298 lines
41 KiB
XML
298 lines
41 KiB
XML
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<mods:title id="74333E60DAD807B5580FE97EC37D276F">Ferussina Petofiana Sp. N. (Gastropoda, Caenogastropoda, Cyclophoridae), The Oldest Representative Of Its Subfamily From The Late Cretaceous Of Romania</mods:title>
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<mods:namePart id="97C840645D885B21DC09EB0256F9A872">Tang, Jin</mods:namePart>
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<mods:namePart id="8B3304B420EEEB5F47FFFC06C84A6D3F">Jiang, Jiamei</mods:namePart>
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<mods:namePart id="8C11FBCB8CF5FB4A3984468D38CFC2A6">Pan, Hongbo</mods:namePart>
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<heading id="C6777A34FFFE38726BC6C86BFC15C544" box="[395,905,569,596]" centered="true" fontSize="11" level="2" pageId="5" pageNumber="342" reason="2">
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<taxonomicName id="5A80B6DBFFFE38726BC6C86BFCA0C543" ID-CoL="BV4LW" authority="Pall-Gergely" authorityName="Pall-Gergely" authorityYear="2023" box="[395,828,569,595]" class="Gastropoda" family="Cyclophoridae" genus="Ferussina" kingdom="Animalia" order="Architaenioglossa" pageId="5" pageNumber="342" phylum="Mollusca" rank="species" species="petofiana" status="sp. nov.">
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<emphasis id="AFF4114AFFFE38726BC6C86BFD0EC543" bold="true" box="[395,658,569,595]" pageId="5" pageNumber="342">Ferussina petofiana</emphasis>
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Páll-Gergely
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<taxonomicNameLabel id="B4C7AC31FFFE38726909C868FC15C544" box="[836,905,570,596]" pageId="5" pageNumber="342" rank="species">sp. n.</taxonomicNameLabel>
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<paragraph id="9D3FCD58FFFE38726ABCC80EFD55C58A" blockId="5.[241,1060,604,666]" pageId="5" pageNumber="342">
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https://zoobank.org/
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(
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<figureCitation id="05BBD1DDFFFE38726819C8D2FD23C58A" box="[596,703,640,666]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="6.[136,172,1500,1522]" captionStartId-1="7.[216,252,1553,1575]" captionTargetBox-0="[136,1087,189,1477]" captionTargetBox-1="[191,1109,885,1534]" captionTargetId-0="figure-17@6.[136,1087,189,1477]" captionTargetId-1="figure-279@7.[191,1109,877,1534]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="Fig. 2. Holotype and only specimen of Ferussina petofiana Páll-Gergely, sp. n.: A–D, F = various views of the shell; E = aperture; G: protoconch and first teleoconch whorls; H = Arrows" captionText-1="Fig. 3. Artistic reconstruction of Ferussina petofiana Páll-Gergely, sp. n. (artwork by Márton Zsoldos)" figureDoi-0="http://doi.org/10.5281/zenodo.10272172" figureDoi-1="http://doi.org/10.5281/zenodo.10272176" httpUri-0="https://zenodo.org/record/10272172/files/figure.png" httpUri-1="https://zenodo.org/record/10272176/files/figure.png" pageId="5" pageNumber="342">Figs 2–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="D59A9ED3FFFE387F6AA7C8EAFD14C68D" lastPageId="8" lastPageNumber="345" pageId="5" pageNumber="342" type="materials_examined">
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<paragraph id="9D3FCD58FFFE38726AA7C8EAFE7FC434" blockId="5.[174,1127,695,919]" pageId="5" pageNumber="342">
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Type material:
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<materialsCitation id="2DE8C705FFFE38726BC1C8E5FE7FC434" collectionCode="III" country="Romania" county="The" location="Faculty of Geology" municipality="University" pageId="5" pageNumber="342" specimenCount="1" stateProvince="Bucuresti" typeStatus="holotype">
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<typeStatus id="423B73FAFFFE38726BC1C8E5FE6CC5DD" box="[396,496,695,717]" pageId="5" pageNumber="342" type="holotype">Holotype</typeStatus>
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(
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<figureCitation id="05BBD1DDFFFE38726BB3C8E5FDA0C5DD" box="[510,572,695,717]" captionStart="Fig" captionStartId="6.[136,172,1500,1522]" captionTargetBox="[136,1087,189,1477]" captionTargetId="figure-17@6.[136,1087,189,1477]" captionTargetPageId="6" captionText="Fig. 2. Holotype and only specimen of Ferussina petofiana Páll-Gergely, sp. n.: A–D, F = various views of the shell; E = aperture; G: protoconch and first teleoconch whorls; H = Arrows" figureDoi="http://doi.org/10.5281/zenodo.10272172" httpUri="https://zenodo.org/record/10272172/files/figure.png" pageId="5" pageNumber="342">Fig. 2</figureCitation>
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). LPB
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<collectionCode id="FB91559DFFFE387268CCC8EAFD05C5DD" box="[641,665,696,717]" pageId="5" pageNumber="342">III</collectionCode>
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g 10021, in the collections of the
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<location id="985F9B83FFFE387269B1C8E5FE94C5FA" LSID="urn:lsid:plazi:treatment:15297C4EFFFE387F6BC6C86BFD14C68D:985F9B83FFFE387269B1C8E5FE94C5FA" country="Romania" county="The" municipality="University" name="Faculty of Geology" pageId="5" pageNumber="342" stateProvince="Bucuresti">Faculty of Geology</location>
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and
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<location id="985F9B83FFFE38726B72C886FE20C5FA" LSID="urn:lsid:plazi:treatment:15297C4EFFFE387F6BC6C86BFD14C68D:985F9B83FFFE38726B72C886FE20C5FA" box="[319,444,724,746]" country="Romania" county="The" municipality="University" name="Geophysics" pageId="5" pageNumber="342" stateProvince="Bucuresti">Geophysics</location>
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,
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<collectingMunicipality id="7D5B5722FFFE38726B85C886FDA4C5FA" box="[456,568,724,746]" pageId="5" pageNumber="342">University</collectingMunicipality>
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of
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<collectingRegion id="5F4403BAFFFE38726811C886FD5BC5FA" box="[604,711,724,746]" country="Romania" name="Bucuresti" pageId="5" pageNumber="342">Bucharest</collectingRegion>
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,
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<collectingCountry id="E5978DC8FFFE3872689FC886FCA8C5FA" box="[722,820,724,746]" name="Romania" pageId="5" pageNumber="342">Romania</collectingCountry>
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.
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<collectingCounty id="745EB5D4FFFE38726972C886FCFBC5FA" box="[831,871,724,746]" pageId="5" pageNumber="342">The</collectingCounty>
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single available shell is relatively well preserved, but the dorsal side, and partly the ventral side of ca. half whorl of the body whorl is missing.
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</materialsCitation>
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</paragraph>
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<paragraph id="9D3FCD58FFFE38726AA7C979FBD5C487" blockId="5.[174,1127,695,919]" pageId="5" pageNumber="342">
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<typeStatus id="423B73FAFFFE38726AA7C979FEBCC450" box="[234,288,811,832]" pageId="5" pageNumber="342">Type</typeStatus>
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locality/stratum/age: K3 fossiliferous site, Neagului Creek, Vălioara, northwestern Hațeg Basin,
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<collectingRegion id="5F4403BAFFFE38726BDAC915FDF5C44D" box="[407,617,839,861]" country="Romania" name="Hunedoara" pageId="5" pageNumber="342">Hunedoara County</collectingRegion>
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,
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<collectingCountry id="E5978DC8FFFE38726839C915FD4FC44D" box="[628,723,839,861]" name="Romania" pageId="5" pageNumber="342">Romania</collectingCountry>
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(see
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<bibRefCitation id="F911B0A9FFFE38726946C91AFC75C44D" author="BOTFALVAI, G. & CSIKI-SAVA, Z. & KOCSIS, L. & ALBERT, G. & MAGYAR, J. & BODOR, E. R. & ULYANOV, A. & MAKADI, L." box="[779,1001,839,861]" pageId="5" pageNumber="342" pagination="104781" refId="ref5745" refString="BOTFALVAI, G., CSIKI-SAVA, Z., KOCSIS, L., ALBERT, G., MAGYAR, J., BODOR, E. R., ȚABARA, D., ULYANOV, A. & MAKADI, L. (2021): ' X' marks the spot! Sedimentological, geochemical and palaeontological investigations of Upper Cretaceous (Maastrichtian) vertebrate fossil localities from the Valioara Valley (Densus-Ciula Formation, Hațeg Basin, Romania). - Cretaceous Research 123: 104781. https: // doi. org / 10.1016 / j. cretres. 2021.104781" type="journal article" year="2021">
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BOTFALVAI
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<emphasis id="AFF4114AFFFE38726932C91AFC34C44C" box="[895,936,839,861]" italics="true" pageId="5" pageNumber="342">et al</emphasis>
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||
. 2021
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||
</bibRefCitation>
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||
). The site is located in the basal part of the Middle Member of the Densuș-Ciula Formation (
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<bibRefCitation id="F911B0A9FFFE38726E5CC936FE89C486" author="GRIGORESCU, D." pageId="5" pageNumber="342" pagination="142 - 164" refId="ref6673" refString="GRIGORESCU, D. (1992): Nonmarine Cretaceous formations of Romania. Pp. 142 - 164. In: MATTER, N. J. & CHEN, P. - J. (eds): Aspects of nonmarine Cretaceous geology. - China Ocean Press, Beijing." type="book chapter" year="1992">GRIGORESCU 1992</bibRefCitation>
|
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,
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||
<bibRefCitation id="F911B0A9FFFE38726B52C9D3FD9EC487" author="CSIKI-SAVA, Z. & VREMIR, M. & VASILE, S. & BRUSATTE, S. L. & DYKE, G. & NAISH, D. & NORELL, M. A. & TOTOIANU, R." box="[287,514,897,919]" pageId="5" pageNumber="342" pagination="662 - 698" refId="ref6138" refString="CSIKI-SAVA, Z., VREMIR, M., VASILE, S., BRUSATTE, S. L., DYKE, G., NAISH, D., NORELL, M. A. & TOTOIANU, R. (2016): The East Side Story - The Transylvanian latest Cretaceous continental vertebrate record and its implications for understanding Cretaceous-Paleogene boundary events. - Cretaceous Research 57: 662 - 698. https: // doi. org / 10.1016 / j. cretres. 2015.09.003" type="journal article" year="2016">
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CSIKI-SAVA
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<emphasis id="AFF4114AFFFE38726BD4C9D0FE5DC486" box="[409,449,897,919]" italics="true" pageId="5" pageNumber="342">et al</emphasis>
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||
. 2016
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||
</bibRefCitation>
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), estimated to have an age of early early Maastrichtian.
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</paragraph>
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<paragraph id="9D3FCD58FFFE38726AA7C9EEFBB7C4E7" blockId="5.[174,1126,956,1016]" pageId="5" pageNumber="342">
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Diagnosis: A small
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<taxonomicName id="5A80B6DBFFFE38726B98C9EFFD3DC4C6" box="[469,673,956,982]" pageId="5" pageNumber="342">
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<emphasis id="AFF4114AFFFE38726B98C9EFFDDFC4C6" box="[469,579,957,982]" italics="true" pageId="5" pageNumber="342">Ferussina</emphasis>
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species
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</taxonomicName>
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with a predominantly smooth sculpture, an oval aperture, and three sharp spiral carinae on the ventral side.
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</paragraph>
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<paragraph id="9D3FCD58FFFE38726AA7CE49FD20C2B7" blockId="5.[174,1127,1051,1476]" pageId="5" pageNumber="342">
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Description: Shell depressed with flat base, domed dorsal surface, and rounded or slightly shouldered body whorl; last ca. quarter whorl turns upright, elevating higher than apex; whorls 4.25, separated by a deep suture; protoconch (
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<figureCitation id="05BBD1DDFFFE38726960CE06FCE0C379" box="[813,892,1108,1130]" captionStart="Fig" captionStartId="6.[136,172,1500,1522]" captionTargetBox="[136,1087,189,1477]" captionTargetId="figure-17@6.[136,1087,189,1477]" captionTargetPageId="6" captionText="Fig. 2. Holotype and only specimen of Ferussina petofiana Páll-Gergely, sp. n.: A–D, F = various views of the shell; E = aperture; G: protoconch and first teleoconch whorls; H = Arrows" figureDoi="http://doi.org/10.5281/zenodo.10272172" httpUri="https://zenodo.org/record/10272172/files/figure.png" pageId="5" pageNumber="342">Fig. 2G</figureCitation>
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) consisting of ca. 1.25 whorls, smooth, protoconch-teleoconch boundary conspicuous, teleoconch predominantly smooth, with inconspicuous, irregular radial growth lines, last ca. quarter whorl near the suture finely ribbed, and this fine ribbing is also visible between the spiral carinae on the ventral side (
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<figureCitation id="05BBD1DDFFFE38726B75CE95FE15C3CD" box="[312,393,1223,1245]" captionStart="Fig" captionStartId="6.[136,172,1500,1522]" captionTargetBox="[136,1087,189,1477]" captionTargetId="figure-17@6.[136,1087,189,1477]" captionTargetPageId="6" captionText="Fig. 2. Holotype and only specimen of Ferussina petofiana Páll-Gergely, sp. n.: A–D, F = various views of the shell; E = aperture; G: protoconch and first teleoconch whorls; H = Arrows" figureDoi="http://doi.org/10.5281/zenodo.10272172" httpUri="https://zenodo.org/record/10272172/files/figure.png" pageId="5" pageNumber="342">Fig. 2H</figureCitation>
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); umbilicus blocked with sediment that could not be removed, therefore it is not clear whether it is open or closed (but very probably open); last ca. half whorl strongly enlarged in diameter, bears three elevated, main spiral carinae inside umbilicus (
|
||
<figureCitation id="05BBD1DDFFFE38726AF8CF4CFEB5C223" box="[181,297,1310,1332]" captionStart="Fig" captionStartId="6.[136,172,1500,1522]" captionTargetBox="[136,1087,189,1477]" captionTargetId="figure-17@6.[136,1087,189,1477]" captionTargetPageId="6" captionText="Fig. 2. Holotype and only specimen of Ferussina petofiana Páll-Gergely, sp. n.: A–D, F = various views of the shell; E = aperture; G: protoconch and first teleoconch whorls; H = Arrows" figureDoi="http://doi.org/10.5281/zenodo.10272172" httpUri="https://zenodo.org/record/10272172/files/figure.png" pageId="5" pageNumber="342">Fig. 2D, H</figureCitation>
|
||
); carinae continuous, run until peristome (see
|
||
<figureCitation id="05BBD1DDFFFE3872696BCF4CFCE8C223" box="[806,884,1310,1332]" captionStart="Fig" captionStartId="6.[136,172,1500,1522]" captionTargetBox="[136,1087,189,1477]" captionTargetId="figure-17@6.[136,1087,189,1477]" captionTargetPageId="6" captionText="Fig. 2. Holotype and only specimen of Ferussina petofiana Páll-Gergely, sp. n.: A–D, F = various views of the shell; E = aperture; G: protoconch and first teleoconch whorls; H = Arrows" figureDoi="http://doi.org/10.5281/zenodo.10272172" httpUri="https://zenodo.org/record/10272172/files/figure.png" pageId="5" pageNumber="342">Fig. 2E</figureCitation>
|
||
), not interrupted with radial ribs; area between the outermost and the middle carinae finely ribbed (
|
||
<figureCitation id="05BBD1DDFFFE387269B8CF69FBDBC240" box="[1013,1095,1339,1361]" captionStart="Fig" captionStartId="6.[136,172,1500,1522]" captionTargetBox="[136,1087,189,1477]" captionTargetId="figure-17@6.[136,1087,189,1477]" captionTargetPageId="6" captionText="Fig. 2. Holotype and only specimen of Ferussina petofiana Páll-Gergely, sp. n.: A–D, F = various views of the shell; E = aperture; G: protoconch and first teleoconch whorls; H = Arrows" figureDoi="http://doi.org/10.5281/zenodo.10272172" httpUri="https://zenodo.org/record/10272172/files/figure.png" pageId="5" pageNumber="342">Fig. 2H</figureCitation>
|
||
); a fourth, much lower carina is also visible; aperture oval (
|
||
<figureCitation id="05BBD1DDFFFE3872694FCF05FCD3C27D" box="[770,847,1367,1389]" captionStart="Fig" captionStartId="6.[136,172,1500,1522]" captionTargetBox="[136,1087,189,1477]" captionTargetId="figure-17@6.[136,1087,189,1477]" captionTargetPageId="6" captionText="Fig. 2. Holotype and only specimen of Ferussina petofiana Páll-Gergely, sp. n.: A–D, F = various views of the shell; E = aperture; G: protoconch and first teleoconch whorls; H = Arrows" figureDoi="http://doi.org/10.5281/zenodo.10272172" httpUri="https://zenodo.org/record/10272172/files/figure.png" pageId="5" pageNumber="342">Fig. 2E</figureCitation>
|
||
), parietal side convex (not concave due to penultimate whorl); peristome is incomplete (the adult peristome is missing), therefore its morphology remains unknown.
|
||
</paragraph>
|
||
<paragraph id="9D3FCD58FFFE38726AA7CFFCFCE7C2D3" blockId="5.[174,1127,1051,1476]" box="[234,891,1454,1476]" pageId="5" pageNumber="342">
|
||
Measurements: Shell diameter:
|
||
<quantity id="5A7860BDFFFE38726875CFFCFD08C2D3" box="[568,660,1454,1475]" metricMagnitude="-2" metricUnit="m" metricValue="1.08" pageId="5" pageNumber="342" unit="mm" value="10.8">10.8 mm</quantity>
|
||
, shell height:
|
||
<quantity id="5A7860BDFFFE3872696BCFFCFCEBC2D3" box="[806,887,1454,1475]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" pageId="5" pageNumber="342" unit="mm" value="4.4">4.4 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="9D3FCD58FFFE38726AA7CFBBFBFAC156" blockId="5.[174,1127,1513,1607]" pageId="5" pageNumber="342">
|
||
Differential diagnosis: All currently known
|
||
<taxonomicName id="5A80B6DBFFFE3872695ECFB8FC7FC113" box="[787,995,1513,1539]" pageId="5" pageNumber="342">
|
||
<emphasis id="AFF4114AFFFE3872695ECFB8FC1DC113" box="[787,897,1514,1539]" italics="true" pageId="5" pageNumber="342">Ferussina</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
are larger than
|
||
<quantity id="5A7860BDFFFE38726AA1CC58FEB9C135" box="[236,293,1546,1573]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="342" unit="cm" value="2.0">2 cm</quantity>
|
||
(
|
||
<bibRefCitation id="F911B0A9FFFE38726B78CC58FE63C134" author="GRATELOUP, J. - P. S. DE" box="[309,511,1546,1572]" pageId="5" pageNumber="342" pagination="3 - 26" refId="ref6587" refString="GRATELOUP, J. - P. S. DE (1827): Description de plusieurs especes de coquilles fossiles des environs de Dax (Landes). - Bulletin d'Histoire Naturelle de la Societe Linneenne de Bordeaux 2 (7): 3 - 26." type="journal article" year="1827">GRATELOUP 1827</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="F911B0A9FFFE38726844CC58FD23C134" author="DESHAYES, G. P." box="[521,703,1546,1572]" pageId="5" pageNumber="342" pagination="282 - 287" refId="ref6312" refString="DESHAYES, G. P. (1828): Memoire sur le Strophostome, nouveau genre de coquilles fossiles de la famille des Helices. - Annales des sciences naturelles 13: 282 - 287." type="journal article" year="1828">DESHAYES 1828</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="F911B0A9FFFE38726887CC58FCEDC134" author="LEUFROY, A." box="[714,881,1546,1572]" pageId="5" pageNumber="342" pagination="401 - 405" refId="ref7103" refString="LEUFROY, A. (1828): Memoire sur une nouvelle espece de coquille fossile du genre Ferussine (Grateloup), Strophostome (Deshayes). - Annales des Sciences naturelles 15: 401 - 405." type="journal article" year="1828">LEUFROY 1828</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="F911B0A9FFFE38726936CC58FB8EC134" author="DUMAS, E." box="[891,1042,1546,1572]" pageId="5" pageNumber="342" refId="ref6349" refString="DUMAS, E. (1876): Statistique geologique, mineralogique, metallurgique et paleontologique du departement du Gard. Deuxieme partie. Arthus Bertrand, Peyrot-Tinel, Brugueirolle. - Paris, Nimes, Alais, 735 pp." type="book" year="1876">DUMAS 1876</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="F911B0A9FFFE38726E50CC58FEDEC157" author="SANDBERGER, C. L. F." pageId="5" pageNumber="342" refId="ref8551" refString="SANDBERGER, C. L. F. (1870 - 1875): Die Land- und Susswasser-Conchylien der Vorwelt. - C. W. Kreidel, Wiesbaden, 1000 pp." type="book" year="1870">SANDBERGER 1870</bibRefCitation>
|
||
–1875,
|
||
<bibRefCitation id="F911B0A9FFFE38726BD8CC7EFDBEC156" author="SACCO, F." box="[405,546,1580,1606]" pageId="5" pageNumber="342" pagination="427 - 476" refId="ref8444" refString="SACCO, F. (1886): Nuove species terziarie di molluschi terrestri e d'aqua dolce e salmastra del Piemonte. - Atti della Societa Italiana di Scienze Naturali 29: 427 - 476." type="journal article" year="1886">SACCO 1886</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="F911B0A9FFFE38726860CC7EFD54C156" author="ROMAN, F." box="[557,712,1580,1606]" pageId="5" pageNumber="342" pagination="546 - 616" refId="ref8407" refString="ROMAN, F. (1904): Contributions a l'etude des bassins lacustres de l'Eocene et de l'Oligocene du Languedoc. - Bulletin de la Societe Geologique, quatrieme serie 3: 546 - 616." type="journal article" year="1904">ROMAN 1904</bibRefCitation>
|
||
,
|
||
<collectionCode id="FB91559DFFFE3872689ECC7EFC87C156" box="[723,795,1580,1606]" pageId="5" pageNumber="342">WENZ</collectionCode>
|
||
1938–1944,
|
||
<collectingCountry id="E5978DC8FFFE387269E2CC7EFBB5C156" box="[943,1065,1580,1606]" name="El Salvador" pageId="5" pageNumber="342">SALVADOR</collectingCountry>
|
||
<emphasis id="AFF4114AFFFE38726E7DCC7FFBC3C156" box="[1072,1119,1580,1606]" italics="true" pageId="5" pageNumber="342">et al</emphasis>
|
||
.
|
||
</paragraph>
|
||
<caption id="C9FF9DD0FFFD38716AC5CF8EFBDDC11E" ID-DOI="http://doi.org/10.5281/zenodo.10272172" ID-Zenodo-Dep="10272172" httpUri="https://zenodo.org/record/10272172/files/figure.png" pageId="6" pageNumber="343" startId="6.[136,172,1500,1522]" targetBox="[136,1087,189,1477]" targetPageId="6" targetType="figure">
|
||
<paragraph id="9D3FCD58FFFD38716AC5CF8EFBDDC11E" blockId="6.[136,1089,1500,1580]" pageId="6" pageNumber="343">
|
||
<emphasis id="AFF4114AFFFD38716AC5CF8EFF54C2E1" bold="true" box="[136,200,1500,1522]" pageId="6" pageNumber="343">Fig. 2.</emphasis>
|
||
Holotype and only specimen of
|
||
<taxonomicName id="5A80B6DBFFFD38716851CF8FFCF8C2E2" authority="Pall-Gergely" authorityName="Pall-Gergely" authorityYear="2023" box="[540,868,1500,1522]" class="Gastropoda" family="Cyclophoridae" genus="Ferussina" kingdom="Animalia" order="Architaenioglossa" pageId="6" pageNumber="343" phylum="Mollusca" rank="species" species="petofiana" status="sp. nov.">
|
||
<emphasis id="AFF4114AFFFD38716851CF8FFD44C2E1" box="[540,728,1500,1522]" italics="true" pageId="6" pageNumber="343">Ferussina petofiana</emphasis>
|
||
Páll-Gergely
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicNameLabel id="B4C7AC31FFFD38716920CF8FFC3BC2E2" box="[877,935,1501,1522]" pageId="6" pageNumber="343" rank="species">sp. n.</taxonomicNameLabel>
|
||
: A–D, F = various views of the shell; E = aperture; G: protoconch and first teleoconch whorls; H = Arrows
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="9D3FCD58FFFD38716B1BCC45FCEFC13C" blockId="6.[136,1089,1500,1580]" box="[342,883,1558,1580]" pageId="6" pageNumber="343">on Fig. E indicate the positions of the spiral carinae</paragraph>
|
||
<paragraph id="9D3FCD58FFFC38706AE3CAEFFCD0C6B1" blockId="7.[174,1127,189,653]" pageId="7" pageNumber="344">
|
||
2016,
|
||
<bibRefCitation id="F911B0A9FFFC38706AB9CAEFFD1FC7C8" author="PALL-GERGELY, B. & NEUBAUER, T. A." box="[244,643,189,217]" pageId="7" pageNumber="344" pagination="29 - 40" refId="ref7994" refString="PALL-GERGELY, B. & NEUBAUER, T. A. (2020): The value of a single character: the Paleogene European land snail Ferussina Grateloup, 1827 is likely a cyclophorid (Gastropoda, Caenogastropoda). - ZooKeys 918: 29 - 40. https: // doi. org / 10.3897 / zookeys. 918.50135" type="journal article" year="2020">PÁLL-GERGELY & NEUBAUER 2020</bibRefCitation>
|
||
), with the exception of
|
||
<taxonomicName id="5A80B6DBFFFC387069ECCAECFF7CC7E9" baseAuthorityName="M. Braun" baseAuthorityYear="1838" class="Gastropoda" family="Cyclophoridae" genus="Ferussina" kingdom="Animalia" order="Architaenioglossa" pageId="7" pageNumber="344" phylum="Mollusca" rank="species" species="tricarinata">
|
||
<emphasis id="AFF4114AFFFC387069ECCAECFF7CC7E9" italics="true" pageId="7" pageNumber="344">Ferussina tricarinata</emphasis>
|
||
</taxonomicName>
|
||
from the Upper Oligocene in
|
||
<collectingCountry id="E5978DC8FFFC3870681DCA8DFD58C7E9" box="[592,708,223,249]" name="Germany" pageId="7" pageNumber="344">Germany</collectingCountry>
|
||
(see
|
||
<bibRefCitation id="F911B0A9FFFC3870694FCA8DFC22C7E9" author="KADOLSKY, D." box="[770,958,223,249]" pageId="7" pageNumber="344" pagination="103 - 137" refId="ref7039" refString="KADOLSKY, D. (2008): Mollusks from the Late Oligocene of Oberleichtersbach (Rhon Mountains, Germany). Part 2: Gastropoda. Neritimorpha and Caenogastropoda. - Courier Forschungsinstitut Senckenberg 260: 103 - 137." type="journal article" year="2008">KADOLSKY 2008</bibRefCitation>
|
||
), which is the only known species that can be as small as ca.
|
||
<quantity id="5A7860BDFFFC387068ACCB53FC81C60B" box="[737,797,257,283]" metricMagnitude="-2" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="344" unit="cm" value="1.0">1 cm</quantity>
|
||
. However, that species has a stronger radial sculpture, presents only a single elevated spiral carina inside the umbilicus, and it also has one carina running along the edge of the body whorl (
|
||
<bibRefCitation id="F911B0A9FFFC38706B41CB34FE57C690" author="KADOLSKY, D." box="[268,459,358,384]" pageId="7" pageNumber="344" pagination="103 - 137" refId="ref7039" refString="KADOLSKY, D. (2008): Mollusks from the Late Oligocene of Oberleichtersbach (Rhon Mountains, Germany). Part 2: Gastropoda. Neritimorpha and Caenogastropoda. - Courier Forschungsinstitut Senckenberg 260: 103 - 137." type="journal article" year="2008">KADOLSKY 2008</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="F911B0A9FFFC38706B97CB37FCE9C690" author="PALL-GERGELY, B. & NEUBAUER, T. A." box="[474,885,357,385]" pageId="7" pageNumber="344" pagination="29 - 40" refId="ref7994" refString="PALL-GERGELY, B. & NEUBAUER, T. A. (2020): The value of a single character: the Paleogene European land snail Ferussina Grateloup, 1827 is likely a cyclophorid (Gastropoda, Caenogastropoda). - ZooKeys 918: 29 - 40. https: // doi. org / 10.3897 / zookeys. 918.50135" type="journal article" year="2020">PÁLL-GERGELY & NEUBAUER 2020</bibRefCitation>
|
||
), differing in these respects from the Vălioara specimen discussed herein.
|
||
</paragraph>
|
||
<paragraph id="9D3FCD58FFFC38706AA7CBFBFBD6C59C" blockId="7.[174,1127,189,653]" pageId="7" pageNumber="344">
|
||
The species of the genus
|
||
<taxonomicName id="5A80B6DBFFFC3870687FCBFBFC25C6D3" authority="P. Fischer, 1883" authorityName="P. Fischer" authorityYear="1883" box="[562,953,425,451]" class="Gastropoda" family="Anostomopsidae" genus="Strophostomella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="344" phylum="Mollusca" rank="genus">
|
||
<emphasis id="AFF4114AFFFC3870687FCBFBFD7AC6D3" box="[562,742,425,451]" italics="true" pageId="7" pageNumber="344">Strophostomella</emphasis>
|
||
P. Fischer, 1883
|
||
</taxonomicName>
|
||
, known from Hungarian [
|
||
<taxonomicName id="5A80B6DBFFFC38706B0ECB99FDF1C6F4" authority="(Tausch, 1886)" baseAuthorityName="Tausch" baseAuthorityYear="1886" box="[323,621,458,485]" class="Gastropoda" family="Anostomopsidae" genus="Strophostomella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="344" phylum="Mollusca" rank="species" species="cretacea">
|
||
<emphasis id="AFF4114AFFFC38706B0ECB99FE25C6F4" box="[323,441,459,485]" italics="true" pageId="7" pageNumber="344">S. cretacea</emphasis>
|
||
(
|
||
<bibRefCitation id="F911B0A9FFFC38706B8BCB98FDF9C6F4" author="TAUSCH, L." box="[454,613,458,484]" pageId="7" pageNumber="344" pagination="1 - 32" refId="ref8922" refString="TAUSCH, L. (1886): Ueber die Fauna der nicht marinen Ablagerungen der oberen Kreide des Csingerthales bei Ajka im Bakony (Veszprimer Comitat, Ungarn) und uber einige Conchylien der Gosaumergel von Aigen bei Salzburg. - Abhandlungen der k. - k. geologischen Reichsanstalt 12 (1): 1 - 32, 3 pls." type="journal article" year="1886">Tausch, 1886</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="5A80B6DBFFFC3870683ACB99FC0AC6F4" authority="(Tausch, 1886)" baseAuthorityName="Tausch" baseAuthorityYear="1886" box="[631,918,458,485]" class="Gastropoda" family="Anostomopsidae" genus="Strophostomella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="344" phylum="Mollusca" rank="species" species="fragilis">
|
||
<emphasis id="AFF4114AFFFC3870683ACB99FD7DC6F4" box="[631,737,458,485]" italics="true" pageId="7" pageNumber="344">S. fragilis</emphasis>
|
||
(
|
||
<bibRefCitation id="F911B0A9FFFC387068BDCB98FC12C6F4" author="TAUSCH, L." box="[752,910,458,484]" pageId="7" pageNumber="344" pagination="1 - 32" refId="ref8922" refString="TAUSCH, L. (1886): Ueber die Fauna der nicht marinen Ablagerungen der oberen Kreide des Csingerthales bei Ajka im Bakony (Veszprimer Comitat, Ungarn) und uber einige Conchylien der Gosaumergel von Aigen bei Salzburg. - Abhandlungen der k. - k. geologischen Reichsanstalt 12 (1): 1 - 32, 3 pls." type="journal article" year="1886">Tausch, 1886</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
] and Austrian [
|
||
<taxonomicName id="5A80B6DBFFFC38706E1DCB99FE22C516" authority="(Stoliczka, 1860)" baseAuthorityName="Stoliczka" baseAuthorityYear="1860" class="Gastropoda" family="Anostomopsidae" genus="Strophostomella" kingdom="Animalia" order="Stylommatophora" pageId="7" pageNumber="344" phylum="Mollusca" rank="species" species="reussi">
|
||
<emphasis id="AFF4114AFFFC38706E1DCB99FF6CC516" italics="true" pageId="7" pageNumber="344">S. reussi</emphasis>
|
||
(
|
||
<bibRefCitation id="F911B0A9FFFC38706AB3CBBEFE2AC516" author="STOLICZKA, F." box="[254,438,492,518]" pageId="7" pageNumber="344" pagination="482 - 496" refId="ref8887" refString="STOLICZKA, F. (1860): Uber eine der Kreideformation angehorige Susswasserbildung in den nordostlichen Alpen. - Sitzungsberichte der mathematisch-naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften 38: 482 - 496." type="journal article" year="1860">Stoliczka, 1860</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
] uppermost Cretaceous deposits, possess semilunar apertures (
|
||
<bibRefCitation id="F911B0A9FFFC38706B5EC85CFE52C537" author="STOLICZKA, F." box="[275,462,525,552]" pageId="7" pageNumber="344" pagination="482 - 496" refId="ref8887" refString="STOLICZKA, F. (1860): Uber eine der Kreideformation angehorige Susswasserbildung in den nordostlichen Alpen. - Sitzungsberichte der mathematisch-naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften 38: 482 - 496." type="journal article" year="1860">STOLICZKA 1860</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="F911B0A9FFFC38706B94C85CFDE8C537" author="TAUSCH, L." box="[473,628,525,552]" pageId="7" pageNumber="344" pagination="1 - 32" refId="ref8922" refString="TAUSCH, L. (1886): Ueber die Fauna der nicht marinen Ablagerungen der oberen Kreide des Csingerthales bei Ajka im Bakony (Veszprimer Comitat, Ungarn) und uber einige Conchylien der Gosaumergel von Aigen bei Salzburg. - Abhandlungen der k. - k. geologischen Reichsanstalt 12 (1): 1 - 32, 3 pls." type="journal article" year="1886">TAUSCH 1886</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="F911B0A9FFFC38706832C85CFC0EC538" author="BANDEL, K. & RIEDEL, F." box="[639,914,526,552]" pageId="7" pageNumber="344" pagination="1 - 65" refId="ref5533" refString="BANDEL, K. & RIEDEL, F. (1994): The late Cretaceous gastropod fauna from Ajka (Bakony Mountains, Hungary): a revision. - Annalen des Naturhistorischen Museums in Wien, Serie A 96: 1 - 65." type="journal article" year="1994">BANDEL & RIEDEL 1994</bibRefCitation>
|
||
). Moreover, none of the three known
|
||
<taxonomicName id="5A80B6DBFFFC38706BD6C87DFD31C559" box="[411,685,559,585]" pageId="7" pageNumber="344">
|
||
<emphasis id="AFF4114AFFFC38706BD6C87DFDD3C559" box="[411,591,559,585]" italics="true" pageId="7" pageNumber="344">Strophostomella</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
possesses elevated spiral carinae on their ventral (=umbilical) sides, and they have much stronger radial sculpture than the new species (
|
||
<bibRefCitation id="F911B0A9FFFC38706BF1C820FDEBC59C" author="STOLICZKA, F." box="[444,631,626,652]" pageId="7" pageNumber="344" pagination="482 - 496" refId="ref8887" refString="STOLICZKA, F. (1860): Uber eine der Kreideformation angehorige Susswasserbildung in den nordostlichen Alpen. - Sitzungsberichte der mathematisch-naturwissenschaftlichen Classe der Kaiserlichen Akademie der Wissenschaften 38: 482 - 496." type="journal article" year="1860">STOLICZKA 1860</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="F911B0A9FFFC387068CEC820FC82C59C" author="TAUSCH, L." box="[643,798,626,652]" pageId="7" pageNumber="344" pagination="1 - 32" refId="ref8922" refString="TAUSCH, L. (1886): Ueber die Fauna der nicht marinen Ablagerungen der oberen Kreide des Csingerthales bei Ajka im Bakony (Veszprimer Comitat, Ungarn) und uber einige Conchylien der Gosaumergel von Aigen bei Salzburg. - Abhandlungen der k. - k. geologischen Reichsanstalt 12 (1): 1 - 32, 3 pls." type="journal article" year="1886">TAUSCH 1886</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="F911B0A9FFFC38706964C821FBA2C59C" author="BANDEL, K. & RIEDEL, F." box="[809,1086,626,653]" pageId="7" pageNumber="344" pagination="1 - 65" refId="ref5533" refString="BANDEL, K. & RIEDEL, F. (1994): The late Cretaceous gastropod fauna from Ajka (Bakony Mountains, Hungary): a revision. - Annalen des Naturhistorischen Museums in Wien, Serie A 96: 1 - 65." type="journal article" year="1994">BANDEL & RIEDEL 1994</bibRefCitation>
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).
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</paragraph>
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<paragraph id="9D3FCD58FFFC387F6AA7C8FDFD14C68D" blockId="7.[174,1127,687,855]" lastBlockId="8.[136,1089,190,414]" lastPageId="8" lastPageNumber="345" pageId="7" pageNumber="344">
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Etymology: This new species is dedicated to and named after SÁNDOR PETŐFI (1823– 1849), the National Poet of
|
||
<collectingCountry id="E5978DC8FFFC38706B86C89FFDB2C5F2" box="[459,558,717,738]" name="Hungary" pageId="7" pageNumber="344">Hungary</collectingCountry>
|
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, to commemorate the bicentenary of his birth but also his presence in the Hațeg Basin roughly 174 years before the identification and description of the new taxon reported in this contribution. Petőfi apparently visited Hațeg in
|
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<date id="E93EEB98FFFC38706E5DC954FF42C428" pageId="7" pageNumber="344" value="1849-04-14">14 April 1849</date>
|
||
(
|
||
<bibRefCitation id="F911B0A9FFFC38706AA6C970FEEAC428" author="KRISTOF, GY" box="[235,374,802,826]" pageId="7" pageNumber="344" pagination="20 - 43" refId="ref7080" refString="KRISTOF, GY. (1944): Miklos Miklos. - Irodalomtorteneti Kozlemenyek 54 (1): 20 - 43." type="journal article" year="1944">KRISTÓF 1944</bibRefCitation>
|
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), and even wrote a poem while being there (Vajdahunyadon = At Hunedoara, transl. from Hungarian in original) (
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<bibRefCitation id="F911B0A9FFFC38706839C96DFCEBC445" author="DAVID, GY & MIKO, I." box="[628,887,831,855]" pageId="7" pageNumber="344" refId="ref6229" refString="DAVID, GY. & MIKO, I. (1972): Petofi Erdelyben. - Kriterion, Bukarest, 325 pp." type="book" year="1972">DÁVID & MIKÓ 1972: 223</bibRefCitation>
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). It is worth emphasizing that lines of this poem (albeit otherwise referring to an important Medieval historical character from Transylvania, János Hunyadi) are incidentally also highly evocative of the site of discovery of
|
||
<taxonomicName id="5A80B6DBFFF3387F6B1FCAAAFE5EC61D" authorityName="Pall-Gergely" authorityYear="2023" box="[338,450,248,269]" class="Gastropoda" family="Cyclophoridae" genus="Ferussina" kingdom="Animalia" order="Architaenioglossa" pageId="8" pageNumber="345" phylum="Mollusca" rank="species" species="petofiana">
|
||
<emphasis id="AFF4114AFFF3387F6B1FCAAAFEC3C61D" box="[338,351,248,269]" italics="true" pageId="8" pageNumber="345">F</emphasis>
|
||
.
|
||
<emphasis id="AFF4114AFFF3387F6B27CAAAFE5EC61D" box="[362,450,248,269]" italics="true" pageId="8" pageNumber="345">petofiana</emphasis>
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</taxonomicName>
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||
. These allude to a setting ‘from where I’m staring at the past’ and which ‘Is hidden from the eyes of the world/… not seen by others than,/from far away, the white-capped head of/ the mountains’ elder, the ancient Retezat’ (transl. from Hungarian in original), altogether forming a remarkably fitting description of site K3 near Vălioara where the
|
||
<typeStatus id="423B73FAFFF3387F6ABACB39FEC9C691" box="[247,341,363,385]" pageId="8" pageNumber="345" type="holotype">holotype</typeStatus>
|
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specimen was excavated, close to the foothills of, and overlooked from the distance by, the Retezat Mountains (
|
||
<figureCitation id="05BBD1DDFFF3387F687DCBDAFDE7C68D" box="[560,635,392,414]" captionStart="Fig" captionStartId="2.[136,171,1059,1081]" captionTargetBox="[136,1088,350,1034]" captionTargetId="figure-59@2.[136,1088,350,1035]" captionTargetPageId="2" captionText="Fig. 1. Geological setting of the type locality of Ferussina petofiana. A = Position of the Hațeg Basin (rectangle; HB) within Romania.B = Simplified geological map of the Hațeg Basin, highlighting the distribution of the uppermost Cretaceous continental deposits (shades of green). Legend: 1 – surrounding metamorphic basement, 2 – units of the sedimentary basin infill (mostly marine), 3-5 – fossiliferous Maastrichtian continental deposits, with 3 – Sînpetru Formation, 4 – Densuș-Ciula Formation (v – volcaniclastic beds), and 5 – Sînpetru Formation-correlative units (see CSIKI-SAVA et al. 2016), 6 – Quaternary cover, 7 – sites with gastropod assemblages (a –SCHAFARZIK 1909,b –RĂDULESCU et al. 1976, c –ANTONESCU et al. 1983, d –GRIGORESCU et al. 1985, e – PANĂ et al. 2002, f – CSIKI et al. 2008, g – VASILE et al. 2011; see text for details), 8 – site K3 of Vălioara, the type locality of Ferussina petofiana (from BOTFALVAI et al. 2021). C = Synthetic lithological column near site K3 (marked by red arrow), modified from BOTFALVAI et al. (2021). Lithofacies abbreviations: BChD – braided channel deposits, mainly conglomerates, ChL – sandstone channel lag, SSp – sandstone sheet-splay, PdFP – poorly drained floodplain deposits, mainly grey-greenish silts and muds, WdFP – well-drained floodplain deposits, mainly red silts and muds. For more details, see BOTFALVAI et al. (2021). D. View of site K3 in the Pârâul Neagului ravine, located in grey-greenish fine-grained floodplain deposits (near handle of hammer). E. The Retezat Mountains overlooking the Hațeg Basin from the south (photo by Gergő Konecsni), mentioned by the Petőfi poem written during his 1849 visit in the" figureDoi="http://doi.org/10.5281/zenodo.10272170" httpUri="https://zenodo.org/record/10272170/files/figure.png" pageId="8" pageNumber="345">Fig. 1E</figureCitation>
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).
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</paragraph>
|
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</subSubSection>
|
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</treatment>
|
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</document> |