156 lines
15 KiB
XML
156 lines
15 KiB
XML
<document id="D5568D72DA818A55578FBC9A62708F47" ID-CLB-Dataset="42309" ID-DOI="10.11646/zootaxa.3878.5.4" ID-GBIF-Dataset="697c91ee-9d8b-4095-bd15-e55709b2b6bd" ID-ISSN="1175-5326" ID-Zenodo-Dep="229037" ID-ZooBank="377F9037-D039-44A5-B562-051560CBC9E0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460949457252" checkinUser="plazi" docAuthor="Coty, David, Háva, Jiří, Prokop, Jakub, Roques, Patrick & Nel, André" docDate="2014" docId="03923C218625FF8FFF45FE8400A1FBD7" docLanguage="en" docName="zt03878p470.pdf" docOrigin="Zootaxa 3878 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tococladus" docType="treatment" docVersion="9" lastPageNumber="464" masterDocId="FFAB44598627FF8DFFD2FFED015AFFFB" masterDocTitle="New archaeorthopteran insects from the Late Carboniferous of the Nord and Pasde-Calais basins in northern France (Insecta: Cnemidolestodea, Panorthoptera)" masterLastPageNumber="470" masterPageNumber="462" pageNumber="464" updateTime="1698585716371" updateUser="ExternalLinkService">
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<mods:mods id="8D82ED186156E500469F7627F5E86EDB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="8F9C633939954590099CF6D77364D333">
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<mods:title id="585A4DC61E305936FB9719DC07218F79">New archaeorthopteran insects from the Late Carboniferous of the Nord and Pasde-Calais basins in northern France (Insecta: Cnemidolestodea, Panorthoptera)</mods:title>
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<mods:name id="3E13911A2B0949D2B8FEE220C058879A" type="personal">
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<mods:roleTerm id="DC6C09C5B3170B05813CAE26647516BE">Author</mods:roleTerm>
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<mods:namePart id="E1168E52941B45BEEDE70FC0D5B8483A">Coty, David</mods:namePart>
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<mods:name id="8D64C3877F9E66D77ACB43EC50F86258" type="personal">
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<mods:roleTerm id="EC895C063C012FE9DFFE0E3FAD41EBEC">Author</mods:roleTerm>
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<mods:namePart id="498C45EAED9E5ED8D48646CCD1D8CF52">Háva, Jiří</mods:namePart>
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</mods:name>
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<mods:name id="A0786561DC9C1AFF85A31DE093B3FB63" type="personal">
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<mods:roleTerm id="328A7DB6EECB972FFC59DA440868DCE5">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="FA9B8E480C20D8E145D48C7D7820CE47">Prokop, Jakub</mods:namePart>
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<mods:name id="70F9920B3A865A1EEF9AAFFF46B88EFB" type="personal">
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<mods:roleTerm id="E077ED9D4292A93897736DD5C29B79F6">Author</mods:roleTerm>
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<mods:namePart id="9FC717B9F661DA5D36BDE9D5E4E52845">Roques, Patrick</mods:namePart>
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</mods:name>
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<mods:name id="0236F68E7042A8F151F841697AD50993" type="personal">
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<mods:roleTerm id="736FB8FAF7D03B08B968849578FE41E9">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="BE1FA6BBE2775767D63D192143CADAAE">Nel, André</mods:namePart>
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<mods:typeOfResource id="2B4BA174C24D34CFCC03CC629C57A89E">text</mods:typeOfResource>
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<mods:titleInfo id="5C5507DEEA1AF676FA99EAA90C7C1B54">
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<mods:title id="08280AA3C4C78E1FF5EF04B9145B8EAD">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="BE4C6BAF96FFAC5B32C8D352F0E06C67">
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<mods:date id="53DF8E3D046D3688966B480BCDCEDA6C">2014</mods:date>
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<mods:detail id="457A829446D5C35E39F3963C4A26DEE8" type="volume">
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<mods:number id="EEC50D3FE3FDA4237F95CEC5C3A25BC8">3878</mods:number>
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</mods:detail>
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<mods:detail id="8C707A7ACFBC5938EDE7DEC70A45DADE" type="issue">
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<mods:number id="ED9D8CD19C60A0AE9EF60C45C7DA5285">5</mods:number>
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<mods:start id="EF25DB73F20BE4FA77A1E1FB1ACA3B6F">462</mods:start>
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<mods:end id="C73A5A384732FAB901582478AE62936D">470</mods:end>
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<mods:classification id="65D8FFE42C0872AE00FCFF6EB3345AA1">journal article</mods:classification>
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<mods:identifier id="CDDF6CF899ABE55CD04FE0B37A0AF6DC" type="CLB-Dataset">42309</mods:identifier>
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<mods:identifier id="1DA966E32A1C218C0CC08B6302B2AD94" type="DOI">10.11646/zootaxa.3878.5.4</mods:identifier>
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<mods:identifier id="3EA23259F491E195C43CD747B3FB33C9" type="GBIF-Dataset">697c91ee-9d8b-4095-bd15-e55709b2b6bd</mods:identifier>
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<mods:identifier id="E2B150E286FC4939692AA72EF37FBDF5" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="E9DB1CBFF638010AA482E1997440821A" type="Zenodo-Dep">229037</mods:identifier>
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<mods:identifier id="6DD0408DF396B08F34649D1B1C880E37" type="ZooBank">377F9037-D039-44A5-B562-051560CBC9E0</mods:identifier>
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<treatment id="03923C218625FF8FFF45FE8400A1FBD7" ID-DOI="http://doi.org/10.5281/zenodo.6142659" ID-GBIF-Taxon="119591891" ID-Zenodo-Dep="6142659" LSID="urn:lsid:plazi:treatment:03923C218625FF8FFF45FE8400A1FBD7" httpUri="http://treatment.plazi.org/id/03923C218625FF8FFF45FE8400A1FBD7" lastPageNumber="464" pageId="2" pageNumber="464">
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<subSubSection id="C321DEBC8625FF8FFF45FE84033BFE78" box="[151,609,361,387]" pageId="2" pageNumber="464" type="nomenclature">
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<paragraph id="8B848D378625FF8FFF45FE84033BFE78" blockId="2.[151,609,361,421]" box="[151,609,361,387]" pageId="2" pageNumber="464">
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<heading id="D0CC3A5B8625FF8FFF45FE84033BFE78" bold="true" box="[151,609,361,387]" fontSize="11" level="1" pageId="2" pageNumber="464" reason="1">
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<emphasis id="B94F51258625FF8FFF45FE84033BFE78" bold="true" box="[151,609,361,387]" pageId="2" pageNumber="464">
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Cf.
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<taxonomicName id="4C3BF6B48625FF8FFF16FE840016FE78" box="[196,332,361,387]" class="Insecta" family="Tococladidae" genus="Tococladus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="464" phylum="Arthropoda" rank="genus">
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<emphasis id="B94F51258625FF8FFF16FE840016FE78" bold="true" box="[196,332,361,387]" italics="true" pageId="2" pageNumber="464">Tococladus</emphasis>
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</taxonomicName>
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species undetermined
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C321DEBC8625FF8FFF45FE6101B1FE5F" box="[151,235,396,421]" pageId="2" pageNumber="464" type="description">
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<paragraph id="8B848D378625FF8FFF45FE6101B1FE5F" blockId="2.[151,609,361,421]" box="[151,235,396,421]" pageId="2" pageNumber="464">
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(
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<figureCitation id="130091B28625FF8FFF4DFE6101BAFE5F" box="[159,224,396,421]" captionStart="FIGURE 1" captionStartId="1.[151,250,1786,1808]" captionTargetBox="[356,1230,483,1756]" captionTargetId="figure@1.[356,1231,483,1761]" captionTargetPageId="1" captionText="FIGURE 1. cf. Tococladus sp. specimen Avion 0 0 0 13 stored in MNHN coll. finally deposited in the Musée Géologique Pierre Vetter, Decazeville, France. A, photograph of wing apex venation; B, line drawing of apical part of wing. Scale bar = 5 mm." httpUri="https://zenodo.org/record/229038/files/figure.png" pageId="2" pageNumber="464">Fig. 1</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C321DEBC8625FF8FFF45FE39032DFDAC" pageId="2" pageNumber="464" type="materials_examined">
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<paragraph id="8B848D378625FF8FFF45FE3903FAFDF4" blockId="2.[151,1436,468,1069]" pageId="2" pageNumber="464">
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<emphasis id="B94F51258625FF8FFF45FE390030FE16" bold="true" box="[151,362,468,493]" pageId="2" pageNumber="464">Material studied:</emphasis>
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Specimen 0 0 13 (print and counterprint), coll. Patrick Roques, to be deposited in the Musée Géologique Pierre Vetter, Decazeville,
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<collectingCountry id="F32CCDA78625FF8FFD9EFE1503C6FDF4" box="[588,668,504,527]" name="France" pageId="2" pageNumber="464">France</collectingCountry>
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.
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</paragraph>
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<paragraph id="8B848D378625FF8FFF15FDF1032DFDAC" blockId="2.[151,1436,468,1069]" pageId="2" pageNumber="464">
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<emphasis id="B94F51258625FF8FFF15FDF100D6FDCE" bold="true" box="[199,396,540,565]" pageId="2" pageNumber="464">Age and outcrop</emphasis>
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.
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<typeStatus id="548033958625FF8FFE49FDF00089FDCE" box="[411,467,541,565]" pageId="2" pageNumber="464">Type</typeStatus>
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locality and horizon. Moscovian (Westphalian C/D equivalent to Bolsovian/Asturian), ‘Terril N°7’, Avion, Pas-de-Calais,
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<collectingCountry id="F32CCDA78625FF8FFDF6FDAD032EFDAC" box="[548,628,576,599]" name="France" pageId="2" pageNumber="464">France</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C321DEBC8625FF8FFF15FD890577FCF7" pageId="2" pageNumber="464" type="description">
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<paragraph id="8B848D378625FF8FFF15FD890577FCF7" blockId="2.[151,1436,468,1069]" pageId="2" pageNumber="464">
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<emphasis id="B94F51258625FF8FFF15FD89000BFD86" bold="true" box="[199,337,612,637]" pageId="2" pageNumber="464">Description</emphasis>
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. A fragment of wing corresponding to the anterior part of the apex, fragment
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<quantity id="4CC320D28625FF8FFB12FD89047EFD87" box="[1216,1316,612,636]" metricMagnitude="-2" metricUnit="m" metricValue="1.94" pageId="2" pageNumber="464" unit="mm" value="19.4">19.4 mm</quantity>
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long,
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<quantity id="4CC320D28625FF8FFABEFD890199FD64" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="2" pageNumber="464" unit="mm" value="11.7">11.7 mm</quantity>
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wide; convex RA with a series of sigmoidal anterior veinlets, RA distally fused with concave main branch of
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<collectingCountry id="F32CCDA78625FF8FFF45FD4001E3FD3F" box="[151,185,685,708]" name="Poland" pageId="2" pageNumber="464">RP</collectingCountry>
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, near wing apex, common branch
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<quantity id="4CC320D28625FF8FFDECFD4103CEFD3F" box="[574,660,684,709]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="464" unit="mm" value="2.6">2.6 mm</quantity>
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long; a series of straight crossveins in area between RA and
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<collectingCountry id="F32CCDA78625FF8FFA9AFD400434FD3F" box="[1352,1390,685,708]" name="Poland" pageId="2" pageNumber="464">RP</collectingCountry>
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;
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<collectingCountry id="F32CCDA78625FF8FFAA8FD4004C6FD3F" box="[1402,1436,685,708]" name="Poland" pageId="2" pageNumber="464">RP</collectingCountry>
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with seven to nine parallel posterior branches with a series of simple straight crossveins between them; apical parts of branches of M and possibly CuA also preserved, parallel to the branches of
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<collectingCountry id="F32CCDA78625FF8FFBD4FD180573FCF7" box="[1030,1065,757,780]" name="Poland" pageId="2" pageNumber="464">RP</collectingCountry>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C321DEBC8625FF8FFF15FCFA00A1FBD7" pageId="2" pageNumber="464" type="discussion">
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<paragraph id="8B848D378625FF8FFF15FCFA00A1FBD7" blockId="2.[151,1436,468,1069]" pageId="2" pageNumber="464">
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<emphasis id="B94F51258625FF8FFF15FCFA001FFCCB" bold="true" box="[199,325,791,816]" pageId="2" pageNumber="464">Discussion</emphasis>
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. Although very incomplete, this wing fragment fits quite well with the equivalent portion of a forewing of
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<taxonomicName id="4C3BF6B48625FF8FFEF8FCD103E9FCAE" authority="Carpenter, 1966" authorityName="Carpenter" authorityYear="1966" box="[298,691,828,853]" class="Insecta" family="Tococladidae" genus="Tococladus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="464" phylum="Arthropoda" rank="species" species="ralus">
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<emphasis id="B94F51258625FF8FFEF8FCD100B4FCAE" box="[298,494,828,853]" italics="true" pageId="2" pageNumber="464">Tococladus ralus</emphasis>
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Carpenter, 1966
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</taxonomicName>
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,
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<typeStatus id="548033958625FF8FFD11FCD003A9FCAE" box="[707,755,829,853]" pageId="2" pageNumber="464">type</typeStatus>
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species of the Permian genus
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<taxonomicName id="4C3BF6B48625FF8FFBB2FCD10585FCAE" box="[1120,1247,828,853]" class="Insecta" family="Tococladidae" genus="Tococladus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="464" phylum="Arthropoda" rank="genus">
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<emphasis id="B94F51258625FF8FFBB2FCD10585FCAE" box="[1120,1247,828,853]" italics="true" pageId="2" pageNumber="464">Tococladus</emphasis>
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</taxonomicName>
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. It is especially remarkable in the distal fusion of RA with
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<collectingCountry id="F32CCDA78625FF8FFDAEFC8D03C4FC8C" box="[636,670,864,887]" name="Poland" pageId="2" pageNumber="464">RP</collectingCountry>
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near wing apex and the numerous parallel posterior branches of
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<collectingCountry id="F32CCDA78625FF8FFAA8FC8D04C6FC8C" box="[1402,1436,864,887]" name="Poland" pageId="2" pageNumber="464">RP</collectingCountry>
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with short crossveins between them (see Béthoux et al., 2003: fig. 1). Of course an attribution to the genus
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<taxonomicName id="4C3BF6B48625FF8FFF45FC4A004CFC3B" box="[151,278,935,960]" class="Insecta" family="Tococladidae" genus="Tococladus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="464" phylum="Arthropoda" rank="genus" sensu="stricto">
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<emphasis id="B94F51258625FF8FFF45FC4A004CFC3B" box="[151,278,935,960]" italics="true" pageId="2" pageNumber="464">Tococladus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A27CEC5E8625FF8FFECFFC4500F0FC3B" box="[285,426,935,960]" pageId="2" pageNumber="464" sensu="stricto">sensu stricto</taxonomicNameLabel>
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is impossible to establish. The main interest of this fossil is that it represents the second
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<taxonomicName id="4C3BF6B48625FF8FFF45FC210000FC1E" box="[151,346,972,997]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="464" phylum="Arthropoda" rank="order">Cnemidolestodea</taxonomicName>
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from the outcrop of Avion, the first one being
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<emphasis id="B94F51258625FF8FFCAEFC210529FC1F" box="[892,1139,972,997]" italics="true" pageId="2" pageNumber="464">Aviocladus pectinatus</emphasis>
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<bibRefCitation id="EFAAF0C68625FF8FFBAEFC210188FBF3" author="Prokop" pageId="2" pageNumber="464" refString="Prokop, J., Roques, P. & Nel, A. (2014) New non-holometabolous insects from Pennsylvanian of Avion locality in Pas-de- Calais, France (Insecta: ' Exopterygota'). Alcheringa, 38, 155 - 169. http: // dx. doi. org / 10.1080 / 03115518.2014.848620" type="journal article" year="2014">Prokop, Roques and Nel, 2014</bibRefCitation>
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, from which it differs in the more numerous branches of
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<collectingCountry id="F32CCDA78625FF8FFC9CFC1D022AFBFC" box="[846,880,1008,1031]" name="Poland" pageId="2" pageNumber="464">RP</collectingCountry>
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and veinlets between RA and costal margin more sigmoidal (
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<bibRefCitation id="EFAAF0C68625FF8FFEC6FBF900B5FBD7" author="Prokop" box="[276,495,1044,1069]" pageId="2" pageNumber="464" refString="Prokop, J., Roques, P. & Nel, A. (2014) New non-holometabolous insects from Pennsylvanian of Avion locality in Pas-de- Calais, France (Insecta: ' Exopterygota'). Alcheringa, 38, 155 - 169. http: // dx. doi. org / 10.1080 / 03115518.2014.848620" type="journal article" year="2014">Prokop et al., 2014</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |