetPageId="6" captionText="Figures 12–15: Maps showing the known geographical distribution of the Sri Lankan species." figureDoi="http://doi.org/10.5281/zenodo.7124476" httpUri="https://zenodo.org/record/7124476/files/figure.png" pageId="54" pageNumber="53">Figs. 14</figureCitation>
, 
<figureCitation id="A0E32A4CA821FFA6FEFAF9FC9F06F9FF" box="[314,356,1579,1602]" captionStart="Figures 193–200" captionStartId="48.[192,278,1070,1094]" captionTargetBox="[213,1446,171,1037]" captionTargetId="figure-11@48.[205,1452,168,1041]" captionTargetPageId="48" captionText="Figures 193–200: Distal segments of leg IV, retroventral view of Sri Lankan Buthidae and Chaerilidae genera. Buthoscorpio sarasinorum, male from locality 15CF (193), Charmus laneus, male from locality 15CO (194), Hottentotta tamulus, male from locality 15CK (195), Isometrus maculatus, female from locality 15CI (196), Isometrus thwaitesi, female from locality 15CO (197), Lychas srilankensis, male from locality 15CJ (198), Reddyanus basilicus, male from locality 15CS (199), and Chaerilus ceylonensis, male from locality 15CD (200)." figureDoi="http://doi.org/10.5281/zenodo.7124608" httpUri="https://zenodo.org/record/7124608/files/figure.png" pageId="54" pageNumber="53">199</figureCitation>
, 
<figureCitation id="A0E32A4CA821FFA6FEAFF9FC9FADF9FF" box="[367,463,1579,1602]" captionStart-0="Figures 201–208" captionStart-1="Figures 209–224" captionStartId-0="49.[192,278,1057,1081]" captionStartId-1="50.[192,278,1736,1760]" captionTargetBox-0="[204,1438,173,1022]" captionTargetBox-1="[208,1434,174,1703]" captionTargetId-0="figure-6@49.[192,1439,168,1028]" captionTargetId-1="figure-11@50.[205,1452,168,1707]" captionTargetPageId-0="49" captionTargetPageId-1="50" captionText-0="Figures 201–208: Distal segments of legs, retroventral view of Sri Lankan Reddyanus. Figures 201–204. R. ceylonensis sp. n., male holotype from locality 15CI, legs I–IV. Figures 205–208. Leg IV. R. besucheti, male from locality 15CD (205), R. jayarathnei sp. n., male paratype (206), R. loebli, male from locality 15CG (207), and R. ranawanai sp. n., male holotype (208)." captionText-1="Figures 209–224: Metasomal segments II–III, lateral view of Sri Lankan Reddyanus and Isometrus species. Figures 209–210. Reddyanus basilicus, male (209) and female (210) from locality 15CR. Figures 211–212. R. besucheti, male holotype (211) and female from locality 15CG (212). Figures 213–214. R. ceylonensis sp. n., male holotype (213) and female paratype from locality 15CI (214). Figures 215–216. R. jayarathnei sp. n., male (215) and female (216) paratypes. Figures 217–218. R. loebli, male (217) and female (218) from locality 15CG. Figures 219–220. R. ranawanai sp. n., male holotype (219) and female paratype (220). Figures 221–222. Isometrus maculatus, male from locality 15CP (221) and female from locality 15CI (222). Figures 223– 224. I. thwaitesi, male from locality 15CH (223) and female from locality 15CO (224)." figureDoi-0="http://doi.org/10.5281/zenodo.7124612" figureDoi-1="http://doi.org/10.5281/zenodo.7124622" httpUri-0="https://zenodo.org/record/7124612/files/figure.png" httpUri-1="https://zenodo.org/record/7124622/files/figure.png" pageId="54" pageNumber="53">201–220</figureCitation>
, 
<figureCitation id="A0E32A4CA821FFA6FE1AF9FC9C58F9FF" box="[474,570,1579,1602]" captionStart="Figures 225–240" captionStartId="51.[192,278,1493,1517]" captionTargetBox="[193,1432,180,1451]" captionTargetId="figure-6@51.[192,1439,179,1464]" captionTargetPageId="51" captionText="Figures 225–240: Pedipalp, dorsal view of Sri Lankan Reddyanus and Isometrus species. Figures 225–226. Reddyanus basilicus, male (225) and female (226) from locality 15CR. Figures 227–228. R. besucheti, male from locality 15CD (227) and female from locality 15CG (228). Figures 229–230. R. ceylonensis sp. n., male holotype (229) and female paratype from locality 15CI (230). Figures 231–232. R. jayarathnei sp. n., male (231) and female (232) paratypes. Figures 233–234. R. loebli, male (233) and female (234) from locality 15CG. Figures 235–236. R. ranawanai sp. n., male holotype (235) and female paratype (236). Figures 237–238. Isometrus maculatus, male from locality 15CP (237) and female from locality 15CI (238). Figures 239–240. I. thwaitesi, male from locality 15CH (239) and female from locality 15CO (240)." figureDoi="http://doi.org/10.5281/zenodo.7124624" httpUri="https://zenodo.org/record/7124624/files/figure.png" pageId="54" pageNumber="53">225–236</figureCitation>
, 
<figureCitation id="A0E32A4CA821FFA6FD85F9FC9D6DF9FF" box="[581,783,1579,1602]" captionStart-0="Figures 241–259" captionStart-1="Figures 260–263" captionStart-10="Figures 307–312" captionStart-11="Figures 313–318" captionStart-12="Figures 319–326" captionStart-13="Figures 327–330" captionStart-14="Figures 331–333" captionStart-15="Figures 334–337" captionStart-16="Figures 338–348" captionStart-17="Figures 349–354" captionStart-18="Figures 355–357" captionStart-19="Figures 358–361" captionStart-2="Figures 264–269" captionStart-20="Figures 362–365" captionStart-21="Figures 366–371" captionStart-22="Figures 372–375" captionStart-23="Figures 376–377" captionStart-24="Figures 378–381" captionStart-25="Figures 382–386" captionStart-26="Figures 387–392" captionStart-27="Figures 393–400" captionStart-28="Figures 401–402" captionStart-3="Figures 270–275" captionStart-4="Figures 276–282" captionStart-5="Figures 283–286" captionStart-6="Figures 287–290" captionStart-7="Figures 295–300" captionStart-8="Figures 301–302" captionStart-9="Figures 303–306" captionStartId-0="52.[192,278,1733,1757]" captionStartId-1="53.[192,278,1632,1656]" captionStartId-10="64.[192,278,1655,1679]" captionStartId-11="66.[192,278,1804,1828]" captionStartId-12="67.[192,278,865,889]" captionStartId-13="68.[192,278,1757,1781]" captionStartId-14="69.[192,278,1493,1517]" captionStartId-15="70.[192,278,1786,1810]" captionStartId-16="71.[192,278,1729,1753]" captionStartId-17="72.[192,278,1600,1624]" captionStartId-18="73.[192,278,1872,1896]" captionStartId-19="74.[192,278,1612,1636]" captionStartId-2="55.[192,278,1408,1432]" captionStartId-20="75.[192,278,1839,1863]" captionStartId-21="76.[192,278,1840,1864]" captionStartId-22="77.[192,278,1141,1165]" captionStartId-23="78.[192,278,1882,1906]" captionStartId-24="79.[192,278,1520,1544]" captionStartId-25="80.[192,278,1371,1395]" captionStartId-26="81.[192,278,1587,1611]" captionStartId-27="82.[192,278,965,989]" captionStartId-28="83.[192,278,1888,1912]" captionStartId-3="56.[192,278,1839,1863]" captionStartId-4="57.[192,278,892,916]" captionStartId-5="58.[192,278,1625,1649]" captionStartId-6="59.[192,278,1867,1891]" captionStartId-7="61.[192,278,1771,1795]" captionStartId-8="62.[192,278,1536,1560]" captionStartId-9="63.[192,278,1848,1872]" captionTargetBox-0="[238,1387,153,1699]" captionTargetBox-1="[196,1427,148,1596]" captionTargetBox-10="[191,1440,191,1626]" captionTargetBox-11="[207,1424,220,1766]" captionTargetBox-12="[194,1429,176,822]" captionTargetBox-13="[205,1429,203,1718]" captionTargetBox-14="[191,1440,175,1465]" captionTargetBox-15="[201,1437,224,1752]" captionTargetBox-16="[191,1440,191,1701]" captionTargetBox-17="[206,1421,175,1560]" captionTargetBox-18="[191,1440,167,1843]" captionTargetBox-19="[210,1425,149,1578]" captionTargetBox-2="[191,1440,167,1380]" captionTargetBox-20="[213,1417,167,1810]" captionTargetBox-21="[205,1406,172,1808]" captionTargetBox-22="[202,1423,152,1109]" captionTargetBox-23="[191,1440,167,1853]" captionTargetBox-24="[198,1432,149,1474]" captionTargetBox-25="[191,1440,191,1342]" captionTargetBox-26="[192,1399,172,1554]" captionTargetBox-27="[216,1425,181,915]" captionTargetBox-28="[191,1440,191,1859]" captionTargetBox-3="[215,1432,184,1819]" captionTargetBox-4="[198,1431,222,869]" captionTargetBox-5="[201,1430,178,1594]" captionTargetBox-6="[191,1440,167,1838]" captionTargetBox-7="[204,1426,228,1719]" captionTargetBox-8="[191,1440,167,1508]" captionTargetBox-9="[210,1426,227,1812]" captionTargetId-0="figure-11@52.[234,1392,152,1704]" captionTargetId-1="figure-62@53.[192,1439,144,1603]" captionTargetId-10="figure-249@64.[192,1439,192,1626]" captionTargetId-11="figure-11@66.[192,1439,216,1775]" captionTargetId-12="figure-6@67.[192,1439,168,836]" captionTargetId-13="figure-13@68.[192,1439,192,1728]" captionTargetId-14="figure-408@69.[192,1439,176,1464]" captionTargetId-15="figure-11@70.[192,1439,216,1757]" captionTargetId-16="figure-68@71.[192,1439,192,1700]" captionTargetId-17="figure-11@72.[192,1439,168,1571]" captionTargetId-18="figure-59@73.[192,1439,168,1842]" captionTargetId-19="figure-11@74.[192,1439,144,1583]" captionTargetId-2="figure-377@55.[192,1439,168,1379]" captionTargetId-20="figure-19@75.[214,1416,168,1809]" captionTargetId-21="figure-11@76.[180,1427,168,1812]" captionTargetId-22="figure-903@77.[192,1439,144,1112]" captionTargetId-23="figure-25@78.[192,1439,168,1852]" captionTargetId-24="figure-6@79.[192,1439,144,1491]" captionTargetId-25="figure-654@80.[192,1439,192,1341]" captionTargetId-26="figure-6@81.[180,1427,168,1558]" captionTargetId-27="figure-11@82.[192,1439,168,936]" captionTargetId-28="figure-6@83.[192,1439,192,1859]" captionTargetId-3="figure-11@56.[192,1439,173,1824]" captionTargetId-4="figure-6@57.[192,1439,199,888]" captionTargetId-5="figure-11@58.[192,1439,168,1596]" captionTargetId-6="figure-47@59.[192,1439,168,1837]" captionTargetId-7="figure-81@61.[192,1439,215,1728]" captionTargetId-8="figure-417@62.[192,1439,168,1507]" captionTargetId-9="figure-63@63.[192,1439,216,1819]" captionTargetPageId-0="52" captionTargetPageId-1="53" captionTargetPageId-10="64" captionTargetPageId-11="66" captionTargetPageId-12="67" captionTargetPageId-13="68" captionTargetPageId-14="69" captionTargetPageId-15="70" captionTargetPageId-16="71" captionTargetPageId-17="72" captionTargetPageId-18="73" captionTargetPageId-19="74" captionTargetPageId-2="55" captionTargetPageId-20="75" captionTargetPageId-21="76" captionTargetPageId-22="77" captionTargetPageId-23="78" captionTargetPageId-24="79" captionTargetPageId-25="80" captionTargetPageId-26="81" captionTargetPageId-27="82" captionTargetPageId-28="83" captionTargetPageId-3="56" captionTargetPageId-4="57" captionTargetPageId-5="58" captionTargetPageId-6="59" captionTargetPageId-7="61" captionTargetPageId-8="62" captionTargetPageId-9="63" captionText-0="Figures 241–259: Pedipalp chela dorsal (241), movable finger (242–250), and fixed finger (251–259) of Sri Lankan Isometrus, Lychas, and Reddyanus species. Figures 241–242, 252. Isometrus maculatus, female from locality 15CO (241) and male from locality 15CP (242, 252). Figures 243–244, 253. I. thwaitesi, male from locality 15CH (243) and female from locality 15CO (244, 253). Figures 245, 254. Reddyanus basilicus, male from locality 15CS. Figures 246, 255. R. besucheti, male from locality 15CD. Figures 247, 256. R. ceylonensis sp. n., male holotype. Figures 248, 257. R. jayarathnei sp. n., male paratype. Figures 249, 258. R. loebli, male from locality 15CG. Figures 250, 259. R. ranawanai sp. n., male holotype. Figure 251. Lychas srilankensis, male from locality 15CO." captionText-1="Figures 260–263: Reddyanus basilicus from locality 15CS. Figures 260–261. Male in dorsal (260) and ventral (261) views. Figures 262–263. Female in dorsal (262) and ventral (263) views. Scale bar: 10 mm." captionText-10="Figures 307–312: Reddyanus ceylonensis sp. n. from locality 15CI. Figures 307, 309, 311–312. Male holotype, chelicerae, carapace and tergites I–III (307), sternopectinal region and sternites III–IV (309), and chelicera in dorsal (311) and ventral (312) views. Figures 308, 310. Female paratype, chelicerae, carapace and tergites I–III (308), and sternopectinal region and sternites III–IV (310)." captionText-11="Figures 313–318: Reddyanus ceylonensis sp. n. from locality 15CI. Figures 313–315. Male holotype, metasoma and telson, lateral (313), ventral (314), and dorsal (315) views. Figures 316–318. Female paratype, metasoma and telson, lateral (316), ventral (317), and dorsal (318) views." captionText-12="Figures 319–326: Reddyanus ceylonensis sp. n. from locality 15CI. Figures 319. Female partatype, pedipalp chela, dorsal view. Figures 320–326. Male holotype. Pedipalp chela, dorsal (320), external (321) and ventral (322) views. Pedipalp patella, dorsal (323) and external (324) views. Pedipalp femur and trochanter internal (325) and dorsal (326) views. The trichobothrial pattern is indicated in Figures 320–326." captionText-13="Figures 327–330: Left hemispermatophore of Reddyanus ceylonensis sp. n. male paratype from locality 15CI. External (327), dorsal (328), internal (329) and ventral (330) views. Extracted after 4th ecdysis. A second hemispermatophore examined from a second male after 5th ecdysis was similar. Scale bar: 250 µm. Abbreviations: bl, basal lobe; f, flagellum; ml, median lobe; mlc, median lobe carina; t, trunk." captionText-14="Figures 331–333: Reddyanus ceylonensis sp. n., female from locality 15CH (331), female from locality 15CI with newborns before first ecdysis (332), and male from locality 15CI (333)." captionText-15="Figures 334–337: Reddyanus jayarathnei sp. n., paratypes. Figures 334–335. Male in dorsal (334) and ventral (335) views. Figures 336–337. Female in dorsal (336) and ventral (337) views. Scale bar: 10 mm." captionText-16="Figures 338–348: Reddyanus jayarathnei sp. n., paratypes. Figures 338, 340, 343–348. Male, chelicerae, carapace and tergites I–III (338), sternopectinal region (340). Pedipalp chela, dorsal (343), external (344) and ventral (345) views. Pedipalp patella, dorsal (346) and external (347) views. Pedipalp femur and trochanter dorsal (348) view. The trichobothrial pattern is indicated in Figures 344–348. Figures 339, 341–342. Female, chelicerae, carapace and tergites I–III (339), sternopectinal region and sternite III (341), and pedipalp chela dorsal view (342)." captionText-17="Figures 349–354: Reddyanus jayarathnei sp. n., paratypes. Figures 349–351. Male, metasoma and telson, lateral (349), ventral (350), and dorsal (351) views. Figures 352–354. Female, metasoma and telson, lateral (352), ventral (353), and dorsal (354) views. Scale bar: 10 mm." captionText-18="Figures 355–357: Reddyanus jayarathnei sp. n., female paratype (355) and male holotype (356) at the type locality (357), Galle district, Kanneliya Rain Forest, 06°15'04&quot; N 80°20'18&quot; E." captionText-19="Figures 358–361: Reddyanus loebli from locality 15CG. Figures 358–359. Male in dorsal (358) and ventral (359) views. Figures 360–361. Female in dorsal (360) and ventral (361) views." captionText-2="Figures 264–269: Reddyanus basilicus. Figures 264, 266. Male from locality 15CS, chelicerae, carapace and tergites I–III (264), and sternopectinal region and sternite III (266). Figures 265, 267. Female from locality 15CS, chelicerae, carapace and tergites I–III (265) and sternopectinal region and sternites III–IV (267). Figures 268–269. Male from locality 15CR, chelicera dorsal (268) and ventral (269) aspects." captionText-20="Figures 362–365: Reddyanus loebli from locality 15CG. Figures 362, 364. Male, chelicerae, carapace and tergites I–III (362), sternopectinal region and sternite III (364). Figures 363, 365. Female, chelicerae, carapace and tergites I–III (363), and sternopectinal region (365)." captionText-21="Figures 366–371: Reddyanus loebli from locality 15CG. Figures 366–368. Male, metasoma and telson, lateral (366), ventral (367), and dorsal (368) views. Figures 369–371. Female, metasoma and telson, lateral (369), ventral (370), and dorsal (371) views." captionText-22="Figures 372–375: Distal trunk, capsule region and flagellum of left hemispermatophore of Reddyanus loebli from locality 15CH. External (372), dorsal (373), internal (374) and ventral (375) views. Extracted after 5th ecdysis. Note: this hemispermatopore was folded along a dorsal line and flattened; dorsal and ventral views show it as unfolded and flat, hence the broad appearance of the trunk. Compare with Figs. 283–286 and 327–330 which show typical Reddyanus hemispermatophores with trunk in natural curved conformation. Scale bar: 500 µm. Abbreviations: bl, basal lobe; f, flagellum; ml, median lobe; mlc, median lobe carina; t, trunk." captionText-23="Figures 376–377: Reddyanus loebli from locality 15CH, male (376) and female with newborns (377)." captionText-24="Figures 378–381: Reddyanus ranawanai sp. n. Figures 378–379. Male holotype in dorsal (378) and ventral (379) views. Figures 380–381. Female paratype in dorsal (380) and ventral (381) views." captionText-25="Figures 382–386: Reddyanus ranawanai sp. n. Figures 382, 384–385. Male holotype, chelicerae, carapace and tergites I–III (382), sternites IV–VII (384), and sternopectinal region (385). Figures 383, 386. Female paratype, chelicerae, carapace and tergites I–III (383), and sternopectinal region with sternite III (386)." captionText-26="Figures 387–392: Reddyanus ranawanai sp. n. Figures 387–389. Male holotype, metasoma and telson, lateral (387), ventral (388), and dorsal (389) views. Figures 390–392. Female paratype, metasoma and telson, lateral (390), ventral (391), and dorsal (392) views. Scale bar: 10 mm." captionText-27="Figures 393–400: Reddyanus ranawanai sp. n. Figure 393. Female paratype, pedipalp chela dorsal. Figures 394–400. Male holotype. Pedipalp chela, dorsal (394), external (395) and ventral (396) views. Pedipalp patella, dorsal (397) and external (398) views. Pedipalp femur internal (399) and trochanter and femur dorsal (400) views. The trichobothrial pattern is indicated in Figures 395–400." captionText-28="Figures 401–402: Reddyanus ranawanai sp. n., female paratype (401) and male holotype (402)." captionText-3="Figures 270–275: Reddyanus basilicus from locality 15CS. Figures 270–272. Male, metasoma and telson, lateral (270), ventral (271), and dorsal (272) views. Figures 273–275. Female, metasoma and telson, lateral (273), ventral (274), and dorsal (275) views. Scale bar: 10 mm." captionText-4="Figures 276–282: Reddyanus basilicus from locality 15CS. Figures 276–277. Female. Pedipalp chela, dorsal (276) and external (277) views. Figures 278–282. Male. Pedipalp chela, dorsal (278) and external (279) views. Pedipalp patella, dorsal (280) and external (281) views. Pedipalp femur and trochanter dorsal view (282). The trichobothrial pattern is indicated in Figures 279–282." captionText-5="Figures 283–286: Left hemispermatophore of Reddyanus basilicus from locality 15CR. External (283), dorsal (284), internal (285) and ventral (286) views. Scale bar: 500 µm. Abbreviations: bl, basal lobe; f, flagellum; ml, median lobe; mlc, median lobe carina; t, trunk." captionText-6="Figures 287–290: Reddyanus basilicus from locality 15CS, female (287), female with newborns before first ecdysis (288), female with juveniles after first ecdysis (289), and male (290)." captionText-7="Figures 295–300: Reddyanus besucheti. Figures 295–297. Male from locality 15CD, metasoma and telson, lateral (295), ventral (296), and dorsal (297) views. Figures 298–300. Female from locality 15CG, metasoma and telson, lateral (298), ventral (299), and dorsal (300) views." captionText-8="Figures 301–302: Reddyanus besucheti, male at locality 15CF (301) and female at locality 15CG (302)." captionText-9="Figures 303–306: Reddyanus ceylonensis sp. n. from locality 15CI. Figures 303–304. Male holotype in dorsal (303) and ventral (304) views. Figures 305–306. Female paratype in dorsal (305) and ventral (306) views." figureDoi-0="http://doi.org/10.5281/zenodo.7124626" figureDoi-1="http://doi.org/10.5281/zenodo.7124628" figureDoi-10="http://doi.org/10.5281/zenodo.7124649" figureDoi-11="http://doi.org/10.5281/zenodo.7124651" figureDoi-12="http://doi.org/10.5281/zenodo.7124655" figureDoi-13="http://doi.org/10.5281/zenodo.7124657" figureDoi-14="http://doi.org/10.5281/zenodo.7124659" figureDoi-15="http://doi.org/10.5281/zenodo.7124661" figureDoi-16="http://doi.org/10.5281/zenodo.7124663" figureDoi-17="http://doi.org/10.5281/zenodo.7124665" figureDoi-18="http://doi.org/10.5281/zenodo.7124668" figureDoi-19="http://doi.org/10.5281/zenodo.7124670" figureDoi-2="http://doi.org/10.5281/zenodo.7124630" figureDoi-20="http://doi.org/10.5281/zenodo.7124672" figureDoi-21="http://doi.org/10.5281/zenodo.7124674" figureDoi-22="http://doi.org/10.5281/zenodo.7124676" figureDoi-23="http://doi.org/10.5281/zenodo.7124678" figureDoi-24="http://doi.org/10.5281/zenodo.7124680" figureDoi-25="http://doi.org/10.5281/zenodo.7124684" figureDoi-26="http://doi.org/10.5281/zenodo.7124686" figureDoi-27="http://doi.org/10.5281/zenodo.7124690" figureDoi-28="http://doi.org/10.5281/zenodo.7124692" figureDoi-3="http://doi.org/10.5281/zenodo.7124632" figureDoi-4="http://doi.org/10.5281/zenodo.7124634" figureDoi-5="http://doi.org/10.5281/zenodo.7124636" figureDoi-6="http://doi.org/10.5281/zenodo.7124638" figureDoi-7="http://doi.org/10.5281/zenodo.7124642" figureDoi-8="http://doi.org/10.5281/zenodo.7124645" figureDoi-9="http://doi.org/10.5281/zenodo.7124647" httpUri-0="https://zenodo.org/record/7124626/files/figure.png" httpUri-1="https://zenodo.org/record/7124628/files/figure.png" httpUri-10="https://zenodo.org/record/7124649/files/figure.png" httpUri-11="https://zenodo.org/record/7124651/files/figure.png" httpUri-12="https://zenodo.org/record/7124655/files/figure.png" httpUri-13="https://zenodo.org/record/7124657/files/figure.png" httpUri-14="https://zenodo.org/record/7124659/files/figure.png" httpUri-15="https://zenodo.org/record/7124661/files/figure.png" httpUri-16="https://zenodo.org/record/7124663/files/figure.png" httpUri-17="https://zenodo.org/record/7124665/files/figure.png" httpUri-18="https://zenodo.org/record/7124668/files/figure.png" httpUri-19="https://zenodo.org/record/7124670/files/figure.png" httpUri-2="https://zenodo.org/record/7124630/files/figure.png" httpUri-20="https://zenodo.org/record/7124672/files/figure.png" httpUri-21="https://zenodo.org/record/7124674/files/figure.png" httpUri-22="https://zenodo.org/record/7124676/files/figure.png" httpUri-23="https://zenodo.org/record/7124678/files/figure.png" httpUri-24="https://zenodo.org/record/7124680/files/figure.png" httpUri-25="https://zenodo.org/record/7124684/files/figure.png" httpUri-26="https://zenodo.org/record/7124686/files/figure.png" httpUri-27="https://zenodo.org/record/7124690/files/figure.png" httpUri-28="https://zenodo.org/record/7124692/files/figure.png" httpUri-3="https://zenodo.org/record/7124632/files/figure.png" httpUri-4="https://zenodo.org/record/7124634/files/figure.png" httpUri-5="https://zenodo.org/record/7124636/files/figure.png" httpUri-6="https://zenodo.org/record/7124638/files/figure.png" httpUri-7="https://zenodo.org/record/7124642/files/figure.png" httpUri-8="https://zenodo.org/record/7124645/files/figure.png" httpUri-9="https://zenodo.org/record/7124647/files/figure.png" pageId="54" pageNumber="53">245–250, 254–402</figureCitation>
, 
<figureCitation id="A0E32A4CA821FFA6FED1F99D9F13F9DC" box="[273,369,1610,1633]" captionStart="Figures 403–429" captionStartId="85.[192,278,744,768]" captionTargetId="figure-11@84.[172,1464,185,1878]" captionTargetPageId="84" captionText="Figures 403–429: Telson, lateral view of Sri Lankan Buthidae and Chaerilidae species. Figures 403–404. Isometrus maculatus, male from locality 15CP (403) and female from locality 15CI (404). Figures 405–406. I. thwaitesi, male from locality 15CH (405) and female from locality 15CO (406). Figures 407–408. Lychas srilankensis, male (407) and female (408) from locality 15CJ. Figures 409–410. Reddyanus basilicus, male (409) and female (410) from locality 15CR. Figures 411–412. R. besucheti, male holotype (411) and female from locality 15CG (412). Figures 413–414. R. ceylonensis sp. n., male holotype (413) and female paratype (414) from locality 15CI. Figures 415–416. R. jayarathnei sp. n., male (415) and female (416) paratypes. Figures 417–418. R. loebli, male (417) and female (418) from locality 15CG. Figures 419–420. R. ranawanai sp. n., male holotype (419) and female paratype (420). Figures 421–422: Buthoscorpio sarasinorum, male from locality 15CF (421) and female from locality 15CG (422). Figures 423–424. Charmus laneus, male (423) and female (424) from locality 15CO. Figures 425–426. Charmus saradieli sp. n. male paratype (425) (MHNG) and female holotype (426). Figurs 427. Hottentotta tamulus, male from locality 15CK (427). Figures 428–429. Chaerilus ceylonensis, male (428) and female (429) from locality 15CD." figureDoi="http://doi.org/10.5281/zenodo.7124696" httpUri="https://zenodo.org/record/7124696/files/figure.png" pageId="54" pageNumber="53">409–420</figureCitation>
, 
<figureCitation id="A0E32A4CA821FFA6FEBCF99D9FBEF9DC" box="[380,476,1610,1633]" captionStart="Figures 547–554" captionStartId="114.[192,278,810,834]" captionTargetBox="[191,1440,191,782]" captionTargetId="figure-538@114.[192,1440,192,781]" captionTargetPageId="114" captionText="Figures 547–554: The mitotic metaphases of buthid males from Sri Lanka. Buthoscorpio sarasinorum from locality 15CF (2n=14) (547), Charmus laneus from locality 15CO (2n=9) (548), Isometrus thwaitesi from locality 15CF (2n=8) (549), Lychas srilankensis from locality 15CN (2n=16) (550), Reddyanus basilicus from locality 15CS (2n=16) (551), Reddyanus basilicus from locality 15CR (2n=15) (552), Reddyanus ceylonensis from locality 15CI (2n=16) (553), Reddyanus loebli from locality 15CH (2n=17) (554). Arrowheads indicate extra large odd chromosomes. Scale bar = 5 µm for 547–554." figureDoi="http://doi.org/10.5281/zenodo.7124765" httpUri="https://zenodo.org/record/7124765/files/figure.png" pageId="54" pageNumber="53">551–554</figureCitation>
, 
<figureCitation id="A0E32A4CA821FFA6FE27F99D9C25F9DC" box="[487,583,1610,1633]" captionStart="Figures 555–570" captionStartId="115.[192,278,1733,1757]" captionTargetBox="[233,1393,170,1705]" captionTargetId="figure-151@115.[234,1392,171,1704]" captionTargetPageId="115" captionText="Figures 555–570: Key characters for distinguishing males of Sri Lankan Isometrus and Reddyanus. Figures 555–556, 565– 566. Pedipalp chela dorsal view, I. maculatus from locality 15CP (555), I. thwaitesi from locality 15CH (556), R. basilicus, holotype (565), and R. besucheti, holotype (566). Figures 557–558. Sternopectinal area, I. maculatus from locality 15CP (557), and I. thwaitesi from locality 15CH (558). Figures 559–560, 567–570. Sternite V, I. maculatus from locality 15CP (559), I. thwaitesi from locality 15CH (560), R. besucheti, from locality 15CF (567), R. ceylonensis sp. n., holotype (568), R. jayarathnei sp. n., paratype (569), and R. ranawanai sp. n., paratype (570). Figures 561–564. Metasomal segment II lateral (561–562) and pedipalp patella dorsal (563–564), R. loebli from locality 15CG (561, 563) and R. besucheti, from locality 15CF (562, 564)." figureDoi="http://doi.org/10.5281/zenodo.7124767" httpUri="https://zenodo.org/record/7124767/files/figure.png" pageId="54" pageNumber="53">561–570</figureCitation>
, 
<tableCitation id="755A0372A821FFA6FD91F99D9CAAF9DC" box="[593,712,1610,1633]" captionStart-0="Table 3" captionStart-1="Table 4" captionStart-2="Table 5" captionStartId-0="86.[192,257,777,801]" captionStartId-1="86.[192,257,1428,1452]" captionStartId-2="86.[192,257,1869,1893]" captionTargetPageId-0="86" captionTargetPageId-1="86" captionTargetPageId-2="86" captionText-0="Table 3: Comparative measurements of adults of Reddyanus basilicus, R. besucheti and R. ceylonensis sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." captionText-1="Table 4: Comparative measurements of adults of Reddyanus ranawanai sp. n. and R. jayarathnei sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to posterior width), depth (D)." captionText-2="Table 5: Comparison among Sri Lankan Reddyanus species (specimens), based upon selected morphometric ratios of adult males. Abbreviations: length (L), width (W), depth (D)." httpUri-0="http://table.plazi.org/id/6CA76641A841FFC6FF00FCDE9DD7FC84" httpUri-1="http://table.plazi.org/id/6CA76641A841FFC6FF00FA439D16FA79" httpUri-2="http://table.plazi.org/id/6CA76641A841FFC6FF00F89A9CBAF8C0" pageId="54" pageNumber="53" tableUuid-0="6CA76641A841FFC6FF00FCDE9DD7FC84" tableUuid-1="6CA76641A841FFC6FF00FA439D16FA79" tableUuid-2="6CA76641A841FFC6FF00F89A9CBAF8C0">Tables 3–5</tableCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="70C26542A821FFA6FF00F9509D63F8C8" pageId="54" pageNumber="53" type="reference_group">
<paragraph id="386736C9A821FFA6FF00F9509F97F88A" blockId="54.[192,802,1671,1909]" pageId="54" pageNumber="53">
<treatmentCitationGroup id="18C811E7A821FFA6FF00F9509F97F88A" pageId="54" pageNumber="53">
<taxonomicName id="FFD84D4AA821FFA6FF00F9509FDDF923" authority="" authorityName="Vachon" authorityYear="1972" box="[192,447,1671,1694]" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="subGenus" subGenus="Reddyanus">
<emphasis id="0AACEADBA821FFA6FF00F9509F4BF923" box="[192,297,1671,1694]" italics="true" pageId="54" pageNumber="53">Isometrus</emphasis>
(
<emphasis id="0AACEADBA821FFA6FEFFF9509FD6F923" box="[319,436,1671,1694]" italics="true" pageId="54" pageNumber="53">Reddyanus</emphasis>
)
</taxonomicName>
: 
<treatmentCitation id="B97910D8A821FFA6FE11F9509C80F923" author="VACHON, M." box="[465,738,1671,1694]" page="176" pageId="54" pageNumber="53" year="1972">
<bibRefCitation id="5C494B38A821FFA6FE11F9509C80F923" author="Vachon" box="[465,738,1671,1694]" firstAuthor="Vachon" journalOrPublisher="Cahiers Pacifique" page="176 - 177" pageId="54" pageNumber="53" pagination="169 - 180" part="16" refId="ref45078" refString="VACHON, M. 1972. Remarques sur les scorpions appartenant au genre Isometrus H. et E. (Buthidae) a propos de l`espece Isometrus maculatus (Geer) habitant Ile de Paques. Cahiers Pacifique, 16: 169 - 180." title="Remarques sur les scorpions appartenant au genre Isometrus H. et E. (Buthidae) a propos de l`espece Isometrus maculatus (Geer) habitant Ile de Paques" type="journal article" year="1972">Vachon, 1972: 176–177</bibRefCitation>
</treatmentCitation>
, figs. 14, 16, 18, 20; 
<treatmentCitation id="B97910D8A821FFA6FE6AF9719CCFF900" author="VACHON, M." box="[426,685,1702,1725]" page="90" pageId="54" pageNumber="53" year="1982">
<bibRefCitation id="5C494B38A821FFA6FE6AF9719CCFF900" author="Vachon" box="[426,685,1702,1725]" firstAuthor="Vachon" journalOrPublisher="Revue suisse de Zoologie" page="90 - 101" pageId="54" pageNumber="53" pagination="77 - 114" part="89" refId="ref45190" refString="VACHON, M. 1982. Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d`Histoire naturelle de Geneve III.). Revue suisse de Zoologie, 89: 77 - 114." title="Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d`Histoire naturelle de Geneve III.)" type="journal article" year="1982">Vachon, 1982: 90–101</bibRefCitation>
</treatmentCitation>
, 108–110, figs. 47–67, 88–91; 
<treatmentCitation id="B97910D8A821FFA6FE16F9139D7CF966" author="FET, V. &amp; W. D. SISSOM &amp; G. LOWE &amp; M. E. BRAUNWALDER" box="[470,798,1732,1755]" page="150" pageId="54" pageNumber="53" year="2000">
<bibRefCitation id="5C494B38A821FFA6FE16F9139D7CF966" author="Fet &amp; Lowe" bookContentInfo="689 pp." box="[470,798,1732,1755]" firstAuthor="Fet" journalOrPublisher="New York: The New York Entomological Society" page="150 - 154" pageId="54" pageNumber="53" refId="ref40873" refString="FET, V., W. D. SISSOM, G. LOWE &amp; M. E. BRAUNWALDER. 2000. Catalog of the Scorpions of the World (1758 - 1998). New York: The New York Entomological Society, 689 pp." title="Catalog of the Scorpions of the World (1758 - 1998)" type="book" year="2000">Fet &amp; Lowe, 2000: 150–154</bibRefCitation>
</treatmentCitation>
; 
<treatmentCitation id="B97910D8A821FFA6FF30F9349FB4F947" author="Kovarik" box="[240,470,1763,1786]" page="5 - 14" pageId="54" pageNumber="53" year="2003">
<bibRefCitation id="5C494B38A821FFA6FF30F9349FB4F947" author="Kovarik" box="[240,470,1763,1786]" firstAuthor="Kovarik" page="5 - 14" pageId="54" pageNumber="53" pagination="1 - 19" refId="ref41718" refString="KOVARIK, F. 2003. A review of the genus Isometrus Ehrenberg, 1828 (Scorpiones: Buthidae) with descriptions of four new species from Asia and Australia. Euscorpius, 10: 1 - 19." type="journal article" year="2003">Kovařík, 2003: 5–14</bibRefCitation>
</treatmentCitation>
, figs. 1–8; 
<treatmentCitation id="B97910D8A821FFA6FD97F9349F89F8A4" author="KOVARIK, F &amp; A. A. OJANGUREN AFFILASTRO" page="184" pageId="54" pageNumber="53" year="2013">
<bibRefCitation id="5C494B38A821FFA6FD97F9349F89F8A4" author="Kovarik &amp; Ojanguren" bookContentInfo="400 pp." firstAuthor="Kovarik" journalOrPublisher="Clairon Production, Prague" page="184 - 193" pageId="54" pageNumber="53" refId="ref42080" refString="KOVARIK, F &amp; A. A. OJANGUREN AFFILASTRO. 2013. Illustrated catalog of scorpions Part II. Bothriuridae; Chaerilidae; Buthidae I., genera Compsobuthus, Hottentotta, Isometrus, Lychas, and Sassanidotus. Clairon Production, Prague, 400 pp." title="Illustrated catalog of scorpions Part II. Bothriuridae; Chaerilidae; Buthidae I., genera Compsobuthus, Hottentotta, Isometrus, Lychas, and Sassanidotus" type="book" year="2013">Kovařík &amp; Ojanguren, 2013: 184–193</bibRefCitation>
</treatmentCitation>
, 347–354, 357–360, figs. 1265–1367, 1384–1409.
</treatmentCitationGroup>
</paragraph>
<paragraph id="386736C9A821FFA6FF00F8E89D63F8C8" blockId="54.[192,802,1671,1909]" pageId="54" pageNumber="53">
<treatmentCitationGroup id="18C811E7A821FFA6FF00F8E89D63F8C8" pageId="54" pageNumber="53">
<taxonomicName id="FFD84D4AA821FFA6FF00F8E89F4BF8EB" ID-CoL="55JF" authorityName="Ehrenberg" authorityYear="1828" box="[192,297,1855,1878]" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA821FFA6FF00F8E89F4BF8EB" box="[192,297,1855,1878]" italics="true" pageId="54" pageNumber="53">Isometrus</emphasis>
</taxonomicName>
(
<taxonomicName id="FFD84D4AA821FFA6FE84F8E89FD8F8EB" authorityName="Tikader &amp; Bastawade" authorityYear="1983" box="[324,442,1855,1878]" class="Arachnida" family="Buthidae" genus="Raddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA821FFA6FE84F8E89FD8F8EB" box="[324,442,1855,1878]" italics="true" pageId="54" pageNumber="53">Raddyanus</emphasis>
</taxonomicName>
[sic]): 
<treatmentCitation id="B97910D8A821FFA6FDDDF8E89FF3F8C8" author="TIKADER B. K. &amp; D. B. BASTAWADE" page="254" pageId="54" pageNumber="53" year="1983">
<bibRefCitation id="5C494B38A821FFA6FDDDF8E89FF3F8C8" author="Tikader &amp; Bastawade" bookContentInfo="668 pp." firstAuthor="Tikader" journalOrPublisher="Zoological Survey of India, Calcutta" page="254 - 311" pageId="54" pageNumber="53" refId="ref44789" refString="TIKADER B. K. &amp; D. B. BASTAWADE. 1983. The Fauna of India: Scorpions. Scorpionida, Arachnida. Vol III. Zoological Survey of India, Calcutta, 668 pp." title="The Fauna of India: Scorpions. Scorpionida, Arachnida. Vol III" type="book" year="1983">Tikader &amp; Bastawade, 1983: 254–311</bibRefCitation>
</treatmentCitation>
(in part), figs. 727–770, 841–895.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="70C26542A821FFA6FCFEFF109B06FF63" box="[830,1380,199,222]" pageId="54" pageNumber="53" type="type_taxon">
<paragraph id="386736C9A821FFA6FCFEFF109B06FF63" blockId="54.[830,1380,199,222]" box="[830,1380,199,222]" pageId="54" pageNumber="53">
<typeStatus id="E763886BA821FFA6FCFEFF109D15FF63" box="[830,887,199,222]" pageId="54" pageNumber="53">TYPE</typeStatus>
SPECIES. 
<taxonomicName id="FFD84D4AA821FFA6FC1DFF109B02FF63" authority="Pocock, 1899" authorityName="Pocock" authorityYear="1899" box="[989,1376,199,222]" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="acanthurus">
<emphasis id="0AACEADBA821FFA6FC1DFF109AA7FF63" box="[989,1221,199,222]" italics="true" pageId="54" pageNumber="53">Isometrus acanthurus</emphasis>
Pocock, 1899
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="70C26542A821FFA6FCFEFED39B73FC98" pageId="54" pageNumber="53" type="diagnosis">
<paragraph id="386736C9A821FFA6FCFEFED39B73FC98" blockId="54.[830,1441,260,805]" pageId="54" pageNumber="53">
DIAGNOSIS. Medium sized buthids, adults 
<quantity id="FF209B2CA821FFA6FAE1FED39BFFFEA6" box="[1313,1437,260,283]" metricMagnitude="-2" metricUnit="m" metricValue="4.7" metricValueMax="7.5" metricValueMin="1.9" pageId="54" pageNumber="53" unit="mm" value="47.0" valueMax="75.0" valueMin="19.0">19–75 mm</quantity>
. Sternum 
<typeStatus id="E763886BA821FFA6FC65FEF49DB1FE87" box="[933,979,291,314]" pageId="54" pageNumber="53">type</typeStatus>
1, triangular in shape. Pedipalps orthobothriotaxic, 
<typeStatus id="E763886BA821FFA6FC0FFE959D9FFEE4" box="[975,1021,322,345]" pageId="54" pageNumber="53">type</typeStatus>
Aβ. Chelal trichobothrium 
<emphasis id="0AACEADBA821FFA6FAEFFE959B28FEE4" box="[1327,1354,322,345]" italics="true" pageId="54" pageNumber="53">db</emphasis>
located between 
<emphasis id="0AACEADBA821FFA6FC62FEB79DD7FECA" box="[930,949,352,375]" italics="true" pageId="54" pageNumber="53">et</emphasis>
and 
<emphasis id="0AACEADBA821FFA6FC31FEB79A6DFECA" box="[1009,1039,352,375]" italics="true" pageId="54" pageNumber="53">est</emphasis>
. Three to five pairs of lateral eyes. Tibial spurs absent on all legs. Pedipalp movable finger with six rows of granules, several accessory granules and external and internal accessory granules. Pedipalp fixed finger with seven rows of granules and six external and seven internal accessory granules. Cheliceral fixed finger with a single ventral denticle. Third and fourth legs with tibial spurs. Tibia and tarsomeres of legs I–III with setae not arranged into bristle combs on dorsal surfaces. Tarsomere II of leg IV with two sparse rows of &lt;20 spiniform setae on ventral surface. Mesosoma dorsally with one median carina. Telson with subaculear tooth pointed or more often rounded. Males of most species have longer metasomal segments and often also a wider pedipalp chela manus than females.
</paragraph>
</subSubSection>
<subSubSection id="70C26542A821FFA6FCFEFC9C9D39F900" pageId="54" pageNumber="53" type="discussion">
<paragraph id="386736C9A821FFA6FCFEFC9C9D39F900" blockId="54.[830,1441,843,1725]" pageId="54" pageNumber="53">
SUBORDINATE TAXA. Currently, 27 species are formally included under the genus (cf. 
<bibRefCitation id="5C494B38A821FFA6FB65FCBD9DE4FC22" author="Kovarik &amp; Ojanguren" bookContentInfo="400 pp." firstAuthor="Kovarik" journalOrPublisher="Clairon Production, Prague" pageId="54" pageNumber="53" refId="ref42080" refString="KOVARIK, F &amp; A. A. OJANGUREN AFFILASTRO. 2013. Illustrated catalog of scorpions Part II. Bothriuridae; Chaerilidae; Buthidae I., genera Compsobuthus, Hottentotta, Isometrus, Lychas, and Sassanidotus. Clairon Production, Prague, 400 pp." title="Illustrated catalog of scorpions Part II. Bothriuridae; Chaerilidae; Buthidae I., genera Compsobuthus, Hottentotta, Isometrus, Lychas, and Sassanidotus" type="book" year="2013">Kovařík &amp; Ojanguren (2013)</bibRefCitation>
for information about species not treated here): 
<taxonomicName id="FFD84D4AA821FFA6FCFEFC709ABCFC03" authority="(Pocock, 1899)" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Pocock" baseAuthorityYear="1899" box="[830,1246,935,958]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="acanthurus" status="comb. nov.">
<emphasis id="0AACEADBA821FFA6FCFEFC709A51FC03" box="[830,1075,935,958]" italics="true" pageId="54" pageNumber="53">Reddyanus acanthurus</emphasis>
(Pocock, 1899)
</taxonomicName>
<emphasis id="0AACEADBA821FFA6FB25FC719B24FC03" bold="true" box="[1253,1350,934,958]" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FB25FC719B24FC03" box="[1253,1350,934,958]" pageId="54" pageNumber="53" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="FFD84D4AA821FFA6FA95FC709AFDFC60" authority="(Oates, 1888)" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Oates" baseAuthorityYear="1888" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="assamensis" status="comb. nov.">
<emphasis id="0AACEADBA821FFA6FA95FC709D95FC60" italics="true" pageId="54" pageNumber="53">Reddyanus assamensis</emphasis>
(Oates, 1888)
</taxonomicName>
<emphasis id="0AACEADBA821FFA6FB6DFC129B74FC60" bold="true" box="[1197,1302,965,989]" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FB6DFC129B74FC60" box="[1197,1302,965,989]" pageId="54" pageNumber="53" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="FFD84D4AA821FFA6FAEBFC119A25FC46" authority="(Karsch, 1879)" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Karsch" baseAuthorityYear="1879" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="basilicus" status="comb. nov.">
<emphasis id="0AACEADBA821FFA6FAEBFC119DFFFC46" italics="true" pageId="54" pageNumber="53">Reddyanus basilicus</emphasis>
(
<bibRefCitation id="5C494B38A821FFA6FC6CFC339A5DFC46" author="Karsch" box="[940,1087,996,1019]" firstAuthor="Karsch" pageId="54" pageNumber="53" pagination="97 - 136" refId="ref41417" refString="KARSCH, F. 1879. Skorpionologische Beitrage I. and II. Mitteilungen des Munchener Entomologischen Vereins, 3: 6 - 22, 97 - 136." type="journal article" year="1879">Karsch, 1879</bibRefCitation>
)
</taxonomicName>
<emphasis id="0AACEADBA821FFA6FB8EFC339AD2FC41" bold="true" box="[1102,1200,996,1020]" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FB8EFC339AD2FC41" box="[1102,1200,996,1020]" pageId="54" pageNumber="53" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="FFD84D4AA821FFA6FB7FFC339D8CFBA7" authority="(Vachon, 1982)" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Vachon" baseAuthorityYear="1982" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="besucheti" status="comb. nov.">
<emphasis id="0AACEADBA821FFA6FB7FFC339BC2FC46" box="[1215,1440,996,1019]" italics="true" pageId="54" pageNumber="53">Reddyanus besucheti</emphasis>
(
<bibRefCitation id="5C494B38A821FFA6FC86FBD49D84FBA7" author="Vachon" box="[838,998,1027,1050]" firstAuthor="Vachon" journalOrPublisher="Revue suisse de Zoologie" pageId="54" pageNumber="53" pagination="77 - 114" part="89" refId="ref45190" refString="VACHON, M. 1982. Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d`Histoire naturelle de Geneve III.). Revue suisse de Zoologie, 89: 77 - 114." title="Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d`Histoire naturelle de Geneve III.)" type="journal article" year="1982">Vachon, 1982</bibRefCitation>
)
</taxonomicName>
<emphasis id="0AACEADBA821FFA6FC3BFBD59A01FBA7" bold="true" box="[1019,1123,1026,1050]" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FC3BFBD59A01FBA7" box="[1019,1123,1026,1050]" pageId="54" pageNumber="53" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="FFD84D4AA821FFA6FBB8FBD49D1EFB84" authority="(Kovarik, 2003)" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Kovarik" baseAuthorityYear="2003" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="bilyi" status="comb. nov.">
<emphasis id="0AACEADBA821FFA6FBB8FBD49B49FBA7" box="[1144,1323,1027,1050]" italics="true" pageId="54" pageNumber="53">Reddyanus bilyi</emphasis>
(
<bibRefCitation id="5C494B38A821FFA6FA80FBD49D17FB84" author="Kovarik" firstAuthor="Kovarik" pageId="54" pageNumber="53" pagination="1 - 19" refId="ref41718" refString="KOVARIK, F. 2003. A review of the genus Isometrus Ehrenberg, 1828 (Scorpiones: Buthidae) with descriptions of four new species from Asia and Australia. Euscorpius, 10: 1 - 19." type="journal article" year="2003">Kovařík, 2003</bibRefCitation>
)
</taxonomicName>
<emphasis id="0AACEADBA821FFA6FC4EFBF69D9BFB84" bold="true" box="[910,1017,1057,1081]" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FC4EFBF69D9BFB84" box="[910,1017,1057,1081]" pageId="54" pageNumber="53" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="FFD84D4AA821FFA6FBD2FBF59D1FFBEA" authority="(Pocock, 1899)" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Pocock" baseAuthorityYear="1899" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="brachycentrus" status="comb. nov.">
<emphasis id="0AACEADBA821FFA6FBD2FBF59B52FB84" box="[1042,1328,1058,1081]" italics="true" pageId="54" pageNumber="53">Reddyanus brachycentrus</emphasis>
(Pocock, 1899)
</taxonomicName>
<emphasis id="0AACEADBA821FFA6FC45FB979D85FBE5" bold="true" box="[901,999,1088,1112]" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FC45FB979D85FBE5" box="[901,999,1088,1112]" pageId="54" pageNumber="53" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="FFD84D4AA821FFA6FC36FB979BFEFBCB" authority="Kovarik, Lowe, Ranawana, Hoferek, Jayarathne, Pliskova et Stahlavsky" authorityName="Kovarik, Lowe, Ranawana, Hoferek, Jayarathne, Pliskova et Stahlavsky" authorityYear="2016" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="ceylonensis" status="sp. nov.">
<emphasis id="0AACEADBA821FFA6FC36FB979A8CFBEA" box="[1014,1262,1088,1111]" italics="true" pageId="54" pageNumber="53">Reddyanus ceylonensis</emphasis>
Kovařík, Lowe, Ranawana, Hoferek, Jayarathne, Plíšková et Šťáhlavský
</taxonomicName>
, 
<emphasis id="0AACEADBA821FFA6FCFEFBAA9D1EFB28" bold="true" box="[830,892,1149,1173]" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FCFEFBAA9D1EFB28" box="[830,892,1149,1173]" pageId="54" pageNumber="53" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="FFD84D4AA821FFA6FC4CFBA99BC2FB28" authority="(Tikader et Bastawade, 1983)" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Tikader et Bastawade" baseAuthorityYear="1983" box="[908,1440,1150,1173]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="corbeti" status="comb. nov.">
<emphasis id="0AACEADBA821FFA6FC4CFBA99A34FB28" box="[908,1110,1150,1173]" italics="true" pageId="54" pageNumber="53">Reddyanus corbeti</emphasis>
(
<bibRefCitation id="5C494B38A821FFA6FBA5FBA99BFBFB28" author="Tikader et Bastawade" bookContentInfo="668 pp." box="[1125,1433,1150,1173]" firstAuthor="Tikader" journalOrPublisher="Zoological Survey of India, Calcutta" pageId="54" pageNumber="53" refId="ref44789" refString="TIKADER B. K. &amp; D. B. BASTAWADE. 1983. The Fauna of India: Scorpions. Scorpionida, Arachnida. Vol III. Zoological Survey of India, Calcutta, 668 pp." title="The Fauna of India: Scorpions. Scorpionida, Arachnida. Vol III" type="book" year="1983">Tikader et Bastawade, 1983</bibRefCitation>
)
</taxonomicName>
<emphasis id="0AACEADBA821FFA6FCFEFB4B9DC1FB09" bold="true" box="[830,931,1180,1204]" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FCFEFB4B9DC1FB09" box="[830,931,1180,1204]" pageId="54" pageNumber="53" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="FFD84D4AA821FFA6FC74FB4B9D1FFB6F" authority="(Lourenco et Duhem, 2010)" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Lourenco et Duhem" baseAuthorityYear="2010" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="deharvengi" status="comb. nov.">
<emphasis id="0AACEADBA821FFA6FC74FB4B9AC9FB0E" box="[948,1195,1180,1203]" italics="true" pageId="54" pageNumber="53">Reddyanus deharvengi</emphasis>
(Lourenço et Duhem, 2010)
</taxonomicName>
<emphasis id="0AACEADBA821FFA6FC46FB6D9D8BFB6F" bold="true" box="[902,1001,1210,1234]" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FC46FB6D9D8BFB6F" box="[902,1001,1210,1234]" pageId="54" pageNumber="53" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="FFD84D4AA821FFA6FC3AFB6C9B30FB6F" authority="(Kovarik, 2013)" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Kovarik" baseAuthorityYear="2013" box="[1018,1362,1211,1234]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="feti" status="comb. nov.">
<emphasis id="0AACEADBA821FFA6FC3AFB6C9AF8FB6F" box="[1018,1178,1211,1234]" italics="true" pageId="54" pageNumber="53">Reddyanus feti</emphasis>
(
<bibRefCitation id="5C494B38A821FFA6FB6BFB6C9B29FB6F" author="Kovarik" box="[1195,1355,1211,1234]" firstAuthor="Kovarik" journalOrPublisher="Euscorpius" pageId="54" pageNumber="53" pagination="1 - 9" part="175" refId="ref41834" refString="KOVARIK, F. 2013. Review of Thaicharmus Kovarik, 1995, with description of Thaicharmus indicus sp. n. from India (Scorpiones, Buthidae). Euscorpius, 175: 1 - 9." title="Review of Thaicharmus Kovarik, 1995, with description of Thaicharmus indicus sp. n. from India (Scorpiones, Buthidae)" type="journal article" year="2013">Kovařík, 2013</bibRefCitation>
)
</taxonomicName>
<emphasis id="0AACEADBA821FFA6FA9BFB6D9D36FB4C" bold="true" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FA9BFB6D9D36FB4C" pageId="54" pageNumber="53" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="FFD84D4AA821FFA6FCA5FB0D9AB7FB4C" authority="(Vachon, 1976)" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Vachon" baseAuthorityYear="1976" box="[869,1237,1242,1265]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="heimi" status="comb. nov.">
<emphasis id="0AACEADBA821FFA6FCA5FB0D9A7DFB4C" box="[869,1055,1242,1265]" italics="true" pageId="54" pageNumber="53">Reddyanus heimi</emphasis>
(Vachon, 1976)
</taxonomicName>
<emphasis id="0AACEADBA821FFA6FB1FFB0E9B21FB4C" bold="true" box="[1247,1347,1241,1265]" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FB1FFB0E9B21FB4C" box="[1247,1347,1241,1265]" pageId="54" pageNumber="53" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="FFD84D4AA821FFA6FA95FB0D9A03FAB2" authority="Kovarik" authorityName="Kovarik" authorityYear="2016" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="jayarathnei" status="sp. nov.">
<emphasis id="0AACEADBA821FFA6FA95FB0D9D99FAB2" italics="true" pageId="54" pageNumber="53">Reddyanus jayarathnei</emphasis>
Kovařík
</taxonomicName>
, 
<emphasis id="0AACEADBA821FFA6FBB6FB2F9AD8FAAD" bold="true" box="[1142,1210,1272,1296]" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FBB6FB2F9AD8FAAD" box="[1142,1210,1272,1296]" pageId="54" pageNumber="53" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="FFD84D4AA821FFA6FB0FFB2F9D93FA93" authority="(Kovarik, 2013)" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Kovarik" baseAuthorityYear="2013" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="jendeki" status="comb. nov.">
<emphasis id="0AACEADBA821FFA6FB0FFB2F9BC2FAB2" box="[1231,1440,1272,1295]" italics="true" pageId="54" pageNumber="53">Reddyanus jendeki</emphasis>
(
<bibRefCitation id="5C494B38A821FFA6FC86FAC09D88FA93" author="Kovarik" box="[838,1002,1303,1326]" firstAuthor="Kovarik" journalOrPublisher="Euscorpius" pageId="54" pageNumber="53" pagination="1 - 9" part="175" refId="ref41834" refString="KOVARIK, F. 2013. Review of Thaicharmus Kovarik, 1995, with description of Thaicharmus indicus sp. n. from India (Scorpiones, Buthidae). Euscorpius, 175: 1 - 9." title="Review of Thaicharmus Kovarik, 1995, with description of Thaicharmus indicus sp. n. from India (Scorpiones, Buthidae)" type="journal article" year="2013">Kovařík, 2013</bibRefCitation>
)
</taxonomicName>
<emphasis id="0AACEADBA821FFA6FC3FFAC19A04FA93" bold="true" box="[1023,1126,1302,1326]" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FC3FFAC19A04FA93" box="[1023,1126,1302,1326]" pageId="54" pageNumber="53" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="FFD84D4AA821FFA6FBBBFAC09D99FAF0" authority="(Kovarik, 2003)" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Kovarik" baseAuthorityYear="2003" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="khammamensis" status="comb. nov.">
<emphasis id="0AACEADBA821FFA6FBBBFAC09BC2FA93" box="[1147,1440,1303,1326]" italics="true" pageId="54" pageNumber="53">Reddyanus khammamensis</emphasis>
(
<bibRefCitation id="5C494B38A821FFA6FC86FAE19D96FAF0" author="Kovarik" box="[838,1012,1334,1357]" firstAuthor="Kovarik" pageId="54" pageNumber="53" pagination="1 - 19" refId="ref41718" refString="KOVARIK, F. 2003. A review of the genus Isometrus Ehrenberg, 1828 (Scorpiones: Buthidae) with descriptions of four new species from Asia and Australia. Euscorpius, 10: 1 - 19." type="journal article" year="2003">Kovařík, 2003</bibRefCitation>
)
</taxonomicName>
<emphasis id="0AACEADBA821FFA6FBD3FAE29AE6FAF0" bold="true" box="[1043,1156,1333,1357]" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FBD3FAE29AE6FAF0" box="[1043,1156,1333,1357]" pageId="54" pageNumber="53" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="FFD84D4AA821FFA6FB63FAE19D8CFAD6" authority="(Kovarik, 1998)" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Kovarik" baseAuthorityYear="1998" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="krasenskyi" status="comb. nov.">
<emphasis id="0AACEADBA821FFA6FB63FAE19BC2FAF0" box="[1187,1440,1334,1357]" italics="true" pageId="54" pageNumber="53">Reddyanus krasenskyi</emphasis>
(
<bibRefCitation id="5C494B38A821FFA6FC86FA839D84FAD6" author="Kovarik" box="[838,998,1364,1387]" firstAuthor="Kovarik" pageId="54" pageNumber="53" refId="ref41687" refString="KOVARIK, F. 1998. Stiri [Scorpiones]. Publishing House &quot; Madagaskar &quot;, Jihlava (Czech Republic). 175 pp. (in Czech)." type="book" year="1998">Kovařík, 1998</bibRefCitation>
)
</taxonomicName>
<emphasis id="0AACEADBA821FFA6FC38FA839A39FAD6" bold="true" box="[1016,1115,1364,1387]" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FC38FA839A39FAD6" box="[1016,1115,1364,1387]" pageId="54" pageNumber="53" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="FFD84D4AA821FFA6FBACFA839D1FFA37" authority="(Kovarik, 1997)" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Kovarik" baseAuthorityYear="1997" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="kurkai" status="comb. nov.">
<emphasis id="0AACEADBA821FFA6FBACFA839B4DFAD6" box="[1132,1327,1364,1387]" italics="true" pageId="54" pageNumber="53">Reddyanus kurkai</emphasis>
(
<bibRefCitation id="5C494B38A821FFA6FA80FA839D17FA37" author="Kovarik" firstAuthor="Kovarik" pageId="54" pageNumber="53" pagination="311 - 371" refId="ref41650" refString="KOVARIK, F. 1997. Revision of the genera Lychas and Hemilychas, with descriptions of six new species (Scorpiones: Buthidae). Acta Societatis Zoologicae Bohemicae, 61: 311 - 371." type="journal article" year="1997">Kovařík, 1997</bibRefCitation>
)
</taxonomicName>
<emphasis id="0AACEADBA821FFA6FC4FFAA59D9EFA37" bold="true" box="[911,1020,1394,1418]" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FC4FFAA59D9EFA37" box="[911,1020,1394,1418]" pageId="54" pageNumber="53" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="FFD84D4AA821FFA6FBD6FAA49BC3FA37" authority="(Vachon, 1982)" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Vachon" baseAuthorityYear="1982" box="[1046,1441,1395,1418]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="loebli" status="comb. nov.">
<emphasis id="0AACEADBA821FFA6FBD6FAA49AB8FA37" box="[1046,1242,1395,1418]" italics="true" pageId="54" pageNumber="53">Reddyanus loebli</emphasis>
(
<bibRefCitation id="5C494B38A821FFA6FB34FAA49BFBFA37" author="Vachon" box="[1268,1433,1395,1418]" firstAuthor="Vachon" journalOrPublisher="Revue suisse de Zoologie" pageId="54" pageNumber="53" pagination="77 - 114" part="89" refId="ref45190" refString="VACHON, M. 1982. Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d`Histoire naturelle de Geneve III.). Revue suisse de Zoologie, 89: 77 - 114." title="Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d`Histoire naturelle de Geneve III.)" type="journal article" year="1982">Vachon, 1982</bibRefCitation>
)
</taxonomicName>
<emphasis id="0AACEADBA821FFA6FCFEFA469DC6FA14" bold="true" box="[830,932,1425,1449]" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FCFEFA469DC6FA14" box="[830,932,1425,1449]" pageId="54" pageNumber="53" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="FFD84D4AA821FFA6FC77FA459BC2FA14" authority="(L. Koch, 1867)" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="L. Koch" baseAuthorityYear="1867" box="[951,1440,1426,1449]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="melanodactylus" status="comb. nov.">
<emphasis id="0AACEADBA821FFA6FC77FA459ABDFA14" box="[951,1247,1426,1449]" italics="true" pageId="54" pageNumber="53">Reddyanus melanodactylus</emphasis>
(L. Koch, 1867)
</taxonomicName>
<emphasis id="0AACEADBA821FFA6FCFEFA679DC2FA75" bold="true" box="[830,928,1456,1480]" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FCFEFA679DC2FA75" box="[830,928,1456,1480]" pageId="54" pageNumber="53" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="FFD84D4AA821FFA6FC6EFA679B53FA7A" authority="(Kovarik, 1998)" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Kovarik" baseAuthorityYear="1998" box="[942,1329,1456,1479]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="navaiae" status="comb. nov.">
<emphasis id="0AACEADBA821FFA6FC6EFA679A1CFA7A" box="[942,1150,1456,1479]" italics="true" pageId="54" pageNumber="53">Reddyanus navaiae</emphasis>
(
<bibRefCitation id="5C494B38A821FFA6FB4CFA679B48FA7A" author="Kovarik" box="[1164,1322,1456,1479]" firstAuthor="Kovarik" pageId="54" pageNumber="53" refId="ref41687" refString="KOVARIK, F. 1998. Stiri [Scorpiones]. Publishing House &quot; Madagaskar &quot;, Jihlava (Czech Republic). 175 pp. (in Czech)." type="book" year="1998">Kovařík, 1998</bibRefCitation>
)
</taxonomicName>
<emphasis id="0AACEADBA821FFA6FAF8FA679BFBFA75" bold="true" box="[1336,1433,1456,1480]" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FAF8FA679BFBFA75" box="[1336,1433,1456,1480]" pageId="54" pageNumber="53" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="FFD84D4AA821FFA6FCFEFA189AD6FA5B" authority="(Kovarik, 2013)" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Kovarik" baseAuthorityYear="2013" box="[830,1204,1487,1510]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="neradi" status="comb. nov.">
<emphasis id="0AACEADBA821FFA6FCFEFA189A63FA5B" box="[830,1025,1487,1510]" italics="true" pageId="54" pageNumber="53">Reddyanus neradi</emphasis>
(
<bibRefCitation id="5C494B38A821FFA6FBCFFA189ACFFA5B" author="Kovarik" box="[1039,1197,1487,1510]" firstAuthor="Kovarik" journalOrPublisher="Euscorpius" pageId="54" pageNumber="53" pagination="1 - 9" part="175" refId="ref41834" refString="KOVARIK, F. 2013. Review of Thaicharmus Kovarik, 1995, with description of Thaicharmus indicus sp. n. from India (Scorpiones, Buthidae). Euscorpius, 175: 1 - 9." title="Review of Thaicharmus Kovarik, 1995, with description of Thaicharmus indicus sp. n. from India (Scorpiones, Buthidae)" type="journal article" year="2013">Kovařík, 2013</bibRefCitation>
)
</taxonomicName>
<emphasis id="0AACEADBA821FFA6FB7BFA199B7FFA5B" bold="true" box="[1211,1309,1486,1510]" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FB7BFA199B7FFA5B" box="[1211,1309,1486,1510]" pageId="54" pageNumber="53" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="FFD84D4AA821FFA6FAEBFA189A00F9B8" authority="(Kovarik, 2003)" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Kovarik" baseAuthorityYear="2003" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="petrzelkai" status="comb. nov.">
<emphasis id="0AACEADBA821FFA6FAEBFA189DC5F9B8" italics="true" pageId="54" pageNumber="53">Reddyanus petrzelkai</emphasis>
(
<bibRefCitation id="5C494B38A821FFA6FC7AFA399A39F9B8" author="Kovarik" box="[954,1115,1518,1541]" firstAuthor="Kovarik" pageId="54" pageNumber="53" pagination="1 - 19" refId="ref41718" refString="KOVARIK, F. 2003. A review of the genus Isometrus Ehrenberg, 1828 (Scorpiones: Buthidae) with descriptions of four new species from Asia and Australia. Euscorpius, 10: 1 - 19." type="journal article" year="2003">Kovařík, 2003</bibRefCitation>
)
</taxonomicName>
<emphasis id="0AACEADBA821FFA6FBAEFA3A9AB1F9B8" bold="true" box="[1134,1235,1517,1541]" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FBAEFA3A9AB1F9B8" box="[1134,1235,1517,1541]" pageId="54" pageNumber="53" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="FFD84D4AA821FFA6FB25FA399A03F99E" authority="(Kovarik, 2003)" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Kovarik" baseAuthorityYear="2003" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="problematicus" status="comb. nov.">
<emphasis id="0AACEADBA821FFA6FB25FA399DC6F99E" italics="true" pageId="54" pageNumber="53">Reddyanus problematicus</emphasis>
(
<bibRefCitation id="5C494B38A821FFA6FC77F9DB9A3BF99E" author="Kovarik" box="[951,1113,1548,1571]" firstAuthor="Kovarik" pageId="54" pageNumber="53" pagination="1 - 19" refId="ref41718" refString="KOVARIK, F. 2003. A review of the genus Isometrus Ehrenberg, 1828 (Scorpiones: Buthidae) with descriptions of four new species from Asia and Australia. Euscorpius, 10: 1 - 19." type="journal article" year="2003">Kovařík, 2003</bibRefCitation>
)
</taxonomicName>
<emphasis id="0AACEADBA821FFA6FBACF9DB9AB0F999" bold="true" box="[1132,1234,1548,1572]" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FBACF9DB9AB0F999" box="[1132,1234,1548,1572]" pageId="54" pageNumber="53" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="FFD84D4AA821FFA6FB24F9DB9D85F9FF" authority="Kovarik" authorityName="Kovarik" authorityYear="2016" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="ranawanai" status="sp. nov.">
<emphasis id="0AACEADBA821FFA6FB24F9DB9D1DF9FF" italics="true" pageId="54" pageNumber="53">Reddyanus ranawanai</emphasis>
Kovařík
</taxonomicName>
, 
<emphasis id="0AACEADBA821FFA6FC39F9FD9A5EF9FF" bold="true" box="[1017,1084,1578,1602]" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FC39F9FD9A5EF9FF" box="[1017,1084,1578,1602]" pageId="54" pageNumber="53" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="FFD84D4AA821FFA6FB91F9FC9D1FF9DC" authority="(Pocock, 1897)" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Pocock" baseAuthorityYear="1897" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="rigidulus" status="comb. nov.">
<emphasis id="0AACEADBA821FFA6FB91F9FC9B56F9FF" box="[1105,1332,1579,1602]" italics="true" pageId="54" pageNumber="53">Reddyanus rigidulus</emphasis>
(
<bibRefCitation id="5C494B38A821FFA6FA89F9FC9D17F9DC" author="Pocock" firstAuthor="Pocock" journalOrPublisher="Journal of the Bombay Natural History Society" pageId="54" pageNumber="53" pagination="102 - 117" part="11" refId="ref43758" refString="POCOCK, R. I. 1897. Descriptions of some new species of scorpions from India. Journal of the Bombay Natural History Society, 11: 102 - 117." title="Descriptions of some new species of scorpions from India" type="journal article" year="1897">Pocock, 1897</bibRefCitation>
)
</taxonomicName>
<emphasis id="0AACEADBA821FFA6FC45F99E9D84F9DC" bold="true" box="[901,998,1609,1633]" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FC45F99E9D84F9DC" box="[901,998,1609,1633]" pageId="54" pageNumber="53" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="FFD84D4AA821FFA6FC36F99D9D1EF9C2" authority="(Lourenco et Zhu, 2008)" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Lourenco et Zhu" baseAuthorityYear="2008" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="tibetanus" status="comb. nov.">
<emphasis id="0AACEADBA821FFA6FC36F99D9AB7F9DC" box="[1014,1237,1610,1633]" italics="true" pageId="54" pageNumber="53">Reddyanus tibetanus</emphasis>
(Lourenço et Zhu, 2008)
</taxonomicName>
<emphasis id="0AACEADBA821FFA6FC4DF9BF9D9AF93D" bold="true" box="[909,1016,1640,1664]" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FC4DF9BF9D9AF93D" box="[909,1016,1640,1664]" pageId="54" pageNumber="53" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
, 
<taxonomicName id="FFD84D4AA821FFA6FBCFF9BF9BC2F9C2" authority="(Pocock, 1900)" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Pocock" baseAuthorityYear="1900" box="[1039,1440,1640,1663]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="vittatus" status="comb. nov.">
<emphasis id="0AACEADBA821FFA6FBCFF9BF9A81F9C2" box="[1039,1251,1640,1663]" italics="true" pageId="54" pageNumber="53">Reddyanus vittatus</emphasis>
(
<bibRefCitation id="5C494B38A821FFA6FB3BF9BF9BFBF9C2" author="Pocock" bookContentInfo="279 pp" box="[1275,1433,1640,1663]" firstAuthor="Pocock" journalOrPublisher="London: W. T. Blandford" pageId="54" pageNumber="53" part="xii" refId="ref43790" refString="POCOCK, R. I. 1900. Arachnida. The fauna of British India, including Ceylon and Burma. Published un- der the authority of the Secretary of State for India in Council. London: W. T. Blandford, xii, 279 pp" title="Arachnida. The fauna of British India, including Ceylon and Burma. Published un- der the authority of the Secretary of State for India in Council" type="book" year="1900">Pocock, 1900</bibRefCitation>
)
</taxonomicName>
<emphasis id="0AACEADBA821FFA6FCFEF9519DC0F923" bold="true" box="[830,930,1670,1694]" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FCFEF9519DC0F923" box="[830,930,1670,1694]" pageId="54" pageNumber="53" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
and 
<taxonomicName id="FFD84D4AA821FFA6FC1CF9509B30F923" authority="(Kovarik, 1994)" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Kovarik" baseAuthorityYear="1994" box="[988,1362,1671,1694]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="species" species="zideki" status="comb. nov.">
<emphasis id="0AACEADBA821FFA6FC1CF9509AFBF923" box="[988,1177,1671,1694]" italics="true" pageId="54" pageNumber="53">Reddyanus zideki</emphasis>
(
<bibRefCitation id="5C494B38A821FFA6FB6AF9509B28F923" author="Kovarik" box="[1194,1354,1671,1694]" firstAuthor="Kovarik" pageId="54" pageNumber="53" pagination="195 - 203" refId="ref41554" refString="KOVARIK, F. 1994. Isometrus zideki sp. n. from Malaysia and Indonesia, and a taxonomic position of Isometrus formosus, I. thurstoni and I. sankariensis (Arachnida: Scorpionida: Buthidae). Acta Societatis Zoologicae Bohemicae, 58: 195 - 203." type="journal article" year="1994">Kovařík, 1994</bibRefCitation>
)
</taxonomicName>
<emphasis id="0AACEADBA821FFA6FA9BF9519D36F900" bold="true" pageId="54" pageNumber="53">
<taxonomicNameLabel id="119F57A0A821FFA6FA9BF9519D36F900" pageId="54" pageNumber="53" rank="species">comb. n.</taxonomicNameLabel>
</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="70C26542A821FFA9FCFEF9349C83FAA2" lastPageId="57" lastPageNumber="56" pageId="54" pageNumber="53" type="discussion">
<paragraph id="386736C9A821FFA9FCFEF9349C83FAA2" blockId="54.[830,1440,1763,1909]" lastBlockId="57.[192,802,1043,1311]" lastPageId="57" lastPageNumber="56" pageId="54" pageNumber="53">
COMMENTS. 
<bibRefCitation id="5C494B38A821FFA6FC09F9349AFDF947" author="Vachon" box="[969,1183,1763,1786]" firstAuthor="Vachon" journalOrPublisher="Cahiers Pacifique" page="177" pageId="54" pageNumber="53" pagination="169 - 180" part="16" refId="ref45078" refString="VACHON, M. 1972. Remarques sur les scorpions appartenant au genre Isometrus H. et E. (Buthidae) a propos de l`espece Isometrus maculatus (Geer) habitant Ile de Paques. Cahiers Pacifique, 16: 169 - 180." title="Remarques sur les scorpions appartenant au genre Isometrus H. et E. (Buthidae) a propos de l`espece Isometrus maculatus (Geer) habitant Ile de Paques" type="journal article" year="1972">Vachon (1972: 177)</bibRefCitation>
described the subgenus 
<taxonomicName id="FFD84D4AA821FFA6FCFEF8D59DD1F8A4" authorityName="Vachon" authorityYear="1972" box="[830,947,1794,1817]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA821FFA6FCFEF8D59DD1F8A4" box="[830,947,1794,1817]" italics="true" pageId="54" pageNumber="53">Reddyanus</emphasis>
</taxonomicName>
and distinguished it from the nominotypic subgenus 
<taxonomicName id="FFD84D4AA821FFA6FC6BF8F79AB0F88A" authority="Ehrenberg, 1828" authorityName="Ehrenberg" authorityYear="1828" box="[939,1234,1824,1847]" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA821FFA6FC6BF8F79A76F88A" box="[939,1044,1824,1847]" italics="true" pageId="54" pageNumber="53">Isometrus</emphasis>
Ehrenberg, 1828
</taxonomicName>
by the position of chelal fixed finger trichobothria (trichobothrium 
<emphasis id="0AACEADBA821FFA6FA9DF8E89B1AF8EB" box="[1373,1400,1855,1878]" italics="true" pageId="54" pageNumber="53">db</emphasis>
located between 
<emphasis id="0AACEADBA821FFA6FC1DF8899D90F8C8" box="[989,1010,1886,1909]" italics="true" pageId="54" pageNumber="53">dt</emphasis>
and 
<emphasis id="0AACEADBA821FFA6FBE6F8899A5BF8C8" box="[1062,1081,1886,1909]" italics="true" pageId="54" pageNumber="53">et</emphasis>
in 
<taxonomicName id="FFD84D4AA821FFA6FBA1F8899AA8F8C8" authorityName="Ehrenberg" authorityYear="1828" box="[1121,1226,1886,1909]" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="54" pageNumber="53" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA821FFA6FBA1F8899AA8F8C8" box="[1121,1226,1886,1909]" italics="true" pageId="54" pageNumber="53">Isometrus</emphasis>
</taxonomicName>
, between 
<emphasis id="0AACEADBA821FFA6FAF8F8899B29F8C8" box="[1336,1355,1886,1909]" italics="true" pageId="54" pageNumber="53">et</emphasis>
and 
<emphasis id="0AACEADBA821FFA6FABFF8899BFFF8C8" box="[1407,1437,1886,1909]" italics="true" pageId="54" pageNumber="53">est</emphasis>
in 
<taxonomicName id="FFD84D4AA820FFA7FF1DF9C09F30F993" authorityName="Vachon" authorityYear="1972" box="[221,338,1559,1582]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA820FFA7FF1DF9C09F30F993" box="[221,338,1559,1582]" italics="true" pageId="55" pageNumber="54">Reddyanus</emphasis>
</taxonomicName>
). Vachon also cited the different positions of the 
<emphasis id="0AACEADBA820FFA7FECBF9E19F75F9F0" box="[267,279,1590,1613]" italics="true" pageId="55" pageNumber="54">e</emphasis>
<subScript id="A45C348CA820FFA7FED7F9979F42F9F2" attach="left" box="[279,288,1600,1615]" fontSize="6" pageId="55" pageNumber="54">2</subScript>
trichobothria on the femur (see figs 
<quantity id="FF209B2CA820FFA7FD7DF9E19D40F9F0" box="[701,802,1590,1613]" metricMagnitude="-1" metricUnit="m" metricValue="3.85064" pageId="55" pageNumber="54" unit="in" value="15.16">15, 16 in</quantity>
<bibRefCitation id="5C494B38A820FFA7FF00F9829FF2F9D1" author="Vachon" box="[192,400,1621,1644]" firstAuthor="Vachon" journalOrPublisher="Cahiers Pacifique" page="176" pageId="55" pageNumber="54" pagination="169 - 180" part="16" refId="ref45078" refString="VACHON, M. 1972. Remarques sur les scorpions appartenant au genre Isometrus H. et E. (Buthidae) a propos de l`espece Isometrus maculatus (Geer) habitant Ile de Paques. Cahiers Pacifique, 16: 169 - 180." title="Remarques sur les scorpions appartenant au genre Isometrus H. et E. (Buthidae) a propos de l`espece Isometrus maculatus (Geer) habitant Ile de Paques" type="journal article" year="1972">Vachon, 1972: 176</bibRefCitation>
). Subsequently, 
<bibRefCitation id="5C494B38A820FFA7FD87F9829C88F9D1" author="Vachon" box="[583,746,1621,1644]" firstAuthor="Vachon" journalOrPublisher="Revue suisse de Zoologie" pageId="55" pageNumber="54" pagination="77 - 114" part="89" refId="ref45190" refString="VACHON, M. 1982. Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d`Histoire naturelle de Geneve III.). Revue suisse de Zoologie, 89: 77 - 114." title="Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d`Histoire naturelle de Geneve III.)" type="journal article" year="1982">Vachon (1982)</bibRefCitation>
added another character for distinguishing the two subgenera: 
<taxonomicName id="FFD84D4AA820FFA7FEDCF9459FE7F914" authorityName="Ehrenberg" authorityYear="1828" box="[284,389,1682,1705]" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA820FFA7FEDCF9459FE7F914" box="[284,389,1682,1705]" italics="true" pageId="55" pageNumber="54">Isometrus</emphasis>
</taxonomicName>
has pedipalp fixed fingers with six rows of granules and six external and internal accessory granules (
<figureCitation id="A0E32A4CA820FFA7FEF1F9189FB2F95B" box="[305,464,1743,1766]" captionStart="Figures 241–259" captionStartId="52.[192,278,1733,1757]" captionTargetBox="[238,1387,153,1699]" captionTargetId="figure-11@52.[234,1392,152,1704]" captionTargetPageId="52" captionText="Figures 241–259: Pedipalp chela dorsal (241), movable finger (242–250), and fixed finger (251–259) of Sri Lankan Isometrus, Lychas, and Reddyanus species. Figures 241–242, 252. Isometrus maculatus, female from locality 15CO (241) and male from locality 15CP (242, 252). Figures 243–244, 253. I. thwaitesi, male from locality 15CH (243) and female from locality 15CO (244, 253). Figures 245, 254. Reddyanus basilicus, male from locality 15CS. Figures 246, 255. R. besucheti, male from locality 15CD. Figures 247, 256. R. ceylonensis sp. n., male holotype. Figures 248, 257. R. jayarathnei sp. n., male paratype. Figures 249, 258. R. loebli, male from locality 15CG. Figures 250, 259. R. ranawanai sp. n., male holotype. Figure 251. Lychas srilankensis, male from locality 15CO." figureDoi="http://doi.org/10.5281/zenodo.7124626" httpUri="https://zenodo.org/record/7124626/files/figure.png" pageId="55" pageNumber="54">Figs. 252–253</figureCitation>
and figs. 
<quantity id="FF209B2CA820FFA7FD8EF9189CD8F95B" box="[590,698,1743,1766]" metricMagnitude="0" metricUnit="m" metricValue="1.64211" pageId="55" pageNumber="54" unit="in" value="64.65">64, 65 in</quantity>
<bibRefCitation id="5C494B38A820FFA7FD08F9189F52F8B8" author="Vachon" firstAuthor="Vachon" journalOrPublisher="Revue suisse de Zoologie" page="100" pageId="55" pageNumber="54" pagination="77 - 114" part="89" refId="ref45190" refString="VACHON, M. 1982. Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d`Histoire naturelle de Geneve III.). Revue suisse de Zoologie, 89: 77 - 114." title="Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d`Histoire naturelle de Geneve III.)" type="journal article" year="1982">Vachon, 1982: 100</bibRefCitation>
), and 
<taxonomicName id="FFD84D4AA820FFA7FEBEF9399F91F8B8" authorityName="Vachon" authorityYear="1972" box="[382,499,1774,1797]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA820FFA7FEBEF9399F91F8B8" box="[382,499,1774,1797]" italics="true" pageId="55" pageNumber="54">Reddyanus</emphasis>
</taxonomicName>
has pedipalp fixed fingers with seven rows of granules and six external and seven internal granules (
<figureCitation id="A0E32A4CA820FFA7FE46F8FC9C41F8FF" box="[390,547,1835,1858]" captionStart="Figures 241–259" captionStartId="52.[192,278,1733,1757]" captionTargetBox="[238,1387,153,1699]" captionTargetId="figure-11@52.[234,1392,152,1704]" captionTargetPageId="52" captionText="Figures 241–259: Pedipalp chela dorsal (241), movable finger (242–250), and fixed finger (251–259) of Sri Lankan Isometrus, Lychas, and Reddyanus species. Figures 241–242, 252. Isometrus maculatus, female from locality 15CO (241) and male from locality 15CP (242, 252). Figures 243–244, 253. I. thwaitesi, male from locality 15CH (243) and female from locality 15CO (244, 253). Figures 245, 254. Reddyanus basilicus, male from locality 15CS. Figures 246, 255. R. besucheti, male from locality 15CD. Figures 247, 256. R. ceylonensis sp. n., male holotype. Figures 248, 257. R. jayarathnei sp. n., male paratype. Figures 249, 258. R. loebli, male from locality 15CG. Figures 250, 259. R. ranawanai sp. n., male holotype. Figure 251. Lychas srilankensis, male from locality 15CO." figureDoi="http://doi.org/10.5281/zenodo.7124626" httpUri="https://zenodo.org/record/7124626/files/figure.png" pageId="55" pageNumber="54">Figs. 254–259</figureCitation>
and figs. 66 and 
<quantity id="FF209B2CA820FFA7FD29F8FC9D40F8FF" box="[745,802,1835,1858]" metricMagnitude="0" metricUnit="m" metricValue="1.7018" pageId="55" pageNumber="54" unit="in" value="67.0">67 in</quantity>
<bibRefCitation id="5C494B38A820FFA7FF00F89D9FFAF8DC" author="Vachon" box="[192,408,1866,1889]" firstAuthor="Vachon" journalOrPublisher="Revue suisse de Zoologie" page="100" pageId="55" pageNumber="54" pagination="77 - 114" part="89" refId="ref45190" refString="VACHON, M. 1982. Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d`Histoire naturelle de Geneve III.). Revue suisse de Zoologie, 89: 77 - 114." title="Les scorpions de Sri Lanka (Recherches sur les scorpions appartenant ou deposes au Museum d`Histoire naturelle de Geneve III.)" type="journal article" year="1982">Vachon, 1982: 100</bibRefCitation>
). Here we propose two additional characters for distinguishing these taxa: (i) ventral aspect of tarsomere II of leg IV with two dense rows of&gt; 30 longer, more filamentous setae in 
<taxonomicName id="FFD84D4AA820FFA7FB75F9829B7EF9D1" authorityName="Ehrenberg" authorityYear="1828" box="[1205,1308,1621,1644]" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA820FFA7FB75F9829B7EF9D1" box="[1205,1308,1621,1644]" italics="true" pageId="55" pageNumber="54">Isometrus</emphasis>
</taxonomicName>
(
<figureCitation id="A0E32A4CA820FFA7FAEEF9829D04F937" captionStart="Figures 193–200" captionStartId="48.[192,278,1070,1094]" captionTargetBox="[213,1446,171,1037]" captionTargetId="figure-11@48.[205,1452,168,1041]" captionTargetPageId="48" captionText="Figures 193–200: Distal segments of leg IV, retroventral view of Sri Lankan Buthidae and Chaerilidae genera. Buthoscorpio sarasinorum, male from locality 15CF (193), Charmus laneus, male from locality 15CO (194), Hottentotta tamulus, male from locality 15CK (195), Isometrus maculatus, female from locality 15CI (196), Isometrus thwaitesi, female from locality 15CO (197), Lychas srilankensis, male from locality 15CJ (198), Reddyanus basilicus, male from locality 15CS (199), and Chaerilus ceylonensis, male from locality 15CD (200)." figureDoi="http://doi.org/10.5281/zenodo.7124608" httpUri="https://zenodo.org/record/7124608/files/figure.png" pageId="55" pageNumber="54">Figs. 196– 197</figureCitation>
), vs. two sparse rows of &lt;20 shorter, thicker, more spiniform setae in 
<taxonomicName id="FFD84D4AA820FFA7FBCCF9459AE3F914" authorityName="Vachon" authorityYear="1972" box="[1036,1153,1682,1705]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA820FFA7FBCCF9459AE3F914" box="[1036,1153,1682,1705]" italics="true" pageId="55" pageNumber="54">Reddyanus</emphasis>
</taxonomicName>
(
<figureCitation id="A0E32A4CA820FFA7FB53F9459A9BF914" box="[1171,1273,1682,1705]" captionStart="Figures 193–200" captionStartId="48.[192,278,1070,1094]" captionTargetBox="[213,1446,171,1037]" captionTargetId="figure-11@48.[205,1452,168,1041]" captionTargetPageId="48" captionText="Figures 193–200: Distal segments of leg IV, retroventral view of Sri Lankan Buthidae and Chaerilidae genera. Buthoscorpio sarasinorum, male from locality 15CF (193), Charmus laneus, male from locality 15CO (194), Hottentotta tamulus, male from locality 15CK (195), Isometrus maculatus, female from locality 15CI (196), Isometrus thwaitesi, female from locality 15CO (197), Lychas srilankensis, male from locality 15CJ (198), Reddyanus basilicus, male from locality 15CS (199), and Chaerilus ceylonensis, male from locality 15CD (200)." figureDoi="http://doi.org/10.5281/zenodo.7124608" httpUri="https://zenodo.org/record/7124608/files/figure.png" pageId="55" pageNumber="54">Figs. 199</figureCitation>
, 
<figureCitation id="A0E32A4CA820FFA7FAC9F9459B05F914" box="[1289,1383,1682,1705]" captionStart="Figures 201–208" captionStartId="49.[192,278,1057,1081]" captionTargetBox="[204,1438,173,1022]" captionTargetId="figure-6@49.[192,1439,168,1028]" captionTargetPageId="49" captionText="Figures 201–208: Distal segments of legs, retroventral view of Sri Lankan Reddyanus. Figures 201–204. R. ceylonensis sp. n., male holotype from locality 15CI, legs I–IV. Figures 205–208. Leg IV. R. besucheti, male from locality 15CD (205), R. jayarathnei sp. n., male paratype (206), R. loebli, male from locality 15CG (207), and R. ranawanai sp. n., male holotype (208)." figureDoi="http://doi.org/10.5281/zenodo.7124612" httpUri="https://zenodo.org/record/7124612/files/figure.png" pageId="55" pageNumber="54">205–208</figureCitation>
); (ii) hemispermatophore with a relatively short, broad trunk, a short, broad, uncoiled flagellum, and a smaller, blunt basal lobe in 
<taxonomicName id="FFD84D4AA820FFA7FC1BF9399A26F8B8" authorityName="Ehrenberg" authorityYear="1828" box="[987,1092,1774,1797]" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA820FFA7FC1BF9399A26F8B8" box="[987,1092,1774,1797]" italics="true" pageId="55" pageNumber="54">Isometrus</emphasis>
</taxonomicName>
(
<figureCitation id="A0E32A4CA820FFA7FB99F9399A9BF8B8" box="[1113,1273,1774,1797]" captionStart="Figures 146–149" captionStartId="38.[192,278,1446,1470]" captionTargetBox="[194,1436,179,1409]" captionTargetId="figure-11@38.[192,1439,168,1417]" captionTargetPageId="38" captionText="Figures 146–149: Left hemispermatophore of Isometrus thwaitesi from locality 15CH. External (146), dorsal (147), internal (148) and ventral (149) views. Note: distal part of flagellum and basal part of trunk (foot) were damaged during dissection. Scale bar: 500 µm. Abbreviations: bl, basal lobe; f, flagellum; ml, median lobe; mlc, median lobe carina; pr, pars recta of flagellum; prf, pars reflecta of flagellum; t, trunk." figureDoi="http://doi.org/10.5281/zenodo.7124574" httpUri="https://zenodo.org/record/7124574/files/figure.png" pageId="55" pageNumber="54">Figs. 146–149</figureCitation>
), vs. a narrow trunk, a long, thin, coiled flagellum and a much enlarged, pointed basal lobe in 
<taxonomicName id="FFD84D4AA820FFA7FB62F8FC9B75F8FF" authorityName="Vachon" authorityYear="1972" box="[1186,1303,1835,1858]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA820FFA7FB62F8FC9B75F8FF" box="[1186,1303,1835,1858]" italics="true" pageId="55" pageNumber="54">Reddyanus</emphasis>
</taxonomicName>
(
<figureCitation id="A0E32A4CA820FFA7FAEBF8FC9D0AF8DC" captionStart="Figures 283–286" captionStartId="58.[192,278,1625,1649]" captionTargetBox="[201,1430,178,1594]" captionTargetId="figure-11@58.[192,1439,168,1596]" captionTargetPageId="58" captionText="Figures 283–286: Left hemispermatophore of Reddyanus basilicus from locality 15CR. External (283), dorsal (284), internal (285) and ventral (286) views. Scale bar: 500 µm. Abbreviations: bl, basal lobe; f, flagellum; ml, median lobe; mlc, median lobe carina; t, trunk." figureDoi="http://doi.org/10.5281/zenodo.7124636" httpUri="https://zenodo.org/record/7124636/files/figure.png" pageId="55" pageNumber="54">Figs. 283– 286</figureCitation>
). The form of the hemispermatophore of 
<taxonomicName id="FFD84D4AA820FFA7FAF0F89D9BFBF8DC" authorityName="Ehrenberg" authorityYear="1828" box="[1328,1433,1866,1889]" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA820FFA7FAF0F89D9BFBF8DC" box="[1328,1433,1866,1889]" italics="true" pageId="55" pageNumber="54">Isometrus</emphasis>
</taxonomicName>
appears more similar to that of 
<taxonomicName id="FFD84D4AA82EFFA9FDDCFBC49C07FB97" authorityName="C. L. Koch" authorityYear="1845" box="[540,613,1043,1066]" class="Arachnida" family="Buthidae" genus="Lychas" kingdom="Animalia" order="Scorpiones" pageId="57" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA82EFFA9FDDCFBC49C07FB97" box="[540,613,1043,1066]" italics="true" pageId="57" pageNumber="56">Lychas</emphasis>
</taxonomicName>
(
<figureCitation id="A0E32A4CA82EFFA9FDB6FBC49D71FB97" box="[630,787,1043,1066]" captionStart="Figures 183–186" captionStartId="45.[192,278,1659,1683]" captionTargetBox="[196,1422,175,1626]" captionTargetId="figure-6@45.[192,1439,168,1630]" captionTargetPageId="45" captionText="Figures 183–186: Right hemispermatophore of Lychas srilankensis from locality 15CN (presented as mirror image for comparison to other species). External (183), dorsal (184), internal (185) and ventral (186) views. Extracted after 3rd ecdysis. Hemispermatophores examined from a second male after 4th ecdysis were similar. Scale bar: 500 µm. Abbreviations: bl, basal lobe; f, flagellum; ml, median lobe; mlc, median lobe carina; pr, pars recta of flagellum; prf, pars reflecta of flagellum; t, trunk." figureDoi="http://doi.org/10.5281/zenodo.7124594" httpUri="https://zenodo.org/record/7124594/files/figure.png" pageId="57" pageNumber="56">Figs. 183–186</figureCitation>
), than to 
<taxonomicName id="FFD84D4AA82EFFA9FEE4FBE59FFBFBF4" authorityName="Vachon" authorityYear="1972" box="[292,409,1074,1097]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="57" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA82EFFA9FEE4FBE59FFBFBF4" box="[292,409,1074,1097]" italics="true" pageId="57" pageNumber="56">Reddyanus</emphasis>
</taxonomicName>
. Differences in trichobothriotaxy, pedipalp finger dentition, tarsal setation and hemispermatophore structure provide characters that are useful for higher level classification of buthid scorpions (
<bibRefCitation id="5C494B38A82EFFA9FF07FB7B9F07FB7E" author="Kovarik" bookContentInfo="170 pp." box="[199,357,1196,1219]" firstAuthor="Kovarik" journalOrPublisher="Clairon Production, Prague" pageId="57" pageNumber="56" refId="ref41793" refString="KOVARIK, F. 2009. Illustrated catalog of scorpions. Part I. Introductory remarks; keys to families and genera; subfamily Scorpioninae with keys to Heterometrus and Pandinus species. Clairon Production, Prague, 170 pp." title="Illustrated catalog of scorpions. Part I. Introductory remarks; keys to families and genera; subfamily Scorpioninae with keys to Heterometrus and Pandinus species" type="book" year="2009">Kovařík, 2009</bibRefCitation>
; 
<bibRefCitation id="5C494B38A82EFFA9FEB5FB7B9C75FB7E" author="Lamoral" box="[373,535,1196,1219]" firstAuthor="Lamoral" pageId="57" pageNumber="56" pagination="497 - 784" refId="ref42453" refString="LAMORAL, B. H. 1979. The scorpions of Namibia (Arachnida: Scorpionida). Annals of the Natal Museum, 23 (3): 497 - 784." type="journal article" year="1979">Lamoral, 1979</bibRefCitation>
; 
<bibRefCitation id="5C494B38A82EFFA9FDE4FB7B9CDBFB7E" author="Sissom" bookContentInfo="587 pp." box="[548,697,1196,1219]" editor="Polis, G. A." firstAuthor="Sissom" journalOrPublisher="Stanford: Stanford University Press" pageId="57" pageNumber="56" pagination="64 - 160" refId="ref44349" refString="SISSOM, W. D. 1990. Systematics, biogeography and paleontology. Pp. 64 - 160. In: Polis, G. A. (ed.), The Biology of Scorpions. Stanford: Stanford University Press, 587 pp." title="Systematics, biogeography and paleontology" type="book chapter" volumeTitle="The Biology of Scorpions" year="1990">Sissom, 1990</bibRefCitation>
; 
<bibRefCitation id="5C494B38A82EFFA9FD05FB7B9E94FB5F" author="Stahnke" firstAuthor="Stahnke" journalOrPublisher="Entomological News" pageId="57" pageNumber="56" pagination="121 - 133" part="83" refId="ref44469" refString="STAHNKE, H. L. 1972. A key to the genera of Buthidae (Scorpionida). Entomological News, 83 (5): 121 - 133." title="A key to the genera of Buthidae (Scorpionida)" type="journal article" year="1972">Stahnke, 1972</bibRefCitation>
; Stockwell, 198; 
<bibRefCitation id="5C494B38A82EFFA9FE00FB1C9C38FB5F" author="Vachon" box="[448,602,1227,1250]" firstAuthor="Vachon" journalOrPublisher="Archives de l'Institut Pasteur d'Algerie" pageId="57" pageNumber="56" pagination="172 - 177" part="30" refId="ref44827" refString="VACHON, M. 1952 a. Complements a la description du petit scorpion Mauritanien Lissothus occidentalis Vachon, 1950 famille des Buthidae E. Simon. Archives de l'Institut Pasteur d'Algerie, 30 (2): 172 - 177." title="Complements a la description du petit scorpion Mauritanien Lissothus occidentalis Vachon, 1950 famille des Buthidae E. Simon" type="journal article" year="1952">Vachon, 1952</bibRefCitation>
). Taken together, these differences support our taxonomic decision to elevate 
<taxonomicName id="FFD84D4AA82EFFA9FF33FADF9F0AFAA2" authorityName="Vachon" authorityYear="1972" box="[243,360,1288,1311]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="57" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA82EFFA9FF33FADF9F0AFAA2" box="[243,360,1288,1311]" italics="true" pageId="57" pageNumber="56">Reddyanus</emphasis>
</taxonomicName>
to a genus distinct from 
<taxonomicName id="FFD84D4AA82EFFA9FDB1FADF9CB8FAA2" authorityName="Ehrenberg" authorityYear="1828" box="[625,730,1288,1311]" class="Arachnida" family="Buthidae" genus="Isometrus" kingdom="Animalia" order="Scorpiones" pageId="57" pageNumber="56" phylum="Arthropoda" rank="genus">
<emphasis id="0AACEADBA82EFFA9FDB1FADF9CB8FAA2" box="[625,730,1288,1311]" italics="true" pageId="57" pageNumber="56">Isometrus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="6CA76641A820FFA7FF00FA579C51FA5F" ID-DOI="http://doi.org/10.5281/zenodo.7124630" ID-Zenodo-Dep="7124630" httpUri="https://zenodo.org/record/7124630/files/figure.png" pageId="55" pageNumber="54" startId="55.[192,278,1408,1432]" targetBox="[191,1440,167,1380]" targetPageId="55">
<paragraph id="386736C9A820FFA7FF00FA579C51FA5F" blockId="55.[192,1441,1408,1506]" pageId="55" pageNumber="54">
<emphasis id="0AACEADBA820FFA7FF00FA579FE4FA25" bold="true" box="[192,390,1408,1432]" pageId="55" pageNumber="54">Figures 264–269:</emphasis>
<taxonomicName id="FFD84D4AA820FFA7FE4FFA549C35FA25" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Karsch" baseAuthorityYear="1879" box="[399,599,1411,1432]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="55" pageNumber="54" phylum="Arthropoda" rank="species" species="basilicus">
<emphasis id="0AACEADBA820FFA7FE4FFA549C35FA25" box="[399,599,1411,1432]" italics="true" pageId="55" pageNumber="54">Reddyanus basilicus</emphasis>
</taxonomicName>
. 
<emphasis id="0AACEADBA820FFA7FDA6FA559D76FA2A" bold="true" box="[614,788,1410,1432]" pageId="55" pageNumber="54">Figures 264, 266</emphasis>
. Male from locality 15CS, chelicerae, carapace and tergites I–III (264), and sternopectinal region and sternite III (266). 
<emphasis id="0AACEADBA820FFA7FD1EFA4C9DE8FA0D" bold="true" box="[734,906,1435,1457]" pageId="55" pageNumber="54">Figures 265, 267</emphasis>
. Female from locality 15CS, chelicerae, carapace and tergites I–III (265) and sternopectinal region and sternites III–IV (267). 
<emphasis id="0AACEADBA820FFA7FC4AFA639A56FA74" bold="true" box="[906,1076,1460,1482]" pageId="55" pageNumber="54">Figures 268–269</emphasis>
. Male from locality 15CR, chelicera dorsal (268) and ventral (269) aspects.
</paragraph>
</caption>
<caption id="6CA76641A82FFFA8FF00F8F89F8CF8C5" ID-DOI="http://doi.org/10.5281/zenodo.7124632" ID-Zenodo-Dep="7124632" httpUri="https://zenodo.org/record/7124632/files/figure.png" pageId="56" pageNumber="55" startId="56.[192,278,1839,1863]" targetBox="[215,1432,184,1819]" targetPageId="56">
<paragraph id="386736C9A82FFFA8FF00F8F89F8CF8C5" blockId="56.[192,1440,1839,1912]" pageId="56" pageNumber="55">
<emphasis id="0AACEADBA82FFFA8FF00F8F89FEAF8FA" bold="true" box="[192,392,1839,1863]" pageId="56" pageNumber="55">Figures 270–275:</emphasis>
<taxonomicName id="FFD84D4AA82FFFA8FE53F8E59C3EF8FA" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Karsch" baseAuthorityYear="1879" box="[403,604,1842,1863]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="56" pageNumber="55" phylum="Arthropoda" rank="species" species="basilicus">
<emphasis id="0AACEADBA82FFFA8FE53F8E59C3EF8FA" box="[403,604,1842,1863]" italics="true" pageId="56" pageNumber="55">Reddyanus basilicus</emphasis>
</taxonomicName>
from locality 15CS. 
<emphasis id="0AACEADBA82FFFA8FCFBF8E69D85F8FB" bold="true" box="[827,999,1841,1863]" pageId="56" pageNumber="55">Figures 270–272</emphasis>
. Male, metasoma and telson, lateral (270), ventral (271), and dorsal (272) views. 
<emphasis id="0AACEADBA82FFFA8FDFEF89D9C85F8E2" bold="true" box="[574,743,1866,1888]" pageId="56" pageNumber="55">Figures 273–275</emphasis>
. Female, metasoma and telson, lateral (273), ventral (274), and dorsal (275) views. Scale bar: 10 mm.
</paragraph>
</caption>
<caption id="6CA76641A82EFFA9FF00FCAB9F0AFC63" ID-DOI="http://doi.org/10.5281/zenodo.7124634" ID-Zenodo-Dep="7124634" httpUri="https://zenodo.org/record/7124634/files/figure.png" pageId="57" pageNumber="56" startId="57.[192,278,892,916]" targetBox="[198,1431,222,869]" targetPageId="57">
<paragraph id="386736C9A82EFFA9FF00FCAB9F0AFC63" blockId="57.[192,1441,892,990]" pageId="57" pageNumber="56">
<emphasis id="0AACEADBA82EFFA9FF00FCAB9FEAFC29" bold="true" box="[192,392,892,916]" pageId="57" pageNumber="56">Figures 276–282:</emphasis>
<taxonomicName id="FFD84D4AA82EFFA9FE54FCA89C3FFC29" authorityName="Kovařík &amp; Lowe &amp; Ranawana &amp; Hoferek &amp; Š" authorityYear="2016" baseAuthorityName="Karsch" baseAuthorityYear="1879" box="[404,605,895,916]" class="Arachnida" family="Buthidae" genus="Reddyanus" kingdom="Animalia" order="Scorpiones" pageId="57" pageNumber="56" phylum="Arthropoda" rank="species" species="basilicus">
<emphasis id="0AACEADBA82EFFA9FE54FCA89C3FFC29" box="[404,605,895,916]" italics="true" pageId="57" pageNumber="56">Reddyanus basilicus</emphasis>
</taxonomicName>
from locality 15CS. 
<emphasis id="0AACEADBA82EFFA9FCFEFCA99D89FC2E" bold="true" box="[830,1003,894,916]" pageId="57" pageNumber="56">Figures 276–277</emphasis>
. Female. Pedipalp chela, dorsal (276) and external (277) views. 
<emphasis id="0AACEADBA82EFFA9FE5AFC409C27FC11" bold="true" box="[410,581,919,941]" pageId="57" pageNumber="56">Figures 278–282</emphasis>
. Male. Pedipalp chela, dorsal (278) and external (279) views. Pedipalp patella, dorsal (280) and external (281) views. Pedipalp femur and trochanter dorsal view (282). The trichobothrial pattern is indicated in Figures 279–282.
</paragraph>
</caption>
<subSubSection id="70C26542A82EFFA9FF00FA919F33FAC6" pageId="57" pageNumber="56" type="distribution">
<paragraph id="386736C9A82EFFA9FF00FA919F33FAC6" blockId="57.[192,802,1350,1403]" pageId="57" pageNumber="56">
DISTRIBUTION. Oriental region from 
<collectingCountry id="40CF7659A82EFFA9FD8CFA919CE1FAE0" box="[588,643,1350,1373]" name="India" pageId="57" pageNumber="56">India</collectingCountry>
, 
<collectingCountry id="40CF7659A82EFFA9FD51FA919CB2FAE0" box="[657,720,1350,1373]" name="China" pageId="57" pageNumber="56">China</collectingCountry>
(
<collectingRegion id="FA1CF82BA82EFFA9FD20FA919D7AFAE0" box="[736,792,1350,1373]" country="China" name="Xizang" pageId="57" pageNumber="56">Tibet</collectingRegion>
) to Melanesia.
</paragraph>
</subSubSection>
</treatment>
</document>