247 lines
32 KiB
XML
247 lines
32 KiB
XML
<document ID-DOI="10.11646/palaeoentomology.5.5.6" ID-ISSN="2624-2834" ID-Zenodo-Dep="7333729" ID-ZooBank="1665808E-F970-48D3-B35D-2FB684E79FA3" approvalRequired="13" approvalRequired_for_textStreams="10" approvalRequired_for_treatments="3" checkinTime="1668767059916" checkinUser="plazi" docAuthor="PARKER, JOSEPH" docDate="2022" docId="0E239B5CFFB11D3D2430B788FBD0DD48" docLanguage="en" docName="palaeoentomology.5.5.6.pdf" docOrigin="Palaeoentomology 5 (5)" docSource="http://dx.doi.org/10.11646/palaeoentomology.5.5.6" docStyle="DocumentStyle:4EE6FD780FF4521D541D0D0600705D0E.2:Palaeontomology.2018-.journal_article" docStyleId="4EE6FD780FF4521D541D0D0600705D0E" docStyleName="Palaeontomology.2018-.journal_article" docStyleVersion="2" docTitle="Yprezethinus grimaldii PARKER 2022, sp. nov." docType="treatment" docVersion="2" lastPageNumber="457" masterDocId="F21AE324FFB51D38271AB275FFDFDE1A" masterDocTitle="Transitional morphology and Afrotropical affinity of a bythinoplectine rove beetle from the early Eocene of India (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="460" masterPageNumber="452" pageNumber="456" updateTime="1668767449493" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Transitional morphology and Afrotropical affinity of a bythinoplectine rove beetle from the early Eocene of India (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>PARKER, JOSEPH</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Palaeoentomology</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-09-22</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>5</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>452</mods:start>
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<mods:end>460</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.11646/palaeoentomology.5.5.6</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/palaeoentomology.5.5.6</mods:identifier>
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<mods:identifier type="ISSN">2624-2834</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7333729</mods:identifier>
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<mods:identifier type="ZooBank">1665808E-F970-48D3-B35D-2FB684E79FA3</mods:identifier>
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</mods:mods>
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<treatment LSID="urn:lsid:plazi:treatment:0E239B5CFFB11D3D2430B788FBD0DD48" httpUri="http://treatment.plazi.org/id/0E239B5CFFB11D3D2430B788FBD0DD48" lastPageId="5" lastPageNumber="457" pageId="4" pageNumber="456">
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<subSubSection pageId="4" pageNumber="456" type="nomenclature">
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<paragraph blockId="4.[810,1451,1533,1631]" box="[810,1168,1533,1560]" pageId="4" pageNumber="456">
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<heading bold="true" box="[810,1168,1533,1560]" fontSize="10" level="2" pageId="4" pageNumber="456" reason="3">
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<emphasis bold="true" box="[810,1168,1533,1560]" pageId="4" pageNumber="456">
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<taxonomicName authority="PARKER, 2022" authorityName="PARKER" authorityYear="2022" box="[810,1072,1533,1560]" class="Insecta" family="Staphylinidae" genus="Yprezethinus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="456" phylum="Arthropoda" rank="species" species="grimaldii" status="sp. nov.">
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<emphasis bold="true" box="[810,1072,1533,1560]" italics="true" pageId="4" pageNumber="456">Yprezethinus grimaldii</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[1078,1168,1534,1560]" pageId="4" pageNumber="456" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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||
</heading>
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||
</paragraph>
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||
<paragraph blockId="4.[810,1451,1533,1631]" pageId="4" pageNumber="456">
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<uri pageId="4" pageNumber="456">
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urn:lsid:zoobank.org:act:
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<uuid pageId="4" pageNumber="456">6657B0A8-4BF3-4A3A-88DD- F943C01A9967</uuid>
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</uri>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="4" pageNumber="456" type="materials_examined">
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<paragraph blockId="4.[810,1452,1676,2092]" pageId="4" pageNumber="456">
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<materialsCitation collectionCode="AMBER" country="India" latitude="21.356667" location="Label" longLatPrecision="1" longitude="73.07553" municipality="Sex" pageId="4" pageNumber="456" specimenCount="1" stateProvince="Gujarat" typeStatus="holotype">
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<emphasis bold="true" box="[810,926,1676,1702]" pageId="4" pageNumber="456">
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<typeStatus box="[810,921,1676,1702]" pageId="4" pageNumber="456">Holotype</typeStatus>
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.
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</emphasis>
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<collectingMunicipality box="[934,976,1676,1702]" pageId="4" pageNumber="456">Sex</collectingMunicipality>
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unknown.
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<location LSID="urn:lsid:plazi:treatment:0E239B5CFFB11D3D2430B788FBD0DD48:83557C91FFB11D3C2355B4F9FB50D8BC" box="[1103,1167,1676,1702]" country="India" latitude="21.356667" longLatPrecision="1" longitude="73.07553" municipality="Sex" name="Label" pageId="4" pageNumber="456" stateProvince="Gujarat">Label</location>
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data: “
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<collectionCode box="[1247,1353,1676,1702]" pageId="4" pageNumber="456">AMBER</collectionCode>
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:
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<collectingCountry box="[1364,1448,1676,1702]" name="India" pageId="4" pageNumber="456">INDIA</collectingCountry>
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:
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<collectingRegion box="[810,894,1711,1737]" country="India" name="Gujarat" pageId="4" pageNumber="456">Gujarat</collectingRegion>
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/ Tadkeshwar lignite mine / Cambay Form. (Paleo- Eocene) /
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<geoCoordinate box="[927,1072,1748,1775]" degrees="21" direction="north" minutes="21.400" orientation="latitude" pageId="4" pageNumber="456" precision="1" value="21.356667">21°21.400′N</geoCoordinate>
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,
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<geoCoordinate box="[1086,1211,1748,1775]" degrees="73" direction="east" minutes="4.532" orientation="longitude" pageId="4" pageNumber="456" precision="1" value="73.07553">73°4.532′E</geoCoordinate>
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/ I/7-12/09 Grimaldi & Nascimbene / No. Tad-130”
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</materialsCitation>
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</paragraph>
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<paragraph blockId="4.[810,1452,1676,2092]" box="[863,1345,1818,1844]" pageId="4" pageNumber="456">
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“Tad-130: / 1
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<taxonomicName authorityName="Linnaeus" authorityYear="1758" box="[1019,1146,1818,1844]" class="Insecta" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="456" phylum="Arthropoda" rank="order">Coleoptera</taxonomicName>
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:
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<taxonomicName authorityName="Leach" authorityYear="1815" box="[1157,1325,1818,1844]" class="Insecta" family="Scydmaenidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="456" phylum="Arthropoda" rank="family">Scydmaenidae</taxonomicName>
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?”
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</paragraph>
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<paragraph blockId="4.[810,1452,1676,2092]" box="[863,1101,1853,1880]" pageId="4" pageNumber="456">
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Specimen in
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<collectionCode box="[1008,1097,1854,1880]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34925" lsid="urn:lsid:biocol.org:col:34925" name="American Museum of Natural History" pageId="4" pageNumber="456">AMNH</collectionCode>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="4" pageNumber="456" type="etymology">
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<paragraph blockId="4.[810,1452,1676,2092]" pageId="4" pageNumber="456">
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<emphasis bold="true" box="[863,998,1889,1915]" pageId="4" pageNumber="456">Etymology.</emphasis>
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The specific name honors entomologist Dr David Grimaldi—friend, mentor, colleague and collaborator—on his 65
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<superScript attach="left" box="[1080,1093,1960,1974]" fontSize="6" pageId="4" pageNumber="456">th</superScript>
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birthday, and in recognition of his wide-ranging contributions to the study of fossilized insects (including pioneering the investigation of
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<collectingCountry box="[1358,1451,2031,2057]" name="India" pageId="4" pageNumber="456">Cambay</collectingCountry>
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amber).
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</paragraph>
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</subSubSection>
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<subSubSection box="[189,468,158,185]" pageId="5" pageNumber="457" type="diagnosis">
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<paragraph blockId="5.[136,778,158,580]" box="[189,468,158,185]" pageId="5" pageNumber="457">
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<emphasis bold="true" box="[189,312,158,184]" pageId="5" pageNumber="457">Diagnosis.</emphasis>
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As for genus.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="457" type="materials_examined">
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<paragraph blockId="5.[136,778,158,580]" pageId="5" pageNumber="457">
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<emphasis bold="true" box="[189,560,194,220]" pageId="5" pageNumber="457">Type locality and horizon.</emphasis>
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<materialsCitation country="India" location="Gujarat state" pageId="5" pageNumber="457" specimenCount="1" stateProvince="Gujarat">
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<collectingCountry box="[587,680,194,220]" name="India" pageId="5" pageNumber="457">Cambay</collectingCountry>
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amber preserving
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<taxonomicName authorityName="PARKER" authorityYear="2022" box="[272,413,230,256]" class="Insecta" family="Staphylinidae" genus="Yprezethinus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="457" phylum="Arthropoda" rank="genus">
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<emphasis box="[272,413,230,256]" italics="true" pageId="5" pageNumber="457">Yprezethinus</emphasis>
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</taxonomicName>
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was collected from outcrops inside lignite mines in
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<collectingRegion box="[432,587,266,293]" country="India" name="Gujarat" pageId="5" pageNumber="457">Gujarat state</collectingRegion>
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, western
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</materialsCitation>
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<materialsCitation country="India" latitude="21.356667" location="Tadkeshwar" longLatPrecision="1" longitude="73.07553" pageId="5" pageNumber="457" specimenCount="1">
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<collectingCountry box="[713,773,266,292]" name="India" pageId="5" pageNumber="457">India</collectingCountry>
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,
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<location LSID="urn:lsid:plazi:treatment:0E239B5CFFB11D3D2430B788FBD0DD48:83557C91FFB01D3D2792B35BFED0DF52" box="[136,271,302,328]" country="India" latitude="21.356667" longLatPrecision="1" longitude="73.07553" name="Tadkeshwar" pageId="5" pageNumber="457">Tadkeshwar</location>
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(
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<geoCoordinate box="[291,437,304,331]" degrees="21" direction="north" minutes="21.400" orientation="latitude" pageId="5" pageNumber="457" precision="1" value="21.356667">21°21.400′N</geoCoordinate>
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,
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<geoCoordinate box="[453,584,304,331]" degrees="73" direction="east" minutes="4.532" orientation="longitude" pageId="5" pageNumber="457" precision="1" value="73.07553">73°4.532′E</geoCoordinate>
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). The horizon is estimated to be Early Ypresian (
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<emphasis box="[493,527,339,364]" italics="true" pageId="5" pageNumber="457">ca.</emphasis>
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54.5 Ma; see
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<bibRefCitation author="Smith, T. & Kumar, K. & Rana, R. S. & Folie, A. & Sole, F. & Noiret, C. & Steeman, T. & Sahni, A. & Rose, K. D." pageId="5" pageNumber="457" pagination="969 - 1001" refId="ref7423" refString="Smith, T., Kumar, K., Rana, R. S., Folie, A., Sole, F., Noiret, C., Steeman, T., Sahni, A. & Rose, K. D. (2016) New early Eocene vertebrate assemblage from western India reveals a mixed fauna of European and Gondwana affinities. Geoscience Frontiers, 7, 969 - 1001. https: // doi. org / 10.1016 / j. gsf. 2016.05.001" type="journal article" year="2016">
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Smith
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<emphasis italics="true" pageId="5" pageNumber="457">et al.</emphasis>
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, 2016
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</bibRefCitation>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="5" pageNumber="457" type="description">
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<paragraph blockId="5.[136,778,158,580]" pageId="5" pageNumber="457">
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<emphasis bold="true" box="[189,335,410,436]" pageId="5" pageNumber="457">Description.</emphasis>
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Body length: ~
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<quantity box="[507,594,410,437]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="5" pageNumber="457" unit="mm" value="0.8">0.8 mm</quantity>
|
||
(
|
||
<figureCitation box="[608,697,410,436]" captionStart="FIGURE 1" captionStartId="2.[136,235,1800,1824]" captionTargetBox="[152,1437,189,1777]" captionTargetId="figure-15@2.[152,1437,189,1777]" captionTargetPageId="2" captionText="FIGURE 1. Yprezethinus grimaldii Holotype AMNH Tad-130. A, Habitus with visible abdominal sternites labelled. B, Head and pronotum with segments of maxillary palpomeres numbered (MP2–MP4). Bythinoplectina-type lateral excavation, eye, apical pseudosegment of maxillary palp (APs) and mandible (Md) indicated. C, Right antenna with antennomeres numbered. Note the club formed from tightly appressed, hemisphere-shaped antennomeres 10 and 11. D, Close-up view of apically globose tubercles of maxillary palpomeres 3 and 4. E, Proximal leg articulation, showing conically projecting metacoxa (MC), short metatrochanter (MT) and femur (F). F, Right metatarsus (F) with bythinoplectine-type enlarged third tarsomere spanning distance between arrowheads, and minute first visible tarsomere." figureDoi="http://doi.org/10.5281/zenodo.7333731" httpUri="https://zenodo.org/record/7333731/files/figure.png" pageId="5" pageNumber="457">Fig. 1A</figureCitation>
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). Body largely glabrous, with smooth, shiny integument. Body color of Tad-130 dark, almost black; antennae also dark, brown-black; other appendages somewhat dark reddish brown.
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</paragraph>
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<paragraph blockId="5.[136,778,626,1876]" box="[136,197,626,652]" pageId="5" pageNumber="457">
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<emphasis box="[136,197,626,652]" italics="true" pageId="5" pageNumber="457">Head</emphasis>
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||
</paragraph>
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||
<paragraph blockId="5.[136,778,626,1876]" pageId="5" pageNumber="457">
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Head length ~
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||
<quantity box="[294,395,662,689]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="457" unit="mm" value="0.13">0.13 mm</quantity>
|
||
(
|
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<figureCitation box="[411,500,662,688]" captionStart="FIGURE 1" captionStartId="2.[136,235,1800,1824]" captionTargetBox="[152,1437,189,1777]" captionTargetId="figure-15@2.[152,1437,189,1777]" captionTargetPageId="2" captionText="FIGURE 1. Yprezethinus grimaldii Holotype AMNH Tad-130. A, Habitus with visible abdominal sternites labelled. B, Head and pronotum with segments of maxillary palpomeres numbered (MP2–MP4). Bythinoplectina-type lateral excavation, eye, apical pseudosegment of maxillary palp (APs) and mandible (Md) indicated. C, Right antenna with antennomeres numbered. Note the club formed from tightly appressed, hemisphere-shaped antennomeres 10 and 11. D, Close-up view of apically globose tubercles of maxillary palpomeres 3 and 4. E, Proximal leg articulation, showing conically projecting metacoxa (MC), short metatrochanter (MT) and femur (F). F, Right metatarsus (F) with bythinoplectine-type enlarged third tarsomere spanning distance between arrowheads, and minute first visible tarsomere." figureDoi="http://doi.org/10.5281/zenodo.7333731" httpUri="https://zenodo.org/record/7333731/files/figure.png" pageId="5" pageNumber="457">Fig. 1A</figureCitation>
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); estimated width across eyes ~
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||
<quantity box="[209,316,698,725]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="5" pageNumber="457" unit="mm" value="0.18">0.18 mm</quantity>
|
||
. Frontal rostrum medially sulcate. Head constrictedsharplybehindeyes,narrowingtoocciput.Gular region of head strongly convex to occipital constriction. Eye with approximately 10 facets. Antennomere 1 partially obscured but appears to be cylindrical-conical, and longer than wide, similar in width to antennomere 2. Antennomere 2 spherical, ~1.3× wider than antennomere 1. Antennomeres 3–9 subequal in width but varied in length and shape (
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||
<figureCitation box="[345,434,986,1012]" captionStart="FIGURE 1" captionStartId="2.[136,235,1800,1824]" captionTargetBox="[152,1437,189,1777]" captionTargetId="figure-15@2.[152,1437,189,1777]" captionTargetPageId="2" captionText="FIGURE 1. Yprezethinus grimaldii Holotype AMNH Tad-130. A, Habitus with visible abdominal sternites labelled. B, Head and pronotum with segments of maxillary palpomeres numbered (MP2–MP4). Bythinoplectina-type lateral excavation, eye, apical pseudosegment of maxillary palp (APs) and mandible (Md) indicated. C, Right antenna with antennomeres numbered. Note the club formed from tightly appressed, hemisphere-shaped antennomeres 10 and 11. D, Close-up view of apically globose tubercles of maxillary palpomeres 3 and 4. E, Proximal leg articulation, showing conically projecting metacoxa (MC), short metatrochanter (MT) and femur (F). F, Right metatarsus (F) with bythinoplectine-type enlarged third tarsomere spanning distance between arrowheads, and minute first visible tarsomere." figureDoi="http://doi.org/10.5281/zenodo.7333731" httpUri="https://zenodo.org/record/7333731/files/figure.png" pageId="5" pageNumber="457">Fig. 1C</figureCitation>
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). Antennomeres 3–6 subequal in length, twice as wide as long; antennomere 1 transverse obconical with distinct pedicel, 4–6 transverse cylindrical with concealed pedicels. Antennomeres 7–9 subequal in length, 1.5× as wide as long; antennomere 7 roundedcylindrical, 8 and 9 weakly obconical. Ovoid club formed by antennomeres 10 and 11 (
|
||
<figureCitation box="[466,554,1202,1228]" captionStart="FIGURE 1" captionStartId="2.[136,235,1800,1824]" captionTargetBox="[152,1437,189,1777]" captionTargetId="figure-15@2.[152,1437,189,1777]" captionTargetPageId="2" captionText="FIGURE 1. Yprezethinus grimaldii Holotype AMNH Tad-130. A, Habitus with visible abdominal sternites labelled. B, Head and pronotum with segments of maxillary palpomeres numbered (MP2–MP4). Bythinoplectina-type lateral excavation, eye, apical pseudosegment of maxillary palp (APs) and mandible (Md) indicated. C, Right antenna with antennomeres numbered. Note the club formed from tightly appressed, hemisphere-shaped antennomeres 10 and 11. D, Close-up view of apically globose tubercles of maxillary palpomeres 3 and 4. E, Proximal leg articulation, showing conically projecting metacoxa (MC), short metatrochanter (MT) and femur (F). F, Right metatarsus (F) with bythinoplectine-type enlarged third tarsomere spanning distance between arrowheads, and minute first visible tarsomere." figureDoi="http://doi.org/10.5281/zenodo.7333731" httpUri="https://zenodo.org/record/7333731/files/figure.png" pageId="5" pageNumber="457">Fig. 1C</figureCitation>
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||
); club at maximum width 3× wider than antennomeres 3–9. Antennomere 11 approximately 1.5× as long as 10. Segments of antennal club, especially antennomere 11, decorated in long, dense sensory setae (preceding antennal segments without discernable setation) (
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<figureCitation box="[478,566,1382,1408]" captionStart="FIGURE 1" captionStartId="2.[136,235,1800,1824]" captionTargetBox="[152,1437,189,1777]" captionTargetId="figure-15@2.[152,1437,189,1777]" captionTargetPageId="2" captionText="FIGURE 1. Yprezethinus grimaldii Holotype AMNH Tad-130. A, Habitus with visible abdominal sternites labelled. B, Head and pronotum with segments of maxillary palpomeres numbered (MP2–MP4). Bythinoplectina-type lateral excavation, eye, apical pseudosegment of maxillary palp (APs) and mandible (Md) indicated. C, Right antenna with antennomeres numbered. Note the club formed from tightly appressed, hemisphere-shaped antennomeres 10 and 11. D, Close-up view of apically globose tubercles of maxillary palpomeres 3 and 4. E, Proximal leg articulation, showing conically projecting metacoxa (MC), short metatrochanter (MT) and femur (F). F, Right metatarsus (F) with bythinoplectine-type enlarged third tarsomere spanning distance between arrowheads, and minute first visible tarsomere." figureDoi="http://doi.org/10.5281/zenodo.7333731" httpUri="https://zenodo.org/record/7333731/files/figure.png" pageId="5" pageNumber="457">Fig. 1C</figureCitation>
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). Estimated length of antenna
|
||
<quantity box="[276,387,1418,1444]" metricMagnitude="-4" metricUnit="m" metricValue="1.9" pageId="5" pageNumber="457" unit="mm" value="0.19">0.19 mm</quantity>
|
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. Maxillary palpi (
|
||
<figureCitation box="[611,704,1418,1444]" captionStart="FIGURE 1" captionStartId="2.[136,235,1800,1824]" captionTargetBox="[152,1437,189,1777]" captionTargetId="figure-15@2.[152,1437,189,1777]" captionTargetPageId="2" captionText="FIGURE 1. Yprezethinus grimaldii Holotype AMNH Tad-130. A, Habitus with visible abdominal sternites labelled. B, Head and pronotum with segments of maxillary palpomeres numbered (MP2–MP4). Bythinoplectina-type lateral excavation, eye, apical pseudosegment of maxillary palp (APs) and mandible (Md) indicated. C, Right antenna with antennomeres numbered. Note the club formed from tightly appressed, hemisphere-shaped antennomeres 10 and 11. D, Close-up view of apically globose tubercles of maxillary palpomeres 3 and 4. E, Proximal leg articulation, showing conically projecting metacoxa (MC), short metatrochanter (MT) and femur (F). F, Right metatarsus (F) with bythinoplectine-type enlarged third tarsomere spanning distance between arrowheads, and minute first visible tarsomere." figureDoi="http://doi.org/10.5281/zenodo.7333731" httpUri="https://zenodo.org/record/7333731/files/figure.png" pageId="5" pageNumber="457">Fig. 1B</figureCitation>
|
||
) with short palpomere 1 and enlarged, clavate palpomere 2. Palpomere 3 broadly triangular in lateral view, twice as long as wide and approximately two thirds palpomere 2 length, with narrow, elongate, apically globose tubercle emerging dorsally at ~2/5 palpomere length (
|
||
<figureCitation box="[642,765,1598,1624]" captionStart="FIGURE 1" captionStartId="2.[136,235,1800,1824]" captionTargetBox="[152,1437,189,1777]" captionTargetId="figure-15@2.[152,1437,189,1777]" captionTargetPageId="2" captionText="FIGURE 1. Yprezethinus grimaldii Holotype AMNH Tad-130. A, Habitus with visible abdominal sternites labelled. B, Head and pronotum with segments of maxillary palpomeres numbered (MP2–MP4). Bythinoplectina-type lateral excavation, eye, apical pseudosegment of maxillary palp (APs) and mandible (Md) indicated. C, Right antenna with antennomeres numbered. Note the club formed from tightly appressed, hemisphere-shaped antennomeres 10 and 11. D, Close-up view of apically globose tubercles of maxillary palpomeres 3 and 4. E, Proximal leg articulation, showing conically projecting metacoxa (MC), short metatrochanter (MT) and femur (F). F, Right metatarsus (F) with bythinoplectine-type enlarged third tarsomere spanning distance between arrowheads, and minute first visible tarsomere." figureDoi="http://doi.org/10.5281/zenodo.7333731" httpUri="https://zenodo.org/record/7333731/files/figure.png" pageId="5" pageNumber="457">Fig. 1B, D</figureCitation>
|
||
). Palpomere 4 rounded-conical in shape, ~0.5× palpomere 3 length, dorsally bearing narrow, elongate, apically globose tubercle similar to that emerging from palpomere 3 (
|
||
<figureCitation captionStart="FIGURE 1" captionStartId="2.[136,235,1800,1824]" captionTargetBox="[152,1437,189,1777]" captionTargetId="figure-15@2.[152,1437,189,1777]" captionTargetPageId="2" captionText="FIGURE 1. Yprezethinus grimaldii Holotype AMNH Tad-130. A, Habitus with visible abdominal sternites labelled. B, Head and pronotum with segments of maxillary palpomeres numbered (MP2–MP4). Bythinoplectina-type lateral excavation, eye, apical pseudosegment of maxillary palp (APs) and mandible (Md) indicated. C, Right antenna with antennomeres numbered. Note the club formed from tightly appressed, hemisphere-shaped antennomeres 10 and 11. D, Close-up view of apically globose tubercles of maxillary palpomeres 3 and 4. E, Proximal leg articulation, showing conically projecting metacoxa (MC), short metatrochanter (MT) and femur (F). F, Right metatarsus (F) with bythinoplectine-type enlarged third tarsomere spanning distance between arrowheads, and minute first visible tarsomere." figureDoi="http://doi.org/10.5281/zenodo.7333731" httpUri="https://zenodo.org/record/7333731/files/figure.png" pageId="5" pageNumber="457">Fig. 1B, D</figureCitation>
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||
). Apex of palpomere 4 bearing prominent 5th apical pseudosegment (
|
||
<figureCitation box="[327,417,1778,1804]" captionStart="FIGURE 1" captionStartId="2.[136,235,1800,1824]" captionTargetBox="[152,1437,189,1777]" captionTargetId="figure-15@2.[152,1437,189,1777]" captionTargetPageId="2" captionText="FIGURE 1. Yprezethinus grimaldii Holotype AMNH Tad-130. A, Habitus with visible abdominal sternites labelled. B, Head and pronotum with segments of maxillary palpomeres numbered (MP2–MP4). Bythinoplectina-type lateral excavation, eye, apical pseudosegment of maxillary palp (APs) and mandible (Md) indicated. C, Right antenna with antennomeres numbered. Note the club formed from tightly appressed, hemisphere-shaped antennomeres 10 and 11. D, Close-up view of apically globose tubercles of maxillary palpomeres 3 and 4. E, Proximal leg articulation, showing conically projecting metacoxa (MC), short metatrochanter (MT) and femur (F). F, Right metatarsus (F) with bythinoplectine-type enlarged third tarsomere spanning distance between arrowheads, and minute first visible tarsomere." figureDoi="http://doi.org/10.5281/zenodo.7333731" httpUri="https://zenodo.org/record/7333731/files/figure.png" pageId="5" pageNumber="457">Fig. 1B</figureCitation>
|
||
). Estimated length of extended maxillary palpus
|
||
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. Mandibles prominent, narrow, sickle-shaped, apparently raptorial (
|
||
<figureCitation box="[538,625,1850,1876]" captionStart="FIGURE 1" captionStartId="2.[136,235,1800,1824]" captionTargetBox="[152,1437,189,1777]" captionTargetId="figure-15@2.[152,1437,189,1777]" captionTargetPageId="2" captionText="FIGURE 1. Yprezethinus grimaldii Holotype AMNH Tad-130. A, Habitus with visible abdominal sternites labelled. B, Head and pronotum with segments of maxillary palpomeres numbered (MP2–MP4). Bythinoplectina-type lateral excavation, eye, apical pseudosegment of maxillary palp (APs) and mandible (Md) indicated. C, Right antenna with antennomeres numbered. Note the club formed from tightly appressed, hemisphere-shaped antennomeres 10 and 11. D, Close-up view of apically globose tubercles of maxillary palpomeres 3 and 4. E, Proximal leg articulation, showing conically projecting metacoxa (MC), short metatrochanter (MT) and femur (F). F, Right metatarsus (F) with bythinoplectine-type enlarged third tarsomere spanning distance between arrowheads, and minute first visible tarsomere." figureDoi="http://doi.org/10.5281/zenodo.7333731" httpUri="https://zenodo.org/record/7333731/files/figure.png" pageId="5" pageNumber="457">Fig. 1B</figureCitation>
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||
).
|
||
</paragraph>
|
||
<paragraph blockId="5.[136,778,1922,2056]" box="[136,217,1922,1948]" pageId="5" pageNumber="457">
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||
<emphasis box="[136,217,1922,1948]" italics="true" pageId="5" pageNumber="457">Thorax</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="5.[136,778,1922,2056]" pageId="5" pageNumber="457">
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||
Pronotum with rounded margins, widest in apical half. Pronotum
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long, approximate width at widest point
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<quantity box="[201,290,2030,2056]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="5" pageNumber="457" unit="mm" value="0.2">0.2 mm</quantity>
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.
|
||
</paragraph>
|
||
<paragraph blockId="5.[810,1452,158,360]" box="[810,916,158,184]" pageId="5" pageNumber="457">
|
||
<emphasis box="[810,916,158,184]" italics="true" pageId="5" pageNumber="457">Abdomen</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="5.[810,1452,158,360]" pageId="5" pageNumber="457">
|
||
Ventral abdomen length along midline
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||
<quantity box="[1238,1337,193,220]" metricMagnitude="-4" metricUnit="m" metricValue="2.9" pageId="5" pageNumber="457" unit="mm" value="0.29">0.29 mm</quantity>
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(
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<figureCitation box="[1350,1437,193,219]" captionStart="FIGURE 1" captionStartId="2.[136,235,1800,1824]" captionTargetBox="[152,1437,189,1777]" captionTargetId="figure-15@2.[152,1437,189,1777]" captionTargetPageId="2" captionText="FIGURE 1. Yprezethinus grimaldii Holotype AMNH Tad-130. A, Habitus with visible abdominal sternites labelled. B, Head and pronotum with segments of maxillary palpomeres numbered (MP2–MP4). Bythinoplectina-type lateral excavation, eye, apical pseudosegment of maxillary palp (APs) and mandible (Md) indicated. C, Right antenna with antennomeres numbered. Note the club formed from tightly appressed, hemisphere-shaped antennomeres 10 and 11. D, Close-up view of apically globose tubercles of maxillary palpomeres 3 and 4. E, Proximal leg articulation, showing conically projecting metacoxa (MC), short metatrochanter (MT) and femur (F). F, Right metatarsus (F) with bythinoplectine-type enlarged third tarsomere spanning distance between arrowheads, and minute first visible tarsomere." figureDoi="http://doi.org/10.5281/zenodo.7333731" httpUri="https://zenodo.org/record/7333731/files/figure.png" pageId="5" pageNumber="457">Fig. 1A</figureCitation>
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). Width at widest point (along apical margin of tergite IV)
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. Tergites and sternites largely glabrous, but basal sulcus of sternite IV with apparent patch of dense, short (possibly squamous) setae.
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||
</paragraph>
|
||
<paragraph blockId="5.[810,1452,404,711]" box="[810,863,404,429]" pageId="5" pageNumber="457">
|
||
<emphasis box="[810,863,404,429]" italics="true" pageId="5" pageNumber="457">Legs</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="5.[810,1452,404,711]" pageId="5" pageNumber="457">
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||
Femora and tibiae of all legs thickened in apical half; lacking cuticular modifications. Tibiae apically setose on ventral face, most prominent on protibia. Tarsi ventrally with sparse, long mechanosensory setae. Lengths of leg segments: profemur
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<quantity box="[1105,1215,579,605]" metricMagnitude="-4" metricUnit="m" metricValue="1.7999999999999998" pageId="5" pageNumber="457" unit="mm" value="0.18">0.18 mm</quantity>
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, protibia
|
||
<quantity box="[1337,1447,579,605]" metricMagnitude="-4" metricUnit="m" metricValue="1.3" pageId="5" pageNumber="457" unit="mm" value="0.13">0.13 mm</quantity>
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, protarsus
|
||
<quantity box="[920,1023,614,641]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="5" pageNumber="457" unit="mm" value="0.07">0.07 mm</quantity>
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, mesofemur
|
||
<quantity box="[1170,1273,614,641]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="457" unit="mm" value="0.16">0.16 mm</quantity>
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, mesotibia
|
||
<quantity metricMagnitude="-4" metricUnit="m" metricValue="1.4" pageId="5" pageNumber="457" unit="mm" value="0.14">0.14 mm</quantity>
|
||
, mesotarsus
|
||
<quantity box="[997,1098,649,676]" metricMagnitude="-5" metricUnit="m" metricValue="6.0" pageId="5" pageNumber="457" unit="mm" value="0.06">0.06 mm</quantity>
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, metafemur
|
||
<quantity box="[1236,1337,649,676]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="457" unit="mm" value="0.16">0.16 mm</quantity>
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, metatibia
|
||
<quantity box="[810,912,684,711]" metricMagnitude="-4" metricUnit="m" metricValue="1.6" pageId="5" pageNumber="457" unit="mm" value="0.16">0.16 mm</quantity>
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, metatarsus
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<quantity box="[1050,1153,684,711]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="5" pageNumber="457" unit="mm" value="0.07">0.07 mm</quantity>
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.
|
||
</paragraph>
|
||
<paragraph blockId="5.[810,1451,754,850]" box="[810,880,754,780]" pageId="5" pageNumber="457">
|
||
<emphasis box="[810,880,754,780]" italics="true" pageId="5" pageNumber="457">Elytra</emphasis>
|
||
</paragraph>
|
||
<paragraph blockId="5.[810,1451,754,850]" pageId="5" pageNumber="457">
|
||
Elytral length along margin
|
||
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(
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||
<figureCitation box="[1265,1358,789,815]" captionStart="FIGURE 1" captionStartId="2.[136,235,1800,1824]" captionTargetBox="[152,1437,189,1777]" captionTargetId="figure-15@2.[152,1437,189,1777]" captionTargetPageId="2" captionText="FIGURE 1. Yprezethinus grimaldii Holotype AMNH Tad-130. A, Habitus with visible abdominal sternites labelled. B, Head and pronotum with segments of maxillary palpomeres numbered (MP2–MP4). Bythinoplectina-type lateral excavation, eye, apical pseudosegment of maxillary palp (APs) and mandible (Md) indicated. C, Right antenna with antennomeres numbered. Note the club formed from tightly appressed, hemisphere-shaped antennomeres 10 and 11. D, Close-up view of apically globose tubercles of maxillary palpomeres 3 and 4. E, Proximal leg articulation, showing conically projecting metacoxa (MC), short metatrochanter (MT) and femur (F). F, Right metatarsus (F) with bythinoplectine-type enlarged third tarsomere spanning distance between arrowheads, and minute first visible tarsomere." figureDoi="http://doi.org/10.5281/zenodo.7333731" httpUri="https://zenodo.org/record/7333731/files/figure.png" pageId="5" pageNumber="457">Fig. 1A</figureCitation>
|
||
). Elytra apparently glabrous.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |