259 lines
32 KiB
XML
259 lines
32 KiB
XML
<document id="94E3FBDF50F972CA36F1AF4CCAB9D9F4" ID-CLB-Dataset="29757" ID-DOI="10.11646/zootaxa.4442.3.1" ID-GBIF-Dataset="3b3ac8e3-69fc-487a-9ef3-82e8c26f1bab" ID-ISSN="1175-5326" ID-Zenodo-Dep="1303687" ID-ZooBank="0D4F249C-370E-4CF7-BFB6-27E619198924" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1530598421570" checkinUser="plazi" docAuthor="Reynoso-Velasco, Daniel & Sites, Robert W." docDate="2018" docId="0392B40E4549FFB9FF1F3E61FAC9FCDE" docLanguage="en" docName="zootaxa.4442.3.1.pdf" docOrigin="Zootaxa 4442 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ambrysus Stal" docType="treatment" docVersion="7" lastPageNumber="353" masterDocId="FFABCC764548FFBBFF88381CFF86FFE6" masterDocTitle="Revision of the Ambrysus pulchellus Montandon species complex (Heteroptera: Nepomorpha: Naucoridae)" masterLastPageNumber="377" masterPageNumber="351" pageNumber="352" updateTime="1698552331559" updateUser="ExternalLinkService">
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<mods:mods id="5812E6C0B82BF06F08B653F9D220C2CD" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="0810E4C4E404A63E97CB0FCA4B8CE4A5">
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<mods:title id="D5C99DD06E8F7E270F357338CA821241">Revision of the Ambrysus pulchellus Montandon species complex (Heteroptera: Nepomorpha: Naucoridae)</mods:title>
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<mods:namePart id="CEFAA499C1EDB0B99191C42A56C87E69">Reynoso-Velasco, Daniel</mods:namePart>
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<mods:namePart id="4ADB930E785A6903CCCFB91F69C08523">Sites, Robert W.</mods:namePart>
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<mods:typeOfResource id="4F328952F2A78F6123BD59D129909BCD">text</mods:typeOfResource>
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<mods:title id="406CBD9D92F00619F316D92EC3CA33CF">Zootaxa</mods:title>
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<mods:part id="D3B9FCB9624DC41E448441753C4CCC2E">
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<mods:date id="2404BBC3FBC8F9BBD9A0700DB1F547D6">2018</mods:date>
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<mods:number id="3DCF968DDB6D3559820F2959A17EDA6D">2018-07-03</mods:number>
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<mods:detail id="944C2CE1CB24C27C4B498D17FF8EB356" type="volume">
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<mods:number id="39E1965F2D22A69A2770AD3B34174B0D">4442</mods:number>
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<mods:identifier id="89F33338B75CE459C36B701CE232F40D" type="DOI">10.11646/zootaxa.4442.3.1</mods:identifier>
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<treatment id="0392B40E4549FFB9FF1F3E61FAC9FCDE" ID-DOI="http://doi.org/10.5281/zenodo.5989604" ID-GBIF-Taxon="150830042" ID-Zenodo-Dep="5989604" LSID="urn:lsid:plazi:treatment:0392B40E4549FFB9FF1F3E61FAC9FCDE" httpUri="http://treatment.plazi.org/id/0392B40E4549FFB9FF1F3E61FAC9FCDE" lastPageId="2" lastPageNumber="353" pageId="1" pageNumber="352">
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<subSubSection id="C32156934549FFBAFF1F3E61FE1AF971" box="[151,412,1661,1687]" pageId="1" pageNumber="352" type="nomenclature">
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<paragraph id="8B8405184549FFBAFF1F3E61FE1AF971" blockId="1.[151,412,1661,1687]" box="[151,412,1661,1687]" pageId="1" pageNumber="352">
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<heading id="D0CCB2744549FFBAFF1F3E61FE1AF971" bold="true" box="[151,412,1661,1687]" fontSize="11" level="1" pageId="1" pageNumber="352" reason="1">
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<emphasis id="B94FD90A4549FFBAFF1F3E61FE1AF971" bold="true" box="[151,412,1661,1687]" pageId="1" pageNumber="352">
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Genus
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<taxonomicName id="4C3B7E9B4549FFBAFF663E61FE1AF971" ID-CoL="TM3" authority="Stal" authorityName="Stal" box="[238,412,1661,1687]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="genus">
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<emphasis id="B94FD90A4549FFBAFF663E61FEE0F971" bold="true" box="[238,358,1661,1687]" italics="true" pageId="1" pageNumber="352">Ambrysus</emphasis>
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Stål
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32156934549FFB9FF1F3ED8FCECFD63" lastPageId="2" lastPageNumber="353" pageId="1" pageNumber="352" type="discussion">
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<paragraph id="8B8405184549FFBAFF1F3ED8FCBBF88B" blockId="1.[151,1436,1732,2009]" pageId="1" pageNumber="352">
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<emphasis id="B94FD90A4549FFBAFF1F3ED8FE9AF93B" bold="true" box="[151,284,1732,1757]" pageId="1" pageNumber="352">Discussion.</emphasis>
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IN hIs EARLy cLAssIfIcATION Of
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<emphasis id="B94FD90A4549FFBAFDF03ED8FD68F93A" box="[632,750,1732,1756]" italics="true" pageId="1" pageNumber="352">
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<taxonomicName id="4C3B7E9B4549FFBAFDF03ED8FD6FF93A" box="[632,745,1732,1756]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="genus">Ambrysus</taxonomicName>
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,
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</emphasis>
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LA
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<bibRefCitation id="EFAA78E94549FFBAFCA93ED9FC39F93B" box="[801,959,1732,1757]" pageId="1" pageNumber="352" refId="ref29578" refString="La Rivers, I. (1951) A revision of the genus Ambrysus in the United States (Hemiptera: Naucoridae). University of California Publications in Entomology, 8, 277 - 338." type="journal article">RIvERs (1951)</bibRefCitation>
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dIvIdEd ThE subGENus
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<taxonomicName id="4C3B7E9B4549FFBAFB423ED8FABEF93A" box="[1226,1336,1732,1756]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="genus">
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<emphasis id="B94FD90A4549FFBAFB423ED8FABEF93A" box="[1226,1336,1732,1756]" italics="true" pageId="1" pageNumber="352">Ambrysus</emphasis>
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</taxonomicName>
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INTO ThE
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<emphasis id="B94FD90A4549FFBAFF1F3EF4FE91F8E6" box="[151,279,1768,1792]" italics="true" pageId="1" pageNumber="352">Coalescens</emphasis>
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ANd
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<emphasis id="B94FD90A4549FFBAFEC53EF4FE43F919" box="[333,453,1768,1791]" italics="true" pageId="1" pageNumber="352">Disjunctus</emphasis>
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dIvIsIONs. LATER, ThE fORmER dIvIsION wOuLd bE ELEvATEd TO subGENERIc RANK As
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<emphasis id="B94FD90A4549FFBAFAB93EF4FA1AF8E6" box="[1329,1436,1768,1792]" italics="true" pageId="1" pageNumber="352">Syncollus</emphasis>
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ANd ThE REmINdING 52 spEcIEs IN
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<emphasis id="B94FD90A4549FFBAFD993F11FD0FF8C2" box="[529,649,1805,1828]" italics="true" pageId="1" pageNumber="352">Disjunctus</emphasis>
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AssIGNEd TO ThE subGENus
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<taxonomicName id="4C3B7E9B4549FFBAFC323F10FBAEF8C2" box="[954,1064,1804,1828]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="genus">
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<emphasis id="B94FD90A4549FFBAFC323F10FBAEF8C2" box="[954,1064,1804,1828]" italics="true" pageId="1" pageNumber="352">Ambrysus</emphasis>
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</taxonomicName>
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(LA
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<bibRefCitation id="EFAA78E94549FFBAFBEA3F11FB76F8C3" box="[1122,1264,1804,1829]" pageId="1" pageNumber="352" refId="ref29766" refString="La Rivers, I. (1965) The subgenera of the genus Ambrysus. Biological Society of Nevada Occasional Papers, 4, 1 - 7." type="journal article">RIvERs 1965</bibRefCitation>
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), ExcEpT fOR
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<taxonomicName id="4C3B7E9B4549FFBAFA0C3F10FF75F8AE" authorityName="La Rivers" authorityYear="1949" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="funebris">
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<emphasis id="B94FD90A4549FFBAFA0C3F10FF75F8AE" italics="true" pageId="1" pageNumber="352">A. funebris</emphasis>
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</taxonomicName>
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LA RIvERs ThAT wAs AssIGNEd TO ThE ThEREIN cREATEd subGENus
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<emphasis id="B94FD90A4549FFBAFC2B3F2CFB93F8A1" box="[931,1045,1840,1863]" italics="true" pageId="1" pageNumber="352">
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<taxonomicName id="4C3B7E9B4549FFBAFC2B3F2CFB97F8A1" authorityName="La Rivers" authorityYear="1965" box="[931,1041,1840,1863]" class="Insecta" family="Naucoridae" genus="Acyttarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="genus">Acyttarus</taxonomicName>
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,
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</emphasis>
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ANd
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<taxonomicName id="4C3B7E9B4549FFBAFBC53F2CFA99F8A1" authorityName="La Rivers" authorityYear="1952" box="[1101,1311,1840,1864]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="usingeri">
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<emphasis id="B94FD90A4549FFBAFBC53F2CFA99F8A1" box="[1101,1311,1840,1864]" italics="true" pageId="1" pageNumber="352">Ambrysus usingeri</emphasis>
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</taxonomicName>
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LA RIvERs, pREvIOusLy pLAcEd IN ThE subGENus
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<taxonomicName id="4C3B7E9B4549FFBAFD913F49FDE9F88A" authorityName="La Rivers" authorityYear="1952" box="[537,623,1877,1900]" class="Insecta" family="Naucoridae" genus="Picrops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="genus">
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<emphasis id="B94FD90A4549FFBAFD913F49FDE9F88A" box="[537,623,1877,1900]" italics="true" pageId="1" pageNumber="352">Picrops</emphasis>
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</taxonomicName>
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(LA
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<bibRefCitation id="EFAA78E94549FFBAFD2C3F49FCB6F88B" box="[676,816,1876,1901]" pageId="1" pageNumber="352" refId="ref29615" refString="La Rivers, I. (1952) A new subgenus of Ambrysus from South America (Hemiptera: Naucoridae). Entomological News, 63, 33 - 39." type="journal article">RIvERs 1952</bibRefCitation>
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).
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</paragraph>
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<paragraph id="8B8405184549FFB9FF4F3F64FAB9FEFA" blockId="1.[151,1436,1732,2009]" lastBlockId="2.[151,1436,151,824]" lastPageId="2" lastPageNumber="353" pageId="1" pageNumber="352">
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IN hIs
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<emphasis id="B94FD90A4549FFBAFE953F64FE13F869" box="[285,405,1912,1935]" italics="true" pageId="1" pageNumber="352">Disjunctus</emphasis>
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dIvIsION, LA
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<bibRefCitation id="EFAA78E94549FFBAFDCA3F64FD63F876" box="[578,741,1911,1936]" pageId="1" pageNumber="352" refId="ref29578" refString="La Rivers, I. (1951) A revision of the genus Ambrysus in the United States (Hemiptera: Naucoridae). University of California Publications in Entomology, 8, 277 - 338." type="journal article">RIvERs (1951)</bibRefCitation>
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INcLudEd EIGhT spEcIEs cOmpLExEs (
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<taxonomicName id="4C3B7E9B4549FFBAFB153F64FAA5F876" authorityName=": La Rivers" authorityYear="1949" box="[1181,1315,1912,1936]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="californicus">
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<emphasis id="B94FD90A4549FFBAFB153F64FAA5F876" box="[1181,1315,1912,1936]" italics="true" pageId="1" pageNumber="352">californicus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C3B7E9B4549FFBAFAB13F64FA13F876" authorityName="La Rivers" authorityYear="1949" box="[1337,1429,1912,1936]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="funebris">
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<emphasis id="B94FD90A4549FFBAFAB13F64FA13F876" box="[1337,1429,1912,1936]" italics="true" pageId="1" pageNumber="352">funebris</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C3B7E9B4549FFBAFF1F3F80FEA9F852" box="[151,303,1948,1972]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="melanopterus">
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<emphasis id="B94FD90A4549FFBAFF1F3F80FEA9F852" box="[151,303,1948,1972]" italics="true" pageId="1" pageNumber="352">melanopterus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C3B7E9B4549FFBAFECA3F80FE1FF852" box="[322,409,1948,1972]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="pudicus">
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<emphasis id="B94FD90A4549FFBAFECA3F80FE1FF852" box="[322,409,1948,1972]" italics="true" pageId="1" pageNumber="352">pudicus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C3B7E9B4549FFBAFE243F80FD99F852" authorityName=": Montandon" authorityYear="1909" box="[428,543,1948,1972]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="pulchellus">
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<emphasis id="B94FD90A4549FFBAFE243F80FD99F852" box="[428,543,1948,1972]" italics="true" pageId="1" pageNumber="352">pulchellus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C3B7E9B4549FFBAFDB93F80FD32F852" authorityName="Stål" authorityYear="1876" box="[561,692,1948,1972]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="puncticollis">
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<emphasis id="B94FD90A4549FFBAFDB93F80FD32F852" box="[561,692,1948,1972]" italics="true" pageId="1" pageNumber="352">puncticollis</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C3B7E9B4549FFBAFD4F3F81FCACF852" box="[711,810,1949,1972]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="signoreti">
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<emphasis id="B94FD90A4549FFBAFD4F3F81FCACF852" box="[711,810,1949,1972]" italics="true" pageId="1" pageNumber="352">signoreti</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C3B7E9B4549FFBAFCB43F80FC2AF852" authorityName="Usinger. The" authorityYear="1946" box="[828,940,1948,1972]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="vanduzeei">
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<emphasis id="B94FD90A4549FFBAFCB43F80FC2AF852" box="[828,940,1948,1972]" italics="true" pageId="1" pageNumber="352">vanduzeei</emphasis>
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</taxonomicName>
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) fOR ThE NORTh AmERIcAN spEcIEs, EAch Of whIch wAs mONOTypIc, ExcEpT
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<taxonomicName id="4C3B7E9B4549FFBAFD8D3FDCFDDAF83E" box="[517,604,1984,2008]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="pudicus">
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<emphasis id="B94FD90A4549FFBAFD8D3FDCFDDAF83E" box="[517,604,1984,2008]" italics="true" pageId="1" pageNumber="352">pudicus</emphasis>
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</taxonomicName>
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ANd
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<taxonomicName id="4C3B7E9B4549FFBAFD283FDDFC85F83E" box="[672,771,1985,2008]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="1" pageNumber="352" phylum="Arthropoda" rank="species" species="signoreti">
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<emphasis id="B94FD90A4549FFBAFD283FDDFC85F83E" box="[672,771,1985,2008]" italics="true" pageId="1" pageNumber="352">signoreti</emphasis>
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</taxonomicName>
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. FuRThERmORE, LA
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<bibRefCitation id="EFAA78E94549FFBAFC6C3FDDFB03F83F" box="[996,1157,1984,2009]" pageId="1" pageNumber="352" refId="ref29578" refString="La Rivers, I. (1951) A revision of the genus Ambrysus in the United States (Hemiptera: Naucoridae). University of California Publications in Entomology, 8, 277 - 338." type="journal article">RIvERs (1951)</bibRefCitation>
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AssIGNEd fOuR Of ThEsE cOmpLExEs TO TwO uNNAmEd ANd pOORLy dEfINEd dIvIsIONs. ThE fIRsT dIvIsION INcLudEd mEmbERs wITh ThE pROpLEuRA fusEd OR TOuchING ALONG ThE mIdLINE (
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<taxonomicName id="4C3B7E9B454AFFB9FDC938A1FD25FF32" box="[577,675,189,212]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="signoreti">
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<emphasis id="B94FD90A454AFFB9FDC938A1FD25FF32" box="[577,675,189,212]" italics="true" pageId="2" pageNumber="353">signoreti</emphasis>
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</taxonomicName>
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ANd
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<taxonomicName id="4C3B7E9B454AFFB9FD5638A0FCC8FF32" authorityName="Usinger. The" authorityYear="1946" box="[734,846,188,212]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="vanduzeei">
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<emphasis id="B94FD90A454AFFB9FD5638A0FCC8FF32" box="[734,846,188,212]" italics="true" pageId="2" pageNumber="353">vanduzeei</emphasis>
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</taxonomicName>
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). ThE sEcONd INcLudEd mEmbERs wITh ThE pROpLEuRA NOT TOuchING ALONG ThE mIdLINE (
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<taxonomicName id="4C3B7E9B454AFFB9FDAD38FCFD2DFF1E" authorityName=": La Rivers" authorityYear="1949" box="[549,683,224,248]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="californicus">
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<emphasis id="B94FD90A454AFFB9FDAD38FCFD2DFF1E" box="[549,683,224,248]" italics="true" pageId="2" pageNumber="353">californicus</emphasis>
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</taxonomicName>
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ANd
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|
<taxonomicName id="4C3B7E9B454AFFB9FD7938FCFCE2FF1E" authorityName=": Montandon" authorityYear="1909" box="[753,868,224,248]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="pulchellus">
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<emphasis id="B94FD90A454AFFB9FD7938FCFCE2FF1E" box="[753,868,224,248]" italics="true" pageId="2" pageNumber="353">pulchellus</emphasis>
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</taxonomicName>
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). EvEN LA
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<bibRefCitation id="EFAA78E9454AFFB9FC7038FCFB1CFF1E" box="[1016,1178,223,248]" pageId="2" pageNumber="353" refId="ref29578" refString="La Rivers, I. (1951) A revision of the genus Ambrysus in the United States (Hemiptera: Naucoridae). University of California Publications in Entomology, 8, 277 - 338." type="journal article">RIvERs (1951)</bibRefCitation>
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cONsIdEREd ThE LATTER cLAssIfIcATION TO bE "wEAK" ANd INdIcATEd ThAT ThE RELIAbILITy Of ThE chARAcTERs REquIREd fuRThER ExAmINATION.
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</paragraph>
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<paragraph id="8B840518454AFFB9FF4F3934FDC5FE36" blockId="2.[151,1436,151,824]" pageId="2" pageNumber="353">
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FOR TwO (
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<taxonomicName id="4C3B7E9B454AFFB9FEB93934FE31FEA6" authorityName=": La Rivers" authorityYear="1949" box="[305,439,296,320]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="californicus">
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<emphasis id="B94FD90A454AFFB9FEB93934FE31FEA6" box="[305,439,296,320]" italics="true" pageId="2" pageNumber="353">californicus</emphasis>
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</taxonomicName>
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ANd
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<taxonomicName id="4C3B7E9B454AFFB9FE673934FDE4FEA6" authorityName=": Montandon" authorityYear="1909" box="[495,610,296,320]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="pulchellus">
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<emphasis id="B94FD90A454AFFB9FE673934FDE4FEA6" box="[495,610,296,320]" italics="true" pageId="2" pageNumber="353">pulchellus</emphasis>
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</taxonomicName>
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) Of ThE ThREE spEcIEs cOmpLExEs Of INTEREsT fOR ThIs sTudy, NO dETAILs wERE pROvIdEd TO jusTIfy ThEIR fORmATION ANd mONOTypIc sTATus. IN ThE cAsE Of ThE GROup
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<taxonomicName id="4C3B7E9B454AFFB9FBEF3950FB51FE82" authorityName="Usinger. The" authorityYear="1946" box="[1127,1239,332,356]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="vanduzeei">
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<emphasis id="B94FD90A454AFFB9FBEF3950FB51FE82" box="[1127,1239,332,356]" italics="true" pageId="2" pageNumber="353">vanduzeei</emphasis>
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</taxonomicName>
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, ThE sEmITubuLAR cONdITION Of ThE fEmALE mEdIOsTERNITE VII (subGENITAL pLATE) ANd ThE shApE Of ThE mALE AccEssORy GENITALIc pROcEss Of TERGum VI (AGP) wERE usEd TO jusTIfy ThE spEcIEs cOmpLEx sTATus.
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<taxonomicName id="4C3B7E9B454AFFB9FC1C3988FB33FE4A" authority="UsINGER" authorityName="Usinger" authorityYear="1946" box="[916,1205,404,428]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="355" phylum="Arthropoda" rank="species" species="buenoi">
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<emphasis id="B94FD90A454AFFB9FC1C3988FBD0FE4A" box="[916,1110,404,428]" italics="true" pageId="2" pageNumber="353">Ambrysus buenoi</emphasis>
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UsINGER
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</taxonomicName>
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wAs NEvER ExpLIcITLy AssIGNEd TO A pARTIcuLAR spEcIEs GROup.
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</paragraph>
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<paragraph id="8B840518454AFFB9FF4F39C0FCECFD63" blockId="2.[151,1436,151,824]" pageId="2" pageNumber="353">
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BAsEd ON chARAcTERIsTIcs Of ThE phALLOsOmA ANd shApE Of ThE mEdIAL LObEs Of mALE AbdOmINAL TERGum VIII (psEudOpARAmEREs), ThE spEcIEs cLOsELy RELATEd TO
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<taxonomicName id="4C3B7E9B454AFFB9FD413A1CFCDBFDFE" authorityName=": Montandon" authorityYear="1909" box="[713,861,512,536]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="pulchellus">
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<emphasis id="B94FD90A454AFFB9FD413A1CFCDBFDFE" box="[713,861,512,536]" italics="true" pageId="2" pageNumber="353">A. pulchellus</emphasis>
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</taxonomicName>
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|
ARE:
|
|
<taxonomicName id="4C3B7E9B454AFFB9FC103A1CFB82FDFE" authorityName="Usinger" authorityYear="1946" box="[920,1028,512,536]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="buenoi">
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<emphasis id="B94FD90A454AFFB9FC103A1CFB82FDFE" box="[920,1028,512,536]" italics="true" pageId="2" pageNumber="353">A. buenoi</emphasis>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="4C3B7E9B454AFFB9FB9C3A1CFAC1FDFE" authority="MONTANdON" authorityName=": Montandon" authorityYear="1909" box="[1044,1351,511,536]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="californicus">
|
|
<emphasis id="B94FD90A454AFFB9FB9C3A1CFB3CFDFE" box="[1044,1210,512,536]" italics="true" pageId="2" pageNumber="353">A. californicus</emphasis>
|
|
MONTANdON
|
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</taxonomicName>
|
|
, ANd
|
|
<taxonomicName id="4C3B7E9B454AFFB9FA0C3A1CFEF4FDDA" authority="UsINGER." authorityName="Usinger. The" authorityYear="1946" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="351" phylum="Arthropoda" rank="species" species="vanduzeei">
|
|
<emphasis id="B94FD90A454AFFB9FA0C3A1CFE81FDDA" italics="true" pageId="2" pageNumber="353">A. vanduzeei</emphasis>
|
|
UsINGER.
|
|
</taxonomicName>
|
|
WE REfER TO ThIs GROup As ThE
|
|
<taxonomicName id="4C3B7E9B454AFFB9FD543A38FCF4FDDA" authorityName=": Montandon" authorityYear="1909" box="[732,882,548,572]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="pulchellus">
|
|
<emphasis id="B94FD90A454AFFB9FD543A38FCF4FDDA" box="[732,882,548,572]" italics="true" pageId="2" pageNumber="353">A. pulchellus</emphasis>
|
|
</taxonomicName>
|
|
cOmpLEx bEcAusE ThIs wAs ThE fIRsT Of ThE fOuR cONsTITuENT spEcIEs TO bE dEscRIbEd. ALThOuGh bOTh
|
|
<taxonomicName id="4C3B7E9B454AFFB9FD6A3A54FCF1FD86" authorityName=": Montandon" authorityYear="1909" box="[738,887,584,608]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="pulchellus">
|
|
<emphasis id="B94FD90A454AFFB9FD6A3A54FCF1FD86" box="[738,887,584,608]" italics="true" pageId="2" pageNumber="353">A. pulchellus</emphasis>
|
|
</taxonomicName>
|
|
ANd
|
|
<taxonomicName id="4C3B7E9B454AFFB9FC3A3A54FBDCFD86" authorityName=": Montandon" authorityYear="1909" box="[946,1114,584,608]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="californicus">
|
|
<emphasis id="B94FD90A454AFFB9FC3A3A54FBDCFD86" box="[946,1114,584,608]" italics="true" pageId="2" pageNumber="353">A. californicus</emphasis>
|
|
</taxonomicName>
|
|
wERE dEscRIbEd IN ThE sAmE pApER (
|
|
<bibRefCitation id="EFAA78E9454AFFB9FF693A70FE2FFD62" box="[225,425,620,645]" pageId="2" pageNumber="353" refId="ref29875" refString="Montandon, A. L. (1897) Hemiptera Cryptocerata. Fam. Naucoridae. Sous-fam. Cryptocricinae. Ferhandlungen Zoologisch- Botanischen Gesellschaft in Wien, 47, 6 - 23." type="journal article">MONTANdON 1897</bibRefCitation>
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|
),
|
|
<taxonomicName id="4C3B7E9B454AFFB9FE343A70FDC9FD62" authorityName=": Montandon" authorityYear="1909" box="[444,591,620,644]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="pulchellus">
|
|
<emphasis id="B94FD90A454AFFB9FE343A70FDC9FD62" box="[444,591,620,644]" italics="true" pageId="2" pageNumber="353">A. pulchellus</emphasis>
|
|
</taxonomicName>
|
|
hAs pOsITION pREcEdENcE.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection id="C3215693454AFFB9FF4F3A93FAC9FCDE" pageId="2" pageNumber="353" type="diagnosis">
|
|
<paragraph id="8B840518454AFFB9FF4F3A93FAC9FCDE" blockId="2.[151,1436,151,824]" pageId="2" pageNumber="353">
|
|
<emphasis id="B94FD90A454AFFB9FF4F3A93FEC7FD4E" bold="true" box="[199,321,655,680]" pageId="2" pageNumber="353">Diagnosis.</emphasis>
|
|
SImILAR TO ThE pREvIOus spEcIEs cOmpLExEs pROpOsEd by
|
|
<bibRefCitation id="EFAA78E9454AFFB9FC423A8CFAEDFD4E" box="[970,1387,655,680]" pageId="2" pageNumber="353" refId="ref30257" refString="Reynoso-Velasco, D. & Sites, R. W. (2016 b) Revision of the Ambrysus hybridus Montandon species complex (Heteroptera: Naucoridae: Cryphocricinae) with the description of a new species from Mexico. Zootaxa, 4169 (1), 87 - 114. https: // doi. org / 10.11646 / zootaxa. 4169.1.4" type="journal article">REyNOsO-VELAscO & SITEs (2016A, b)</bibRefCitation>
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|
, ThE mOsT RELIAbLE uNIfyING fEATuREs AmONG spEcIEs IN ThE
|
|
<taxonomicName id="4C3B7E9B454AFFB9FD6E3AA8FCFEFD2A" authorityName=": Montandon" authorityYear="1909" box="[742,888,692,716]" class="Insecta" family="Naucoridae" genus="Ambrysus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="353" phylum="Arthropoda" rank="species" species="pulchellus">
|
|
<emphasis id="B94FD90A454AFFB9FD6E3AA8FCFEFD2A" box="[742,888,692,716]" italics="true" pageId="2" pageNumber="353">A. pulchellus</emphasis>
|
|
</taxonomicName>
|
|
cOmpLEx INcLudE mALEs wITh A LEfT dORsAL fLAp ON ThE phALLOsOmA (E.G.,
|
|
<figureCitation id="1300199D454AFFB9FE0E3AC4FD92FD16" box="[390,532,728,752]" captionStart="FIGURE 1" captionStartId="4.[151,250,1864,1886]" captionTargetBox="[151,1436,424,1858]" captionTargetId="figure@4.[151,1436,424,1858]" captionTargetPageId="4" captionText="FIGURE 1. Structures of Ambrysus buenoi. (A) dorsal habitus of male, (B) 6th and 7th abdominal terga of male, (C) 8th abdominal tergum of male, (D) ventral surface of phallosoma, (E) left side of phallososma, (F) sternum VI and subgenital plate of female (terminal abdominal segments removed; arrow indicates posterior margin of laterosternite VI). agp = accessory genitalic process, al = apical lobe, ldf = left dorsal flap, lvl = left ventral lobe, mc = medial convexity, plc = posterolateral corner, pp = pseudoparamere, sct = scutellum, sgp = subgenital plate. Size bar = 0.5 mm and applies only to figures D and E." httpUri="https://zenodo.org/record/1303689/files/figure.png" pageId="2" pageNumber="353">FIGs. 1D, 1E</figureCitation>
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,
|
|
<figureCitation id="1300199D454AFFB9FD973AC4FDF7FD16" box="[543,625,728,752]" captionStart="FIGURE 4" captionStartId="11.[151,250,1634,1656]" captionTargetBox="[151,1436,188,1622]" captionTargetId="figure@11.[151,1436,188,1622]" captionTargetPageId="11" captionText="FIGURE 4. Structures of Ambrysus pulchellus. (A) dorsal habitus of female, (B) 6th and 7th abdominal terga of male, (C) 8th abdominal tergum of male, (D) ventral surface of phallosoma, (E) left side of phallososma, (F) sternum VI and subgenital plate of female (terminal abdominal segments removed). plc = posterolateral corner. Size bar = 0.5 mm and applies only to figures D and E." httpUri="https://zenodo.org/record/1303695/files/figure.png" pageId="2" pageNumber="353">4D, 4E</figureCitation>
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), ThE LEfT vENTRAL LObE Of ThE phALLOsOmA LATEROfLExEd (E.G.,
|
|
<figureCitation id="1300199D454AFFB9FB743AC4FAD8FD16" box="[1276,1374,728,752]" captionStart="FIGURE 1" captionStartId="4.[151,250,1864,1886]" captionTargetBox="[151,1436,424,1858]" captionTargetId="figure@4.[151,1436,424,1858]" captionTargetPageId="4" captionText="FIGURE 1. Structures of Ambrysus buenoi. (A) dorsal habitus of male, (B) 6th and 7th abdominal terga of male, (C) 8th abdominal tergum of male, (D) ventral surface of phallosoma, (E) left side of phallososma, (F) sternum VI and subgenital plate of female (terminal abdominal segments removed; arrow indicates posterior margin of laterosternite VI). agp = accessory genitalic process, al = apical lobe, ldf = left dorsal flap, lvl = left ventral lobe, mc = medial convexity, plc = posterolateral corner, pp = pseudoparamere, sct = scutellum, sgp = subgenital plate. Size bar = 0.5 mm and applies only to figures D and E." httpUri="https://zenodo.org/record/1303689/files/figure.png" pageId="2" pageNumber="353">FIGs. 1D</figureCitation>
|
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,
|
|
<figureCitation id="1300199D454AFFB9FAE23AC4FA09FD16" box="[1386,1423,728,752]" captionStart="FIGURE 4" captionStartId="11.[151,250,1634,1656]" captionTargetBox="[151,1436,188,1622]" captionTargetId="figure@11.[151,1436,188,1622]" captionTargetPageId="11" captionText="FIGURE 4. Structures of Ambrysus pulchellus. (A) dorsal habitus of female, (B) 6th and 7th abdominal terga of male, (C) 8th abdominal tergum of male, (D) ventral surface of phallosoma, (E) left side of phallososma, (F) sternum VI and subgenital plate of female (terminal abdominal segments removed). plc = posterolateral corner. Size bar = 0.5 mm and applies only to figures D and E." httpUri="https://zenodo.org/record/1303695/files/figure.png" pageId="2" pageNumber="353">4D</figureCitation>
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), ANd AN OvERALL sImILAR shApE Of ThE psEudOpARAmEREs. FEmALEs Of ThIs cOmpLEx ExhIbIT A GREAT dEGREE Of vARIATION Of ThE subGENITAL pLATE, ALThOuGh ThEsE dIffERENcEs pROvIdE ThE mOsT RELIAbLE INTERspEcIfIc dIAGNOsTIc chARAcTERs.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |