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<document id="D5568D72DA818A55578FBC9A62708F47" ID-CLB-Dataset="42309" ID-DOI="10.11646/zootaxa.3878.5.4" ID-GBIF-Dataset="697c91ee-9d8b-4095-bd15-e55709b2b6bd" ID-ISSN="1175-5326" ID-Zenodo-Dep="229037" ID-ZooBank="377F9037-D039-44A5-B562-051560CBC9E0" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460949457252" checkinUser="plazi" docAuthor="Coty, David, Háva, Jiří, Prokop, Jakub, Roques, Patrick &amp; Nel, André" docDate="2014" docId="03923C218625FF8FFF45FE8400A1FBD7" docLanguage="en" docName="zt03878p470.pdf" docOrigin="Zootaxa 3878 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tococladus" docType="treatment" docVersion="9" lastPageNumber="464" masterDocId="FFAB44598627FF8DFFD2FFED015AFFFB" masterDocTitle="New archaeorthopteran insects from the Late Carboniferous of the Nord and Pasde-Calais basins in northern France (Insecta: Cnemidolestodea, Panorthoptera)" masterLastPageNumber="470" masterPageNumber="462" pageNumber="464" updateTime="1698585716371" updateUser="ExternalLinkService">
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<mods:title id="585A4DC61E305936FB9719DC07218F79">New archaeorthopteran insects from the Late Carboniferous of the Nord and Pasde-Calais basins in northern France (Insecta: Cnemidolestodea, Panorthoptera)</mods:title>
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<mods:namePart id="E1168E52941B45BEEDE70FC0D5B8483A">Coty, David</mods:namePart>
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<mods:namePart id="498C45EAED9E5ED8D48646CCD1D8CF52">Háva, Jiří</mods:namePart>
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<mods:namePart id="FA9B8E480C20D8E145D48C7D7820CE47">Prokop, Jakub</mods:namePart>
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<mods:namePart id="9FC717B9F661DA5D36BDE9D5E4E52845">Roques, Patrick</mods:namePart>
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<mods:namePart id="BE1FA6BBE2775767D63D192143CADAAE">Nel, André</mods:namePart>
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<mods:title id="08280AA3C4C78E1FF5EF04B9145B8EAD">Zootaxa</mods:title>
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<mods:date id="53DF8E3D046D3688966B480BCDCEDA6C">2014</mods:date>
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<mods:number id="EEC50D3FE3FDA4237F95CEC5C3A25BC8">3878</mods:number>
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<treatment id="03923C218625FF8FFF45FE8400A1FBD7" ID-DOI="http://doi.org/10.5281/zenodo.6142659" ID-GBIF-Taxon="119591891" ID-Zenodo-Dep="6142659" LSID="urn:lsid:plazi:treatment:03923C218625FF8FFF45FE8400A1FBD7" httpUri="http://treatment.plazi.org/id/03923C218625FF8FFF45FE8400A1FBD7" lastPageNumber="464" pageId="2" pageNumber="464">
<subSubSection id="C321DEBC8625FF8FFF45FE84033BFE78" box="[151,609,361,387]" pageId="2" pageNumber="464" type="nomenclature">
<paragraph id="8B848D378625FF8FFF45FE84033BFE78" blockId="2.[151,609,361,421]" box="[151,609,361,387]" pageId="2" pageNumber="464">
<heading id="D0CC3A5B8625FF8FFF45FE84033BFE78" bold="true" box="[151,609,361,387]" fontSize="11" level="1" pageId="2" pageNumber="464" reason="1">
<emphasis id="B94F51258625FF8FFF45FE84033BFE78" bold="true" box="[151,609,361,387]" pageId="2" pageNumber="464">
Cf.
<taxonomicName id="4C3BF6B48625FF8FFF16FE840016FE78" box="[196,332,361,387]" class="Insecta" family="Tococladidae" genus="Tococladus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="464" phylum="Arthropoda" rank="genus">
<emphasis id="B94F51258625FF8FFF16FE840016FE78" bold="true" box="[196,332,361,387]" italics="true" pageId="2" pageNumber="464">Tococladus</emphasis>
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species undetermined
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C321DEBC8625FF8FFF45FE6101B1FE5F" box="[151,235,396,421]" pageId="2" pageNumber="464" type="description">
<paragraph id="8B848D378625FF8FFF45FE6101B1FE5F" blockId="2.[151,609,361,421]" box="[151,235,396,421]" pageId="2" pageNumber="464">
(
<figureCitation id="130091B28625FF8FFF4DFE6101BAFE5F" box="[159,224,396,421]" captionStart="FIGURE 1" captionStartId="1.[151,250,1786,1808]" captionTargetBox="[356,1230,483,1756]" captionTargetId="figure@1.[356,1231,483,1761]" captionTargetPageId="1" captionText="FIGURE 1. cf. Tococladus sp. specimen Avion 0 0 0 13 stored in MNHN coll. finally deposited in the Musée Géologique Pierre Vetter, Decazeville, France. A, photograph of wing apex venation; B, line drawing of apical part of wing. Scale bar = 5 mm." httpUri="https://zenodo.org/record/229038/files/figure.png" pageId="2" pageNumber="464">Fig. 1</figureCitation>
)
</paragraph>
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<subSubSection id="C321DEBC8625FF8FFF45FE39032DFDAC" pageId="2" pageNumber="464" type="materials_examined">
<paragraph id="8B848D378625FF8FFF45FE3903FAFDF4" blockId="2.[151,1436,468,1069]" pageId="2" pageNumber="464">
<emphasis id="B94F51258625FF8FFF45FE390030FE16" bold="true" box="[151,362,468,493]" pageId="2" pageNumber="464">Material studied:</emphasis>
Specimen 0 0 13 (print and counterprint), coll. Patrick Roques, to be deposited in the Musée Géologique Pierre Vetter, Decazeville,
<collectingCountry id="F32CCDA78625FF8FFD9EFE1503C6FDF4" box="[588,668,504,527]" name="France" pageId="2" pageNumber="464">France</collectingCountry>
.
</paragraph>
<paragraph id="8B848D378625FF8FFF15FDF1032DFDAC" blockId="2.[151,1436,468,1069]" pageId="2" pageNumber="464">
<emphasis id="B94F51258625FF8FFF15FDF100D6FDCE" bold="true" box="[199,396,540,565]" pageId="2" pageNumber="464">Age and outcrop</emphasis>
.
<typeStatus id="548033958625FF8FFE49FDF00089FDCE" box="[411,467,541,565]" pageId="2" pageNumber="464">Type</typeStatus>
locality and horizon. Moscovian (Westphalian C/D equivalent to Bolsovian/Asturian), Terril N°7, Avion, Pas-de-Calais,
<collectingCountry id="F32CCDA78625FF8FFDF6FDAD032EFDAC" box="[548,628,576,599]" name="France" pageId="2" pageNumber="464">France</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C321DEBC8625FF8FFF15FD890577FCF7" pageId="2" pageNumber="464" type="description">
<paragraph id="8B848D378625FF8FFF15FD890577FCF7" blockId="2.[151,1436,468,1069]" pageId="2" pageNumber="464">
<emphasis id="B94F51258625FF8FFF15FD89000BFD86" bold="true" box="[199,337,612,637]" pageId="2" pageNumber="464">Description</emphasis>
. A fragment of wing corresponding to the anterior part of the apex, fragment
<quantity id="4CC320D28625FF8FFB12FD89047EFD87" box="[1216,1316,612,636]" metricMagnitude="-2" metricUnit="m" metricValue="1.94" pageId="2" pageNumber="464" unit="mm" value="19.4">19.4 mm</quantity>
long,
<quantity id="4CC320D28625FF8FFABEFD890199FD64" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="2" pageNumber="464" unit="mm" value="11.7">11.7 mm</quantity>
wide; convex RA with a series of sigmoidal anterior veinlets, RA distally fused with concave main branch of
<collectingCountry id="F32CCDA78625FF8FFF45FD4001E3FD3F" box="[151,185,685,708]" name="Poland" pageId="2" pageNumber="464">RP</collectingCountry>
, near wing apex, common branch
<quantity id="4CC320D28625FF8FFDECFD4103CEFD3F" box="[574,660,684,709]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="2" pageNumber="464" unit="mm" value="2.6">2.6 mm</quantity>
long; a series of straight crossveins in area between RA and
<collectingCountry id="F32CCDA78625FF8FFA9AFD400434FD3F" box="[1352,1390,685,708]" name="Poland" pageId="2" pageNumber="464">RP</collectingCountry>
;
<collectingCountry id="F32CCDA78625FF8FFAA8FD4004C6FD3F" box="[1402,1436,685,708]" name="Poland" pageId="2" pageNumber="464">RP</collectingCountry>
with seven to nine parallel posterior branches with a series of simple straight crossveins between them; apical parts of branches of M and possibly CuA also preserved, parallel to the branches of
<collectingCountry id="F32CCDA78625FF8FFBD4FD180573FCF7" box="[1030,1065,757,780]" name="Poland" pageId="2" pageNumber="464">RP</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C321DEBC8625FF8FFF15FCFA00A1FBD7" pageId="2" pageNumber="464" type="discussion">
<paragraph id="8B848D378625FF8FFF15FCFA00A1FBD7" blockId="2.[151,1436,468,1069]" pageId="2" pageNumber="464">
<emphasis id="B94F51258625FF8FFF15FCFA001FFCCB" bold="true" box="[199,325,791,816]" pageId="2" pageNumber="464">Discussion</emphasis>
. Although very incomplete, this wing fragment fits quite well with the equivalent portion of a forewing of
<taxonomicName id="4C3BF6B48625FF8FFEF8FCD103E9FCAE" authority="Carpenter, 1966" authorityName="Carpenter" authorityYear="1966" box="[298,691,828,853]" class="Insecta" family="Tococladidae" genus="Tococladus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="464" phylum="Arthropoda" rank="species" species="ralus">
<emphasis id="B94F51258625FF8FFEF8FCD100B4FCAE" box="[298,494,828,853]" italics="true" pageId="2" pageNumber="464">Tococladus ralus</emphasis>
Carpenter, 1966
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,
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species of the Permian genus
<taxonomicName id="4C3BF6B48625FF8FFBB2FCD10585FCAE" box="[1120,1247,828,853]" class="Insecta" family="Tococladidae" genus="Tococladus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="464" phylum="Arthropoda" rank="genus">
<emphasis id="B94F51258625FF8FFBB2FCD10585FCAE" box="[1120,1247,828,853]" italics="true" pageId="2" pageNumber="464">Tococladus</emphasis>
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. It is especially remarkable in the distal fusion of RA with
<collectingCountry id="F32CCDA78625FF8FFDAEFC8D03C4FC8C" box="[636,670,864,887]" name="Poland" pageId="2" pageNumber="464">RP</collectingCountry>
near wing apex and the numerous parallel posterior branches of
<collectingCountry id="F32CCDA78625FF8FFAA8FC8D04C6FC8C" box="[1402,1436,864,887]" name="Poland" pageId="2" pageNumber="464">RP</collectingCountry>
with short crossveins between them (see Béthoux et al., 2003: fig. 1). Of course an attribution to the genus
<taxonomicName id="4C3BF6B48625FF8FFF45FC4A004CFC3B" box="[151,278,935,960]" class="Insecta" family="Tococladidae" genus="Tococladus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="464" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B94F51258625FF8FFF45FC4A004CFC3B" box="[151,278,935,960]" italics="true" pageId="2" pageNumber="464">Tococladus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27CEC5E8625FF8FFECFFC4500F0FC3B" box="[285,426,935,960]" pageId="2" pageNumber="464" sensu="stricto">sensu stricto</taxonomicNameLabel>
is impossible to establish. The main interest of this fossil is that it represents the second
<taxonomicName id="4C3BF6B48625FF8FFF45FC210000FC1E" box="[151,346,972,997]" class="Insecta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cnemidolestodea" pageId="2" pageNumber="464" phylum="Arthropoda" rank="order">Cnemidolestodea</taxonomicName>
from the outcrop of Avion, the first one being
<emphasis id="B94F51258625FF8FFCAEFC210529FC1F" box="[892,1139,972,997]" italics="true" pageId="2" pageNumber="464">Aviocladus pectinatus</emphasis>
<bibRefCitation id="EFAAF0C68625FF8FFBAEFC210188FBF3" author="Prokop" pageId="2" pageNumber="464" refString="Prokop, J., Roques, P. &amp; Nel, A. (2014) New non-holometabolous insects from Pennsylvanian of Avion locality in Pas-de- Calais, France (Insecta: ' Exopterygota'). Alcheringa, 38, 155 - 169. http: // dx. doi. org / 10.1080 / 03115518.2014.848620" type="journal article" year="2014">Prokop, Roques and Nel, 2014</bibRefCitation>
, from which it differs in the more numerous branches of
<collectingCountry id="F32CCDA78625FF8FFC9CFC1D022AFBFC" box="[846,880,1008,1031]" name="Poland" pageId="2" pageNumber="464">RP</collectingCountry>
and veinlets between RA and costal margin more sigmoidal (
<bibRefCitation id="EFAAF0C68625FF8FFEC6FBF900B5FBD7" author="Prokop" box="[276,495,1044,1069]" pageId="2" pageNumber="464" refString="Prokop, J., Roques, P. &amp; Nel, A. (2014) New non-holometabolous insects from Pennsylvanian of Avion locality in Pas-de- Calais, France (Insecta: ' Exopterygota'). Alcheringa, 38, 155 - 169. http: // dx. doi. org / 10.1080 / 03115518.2014.848620" type="journal article" year="2014">Prokop et al., 2014</bibRefCitation>
).
</paragraph>
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