treatments-xml/data/03/92/3C/03923C45FF87FF8E30A4FAACFBB6145E.xml
2024-06-21 12:22:17 +02:00

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<document id="BBF86B5BA6361F1512D92D51E3BC5550" ID-CLB-Dataset="7746" ID-DOI="10.5252/geodiversitas2021v43a5" ID-GBIF-Dataset="e42658a2-0e15-4e90-9bc5-0d00cbcbd781" ID-ISSN="1638-9395" ID-Zenodo-Dep="4605963" ID-ZooBank="urn:lsid:zoobank.org:pub:697FC553-E37B-4EF9-97A4-950E4DEE246C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1615828395930" checkinUser="felipe" docAuthor="Zouhri, Samir, Gingerich, Philip D., Khalloufi, Bouziane, Bourdon, Estelle, Adnet, Sylvain, Jouve, Stéphane, Elboudali, Najia, Amane, Ayoub, Rage, Jean-Claude &amp; Tabuce, Rodolphe" docDate="2021" docId="03923C45FF87FF8E30A4FAACFBB6145E" docLanguage="en" docName="Geodiversitas.43.5.121-150.pdf" docOrigin="Geodiversitas 43 (5)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Otodus (Carcharocles) sokolowi" docType="treatment" docVersion="9" lastPageNumber="125" masterDocId="FFAB443DFF81FF883018FFDFFF95166E" masterDocTitle="Middle Eocene vertebrate fauna from the Aridal Formation, Sabkha of Gueran, southwestern Morocco" masterLastPageNumber="150" masterPageNumber="121" pageNumber="125" updateTime="1698933443279" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="DCEED4B5C17086D08BAD5E53FBA0032E">Middle Eocene vertebrate fauna from the Aridal Formation, Sabkha of Gueran, southwestern Morocco</mods:title>
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<mods:namePart id="9B9C1C88F3C5C19415A957FAF414DA23">Gingerich, Philip D.</mods:namePart>
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<mods:namePart id="67FEA0F03E428637E7BC4090A7E795E9">Khalloufi, Bouziane</mods:namePart>
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<mods:namePart id="E67B6FFAF7147CB622D3B90C7299450C">Bourdon, Estelle</mods:namePart>
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<mods:namePart id="D652416D992D25ADECAB678D8081D8F2">Jouve, Stéphane</mods:namePart>
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<mods:namePart id="B3803A7BD6A1604AB2EE3B59370A7B22">Elboudali, Najia</mods:namePart>
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<mods:namePart id="83B617754BD008537B125E26B47464A5">Rage, Jean-Claude</mods:namePart>
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<mods:title id="9C360CE507097B1068CD79231B79BE34">Geodiversitas</mods:title>
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<mods:date id="1F037B1A6DEBC175D2ED3A8840EA5047">2021</mods:date>
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<mods:number id="25156269E6C54AFD371C0599DD2A5E5B">2021-03-11</mods:number>
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<mods:number id="B52709F39DD26631C22072CF4FBDD165">43</mods:number>
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<treatment id="03923C45FF87FF8E30A4FAACFBB6145E" ID-DOI="http://doi.org/10.5281/zenodo.4606609" ID-GBIF-Taxon="180569480" ID-Zenodo-Dep="4606609" LSID="urn:lsid:plazi:treatment:03923C45FF87FF8E30A4FAACFBB6145E" httpUri="http://treatment.plazi.org/id/03923C45FF87FF8E30A4FAACFBB6145E" lastPageNumber="125" pageId="6" pageNumber="125">
<subSubSection id="C321DED8FF87FF8E30A4FAACFD5A13E0" box="[188,719,1395,1422]" pageId="6" pageNumber="125" type="nomenclature">
<paragraph id="8B848D53FF87FF8E30A4FAACFD5A13E0" blockId="6.[188,719,1395,1422]" box="[188,719,1395,1422]" pageId="6" pageNumber="125">
<heading id="D0CC3A3FFF87FF8E30A4FAACFD5A13E0" box="[188,719,1395,1422]" centered="true" fontSize="11" level="2" pageId="6" pageNumber="125" reason="2">
<taxonomicName id="4C3BF6D0FF87FF8E30A4FAACFD5A13E0" authority="(Jaekel, 1895)" baseAuthorityName="Jaekel" baseAuthorityYear="1895" box="[188,719,1395,1422]" class="Elasmobranchii" family="Lamnidae" genus="Otodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lamniformes" pageId="6" pageNumber="125" phylum="Chordata" rank="species" species="sokolowi" subGenus="Carcharocles">
<emphasis id="B94F5141FF87FF8E30A4FAACFE9F13E0" bold="true" box="[188,266,1395,1422]" italics="true" pageId="6" pageNumber="125">Otodus</emphasis>
(
<emphasis id="B94F5141FF87FF8E3102FAACFE3613E0" bold="true" box="[282,419,1395,1422]" italics="true" pageId="6" pageNumber="125">Carcharocles</emphasis>
) cf.
<emphasis id="B94F5141FF87FF8E31CDFAACFDA413E0" bold="true" box="[469,561,1395,1422]" italics="true" pageId="6" pageNumber="125">sokolowi</emphasis>
(Jaekel, 1895)
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</heading>
</paragraph>
</subSubSection>
<subSubSection id="C321DED8FF87FF8E309CFA72FD3813B1" pageId="6" pageNumber="125" type="materials_examined">
<paragraph id="8B848D53FF87FF8E309CFA72FD3813B1" blockId="6.[132,775,1452,1503]" pageId="6" pageNumber="125">
EXAMINED MATERIAL. —
<materialsCitation id="3B53870EFF87FF8E3190FA73FD3C13B1" ID-GBIF-Occurrence="3053068320" pageId="6" pageNumber="125">Around twenty broken isolated teeth, figured material includes FSAC Bouj-320, 321 and 322</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C321DED8FF87FF8E309CF9CCFEB81082" pageId="6" pageNumber="125" type="description">
<paragraph id="8B848D53FF87FF8E309CF9CCFE801045" blockId="6.[132,776,1555,1772]" box="[132,277,1555,1581]" pageId="6" pageNumber="125">DESCRIPTION</paragraph>
<paragraph id="8B848D53FF87FF8E309CF9EDFEB81082" blockId="6.[132,776,1555,1772]" pageId="6" pageNumber="125">
Teeth can reach up to
<quantity id="4CC320B6FF87FF8E3199F9ECFE5D1023" box="[385,456,1587,1613]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="6" pageNumber="125" unit="cm" value="10.0">10 cm</quantity>
in height, displaying a large triangular cusp with well-marked and regular serrations on the cutting edges and a pair of lateral cusplets (
<figureCitation id="130091D6FF87FF8E32BBF9ADFD6E10E2" box="[675,763,1650,1676]" captionStart="FIG" captionStartId="7.[132,143,1872,1889]" captionTargetBox="[132,1455,215,1848]" captionTargetId="figure-15@7.[132,1455,215,1869]" captionTargetPageId="7" captionText="FIG. 2. — Elasmobranchii Lamniformes teeth remains from Sabkha of Gueran collected around Archaeocete whale carcasses: A-C, Otodus (Carcharocles) cf. sokolowi: A, FSAC Bouj-320 lateral upper tooth in (A1) labial and (A2) lingual views; B, FSAC Bouj-321 anterior lower tooth in labial view; C, FSAC Bouj-322 lateral lower tooth in labial view; D, E, Otodus (Carcharocles) sp.: D, FSAC Bouj-323 antero-lateral tooth in (D1) labial and (D2) lingual views; E, FSAC Bouj-324 lateral tooth (E1) labial and (E2) lingual views; F, G, “Carcharias” koerti; F, FSAC Bouj-325 anterolateral upper tooth in (F1) labial and (F2) lingual views; G, FSAC Bouj-326 anterior tooth of juvenile (G1) labial and (G2) lingual views; H-K, Macrorhizodus praecursor: H, FSAC Bouj-327 anterior lower tooth (H1) labial and (H2) lingual views; I, FSAC Bouj-328 anterior upper tooth in (I1) labial and (I2) lingual views; J, FSAC Bouj-329 lateral upper tooth in (J1) labial and (J2) lingual views; K, FSAC Bouj-330 lateral lower tooth (K1) labial and (K2) lingual views. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4605967" httpUri="https://zenodo.org/record/4605967/files/figure.png" pageId="6" pageNumber="125">Fig. 2A</figureCitation>
). Cusplets are not very high when conserved and often divergent in lateral teeth (
<figureCitation id="130091D6FF87FF8E3175F96DFE5510A2" box="[365,448,1714,1740]" captionStart="FIG" captionStartId="7.[132,143,1872,1889]" captionTargetBox="[132,1455,215,1848]" captionTargetId="figure-15@7.[132,1455,215,1869]" captionTargetPageId="7" captionText="FIG. 2. — Elasmobranchii Lamniformes teeth remains from Sabkha of Gueran collected around Archaeocete whale carcasses: A-C, Otodus (Carcharocles) cf. sokolowi: A, FSAC Bouj-320 lateral upper tooth in (A1) labial and (A2) lingual views; B, FSAC Bouj-321 anterior lower tooth in labial view; C, FSAC Bouj-322 lateral lower tooth in labial view; D, E, Otodus (Carcharocles) sp.: D, FSAC Bouj-323 antero-lateral tooth in (D1) labial and (D2) lingual views; E, FSAC Bouj-324 lateral tooth (E1) labial and (E2) lingual views; F, G, “Carcharias” koerti; F, FSAC Bouj-325 anterolateral upper tooth in (F1) labial and (F2) lingual views; G, FSAC Bouj-326 anterior tooth of juvenile (G1) labial and (G2) lingual views; H-K, Macrorhizodus praecursor: H, FSAC Bouj-327 anterior lower tooth (H1) labial and (H2) lingual views; I, FSAC Bouj-328 anterior upper tooth in (I1) labial and (I2) lingual views; J, FSAC Bouj-329 lateral upper tooth in (J1) labial and (J2) lingual views; K, FSAC Bouj-330 lateral lower tooth (K1) labial and (K2) lingual views. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4605967" httpUri="https://zenodo.org/record/4605967/files/figure.png" pageId="6" pageNumber="125">Fig. 2A</figureCitation>
) to less developed in anterior teeth (
<figureCitation id="130091D6FF87FF8E30D4F90DFE8A1082" box="[204,287,1746,1772]" captionStart="FIG" captionStartId="7.[132,143,1872,1889]" captionTargetBox="[132,1455,215,1848]" captionTargetId="figure-15@7.[132,1455,215,1869]" captionTargetPageId="7" captionText="FIG. 2. — Elasmobranchii Lamniformes teeth remains from Sabkha of Gueran collected around Archaeocete whale carcasses: A-C, Otodus (Carcharocles) cf. sokolowi: A, FSAC Bouj-320 lateral upper tooth in (A1) labial and (A2) lingual views; B, FSAC Bouj-321 anterior lower tooth in labial view; C, FSAC Bouj-322 lateral lower tooth in labial view; D, E, Otodus (Carcharocles) sp.: D, FSAC Bouj-323 antero-lateral tooth in (D1) labial and (D2) lingual views; E, FSAC Bouj-324 lateral tooth (E1) labial and (E2) lingual views; F, G, “Carcharias” koerti; F, FSAC Bouj-325 anterolateral upper tooth in (F1) labial and (F2) lingual views; G, FSAC Bouj-326 anterior tooth of juvenile (G1) labial and (G2) lingual views; H-K, Macrorhizodus praecursor: H, FSAC Bouj-327 anterior lower tooth (H1) labial and (H2) lingual views; I, FSAC Bouj-328 anterior upper tooth in (I1) labial and (I2) lingual views; J, FSAC Bouj-329 lateral upper tooth in (J1) labial and (J2) lingual views; K, FSAC Bouj-330 lateral lower tooth (K1) labial and (K2) lingual views. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4605967" httpUri="https://zenodo.org/record/4605967/files/figure.png" pageId="6" pageNumber="125">Fig. 2B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C321DED8FF87FF8E309CF8CDFBB6145E" pageId="6" pageNumber="125" type="discussion">
<paragraph id="8B848D53FF87FF8E309CF8CDFF7D1144" blockId="6.[131,775,1810,2027]" box="[132,232,1810,1836]" pageId="6" pageNumber="125">REMARKS</paragraph>
<paragraph id="8B848D53FF87FF8E309CF8EEFBB6145E" blockId="6.[131,775,1810,2027]" lastBlockId="6.[811,1457,215,560]" pageId="6" pageNumber="125">
Teeth of
<taxonomicName id="4C3BF6D0FF87FF8E30E8F8EEFDFB1125" baseAuthorityName="Jaekel" baseAuthorityYear="1895" box="[240,622,1841,1868]" class="Elasmobranchii" family="Lamnidae" genus="Otodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lamniformes" pageId="6" pageNumber="125" phylum="Chordata" rank="species" species="sokolowi" subGenus="Carcharocles">
<emphasis id="B94F5141FF87FF8E30E8F8EEFEAB1125" box="[240,318,1841,1867]" italics="true" pageId="6" pageNumber="125">Otodus</emphasis>
(
<emphasis id="B94F5141FF87FF8E3157F8EEFE421125" box="[335,471,1841,1867]" italics="true" pageId="6" pageNumber="125">Carcharocles</emphasis>
) cf.
<emphasis id="B94F5141FF87FF8E320BF8EEFDFB1125" box="[531,622,1841,1867]" italics="true" pageId="6" pageNumber="125">sokolowi</emphasis>
</taxonomicName>
are relatively common around the archaeocete carcasses (
<figureCitation id="130091D6FF87FF8E329AF88EFD501102" box="[642,709,1873,1900]" captionStart="FIG" captionStartId="7.[132,143,1872,1889]" captionTargetBox="[132,1455,215,1848]" captionTargetId="figure-15@7.[132,1455,215,1869]" captionTargetPageId="7" captionText="FIG. 2. — Elasmobranchii Lamniformes teeth remains from Sabkha of Gueran collected around Archaeocete whale carcasses: A-C, Otodus (Carcharocles) cf. sokolowi: A, FSAC Bouj-320 lateral upper tooth in (A1) labial and (A2) lingual views; B, FSAC Bouj-321 anterior lower tooth in labial view; C, FSAC Bouj-322 lateral lower tooth in labial view; D, E, Otodus (Carcharocles) sp.: D, FSAC Bouj-323 antero-lateral tooth in (D1) labial and (D2) lingual views; E, FSAC Bouj-324 lateral tooth (E1) labial and (E2) lingual views; F, G, “Carcharias” koerti; F, FSAC Bouj-325 anterolateral upper tooth in (F1) labial and (F2) lingual views; G, FSAC Bouj-326 anterior tooth of juvenile (G1) labial and (G2) lingual views; H-K, Macrorhizodus praecursor: H, FSAC Bouj-327 anterior lower tooth (H1) labial and (H2) lingual views; I, FSAC Bouj-328 anterior upper tooth in (I1) labial and (I2) lingual views; J, FSAC Bouj-329 lateral upper tooth in (J1) labial and (J2) lingual views; K, FSAC Bouj-330 lateral lower tooth (K1) labial and (K2) lingual views. Scale bars: 10 mm." figureDoi="http://doi.org/10.5281/zenodo.4605967" httpUri="https://zenodo.org/record/4605967/files/figure.png" pageId="6" pageNumber="125">Fig. 2</figureCitation>
A-C).
<bibRefCitation id="EFAAF0A2FF87FF8E309CF8AEFE1C11E2" author="CASE G. R. &amp; CAPPETTA H." box="[132,393,1905,1932]" pageId="6" pageNumber="125" pagination="1 - 30" refId="ref20466" refString="CASE G. R. &amp; CAPPETTA H. 1990. - The Eocene Selachians Fauna from the Fayum Depression in Egypt. Palaeontographica Abteilung A 212: 1 - 30." type="journal article" year="1990">Case &amp; Cappetta (1990)</bibRefCitation>
have discussed about the taxonomic distinctness of
<taxonomicName id="4C3BF6D0FF87FF8E3131F84EFDE011C5" baseAuthorityName="Jaekel" baseAuthorityYear="1895" box="[297,629,1937,1964]" class="Elasmobranchii" family="Lamnidae" genus="Otodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lamniformes" pageId="6" pageNumber="125" phylum="Chordata" rank="species" species="sokolowi" subGenus="Carcharocles">
<emphasis id="B94F5141FF87FF8E3131F84EFEE011C5" box="[297,373,1937,1963]" italics="true" pageId="6" pageNumber="125">Otodus</emphasis>
(
<emphasis id="B94F5141FF87FF8E319EF84EFD9911C5" box="[390,524,1937,1963]" italics="true" pageId="6" pageNumber="125">Carcharocles</emphasis>
)
<emphasis id="B94F5141FF87FF8E3203F84EFDE011C5" box="[539,629,1937,1963]" italics="true" pageId="6" pageNumber="125">sokolowi</emphasis>
</taxonomicName>
compared to the other Eocene species of subgenus, and in particularly with the smaller and coeval species
<taxonomicName id="4C3BF6D0FF87FF8E3229F80EFCCC169C" authority="(Carcharocles)" authorityName="Jordan &amp; Hannibal" authorityYear="1923" baseAuthorityName="Carcharocles" box="[561,857,215,2027]" class="Elasmobranchii" family="Lamnidae" genus="Otodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lamniformes" pageId="6" pageNumber="125" phylum="Chordata" rank="genus">
<emphasis id="B94F5141FF87FF8E3229F80EFD141185" box="[561,641,2001,2027]" italics="true" pageId="6" pageNumber="125">Otodus</emphasis>
(
<emphasis id="B94F5141FF87FF8E328EF80EFCC5169F" box="[662,848,215,2027]" italics="true" pageId="6" pageNumber="125">Carcharocles</emphasis>
)
</taxonomicName>
<taxonomicName id="4C3BF6D0FF87FF8E337BFF08FB21169F" authority="(Blainville, 1818)" baseAuthorityName="Blainville" baseAuthorityYear="1818" box="[867,1204,215,241]" class="Elasmobranchii" family="Lamnidae" genus="Otodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lamniformes" pageId="6" pageNumber="125" phylum="Chordata" rank="species" species="auriculatus">
<emphasis id="B94F5141FF87FF8E337BFF08FC49169F" box="[867,988,215,241]" italics="true" pageId="6" pageNumber="125">auriculatus</emphasis>
(Blainville, 1818)
</taxonomicName>
.
<taxonomicName id="4C3BF6D0FF87FF8E34D9FF08FC13177F" baseAuthorityName="Jaekel" baseAuthorityYear="1895" class="Elasmobranchii" family="Lamnidae" genus="Otodus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Lamniformes" pageId="6" pageNumber="125" phylum="Chordata" rank="species" species="sokolowi" subGenus="Carcharocles">
<emphasis id="B94F5141FF87FF8E34D9FF08FA9B169F" box="[1217,1294,215,241]" italics="true" pageId="6" pageNumber="125">Otodus</emphasis>
(
<emphasis id="B94F5141FF87FF8E3507FF08FA33169F" box="[1311,1446,215,241]" italics="true" pageId="6" pageNumber="125">Carcharocles</emphasis>
)
<emphasis id="B94F5141FF87FF8E3335FF28FC13177F" box="[813,902,247,273]" italics="true" pageId="6" pageNumber="125">sokolowi</emphasis>
</taxonomicName>
appears widely distributed and relatively common in the tropical marine realm since the GEA-C (uppermost Bartonian-lowermost Priabonian) and throughout the Priabonian successions (GE D-G, BQ, QS) according to
<bibRefCitation id="EFAAF0A2FF87FF8E3337FEA8FBD517FF" author="UNDERWOOD C. J. &amp; WARD D. J. &amp; KING C. &amp; ANTAR S. M. &amp; MOUT I. S. &amp; GINGERICH P. D." box="[815,1088,375,401]" pageId="6" pageNumber="125" pagination="47 - 66" refId="ref26062" refString="UNDERWOOD C. J., WARD D. J., KING C., ANTAR S. M., ZAL- MOUT I. S. &amp; GINGERICH P. D. 2011. - Shark and ray faunas in the middle and late Eocene of the Fayum area, Egypt. Proceedings of the Geologists' Association 122: 47 - 66. https: // doi. org / 10.1016 / j. pgeola. 20" type="journal article" year="2011">
Underwood
<emphasis id="B94F5141FF87FF8E33A0FEA8FC7917FF" box="[952,1004,375,401]" italics="true" pageId="6" pageNumber="125">et al.</emphasis>
(2011)
</bibRefCitation>
, southwestern
<collectingCountry id="F32CCDC3FF87FF8E34FCFEA8FADD17FF" box="[1252,1352,375,401]" name="Morocco" pageId="6" pageNumber="125">Morocco</collectingCountry>
included (
<bibRefCitation id="EFAAF0A2FF87FF8E332EFE49FC6C17DF" author="ADNET S. &amp; CAPPETTA H. &amp; TABUCE R." box="[822,1017,406,433]" pageId="6" pageNumber="125" pagination="860 - 870" refId="ref18939" refString="ADNET S., CAPPETTA H. &amp; TABUCE R. 2010. - A middle-late Eocene vertebrate fauna (marine fish and mammals) from southwestern Morocco preliminary report: age and palaeobiogeographical implications. Geological Magazine 147: 860 - 870. https: // doi. org / 10.1017 / s 0016756810000348" type="journal article" year="2010">
Adnet
<emphasis id="B94F5141FF87FF8E339AFE48FC2D17DE" box="[898,952,406,432]" italics="true" pageId="6" pageNumber="125">et al.</emphasis>
2010
</bibRefCitation>
). However this species seems relatively discrete in older deposits, as in the MI (
<bibRefCitation id="EFAAF0A2FF87FF8E34F5FE69FCF3179E" author="UNDERWOOD C. J. &amp; WARD D. J. &amp; KING C. &amp; ANTAR S. M. &amp; MOUT I. S. &amp; GINGERICH P. D." pageId="6" pageNumber="125" pagination="47 - 66" refId="ref26062" refString="UNDERWOOD C. J., WARD D. J., KING C., ANTAR S. M., ZAL- MOUT I. S. &amp; GINGERICH P. D. 2011. - Shark and ray faunas in the middle and late Eocene of the Fayum area, Egypt. Proceedings of the Geologists' Association 122: 47 - 66. https: // doi. org / 10.1016 / j. pgeola. 20" type="journal article" year="2011">
Underwood
<emphasis id="B94F5141FF87FF8E3562FE68FA2517BE" box="[1402,1456,438,464]" italics="true" pageId="6" pageNumber="125">et al.</emphasis>
2011
</bibRefCitation>
). Its occurrence in Bartonian of Gueran,
<collectingCountry id="F32CCDC3FF87FF8E355CFE08FA3E179F" box="[1348,1451,471,497]" name="Morocco" pageId="6" pageNumber="125">Morocco</collectingCountry>
, testify of its spatial expansion along the Tethysian coasts during the Bartonian.
</paragraph>
</subSubSection>
</treatment>
</document>