113 lines
14 KiB
XML
113 lines
14 KiB
XML
<document id="C997912F115B05180CE108010D857302" ID-DOI="10.11646/zootaxa.4576.3.6" ID-GBIF-Dataset="7abdc62c-c350-49dd-8060-4ed64da07362" ID-ISSN="1175-5326" ID-Zenodo-Dep="3715772" ID-ZooBank="56BC8573-D4A1-4B18-9BF6-7AB5F7984BFD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584603665960" checkinUser="plazi" docAuthor="Muona, Jyrki" docDate="2019" docId="03D387E9FFAE7777FF6FEC40FF51FB94" docLanguage="en" docName="zootaxa.4576.3.6.pdf" docOrigin="Zootaxa 4576 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tyrannothroscus rex Muona 2019, new species" docType="treatment" docVersion="5" lastPageNumber="530" masterDocId="FFEAFF91FFA7777EFFF8EE40FF81FFA7" masterDocTitle="Throscidae (Coleoptera) relationships, with descriptions of new fossil genera and species" masterLastPageNumber="543" masterPageNumber="521" pageNumber="530" updateTime="1698730058212" updateUser="plazi">
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<mods:title id="6178137EE68CA07ED2BD8CA92A817AE2">Throscidae (Coleoptera) relationships, with descriptions of new fossil genera and species</mods:title>
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<mods:namePart id="40868937E0DD12A88B2E6CF060567BBD">Muona, Jyrki</mods:namePart>
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<mods:title id="6DDF9D828C8079DA3FB9631F6D7BC98C">Zootaxa</mods:title>
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<mods:part id="C9BC54630DCD68AC74890B05CCC99E46">
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<mods:date id="891C494962CBED5C1BABBCB831CA76CB">2019</mods:date>
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<mods:number id="30F7CDC7F0AD638448DC02E9A3842FA3">2019-04-03</mods:number>
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<mods:number id="1945856FC3EC309E0D8CDB77D7DCCF48">4576</mods:number>
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<treatment id="03D387E9FFAE7777FF6FEC40FF51FB94" ID-DOI="http://doi.org/10.5281/zenodo.3716786" ID-GBIF-Taxon="162639249" ID-Zenodo-Dep="3716786" LSID="urn:lsid:plazi:treatment:03D387E9FFAE7777FF6FEC40FF51FB94" httpUri="http://treatment.plazi.org/id/03D387E9FFAE7777FF6FEC40FF51FB94" lastPageNumber="530" pageId="9" pageNumber="530">
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<subSubSection id="C3606574FFAE7777FF6FEC40FDABFDBD" box="[151,554,512,538]" pageId="9" pageNumber="530" type="nomenclature">
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<paragraph id="8BC536FFFFAE7777FF6FEC40FDABFDBD" blockId="9.[151,554,512,571]" box="[151,554,512,538]" pageId="9" pageNumber="530">
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<heading id="D08D8193FFAE7777FF6FEC40FDABFDBD" bold="true" box="[151,554,512,538]" fontSize="11" level="1" pageId="9" pageNumber="530" reason="1">
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<emphasis id="B90EEAEDFFAE7777FF6FEC40FDABFDBD" bold="true" box="[151,554,512,538]" pageId="9" pageNumber="530">
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<taxonomicName id="4C7A4D7CFFAE7777FF6FEC40FE12FDBD" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[151,403,512,538]" class="Insecta" family="Throscidae" genus="Tyrannothroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="genus" status="sp. nov.">
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<emphasis id="B90EEAEDFFAE7777FF6FEC40FE12FDBD" bold="true" box="[151,403,512,538]" italics="true" pageId="9" pageNumber="530">Tyrannothroscus rex</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23D5796FFAE7777FE62EC40FDABFDBD" box="[410,554,512,538]" pageId="9" pageNumber="530" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606574FFAE7777FF6FEC62FEB6FD9C" box="[151,311,546,571]" pageId="9" pageNumber="530" type="description">
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<paragraph id="8BC536FFFFAE7777FF6FEC62FEB6FD9C" blockId="9.[151,554,512,571]" box="[151,311,546,571]" pageId="9" pageNumber="530">
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<figureCitation id="13412A7AFFAE7777FF6FEC62FE91FD9C" box="[151,272,546,571]" captionStart-0="FIGURE 6" captionStart-1="FIGURE 7" captionStart-2="FIGURE 8" captionStart-3="FIGURE 9" captionStart-4="FIGURE 10" captionStartId-0="10.[151,250,873,895]" captionStartId-1="10.[151,250,1949,1971]" captionStartId-2="11.[151,250,1081,1103]" captionStartId-3="11.[151,250,1998,2020]" captionStartId-4="12.[151,250,995,1017]" captionTargetBox-0="[151,1436,193,860]" captionTargetBox-1="[162,1431,968,1932]" captionTargetBox-2="[152,1435,210,1031]" captionTargetBox-3="[151,1436,1177,1948]" captionTargetBox-4="[150,1436,193,974]" captionTargetId-0="figure@10.[151,1436,193,860]" captionTargetId-1="figure@10.[162,1431,968,1932]" captionTargetId-2="figure@11.[152,1435,210,1031]" captionTargetId-3="figure@11.[151,1436,1177,1948]" captionTargetId-4="figure@12.[150,1436,193,974]" captionTargetPageId-0="10" captionTargetPageId-1="10" captionTargetPageId-2="11" captionTargetPageId-3="11" captionTargetPageId-4="12" captionText-0="FIGURE 6. Tyrannothroscus rex n. sp., ventral view. A = antennal groove; MtG = metathoracic groove; AtG = abdominal tarsal groove, SAR = supra-antennal ridge." captionText-1="FIGURE 7. Tyrannothroscus rex n. sp., head, ventrolateral view, slightly frontally. A = antennal groove; E = eye; F = frontoclypeal region; C = labrum; PL = ventrally bent fron margin of prosternum; PP = prosternal process; SAR = supraantennal ridge." captionText-2="FIGURE 8. Tyrannothroscus rex n. sp., metatarsus. t4 = lobed tarsomere 4." captionText-3="FIGURE 9. Tyrannothroscus rex n. sp., dorsal view." captionText-4="FIGURE 10. Tyrannothroscus rex n. sp., occiput and pronotum, frontal view." figureDoi-0="http://doi.org/10.5281/zenodo.3715784" figureDoi-1="http://doi.org/10.5281/zenodo.3715788" figureDoi-2="http://doi.org/10.5281/zenodo.3715790" figureDoi-3="http://doi.org/10.5281/zenodo.3715792" figureDoi-4="http://doi.org/10.5281/zenodo.3715794" httpUri-0="https://zenodo.org/record/3715784/files/figure.png" httpUri-1="https://zenodo.org/record/3715788/files/figure.png" httpUri-2="https://zenodo.org/record/3715790/files/figure.png" httpUri-3="https://zenodo.org/record/3715792/files/figure.png" httpUri-4="https://zenodo.org/record/3715794/files/figure.png" pageId="9" pageNumber="530">Figs. 6–10</figureCitation>
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,
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<figureCitation id="13412A7AFFAE7777FEE3EC63FEB6FD9C" box="[283,311,547,571]" captionStart="FIGURE 23" captionStartId="19.[151,250,1063,1085]" captionTargetBox="[216,1373,183,1042]" captionTargetId="figure@19.[216,1373,183,1042]" captionTargetPageId="19" captionText="FIGURE 23. Tyrannothroscus rex n. sp., holotype." figureDoi="http://doi.org/10.5281/zenodo.3715820" httpUri="https://zenodo.org/record/3715820/files/figure.png" pageId="9" pageNumber="530">23</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606574FFAE7777FF6FEC2AFDF4FD48" pageId="9" pageNumber="530" type="materials_examined">
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<paragraph id="8BC536FFFFAE7777FF6FEC2AFDF4FD48" blockId="9.[151,1437,618,1075]" pageId="9" pageNumber="530">
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<emphasis id="B90EEAEDFFAE7777FF6FEC2AFEC0FD24" bold="true" box="[151,321,618,643]" pageId="9" pageNumber="530">Type material</emphasis>
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.
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<typeStatus id="54C1885DFFAE7777FEABEC2AFE3AFD24" box="[339,443,618,643]" pageId="9" pageNumber="530" type="holotype">Holotype</typeStatus>
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labeled:
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<taxonomicName id="4C7A4D7CFFAE7777FDD1EC2AFC93FD25" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[553,786,618,643]" class="Insecta" family="Throscidae" genus="Tyrannothroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="genus" status="sp. nov.">Tyrannothroscus rex</taxonomicName>
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<emphasis id="B90EEAEDFFAE7777FCE4EC2AFCDEFD24" bold="true" box="[796,863,618,643]" pageId="9" pageNumber="530">
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<taxonomicNameLabel id="A23D5796FFAE7777FCE4EC2AFCDEFD24" box="[796,863,618,643]" pageId="9" pageNumber="530" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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/
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<typeStatus id="54C1885DFFAE7777FC90EC2BFC7EFD24" box="[872,1023,619,643]" pageId="9" pageNumber="530" type="holotype">HOLOTYPE</typeStatus>
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/J.Muona des. 10-2015. Reported to have been found from Jantarny. Sex unknown. Embedded in clear, flat, droplet-shaped Baltic Amber piece,
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<quantity id="4C829B1AFFAE7777FAB6ECCFFA1CFD01" box="[1358,1437,655,679]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="530" unit="mm" value="30.0">30 mm</quantity>
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<date id="FFC4103FFFAE7777FF6FECF3FF48FD6C" box="[151,201,691,715]" pageId="9" pageNumber="530" value="1919-10">
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x
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<quantity id="4C829B1AFFAE7777FF55ECF3FF48FD6C" box="[173,201,691,715]" metricMagnitude="-2" metricUnit="m" metricValue="1.9" pageId="9" pageNumber="530" unit="mm" value="19.0">19</quantity>
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</date>
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mm, narrow end with drilled hole with an attached silver loop for use as a pendant. A small number of tiny plant remains are present as well (
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<figureCitation id="13412A7AFFAE7777FDECEC96FDE8FD48" box="[532,617,726,751]" captionStart="FIGURE 23" captionStartId="19.[151,250,1063,1085]" captionTargetBox="[216,1373,183,1042]" captionTargetId="figure@19.[216,1373,183,1042]" captionTargetPageId="19" captionText="FIGURE 23. Tyrannothroscus rex n. sp., holotype." figureDoi="http://doi.org/10.5281/zenodo.3715820" httpUri="https://zenodo.org/record/3715820/files/figure.png" pageId="9" pageNumber="530">Fig. 23</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606574FFAE7777FF3FECBAFB9EFCB4" box="[199,1055,762,787]" pageId="9" pageNumber="530" type="diagnosis">
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<paragraph id="8BC536FFFFAE7777FF3FECBAFB9EFCB4" blockId="9.[151,1437,618,1075]" box="[199,1055,762,787]" pageId="9" pageNumber="530">
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<emphasis id="B90EEAEDFFAE7777FF3FECBAFEBBFCB4" bold="true" box="[199,314,762,787]" pageId="9" pageNumber="530">Diagnosis</emphasis>
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. Easily recognized by the generic characters and in the key below
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606574FFAE7777FF3FED5EFE95FC60" pageId="9" pageNumber="530" type="description">
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<paragraph id="8BC536FFFFAE7777FF3FED5EFE95FC60" blockId="9.[151,1437,618,1075]" pageId="9" pageNumber="530">
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<emphasis id="B90EEAEDFFAE7777FF3FED5EFED0FC90" bold="true" box="[199,337,798,823]" pageId="9" pageNumber="530">Description</emphasis>
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. Length
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<quantity id="4C829B1AFFAE7777FE44ED5FFD98FC91" box="[444,537,799,823]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="9" pageNumber="530" unit="mm" value="8.2">8.2 mm</quantity>
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. Head and pronotum with relatively sparse and fine punctation, on pronotum punctures of variable size, mostly small, microsculpture dense, isodiametric (
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<figureCitation id="13412A7AFFAE7777FC0AED02FBC4FCFC" box="[1010,1093,834,859]" captionStart="FIGURE 10" captionStartId="12.[151,250,995,1017]" captionTargetBox="[150,1436,193,974]" captionTargetId="figure@12.[150,1436,193,974]" captionTargetPageId="12" captionText="FIGURE 10. Tyrannothroscus rex n. sp., occiput and pronotum, frontal view." figureDoi="http://doi.org/10.5281/zenodo.3715794" httpUri="https://zenodo.org/record/3715794/files/figure.png" pageId="9" pageNumber="530">Fig. 10</figureCitation>
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). Elytral striae sharp, minutely punctate, interstices very wide, anteriorly flat, towards apex convex (
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<figureCitation id="13412A7AFFAE7777FC31ED26FB93FCD8" box="[969,1042,870,895]" captionStart="FIGURE 9" captionStartId="11.[151,250,1998,2020]" captionTargetBox="[151,1436,1177,1948]" captionTargetId="figure@11.[151,1436,1177,1948]" captionTargetPageId="11" captionText="FIGURE 9. Tyrannothroscus rex n. sp., dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3715792" httpUri="https://zenodo.org/record/3715792/files/figure.png" pageId="9" pageNumber="530">Fig. 9</figureCitation>
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). Venter strongly and moderately densely punctate, densest on first and second abdominal segment, vestiture visible on abdomen only, sparse, hairs fairly long.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606574FFAE7777FF3FED92FF51FB94" pageId="9" pageNumber="530" type="etymology">
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<paragraph id="8BC536FFFFAE7777FF3FED92FF51FB94" blockId="9.[151,1437,618,1075]" pageId="9" pageNumber="530">
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<emphasis id="B90EEAEDFFAE7777FF3FED92FEC7FC4C" bold="true" box="[199,326,978,1003]" pageId="9" pageNumber="530">Etymology</emphasis>
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.
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<taxonomicName id="4C7A4D7CFFAE7777FEA4ED92FDC8FC4C" authorityName="Muona" authorityYear="2019" box="[348,585,978,1003]" class="Insecta" family="Throscidae" genus="Tyrannothroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="530" phylum="Arthropoda" rank="genus">
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<emphasis id="B90EEAEDFFAE7777FEA4ED92FDC8FC4C" box="[348,585,978,1003]" italics="true" pageId="9" pageNumber="530">Tyrannothroscus rex</emphasis>
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</taxonomicName>
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is the largest throscid known. This in combination with the massive “eyebrows” creating an aggressive, fierce look and the wide body suggest it was a real monster among its own kind.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |