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<document id="1C4D89601C2FF158C287D87F348CE1D7" ID-CLB-Dataset="8152" ID-DOI="10.2478/aemnp-2018-0035" ID-GBIF-Dataset="92fd172c-9d32-419f-9ea4-924f1cecb1b5" ID-ISSN="1804-6487" ID-Zenodo-Dep="4488739" ID-ZooBank="B1327F4F-90E6-4BE1-83CA-0D5BE068E6C3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612217885138" checkinUser="felipe" docAuthor="Huang, Meng-Chi, Li, Li-Zhen &amp; Yin, Zi-Wei" docDate="2018" docId="03EF20375869FFA1ECD2FE7BFEA85AAF" docLanguage="en" docName="ActaEntMusNatPra.58.2.457-478.pdf" docOrigin="Acta Entomologica Musei Nationalis Pragae 58 (2)" docStyle="DocumentStyle:BA2C5C8540539042E893A65A182CEA85.8:ActaEntMusNatPra.2018-.journal_article" docStyleId="BA2C5C8540539042E893A65A182CEA85" docStyleName="ActaEntMusNatPra.2018-.journal_article" docStyleVersion="8" docTitle="Pselaphodes spinosus Champion 1925" docType="treatment" docVersion="7" lastPageNumber="475" masterDocId="FFD6584F587BFFB3EC22FFA6FF8B5E7F" masterDocTitle="Eleven new species and a new country record of Pselaphodes (Coleoptera: Staphylinidae: Pselaphinae) from China, with a revised checklist of world species" masterLastPageNumber="478" masterPageNumber="457" pageNumber="475" updateTime="1698926012850" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-ND-3.0">
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<mods:title id="0368B078F420D450DD09EF7B254AD71C">Eleven new species and a new country record of Pselaphodes (Coleoptera: Staphylinidae: Pselaphinae) from China, with a revised checklist of world species</mods:title>
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<mods:namePart id="CA4603DD34757318433DB307FAB74613">Huang, Meng-Chi</mods:namePart>
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<mods:namePart id="5CA5B7A039F095401F89B017F6896A79">Li, Li-Zhen</mods:namePart>
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<mods:namePart id="A05C95E77DC8EC67A858ECE35B538F8B">Yin, Zi-Wei</mods:namePart>
<mods:nameIdentifier id="2F996CD3354B985C9A61EC73C18414A9" type="email">pselaphinae@gmail.com</mods:nameIdentifier>
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<mods:title id="60359197F4D86CE0B4B6F2D04306FC61">Acta Entomologica Musei Nationalis Pragae</mods:title>
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<mods:date id="27EE5573B23975CC0813411D6A4067E2">2018</mods:date>
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<mods:number id="85413E1B998B2339E0435861A4EE3CC4">2018-11-14</mods:number>
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<mods:classification id="DD9A8207FF7347B65215DAE3F91A63C9">journal article</mods:classification>
<mods:identifier id="1593F5945C3F9B3D3DB0037EDF483BE1" type="CLB-Dataset">8152</mods:identifier>
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<treatment id="03EF20375869FFA1ECD2FE7BFEA85AAF" ID-DOI="http://doi.org/10.5281/zenodo.4548447" ID-GBIF-Taxon="177423175" ID-Zenodo-Dep="4548447" LSID="urn:lsid:plazi:treatment:03EF20375869FFA1ECD2FE7BFEA85AAF" httpUri="http://treatment.plazi.org/id/03EF20375869FFA1ECD2FE7BFEA85AAF" lastPageNumber="475" pageId="18" pageNumber="475">
<subSubSection id="C35CC2AA5869FFA1ECD2FE7BFD145F8A" box="[240,671,477,501]" pageId="18" pageNumber="475" type="nomenclature">
<paragraph id="8BF991215869FFA1ECD2FE7BFD145F8A" blockId="18.[240,671,477,501]" box="[240,671,477,501]" pageId="18" pageNumber="475">
<heading id="D0B1264D5869FFA1ECD2FE7BFD145F8A" box="[240,671,477,501]" centered="true" fontSize="10" level="2" pageId="18" pageNumber="475" reason="3">
<taxonomicName id="4C46EAA25869FFA1ECD2FE7BFD145F8A" ID-CoL="4NF8J" authority="Champion, 1925" authorityName="Champion" authorityYear="1925" box="[240,671,477,501]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="475" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B9324D335869FFA1ECD2FE7BFD145F8A" bold="true" box="[240,671,477,501]" pageId="18" pageNumber="475">
<emphasis id="B9324D335869FFA1ECD2FE7BFE555F8A" bold="true" box="[240,478,477,501]" italics="true" pageId="18" pageNumber="475">Pselaphodes spinosus</emphasis>
<bibRefCitation id="EFD7ECD05869FFA1EDC6FE78FD145F8A" author="CHAMPION G. C." box="[484,671,478,501]" pageId="18" pageNumber="475" pagination="260 - 273" refId="ref10477" refString="CHAMPION G. C. 1925: Some Indian (and Tibetan) Coleoptera (18). Entomologist's Monthly Magazine 61: 260 - 273." type="journal article" year="1925">Champion, 1925</bibRefCitation>
</emphasis>
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</paragraph>
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<paragraph id="8BF991215869FFA1EDABFE5AFD8D5C70" blockId="18.[151,760,508,615]" box="[393,518,508,527]" pageId="18" pageNumber="475">
(
<figureCitation id="137D8DA45869FFA1EDADFE5AFE6B5C70" box="[399,480,508,527]" captionStart="Fig" captionStartId="16.[151,182,2063,2082]" captionTargetBox="[155,1432,869,2038]" captionTargetId="figure-306@16.[151,1436,867,2044]" captionTargetPageId="16" captionText="Fig. 16. Dorsal habitus of Pselaphodes males. A – P. songxiaobini sp. nov; B – P. spinosus Champion, 1925. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4488805" httpUri="https://zenodo.org/record/4488805/files/figure.png" pageId="18" pageNumber="475">Figs 16B</figureCitation>
,
<figureCitation id="137D8DA45869FFA1EDCBFE5AFE745C70" box="[489,511,508,527]" captionStart="Fig" captionStartId="18.[151,182,2036,2055]" captionTargetBox="[236,1358,1267,2020]" captionTargetId="figure-422@18.[229,1359,1264,2024]" captionTargetPageId="18" captionText="Fig. 18. Male diagnostic features of Pselaphodes spinosus Champion, 1925. A – antennal club; B–C – aedeagus, in dorsal (B), and lateral (C) view. Scale bar: 0.3 mm in A; 0.2 mm in B–C." figureDoi="http://doi.org/10.5281/zenodo.4488813" httpUri="https://zenodo.org/record/4488813/files/figure.png" pageId="18" pageNumber="475">18</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C35CC2AA5869FFA1ECB5FDB8FECD5C18" pageId="18" pageNumber="475" type="reference_group">
<paragraph id="8BF991215869FFA1ECB5FDB8FD9C5C4E" blockId="18.[151,760,508,615]" box="[151,535,542,561]" pageId="18" pageNumber="475">
<treatmentCitationGroup id="AB56B60F5869FFA1ECB5FDB8FD9C5C4E" box="[151,535,542,561]" pageId="18" pageNumber="475">
<treatmentCitation id="0AE7B7305869FFA1ECB5FDB8FD9F5C4E" author="CHAMPION G. C." box="[151,532,542,561]" page="261" pageId="18" pageNumber="475" year="1925">
<taxonomicName id="4C46EAA25869FFA1ECB5FDB8FD9F5C4E" ID-CoL="4NF8J" authority="Champion, 1925: 261" authorityName="Champion" authorityPageNumber="261" authorityYear="1925" box="[151,532,542,561]" class="Insecta" family="Staphylinidae" genus="Pselaphodes" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="475" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B9324D335869FFA1ECB5FDB8FEDA5C4E" box="[151,337,542,561]" italics="true" pageId="18" pageNumber="475">Pselaphodes spinosus</emphasis>
<bibRefCitation id="EFD7ECD05869FFA1ED75FDB8FD9F5C4E" author="CHAMPION G. C." box="[343,532,542,561]" pageId="18" pageNumber="475" pagination="260 - 273" refId="ref10477" refString="CHAMPION G. C. 1925: Some Indian (and Tibetan) Coleoptera (18). Entomologist's Monthly Magazine 61: 260 - 273." type="journal article" year="1925">Champion, 1925: 261</bibRefCitation>
</taxonomicName>
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.
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</paragraph>
<paragraph id="8BF991215869FFA1ECB5FD9FFECD5C18" blockId="18.[151,760,508,615]" pageId="18" pageNumber="475">
<treatmentCitationGroup id="AB56B60F5869FFA1ECB5FD9FFECD5C18" pageId="18" pageNumber="475">
<emphasis id="B9324D335869FFA1ECB5FD9FFF2E5C33" box="[151,165,569,588]" italics="true" pageId="18" pageNumber="475">=</emphasis>
<treatmentCitation id="0AE7B7305869FFA1EC89FD9FFD9E5C33" author="JEANNEL R." box="[171,533,569,588]" page="454" pageId="18" pageNumber="475" year="1960">
<taxonomicName id="4C46EAA25869FFA1EC89FD9FFD9E5C33" ID-CoL="6NPNQ" authority="Jeannel, 1960: 454" authorityName="Jeannel" authorityPageNumber="454" authorityYear="1960" box="[171,533,569,588]" class="Insecta" family="Staphylinidae" genus="Labomimus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="475" phylum="Arthropoda" rank="species" species="championi">
<emphasis id="B9324D335869FFA1EC89FD9FFEE65C33" box="[171,365,569,588]" italics="true" pageId="18" pageNumber="475">Labomimus championi</emphasis>
<bibRefCitation id="EFD7ECD05869FFA1ED50FD9FFD9E5C33" author="JEANNEL R." box="[370,533,569,588]" pageId="18" pageNumber="475" pagination="403 - 456" refId="ref10692" refString="JEANNEL R. 1960: Sur les Pselaphides (Coleoptera) de l'Inde septentrionale. Bulletin of the British Museum (Natural History). Entomology 9: 403 - 456." type="journal article" year="1960">Jeannel, 1960: 454</bibRefCitation>
</taxonomicName>
</treatmentCitation>
. (comb. &amp; syn. nov. in
<treatmentCitation id="0AE7B7305869FFA1EEFAFD9FFEB75C18" author="YIN Z. - W. &amp; LI L. - Z." page="340" pageId="18" pageNumber="475" year="2015">
<bibRefCitation id="EFD7ECD05869FFA1EEFAFD9FFEB75C18" author="YIN Z. - W. &amp; LI L. - Z." pageId="18" pageNumber="475" pagination="331 - 344" refId="ref11092" refString="YIN Z. - W. &amp; LI L. - Z. 2015: Review of some species in the genera Pselaphodes Westwood and Labomimus Sharp (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 4040 (3): 331 - 344." type="journal article" year="2015">YIN &amp; LI 2015: 340</bibRefCitation>
</treatmentCitation>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C35CC2AA5869FFA1ECB5FDD8FDE45D8F" pageId="18" pageNumber="475" type="materials_examined">
<paragraph id="8BF991215869FFA1ECB5FDD8FDD55CEE" blockId="18.[151,761,638,737]" box="[151,606,638,657]" pageId="18" pageNumber="475">
<emphasis id="B9324D335869FFA1ECB5FDD8FEF25CEE" bold="true" box="[151,377,638,657]" pageId="18" pageNumber="475">
<typeStatus id="54FD2F835869FFA1ECB5FDD8FF4F5CEE" box="[151,196,638,657]" pageId="18" pageNumber="475">Type</typeStatus>
material examined.
</emphasis>
See
<bibRefCitation id="EFD7ECD05869FFA1ED83FDD8FDD25CEE" author="YIN Z. - W. &amp; LI L. - Z." box="[417,601,638,657]" pageId="18" pageNumber="475" pagination="331 - 344" refId="ref11092" refString="YIN Z. - W. &amp; LI L. - Z. 2015: Review of some species in the genera Pselaphodes Westwood and Labomimus Sharp (Coleoptera: Staphylinidae: Pselaphinae). Zootaxa 4040 (3): 331 - 344." type="journal article" year="2015">YIN &amp; LI (2015: 340)</bibRefCitation>
.
</paragraph>
<paragraph id="8BF991215869FFA1ECB5FD3EFDA65C9E" blockId="18.[151,761,638,737]" pageId="18" pageNumber="475">
<emphasis id="B9324D335869FFA1ECB5FD3EFE745CD4" bold="true" box="[151,511,664,683]" pageId="18" pageNumber="475">
Additional material examined.
<collectingCountry id="F351D1B15869FFA1ED93FD3EFE705CD4" box="[433,507,664,683]" name="China" pageId="18" pageNumber="475">CHINA</collectingCountry>
:
</emphasis>
2 ♁♁, ‘China:
<collectingRegion id="49825FC35869FFA1EEA3FD3EFD3B5CD4" box="[641,688,664,683]" country="China" name="Xizang" pageId="18" pageNumber="475">Tibet</collectingRegion>
, Rikaze, Jilong,
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,
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,
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,
<date id="FFF8B7E15869FFA1EE6CFD15FD465CB9" box="[590,717,691,710]" pageId="18" pageNumber="475" value="2017-07-29" valueMax="2017-07-31" valueMin="2017-07-29">29-31.vii.2017</date>
, Qiu &amp; Xu leg. [
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<collectionCode id="ED5709E45869FFA1ECDAFD6FFEB85CA1" box="[248,307,713,734]" country="USA" httpUri="http://grbio.org/cool/10w8-eg5b" name="Michigan Department of Health" pageId="18" pageNumber="475">MDH</collectionCode>
ṞNJẘợḦÁIJ
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]’ (
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).
</paragraph>
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<emphasis id="B9324D335869FFA1ECB5FD5FFD355D6F" bold="true" box="[151,702,761,784]" pageId="18" pageNumber="475">Supplementary description of additional material.</emphasis>
<emphasis id="B9324D335869FFA1EEE0FD5FFD735D6F" box="[706,760,761,784]" italics="true" pageId="18" pageNumber="475">Male</emphasis>
(
<figureCitation id="137D8DA45869FFA1ECBDFCBFFF755D4F" box="[159,254,793,816]" captionStart="Fig" captionStartId="16.[151,182,2063,2082]" captionTargetBox="[155,1432,869,2038]" captionTargetId="figure-306@16.[151,1436,867,2044]" captionTargetPageId="16" captionText="Fig. 16. Dorsal habitus of Pselaphodes males. A – P. songxiaobini sp. nov; B – P. spinosus Champion, 1925. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4488805" httpUri="https://zenodo.org/record/4488805/files/figure.png" pageId="18" pageNumber="475">Fig. 16B</figureCitation>
). Antennomere X (
<figureCitation id="137D8DA45869FFA1EDE1FCBFFDAA5D4F" box="[451,545,793,816]" captionStart="Fig" captionStartId="18.[151,182,2036,2055]" captionTargetBox="[236,1358,1267,2020]" captionTargetId="figure-422@18.[229,1359,1264,2024]" captionTargetPageId="18" captionText="Fig. 18. Male diagnostic features of Pselaphodes spinosus Champion, 1925. A – antennal club; B–C – aedeagus, in dorsal (B), and lateral (C) view. Scale bar: 0.3 mm in A; 0.2 mm in B–C." figureDoi="http://doi.org/10.5281/zenodo.4488813" httpUri="https://zenodo.org/record/4488813/files/figure.png" pageId="18" pageNumber="475">Fig. 18A</figureCitation>
) strongly concave at middle, metaventral processes long, emarginated at apices. Length of aedeagus (
<figureCitation id="137D8DA45869FFA1EDA6FCFFFE5C5D0F" box="[388,471,857,880]" captionStart="Fig" captionStartId="18.[151,182,2036,2055]" captionTargetBox="[236,1358,1267,2020]" captionTargetId="figure-422@18.[229,1359,1264,2024]" captionTargetPageId="18" captionText="Fig. 18. Male diagnostic features of Pselaphodes spinosus Champion, 1925. A – antennal club; B–C – aedeagus, in dorsal (B), and lateral (C) view. Scale bar: 0.3 mm in A; 0.2 mm in B–C." figureDoi="http://doi.org/10.5281/zenodo.4488813" httpUri="https://zenodo.org/record/4488813/files/figure.png" pageId="18" pageNumber="475">Figs 18</figureCitation>
B–C)
<quantity id="4CBE3CC45869FFA1EE3CFCFFFD0F5D0F" box="[542,644,857,880]" metricMagnitude="-4" metricUnit="m" metricValue="6.3" pageId="18" pageNumber="475" unit="mm" value="0.63">0.63 mm</quantity>
. Each eye composed of about 40 facets. Measurements: BL
<quantity id="4CBE3CC45869FFA1EEADFCDFFF485DCF" metricMagnitude="-3" metricUnit="m" metricValue="3.645" metricValueMax="3.68" metricValueMin="3.61" pageId="18" pageNumber="475" unit="mm" value="3.645" valueMax="3.68" valueMin="3.61">3.61–3.68 mm</quantity>
, HL
<quantity id="4CBE3CC45869FFA1ECD1FC3FFE055DCF" box="[243,398,921,944]" metricMagnitude="-4" metricUnit="m" metricValue="7.55" metricValueMax="7.8" metricValueMin="7.3" pageId="18" pageNumber="475" unit="mm" value="0.755" valueMax="0.78" valueMin="0.73">0.73–0.78 mm</quantity>
, HW
<quantity id="4CBE3CC45869FFA1EDE5FC3FFDE95DCF" box="[455,610,921,944]" metricMagnitude="-4" metricUnit="m" metricValue="7.1499999999999995" metricValueMax="7.199999999999999" metricValueMin="7.1" pageId="18" pageNumber="475" unit="mm" value="0.715" valueMax="0.72" valueMin="0.71">0.71–0.72 mm</quantity>
,
<collectionCode id="ED5709E45869FFA1EE49FC3FFD025DCF" box="[619,649,921,944]" country="Czech Republic" name="Západoceské muzeum v Plzni" pageId="18" pageNumber="475" type="Museum">PL</collectionCode>
<quantity id="4CBE3CC45869FFA1EEACFC3FFF485DAF" metricMagnitude="-4" metricUnit="m" metricValue="7.75" metricValueMax="7.8" metricValueMin="7.7" pageId="18" pageNumber="475" unit="mm" value="0.775" valueMax="0.78" valueMin="0.77">0.77–0.78 mm</quantity>
,
<collectionCode id="ED5709E45869FFA1ECEEFC1FFF785DAF" box="[204,243,953,976]" country="Thailand" name="Paleontological Collections" pageId="18" pageNumber="475">PW</collectionCode>
<quantity id="4CBE3CC45869FFA1ECDAFC1FFE195DAF" box="[248,402,953,976]" metricMagnitude="-4" metricUnit="m" metricValue="7.199999999999999" metricValueMax="7.3" metricValueMin="7.1" pageId="18" pageNumber="475" unit="mm" value="0.72" valueMax="0.73" valueMin="0.71">0.71–0.73 mm</quantity>
, EL
<quantity id="4CBE3CC45869FFA1ED9CFC1FFDD35DAF" box="[446,600,953,976]" metricMagnitude="-3" metricUnit="m" metricValue="1.1099999999999999" metricValueMax="1.16" metricValueMin="1.06" pageId="18" pageNumber="475" unit="mm" value="1.1099999999999999" valueMax="1.16" valueMin="1.06">1.06–1.16 mm</quantity>
, EW
<quantity id="4CBE3CC45869FFA1EEACFC1FFF485D8F" metricMagnitude="-3" metricUnit="m" metricValue="1.375" metricValueMax="1.4" metricValueMin="1.35" pageId="18" pageNumber="475" unit="mm" value="1.375" valueMax="1.4" valueMin="1.35">1.35–1.40 mm</quantity>
, AL 1.00–
<quantity id="4CBE3CC45869FFA1ED10FC7FFE185D8F" box="[306,403,985,1008]" metricMagnitude="-3" metricUnit="m" metricValue="1.02" pageId="18" pageNumber="475" unit="mm" value="1.02">1.02 mm</quantity>
, AW
<quantity id="4CBE3CC45869FFA1EDEFFC7FFDE05D8F" box="[461,619,985,1008]" metricMagnitude="-3" metricUnit="m" metricValue="1.39" metricValueMax="1.41" metricValueMin="1.37" pageId="18" pageNumber="475" unit="mm" value="1.39" valueMax="1.41" valueMin="1.37">1.37–1.41 mm</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C35CC2AA5869FFA1ECB5FC5FFD645AEF" pageId="18" pageNumber="475" type="discussion">
<paragraph id="8BF991215869FFA1ECB5FC5FFD645AEF" blockId="18.[151,761,761,1232]" pageId="18" pageNumber="475">
<emphasis id="B9324D335869FFA1ECB5FC5FFE9C5A6F" bold="true" box="[151,279,1017,1040]" pageId="18" pageNumber="475">Comments.</emphasis>
Male of this species has a unique antennomere X which is distinctly constricted at its middle. Combined with the nearly identical form of the aedeagus and close distribution, the population from
<collectingRegion id="49825FC35869FFA1EE24FBFFFDD85A0F" box="[518,595,1113,1136]" country="China" name="Xizang" pageId="18" pageNumber="475">Xizang</collectingRegion>
can be readily identified as conspecific with that from the
<typeStatus id="54FD2F835869FFA1EE47FBDFFD185AEF" box="[613,659,1145,1168]" pageId="18" pageNumber="475">type</typeStatus>
locality.
</paragraph>
</subSubSection>
<subSubSection id="C35CC2AA5869FFA1ECB5FB3FFEA85AAF" pageId="18" pageNumber="475" type="distribution">
<paragraph id="8BF991215869FFA1ECB5FB3FFEA85AAF" blockId="18.[151,761,761,1232]" pageId="18" pageNumber="475">
<emphasis id="B9324D335869FFA1ECB5FB3FFEAD5ACF" bold="true" box="[151,294,1177,1200]" pageId="18" pageNumber="475">Distribution.</emphasis>
<collectingCountry id="F351D1B15869FFA1ED09FB3FFEE55ACF" box="[299,366,1177,1200]" name="China" pageId="18" pageNumber="475">China</collectingCountry>
:
<collectingRegion id="49825FC35869FFA1ED57FB3FFE4A5ACF" box="[373,449,1177,1200]" country="China" name="Xizang" pageId="18" pageNumber="475">Xizang</collectingRegion>
(
<emphasis id="B9324D335869FFA1EDECFB3FFD2D5ACF" bold="true" box="[462,678,1177,1200]" pageId="18" pageNumber="475">new country record</emphasis>
);
<collectingCountry id="F351D1B15869FFA1EE99FB3FFD7E5ACF" box="[699,757,1177,1200]" name="India" pageId="18" pageNumber="475">India</collectingCountry>
:
<collectingRegion id="49825FC35869FFA1ECB5FB1FFE955AAF" box="[151,286,1209,1232]" country="India" name="Uttarakhand" pageId="18" pageNumber="475">Uttarakhand</collectingRegion>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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