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<document id="7AC4AD8090AD7778C7B27891D2DDF966" ID-CLB-Dataset="31822" ID-DOI="10.11646/zootaxa.4337.2.1" ID-GBIF-Dataset="bca0e227-e4fb-4f93-b99b-c8794b19de8d" ID-ISSN="1175-5326" ID-Zenodo-Dep="1018631" ID-ZooBank="A712D90D-FCD1-49C9-928F-21C95814815B" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1508314185452" checkinUser="plazi" docAuthor="Behan-Pelletier, Valerie M." docDate="2017" docId="9D02C578FF93FFACFF12C89FFA01F8A0" docLanguage="en" docName="zootaxa.4337.2.1.pdf" docOrigin="Zootaxa 4337 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tegoribates Ewing 1917" docType="treatment" docVersion="7" lastPageNumber="166" masterDocId="613BBD00FF9EFFA3FF85CC7CFF9CFFD0" masterDocTitle="Tegoribatidae of North America, with proposal of Protectoribates gen. nov., and new species (Acari, Oribatida, Tegoribatidae)" pageNumber="164" updateTime="1698469699374" updateUser="ExternalLinkService">
<mods:mods id="63B3928F57E0BB45F4A5D71D5D18CBF7" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="4739B24967D5DE6AE78AA8B14D07F48E">
<mods:title id="645B9A28E5B543F0CEE22843C219B7DE">Tegoribatidae of North America, with proposal of Protectoribates gen. nov., and new species (Acari, Oribatida, Tegoribatidae)</mods:title>
</mods:titleInfo>
<mods:name id="AE9E8A515C8161208149FAADA6519129" type="personal">
<mods:role id="B49F3F6A214B436B0DB7300FE9285F18">
<mods:roleTerm id="2370EAE392E71F2CF03F492132E6BBAD">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="4EDC19DD2EF906963DBA82626059A64E">Behan-Pelletier, Valerie M.</mods:namePart>
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<mods:typeOfResource id="12B84FFAF8D497FDEBC0F512497AAAE2">text</mods:typeOfResource>
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<mods:title id="704A60E857D40B58CD9816C354671233">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="DE6B8ABFD57843E6EF1937069C5E7D8F">
<mods:date id="1FED999D123E66A1FC80D46BFAD7FC66">2017</mods:date>
<mods:detail id="A8D294526B162113936D44ED2E6719BB" type="pubDate">
<mods:number id="7991206F71E8915844E5E730079CB5AD">2017-10-18</mods:number>
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<mods:detail id="2E0FF0D8A3973E986CFC04A50410E662" type="volume">
<mods:number id="2AB5961351ADDB50F9DFFAE62154A511">4337</mods:number>
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<mods:detail id="AE4A363632B79557621A84C648E82E9D" type="issue">
<mods:number id="F388E15FAA8F98A8722BDEADF89CB48D">2</mods:number>
</mods:detail>
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<mods:classification id="399B0F1A021A335CC9AD8D4314FE1848">journal volume</mods:classification>
<mods:identifier id="7F045026743FBA5BD8B6C07125B1EFDB" type="CLB-Dataset">31822</mods:identifier>
<mods:identifier id="F68867842C0DE7B4EE36A84B95DA00BD" type="DOI">10.11646/zootaxa.4337.2.1</mods:identifier>
<mods:identifier id="C21CCCB40E5CECA270659464F4F6B253" type="GBIF-Dataset">bca0e227-e4fb-4f93-b99b-c8794b19de8d</mods:identifier>
<mods:identifier id="4C1010886F6EDAA1A89F619C9ED1B71F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="AAD8BD09A3465830A42A8D6EA36150E6" type="Zenodo-Dep">1018631</mods:identifier>
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</mods:mods>
<treatment id="9D02C578FF93FFACFF12C89FFA01F8A0" ID-GBIF-Taxon="135450335" LSID="urn:lsid:plazi:treatment:9D02C578FF93FFACFF12C89FFA01F8A0" httpUri="http://treatment.plazi.org/id/9D02C578FF93FFACFF12C89FFA01F8A0" lastPageId="15" lastPageNumber="166" pageId="13" pageNumber="164">
<paragraph id="1514746EFF93FFAEFF12C89FFD14FA90" blockId="13.[151,450,1251,1277]" lastBlockId="13.[151,1055,1322,1375]" pageId="13" pageNumber="164">
<subSubSection id="5DB127E5FF93FFAEFF12C89FFE5EFB2D" box="[151,450,1251,1277]" pageId="13" pageNumber="164" type="nomenclature">
<taxonomicName id="D2AB0FEDFF93FFAEFF12C89FFE5EFB2D" ID-CoL="7TBD" authority="Ewing, 1917" authorityName="Ewing" authorityYear="1917" box="[151,450,1251,1277]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF93FFAEFF12C89FFEBEFB2D" bold="true" box="[151,290,1251,1277]" italics="true" pageId="13" pageNumber="164">Tegoribates</emphasis>
Ewing, 1917
</taxonomicName>
</subSubSection>
<subSubSection id="5DB127E5FF93FFAEFF12C956FD14FA90" box="[151,648,1322,1344]" pageId="13" pageNumber="164" type="materials_examined">
<emphasis id="27DFA87CFF93FFAEFF12C956FEBAFA90" bold="true" box="[151,294,1322,1344]" pageId="13" pageNumber="164">Type species:</emphasis>
<taxonomicName id="D2AB0FEDFF93FFAEFEAEC956FD14FA90" authority="Ewing, 1917" authorityName="Ewing" authorityYear="1917" box="[299,648,1322,1344]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="subniger">
<emphasis id="27DFA87CFF93FFAEFEAEC956FD9DFA90" box="[299,513,1322,1344]" italics="true" pageId="13" pageNumber="164">Tegoribates subniger</emphasis>
<bibRefCitation id="713A099FFF93FFAEFD82C956FD14FA90" author="Ewing, H. E." box="[519,648,1322,1344]" journalOrPublisher="Annals of The Entomological Society of America" pageId="13" pageNumber="164" pagination="117 - 132" part="10" publicationUrl="https://doi.org/10.1093/aesa/10.2.117" refString="Ewing, H. E. (1917) A synopsis of the genera of beetle mites with special reference to the North American fauna. Annals of The Entomological Society of America, 10, 117 - 132. https: // doi. org / 10.1093 / aesa / 10.2.117" title="A synopsis of the genera of beetle mites with special reference to the North American fauna" type="journal article" year="1917">Ewing, 1917</bibRefCitation>
</taxonomicName>
</subSubSection>
</paragraph>
<paragraph id="1514746EFF93FFAEFF12C934FB83FA8E" blockId="13.[151,1055,1322,1375]" box="[151,1055,1352,1375]" pageId="13" pageNumber="164">
<emphasis id="27DFA87CFF93FFAEFF12C934FF60FA8E" bold="true" box="[151,252,1352,1374]" pageId="13" pageNumber="164">Synonym</emphasis>
:
<taxonomicName id="D2AB0FEDFF93FFAEFE8CC934FDADFA8E" authority="Sellnick, 1920" authorityName="Sellnick" authorityYear="1920" box="[265,561,1352,1374]" class="Arachnida" family="Tegoribatidae" genus="Lepidoribates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="164" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF93FFAEFE8CC934FE0AFA8E" box="[265,406,1352,1374]" italics="true" pageId="13" pageNumber="164">Lepidoribates</emphasis>
<bibRefCitation id="713A099FFF93FFAEFE19C934FDADFA8E" author="Sellnick, M." box="[412,561,1352,1374]" journalOrPublisher="Schriften der physik. - okonom. Gesellschaft zu Konigsberg, 61 - 62" pageId="13" pageNumber="164" pagination="35 - 42" refString="Sellnick, M. (1920) Neue und seltene Oribatiden aus Deutschland. Schriften der physik. - okonom. Gesellschaft zu Konigsberg, 61 - 62, 35 - 42." title="Neue und seltene Oribatiden aus Deutschland" type="book chapter" year="1920">Sellnick, 1920</bibRefCitation>
</taxonomicName>
. Type species:
<taxonomicName id="D2AB0FEDFF93FFAEFD4FC936FB83FA8E" authority="C. L. Koch, 1844" authorityName="C. L. Koch" authorityYear="1844" box="[714,1055,1352,1375]" class="Arachnida" family="Galumnidae" genus="Zetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="latirostris">
<emphasis id="27DFA87CFF93FFAEFD4FC936FCF5FA8E" box="[714,873,1352,1375]" italics="true" pageId="13" pageNumber="164">Zetes latirostris</emphasis>
<bibRefCitation id="713A099FFF93FFAEFCEAC934FB83FA8E" author="Koch, C. L." bookContentInfo="13 pp." box="[879,1055,1352,1374]" journalOrPublisher="Pustet, Regensburg" pageId="13" pageNumber="164" refString="Koch, C. L. (1844) Deutschlands Crustaceen, Myriapoden und Arachniden. Heft 38. F. Pustet, Regensburg, 13 pp." title="Deutschlands Crustaceen, Myriapoden und Arachniden. Heft 38. F" type="book" year="1844">C. L. Koch, 1844</bibRefCitation>
</taxonomicName>
</paragraph>
<paragraph id="1514746EFF93FFAEFF12C9FAFE17FA4C" blockId="13.[151,1341,1414,1650]" box="[151,395,1414,1436]" pageId="13" pageNumber="164">Included extant species:</paragraph>
<subSubSection id="5DB127E5FF93FFAEFF12C9D8FC84F9A2" pageId="13" pageNumber="164" type="reference_group">
<paragraph id="1514746EFF93FFAEFF12C9D8FC8BFA6A" blockId="13.[151,1341,1414,1650]" box="[151,791,1444,1466]" pageId="13" pageNumber="164">
<taxonomicName id="D2AB0FEDFF93FFAEFF12C9D8FE6BFA6A" ID-CoL="7BMSC" authority="Ewing, 1917" authorityName="Ewing" authorityYear="1917" box="[151,503,1444,1466]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="subniger">
<emphasis id="27DFA87CFF93FFAEFF12C9D8FEF0FA6A" box="[151,364,1444,1466]" italics="true" pageId="13" pageNumber="164">Tegoribates subniger</emphasis>
<bibRefCitation id="713A099FFF93FFAEFEF6C9D8FE6BFA6A" author="Ewing, H. E." box="[371,503,1444,1466]" journalOrPublisher="Annals of The Entomological Society of America" pageId="13" pageNumber="164" pagination="117 - 132" part="10" publicationUrl="https://doi.org/10.1093/aesa/10.2.117" refString="Ewing, H. E. (1917) A synopsis of the genera of beetle mites with special reference to the North American fauna. Annals of The Entomological Society of America, 10, 117 - 132. https: // doi. org / 10.1093 / aesa / 10.2.117" title="A synopsis of the genera of beetle mites with special reference to the North American fauna" type="journal article" year="1917">Ewing, 1917</bibRefCitation>
</taxonomicName>
; known from
<collectingRegion id="D76FBA8CFF93FFAEFD01C9D8FD4EFA6A" box="[644,722,1444,1466]" country="United States of America" name="Indiana" pageId="13" pageNumber="164">Indiana</collectingRegion>
,
<collectingCountry id="6DBC34FEFF93FFAEFD58C9D8FC8FFA6A" box="[733,787,1444,1466]" name="United States of America" pageId="13" pageNumber="164">USA</collectingCountry>
.
</paragraph>
<paragraph id="1514746EFF93FFAEFF12C9BFFCD2FA28" blockId="13.[151,1341,1414,1650]" pageId="13" pageNumber="164">
<taxonomicName id="D2AB0FEDFF93FFAEFF12C9BFFDB6FA09" ID-CoL="7BN4H" authority="Hammer, 1958" authorityName="Hammer. Setae (Roman" authorityYear="1958" box="[151,554,1475,1497]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="27DFA87CFF93FFAEFF12C9BFFE16FA09" box="[151,394,1475,1497]" italics="true" pageId="13" pageNumber="164">Tegoribates americanus</emphasis>
<bibRefCitation id="713A099FFF93FFAEFE15C9B8FDB6FA09" author="Hammer, M." box="[400,554,1475,1497]" journalOrPublisher="Biologiske Skrifter Det Kongelige Danske Videnskabernes Selskab" pageId="13" pageNumber="164" pagination="1 - 129" part="10" refString="Hammer, M. (1958) Investigations on the oribatid fauna of the Andes Mountains. I. The Argentine and Bolivia. Biologiske Skrifter Det Kongelige Danske Videnskabernes Selskab, 10, 1 - 129." title="Investigations on the oribatid fauna of the Andes Mountains. I. The Argentine and Bolivia" type="journal article" year="1958">Hammer, 1958</bibRefCitation>
</taxonomicName>
; known from the
<location id="107422B5FF93FFAEFD59C9B8FC15FA09" LSID="urn:lsid:plazi:treatment:9D02C578FF93FFACFF12C89FFA01F8A0:107422B5FF93FFAEFD59C9B8FC15FA09" box="[732,905,1475,1497]" country="Argentina" name="western Nearctic" pageId="13" pageNumber="164">western Nearctic</location>
, Russian Far East,
<collectingCountry id="6DBC34FEFF93FFAEFBCFC9BFFB33FA09" box="[1098,1199,1475,1497]" name="Argentina" pageId="13" pageNumber="164">Argentina</collectingCountry>
and
<collectingCountry id="6DBC34FEFF93FFAEFB65C9BFFAA5FA09" box="[1248,1337,1475,1497]" name="Vietnam" pageId="13" pageNumber="164">Vietnam</collectingCountry>
.
<taxonomicName id="D2AB0FEDFF93FFAEFF12C99EFD80FA28" ID-CoL="7BMSF" authority="Woolley, 1965" authorityName="Woolley" authorityYear="1965" box="[151,540,1506,1528]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="bryophilus">
<emphasis id="27DFA87CFF93FFAEFF12C99EFE1CFA28" box="[151,384,1506,1528]" italics="true" pageId="13" pageNumber="164">Tegoribates bryophilus</emphasis>
<bibRefCitation id="713A099FFF93FFAEFE03C99EFD80FA28" author="Woolley, T. A." box="[390,540,1506,1528]" journalOrPublisher="Transaction of the American Microscopical Society" pageId="13" pageNumber="164" pagination="409 – 412" part="84" publicationUrl="https://doi.org/10.2307/3224729" refString="Woolley, T. A. (1965) A New Species of Tegoribates from Colorado with Notes on the Genus (Acarina: Oribatei, Tegoribatidae). Transaction of the American Microscopical Society, 84, 409 - 412. https: // doi. org / 10.2307 / 3224729" title="A New Species of Tegoribates from Colorado with Notes on the Genus (Acarina: Oribatei, Tegoribatidae)" type="journal article" year="1965">Woolley, 1965</bibRefCitation>
</taxonomicName>
; known from
<collectingRegion id="D76FBA8CFF93FFAEFD2CC99EFC96FA28" box="[681,778,1506,1528]" country="United States of America" name="Colorado" pageId="13" pageNumber="164">Colorado</collectingRegion>
,
<collectingCountry id="6DBC34FEFF93FFAEFC91C99EFCD6FA28" box="[788,842,1506,1528]" name="United States of America" pageId="13" pageNumber="164">USA</collectingCountry>
.
</paragraph>
<paragraph id="1514746EFF93FFAEFF12CA7CFCA3F9E5" blockId="13.[151,1341,1414,1650]" pageId="13" pageNumber="164">
<taxonomicName id="D2AB0FEDFF93FFAEFF12CA7CFDCDF9C6" ID-CoL="7BN4F" authority="(Koch, 1844) (Zetes)" authorityName="Koch" authorityYear="1844" baseAuthorityName="Koch" baseAuthorityYear="1844" box="[151,593,1536,1559]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="latirostris">
<emphasis id="27DFA87CFF93FFAEFF12CA7CFEEBF9C6" box="[151,375,1536,1558]" italics="true" pageId="13" pageNumber="164">Tegoribates latirostris</emphasis>
(
<bibRefCitation id="713A099FFF93FFAEFE01CA7CFE61F9C6" author="Koch, C. L." bookContentInfo="13 pp." box="[388,509,1536,1558]" journalOrPublisher="Pustet, Regensburg" pageId="13" pageNumber="164" refString="Koch, C. L. (1844) Deutschlands Crustaceen, Myriapoden und Arachniden. Heft 38. F. Pustet, Regensburg, 13 pp." title="Deutschlands Crustaceen, Myriapoden und Arachniden. Heft 38. F" type="book" year="1844">Koch, 1844</bibRefCitation>
) (
<emphasis id="27DFA87CFF93FFAEFD96CA7EFDD4F9C7" box="[531,584,1538,1559]" italics="true" pageId="13" pageNumber="164">Zetes</emphasis>
)
</taxonomicName>
(=
<taxonomicName id="D2AB0FEDFF93FFAEFDE8CA7CFC0CF9C6" authority="Coggi, 1900" authorityName="Coggi" authorityYear="1900" box="[621,912,1536,1558]" class="Arachnida" family="Tegoribatidae" genus="Oribata" higherTaxonomySource="CoL" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="alzanii">
<emphasis id="27DFA87CFF93FFAEFDE8CA7CFC95F9C6" box="[621,777,1536,1558]" italics="true" pageId="13" pageNumber="164">Oribata alzanii</emphasis>
<bibRefCitation id="713A099FFF93FFAEFC8ACA7CFC0CF9C6" author="Coggi, A." box="[783,912,1536,1558]" journalOrPublisher="Bulletino (Bolletino) della Societa Zoologica Italiana" pageId="13" pageNumber="164" pagination="309 - 324" part="32" refString="Coggi, A. (1900) Nuovi Oribatidi italiani. Bulletino (Bolletino) della Societa Zoologica Italiana, 32, 309 - 324." title="Nuovi Oribatidi italiani" type="journal article" year="1900">Coggi, 1900</bibRefCitation>
</taxonomicName>
); known from the Palaearctic.
<taxonomicName id="D2AB0FEDFF93FFAEFF12CA63FDACF9E5" authority="Engelbrecht, 1986" authorityName="Engelbrecht" authorityYear="1986" box="[151,560,1567,1589]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="montana">
<emphasis id="27DFA87CFF93FFAEFF12CA63FEF0F9E5" box="[151,364,1567,1589]" italics="true" pageId="13" pageNumber="164">Tegoribates montana</emphasis>
<bibRefCitation id="713A099FFF93FFAEFEF6CA63FDACF9E5" author="Engelbrecht, C. M." box="[371,560,1567,1589]" journalOrPublisher="Navorsinge van die Nasionale Museum, Bloemfontein" pageId="13" pageNumber="164" pagination="169 - 250" part="5" refString="Engelbrecht, C. M. (1986) New oribatid taxa and distribution records predominantly from Southern Africa (Acari: Oribatei: Oribatelloidea. Navorsinge van die Nasionale Museum, Bloemfontein, 5 (9), 169 - 250." title="New oribatid taxa and distribution records predominantly from Southern Africa (Acari: Oribatei: Oribatelloidea" type="journal article" year="1986">Engelbrecht, 1986</bibRefCitation>
</taxonomicName>
; known from
<collectingCountry id="6DBC34FEFF93FFAEFD39CA63FCA3F9E5" box="[700,831,1567,1589]" name="South Africa" pageId="13" pageNumber="164">South Africa</collectingCountry>
</paragraph>
<paragraph id="1514746EFF93FFAEFF12CA42FCD1F984" blockId="13.[151,1341,1414,1650]" box="[151,845,1598,1620]" pageId="13" pageNumber="164">
<taxonomicName id="D2AB0FEDFF93FFAEFF12CA42FDA2F984" authority="Engelbrecht, 1986" authorityName="Engelbrecht" authorityYear="1986" box="[151,574,1598,1620]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="natalensis">
<emphasis id="27DFA87CFF93FFAEFF12CA42FEE6F984" box="[151,378,1598,1620]" italics="true" pageId="13" pageNumber="164">Tegoribates natalensis</emphasis>
<bibRefCitation id="713A099FFF93FFAEFE04CA42FDA2F984" author="Engelbrecht, C. M." box="[385,574,1598,1620]" journalOrPublisher="Navorsinge van die Nasionale Museum, Bloemfontein" pageId="13" pageNumber="164" pagination="169 - 250" part="5" refString="Engelbrecht, C. M. (1986) New oribatid taxa and distribution records predominantly from Southern Africa (Acari: Oribatei: Oribatelloidea. Navorsinge van die Nasionale Museum, Bloemfontein, 5 (9), 169 - 250." title="New oribatid taxa and distribution records predominantly from Southern Africa (Acari: Oribatei: Oribatelloidea" type="journal article" year="1986">Engelbrecht, 1986</bibRefCitation>
</taxonomicName>
; known from South Africa
</paragraph>
<paragraph id="1514746EFF93FFAEFF12CA20FC84F9A2" blockId="13.[151,1341,1414,1650]" box="[151,792,1628,1650]" pageId="13" pageNumber="164">
<taxonomicName id="D2AB0FEDFF93FFAEFF12CA20FD95F9A2" authority="Engelbrecht, 1986" authorityName="Engelbrecht" authorityYear="1986" box="[151,521,1628,1650]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="164" phylum="Arthropoda" rank="species" species="nuda">
<emphasis id="27DFA87CFF93FFAEFF12CA20FEDBF9A2" box="[151,327,1628,1650]" italics="true" pageId="13" pageNumber="164">Tegoribates nuda</emphasis>
<bibRefCitation id="713A099FFF93FFAEFEC9CA20FD95F9A2" author="Engelbrecht, C. M." box="[332,521,1628,1650]" journalOrPublisher="Navorsinge van die Nasionale Museum, Bloemfontein" pageId="13" pageNumber="164" pagination="169 - 250" part="5" refString="Engelbrecht, C. M. (1986) New oribatid taxa and distribution records predominantly from Southern Africa (Acari: Oribatei: Oribatelloidea. Navorsinge van die Nasionale Museum, Bloemfontein, 5 (9), 169 - 250." title="New oribatid taxa and distribution records predominantly from Southern Africa (Acari: Oribatei: Oribatelloidea" type="journal article" year="1986">Engelbrecht, 1986</bibRefCitation>
</taxonomicName>
; known from South Africa
</paragraph>
</subSubSection>
<subSubSection id="5DB127E5FF93FFADFF12CAE3FA90FE5A" lastPageId="14" lastPageNumber="165" pageId="13" pageNumber="164" type="diagnosis">
<paragraph id="1514746EFF93FFADFF12CAE3FC0BFF05" blockId="13.[151,1437,1695,2008]" lastBlockId="14.[151,1437,151,2018]" lastPageId="14" lastPageNumber="165" pageId="13" pageNumber="164">
<emphasis id="27DFA87CFF93FFAEFF12CAE3FEF5F968" bold="true" box="[151,361,1695,1720]" pageId="13" pageNumber="164">Revised diagnosis</emphasis>
:
<emphasis id="27DFA87CFF93FFAEFEF2CAE3FE22F968" box="[375,446,1695,1720]" italics="true" pageId="13" pageNumber="164">Adults</emphasis>
: With character states of
<taxonomicName id="D2AB0FEDFF93FFAEFD5FCAE3FCE8F968" authorityName="Grandjean" authorityYear="1954" box="[730,884,1695,1720]" class="Arachnida" family="Tegoribatidae" kingdom="Animalia" order="Sarcoptiformes" pageId="13" pageNumber="164" phylum="Arthropoda" rank="family">Tegoribatidae</taxonomicName>
(
<bibRefCitation id="713A099FFF93FFAEFC04CAE3FBDEF968" author="Grandjean, F." box="[897,1090,1695,1720]" journalOrPublisher="Bulletin du Museum national d´Histoire naturelle, 2 e Serie" pageId="13" pageNumber="164" pagination="286 - 293" part="25" refString="Grandjean, F. (1953) Observations sur les Oribates (26 e serie). Bulletin du Museum national d´Histoire naturelle, 2 e Serie, 25 (3), 286 - 293." title="Observations sur les Oribates (26 e serie)" type="journal article" year="1953">Grandjean (1953</bibRefCitation>
,
<bibRefCitation id="713A099FFF93FFAEFBC8CAE3FB15F968" author="Grandjean, F." box="[1101,1161,1695,1720]" journalOrPublisher="Bulletin de la Societe Zoologique de France" pageId="13" pageNumber="164" pagination="421 - 446" part="78" refString="Grandjean, F. (1954) Essai de classification des Oribates (Acariens). Bulletin de la Societe Zoologique de France, 78, 421 - 446." title="Essai de classification des Oribates (Acariens)" type="journal article" year="1954">1954</bibRefCitation>
),
<bibRefCitation id="713A099FFF93FFAEFB19CAE3FAD7F968" author="Woolley, T. A." box="[1180,1355,1695,1720]" journalOrPublisher="Transaction of the American Microscopical Society" pageId="13" pageNumber="164" pagination="409 – 412" part="84" publicationUrl="https://doi.org/10.2307/3224729" refString="Woolley, T. A. (1965) A New Species of Tegoribates from Colorado with Notes on the Genus (Acarina: Oribatei, Tegoribatidae). Transaction of the American Microscopical Society, 84, 409 - 412. https: // doi. org / 10.2307 / 3224729" title="A New Species of Tegoribates from Colorado with Notes on the Genus (Acarina: Oribatei, Tegoribatidae)" type="journal article" year="1965">Woolley (1965)</bibRefCitation>
<bibRefCitation id="713A099FFF93FFAEFAD4CAE3FEF2F90C" author="Fredes, N. A. &amp; Martinez, P. A." journalOrPublisher="Zootaxa" pageId="13" pageNumber="164" pagination="191 - 200" part="4098" publicationUrl="https://doi.org/10.11646/zootaxa.4098.1.10" refString="Fredes, N. A. &amp; Martinez, P. A. (2016) Revision of Physobates (Acari, Oribatida, Tegoribatidae) and redescription of Physobates spinipes Hammer, 1962 with comments on tegoribatid genera. Zootaxa, 4098 (1), 191 - 200. https: // doi. org / 10.11646 / zootaxa. 4098.1.10" title="Revision of Physobates (Acari, Oribatida, Tegoribatidae) and redescription of Physobates spinipes Hammer, 1962 with comments on tegoribatid genera" type="journal article" year="2016">Fredes &amp; Martínez (2016)</bibRefCitation>
, and see below). Prodorsum with lamellae fused medially. Genal tooth absent; open taenidium present in usual position of genal incision, extending to acetabulum I. Humerosejugal organs
<emphasis id="27DFA87CFF93FFAEFB49CA94FABDF8D0" box="[1228,1313,1767,1792]" italics="true" pageId="13" pageNumber="164">Am, Ah</emphasis>
present as porose area;
<emphasis id="27DFA87CFF93FFAEFEB4CB77FED6F8F4" box="[305,330,1803,1828]" italics="true" pageId="13" pageNumber="164">Al</emphasis>
absent. Dorsophragmata fused medially, elongated. Pedotectum I large, extending to base of bothridium. Bothridium cup-shaped, positioned laterally, closely adjacent to lamella, with anterior pointed scale. Notogaster with continuous posterior tectum; pteromorphs with hinge; lenticulus present or absent. Notogastral setation 10 pairs; setae apobasic. Octotaxic system developed as porose areas or saccules or tubules. Genital setation 6 pairs. Postanal porose area present. Subcapitular mentum with complete tectum. Axillary saccule of subcapitulum present. On palptarsus eupathidium
<emphasis id="27DFA87CFF93FFAEFD48CBBCFD60F807" box="[717,764,1984,2007]" italics="true" pageId="13" pageNumber="164">acm</emphasis>
running parallel to solenidion (Acm S of
<bibRefCitation id="713A099FFF93FFAEFB52CBC3FA0CF808" author="Grandjean, F." box="[1239,1424,1983,2008]" journalOrPublisher="Bulletin de la Societe Zoologique de France" pageId="13" pageNumber="164" pagination="421 - 446" part="78" refString="Grandjean, F. (1954) Essai de classification des Oribates (Acariens). Bulletin de la Societe Zoologique de France, 78, 421 - 446." title="Essai de classification des Oribates (Acariens)" type="journal article" year="1954">Grandjean 1954</bibRefCitation>
), but bases slightly separated. Femur I long, with medial, distinct bend. Tibia IV with solenidion. Tibia II and tarsus II without anterodorsal carina. Monodactylous or heterotridactylous.
</paragraph>
<paragraph id="1514746EFF90FFADFF42CC9DFA90FE5A" blockId="14.[151,1437,151,2018]" pageId="14" pageNumber="165">
<emphasis id="27DFA87CFF90FFADFF42CC9DFEA2FF28" box="[199,318,225,248]" italics="true" pageId="14" pageNumber="165">Immatures</emphasis>
: Plicate, eupheredermous, line of dehiscense complete, without hysterosomal macrosclerites or excentrosclerites, at least partially tuberculate; cerotegument as tightly packed platelets,
<quantity id="D253D98BFF90FFADFBF9CD78FB5FFECD" box="[1148,1219,260,285]" metricMagnitude="-2" metricUnit="m" metricValue="3.81" metricValueMax="5.08" metricValueMin="2.54" pageId="14" pageNumber="165" unit="in" value="1.5" valueMax="2.0" valueMin="1.0">1–2 in</quantity>
diameter, covering all of body and leg segments. Prodorsal porose regions present. Gastronotal setation unideficient; larva with 12 pairs, nymphs with 15 pairs of setae (
<emphasis id="27DFA87CFF90FFADFDBECD30FDD7FEB9" box="[571,587,332,361]" italics="true" pageId="14" pageNumber="165">f1</emphasis>
absent). Gastronotal setae dimorphic with those of
<emphasis id="27DFA87CFF90FFADFB08CD31FB2AFEB5" box="[1165,1206,332,357]" italics="true" pageId="14" pageNumber="165">c, d</emphasis>
and
<emphasis id="27DFA87CFF90FFADFB69CD31FB66FEB4" box="[1260,1274,333,356]" italics="true" pageId="14" pageNumber="165">p</emphasis>
series similar, setose; some or all setae of
<emphasis id="27DFA87CFF90FFADFE4ECD0EFE4FFE5B" box="[459,467,370,395]" italics="true" pageId="14" pageNumber="165">l</emphasis>
and
<emphasis id="27DFA87CFF90FFADFD8CCD0EFD8BFE5B" box="[521,535,370,395]" italics="true" pageId="14" pageNumber="165">h</emphasis>
series brush-like. Seta
<emphasis id="27DFA87CFF90FFADFC9ECD0EFCB5FE5B" box="[795,809,370,395]" italics="true" pageId="14" pageNumber="165">d</emphasis>
present on tibiae I to IV and genua I to III.
</paragraph>
</subSubSection>
<subSubSection id="5DB127E5FF90FFADFF42CDEAFE4BF8D9" pageId="14" pageNumber="165" type="description">
<paragraph id="1514746EFF90FFADFF42CDEAFE78FE27" blockId="14.[151,1437,151,2018]" pageId="14" pageNumber="165">
<emphasis id="27DFA87CFF90FFADFF42CDEAFECDFE7F" bold="true" box="[199,337,406,431]" pageId="14" pageNumber="165">Description</emphasis>
.
<emphasis id="27DFA87CFF90FFADFEE7CDEAFDB1FE7F" box="[354,557,406,431]" italics="true" pageId="14" pageNumber="165">Adult. Integument</emphasis>
. Cerotegument present laterally on podosoma, finely and densely granulate. Integument microtuberculate throughout; U-shaped shallow groove extending from coxisternal region to posterior of anal plates (
<figureCitation id="8D9068EBFF90FFADFEB9CDA2FE05FE27" box="[316,409,478,503]" captionStart="FIGURES 10 – 11" captionStartId="21.[151,264,1974,1996]" captionTargetBox="[293,1284,213,1897]" captionTargetId="figure@21.[234,1338,193,1940]" captionTargetPageId="21" captionText="FIGURES 10 – 11. Tegoribates americanus Hammer, adult female, legs removed; 10, ventral aspect; 11, lateral aspect of prodorsum and podosoma, gnathosoma removed, with arrow to taenidium. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018653/files/figure.png" pageId="14" pageNumber="165">Figs. 10</figureCitation>
,
<figureCitation id="8D9068EBFF90FFADFE20CDA3FE4BFE27" box="[421,471,479,503]" captionStart="FIGURE 21" captionStartId="33.[151,250,1886,1908]" captionTargetBox="[225,1362,193,1858]" captionTargetId="figure@33.[225,1362,193,1865]" captionTargetPageId="33" captionText="FIGURE 21. Tegoribates walteri sp. nov., scanning electron microscope images of adult female from California, A. habitus dorsal, with arrow to spine-like seta l ” of genu II; B, habitus ventral, with arrow to ventral depression; C, frontal aspect, with arrows to spine-like setae l ” of genua I and II; D, anterolateral of prodorsum and part of notogaster, with arrow to spine-like seta l ” of genu II; E, ventrofrontal aspect, with arrow to taenidium. Scale bars: A, B = 50 µm; C-E = 25 µm." httpUri="https://zenodo.org/record/1018675/files/figure.png" pageId="14" pageNumber="165">21B</figureCitation>
).
</paragraph>
<paragraph id="1514746EFF90FFADFF42CE7EFDCAFCEB" blockId="14.[151,1437,151,2018]" pageId="14" pageNumber="165">
<emphasis id="27DFA87CFF90FFADFF42CE7EFED8FDCB" box="[199,324,514,539]" italics="true" pageId="14" pageNumber="165">Prodorsum</emphasis>
. Rostrum with smooth margin, without medial incision; weak medial keel extending posteriorly to underneath lamella (
<figureCitation id="8D9068EBFF90FFADFEF9CE5AFD98FDEF" box="[380,516,550,575]" captionStart="FIGURE 21" captionStartId="33.[151,250,1886,1908]" captionTargetBox="[225,1362,193,1858]" captionTargetId="figure@33.[225,1362,193,1865]" captionTargetPageId="33" captionText="FIGURE 21. Tegoribates walteri sp. nov., scanning electron microscope images of adult female from California, A. habitus dorsal, with arrow to spine-like seta l ” of genu II; B, habitus ventral, with arrow to ventral depression; C, frontal aspect, with arrows to spine-like setae l ” of genua I and II; D, anterolateral of prodorsum and part of notogaster, with arrow to spine-like seta l ” of genu II; E, ventrofrontal aspect, with arrow to taenidium. Scale bars: A, B = 50 µm; C-E = 25 µm." httpUri="https://zenodo.org/record/1018675/files/figure.png" pageId="14" pageNumber="165">Fig. 21C, D</figureCitation>
). Rostral setae heavily barbed laterally, directed anteromedially (
<figureCitation id="8D9068EBFF90FFADFB5FCE5AFAB7FDEF" box="[1242,1323,550,575]" captionStart="FIGURES 10 – 11" captionStartId="21.[151,264,1974,1996]" captionTargetBox="[293,1284,213,1897]" captionTargetId="figure@21.[234,1338,193,1940]" captionTargetPageId="21" captionText="FIGURES 10 – 11. Tegoribates americanus Hammer, adult female, legs removed; 10, ventral aspect; 11, lateral aspect of prodorsum and podosoma, gnathosoma removed, with arrow to taenidium. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018653/files/figure.png" pageId="14" pageNumber="165">Fig. 11</figureCitation>
). Lamella long, pair fused, covering all of prodorsum in dorsal aspect (
<figureCitation id="8D9068EBFF90FFADFCCCCE36FC13FDB3" box="[841,911,586,611]" captionStart="FIGURE 9" captionStartId="20.[151,250,1982,2004]" captionTargetBox="[250,1356,207,1944]" captionTargetId="figure@20.[210,1367,193,1949]" captionTargetPageId="20" captionText="FIGURE 9. Tegoribates americanus Hammer, adult female, dorsal aspect, legs not illustrated, with arrow to fused dorsophragmata. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018649/files/figure.png" pageId="14" pageNumber="165">Fig. 9</figureCitation>
), with faint longitudinal line medially (visible with light microscopy), with almost parallel areas accommodating nerve canal for seta
<emphasis id="27DFA87CFF90FFADFBEECE12FB1CFD57" box="[1131,1152,622,647]" italics="true" pageId="14" pageNumber="165">le</emphasis>
(
<figureCitation id="8D9068EBFF90FFADFB0ACE12FB70FD57" box="[1167,1260,622,647]" captionStart="FIGURE 8" captionStartId="17.[151,250,1857,1879]" captionTargetBox="[219,1314,198,1804]" captionTargetId="figure@17.[219,1314,193,1804]" captionTargetPageId="17" captionText="FIGURE 8 A – H. Tegoribates subniger Ewing, differential interference contrast microscope images of adult. A, holotype specimen, Type No. 2009427, housed in USNM; B, lateral of prodorsum with arrow to taenidium; C, preanal organ indicated by arrow; D, region of tubule T 1, h 3 and gla, with small arrows to tubule and large arrow to its opening (3 layers); E, lamella (5 layers), with arrow to nerve canal for seta le, lateral of podosoma displaced so Ah visible, black arrow to fused dorsophragmata; F, ventral region showing discidium (dis) and circumpedal carina (cc), with arrow to minute custodium (3 layers); G, pedipalp (5 layers), with arrow to solenidion ω. Scale bars: A = 100 µm; C, D, F = 25 µm; E = 40 µm; G = 30 µm." httpUri="https://zenodo.org/record/1018647/files/figure.png" pageId="14" pageNumber="165">Figs. 8F</figureCitation>
,
<figureCitation id="8D9068EBFF90FFADFB7DCE13FAB1FD57" box="[1272,1325,623,647]" captionStart="FIGURE 22" captionStartId="34.[151,250,1827,1849]" captionTargetBox="[222,1338,193,1774]" captionTargetId="figure@34.[222,1338,193,1794]" captionTargetPageId="34" captionText="FIGURE 22. Differential interference contrast microscope images of adults; A – D, Tegoribates walteri sp. nov., A, part of prodorsum (4 layers) showing lamella, bothridium, bothridial seta, interlamellar seta, with arrow to fused dorsophragmata; B, hinged pteromorph (3 layers); C, coxisternal region showing custodium (arrow) and pedotectum II (3 layers); D, porose area Aa and surrounding muscle sigillae (5 layers). E, Tegoribates subniger Ewing, lateral of podosoma, showing elongated, bent femur I (indicated by arrow); F, G, Protectoribates occidentalis sp. nov., detail of preanal organ; H, Hypozetes laysanensis Aoki, detail of preanal organ; I, Tectoribates borealis Behan-Pelletier &amp; Walter, detail of preanal organ. Scale bars: A, B, E = 25 µm; C, D = 20 µm." httpUri="https://zenodo.org/record/1018677/files/figure.png" pageId="14" pageNumber="165">22A</figureCitation>
). Lamella with short, V or U-shaped indentation anteriorly; concave posterolaterally, accommodating bothridium (
<figureCitation id="8D9068EBFF90FFADFAB0CEEEFA0BFD7B" box="[1333,1431,658,683]" captionStart="FIGURE 8" captionStartId="17.[151,250,1857,1879]" captionTargetBox="[219,1314,198,1804]" captionTargetId="figure@17.[219,1314,193,1804]" captionTargetPageId="17" captionText="FIGURE 8 A – H. Tegoribates subniger Ewing, differential interference contrast microscope images of adult. A, holotype specimen, Type No. 2009427, housed in USNM; B, lateral of prodorsum with arrow to taenidium; C, preanal organ indicated by arrow; D, region of tubule T 1, h 3 and gla, with small arrows to tubule and large arrow to its opening (3 layers); E, lamella (5 layers), with arrow to nerve canal for seta le, lateral of podosoma displaced so Ah visible, black arrow to fused dorsophragmata; F, ventral region showing discidium (dis) and circumpedal carina (cc), with arrow to minute custodium (3 layers); G, pedipalp (5 layers), with arrow to solenidion ω. Scale bars: A = 100 µm; C, D, F = 25 µm; E = 40 µm; G = 30 µm." httpUri="https://zenodo.org/record/1018647/files/figure.png" pageId="14" pageNumber="165">Figs. 8E</figureCitation>
,
<figureCitation id="8D9068EBFF90FFADFF12CECBFF5BFD1F" box="[151,199,695,719]" captionStart="FIGURE 13" captionStartId="23.[151,250,1906,1928]" captionTargetBox="[213,1359,193,1878]" captionTargetId="figure@23.[213,1359,193,1878]" captionTargetPageId="23" captionText="FIGURE 13. Tegoribates americanus Hammer, differential interference contrast microscope images of adult. A, lateral of prodorsum (6 layers) with arrows to taenidium; B, bothridial seta and bothridium, with arrow to anterior scale; C, chelicera, pedipalp and rutellum showing size relationship, supracoxal seta e, and with arrow to axillary saccule (5 layers); D, tarsus II, showing shape of seta pv ”; E, postanal porose area. Scale bars: A, B = 50 µm; C = 25 µm." httpUri="https://zenodo.org/record/1018657/files/figure.png" pageId="14" pageNumber="165">13B</figureCitation>
,
<figureCitation id="8D9068EBFF90FFADFF56CECBFE9BFD1F" box="[211,263,695,719]" captionStart="FIGURE 21" captionStartId="33.[151,250,1886,1908]" captionTargetBox="[225,1362,193,1858]" captionTargetId="figure@33.[225,1362,193,1865]" captionTargetPageId="33" captionText="FIGURE 21. Tegoribates walteri sp. nov., scanning electron microscope images of adult female from California, A. habitus dorsal, with arrow to spine-like seta l ” of genu II; B, habitus ventral, with arrow to ventral depression; C, frontal aspect, with arrows to spine-like setae l ” of genua I and II; D, anterolateral of prodorsum and part of notogaster, with arrow to spine-like seta l ” of genu II; E, ventrofrontal aspect, with arrow to taenidium. Scale bars: A, B = 50 µm; C-E = 25 µm." httpUri="https://zenodo.org/record/1018675/files/figure.png" pageId="14" pageNumber="165">21D</figureCitation>
). Seta
<emphasis id="27DFA87CFF90FFADFED4CECAFEFAFD1F" box="[337,358,694,719]" italics="true" pageId="14" pageNumber="165">le</emphasis>
arising dorsally on lamella (
<figureCitation id="8D9068EBFF90FFADFD26CECAFD6DFD1F" box="[675,753,694,719]" captionStart="FIGURE 9" captionStartId="20.[151,250,1982,2004]" captionTargetBox="[250,1356,207,1944]" captionTargetId="figure@20.[210,1367,193,1949]" captionTargetPageId="20" captionText="FIGURE 9. Tegoribates americanus Hammer, adult female, dorsal aspect, legs not illustrated, with arrow to fused dorsophragmata. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018649/files/figure.png" pageId="14" pageNumber="165">Figs. 9</figureCitation>
,
<figureCitation id="8D9068EBFF90FFADFD78CECBFC87FD1F" box="[765,795,695,719]" captionStart="FIGURES 17 – 18" captionStartId="29.[151,264,1910,1932]" captionTargetBox="[173,1413,224,1840]" captionTargetId="figure@29.[161,1426,193,1888]" captionTargetPageId="29" captionText="FIGURES 17 – 18. Tegoribates bryophilus Woolley, adult female (non-type specimen), 17, dorsal aspect, 18, ventral aspect; legs not illustrated. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018665/files/figure.png" pageId="14" pageNumber="165">17</figureCitation>
,
<figureCitation id="8D9068EBFF90FFADFCA3CECBFCDAFD1F" box="[806,838,695,719]" captionStart="FIGURE 20" captionStartId="32.[151,250,1976,1998]" captionTargetBox="[244,1353,224,1923]" captionTargetId="figure@32.[202,1385,193,1955]" captionTargetPageId="32" captionText="FIGURE 20. Tegoribates walteri sp. nov., adult male, dorsal aspect, legs not illustrated. Scale bar = 100 µm." httpUri="https://zenodo.org/record/1018673/files/figure.png" pageId="14" pageNumber="165">20</figureCitation>
). Interlamellar setae subequal in thickness to lamellar setae. Exobothridial seta present, or absent; alveolus usually present. Bothridial wall expanded anterodorsally with pointed scale, otherwise cup-shaped, without lateral indentation (
<figureCitation id="8D9068EBFF90FFADFC20CE82FB80FCC7" box="[933,1052,766,791]" captionStart="FIGURE 13" captionStartId="23.[151,250,1906,1928]" captionTargetBox="[213,1359,193,1878]" captionTargetId="figure@23.[213,1359,193,1878]" captionTargetPageId="23" captionText="FIGURE 13. Tegoribates americanus Hammer, differential interference contrast microscope images of adult. A, lateral of prodorsum (6 layers) with arrows to taenidium; B, bothridial seta and bothridium, with arrow to anterior scale; C, chelicera, pedipalp and rutellum showing size relationship, supracoxal seta e, and with arrow to axillary saccule (5 layers); D, tarsus II, showing shape of seta pv ”; E, postanal porose area. Scale bars: A, B = 50 µm; C = 25 µm." httpUri="https://zenodo.org/record/1018657/files/figure.png" pageId="14" pageNumber="165">Figs. 13B</figureCitation>
,
<figureCitation id="8D9068EBFF90FFADFBB5CE83FBF8FCC7" box="[1072,1124,767,791]" captionStart="FIGURE 22" captionStartId="34.[151,250,1827,1849]" captionTargetBox="[222,1338,193,1774]" captionTargetId="figure@34.[222,1338,193,1794]" captionTargetPageId="34" captionText="FIGURE 22. Differential interference contrast microscope images of adults; A – D, Tegoribates walteri sp. nov., A, part of prodorsum (4 layers) showing lamella, bothridium, bothridial seta, interlamellar seta, with arrow to fused dorsophragmata; B, hinged pteromorph (3 layers); C, coxisternal region showing custodium (arrow) and pedotectum II (3 layers); D, porose area Aa and surrounding muscle sigillae (5 layers). E, Tegoribates subniger Ewing, lateral of podosoma, showing elongated, bent femur I (indicated by arrow); F, G, Protectoribates occidentalis sp. nov., detail of preanal organ; H, Hypozetes laysanensis Aoki, detail of preanal organ; I, Tectoribates borealis Behan-Pelletier &amp; Walter, detail of preanal organ. Scale bars: A, B, E = 25 µm; C, D = 20 µm." httpUri="https://zenodo.org/record/1018677/files/figure.png" pageId="14" pageNumber="165">22A</figureCitation>
). Porose area
<emphasis id="27DFA87CFF90FFADFA93CE82FAA9FCC7" box="[1302,1333,766,791]" italics="true" pageId="14" pageNumber="165">Ad</emphasis>
present. Dorsophragmata fused (
<figureCitation id="8D9068EBFF90FFADFE21CF5EFE68FCEB" box="[420,500,802,827]" captionStart="FIGURE 9" captionStartId="20.[151,250,1982,2004]" captionTargetBox="[250,1356,207,1944]" captionTargetId="figure@20.[210,1367,193,1949]" captionTargetPageId="20" captionText="FIGURE 9. Tegoribates americanus Hammer, adult female, dorsal aspect, legs not illustrated, with arrow to fused dorsophragmata. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018649/files/figure.png" pageId="14" pageNumber="165">Figs. 9</figureCitation>
,
<figureCitation id="8D9068EBFF90FFADFD85CF5FFD82FCEB" box="[512,542,803,827]" captionStart="FIGURES 17 – 18" captionStartId="29.[151,264,1910,1932]" captionTargetBox="[173,1413,224,1840]" captionTargetId="figure@29.[161,1426,193,1888]" captionTargetPageId="29" captionText="FIGURES 17 – 18. Tegoribates bryophilus Woolley, adult female (non-type specimen), 17, dorsal aspect, 18, ventral aspect; legs not illustrated. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018665/files/figure.png" pageId="14" pageNumber="165">17</figureCitation>
,
<figureCitation id="8D9068EBFF90FFADFDAFCF5FFDD6FCEB" box="[554,586,803,827]" captionStart="FIGURE 20" captionStartId="32.[151,250,1976,1998]" captionTargetBox="[244,1353,224,1923]" captionTargetId="figure@32.[202,1385,193,1955]" captionTargetPageId="32" captionText="FIGURE 20. Tegoribates walteri sp. nov., adult male, dorsal aspect, legs not illustrated. Scale bar = 100 µm." httpUri="https://zenodo.org/record/1018673/files/figure.png" pageId="14" pageNumber="165">20</figureCitation>
).
</paragraph>
<paragraph id="1514746EFF90FFADFF42CF34FE4EFC3F" blockId="14.[151,1437,151,2018]" pageId="14" pageNumber="165">
<emphasis id="27DFA87CFF90FFADFF42CF34FEDEFC8F" box="[199,322,840,863]" italics="true" pageId="14" pageNumber="165">Notogaster</emphasis>
. Longer than wide; with hinged pteromorphs having smooth margins without dens anteroventrally (
<figureCitation id="8D9068EBFF90FFADFF1ACF16FE89FC53" box="[159,277,874,899]" captionStart="FIGURE 19" captionStartId="30.[151,250,1905,1927]" captionTargetBox="[213,1359,193,1849]" captionTargetId="figure@30.[213,1359,193,1878]" captionTargetPageId="30" captionText="FIGURE 19. Tegoribates bryophilus Woolley, differential interference contrast microscope images of adult type material. A. anterior of prodorsum (note, specimen broken), with arrow to taenidium; B, pteromorph; C, saccule Sa and seta lm; D, posterior of prodorsum with arrow to fused dorsophragmata (3 layers); E, lateral of coxisternal region showing circumpedal carina and discidium; F, lenticulus; G, seta l ” of genu II. Scale bars: A, B, E = 50 µm; C, = 10 µm; D = 35 µm." httpUri="https://zenodo.org/record/1018669/files/figure.png" pageId="14" pageNumber="165">Figs. 19B</figureCitation>
,
<figureCitation id="8D9068EBFF90FFADFEA3CF17FEC4FC53" box="[294,344,875,899]" captionStart="FIGURE 21" captionStartId="33.[151,250,1886,1908]" captionTargetBox="[225,1362,193,1858]" captionTargetId="figure@33.[225,1362,193,1865]" captionTargetPageId="33" captionText="FIGURE 21. Tegoribates walteri sp. nov., scanning electron microscope images of adult female from California, A. habitus dorsal, with arrow to spine-like seta l ” of genu II; B, habitus ventral, with arrow to ventral depression; C, frontal aspect, with arrows to spine-like setae l ” of genua I and II; D, anterolateral of prodorsum and part of notogaster, with arrow to spine-like seta l ” of genu II; E, ventrofrontal aspect, with arrow to taenidium. Scale bars: A, B = 50 µm; C-E = 25 µm." httpUri="https://zenodo.org/record/1018675/files/figure.png" pageId="14" pageNumber="165">21D</figureCitation>
,
<figureCitation id="8D9068EBFF90FFADFEECCF17FE00FC53" box="[361,412,875,899]" captionStart="FIGURE 22" captionStartId="34.[151,250,1827,1849]" captionTargetBox="[222,1338,193,1774]" captionTargetId="figure@34.[222,1338,193,1794]" captionTargetPageId="34" captionText="FIGURE 22. Differential interference contrast microscope images of adults; A – D, Tegoribates walteri sp. nov., A, part of prodorsum (4 layers) showing lamella, bothridium, bothridial seta, interlamellar seta, with arrow to fused dorsophragmata; B, hinged pteromorph (3 layers); C, coxisternal region showing custodium (arrow) and pedotectum II (3 layers); D, porose area Aa and surrounding muscle sigillae (5 layers). E, Tegoribates subniger Ewing, lateral of podosoma, showing elongated, bent femur I (indicated by arrow); F, G, Protectoribates occidentalis sp. nov., detail of preanal organ; H, Hypozetes laysanensis Aoki, detail of preanal organ; I, Tectoribates borealis Behan-Pelletier &amp; Walter, detail of preanal organ. Scale bars: A, B, E = 25 µm; C, D = 20 µm." httpUri="https://zenodo.org/record/1018677/files/figure.png" pageId="14" pageNumber="165">22B</figureCitation>
). Anterior margin of notogaster straight, transverse, without ridges in region lateral to bothridium (
<figureCitation id="8D9068EBFF90FFADFEACCFF2FEEFFC77" box="[297,371,910,935]" captionStart="FIGURE 9" captionStartId="20.[151,250,1982,2004]" captionTargetBox="[250,1356,207,1944]" captionTargetId="figure@20.[210,1367,193,1949]" captionTargetPageId="20" captionText="FIGURE 9. Tegoribates americanus Hammer, adult female, dorsal aspect, legs not illustrated, with arrow to fused dorsophragmata. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018649/files/figure.png" pageId="14" pageNumber="165">Fig. 9</figureCitation>
). Anterior of notogaster without hexagonal pattern in transmitted light. Octotaxic system expressed as 4 pairs of either porose areas, saccules or tubules (
<figureCitation id="8D9068EBFF90FFADFCEBCFCEFC48FC1B" box="[878,980,946,971]" captionStart="FIGURE 8" captionStartId="17.[151,250,1857,1879]" captionTargetBox="[219,1314,198,1804]" captionTargetId="figure@17.[219,1314,193,1804]" captionTargetPageId="17" captionText="FIGURE 8 A – H. Tegoribates subniger Ewing, differential interference contrast microscope images of adult. A, holotype specimen, Type No. 2009427, housed in USNM; B, lateral of prodorsum with arrow to taenidium; C, preanal organ indicated by arrow; D, region of tubule T 1, h 3 and gla, with small arrows to tubule and large arrow to its opening (3 layers); E, lamella (5 layers), with arrow to nerve canal for seta le, lateral of podosoma displaced so Ah visible, black arrow to fused dorsophragmata; F, ventral region showing discidium (dis) and circumpedal carina (cc), with arrow to minute custodium (3 layers); G, pedipalp (5 layers), with arrow to solenidion ω. Scale bars: A = 100 µm; C, D, F = 25 µm; E = 40 µm; G = 30 µm." httpUri="https://zenodo.org/record/1018647/files/figure.png" pageId="14" pageNumber="165">Figs. 8D</figureCitation>
,
<figureCitation id="8D9068EBFF90FFADFC64CFCFFC6DFC1B" box="[993,1009,947,971]" captionStart="FIGURE 9" captionStartId="20.[151,250,1982,2004]" captionTargetBox="[250,1356,207,1944]" captionTargetId="figure@20.[210,1367,193,1949]" captionTargetPageId="20" captionText="FIGURE 9. Tegoribates americanus Hammer, adult female, dorsal aspect, legs not illustrated, with arrow to fused dorsophragmata. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018649/files/figure.png" pageId="14" pageNumber="165">9</figureCitation>
,
<figureCitation id="8D9068EBFF90FFADFC7ACFCFFB81FC1B" box="[1023,1053,947,971]" captionStart="FIGURES 17 – 18" captionStartId="29.[151,264,1910,1932]" captionTargetBox="[173,1413,224,1840]" captionTargetId="figure@29.[161,1426,193,1888]" captionTargetPageId="29" captionText="FIGURES 17 – 18. Tegoribates bryophilus Woolley, adult female (non-type specimen), 17, dorsal aspect, 18, ventral aspect; legs not illustrated. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018665/files/figure.png" pageId="14" pageNumber="165">17</figureCitation>
,
<figureCitation id="8D9068EBFF90FFADFBAFCFCFFBD6FC1B" box="[1066,1098,947,971]" captionStart="FIGURE 20" captionStartId="32.[151,250,1976,1998]" captionTargetBox="[244,1353,224,1923]" captionTargetId="figure@32.[202,1385,193,1955]" captionTargetPageId="32" captionText="FIGURE 20. Tegoribates walteri sp. nov., adult male, dorsal aspect, legs not illustrated. Scale bar = 100 µm." httpUri="https://zenodo.org/record/1018673/files/figure.png" pageId="14" pageNumber="165">20</figureCitation>
); without sexual dimorphism. With 10 pairs of short setae.
</paragraph>
<paragraph id="1514746EFF90FFADFF42CF86FB0EFB3B" blockId="14.[151,1437,151,2018]" pageId="14" pageNumber="165">
<emphasis id="27DFA87CFF90FFADFF42CF86FD90FBC3" box="[199,524,1018,1043]" italics="true" pageId="14" pageNumber="165">Lateral Region of Podosoma</emphasis>
. Genal incision and genal tooth absent; open taenidium present in usual position of genal incision, extending to acetabulum I (
<figureCitation id="8D9068EBFF90FFADFD1CC862FD60FBE7" box="[665,764,1054,1079]" captionStart="FIGURE 8" captionStartId="17.[151,250,1857,1879]" captionTargetBox="[219,1314,198,1804]" captionTargetId="figure@17.[219,1314,193,1804]" captionTargetPageId="17" captionText="FIGURE 8 A – H. Tegoribates subniger Ewing, differential interference contrast microscope images of adult. A, holotype specimen, Type No. 2009427, housed in USNM; B, lateral of prodorsum with arrow to taenidium; C, preanal organ indicated by arrow; D, region of tubule T 1, h 3 and gla, with small arrows to tubule and large arrow to its opening (3 layers); E, lamella (5 layers), with arrow to nerve canal for seta le, lateral of podosoma displaced so Ah visible, black arrow to fused dorsophragmata; F, ventral region showing discidium (dis) and circumpedal carina (cc), with arrow to minute custodium (3 layers); G, pedipalp (5 layers), with arrow to solenidion ω. Scale bars: A = 100 µm; C, D, F = 25 µm; E = 40 µm; G = 30 µm." httpUri="https://zenodo.org/record/1018647/files/figure.png" pageId="14" pageNumber="165">Figs. 8B</figureCitation>
,
<figureCitation id="8D9068EBFF90FFADFC8CC863FCA0FBE7" box="[777,828,1055,1079]" captionStart="FIGURE 13" captionStartId="23.[151,250,1906,1928]" captionTargetBox="[213,1359,193,1878]" captionTargetId="figure@23.[213,1359,193,1878]" captionTargetPageId="23" captionText="FIGURE 13. Tegoribates americanus Hammer, differential interference contrast microscope images of adult. A, lateral of prodorsum (6 layers) with arrows to taenidium; B, bothridial seta and bothridium, with arrow to anterior scale; C, chelicera, pedipalp and rutellum showing size relationship, supracoxal seta e, and with arrow to axillary saccule (5 layers); D, tarsus II, showing shape of seta pv ”; E, postanal porose area. Scale bars: A, B = 50 µm; C = 25 µm." httpUri="https://zenodo.org/record/1018657/files/figure.png" pageId="14" pageNumber="165">13A</figureCitation>
,
<figureCitation id="8D9068EBFF90FFADFCCDC863FCE5FBE7" box="[840,889,1055,1079]" captionStart="FIGURE 21" captionStartId="33.[151,250,1886,1908]" captionTargetBox="[225,1362,193,1858]" captionTargetId="figure@33.[225,1362,193,1865]" captionTargetPageId="33" captionText="FIGURE 21. Tegoribates walteri sp. nov., scanning electron microscope images of adult female from California, A. habitus dorsal, with arrow to spine-like seta l ” of genu II; B, habitus ventral, with arrow to ventral depression; C, frontal aspect, with arrows to spine-like setae l ” of genua I and II; D, anterolateral of prodorsum and part of notogaster, with arrow to spine-like seta l ” of genu II; E, ventrofrontal aspect, with arrow to taenidium. Scale bars: A, B = 50 µm; C-E = 25 µm." httpUri="https://zenodo.org/record/1018675/files/figure.png" pageId="14" pageNumber="165">21E</figureCitation>
). Tutorium narrow, lamelliform, cusp triangular, lying parallel to dorsal contour of prodorsum in lateral aspect, extending anterior of insertion of rostral seta (
<figureCitation id="8D9068EBFF90FFADFAE1C83EFF28FBAF" captionStart="FIGURES 10 – 11" captionStartId="21.[151,264,1974,1996]" captionTargetBox="[293,1284,213,1897]" captionTargetId="figure@21.[234,1338,193,1940]" captionTargetPageId="21" captionText="FIGURES 10 – 11. Tegoribates americanus Hammer, adult female, legs removed; 10, ventral aspect; 11, lateral aspect of prodorsum and podosoma, gnathosoma removed, with arrow to taenidium. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018653/files/figure.png" pageId="14" pageNumber="165">Figs. 11</figureCitation>
,
<figureCitation id="8D9068EBFF90FFADFF43C81BFF66FBAF" box="[198,250,1127,1151]" captionStart="FIGURE 13" captionStartId="23.[151,250,1906,1928]" captionTargetBox="[213,1359,193,1878]" captionTargetId="figure@23.[213,1359,193,1878]" captionTargetPageId="23" captionText="FIGURE 13. Tegoribates americanus Hammer, differential interference contrast microscope images of adult. A, lateral of prodorsum (6 layers) with arrows to taenidium; B, bothridial seta and bothridium, with arrow to anterior scale; C, chelicera, pedipalp and rutellum showing size relationship, supracoxal seta e, and with arrow to axillary saccule (5 layers); D, tarsus II, showing shape of seta pv ”; E, postanal porose area. Scale bars: A, B = 50 µm; C = 25 µm." httpUri="https://zenodo.org/record/1018657/files/figure.png" pageId="14" pageNumber="165">13A</figureCitation>
). Pedotectum I flattened or rounded dorsally, pointed distally, without ventral depression (
<figureCitation id="8D9068EBFF90FFADFAB3C81AFA12FBAF" box="[1334,1422,1126,1151]" captionStart="FIGURES 10 – 11" captionStartId="21.[151,264,1974,1996]" captionTargetBox="[293,1284,213,1897]" captionTargetId="figure@21.[234,1338,193,1940]" captionTargetPageId="21" captionText="FIGURES 10 – 11. Tegoribates americanus Hammer, adult female, legs removed; 10, ventral aspect; 11, lateral aspect of prodorsum and podosoma, gnathosoma removed, with arrow to taenidium. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018653/files/figure.png" pageId="14" pageNumber="165">Fig. 11</figureCitation>
). Pedotectum II present, without tubercle close to body wall. Custodium present as very short extension to circumpedal carina (
<figureCitation id="8D9068EBFF90FFADFE05C8D2FE43FB17" box="[384,479,1198,1223]" captionStart="FIGURE 8" captionStartId="17.[151,250,1857,1879]" captionTargetBox="[219,1314,198,1804]" captionTargetId="figure@17.[219,1314,193,1804]" captionTargetPageId="17" captionText="FIGURE 8 A – H. Tegoribates subniger Ewing, differential interference contrast microscope images of adult. A, holotype specimen, Type No. 2009427, housed in USNM; B, lateral of prodorsum with arrow to taenidium; C, preanal organ indicated by arrow; D, region of tubule T 1, h 3 and gla, with small arrows to tubule and large arrow to its opening (3 layers); E, lamella (5 layers), with arrow to nerve canal for seta le, lateral of podosoma displaced so Ah visible, black arrow to fused dorsophragmata; F, ventral region showing discidium (dis) and circumpedal carina (cc), with arrow to minute custodium (3 layers); G, pedipalp (5 layers), with arrow to solenidion ω. Scale bars: A = 100 µm; C, D, F = 25 µm; E = 40 µm; G = 30 µm." httpUri="https://zenodo.org/record/1018647/files/figure.png" pageId="14" pageNumber="165">Figs. 8F</figureCitation>
,
<figureCitation id="8D9068EBFF90FFADFE68C8D3FD96FB17" box="[493,522,1199,1223]" captionStart="FIGURES 10 – 11" captionStartId="21.[151,264,1974,1996]" captionTargetBox="[293,1284,213,1897]" captionTargetId="figure@21.[234,1338,193,1940]" captionTargetPageId="21" captionText="FIGURES 10 – 11. Tegoribates americanus Hammer, adult female, legs removed; 10, ventral aspect; 11, lateral aspect of prodorsum and podosoma, gnathosoma removed, with arrow to taenidium. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018653/files/figure.png" pageId="14" pageNumber="165">11</figureCitation>
,
<figureCitation id="8D9068EBFF90FFADFD92C8D3FDDBFB17" box="[535,583,1199,1223]" captionStart="FIGURE 19" captionStartId="30.[151,250,1905,1927]" captionTargetBox="[213,1359,193,1849]" captionTargetId="figure@30.[213,1359,193,1878]" captionTargetPageId="30" captionText="FIGURE 19. Tegoribates bryophilus Woolley, differential interference contrast microscope images of adult type material. A. anterior of prodorsum (note, specimen broken), with arrow to taenidium; B, pteromorph; C, saccule Sa and seta lm; D, posterior of prodorsum with arrow to fused dorsophragmata (3 layers); E, lateral of coxisternal region showing circumpedal carina and discidium; F, lenticulus; G, seta l ” of genu II. Scale bars: A, B, E = 50 µm; C, = 10 µm; D = 35 µm." httpUri="https://zenodo.org/record/1018669/files/figure.png" pageId="14" pageNumber="165">19E</figureCitation>
,
<figureCitation id="8D9068EBFF90FFADFDD0C8D3FD14FB17" box="[597,648,1199,1223]" captionStart="FIGURE 22" captionStartId="34.[151,250,1827,1849]" captionTargetBox="[222,1338,193,1774]" captionTargetId="figure@34.[222,1338,193,1794]" captionTargetPageId="34" captionText="FIGURE 22. Differential interference contrast microscope images of adults; A – D, Tegoribates walteri sp. nov., A, part of prodorsum (4 layers) showing lamella, bothridium, bothridial seta, interlamellar seta, with arrow to fused dorsophragmata; B, hinged pteromorph (3 layers); C, coxisternal region showing custodium (arrow) and pedotectum II (3 layers); D, porose area Aa and surrounding muscle sigillae (5 layers). E, Tegoribates subniger Ewing, lateral of podosoma, showing elongated, bent femur I (indicated by arrow); F, G, Protectoribates occidentalis sp. nov., detail of preanal organ; H, Hypozetes laysanensis Aoki, detail of preanal organ; I, Tectoribates borealis Behan-Pelletier &amp; Walter, detail of preanal organ. Scale bars: A, B, E = 25 µm; C, D = 20 µm." httpUri="https://zenodo.org/record/1018677/files/figure.png" pageId="14" pageNumber="165">22C</figureCitation>
). Discidium strongly curved lamina (
<figureCitation id="8D9068EBFF90FFADFBB7C8D2FB15FB17" box="[1074,1161,1198,1223]" captionStart="FIGURE 8" captionStartId="17.[151,250,1857,1879]" captionTargetBox="[219,1314,198,1804]" captionTargetId="figure@17.[219,1314,193,1804]" captionTargetPageId="17" captionText="FIGURE 8 A – H. Tegoribates subniger Ewing, differential interference contrast microscope images of adult. A, holotype specimen, Type No. 2009427, housed in USNM; B, lateral of prodorsum with arrow to taenidium; C, preanal organ indicated by arrow; D, region of tubule T 1, h 3 and gla, with small arrows to tubule and large arrow to its opening (3 layers); E, lamella (5 layers), with arrow to nerve canal for seta le, lateral of podosoma displaced so Ah visible, black arrow to fused dorsophragmata; F, ventral region showing discidium (dis) and circumpedal carina (cc), with arrow to minute custodium (3 layers); G, pedipalp (5 layers), with arrow to solenidion ω. Scale bars: A = 100 µm; C, D, F = 25 µm; E = 40 µm; G = 30 µm." httpUri="https://zenodo.org/record/1018647/files/figure.png" pageId="14" pageNumber="165">Fig. 8F</figureCitation>
). Porose area
<emphasis id="27DFA87CFF90FFADFAA9C8D2FAD9FB17" box="[1324,1349,1198,1223]" italics="true" pageId="14" pageNumber="165">Al</emphasis>
absent. Humerosejugal porose organs
<emphasis id="27DFA87CFF90FFADFE69C8AFFD8DFB3A" box="[492,529,1235,1258]" italics="true" pageId="14" pageNumber="165">Am</emphasis>
and
<emphasis id="27DFA87CFF90FFADFDC2C8AEFDFAFB3B" box="[583,614,1234,1259]" italics="true" pageId="14" pageNumber="165">Ah</emphasis>
present, expressed as porose areas (
<figureCitation id="8D9068EBFF90FFADFC7EC8AEFBC7FB3B" box="[1019,1115,1234,1259]" captionStart="FIGURE 8" captionStartId="17.[151,250,1857,1879]" captionTargetBox="[219,1314,198,1804]" captionTargetId="figure@17.[219,1314,193,1804]" captionTargetPageId="17" captionText="FIGURE 8 A – H. Tegoribates subniger Ewing, differential interference contrast microscope images of adult. A, holotype specimen, Type No. 2009427, housed in USNM; B, lateral of prodorsum with arrow to taenidium; C, preanal organ indicated by arrow; D, region of tubule T 1, h 3 and gla, with small arrows to tubule and large arrow to its opening (3 layers); E, lamella (5 layers), with arrow to nerve canal for seta le, lateral of podosoma displaced so Ah visible, black arrow to fused dorsophragmata; F, ventral region showing discidium (dis) and circumpedal carina (cc), with arrow to minute custodium (3 layers); G, pedipalp (5 layers), with arrow to solenidion ω. Scale bars: A = 100 µm; C, D, F = 25 µm; E = 40 µm; G = 30 µm." httpUri="https://zenodo.org/record/1018647/files/figure.png" pageId="14" pageNumber="165">Figs. 8E</figureCitation>
,
<figureCitation id="8D9068EBFF90FFADFBE2C8AFFB19FB3B" box="[1127,1157,1235,1259]" captionStart="FIGURES 10 – 11" captionStartId="21.[151,264,1974,1996]" captionTargetBox="[293,1284,213,1897]" captionTargetId="figure@21.[234,1338,193,1940]" captionTargetPageId="21" captionText="FIGURES 10 – 11. Tegoribates americanus Hammer, adult female, legs removed; 10, ventral aspect; 11, lateral aspect of prodorsum and podosoma, gnathosoma removed, with arrow to taenidium. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018653/files/figure.png" pageId="14" pageNumber="165">11</figureCitation>
).
</paragraph>
<paragraph id="1514746EFF90FFADFF42C88AFA8BFAAB" blockId="14.[151,1437,151,2018]" pageId="14" pageNumber="165">
<emphasis id="27DFA87CFF90FFADFF42C88AFE89FADF" box="[199,277,1270,1295]" italics="true" pageId="14" pageNumber="165">Ventral</emphasis>
. Epimere I without necklace of small tubercles anteriorly. Epimeral setal formula 3-1-3-3. Genital plates with 6 setae, 3 setae on anterior margin; 1 pair aggenital setae; 3 pairs adanal setae and 2 pairs anal setae (
<figureCitation id="8D9068EBFF90FFADFF1AC942FF6FFA87" box="[159,243,1342,1367]" captionStart="FIGURES 10 – 11" captionStartId="21.[151,264,1974,1996]" captionTargetBox="[293,1284,213,1897]" captionTargetId="figure@21.[234,1338,193,1940]" captionTargetPageId="21" captionText="FIGURES 10 – 11. Tegoribates americanus Hammer, adult female, legs removed; 10, ventral aspect; 11, lateral aspect of prodorsum and podosoma, gnathosoma removed, with arrow to taenidium. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018653/files/figure.png" pageId="14" pageNumber="165">Fig. 10</figureCitation>
). Lyrifissure
<emphasis id="27DFA87CFF90FFADFE02C942FE37FA87" box="[391,427,1342,1367]" italics="true" pageId="14" pageNumber="165">iad</emphasis>
at anterolateral edge of anal plate (
<figureCitation id="8D9068EBFF90FFADFCB0C942FC1AFA87" box="[821,902,1342,1367]" captionStart="FIGURES 10 – 11" captionStartId="21.[151,264,1974,1996]" captionTargetBox="[293,1284,213,1897]" captionTargetId="figure@21.[234,1338,193,1940]" captionTargetPageId="21" captionText="FIGURES 10 – 11. Tegoribates americanus Hammer, adult female, legs removed; 10, ventral aspect; 11, lateral aspect of prodorsum and podosoma, gnathosoma removed, with arrow to taenidium. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018653/files/figure.png" pageId="14" pageNumber="165">Fig. 11</figureCitation>
). Postanal porose area present (
<figureCitation id="8D9068EBFF90FFADFB6DC942FAD1FA87" box="[1256,1357,1342,1367]" captionStart="FIGURE 13" captionStartId="23.[151,250,1906,1928]" captionTargetBox="[213,1359,193,1878]" captionTargetId="figure@23.[213,1359,193,1878]" captionTargetPageId="23" captionText="FIGURE 13. Tegoribates americanus Hammer, differential interference contrast microscope images of adult. A, lateral of prodorsum (6 layers) with arrows to taenidium; B, bothridial seta and bothridium, with arrow to anterior scale; C, chelicera, pedipalp and rutellum showing size relationship, supracoxal seta e, and with arrow to axillary saccule (5 layers); D, tarsus II, showing shape of seta pv ”; E, postanal porose area. Scale bars: A, B = 50 µm; C = 25 µm." httpUri="https://zenodo.org/record/1018657/files/figure.png" pageId="14" pageNumber="165">Fig. 13E</figureCitation>
). Band of darker integument absent between genital and anal plates. Preanal organ large cup-shaped (
<figureCitation id="8D9068EBFF90FFADFB37C91EFA96FAAB" box="[1202,1290,1378,1403]" captionStart="FIGURE 8" captionStartId="17.[151,250,1857,1879]" captionTargetBox="[219,1314,198,1804]" captionTargetId="figure@17.[219,1314,193,1804]" captionTargetPageId="17" captionText="FIGURE 8 A – H. Tegoribates subniger Ewing, differential interference contrast microscope images of adult. A, holotype specimen, Type No. 2009427, housed in USNM; B, lateral of prodorsum with arrow to taenidium; C, preanal organ indicated by arrow; D, region of tubule T 1, h 3 and gla, with small arrows to tubule and large arrow to its opening (3 layers); E, lamella (5 layers), with arrow to nerve canal for seta le, lateral of podosoma displaced so Ah visible, black arrow to fused dorsophragmata; F, ventral region showing discidium (dis) and circumpedal carina (cc), with arrow to minute custodium (3 layers); G, pedipalp (5 layers), with arrow to solenidion ω. Scale bars: A = 100 µm; C, D, F = 25 µm; E = 40 µm; G = 30 µm." httpUri="https://zenodo.org/record/1018647/files/figure.png" pageId="14" pageNumber="165">Fig. 8C</figureCitation>
).
</paragraph>
<paragraph id="1514746EFF90FFADFF42C9FAFAD2FA37" blockId="14.[151,1437,151,2018]" pageId="14" pageNumber="165">
<taxonomicName id="D2AB0FEDFF90FFADFF42C9FAFECAFA4F" box="[199,342,1414,1439]" class="Secernentea" family="Gnathostomatidae" genus="Gnathosoma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirurida" pageId="14" pageNumber="165" phylum="Nematoda" rank="genus">
<emphasis id="27DFA87CFF90FFADFF42C9FAFECAFA4F" box="[199,342,1414,1439]" italics="true" pageId="14" pageNumber="165">Gnathosoma</emphasis>
</taxonomicName>
. Axillary saccule present at base of palp. Chelicera chelate-dentate (
<figureCitation id="8D9068EBFF90FFADFBFBC9FAFB74FA4F" box="[1150,1256,1414,1439]" captionStart="FIGURE 13" captionStartId="23.[151,250,1906,1928]" captionTargetBox="[213,1359,193,1878]" captionTargetId="figure@23.[213,1359,193,1878]" captionTargetPageId="23" captionText="FIGURE 13. Tegoribates americanus Hammer, differential interference contrast microscope images of adult. A, lateral of prodorsum (6 layers) with arrows to taenidium; B, bothridial seta and bothridium, with arrow to anterior scale; C, chelicera, pedipalp and rutellum showing size relationship, supracoxal seta e, and with arrow to axillary saccule (5 layers); D, tarsus II, showing shape of seta pv ”; E, postanal porose area. Scale bars: A, B = 50 µm; C = 25 µm." httpUri="https://zenodo.org/record/1018657/files/figure.png" pageId="14" pageNumber="165">Fig. 13C</figureCitation>
). Mentum with tectum covering camerostome when elevated (
<figureCitation id="8D9068EBFF90FFADFD1AC9D6FD61FA13" box="[671,765,1450,1475]" captionStart="FIGURES 10 – 11" captionStartId="21.[151,264,1974,1996]" captionTargetBox="[293,1284,213,1897]" captionTargetId="figure@21.[234,1338,193,1940]" captionTargetPageId="21" captionText="FIGURES 10 – 11. Tegoribates americanus Hammer, adult female, legs removed; 10, ventral aspect; 11, lateral aspect of prodorsum and podosoma, gnathosoma removed, with arrow to taenidium. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018653/files/figure.png" pageId="14" pageNumber="165">Figs. 10</figureCitation>
,
<figureCitation id="8D9068EBFF90FFADFC8CC9D7FCC5FA13" box="[777,857,1451,1475]" captionStart="FIGURE 21" captionStartId="33.[151,250,1886,1908]" captionTargetBox="[225,1362,193,1858]" captionTargetId="figure@33.[225,1362,193,1865]" captionTargetPageId="33" captionText="FIGURE 21. Tegoribates walteri sp. nov., scanning electron microscope images of adult female from California, A. habitus dorsal, with arrow to spine-like seta l ” of genu II; B, habitus ventral, with arrow to ventral depression; C, frontal aspect, with arrows to spine-like setae l ” of genua I and II; D, anterolateral of prodorsum and part of notogaster, with arrow to spine-like seta l ” of genu II; E, ventrofrontal aspect, with arrow to taenidium. Scale bars: A, B = 50 µm; C-E = 25 µm." httpUri="https://zenodo.org/record/1018675/files/figure.png" pageId="14" pageNumber="165">21C, E</figureCitation>
). Palp setal formula 0-2-1-3-9(1); eupathidium
<emphasis id="27DFA87CFF90FFADFAE8C9D0FA00FA13" box="[1389,1436,1452,1475]" italics="true" pageId="14" pageNumber="165">acm</emphasis>
subequal to solenidion, forming double horn with solenidion along length, bases slightly separate (
<figureCitation id="8D9068EBFF90FFADFB62C9B2FADDFA37" box="[1255,1345,1486,1511]" captionStart="FIGURE 8" captionStartId="17.[151,250,1857,1879]" captionTargetBox="[219,1314,198,1804]" captionTargetId="figure@17.[219,1314,193,1804]" captionTargetPageId="17" captionText="FIGURE 8 A – H. Tegoribates subniger Ewing, differential interference contrast microscope images of adult. A, holotype specimen, Type No. 2009427, housed in USNM; B, lateral of prodorsum with arrow to taenidium; C, preanal organ indicated by arrow; D, region of tubule T 1, h 3 and gla, with small arrows to tubule and large arrow to its opening (3 layers); E, lamella (5 layers), with arrow to nerve canal for seta le, lateral of podosoma displaced so Ah visible, black arrow to fused dorsophragmata; F, ventral region showing discidium (dis) and circumpedal carina (cc), with arrow to minute custodium (3 layers); G, pedipalp (5 layers), with arrow to solenidion ω. Scale bars: A = 100 µm; C, D, F = 25 µm; E = 40 µm; G = 30 µm." httpUri="https://zenodo.org/record/1018647/files/figure.png" pageId="14" pageNumber="165">Fig. 8G</figureCitation>
).
</paragraph>
<paragraph id="1514746EFF90FFADFF42C988FE4BF8D9" blockId="14.[151,1437,151,2018]" pageId="14" pageNumber="165">
<emphasis id="27DFA87CFF90FFADFF42C988FF60F9DB" box="[199,252,1524,1547]" italics="true" pageId="14" pageNumber="165">Legs</emphasis>
. Monodactylous or heterotridactylous. Leg setal formulas I–IV, solenidia in parentheses: I: 1-5-3(1)-4(2)- 20(2); II: 1-5-3(1)-4(1)-15(2); III: 2-3-1(1)-3(1)-15; IV: 1-2-2-3(1)-12. Genua I and II without tooth ventrally (
<figureCitation id="8D9068EBFF90FFADFAEFCA6AFF77F983" captionStart="FIGURE 12" captionStartId="22.[151,250,1970,1992]" captionTargetBox="[236,1371,232,1949]" captionTargetId="figure@22.[204,1382,193,1949]" captionTargetPageId="22" captionText="FIGURE 12 A – C. Tegoribates americanus Hammer, adult female, legs, all antiaxial aspect (trochanter not shown in A, B): A, leg I, with arrow to bend in femur; B, leg II (femur to tibia), C, leg III. Scale bar = 20 µm." httpUri="https://zenodo.org/record/1018655/files/figure.png" pageId="14" pageNumber="165">Figs 12A, B</figureCitation>
). Porose areas on femora I to IV and trochanters III and IV. Femur I with obtuse bend medially ventral to seta
<emphasis id="27DFA87CFF90FFADFF4FCA22FF44F9A7" box="[202,216,1630,1655]" italics="true" pageId="14" pageNumber="165">d</emphasis>
(
<figureCitation id="8D9068EBFF90FFADFF6CCA22FECCF9A7" box="[233,336,1630,1655]" captionStart="FIGURE 12" captionStartId="22.[151,250,1970,1992]" captionTargetBox="[236,1371,232,1949]" captionTargetId="figure@22.[204,1382,193,1949]" captionTargetPageId="22" captionText="FIGURE 12 A – C. Tegoribates americanus Hammer, adult female, legs, all antiaxial aspect (trochanter not shown in A, B): A, leg I, with arrow to bend in femur; B, leg II (femur to tibia), C, leg III. Scale bar = 20 µm." httpUri="https://zenodo.org/record/1018655/files/figure.png" pageId="14" pageNumber="165">Fig. 12A</figureCitation>
,
<figureCitation id="8D9068EBFF90FFADFEDBCA22FE58F9A7" box="[350,452,1630,1655]" captionStart="FIGURE 22" captionStartId="34.[151,250,1827,1849]" captionTargetBox="[222,1338,193,1774]" captionTargetId="figure@34.[222,1338,193,1794]" captionTargetPageId="34" captionText="FIGURE 22. Differential interference contrast microscope images of adults; A – D, Tegoribates walteri sp. nov., A, part of prodorsum (4 layers) showing lamella, bothridium, bothridial seta, interlamellar seta, with arrow to fused dorsophragmata; B, hinged pteromorph (3 layers); C, coxisternal region showing custodium (arrow) and pedotectum II (3 layers); D, porose area Aa and surrounding muscle sigillae (5 layers). E, Tegoribates subniger Ewing, lateral of podosoma, showing elongated, bent femur I (indicated by arrow); F, G, Protectoribates occidentalis sp. nov., detail of preanal organ; H, Hypozetes laysanensis Aoki, detail of preanal organ; I, Tectoribates borealis Behan-Pelletier &amp; Walter, detail of preanal organ. Scale bars: A, B, E = 25 µm; C, D = 20 µm." httpUri="https://zenodo.org/record/1018677/files/figure.png" pageId="14" pageNumber="165">Fig. 22E</figureCitation>
). Tibia II without anterodorsal dentate carina. Seta
<emphasis id="27DFA87CFF90FFADFB8FCA1CFB89F9A7" box="[1034,1045,1632,1655]" italics="true" pageId="14" pageNumber="165">s</emphasis>
on tarsus I eupathidial, positioned distal of setae (
<emphasis id="27DFA87CFF90FFADFEC7CAF8FECCF94B" box="[322,336,1668,1691]" italics="true" pageId="14" pageNumber="165">a</emphasis>
). Solenidia on tarsus II shorter than setae on segment;
<emphasis id="27DFA87CFF90FFADFC3FCAF8FC4AF94F" box="[954,982,1668,1695]" italics="true" pageId="14" pageNumber="165">ω1</emphasis>
not most proximal setiform structure on segment;. Genua I and II with distinctive and spinous seta
<emphasis id="27DFA87CFF90FFADFCB6CAD5FCD7F912" box="[819,843,1705,1730]" italics="true" pageId="14" pageNumber="165">l”</emphasis>
(
<figureCitation id="8D9068EBFF90FFADFCDECAD5FC51F912" box="[859,973,1705,1730]" captionStart="FIGURE 12" captionStartId="22.[151,250,1970,1992]" captionTargetBox="[236,1371,232,1949]" captionTargetId="figure@22.[204,1382,193,1949]" captionTargetPageId="22" captionText="FIGURE 12 A – C. Tegoribates americanus Hammer, adult female, legs, all antiaxial aspect (trochanter not shown in A, B): A, leg I, with arrow to bend in femur; B, leg II (femur to tibia), C, leg III. Scale bar = 20 µm." httpUri="https://zenodo.org/record/1018655/files/figure.png" pageId="14" pageNumber="165">Figs. 12B</figureCitation>
,
<figureCitation id="8D9068EBFF90FFADFC5FCAD6FB9AF912" box="[986,1030,1706,1730]" captionStart="FIGURE 19" captionStartId="30.[151,250,1905,1927]" captionTargetBox="[213,1359,193,1849]" captionTargetId="figure@30.[213,1359,193,1878]" captionTargetPageId="30" captionText="FIGURE 19. Tegoribates bryophilus Woolley, differential interference contrast microscope images of adult type material. A. anterior of prodorsum (note, specimen broken), with arrow to taenidium; B, pteromorph; C, saccule Sa and seta lm; D, posterior of prodorsum with arrow to fused dorsophragmata (3 layers); E, lateral of coxisternal region showing circumpedal carina and discidium; F, lenticulus; G, seta l ” of genu II. Scale bars: A, B, E = 50 µm; C, = 10 µm; D = 35 µm." httpUri="https://zenodo.org/record/1018669/files/figure.png" pageId="14" pageNumber="165">19F</figureCitation>
, insert, 21C). Femur III with seta
<emphasis id="27DFA87CFF90FFADFA0ECAD5FA00F912" box="[1419,1436,1705,1730]" italics="true" pageId="14" pageNumber="165">l’</emphasis>
present; seta
<emphasis id="27DFA87CFF90FFADFEADCAB2FEA7F935" box="[296,315,1742,1765]" italics="true" pageId="14" pageNumber="165">v'</emphasis>
of genua I and II present. On tibiae I and II seta
<emphasis id="27DFA87CFF90FFADFCD3CAB1FCF2F936" box="[854,878,1741,1766]" italics="true" pageId="14" pageNumber="165">l”</emphasis>
slightly more spinous than
<emphasis id="27DFA87CFF90FFADFB27CAB1FB2FF936" box="[1186,1203,1741,1766]" italics="true" pageId="14" pageNumber="165">l’</emphasis>
. Anterodorsal knobs or spines absent from tibia I.
</paragraph>
</subSubSection>
<subSubSection id="5DB127E5FF90FFACFF42CB69FA01F8A0" lastPageId="15" lastPageNumber="166" pageId="14" pageNumber="165" type="discussion">
<paragraph id="1514746EFF90FFADFF42CB69FA0AF882" blockId="14.[151,1437,151,2018]" pageId="14" pageNumber="165">
<emphasis id="27DFA87CFF90FFADFF42CB69FEA8F8FE" bold="true" box="[199,308,1813,1838]" pageId="14" pageNumber="165">Remarks</emphasis>
. 1.
<bibRefCitation id="713A099FFF90FFADFEE0CB69FD8AF8FE" author="Sellnick, M." box="[357,534,1813,1838]" journalOrPublisher="Schriften der physik. - okonom. Gesellschaft zu Konigsberg, 61 - 62" pageId="14" pageNumber="165" pagination="35 - 42" refString="Sellnick, M. (1920) Neue und seltene Oribatiden aus Deutschland. Schriften der physik. - okonom. Gesellschaft zu Konigsberg, 61 - 62, 35 - 42." title="Neue und seltene Oribatiden aus Deutschland" type="book chapter" year="1920">Sellnick (1920)</bibRefCitation>
proposed
<taxonomicName id="D2AB0FEDFF90FFADFD16CB69FCB3F8FE" authorityName="Sellnick" authorityYear="1920" box="[659,815,1813,1838]" class="Arachnida" family="Tegoribatidae" genus="Lepidoribates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="165" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF90FFADFD16CB69FCB3F8FE" box="[659,815,1813,1838]" italics="true" pageId="14" pageNumber="165">Lepidoribates</emphasis>
</taxonomicName>
to accommodate
<taxonomicName id="D2AB0FEDFF90FFADFB85CB6AFA00F8FE" authority="Koch. Jacot (1929)" authorityName="Koch. Jacot" authorityYear="1929" box="[1024,1436,1813,1838]" class="Arachnida" family="Galumnidae" genus="Zetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="165" phylum="Arthropoda" rank="species" species="latirostris">
<emphasis id="27DFA87CFF90FFADFB85CB6AFB2FF8FE" box="[1024,1203,1813,1838]" italics="true" pageId="14" pageNumber="165">Zetes latirostris</emphasis>
Koch.
<bibRefCitation id="713A099FFF90FFADFA88CB6AFA00F8FE" author="Jacot, A. P." box="[1293,1436,1814,1838]" journalOrPublisher="Transactions of the American Microscopical Society" pageId="14" pageNumber="165" pagination="416 - 430" part="48" publicationUrl="https://doi.org/10.2307/3222059" refString="Jacot, A. P. (1929) Genera of Pterogasterine Oribatidae (Acarina). Transactions of the American Microscopical Society, 48, 416 - 430. https: // doi. org / 10.2307 / 3222059" title="Genera of Pterogasterine Oribatidae (Acarina)" type="journal article" year="1929">Jacot (1929)</bibRefCitation>
</taxonomicName>
considered it a junior synonym of
<taxonomicName id="D2AB0FEDFF90FFADFD93CB45FD05F882" authorityName="Ewing" authorityYear="1917" box="[534,665,1849,1874]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="165" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF90FFADFD93CB45FD05F882" box="[534,665,1849,1874]" italics="true" pageId="14" pageNumber="165">Tegoribates</emphasis>
</taxonomicName>
and this was recognized by
<bibRefCitation id="713A099FFF90FFADFC50CB45FB1EF882" author="Sellnick, M." box="[981,1154,1849,1874]" journalOrPublisher="Entomologiske Tidskrift" pageId="14" pageNumber="165" pagination="60 - 63" part="76" refString="Sellnick, M. (1955) Berichtigungen und Klarstellungen zu einigen meiner bisher beschriebenen Gattungen und Arten der Acari. Entomologiske Tidskrift, 76, 60 - 63." title="Berichtigungen und Klarstellungen zu einigen meiner bisher beschriebenen Gattungen und Arten der Acari" type="journal article" year="1955">Sellnick (1955)</bibRefCitation>
and subsequent authors.
</paragraph>
<paragraph id="1514746EFF90FFADFF42CB22FADCF832" blockId="14.[151,1437,151,2018]" pageId="14" pageNumber="165">
2. Other than a brief comment on larva and nymphs of
<taxonomicName id="D2AB0FEDFF90FFADFCCCCB23FB6BF8A6" authority="(Behan-Pelletier 2001)" authorityName="Behan-Pelletier" authorityYear="2001" baseAuthorityName="Behan-Pelletier" baseAuthorityYear="2001" box="[841,1271,1885,1910]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="165" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="27DFA87CFF90FFADFCCCCB23FC75F8A5" box="[841,1001,1886,1910]" italics="true" pageId="14" pageNumber="165">T. americanus</emphasis>
(
<bibRefCitation id="713A099FFF90FFADFC7FCB21FB73F8A6" author="Behan-Pelletier, V. M." box="[1018,1263,1885,1910]" editor="Halliday, R. B." journalOrPublisher="CSIRO Publishing, Melbourne" pageId="14" pageNumber="165" pagination="50 - 57" refString="Behan-Pelletier, V. M. (2001) Phylogenetic relationships of Hypozetes (Acari: Tegoribatidae). In: Halliday, R. B., Walter, D. E., Proctor, H. C., Norton, R. A. &amp; Colloff, M. J. (Eds.), Acarology: Proceedings of the 10 th International Congress. CSIRO Publishing, Melbourne, pp. 50 - 57." title="Phylogenetic relationships of Hypozetes (Acari: Tegoribatidae)" type="book chapter" volumeTitle="Acarology: Proceedings of the 10 th International Congress" year="2001">Behan-Pelletier 2001</bibRefCitation>
)
</taxonomicName>
, immatures of
<taxonomicName id="D2AB0FEDFF90FFADFF12CBFDFE86F84A" authorityName="Ewing" authorityYear="1917" box="[151,282,1921,1946]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="165" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF90FFADFF12CBFDFE86F84A" box="[151,282,1921,1946]" italics="true" pageId="14" pageNumber="165">Tegoribates</emphasis>
</taxonomicName>
have been known only for
<taxonomicName id="D2AB0FEDFF90FFADFDD0CBFEFD43F84A" authorityName="Koch. Jacot" authorityYear="1929" box="[597,735,1921,1946]" class="Arachnida" family="Galumnidae" genus="Zetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="165" phylum="Arthropoda" rank="species" species="latirostris">
<emphasis id="27DFA87CFF90FFADFDD0CBFEFD43F84A" box="[597,735,1921,1946]" italics="true" pageId="14" pageNumber="165">T. latirostris</emphasis>
</taxonomicName>
(larva and nymph;
<bibRefCitation id="713A099FFF90FFADFC45CBFEFBD6F84A" author="Tuxen, S. L." box="[960,1098,1922,1946]" journalOrPublisher="Entomologiske Meddelleilser" pageId="14" pageNumber="165" pagination="321 - 336" part="23" refString="Tuxen, S. L. (1943) Die zeitliche und raumliche Verteilung der Oribatiden-Fauna (Acar.) bei Maelifell, Nord-Island. Entomologiske Meddelleilser, 23, 321 - 336." title="Die zeitliche und raumliche Verteilung der Oribatiden-Fauna (Acar.) bei Maelifell, Nord-Island" type="journal article" year="1943">Tuxen 1943</bibRefCitation>
,
<bibRefCitation id="713A099FFF90FFADFBD2CBFDFA93F84A" author="Grandjean, F." box="[1111,1295,1921,1946]" journalOrPublisher="Bulletin de la Societe Zoologique de France" pageId="14" pageNumber="165" pagination="421 - 446" part="78" refString="Grandjean, F. (1954) Essai de classification des Oribates (Acariens). Bulletin de la Societe Zoologique de France, 78, 421 - 446." title="Essai de classification des Oribates (Acariens)" type="journal article" year="1954">Grandjean 1954</bibRefCitation>
,
<bibRefCitation id="713A099FFF90FFADFA9ECBFEFEAAF86E" author="Norton, R. A. &amp; Ermilov, S. G." journalOrPublisher="Zootaxa" pageId="14" pageNumber="165" pagination="1 - 132" part="3833" publicationUrl="https://doi.org/10.11646/zootaxa.3833.1.1" refString="Norton, R. A. &amp; Ermilov, S. G. (2014) Catalogue and historical overview of juvenile instars of oribatid mites (Acari: Oribatida). Zootaxa, 3833 (1), 1 - 132. https: // doi. org / 10.11646 / zootaxa. 3833.1.1" title="Catalogue and historical overview of juvenile instars of oribatid mites (Acari: Oribatida)" type="journal article" year="2014">Norton and Ermilov 2014</bibRefCitation>
).
<bibRefCitation id="713A099FFF90FFADFECDCBDAFDF8F86E" author="Tuxen, S. L." box="[328,612,1957,1982]" journalOrPublisher="Entomologiske Meddelleilser" pageId="14" pageNumber="165" pagination="321 - 336" part="23" refString="Tuxen, S. L. (1943) Die zeitliche und raumliche Verteilung der Oribatiden-Fauna (Acar.) bei Maelifell, Nord-Island. Entomologiske Meddelleilser, 23, 321 - 336." title="Die zeitliche und raumliche Verteilung der Oribatiden-Fauna (Acar.) bei Maelifell, Nord-Island" type="journal article" year="1943">
Tuxen (1943
<figureCitation id="8D9068EBFF90FFADFE58CBD9FDAFF86E" box="[477,563,1957,1982]" captionStart="FIGURE 12" captionStartId="22.[151,250,1970,1992]" captionTargetBox="[236,1371,232,1949]" captionTargetId="figure@22.[204,1382,193,1949]" captionTargetPageId="22" captionText="FIGURE 12 A – C. Tegoribates americanus Hammer, adult female, legs, all antiaxial aspect (trochanter not shown in A, B): A, leg I, with arrow to bend in femur; B, leg II (femur to tibia), C, leg III. Scale bar = 20 µm." httpUri="https://zenodo.org/record/1018655/files/figure.png" pageId="14" pageNumber="165">Figs 12</figureCitation>
,
<figureCitation id="8D9068EBFF90FFADFDBBCBDAFDC7F86E" box="[574,603,1958,1982]" captionStart="FIGURE 13" captionStartId="23.[151,250,1906,1928]" captionTargetBox="[213,1359,193,1878]" captionTargetId="figure@23.[213,1359,193,1878]" captionTargetPageId="23" captionText="FIGURE 13. Tegoribates americanus Hammer, differential interference contrast microscope images of adult. A, lateral of prodorsum (6 layers) with arrows to taenidium; B, bothridial seta and bothridium, with arrow to anterior scale; C, chelicera, pedipalp and rutellum showing size relationship, supracoxal seta e, and with arrow to axillary saccule (5 layers); D, tarsus II, showing shape of seta pv ”; E, postanal porose area. Scale bars: A, B = 50 µm; C = 25 µm." httpUri="https://zenodo.org/record/1018657/files/figure.png" pageId="14" pageNumber="165">13</figureCitation>
)
</bibRefCitation>
described a “spinous knob” on the dorsal of the hysterosoma of the larva; this is the hysterosomal region bearing setae
<emphasis id="27DFA87CFF90FFADFD0BCBB5FD36F832" box="[654,682,1993,2018]" italics="true" pageId="14" pageNumber="165">dp</emphasis>
, as found in other species of
<taxonomicName id="D2AB0FEDFF90FFADFC77CBB5FBE9F832" authorityName="Ewing" authorityYear="1917" box="[1010,1141,1993,2018]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="14" pageNumber="165" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF90FFADFC77CBB5FBE9F832" box="[1010,1141,1993,2018]" italics="true" pageId="14" pageNumber="165">Tegoribates</emphasis>
</taxonomicName>
, described below.
</paragraph>
<paragraph id="1514746EFF91FFACFF4BCCE4FC66FD54" blockId="15.[151,1437,151,1904]" pageId="15" pageNumber="166">
3.
<bibRefCitation id="713A099FFF91FFACFF6ECCEBFE2EFF60" author="Grandjean, F." box="[235,434,151,176]" journalOrPublisher="Bulletin de la Societe Zoologique de France" pageId="15" pageNumber="166" pagination="421 - 446" part="78" refString="Grandjean, F. (1954) Essai de classification des Oribates (Acariens). Bulletin de la Societe Zoologique de France, 78, 421 - 446." title="Essai de classification des Oribates (Acariens)" type="journal article" year="1954">Grandjean (1954)</bibRefCitation>
diagnosed
<taxonomicName id="D2AB0FEDFF91FFACFDB7CCEBFD29FF60" authorityName="Ewing" authorityYear="1917" box="[562,693,151,176]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF91FFACFDB7CCEBFD29FF60" box="[562,693,151,176]" italics="true" pageId="15" pageNumber="166">Tegoribates</emphasis>
</taxonomicName>
as lacking a genal notch. This is the case in all described species. However, a taenidium is present in the same region in all members of this genus, running from close to the border of the rostrum posteriorly to acetabulum I. Following the terminology of
<bibRefCitation id="713A099FFF91FFACFC5ECCA3FB00FF28" author="Grandjean, F." box="[987,1180,223,248]" journalOrPublisher="Acarologia" pageId="15" pageNumber="166" pagination="357 - 391" part="10" refString="Grandjean, F. (1968) Nouvelles observations sur les Oribates (6 e serie). Acarologia, 10, 357 - 391" title="Nouvelles observations sur les Oribates (6 e serie)" type="journal article" year="1968">Grandjean (1968</bibRefCitation>
; see also
<bibRefCitation id="713A099FFF91FFACFA8ECC9CFA0CFF28" author="Trave, J." box="[1291,1424,223,248]" journalOrPublisher="Acarologia" pageId="15" pageNumber="166" pagination="85 - 94" part="27" refString="Trave, J. (1986) Les taenidies respiratoires des Oribates. Acarologia, 27 (1), 85 - 94." title="Les taenidies respiratoires des Oribates" type="journal article" year="1986">Travé 1986</bibRefCitation>
), this appears to be a “respiratory taenidium”, which he defined as a furrow, almost closed or not, banded or smooth, hollowed into the integument, and communicating with the stigma of trachea I or with those of tracheae I, sejugal and III. The taenidium of
<taxonomicName id="D2AB0FEDFF91FFACFE4ACD30FDCEFEB5" authorityName="Ewing" authorityYear="1917" box="[463,594,332,357]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF91FFACFE4ACD30FDCEFEB5" box="[463,594,332,357]" italics="true" pageId="15" pageNumber="166">Tegoribates</emphasis>
</taxonomicName>
extends to acetabulum I, and possibly communicates with trachea I. A taenidium, with a similar morphology, is found in the same region in the licneremaeoid genus
<emphasis id="27DFA87CFF91FFACFB3BCD13FAD5FE58" box="[1214,1353,367,392]" italics="true" pageId="15" pageNumber="166">Phylleremus</emphasis>
, which is a unique expression in the superfamily (
<bibRefCitation id="713A099FFF91FFACFDEACDE8FC4DFE7C" author="Behan-Pelletier, V. M. &amp; Walter, D. E." box="[623,977,404,429]" journalOrPublisher="Zootaxa" pageId="15" pageNumber="166" pagination="1 - 17" part="1386" refString="Behan-Pelletier, V. M. &amp; Walter, D. E. (2007) Phylleremus n. gen. from leaves of deciduous trees in eastern Australia (Oribatida: Licneremaeoidea), Zootaxa, 1386, 1 - 17." title="Phylleremus n. gen. from leaves of deciduous trees in eastern Australia (Oribatida: Licneremaeoidea)" type="journal article" year="2007">Behan-Pelletier &amp; Walter 2007</bibRefCitation>
). A ‘taenidial gutter’ is also found on the prodorsum of the oppioid
<taxonomicName id="D2AB0FEDFF91FFACFE45CDC5FC68FE00" authority="Wauthy &amp; Ducarme, 2006" authorityName="Wauthy &amp; Ducarme" authorityYear="2006" box="[448,1012,439,464]" class="Arachnida" family="Oppiidae" genus="Hypogeoppia" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="belgicae">
<emphasis id="27DFA87CFF91FFACFE45CDC5FD22FE00" box="[448,702,439,464]" italics="true" pageId="15" pageNumber="166">Hypogeoppia belgicae</emphasis>
<bibRefCitation id="713A099FFF91FFACFD43CDCBFC68FE00" author="Wauthy, G. &amp; Ducarme, X." box="[710,1012,439,464]" journalOrPublisher="Belgian Journal Zoology" pageId="15" pageNumber="166" pagination="203 - 218" part="136" refString="Wauthy, G. &amp; Ducarme, X. (2006) Description of Hypogeoppia belgicae, a new species of cave mites (Acari, Oribatida), and comments on some characters. Belgian Journal Zoology, 136 (2), 203 - 218." title="Description of Hypogeoppia belgicae, a new species of cave mites (Acari, Oribatida), and comments on some characters" type="journal article" year="2006">Wauthy &amp; Ducarme, 2006</bibRefCitation>
</taxonomicName>
.
<bibRefCitation id="713A099FFF91FFACFC7ACDC4FB0FFE00" author="Trave, J." box="[1023,1171,439,464]" journalOrPublisher="Acarologia" pageId="15" pageNumber="166" pagination="85 - 94" part="27" refString="Trave, J. (1986) Les taenidies respiratoires des Oribates. Acarologia, 27 (1), 85 - 94." title="Les taenidies respiratoires des Oribates" type="journal article" year="1986">Travé (1986)</bibRefCitation>
noted how easy it is to overlook this structure. When viewed with incident illumination of flattened, slide-mounted specimens the taenidium appears as in
<figureCitation id="8D9068EBFF91FFACFE32CD83FD81FDC8" box="[439,541,511,536]" captionStart="FIGURE 8" captionStartId="17.[151,250,1857,1879]" captionTargetBox="[219,1314,198,1804]" captionTargetId="figure@17.[219,1314,193,1804]" captionTargetPageId="17" captionText="FIGURE 8 A – H. Tegoribates subniger Ewing, differential interference contrast microscope images of adult. A, holotype specimen, Type No. 2009427, housed in USNM; B, lateral of prodorsum with arrow to taenidium; C, preanal organ indicated by arrow; D, region of tubule T 1, h 3 and gla, with small arrows to tubule and large arrow to its opening (3 layers); E, lamella (5 layers), with arrow to nerve canal for seta le, lateral of podosoma displaced so Ah visible, black arrow to fused dorsophragmata; F, ventral region showing discidium (dis) and circumpedal carina (cc), with arrow to minute custodium (3 layers); G, pedipalp (5 layers), with arrow to solenidion ω. Scale bars: A = 100 µm; C, D, F = 25 µm; E = 40 µm; G = 30 µm." httpUri="https://zenodo.org/record/1018647/files/figure.png" pageId="15" pageNumber="166">Figs. 8B</figureCitation>
,
<figureCitation id="8D9068EBFF91FFACFDABCE7CFDFCFDC8" box="[558,608,512,536]" captionStart="FIGURE 13" captionStartId="23.[151,250,1906,1928]" captionTargetBox="[213,1359,193,1878]" captionTargetId="figure@23.[213,1359,193,1878]" captionTargetPageId="23" captionText="FIGURE 13. Tegoribates americanus Hammer, differential interference contrast microscope images of adult. A, lateral of prodorsum (6 layers) with arrows to taenidium; B, bothridial seta and bothridium, with arrow to anterior scale; C, chelicera, pedipalp and rutellum showing size relationship, supracoxal seta e, and with arrow to axillary saccule (5 layers); D, tarsus II, showing shape of seta pv ”; E, postanal porose area. Scale bars: A, B = 50 µm; C = 25 µm." httpUri="https://zenodo.org/record/1018657/files/figure.png" pageId="15" pageNumber="166">13A</figureCitation>
, and
<figureCitation id="8D9068EBFF91FFACFD21CE7CFD4BFDC8" box="[676,727,512,536]" captionStart="FIGURE 19" captionStartId="30.[151,250,1905,1927]" captionTargetBox="[213,1359,193,1849]" captionTargetId="figure@30.[213,1359,193,1878]" captionTargetPageId="30" captionText="FIGURE 19. Tegoribates bryophilus Woolley, differential interference contrast microscope images of adult type material. A. anterior of prodorsum (note, specimen broken), with arrow to taenidium; B, pteromorph; C, saccule Sa and seta lm; D, posterior of prodorsum with arrow to fused dorsophragmata (3 layers); E, lateral of coxisternal region showing circumpedal carina and discidium; F, lenticulus; G, seta l ” of genu II. Scale bars: A, B, E = 50 µm; C, = 10 µm; D = 35 µm." httpUri="https://zenodo.org/record/1018669/files/figure.png" pageId="15" pageNumber="166">19A</figureCitation>
; on heavily sclerotized whole-mounts, it has the appearance indicated in
<figureCitation id="8D9068EBFF91FFACFEA9CE58FE51FDEC" box="[300,461,548,573]" captionStart="FIGURE 21" captionStartId="33.[151,250,1886,1908]" captionTargetBox="[225,1362,193,1858]" captionTargetId="figure@33.[225,1362,193,1865]" captionTargetPageId="33" captionText="FIGURE 21. Tegoribates walteri sp. nov., scanning electron microscope images of adult female from California, A. habitus dorsal, with arrow to spine-like seta l ” of genu II; B, habitus ventral, with arrow to ventral depression; C, frontal aspect, with arrows to spine-like setae l ” of genua I and II; D, anterolateral of prodorsum and part of notogaster, with arrow to spine-like seta l ” of genu II; E, ventrofrontal aspect, with arrow to taenidium. Scale bars: A, B = 50 µm; C-E = 25 µm." httpUri="https://zenodo.org/record/1018675/files/figure.png" pageId="15" pageNumber="166">Figs 21 D, E</figureCitation>
. This is the lateral structure outlined by
<bibRefCitation id="713A099FFF91FFACFC38CE58FB07FDED" author="Engelbrecht, C. M." box="[957,1179,548,573]" journalOrPublisher="Navorsinge van die Nasionale Museum, Bloemfontein" pageId="15" pageNumber="166" pagination="169 - 250" part="5" refString="Engelbrecht, C. M. (1986) New oribatid taxa and distribution records predominantly from Southern Africa (Acari: Oribatei: Oribatelloidea. Navorsinge van die Nasionale Museum, Bloemfontein, 5 (9), 169 - 250." title="New oribatid taxa and distribution records predominantly from Southern Africa (Acari: Oribatei: Oribatelloidea" type="journal article" year="1986">Engelbrecht (1986)</bibRefCitation>
on the prodorsum of
<taxonomicName id="D2AB0FEDFF91FFACFF12CE3BFE09FDB0" authorityName="Engelbrecht" authorityYear="1986" box="[151,405,583,608]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="natalensis">
<emphasis id="27DFA87CFF91FFACFF12CE3BFE09FDB0" box="[151,405,583,608]" italics="true" pageId="15" pageNumber="166">Tegoribates natalensis</emphasis>
</taxonomicName>
(his Fig. 65).
<taxonomicName id="D2AB0FEDFF91FFACFDBCCE35FD30FDB0" authorityName="Balogh" authorityYear="1959" box="[569,684,585,608]" class="Arachnida" family="Austrachipteriidae" genus="Hypozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF91FFACFDBCCE35FD30FDB0" box="[569,684,585,608]" italics="true" pageId="15" pageNumber="166">Hypozetes</emphasis>
</taxonomicName>
also lacks a genal notch, and while the rostral ridge has a narrow tectum there is no evidence of a taenidium (
<bibRefCitation id="713A099FFF91FFACFD05CE10FC70FD54" author="Behan-Pelletier, V. M." box="[640,1004,620,645]" editor="Halliday, R. B." journalOrPublisher="CSIRO Publishing, Melbourne" pageId="15" pageNumber="166" pagination="50 - 57" refString="Behan-Pelletier, V. M. (2001) Phylogenetic relationships of Hypozetes (Acari: Tegoribatidae). In: Halliday, R. B., Walter, D. E., Proctor, H. C., Norton, R. A. &amp; Colloff, M. J. (Eds.), Acarology: Proceedings of the 10 th International Congress. CSIRO Publishing, Melbourne, pp. 50 - 57." title="Phylogenetic relationships of Hypozetes (Acari: Tegoribatidae)" type="book chapter" volumeTitle="Acarology: Proceedings of the 10 th International Congress" year="2001">
Behan-Pelletier 2001, her
<figureCitation id="8D9068EBFF91FFACFC2DCE10FC70FD54" box="[936,1004,620,645]" captionStart="FIGURES 2 – 3" captionStartId="7.[151,264,1946,1968]" captionTargetBox="[335,1280,208,1896]" captionTargetId="figure@7.[253,1333,193,1924]" captionTargetPageId="7" captionText="FIGURES 2 – 3. Protectoribates occidentalis sp. nov., adult female, legs not illustrated; 2, ventral aspect; 3, lateral aspect, only alveolus of seta ex illustrated, gnathosoma not illustrated. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018635/files/figure.png" pageId="15" pageNumber="166">Fig. 3</figureCitation>
</bibRefCitation>
).
</paragraph>
<paragraph id="1514746EFF91FFACFF42CEECFAC8FB88" blockId="15.[151,1437,151,1904]" pageId="15" pageNumber="166">
4. The octotaxic system in
<taxonomicName id="D2AB0FEDFF91FFACFE79CEF3FDE3FD78" authorityName="Ewing" authorityYear="1917" box="[508,639,655,680]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF91FFACFE79CEF3FDE3FD78" box="[508,639,655,680]" italics="true" pageId="15" pageNumber="166">Tegoribates</emphasis>
</taxonomicName>
is developed in various forms.
<bibRefCitation id="713A099FFF91FFACFC6CCEF3FB35FD78" author="Grandjean, F." box="[1001,1193,655,680]" journalOrPublisher="Bulletin du Museum national d´Histoire naturelle, 2 e Serie" pageId="15" pageNumber="166" pagination="286 - 293" part="25" refString="Grandjean, F. (1953) Observations sur les Oribates (26 e serie). Bulletin du Museum national d´Histoire naturelle, 2 e Serie, 25 (3), 286 - 293." title="Observations sur les Oribates (26 e serie)" type="journal article" year="1953">Grandjean (1953</bibRefCitation>
,
<bibRefCitation id="713A099FFF91FFACFB32CEF3FB6CFD78" author="Grandjean, F." box="[1207,1264,655,680]" journalOrPublisher="Bulletin de la Societe Zoologique de France" pageId="15" pageNumber="166" pagination="421 - 446" part="78" refString="Grandjean, F. (1954) Essai de classification des Oribates (Acariens). Bulletin de la Societe Zoologique de France, 78, 421 - 446." title="Essai de classification des Oribates (Acariens)" type="journal article" year="1954">1954</bibRefCitation>
) noted that the octotaxic system is represented by 4 pairs of small “trachea”, called ‘tubules’ by
<bibRefCitation id="713A099FFF91FFACFBA2CEC9FA9EFD1C" author="Alberti, G. &amp; Norton, R. A. &amp; Adis, J. &amp; Fernandez, N. A. &amp; Franklin, E. &amp; Kratzmann, M. &amp; Moreno, A. &amp; Weigmann, G. &amp; Woas, S." box="[1063,1282,692,717]" journalOrPublisher="Zoologica, Stuttgart" pageId="15" pageNumber="166" pagination="33 - 114" part="146" refString="Alberti, G., Norton, R. A., Adis, J., Fernandez, N. A., Franklin, E., Kratzmann, M., Moreno, A. I., Weigmann, G. &amp; Woas, S. (1997) Porose integumental organs of oribatid mites (Acari, Oribatida). 2. Fine structure. Zoologica, Stuttgart, 146, 33 - 114." title="Porose integumental organs of oribatid mites (Acari, Oribatida). 2. Fine structure" type="journal article" year="1997">
Norton
<emphasis id="27DFA87CFF91FFACFBFBCEC9FB33FD1D" box="[1150,1199,692,717]" italics="true" pageId="15" pageNumber="166">et al</emphasis>
. (1997
</bibRefCitation>
; their
<figureCitation id="8D9068EBFF91FFACFACCCEC8FA0FFD1D" box="[1353,1427,692,717]" captionStart="FIGURE 5" captionStartId="9.[151,250,1917,1939]" captionTargetBox="[207,1364,193,1894]" captionTargetId="figure@9.[207,1364,193,1894]" captionTargetPageId="9" captionText="FIGURE 5 A – E. Protectoribates occidentalis sp. nov., scanning electron microscope images of adult female from California; A. habitus dorsal; B, habitus ventral, with arrow to custodium; C, frontal aspect; D, dorsolateral of prodorsum, with arrow to spindle end of bothridial seta (bo), second arrow to genal incision and third arrow to carina on tibia II; E, ventrolateral of prodorsum, with arrow to ridge on genal tooth. Scale bars: A, B = 25 µm, C – E = 15 µm." httpUri="https://zenodo.org/record/1018639/files/figure.png" pageId="15" pageNumber="166">Fig. 5l</figureCitation>
) and herein. This is the expression of the octotaxic system in
<taxonomicName id="D2AB0FEDFF91FFACFCD5CEA5FC47FD20" authorityName="Koch. Jacot" authorityYear="1929" box="[848,987,727,752]" class="Arachnida" family="Galumnidae" genus="Zetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="latirostris">
<emphasis id="27DFA87CFF91FFACFCD5CEA5FC47FD20" box="[848,987,727,752]" italics="true" pageId="15" pageNumber="166">T. latirostris</emphasis>
</taxonomicName>
and
<taxonomicName id="D2AB0FEDFF91FFACFB90CEA5FB09FD20" authorityName="Ewing" authorityYear="1917" box="[1045,1173,727,752]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="subniger">
<emphasis id="27DFA87CFF91FFACFB90CEA5FB09FD20" box="[1045,1173,727,752]" italics="true" pageId="15" pageNumber="166">T. subniger</emphasis>
</taxonomicName>
(see below), and these tubules are closely associated with setae. I follow
<bibRefCitation id="713A099FFF91FFACFD59CE81FC5FFCC4" author="Alberti, G. &amp; Norton, R. A. &amp; Adis, J. &amp; Fernandez, N. A. &amp; Franklin, E. &amp; Kratzmann, M. &amp; Moreno, A. &amp; Weigmann, G. &amp; Woas, S." box="[732,963,764,789]" journalOrPublisher="Zoologica, Stuttgart" pageId="15" pageNumber="166" pagination="33 - 114" part="146" refString="Alberti, G., Norton, R. A., Adis, J., Fernandez, N. A., Franklin, E., Kratzmann, M., Moreno, A. I., Weigmann, G. &amp; Woas, S. (1997) Porose integumental organs of oribatid mites (Acari, Oribatida). 2. Fine structure. Zoologica, Stuttgart, 146, 33 - 114." title="Porose integumental organs of oribatid mites (Acari, Oribatida). 2. Fine structure" type="journal article" year="1997">
Norton
<emphasis id="27DFA87CFF91FFACFCB1CE81FCF4FCC5" box="[820,872,764,789]" italics="true" pageId="15" pageNumber="166">et al</emphasis>
. (1997)
</bibRefCitation>
and use the notation
<emphasis id="27DFA87CFF91FFACFB3ACE81FAB3FCC4" box="[1215,1327,764,788]" italics="true" pageId="15" pageNumber="166">Ta, T1-T3</emphasis>
for these structures, corresponding to the notation for porose areas (
<emphasis id="27DFA87CFF91FFACFCC7CF5CFC23FCE8" box="[834,959,800,824]" italics="true" pageId="15" pageNumber="166">Aa, A1–A3</emphasis>
) and saccules (
<emphasis id="27DFA87CFF91FFACFBF2CF5CFB77FCE8" box="[1143,1259,800,824]" italics="true" pageId="15" pageNumber="166">Sa, S1–S3</emphasis>
).
<bibRefCitation id="713A099FFF91FFACFA83CF5CFF7DFC8C" author="Alberti, G. &amp; Norton, R. A. &amp; Adis, J. &amp; Fernandez, N. A. &amp; Franklin, E. &amp; Kratzmann, M. &amp; Moreno, A. &amp; Weigmann, G. &amp; Woas, S." journalOrPublisher="Zoologica, Stuttgart" pageId="15" pageNumber="166" pagination="33 - 114" part="146" refString="Alberti, G., Norton, R. A., Adis, J., Fernandez, N. A., Franklin, E., Kratzmann, M., Moreno, A. I., Weigmann, G. &amp; Woas, S. (1997) Porose integumental organs of oribatid mites (Acari, Oribatida). 2. Fine structure. Zoologica, Stuttgart, 146, 33 - 114." title="Porose integumental organs of oribatid mites (Acari, Oribatida). 2. Fine structure" type="journal article" year="1997">
Norton
<emphasis id="27DFA87CFF91FFACFAE5CF5DFA08FCE8" box="[1376,1428,799,824]" italics="true" pageId="15" pageNumber="166">et al</emphasis>
. (1997)
</bibRefCitation>
also noted the presence of tubules in
<emphasis id="27DFA87CFF91FFACFD13CF38FCB1FC8D" box="[662,813,836,861]" italics="true" pageId="15" pageNumber="166">
<taxonomicName id="D2AB0FEDFF91FFACFD13CF38FCB5FC8D" authorityName="Popp" authorityYear="1960" box="[662,809,836,861]" class="Arachnida" family="Achipteriidae" genus="Plakoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">Plakoribates</taxonomicName>
.
</emphasis>
Fine tubules also are found in the ceratozetoid genera
<taxonomicName id="D2AB0FEDFF91FFACFF12CF1BFD9FFC50" authority=", Paralamellobates" authorityName="Paralamellobates" box="[151,515,871,896]" class="Arachnida" family="Austrachipteriidae" genus="Lamellobates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF91FFACFF12CF1BFD9FFC50" box="[151,515,871,896]" italics="true" pageId="15" pageNumber="166">Lamellobates, Paralamellobates</emphasis>
</taxonomicName>
and
<taxonomicName id="D2AB0FEDFF91FFACFDBECF1BFD56FC50" box="[571,714,871,896]" class="Arachnida" family="Austrachipteriidae" genus="Sacculozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF91FFACFDBECF1BFD56FC50" box="[571,714,871,896]" italics="true" pageId="15" pageNumber="166">Sacculozetes</emphasis>
</taxonomicName>
, but these are not closely associated with setae.
<bibRefCitation id="713A099FFF91FFACFB69CF1BFA00FC50" author="Woolley, T. A." box="[1260,1436,871,896]" journalOrPublisher="Transaction of the American Microscopical Society" pageId="15" pageNumber="166" pagination="409 – 412" part="84" publicationUrl="https://doi.org/10.2307/3224729" refString="Woolley, T. A. (1965) A New Species of Tegoribates from Colorado with Notes on the Genus (Acarina: Oribatei, Tegoribatidae). Transaction of the American Microscopical Society, 84, 409 - 412. https: // doi. org / 10.2307 / 3224729" title="A New Species of Tegoribates from Colorado with Notes on the Genus (Acarina: Oribatei, Tegoribatidae)" type="journal article" year="1965">Woolley (1965)</bibRefCitation>
expanded the diagnosis of
<taxonomicName id="D2AB0FEDFF91FFACFE3ACFF0FDDEFC75" authorityName="Ewing" authorityYear="1917" box="[447,578,908,933]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF91FFACFE3ACFF0FDDEFC75" box="[447,578,908,933]" italics="true" pageId="15" pageNumber="166">Tegoribates</emphasis>
</taxonomicName>
to include
<taxonomicName id="D2AB0FEDFF91FFACFD3ACFF1FCDBFC75" box="[703,839,908,933]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="bryophilu">
<emphasis id="27DFA87CFF91FFACFD3ACFF1FCDBFC75" box="[703,839,908,933]" italics="true" pageId="15" pageNumber="166">T. bryophilu</emphasis>
</taxonomicName>
s with the octotaxic system of saccules, an expression also found in
<taxonomicName id="D2AB0FEDFF91FFACFEBFCFCDFE40FC18" baseAuthorityName="Behan-Pelletier" baseAuthorityYear="2001" box="[314,476,945,968]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="americanus">
<emphasis id="27DFA87CFF91FFACFEBFCFCDFE40FC18" box="[314,476,945,968]" italics="true" pageId="15" pageNumber="166">T. americanus</emphasis>
</taxonomicName>
. The expression of the octotaxic system is further expanded in
<taxonomicName id="D2AB0FEDFF91FFACFB43CFCDFAA9FC18" authority="Behan-Pelletier, 2017" authorityName="Behan-Pelletier" authorityYear="2017" box="[1222,1333,943,968]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="walteri" status="sp. nov.">
<emphasis id="27DFA87CFF91FFACFB43CFCDFAA9FC18" box="[1222,1333,943,968]" italics="true" pageId="15" pageNumber="166">T. walteri</emphasis>
</taxonomicName>
<taxonomicNameLabel id="3CEC1507FF91FFACFABACFD3FA00FC18" box="[1343,1436,943,968]" pageId="15" pageNumber="166" rank="species">
<emphasis id="27DFA87CFF91FFACFABACFD3FA00FC18" bold="true" box="[1343,1436,943,968]" pageId="15" pageNumber="166">sp. nov.</emphasis>
</taxonomicNameLabel>
(described below) to include porose areas. The close association of the octotaxic system with setae is most evident in
<taxonomicName id="D2AB0FEDFF91FFACFF31CF85FEA0FBC0" authorityName="Koch. Jacot" authorityYear="1929" box="[180,316,1015,1040]" class="Arachnida" family="Galumnidae" genus="Zetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="latirostris">
<emphasis id="27DFA87CFF91FFACFF31CF85FEA0FBC0" box="[180,316,1015,1040]" italics="true" pageId="15" pageNumber="166">T. latirostris</emphasis>
</taxonomicName>
and
<taxonomicName id="D2AB0FEDFF91FFACFEF6CF85FE73FBC0" authorityName="Ewing" authorityYear="1917" box="[371,495,1015,1040]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="subniger">
<emphasis id="27DFA87CFF91FFACFEF6CF85FE73FBC0" box="[371,495,1015,1040]" italics="true" pageId="15" pageNumber="166">T. subniger</emphasis>
</taxonomicName>
. Other genera of poronotic
<taxonomicName id="D2AB0FEDFF91FFACFCA3CF8BFC21FBC0" authorityName="Hull" authorityYear="1918" box="[806,957,1015,1040]" class="Arachnida" infraOrder="Brachypylina" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="infraOrder">Brachypylina</taxonomicName>
have the octotaxic system as either porose areas or saccules, according to species (e.g.,
<taxonomicName id="D2AB0FEDFF91FFACFD63C860FC19FBE5" authorityName="Jacot" authorityYear="1925" box="[742,901,1052,1077]" class="Arachnida" family="Ceratozetidae" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="family">Ceratozetidae</taxonomicName>
:
<taxonomicName id="D2AB0FEDFF91FFACFC19C860FBAEFBE5" authorityName="Berlese" authorityYear="1910" box="[924,1074,1052,1077]" class="Arachnida" family="Ceratozetidae" genus="Trichoribates" higherTaxonomySource="GBIF" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF91FFACFC19C860FBAEFBE5" box="[924,1074,1052,1077]" italics="true" pageId="15" pageNumber="166">Trichoribates</emphasis>
</taxonomicName>
,
<taxonomicName id="D2AB0FEDFF91FFACFBCBC860FB77FBE5" authorityName="Thor" authorityYear="1929" box="[1102,1259,1052,1077]" class="Arachnida" family="Achipteriidae" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="family">Achipteriidae</taxonomicName>
:
<taxonomicName id="D2AB0FEDFF91FFACFA86C860FA09FBE5" box="[1283,1429,1052,1077]" class="Arachnida" family="Achipteriidae" genus="Anachipteria" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF91FFACFA86C860FA09FBE5" box="[1283,1429,1052,1077]" italics="true" pageId="15" pageNumber="166">Anachipteria</emphasis>
</taxonomicName>
,
<taxonomicName id="D2AB0FEDFF91FFACFF12C843FEA9FB88" authorityName="Grandjean" authorityYear="1954" box="[151,309,1087,1112]" class="Arachnida" family="Tegoribatidae" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="family">Tegoribatidae</taxonomicName>
:
<taxonomicName id="D2AB0FEDFF91FFACFEBAC843FE5AFB88" authorityName="Behan-Pelletier &amp; Walter" authorityYear="2013" box="[319,454,1087,1112]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF91FFACFEBAC843FE5AFB88" box="[319,454,1087,1112]" italics="true" pageId="15" pageNumber="166">Tectoribates</emphasis>
</taxonomicName>
); only
<taxonomicName id="D2AB0FEDFF91FFACFD9CC843FD06FB88" authorityName="Ewing" authorityYear="1917" box="[537,666,1087,1112]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF91FFACFD9CC843FD06FB88" box="[537,666,1087,1112]" italics="true" pageId="15" pageNumber="166">Tegoribates</emphasis>
</taxonomicName>
includes species with either porose areas, saccules or tubules.
</paragraph>
<paragraph id="1514746EFF91FFACFF42C818FCE6FAE0" blockId="15.[151,1437,151,1904]" pageId="15" pageNumber="166">
5. A distinct apomorphy of all known species of
<taxonomicName id="D2AB0FEDFF91FFACFD6AC818FCEEFBAD" authorityName="Ewing" authorityYear="1917" box="[751,882,1124,1149]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF91FFACFD6AC818FCEEFBAD" box="[751,882,1124,1149]" italics="true" pageId="15" pageNumber="166">Tegoribates</emphasis>
</taxonomicName>
is femur I distinctly bent to an obtuse angle near the level of seta
<emphasis id="27DFA87CFF91FFACFEDDC8FBFEFAFB70" box="[344,358,1159,1184]" italics="true" pageId="15" pageNumber="166">d</emphasis>
. Femur I is also relatively longer than in other genera of
<taxonomicName id="D2AB0FEDFF91FFACFB87C8FBFB00FB70" authorityName="Grandjean" authorityYear="1954" box="[1026,1180,1159,1184]" class="Arachnida" family="Tegoribatidae" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="family">Tegoribatidae</taxonomicName>
where legs have been studied; e.g., compare
<figureCitation id="8D9068EBFF91FFACFE1CC8D0FD98FB14" box="[409,516,1196,1221]" captionStart="FIGURE 12" captionStartId="22.[151,250,1970,1992]" captionTargetBox="[236,1371,232,1949]" captionTargetId="figure@22.[204,1382,193,1949]" captionTargetPageId="22" captionText="FIGURE 12 A – C. Tegoribates americanus Hammer, adult female, legs, all antiaxial aspect (trochanter not shown in A, B): A, leg I, with arrow to bend in femur; B, leg II (femur to tibia), C, leg III. Scale bar = 20 µm." httpUri="https://zenodo.org/record/1018655/files/figure.png" pageId="15" pageNumber="166">Figs 12A</figureCitation>
,
<figureCitation id="8D9068EBFF91FFACFD97C8D0FDA3FB14" box="[530,575,1196,1220]" captionStart="FIGURE 22" captionStartId="34.[151,250,1827,1849]" captionTargetBox="[222,1338,193,1774]" captionTargetId="figure@34.[222,1338,193,1794]" captionTargetPageId="34" captionText="FIGURE 22. Differential interference contrast microscope images of adults; A – D, Tegoribates walteri sp. nov., A, part of prodorsum (4 layers) showing lamella, bothridium, bothridial seta, interlamellar seta, with arrow to fused dorsophragmata; B, hinged pteromorph (3 layers); C, coxisternal region showing custodium (arrow) and pedotectum II (3 layers); D, porose area Aa and surrounding muscle sigillae (5 layers). E, Tegoribates subniger Ewing, lateral of podosoma, showing elongated, bent femur I (indicated by arrow); F, G, Protectoribates occidentalis sp. nov., detail of preanal organ; H, Hypozetes laysanensis Aoki, detail of preanal organ; I, Tectoribates borealis Behan-Pelletier &amp; Walter, detail of preanal organ. Scale bars: A, B, E = 25 µm; C, D = 20 µm." httpUri="https://zenodo.org/record/1018677/files/figure.png" pageId="15" pageNumber="166">22E</figureCitation>
with
<figureCitation id="8D9068EBFF91FFACFD07C8D0FD46FB14" box="[642,730,1196,1221]" captionStart="FIGURE 4" captionStartId="8.[151,250,1974,1996]" captionTargetBox="[313,1242,203,1931]" captionTargetId="figure@8.[236,1331,193,1943]" captionTargetPageId="8" captionText="FIGURE 4. Protectoribates occidentalis sp. nov., adult female, A, leg I, B, leg II, C, leg IV, all antiaxial aspect. Legs I and II, trochanter not illustrated. Scale bar = 20 µm." httpUri="https://zenodo.org/record/1018637/files/figure.png" pageId="15" pageNumber="166">Fig. 4A</figureCitation>
herein, and
<figureCitation id="8D9068EBFF91FFACFCECC8D0FC5CFB14" box="[873,960,1196,1221]" captionStart="FIGURE 7" captionStartId="11.[151,250,1861,1883]" captionTargetBox="[207,1359,193,1809]" captionTargetId="figure@11.[207,1359,193,1828]" captionTargetPageId="11" captionText="FIGURE 7. Protectoribates occidentalis sp. nov., differential interference contrast microscope images, A – E adults; A, bothridium and bothridial seta (5 layers), with arrow to nerve canal of seta le on lamella; insert of bothridium showing scales; B, custodium and discidium (3 layers), with arrow to broad, rounded anterior edge of custodium; C, porose organ Ah, lateral view, with arrow to surrounding cerotegument (3 layers); D, dissected specimen with Ah in direct view (2 layers); E, genu, tibia and tarsus II, with white arrow to dentate carina on tibia (2 layers) and narrow black arrow to seta pv ”; F, protonymph (2 layers), dorsal view. Scale bar: A = 25 µm; B = 10 µm; D = 7.5 µm; F = 50 µm." httpUri="https://zenodo.org/record/1018645/files/figure.png" pageId="15" pageNumber="166">Fig. 7A</figureCitation>
(
<bibRefCitation id="713A099FFF91FFACFC55C8D0FAAAFB14" author="Behan-Pelletier, V. M. &amp; Walter, D. E." box="[976,1334,1196,1221]" journalOrPublisher="Zootaxa" pageId="15" pageNumber="166" pagination="459 - 489" part="3741" publicationUrl="https://doi.org/10.11646/zootaxa.3741.4.2" refString="Behan-Pelletier, V. M. &amp; Walter, D. E. (2013) Phylogenetic relationships of Tectoribates: nymphal characters of new North American species place the genus in Tegoribatidae (Acari, Oribatida). Zootaxa, 3741 (4), 459 - 489. https: // doi. org / 10.11646 / zootaxa. 3741.4.2" title="Phylogenetic relationships of Tectoribates: nymphal characters of new North American species place the genus in Tegoribatidae (Acari, Oribatida)" type="journal article" year="2013">Behan-Pelletier &amp; Walter 2013</bibRefCitation>
),
<figureCitation id="8D9068EBFF91FFACFACEC8D0FA00FB14" box="[1355,1436,1196,1221]" captionStart="FIGURE 21" captionStartId="33.[151,250,1886,1908]" captionTargetBox="[225,1362,193,1858]" captionTargetId="figure@33.[225,1362,193,1865]" captionTargetPageId="33" captionText="FIGURE 21. Tegoribates walteri sp. nov., scanning electron microscope images of adult female from California, A. habitus dorsal, with arrow to spine-like seta l ” of genu II; B, habitus ventral, with arrow to ventral depression; C, frontal aspect, with arrows to spine-like setae l ” of genua I and II; D, anterolateral of prodorsum and part of notogaster, with arrow to spine-like seta l ” of genu II; E, ventrofrontal aspect, with arrow to taenidium. Scale bars: A, B = 50 µm; C-E = 25 µm." httpUri="https://zenodo.org/record/1018675/files/figure.png" pageId="15" pageNumber="166">Fig. 21</figureCitation>
(
<bibRefCitation id="713A099FFF91FFACFF1BC8B3FE45FB38" author="Ermilov, S. G. &amp; Anichkin, A. E." box="[158,473,1231,1256]" journalOrPublisher="Systematic &amp; Applied Acarology" pageId="15" pageNumber="166" pagination="137 - 144" part="18" publicationUrl="https://doi.org/10.11158/saa.18.2.6" refString="Ermilov, S. G. &amp; Anichkin, A. E. (2013) A new species of Plakoribates (Acari: Oribatida: Achipteriidae) from Vietnam. Systematic &amp; Applied Acarology, 18 (2), 137 - 144. https: // doi. org / 10.11158 / saa. 18.2.6" title="A new species of Plakoribates (Acari: Oribatida: Achipteriidae) from Vietnam" type="journal article" year="2013">Ermilov &amp; Anichkin 2013</bibRefCitation>
),
<figureCitation id="8D9068EBFF91FFACFE77C8B3FDD0FB38" box="[498,588,1231,1256]" captionStart="FIGURES 2 – 3" captionStartId="7.[151,264,1946,1968]" captionTargetBox="[335,1280,208,1896]" captionTargetId="figure@7.[253,1333,193,1924]" captionTargetPageId="7" captionText="FIGURES 2 – 3. Protectoribates occidentalis sp. nov., adult female, legs not illustrated; 2, ventral aspect; 3, lateral aspect, only alveolus of seta ex illustrated, gnathosoma not illustrated. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018635/files/figure.png" pageId="15" pageNumber="166">Fig. 3A</figureCitation>
(
<bibRefCitation id="713A099FFF91FFACFDDAC8B3FCEAFB38" author="Ermilov, S. G. &amp; Minor, M. A." box="[607,886,1231,1256]" journalOrPublisher="Acarologia" pageId="15" pageNumber="166" pagination="285 - 295" part="55" publicationUrl="https://doi.org/10.1051/acarologia/20152171" refString="Ermilov, S. G. &amp; Minor, M. A. (2015) Two new species of Neophysobates (Acari, Oribatida, Tegoribatidae) from New Zealand. Acarologia, 55 (3), 285 - 295. https: // doi. org / 10.1051 / acarologia / 20152171" title="Two new species of Neophysobates (Acari, Oribatida, Tegoribatidae) from New Zealand" type="journal article" year="2015">Ermilov &amp; Minor 2015</bibRefCitation>
) and
<figureCitation id="8D9068EBFF91FFACFC38C8B3FB84FB38" box="[957,1048,1231,1256]" captionStart="FIGURES 2 – 3" captionStartId="7.[151,264,1946,1968]" captionTargetBox="[335,1280,208,1896]" captionTargetId="figure@7.[253,1333,193,1924]" captionTargetPageId="7" captionText="FIGURES 2 – 3. Protectoribates occidentalis sp. nov., adult female, legs not illustrated; 2, ventral aspect; 3, lateral aspect, only alveolus of seta ex illustrated, gnathosoma not illustrated. Scale bar = 50 µm." httpUri="https://zenodo.org/record/1018635/files/figure.png" pageId="15" pageNumber="166">Fig. 2A</figureCitation>
(
<bibRefCitation id="713A099FFF91FFACFBAEC8B3FACFFB38" author="Fredes, N. A. &amp; Martinez, P. A." box="[1067,1363,1231,1256]" journalOrPublisher="Zootaxa" pageId="15" pageNumber="166" pagination="191 - 200" part="4098" publicationUrl="https://doi.org/10.11646/zootaxa.4098.1.10" refString="Fredes, N. A. &amp; Martinez, P. A. (2016) Revision of Physobates (Acari, Oribatida, Tegoribatidae) and redescription of Physobates spinipes Hammer, 1962 with comments on tegoribatid genera. Zootaxa, 4098 (1), 191 - 200. https: // doi. org / 10.11646 / zootaxa. 4098.1.10" title="Revision of Physobates (Acari, Oribatida, Tegoribatidae) and redescription of Physobates spinipes Hammer, 1962 with comments on tegoribatid genera" type="journal article" year="2016">Fredes &amp; Martínez 2016</bibRefCitation>
). This elongated and bent femur I could possibly be a modification to accommodate the depth of the prosoma of
<taxonomicName id="D2AB0FEDFF91FFACFF12C96BFE86FAE0" authorityName="Ewing" authorityYear="1917" box="[151,282,1303,1328]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF91FFACFF12C96BFE86FAE0" box="[151,282,1303,1328]" italics="true" pageId="15" pageNumber="166">Tegoribates</emphasis>
</taxonomicName>
, and the associated length of pedotectum I (
<figureCitation id="8D9068EBFF91FFACFC8DC96BFCF2FAE0" box="[776,878,1303,1328]" captionStart="FIGURE 22" captionStartId="34.[151,250,1827,1849]" captionTargetBox="[222,1338,193,1774]" captionTargetId="figure@34.[222,1338,193,1794]" captionTargetPageId="34" captionText="FIGURE 22. Differential interference contrast microscope images of adults; A – D, Tegoribates walteri sp. nov., A, part of prodorsum (4 layers) showing lamella, bothridium, bothridial seta, interlamellar seta, with arrow to fused dorsophragmata; B, hinged pteromorph (3 layers); C, coxisternal region showing custodium (arrow) and pedotectum II (3 layers); D, porose area Aa and surrounding muscle sigillae (5 layers). E, Tegoribates subniger Ewing, lateral of podosoma, showing elongated, bent femur I (indicated by arrow); F, G, Protectoribates occidentalis sp. nov., detail of preanal organ; H, Hypozetes laysanensis Aoki, detail of preanal organ; I, Tectoribates borealis Behan-Pelletier &amp; Walter, detail of preanal organ. Scale bars: A, B, E = 25 µm; C, D = 20 µm." httpUri="https://zenodo.org/record/1018677/files/figure.png" pageId="15" pageNumber="166">Fig. 22E</figureCitation>
).
</paragraph>
<paragraph id="1514746EFF91FFACFF42C940FD1DF9A4" blockId="15.[151,1437,151,1904]" pageId="15" pageNumber="166">
6. The preanal organ of
<taxonomicName id="D2AB0FEDFF91FFACFE5EC940FDC0FA85" authorityName="Ewing" authorityYear="1917" box="[475,604,1340,1365]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF91FFACFE5EC940FDC0FA85" box="[475,604,1340,1365]" italics="true" pageId="15" pageNumber="166">Tegoribates</emphasis>
</taxonomicName>
(
<figureCitation id="8D9068EBFF91FFACFDEAC940FD5AFA84" box="[623,710,1340,1365]" captionStart="FIGURE 8" captionStartId="17.[151,250,1857,1879]" captionTargetBox="[219,1314,198,1804]" captionTargetId="figure@17.[219,1314,193,1804]" captionTargetPageId="17" captionText="FIGURE 8 A – H. Tegoribates subniger Ewing, differential interference contrast microscope images of adult. A, holotype specimen, Type No. 2009427, housed in USNM; B, lateral of prodorsum with arrow to taenidium; C, preanal organ indicated by arrow; D, region of tubule T 1, h 3 and gla, with small arrows to tubule and large arrow to its opening (3 layers); E, lamella (5 layers), with arrow to nerve canal for seta le, lateral of podosoma displaced so Ah visible, black arrow to fused dorsophragmata; F, ventral region showing discidium (dis) and circumpedal carina (cc), with arrow to minute custodium (3 layers); G, pedipalp (5 layers), with arrow to solenidion ω. Scale bars: A = 100 µm; C, D, F = 25 µm; E = 40 µm; G = 30 µm." httpUri="https://zenodo.org/record/1018647/files/figure.png" pageId="15" pageNumber="166">Fig. 8C</figureCitation>
) is most similar to that of
<taxonomicName id="D2AB0FEDFF91FFACFC7CC940FEAEFAA8" authority="Grandjean (1955)" authorityName="Grandjean" authorityYear="1955" class="Arachnida" family="Ameronothridae" genus="Podacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="auberti">
<emphasis id="27DFA87CFF91FFACFC7CC940FB57FA85" box="[1017,1227,1340,1365]" italics="true" pageId="15" pageNumber="166">Podacarus auberti</emphasis>
<bibRefCitation id="713A099FFF91FFACFB51C940FA00FA85" author="Grandjean, F." box="[1236,1436,1340,1365]" journalOrPublisher="Podacarus Auberti (Oribate). Memoires du Museum nationale d'Histoire Naturelle, New Series, Series A, Zoologie" pageId="15" pageNumber="166" pagination="109 - 150" part="8" refString="Grandjean, F. (1955) Sur un Acarien des iles Kerguelen. Podacarus Auberti (Oribate). Memoires du Museum nationale d'Histoire Naturelle, New Series, Series A, Zoologie, 8, 109 - 150." title="Sur un Acarien des iles Kerguelen" type="journal article" year="1955">Grandjean (1955)</bibRefCitation>
(Podacaridae)
</taxonomicName>
with a large, wide cup, the anterior face of which serves for attachment of muscles for the genital aperture, and protuberance
<emphasis id="27DFA87CFF91FFACFE52C9F8FE6FFA4D" box="[471,499,1412,1437]" italics="true" pageId="15" pageNumber="166">ob</emphasis>
. It differs in that the struts for
<taxonomicName id="D2AB0FEDFF91FFACFCF5C9F9FC7DFA4D" authorityName="Grandjean" authorityYear="1955" box="[880,993,1412,1437]" class="Arachnida" family="Ameronothridae" genus="Podacarus" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="auberti">
<emphasis id="27DFA87CFF91FFACFCF5C9F9FC7DFA4D" box="[880,993,1412,1437]" italics="true" pageId="15" pageNumber="166">P. auberti</emphasis>
</taxonomicName>
are parallel; those of
<taxonomicName id="D2AB0FEDFF91FFACFB6EC9F8FAF2FA4D" authorityName="Ewing" authorityYear="1917" box="[1259,1390,1412,1437]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF91FFACFB6EC9F8FAF2FA4D" box="[1259,1390,1412,1437]" italics="true" pageId="15" pageNumber="166">Tegoribates</emphasis>
</taxonomicName>
are directed laterally at an obtuse angle. The preanal organ has not been studied in much detail in poronotic
<taxonomicName id="D2AB0FEDFF91FFACFF12C9B0FEB2FA35" authorityName="Hull" authorityYear="1918" box="[151,302,1484,1509]" class="Arachnida" infraOrder="Brachypylina" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="infraOrder">Brachypylina</taxonomicName>
other than by Grandjean (especially
<bibRefCitation id="713A099FFF91FFACFD57C9B0FC17FA35" author="Grandjean, F." box="[722,907,1484,1509]" journalOrPublisher="Acarologia" pageId="15" pageNumber="166" pagination="317 - 349" part="11" refString="Grandjean, F. (1969) Observation sur les muscles de fermeture des volets anaux et genitaux et sur la structure progenitale chez les Oribates superieures adultes. Acarologia, 11, 317 - 349." title="Observation sur les muscles de fermeture des volets anaux et genitaux et sur la structure progenitale chez les Oribates superieures adultes" type="journal article" year="1969">Grandjean 1969</bibRefCitation>
). This character seems to be highly variable in
<taxonomicName id="D2AB0FEDFF91FFACFF12C993FEA8F9D8" authorityName="Grandjean" authorityYear="1954" box="[151,308,1519,1544]" class="Arachnida" family="Tegoribatidae" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="family">Tegoribatidae</taxonomicName>
, with that of
<taxonomicName id="D2AB0FEDFF91FFACFE4FC993FDEBF9D8" authority="Behan-Pelletier, 2017" authorityName="Behan-Pelletier" authorityYear="2017" box="[458,631,1519,1544]" class="Arachnida" family="Tegoribatidae" genus="Protectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="194" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="27DFA87CFF91FFACFE4FC993FDEBF9D8" box="[458,631,1519,1544]" italics="true" pageId="15" pageNumber="166">Protectoribates</emphasis>
</taxonomicName>
(
<figureCitation id="8D9068EBFF91FFACFD0DC993FC8AF9D8" box="[648,790,1519,1544]" captionStart="FIGURE 22" captionStartId="34.[151,250,1827,1849]" captionTargetBox="[222,1338,193,1774]" captionTargetId="figure@34.[222,1338,193,1794]" captionTargetPageId="34" captionText="FIGURE 22. Differential interference contrast microscope images of adults; A – D, Tegoribates walteri sp. nov., A, part of prodorsum (4 layers) showing lamella, bothridium, bothridial seta, interlamellar seta, with arrow to fused dorsophragmata; B, hinged pteromorph (3 layers); C, coxisternal region showing custodium (arrow) and pedotectum II (3 layers); D, porose area Aa and surrounding muscle sigillae (5 layers). E, Tegoribates subniger Ewing, lateral of podosoma, showing elongated, bent femur I (indicated by arrow); F, G, Protectoribates occidentalis sp. nov., detail of preanal organ; H, Hypozetes laysanensis Aoki, detail of preanal organ; I, Tectoribates borealis Behan-Pelletier &amp; Walter, detail of preanal organ. Scale bars: A, B, E = 25 µm; C, D = 20 µm." httpUri="https://zenodo.org/record/1018677/files/figure.png" pageId="15" pageNumber="166">Figs. 22F, G</figureCitation>
) different from any described to date. In contrast, that of
<taxonomicName id="D2AB0FEDFF91FFACFF12CA68FE82F9FD" authorityName="Behan-Pelletier &amp; Walter" authorityYear="2013" box="[151,286,1556,1581]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF91FFACFF12CA68FE82F9FD" box="[151,286,1556,1581]" italics="true" pageId="15" pageNumber="166">Tectoribates</emphasis>
</taxonomicName>
(
<figureCitation id="8D9068EBFF91FFACFEB5CA68FE10F9FC" box="[304,396,1556,1581]" captionStart="FIGURE 22" captionStartId="34.[151,250,1827,1849]" captionTargetBox="[222,1338,193,1774]" captionTargetId="figure@34.[222,1338,193,1794]" captionTargetPageId="34" captionText="FIGURE 22. Differential interference contrast microscope images of adults; A – D, Tegoribates walteri sp. nov., A, part of prodorsum (4 layers) showing lamella, bothridium, bothridial seta, interlamellar seta, with arrow to fused dorsophragmata; B, hinged pteromorph (3 layers); C, coxisternal region showing custodium (arrow) and pedotectum II (3 layers); D, porose area Aa and surrounding muscle sigillae (5 layers). E, Tegoribates subniger Ewing, lateral of podosoma, showing elongated, bent femur I (indicated by arrow); F, G, Protectoribates occidentalis sp. nov., detail of preanal organ; H, Hypozetes laysanensis Aoki, detail of preanal organ; I, Tectoribates borealis Behan-Pelletier &amp; Walter, detail of preanal organ. Scale bars: A, B, E = 25 µm; C, D = 20 µm." httpUri="https://zenodo.org/record/1018677/files/figure.png" pageId="15" pageNumber="166">
Fig.
<date id="611552AEFF91FFACFEE3CA68FE10F9FC" box="[358,396,1556,1580]" pageId="15" pageNumber="166" value="1922-01">22I</date>
</figureCitation>
) is similar to that of described Ceratozetoidea (e.g.,
<emphasis id="27DFA87CFF91FFACFC67CA69FBDCF9FC" box="[994,1088,1557,1580]" italics="true" pageId="15" pageNumber="166">
<taxonomicName id="D2AB0FEDFF91FFACFC67CA69FBA6F9FC" authorityName="Berlese" authorityYear="1908" box="[994,1082,1557,1580]" class="Arachnida" family="Ceratozetidae" genus="Euzetes" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">Euzetes</taxonomicName>
;
</emphasis>
<bibRefCitation id="713A099FFF91FFACFBCCCA68FA9DF9FD" author="Grandjean, F." box="[1097,1281,1556,1581]" journalOrPublisher="Acarologia" pageId="15" pageNumber="166" pagination="317 - 349" part="11" refString="Grandjean, F. (1969) Observation sur les muscles de fermeture des volets anaux et genitaux et sur la structure progenitale chez les Oribates superieures adultes. Acarologia, 11, 317 - 349." title="Observation sur les muscles de fermeture des volets anaux et genitaux et sur la structure progenitale chez les Oribates superieures adultes" type="journal article" year="1969">Grandjean 1969</bibRefCitation>
). The preanal organ of
<taxonomicName id="D2AB0FEDFF91FFACFE87CA45FEE9F980" authorityName="Balogh" authorityYear="1959" box="[258,373,1593,1616]" class="Arachnida" family="Austrachipteriidae" genus="Hypozetes" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF91FFACFE87CA45FEE9F980" box="[258,373,1593,1616]" italics="true" pageId="15" pageNumber="166">Hypozetes</emphasis>
</taxonomicName>
is similar to that of
<emphasis id="27DFA87CFF91FFACFDE9CA4BFD60F980" box="[620,764,1591,1616]" italics="true" pageId="15" pageNumber="166">
<taxonomicName id="D2AB0FEDFF91FFACFDE9CA4BFD64F980" authorityName="Behan-Pelletier &amp; Walter" authorityYear="2013" box="[620,760,1591,1616]" class="Arachnida" family="Oribatellidae" genus="Tectoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">Tectoribates</taxonomicName>
,
</emphasis>
except that the cup is very wide and the struts are thick, strongly sclerotized and parallel (
<figureCitation id="8D9068EBFF91FFACFD88CA20FDE9F9A4" box="[525,629,1628,1653]" captionStart="FIGURE 22" captionStartId="34.[151,250,1827,1849]" captionTargetBox="[222,1338,193,1774]" captionTargetId="figure@34.[222,1338,193,1794]" captionTargetPageId="34" captionText="FIGURE 22. Differential interference contrast microscope images of adults; A – D, Tegoribates walteri sp. nov., A, part of prodorsum (4 layers) showing lamella, bothridium, bothridial seta, interlamellar seta, with arrow to fused dorsophragmata; B, hinged pteromorph (3 layers); C, coxisternal region showing custodium (arrow) and pedotectum II (3 layers); D, porose area Aa and surrounding muscle sigillae (5 layers). E, Tegoribates subniger Ewing, lateral of podosoma, showing elongated, bent femur I (indicated by arrow); F, G, Protectoribates occidentalis sp. nov., detail of preanal organ; H, Hypozetes laysanensis Aoki, detail of preanal organ; I, Tectoribates borealis Behan-Pelletier &amp; Walter, detail of preanal organ. Scale bars: A, B, E = 25 µm; C, D = 20 µm." httpUri="https://zenodo.org/record/1018677/files/figure.png" pageId="15" pageNumber="166">Fig. 22H</figureCitation>
),
</paragraph>
<paragraph id="1514746EFF91FFACFF42CAFCFEF3F8F8" blockId="15.[151,1437,151,1904]" pageId="15" pageNumber="166">
7.
<bibRefCitation id="713A099FFF91FFACFF63CA03FD85F948" author="Ermilov, S. G. &amp; Minor, M. A." box="[230,537,1663,1688]" journalOrPublisher="Acarologia" pageId="15" pageNumber="166" pagination="285 - 295" part="55" publicationUrl="https://doi.org/10.1051/acarologia/20152171" refString="Ermilov, S. G. &amp; Minor, M. A. (2015) Two new species of Neophysobates (Acari, Oribatida, Tegoribatidae) from New Zealand. Acarologia, 55 (3), 285 - 295. https: // doi. org / 10.1051 / acarologia / 20152171" title="Two new species of Neophysobates (Acari, Oribatida, Tegoribatidae) from New Zealand" type="journal article" year="2015">Ermilov and Minor (2015)</bibRefCitation>
disagreed with the transfer of
<taxonomicName id="D2AB0FEDFF91FFACFC04CA03FAD7F948" authority="Engelbrecht, 1986" authorityName="Engelbrecht" authorityYear="1986" box="[897,1355,1663,1688]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="montana">
<emphasis id="27DFA87CFF91FFACFC04CA03FBECF948" box="[897,1136,1663,1688]" italics="true" pageId="15" pageNumber="166">Tegoribates montana</emphasis>
<bibRefCitation id="713A099FFF91FFACFBFECA03FAD7F948" author="Engelbrecht, C. M." box="[1147,1355,1663,1688]" journalOrPublisher="Navorsinge van die Nasionale Museum, Bloemfontein" pageId="15" pageNumber="166" pagination="169 - 250" part="5" refString="Engelbrecht, C. M. (1986) New oribatid taxa and distribution records predominantly from Southern Africa (Acari: Oribatei: Oribatelloidea. Navorsinge van die Nasionale Museum, Bloemfontein, 5 (9), 169 - 250." title="New oribatid taxa and distribution records predominantly from Southern Africa (Acari: Oribatei: Oribatelloidea" type="journal article" year="1986">Engelbrecht, 1986</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="D2AB0FEDFF91FFACFA02CAFDFE39F96D" authority="Engelbrecht, 1986" authorityName="Engelbrecht" authorityYear="1986" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="nuda">
<emphasis id="27DFA87CFF91FFACFA02CAFDFF53F96D" italics="true" pageId="15" pageNumber="166">T. nuda</emphasis>
<bibRefCitation id="713A099FFF91FFACFF52CAD8FE39F96D" author="Engelbrecht, C. M." box="[215,421,1700,1725]" journalOrPublisher="Navorsinge van die Nasionale Museum, Bloemfontein" pageId="15" pageNumber="166" pagination="169 - 250" part="5" refString="Engelbrecht, C. M. (1986) New oribatid taxa and distribution records predominantly from Southern Africa (Acari: Oribatei: Oribatelloidea. Navorsinge van die Nasionale Museum, Bloemfontein, 5 (9), 169 - 250." title="New oribatid taxa and distribution records predominantly from Southern Africa (Acari: Oribatei: Oribatelloidea" type="journal article" year="1986">Engelbrecht, 1986</bibRefCitation>
</taxonomicName>
(both from
<collectingCountry id="6DBC34FEFF91FFACFDABCAD8FD5DF96D" box="[558,705,1700,1725]" name="South Africa" pageId="15" pageNumber="166">South Africa</collectingCountry>
) to
<taxonomicName id="D2AB0FEDFF91FFACFD6BCAD8FC04F96D" authorityName="Luxton" authorityYear="1987" box="[750,920,1700,1725]" class="Arachnida" family="Tegoribatidae" genus="Neophysobates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF91FFACFD6BCAD8FC04F96D" box="[750,920,1700,1725]" italics="true" pageId="15" pageNumber="166">Neophysobates</emphasis>
</taxonomicName>
by
<bibRefCitation id="713A099FFF91FFACFC46CAD8FBC6F96C" author="Subias, L. S." box="[963,1114,1700,1725]" journalOrPublisher="Graellsia" pageId="15" pageNumber="166" pagination="3 - 305" part="60" publicationUrl="https://doi.org/10.3989/graellsia.2004.v60.iExtra.218" refString="Subias, L. S. (2004) Listado sistematico, sinonimico y biogeografico de los acaros oribatidos (Acariformes, Oribatida) de mundo (1758 - 2002). Graellsia, 60, 3 - 305. https: // doi. org / 10.3989 / graellsia. 2004. v 60. iExtra. 218" title="Listado sistematico, sinonimico y biogeografico de los acaros oribatidos (Acariformes, Oribatida) de mundo (1758 - 2002)" type="journal article" year="2004">Subías (2004</bibRefCitation>
,
<bibRefCitation id="713A099FFF91FFACFBE3CAD8FB3CF96C" author="Subias, L. S." box="[1126,1184,1700,1724]" journalOrPublisher="sinonimico y biogeografico de los acaros oribatidos (Acariformes, Oribatida) del mundo (Excepto fosiles). (12 ª actualizacion) Available from" pageId="15" pageNumber="166" pagination="2017" part="30" publicationUrl="http://bba.bioucm.es/cont/docs/RO_1.pdf" refString="Subias, L. S. (2017) Listado sistematico, sinonimico y biogeografico de los acaros oribatidos (Acariformes, Oribatida) del mundo (Excepto fosiles). (12 ª actualizacion) Available from: http: // bba. bioucm. es / cont / docs / RO _ 1. pdf (accessed 30 April 2017)" title="Listado sistematico" type="journal article" year="2017">2017</bibRefCitation>
) and these species are retained in
<taxonomicName id="D2AB0FEDFF91FFACFE9FCABBFE01F930" authorityName="Ewing" authorityYear="1917" box="[282,413,1735,1760]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF91FFACFE9FCABBFE01F930" box="[282,413,1735,1760]" italics="true" pageId="15" pageNumber="166">Tegoribates</emphasis>
</taxonomicName>
herein. Similarly,
<bibRefCitation id="713A099FFF91FFACFDF0CABBFC29F930" author="Fredes, N. A. &amp; Martinez, P. A." box="[629,949,1735,1760]" journalOrPublisher="Zootaxa" pageId="15" pageNumber="166" pagination="191 - 200" part="4098" publicationUrl="https://doi.org/10.11646/zootaxa.4098.1.10" refString="Fredes, N. A. &amp; Martinez, P. A. (2016) Revision of Physobates (Acari, Oribatida, Tegoribatidae) and redescription of Physobates spinipes Hammer, 1962 with comments on tegoribatid genera. Zootaxa, 4098 (1), 191 - 200. https: // doi. org / 10.11646 / zootaxa. 4098.1.10" title="Revision of Physobates (Acari, Oribatida, Tegoribatidae) and redescription of Physobates spinipes Hammer, 1962 with comments on tegoribatid genera" type="journal article" year="2016">Fredes and Martínez (2016)</bibRefCitation>
questioned the change in classification of
<taxonomicName id="D2AB0FEDFF91FFACFF12CA90FDF5F8D5" authority="Engelbrecht, 1986" authorityName="Engelbrecht" authorityYear="1986" box="[151,617,1772,1797]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="species" species="natalensis">
<emphasis id="27DFA87CFF91FFACFF12CA90FE08F8D5" box="[151,404,1772,1797]" italics="true" pageId="15" pageNumber="166">Tegoribates natalensis</emphasis>
<bibRefCitation id="713A099FFF91FFACFE1ECA90FDF5F8D5" author="Engelbrecht, C. M." box="[411,617,1772,1797]" journalOrPublisher="Navorsinge van die Nasionale Museum, Bloemfontein" pageId="15" pageNumber="166" pagination="169 - 250" part="5" refString="Engelbrecht, C. M. (1986) New oribatid taxa and distribution records predominantly from Southern Africa (Acari: Oribatei: Oribatelloidea. Navorsinge van die Nasionale Museum, Bloemfontein, 5 (9), 169 - 250." title="New oribatid taxa and distribution records predominantly from Southern Africa (Acari: Oribatei: Oribatelloidea" type="journal article" year="1986">Engelbrecht, 1986</bibRefCitation>
</taxonomicName>
to
<taxonomicName id="D2AB0FEDFF91FFACFD0ACA90FCDFF8D5" authorityName="Mahunka" authorityYear="1985" box="[655,835,1772,1797]" class="Arachnida" family="Tegoribatidae" genus="Paraphysobates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF91FFACFD0ACA90FCDFF8D5" box="[655,835,1772,1797]" italics="true" pageId="15" pageNumber="166">Paraphysobates</emphasis>
</taxonomicName>
by
<bibRefCitation id="713A099FFF91FFACFCEACA90FB93F8D4" author="Subias, L. S." box="[879,1039,1772,1797]" journalOrPublisher="Graellsia" pageId="15" pageNumber="166" pagination="3 - 305" part="60" publicationUrl="https://doi.org/10.3989/graellsia.2004.v60.iExtra.218" refString="Subias, L. S. (2004) Listado sistematico, sinonimico y biogeografico de los acaros oribatidos (Acariformes, Oribatida) de mundo (1758 - 2002). Graellsia, 60, 3 - 305. https: // doi. org / 10.3989 / graellsia. 2004. v 60. iExtra. 218" title="Listado sistematico, sinonimico y biogeografico de los acaros oribatidos (Acariformes, Oribatida) de mundo (1758 - 2002)" type="journal article" year="2004">Subías (2004)</bibRefCitation>
, and this species is also retained in
<taxonomicName id="D2AB0FEDFF91FFACFF12CB73FE86F8F8" authorityName="Ewing" authorityYear="1917" box="[151,282,1807,1832]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF91FFACFF12CB73FE86F8F8" box="[151,282,1807,1832]" italics="true" pageId="15" pageNumber="166">Tegoribates</emphasis>
</taxonomicName>
herein.
</paragraph>
<paragraph id="1514746EFF91FFACFF42CB48FA01F8A0" blockId="15.[151,1437,151,1904]" pageId="15" pageNumber="166">
8. A tutorium is present, but difficult to see, in
<taxonomicName id="D2AB0FEDFF91FFACFD52CB48FCC6F89D" authorityName="Ewing" authorityYear="1917" box="[727,858,1844,1869]" class="Arachnida" family="Tegoribatidae" genus="Tegoribates" kingdom="Animalia" order="Sarcoptiformes" pageId="15" pageNumber="166" phylum="Arthropoda" rank="genus">
<emphasis id="27DFA87CFF91FFACFD52CB48FCC6F89D" box="[727,858,1844,1869]" italics="true" pageId="15" pageNumber="166">Tegoribates</emphasis>
</taxonomicName>
species as it is overlain by the lamella dorsally and the pedotectum laterally. This possibly explains the absence of this character from previous descriptions of species.
</paragraph>
</subSubSection>
</treatment>
</document>
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