204 lines
28 KiB
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204 lines
28 KiB
XML
<document id="58323D90E36004340F29DC893788DC8A" ID-CLB-Dataset="7746" ID-DOI="10.5252/geodiversitas2021v43a5" ID-GBIF-Dataset="e42658a2-0e15-4e90-9bc5-0d00cbcbd781" ID-ISSN="1638-9395" ID-Zenodo-Dep="4605963" ID-ZooBank="urn:lsid:zoobank.org:pub:697FC553-E37B-4EF9-97A4-950E4DEE246C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615828395930" checkinUser="felipe" docAuthor="Zouhri, Samir, Gingerich, Philip D., Khalloufi, Bouziane, Bourdon, Estelle, Adnet, Sylvain, Jouve, Stéphane, Elboudali, Najia, Amane, Ayoub, Rage, Jean-Claude & Tabuce, Rodolphe" docDate="2021" docId="03923C45FF92FF9B30E4FB0BFA821043" docLanguage="en" docName="Geodiversitas.43.5.121-150.pdf" docOrigin="Geodiversitas 43 (5)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Gavialoidea Hay 1930" docType="treatment" docVersion="7" lastPageNumber="138" masterDocId="FFAB443DFF81FF883018FFDFFF95166E" masterDocTitle="Middle Eocene vertebrate fauna from the Aridal Formation, Sabkha of Gueran, southwestern Morocco" masterLastPageNumber="150" masterPageNumber="121" pageNumber="138" updateTime="1698933443279" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:titleInfo id="307FA42B2EC77B6C3559DA7F9103A920">
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<mods:title id="1C6B0D5062636B345706FBA5447DD359">Middle Eocene vertebrate fauna from the Aridal Formation, Sabkha of Gueran, southwestern Morocco</mods:title>
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<mods:namePart id="1F9B3785D64F798CAF6964170247ACA0">Zouhri, Samir</mods:namePart>
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<mods:namePart id="4C5AC9551BCFE4FBD7A2162404FDE86E">Gingerich, Philip D.</mods:namePart>
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<mods:namePart id="95BC7A052AF6AEFC65744CAACFA395E6">Khalloufi, Bouziane</mods:namePart>
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<mods:namePart id="1F2BA51AB509215658F5523F36FCE818">Bourdon, Estelle</mods:namePart>
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<mods:namePart id="16F453182B1AF9AAD5B81411F7C548B8">Adnet, Sylvain</mods:namePart>
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<mods:namePart id="9DCCCB4719A1C60BB8F5CDCEF1EDABCB">Jouve, Stéphane</mods:namePart>
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<mods:namePart id="3B602E6F5913F2898CB64E8088D0A459">Elboudali, Najia</mods:namePart>
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<mods:namePart id="5FD99AA2938407CA57FB566FA875BA24">Amane, Ayoub</mods:namePart>
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<mods:namePart id="726D57F79F389A95CBEE8E0AEFBF8055">Rage, Jean-Claude</mods:namePart>
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<mods:namePart id="ECE08E502E436C6A4E0395C07FB0ED2F">Tabuce, Rodolphe</mods:namePart>
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<mods:typeOfResource id="EBF5A8BA831154F2518329FF61D1A807">text</mods:typeOfResource>
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<mods:titleInfo id="211A82FB8BFB694357F8BA43E24DA0DD">
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<mods:title id="0416041B53213D105F27AEE9EF9E72C5">Geodiversitas</mods:title>
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<mods:part id="D51D8A7304779F1D6AC4477488494F75">
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<mods:date id="5E847971E3601E79729AB815E83E1602">2021</mods:date>
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<mods:detail id="E2A2DE7B301C5D1BDBB606E4CE570A5C" type="pubDate">
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<mods:number id="5D9B66135A4FCD2BDFEA7DA2E67058A5">2021-03-11</mods:number>
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<mods:detail id="8F3F8DD66DA1A8F9C6CEB0A633E23C9C" type="volume">
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<mods:number id="DAAE6E7544528D72D0ACE40BABD59FFA">43</mods:number>
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<mods:detail id="38DB16DBECD88713970ADA8FE425AEFD" type="issue">
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<mods:number id="5228980478E891C8B1A963B5D8D3D68F">5</mods:number>
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<mods:start id="344B333024C41F2163FFD51D4B90408D">121</mods:start>
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<mods:end id="0751A4CAB3E4828C808FAF2D6245F5F5">150</mods:end>
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<mods:classification id="3D358E8CD03B98F634B5F4776C7B7027">journal article</mods:classification>
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<mods:identifier id="13A1E6EE606323300591B5C34B938D27" type="CLB-Dataset">7746</mods:identifier>
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<mods:identifier id="091104221F0AEE1A1FC03DA9F56B6302" type="DOI">10.5252/geodiversitas2021v43a5</mods:identifier>
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<mods:identifier id="B17D0C0796A9647CFE7C2480F7FCF6C7" type="GBIF-Dataset">e42658a2-0e15-4e90-9bc5-0d00cbcbd781</mods:identifier>
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<mods:identifier id="597C31DC46691F51238101AB005611A5" type="ISSN">1638-9395</mods:identifier>
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<mods:identifier id="A561217D01E04F091BF6141FB634E5EF" type="Zenodo-Dep">4605963</mods:identifier>
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<mods:identifier id="C1226E1E7678D8A9A9B1D64E6A98F9EE" type="ZooBank">urn:lsid:zoobank.org:pub:697FC553-E37B-4EF9-97A4-950E4DEE246C</mods:identifier>
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<treatment id="03923C45FF92FF9B30E4FB0BFA821043" ID-DOI="http://doi.org/10.5281/zenodo.4606651" ID-GBIF-Taxon="180569433" ID-Zenodo-Dep="4606651" LSID="urn:lsid:plazi:treatment:03923C45FF92FF9B30E4FB0BFA821043" httpUri="http://treatment.plazi.org/id/03923C45FF92FF9B30E4FB0BFA821043" lastPageNumber="138" pageId="19" pageNumber="138">
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<subSubSection id="C321DED8FF92FF9B30E4FB0BFD1A1280" box="[252,655,1236,1262]" pageId="19" pageNumber="138" type="nomenclature">
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<paragraph id="8B848D53FF92FF9B30E4FB0BFD1A1280" blockId="19.[252,655,1236,1262]" box="[252,655,1236,1262]" pageId="19" pageNumber="138">
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<heading id="D0CC3A3FFF92FF9B30E4FB0BFD1A1280" box="[252,655,1236,1262]" centered="true" fontSize="9" level="2" pageId="19" pageNumber="138" reason="7">
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Superfamily
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<taxonomicName id="4C3BF6D0FF92FF9B319DFB0BFD1A1280" authority="Hay, 1930" authorityName="Hay" authorityYear="1930" box="[389,655,1236,1262]" kingdom="Animalia" pageId="19" pageNumber="138" rank="superFamily" superFamily="Gavialoidea">GAVIALOIDEA Hay, 1930</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C321DED8FF92FF9B317BFACCFDBC1343" box="[355,553,1299,1325]" pageId="19" pageNumber="138" type="description">
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<paragraph id="8B848D53FF92FF9B317BFACCFDBC1343" blockId="19.[355,553,1299,1325]" box="[355,553,1299,1325]" pageId="19" pageNumber="138">
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<taxonomicName id="4C3BF6D0FF92FF9B317BFACCFE771343" authorityName="Hay" authorityYear="1930" box="[355,482,1299,1325]" kingdom="Animalia" pageId="19" pageNumber="138" rank="superFamily" superFamily="Gavialoidea">Gavialoidea</taxonomicName>
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indet.
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</paragraph>
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</subSubSection>
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<subSubSection id="C321DED8FF92FF9B309CFA92FD6B13DA" pageId="19" pageNumber="138" type="materials_examined">
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<paragraph id="8B848D53FF92FF9B309CFA92FD6B13DA" blockId="19.[131,776,1356,1460]" pageId="19" pageNumber="138">
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EXAMINED MATERIAL. —
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<materialsCitation id="3B53870EFF92FF9B319AFA93FD6B13DA" ID-GBIF-Occurrence="3053068321" pageId="19" pageNumber="138">FSACBouj-401, 403 and 404, anterior and posterior portion of left maxilla; 407, posterior portion of a left maxilla; 402, mid-portion of a right maxilla. All these specimens are from the same individual. Also 405, portion of a left dentary.</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C321DED8FF92FF9B309CFA0CFB83173F" pageId="19" pageNumber="138" type="description">
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<paragraph id="8B848D53FF92FF9B309CFA0CFE801385" blockId="19.[130,777,1491,2028]" box="[132,277,1491,1517]" pageId="19" pageNumber="138">DESCRIPTION</paragraph>
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<paragraph id="8B848D53FF92FF9B309CFA2CFDEC1185" blockId="19.[130,777,1491,2028]" pageId="19" pageNumber="138">
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The reconstruction of the maxillae shows a slender snouted form with 16 preserved teeth but more teeth were probably present (
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<figureCitation id="130091D6FF92FF9B3095F9EDFF591023" box="[141,204,1586,1613]" captionStart="FIG" captionStartId="20.[132,143,1407,1424]" captionTargetBox="[132,1455,215,1365]" captionTargetId="figure-294@20.[698,1267,789,1366]" captionTargetPageId="20" captionText="FIG. 7. — Crocodyliformes remains from Gueran: A-J, Eusuchia indet.: A, FSAC BOUJ-410, last cervical (ninth) vertebra in left lateral view; B, FSAC BOUJ- 1b, anterior cervical vertebra in left lateral view; C, FSAC BOUJ-400, posterior dorsal vertebra in left lateral view; D, FSAC BOUJ-1a, first caudal vertebra in left lateral view; E, FSAC BOUJ-124, caudal vertebra in left lateral view; F, FSAC BOUJ-94, osteoderm in dorsal view; G, FSAC BOUJ-96, posterior portion of a dorsal osteoderm; H, FSAC BOUJ-96, lateral portion of a dorsal osteoderm in dorsal views; I, J, Crocodyliformes indet.: I, FSAC BOUJ-355, mid portion of a dentary in dorsal view; J, FSAC BOUJ-406, posterior fragment of a left mandibular ramus in dorsal view; K-P, Gavialoidea indet.: K, FSAC BOUJ-405, portion of a left dentary in dorsal view; L, FSAC BOUJ-402, mid portion of a right maxilla in ventral view; M, N, FSAC BOUJ-404, FSAC BOUJ-403 and FSAC BOUJ-401, anterior (M) and mid portion (N) of left maxilla in ventral views; O, P, FSAC BOUJ-407, posterior portion of a left maxilla in lateral (O) and ventral (P) views. Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4605979" httpUri="https://zenodo.org/record/4605979/files/figure.png" pageId="19" pageNumber="138">Fig. 7</figureCitation>
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L-N). The snout is wider than high and the palate is lower than the tooth row, so that the tooth row is underlined. The diameter of the alveoli is nearly constant along the tooth row and the interalveolus distances are equal or slightly longer than the alveolus diameter. The lateral margin of the maxilla is marked with shallow grooves visible in dorsal view for the occlusion of dentary teeth. FSAC Bouj-405 consists of a left portion of dentary, (
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<figureCitation id="130091D6FF92FF9B3154F8CDFE0F1142" box="[332,410,1810,1836]" captionStart="FIG" captionStartId="20.[132,143,1407,1424]" captionTargetBox="[132,1455,215,1365]" captionTargetId="figure-294@20.[698,1267,789,1366]" captionTargetPageId="20" captionText="FIG. 7. — Crocodyliformes remains from Gueran: A-J, Eusuchia indet.: A, FSAC BOUJ-410, last cervical (ninth) vertebra in left lateral view; B, FSAC BOUJ- 1b, anterior cervical vertebra in left lateral view; C, FSAC BOUJ-400, posterior dorsal vertebra in left lateral view; D, FSAC BOUJ-1a, first caudal vertebra in left lateral view; E, FSAC BOUJ-124, caudal vertebra in left lateral view; F, FSAC BOUJ-94, osteoderm in dorsal view; G, FSAC BOUJ-96, posterior portion of a dorsal osteoderm; H, FSAC BOUJ-96, lateral portion of a dorsal osteoderm in dorsal views; I, J, Crocodyliformes indet.: I, FSAC BOUJ-355, mid portion of a dentary in dorsal view; J, FSAC BOUJ-406, posterior fragment of a left mandibular ramus in dorsal view; K-P, Gavialoidea indet.: K, FSAC BOUJ-405, portion of a left dentary in dorsal view; L, FSAC BOUJ-402, mid portion of a right maxilla in ventral view; M, N, FSAC BOUJ-404, FSAC BOUJ-403 and FSAC BOUJ-401, anterior (M) and mid portion (N) of left maxilla in ventral views; O, P, FSAC BOUJ-407, posterior portion of a left maxilla in lateral (O) and ventral (P) views. Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4605979" httpUri="https://zenodo.org/record/4605979/files/figure.png" pageId="19" pageNumber="138">Fig. 7K</figureCitation>
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).Its lateral margin is marked by deep natural notches that indicate occlusal grooves for the maxillary teeth. The mandible was more than twice wider than high, and the symphysis was probably very long. Its morphology suggests than it is probably from the same species as the maxillary fragments. FSAC Bouj-407 is a fragment of the posterior portion of the left maxilla and with two complete teeth.
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</paragraph>
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<paragraph id="8B848D53FF92FF9B335BFF08FB83173F" blockId="19.[812,1455,215,337]" pageId="19" pageNumber="138">Teeth are preserved. They are moderately long, circular in cross section (posteriormost being slightly compressed lateromedially) and their surfaces are smooth and bear anterior and posterior carinae.</paragraph>
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</subSubSection>
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<subSubSection id="C321DED8FF92FF9B3335FEA8FA821043" pageId="19" pageNumber="138" type="discussion">
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<paragraph id="8B848D53FF92FF9B3335FEA8FBE017E1" blockId="19.[811,1460,375,1581]" box="[813,1141,375,401]" pageId="19" pageNumber="138">COMPARISON AND DISCUSSION</paragraph>
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<paragraph id="8B848D53FF92FF9B3335FE49FA821043" blockId="19.[811,1460,375,1581]" pageId="19" pageNumber="138">
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All recovered vertebrae are procoelous, suggesting eusuchian affinity. The material belongs to at least two species, and both are longirostrine forms. FSAC Bouj-355 has a different morphology from other mandibular and maxillary fragments (
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<figureCitation id="130091D6FF92FF9B332EFDC9FC11145F" box="[822,900,534,561]" captionStart="FIG" captionStartId="20.[132,143,1407,1424]" captionTargetBox="[132,1455,215,1365]" captionTargetId="figure-294@20.[698,1267,789,1366]" captionTargetPageId="20" captionText="FIG. 7. — Crocodyliformes remains from Gueran: A-J, Eusuchia indet.: A, FSAC BOUJ-410, last cervical (ninth) vertebra in left lateral view; B, FSAC BOUJ- 1b, anterior cervical vertebra in left lateral view; C, FSAC BOUJ-400, posterior dorsal vertebra in left lateral view; D, FSAC BOUJ-1a, first caudal vertebra in left lateral view; E, FSAC BOUJ-124, caudal vertebra in left lateral view; F, FSAC BOUJ-94, osteoderm in dorsal view; G, FSAC BOUJ-96, posterior portion of a dorsal osteoderm; H, FSAC BOUJ-96, lateral portion of a dorsal osteoderm in dorsal views; I, J, Crocodyliformes indet.: I, FSAC BOUJ-355, mid portion of a dentary in dorsal view; J, FSAC BOUJ-406, posterior fragment of a left mandibular ramus in dorsal view; K-P, Gavialoidea indet.: K, FSAC BOUJ-405, portion of a left dentary in dorsal view; L, FSAC BOUJ-402, mid portion of a right maxilla in ventral view; M, N, FSAC BOUJ-404, FSAC BOUJ-403 and FSAC BOUJ-401, anterior (M) and mid portion (N) of left maxilla in ventral views; O, P, FSAC BOUJ-407, posterior portion of a left maxilla in lateral (O) and ventral (P) views. Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4605979" httpUri="https://zenodo.org/record/4605979/files/figure.png" pageId="19" pageNumber="138">Fig. 7I</figureCitation>
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). It has large alveoli and its symphysis is slightly wider than high. The second form, represented by maxillae and portions of left and right dentaries (
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<figureCitation id="130091D6FF92FF9B34FBFD89FAB4141E" box="[1251,1313,598,624]" captionStart="FIG" captionStartId="20.[132,143,1407,1424]" captionTargetBox="[132,1455,215,1365]" captionTargetId="figure-294@20.[698,1267,789,1366]" captionTargetPageId="20" captionText="FIG. 7. — Crocodyliformes remains from Gueran: A-J, Eusuchia indet.: A, FSAC BOUJ-410, last cervical (ninth) vertebra in left lateral view; B, FSAC BOUJ- 1b, anterior cervical vertebra in left lateral view; C, FSAC BOUJ-400, posterior dorsal vertebra in left lateral view; D, FSAC BOUJ-1a, first caudal vertebra in left lateral view; E, FSAC BOUJ-124, caudal vertebra in left lateral view; F, FSAC BOUJ-94, osteoderm in dorsal view; G, FSAC BOUJ-96, posterior portion of a dorsal osteoderm; H, FSAC BOUJ-96, lateral portion of a dorsal osteoderm in dorsal views; I, J, Crocodyliformes indet.: I, FSAC BOUJ-355, mid portion of a dentary in dorsal view; J, FSAC BOUJ-406, posterior fragment of a left mandibular ramus in dorsal view; K-P, Gavialoidea indet.: K, FSAC BOUJ-405, portion of a left dentary in dorsal view; L, FSAC BOUJ-402, mid portion of a right maxilla in ventral view; M, N, FSAC BOUJ-404, FSAC BOUJ-403 and FSAC BOUJ-401, anterior (M) and mid portion (N) of left maxilla in ventral views; O, P, FSAC BOUJ-407, posterior portion of a left maxilla in lateral (O) and ventral (P) views. Scale bar: 1 cm." figureDoi="http://doi.org/10.5281/zenodo.4605979" httpUri="https://zenodo.org/record/4605979/files/figure.png" pageId="19" pageNumber="138">Fig. 7</figureCitation>
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K-P), has its symphysis much wider than high with smaller alveoli than the first species.Two groups of longirostrine eusuchians have been described from the late Eocene: the gavialoids and the tomistomines (
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<bibRefCitation id="EFAAF0A2FF92FF9B3382FD0AFBB3149E" author="BROCHU C. A." box="[922,1062,725,752]" pageId="19" pageNumber="138" pagination="357 - 397" refId="ref20017" refString="BROCHU C. A. 2003. - Phylogenetic approaches toward crocodylian history. Annual Review of Earth and Planetary Sciences 31: 357 - 397. https: // doi. org / 10.1146 / annurev. earth. 31.100901.141308" type="journal article" year="2003">Brochu 2003</bibRefCitation>
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). FSAC Bouj-355 is too fragmentary to be attributed with certainty to any group, but its symphysis slightly wider than high with a straight lateral margin, the short distance between the left and right alveoli and their offset margins clearly differ from what is found in gavialoids and tomistomines. Even if no amphycoelous vertebra has been found, it cannot be excluded that this mandible pertains to a dyrosaurid, a group of non-eusuchian
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<taxonomicName id="4C3BF6D0FF92FF9B34E2FC6AFA2415A1" authorityName="Hay, 1930 sensu Benton & Clark" authorityYear="1988" box="[1274,1457,949,975]" order="Crocodyliformes" pageId="19" pageNumber="137" rank="order">crocodyliformes</taxonomicName>
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, in which previously cited characters are present (
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<bibRefCitation id="EFAAF0A2FF92FF9B3521FC0AFCF01261" author="JOUVE S. & KHALLOUFI B. & ZOUHRI S." pageId="19" pageNumber="138" pagination="1617723" refId="ref22266" refString="JOUVE S., KHALLOUFI B. & ZOUHRI S. 2019. - New Longirostrine crocodyliforms from the Bartonian of Morocco and Paleogene climatic and sea-level oscillations in peri-Tethys area. Journal of Vertebrate Paleontology 39 (3): e 1617723 https: // doi. org / 10. 1080 / 02724634.2019.1617723" type="journal article" year="2019">
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Jouve
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<emphasis id="B94F5141FF92FF9B3564FC0AFA251580" box="[1404,1456,980,1006]" italics="true" pageId="19" pageNumber="138">et al.</emphasis>
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2019
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</bibRefCitation>
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). These neosuchians survived to the Lutetian in Africa and
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<collectingCountry id="F32CCDC3FF92FF9B3341FBCAFC341241" box="[857,929,1045,1071]" name="Myanmar" pageId="19" pageNumber="138">Burma</collectingCountry>
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(
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<bibRefCitation id="EFAAF0A2FF92FF9B33B7FBCBFBDB1240" author="BUFFETAUT E." box="[943,1102,1044,1070]" pageId="19" pageNumber="138" pagination="273 - 281" refId="ref20102" refString="BUFFETAUT E. 1978. - A dyrosaurid (Crocodylia, Mesosuchia) from the Upper Eocene of Burma. Neues Jahrbuch Fur Geologie Und Palaontologie, Monatshefte 5: 273 - 281." type="journal article" year="1978">Buffetaut 1978</bibRefCitation>
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). Awaiting more diagnostic material, FSAC Bouj-355 is thus considered as
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<taxonomicName id="4C3BF6D0FF92FF9B34E2FBEBFA3A1220" authorityName="Hay, 1930 sensu Benton & Clark" authorityYear="1988" box="[1274,1455,1076,1102]" order="Crocodyliformes" pageId="19" pageNumber="137" rank="order">Crocodyliformes</taxonomicName>
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indet. The second mandible has laterally opened alveoli and its alveolar margin is not leveled with the palate, characters that are found in gavialoids (
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<bibRefCitation id="EFAAF0A2FF92FF9B3478FB4BFAB812C0" author="HUA S. & JOUVE S." box="[1120,1325,1172,1198]" pageId="19" pageNumber="138" pagination="344 - 353" refId="ref21960" refString="HUA S. & JOUVE S. 2004. - A primitive marine gavialoid from the Paleocene of Morocco. Journal of Vertebrate Paleontology 24: 344 - 353. https: // doi. org / 10.1671 / 1104" type="journal article" year="2004">Hua & Jouve 2004</bibRefCitation>
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;
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<bibRefCitation id="EFAAF0A2FF92FF9B3523FB4BFCF012A0" author="JOUVE S. & IAROCHENE M. & BOUYA B. & AMAGHZAZ M." pageId="19" pageNumber="138" pagination="817 - 832" refId="ref22075" refString="JOUVE S., IAROCHENE M., BOUYA B. & AMAGHZAZ M. 2006. - New material of Argochampsa krebsi (Eusuchia: Gavialoidea) and phylogenetic implications. Geobios 39 (6): 817 - 832. https: // doi. org / 10.1016 / j. geobios. 2005.07.003" type="journal article" year="2006">
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Jouve
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<emphasis id="B94F5141FF92FF9B3565FB4AFA2512C0" box="[1405,1456,1172,1198]" italics="true" pageId="19" pageNumber="138">et al.</emphasis>
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2006
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</bibRefCitation>
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,
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<bibRefCitation id="EFAAF0A2FF92FF9B3369FB6BFC3C12A0" author="JOUVE S. & BOUYA B. & AMAGHZAZ M. & MESLOUH S." box="[881,937,1204,1230]" pageId="19" pageNumber="138" pagination="1 - 25" refId="ref22203" refString="JOUVE S., BOUYA B., AMAGHZAZ M. & MESLOUH S. 2014. - Maroccosuchus zennaroi (Crocodylia: Tomistominae) from the Eocene of Morocco: phylogenetic and palaeobiogeographical implications of the basalmost tomistomine. Journal of Systematic Palaeontology 12: 1 - 25. https: // doi. org / 10.1080 / 1477201 9.2014. 913078" type="journal article" year="2014">2014</bibRefCitation>
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). Gavialoids are particularly scarce in the Eocene and Oligocene of the Peri-Tethys deposits, and only three gavialoids are known: “
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<taxonomicName id="4C3BF6D0FF92FF9B3434FB2CFB171363" authorityName="Oppel" authorityYear="1811" box="[1068,1154,1267,1293]" class="Reptilia" family="Gavialidae" genus="Gavialis" kingdom="Animalia" order="Crocodylia" pageId="19" pageNumber="138" phylum="Chordata" rank="genus">
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<emphasis id="B94F5141FF92FF9B3434FB2CFB171363" box="[1068,1154,1267,1293]" italics="true" pageId="19" pageNumber="138">Gavialis</emphasis>
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</taxonomicName>
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”
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<emphasis id="B94F5141FF92FF9B348EFB2CFB4D1363" box="[1174,1240,1267,1293]" italics="true" pageId="19" pageNumber="138">dixoni</emphasis>
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Owen, 1849, from the early-middle Eocene of
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<collectingCountry id="F32CCDC3FF92FF9B3470FACCFB501343" box="[1128,1221,1299,1325]" name="United Kingdom" pageId="19" pageNumber="138">England</collectingCountry>
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and now considered as a
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<emphasis id="B94F5141FF92FF9B3347FAEBFC6A1323" box="[863,1023,1331,1357]" italics="true" pageId="19" pageNumber="138">nomen dubium</emphasis>
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(
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<bibRefCitation id="EFAAF0A2FF92FF9B3417FAECFB0A1323" author="BROCHU C. A." box="[1039,1183,1331,1357]" pageId="19" pageNumber="138" pagination="917 - 928" refId="ref20058" refString="BROCHU C. A. 2007. - Systematics and taxonomy of Eocene tomistomine crocodylians from Britain and Northern Europe. Palaeontology 50: 917 - 928. https: // doi. org / 10.1111 / j. 1475 - 4983.2007.00679. x" type="journal article" year="2007">Brochu 2007</bibRefCitation>
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), unidentified Bartonian gavialoid remains from Dur At-Talah (Southern
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<collectingCountry id="F32CCDC3FF92FF9B352EFA8CFAE11300" box="[1334,1396,1363,1390]" name="Libya" pageId="19" pageNumber="138">Libya</collectingCountry>
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) (
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<bibRefCitation id="EFAAF0A2FF92FF9B3591FA8CFC7513E3" author="LLINAS AGRASAR E. L." pageId="19" pageNumber="138" pagination="209 - 222" refId="ref22865" refString="LLINAS AGRASAR E. L. 2004. - Crocodilian remains from the upper Eocene of Dor-El-Talha, Libya. Annales de Paleontologie 90: 209 - 222. https: // doi. org / 10.1016 / j. annpal. 2004.05.001" type="journal article" year="2004">Llinas Agrasar 2004</bibRefCitation>
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), and
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<taxonomicName id="4C3BF6D0FF92FF9B3438FAACFA3A13E3" authority="(Andrews, 1901)" baseAuthorityName="Andrews" baseAuthorityYear="1901" box="[1056,1455,1395,1421]" class="Reptilia" family="Gavialidae" genus="Eogavialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" pageNumber="138" phylum="Chordata" rank="species" species="africanum">
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<emphasis id="B94F5141FF92FF9B3438FAACFB6313E3" box="[1056,1270,1395,1421]" italics="true" pageId="19" pageNumber="138">Eogavialis africanum</emphasis>
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(
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<bibRefCitation id="EFAAF0A2FF92FF9B351CFAACFA3213E3" author="ANDREWS C. W." box="[1284,1447,1395,1421]" pageId="19" pageNumber="138" pagination="436 - 444" refId="ref18994" refString="ANDREWS C. W. 1901. - Preliminary note on some recently discovered extinct vertebrates from Egypt. Geological Magazine 4: 436 - 444. https: // doi. org / 10.1017 / S 0016756800179750" type="journal article" year="1901">Andrews, 1901</bibRefCitation>
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)
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</taxonomicName>
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from the Priabonian and Rupelian of Fayum
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<collectingCountry id="F32CCDC3FF92FF9B3516FA4BFAC413C0" box="[1294,1361,1428,1454]" name="Egypt" pageId="19" pageNumber="138">Egypt</collectingCountry>
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, (
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<bibRefCitation id="EFAAF0A2FF92FF9B357EFA4CFCF113A3" author="MULLER L." pageId="19" pageNumber="138" pagination="1 - 96" refId="ref24057" refString="MULLER L. 1927. - Ergebnisse der Forschungsreisen Prof. E. Stromers in den Wusten Agyptens, V. Tertiare Wirbeltiere, 1. Beitrage zur Kenntnis der Krokodilier des agyptischen Tertiars. Abhandlungen der Bayerischen Akademie der Wissenschaften, mathematisch-naturwissenschaftliche Abteilung 31 (2): 1 - 96." type="journal article" year="1927">Müller 1927</bibRefCitation>
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). The remains from Gueran strongly resemble
|
|
<taxonomicName id="4C3BF6D0FF92FF9B355FFA6CFC0D1383" baseAuthorityName="Andrews" baseAuthorityYear="1901" class="Reptilia" family="Gavialidae" genus="Eogavialis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Crocodylia" pageId="19" pageNumber="138" phylum="Chordata" rank="species" species="africanum">
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<emphasis id="B94F5141FF92FF9B355FFA6CFC0D1383" italics="true" pageId="19" pageNumber="138">Eogavialis africanum</emphasis>
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</taxonomicName>
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|
, but they are too poorly preserved for an in-depth comparison. Therefore, the Gueran gavialoid is here considered as
|
|
<taxonomicName id="4C3BF6D0FF92FF9B3364F9CDFC6E1042" authorityName="Hay" authorityYear="1930" box="[892,1019,1554,1580]" kingdom="Animalia" pageId="19" pageNumber="138" rank="superFamily" superFamily="Gavialoidea">Gavialoidea</taxonomicName>
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indet. (
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<bibRefCitation id="EFAAF0A2FF92FF9B344AF9CCFA921043" author="JOUVE S. & KHALLOUFI B. & ZOUHRI S." box="[1106,1287,1554,1581]" pageId="19" pageNumber="138" pagination="1617723" refId="ref22266" refString="JOUVE S., KHALLOUFI B. & ZOUHRI S. 2019. - New Longirostrine crocodyliforms from the Bartonian of Morocco and Paleogene climatic and sea-level oscillations in peri-Tethys area. Journal of Vertebrate Paleontology 39 (3): e 1617723 https: // doi. org / 10. 1080 / 02724634.2019.1617723" type="journal article" year="2019">
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Jouve
|
|
<emphasis id="B94F5141FF92FF9B348DF9CCFB5D1042" box="[1173,1224,1554,1580]" italics="true" pageId="19" pageNumber="138">et al.</emphasis>
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2019
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</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |