153 lines
17 KiB
XML
153 lines
17 KiB
XML
<document id="D14771CC8F215D9E8DE02445D79C2F3F" ID-DOI="10.11646/zootaxa.4576.3.6" ID-GBIF-Dataset="7abdc62c-c350-49dd-8060-4ed64da07362" ID-ISSN="1175-5326" ID-Zenodo-Dep="3715772" ID-ZooBank="56BC8573-D4A1-4B18-9BF6-7AB5F7984BFD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1584603665960" checkinUser="plazi" docAuthor="Muona, Jyrki" docDate="2019" docId="03D387E9FFA07776FF6FE86BFD40FAF9" docLanguage="en" docName="zootaxa.4576.3.6.pdf" docOrigin="Zootaxa 4576 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pseudothroscus balticus Muona 2019, new species" docType="treatment" docVersion="5" lastPageNumber="529" masterDocId="FFEAFF91FFA7777EFFF8EE40FF81FFA7" masterDocTitle="Throscidae (Coleoptera) relationships, with descriptions of new fossil genera and species" masterLastPageNumber="543" masterPageNumber="521" pageNumber="528" updateTime="1698730058212" updateUser="plazi">
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<mods:title id="7F9B15FCE76FF26372E9EF3D702CD944">Throscidae (Coleoptera) relationships, with descriptions of new fossil genera and species</mods:title>
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<mods:namePart id="B645DFACC83E29C44C2B9825949B8304">Muona, Jyrki</mods:namePart>
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<mods:title id="DCCCF24AE7B00E2D1D1DB3FDCFCF9E26">Zootaxa</mods:title>
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<mods:date id="370D42E88BA37242948FEAF668201A57">2019</mods:date>
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<mods:number id="90FB9264FF77BB9638A2FA0209405C9A">2019-04-03</mods:number>
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<mods:number id="60B01C467C12E3AC6416CAC8095F21CC">4576</mods:number>
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<treatment id="03D387E9FFA07776FF6FE86BFD40FAF9" ID-DOI="http://doi.org/10.5281/zenodo.3716778" ID-GBIF-Taxon="162639255" ID-Zenodo-Dep="3716778" LSID="urn:lsid:plazi:treatment:03D387E9FFA07776FF6FE86BFD40FAF9" httpUri="http://treatment.plazi.org/id/03D387E9FFA07776FF6FE86BFD40FAF9" lastPageId="8" lastPageNumber="529" pageId="7" pageNumber="528">
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<subSubSection id="C3606574FFA07779FF6FE86BFDD3F9E2" box="[151,594,1579,1605]" pageId="7" pageNumber="528" type="nomenclature">
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<paragraph id="8BC536FFFFA07779FF6FE86BFDD3F9E2" blockId="7.[151,594,1579,1638]" box="[151,594,1579,1605]" pageId="7" pageNumber="528">
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<heading id="D08D8193FFA07779FF6FE86BFDD3F9E2" bold="true" box="[151,594,1579,1605]" fontSize="11" level="1" pageId="7" pageNumber="528" reason="1">
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<emphasis id="B90EEAEDFFA07779FF6FE86BFDD3F9E2" bold="true" box="[151,594,1579,1605]" pageId="7" pageNumber="528">
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<taxonomicName id="4C7A4D7CFFA07779FF6FE86BFE3AF9E2" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[151,443,1579,1605]" class="Insecta" family="Throscidae" genus="Pseudothroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="genus" status="sp. nov.">
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<emphasis id="B90EEAEDFFA07779FF6FE86BFE3AF9E2" bold="true" box="[151,443,1579,1605]" italics="true" pageId="7" pageNumber="528">Pseudothroscus balticus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23D5796FFA07779FE3AE86BFDD3F9E2" box="[450,594,1579,1605]" pageId="7" pageNumber="528" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<subSubSection id="C3606574FFA07779FF6FE80DFEA8F9C1" box="[151,297,1613,1638]" pageId="7" pageNumber="528" type="description">
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<paragraph id="8BC536FFFFA07779FF6FE80DFEA8F9C1" blockId="7.[151,594,1579,1638]" box="[151,297,1613,1638]" pageId="7" pageNumber="528">
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<figureCitation id="13412A7AFFA07779FF6FE80DFF64F9C1" box="[151,229,1613,1638]" captionStart="FIGURE 4" captionStartId="7.[151,250,1482,1504]" captionTargetBox="[150,1436,802,1467]" captionTargetId="figure@7.[150,1436,802,1467]" captionTargetPageId="7" captionText="FIGURE 4. Pseudothroscus balticus n. sp., dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3715780" httpUri="https://zenodo.org/record/3715780/files/figure.png" pageId="7" pageNumber="528">Figs. 4</figureCitation>
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,
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<figureCitation id="13412A7AFFA07779FF09E80DFE80F9C1" box="[241,257,1613,1638]" captionStart="FIGURE 5" captionStartId="8.[151,250,1071,1093]" captionTargetBox="[150,1436,193,1050]" captionTargetId="figure@8.[150,1436,193,1050]" captionTargetPageId="8" captionText="FIGURE 5. Pseudothroscus balticus n. sp., ventral view. AC = antennal club; AF = antennal pit boarded by delicate ridge; L = labrum; LC = complete lateral pronotal ridge; MsT = simple mesotibia." figureDoi="http://doi.org/10.5281/zenodo.3715782" httpUri="https://zenodo.org/record/3715782/files/figure.png" pageId="7" pageNumber="528">5</figureCitation>
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,
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<figureCitation id="13412A7AFFA07779FEF5E80EFEA8F9C1" box="[269,297,1614,1638]" captionStart="FIGURE 22" captionStartId="18.[151,250,1887,1909]" captionTargetBox="[178,1417,1018,1876]" captionTargetId="figure@18.[178,1417,1018,1876]" captionTargetPageId="18" captionText="FIGURE 22. Pseudothroscus balticus n. sp., holotype." figureDoi="http://doi.org/10.5281/zenodo.3715818" httpUri="https://zenodo.org/record/3715818/files/figure.png" pageId="7" pageNumber="528">22</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C3606574FFA07779FF6FE8D5FCC5F821" pageId="7" pageNumber="528" type="materials_examined">
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||
<paragraph id="8BC536FFFFA07779FF6FE8D5FA82F909" blockId="7.[151,1437,1685,1998]" box="[151,1283,1685,1710]" pageId="7" pageNumber="528">
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<emphasis id="B90EEAEDFFA07779FF6FE8D5FEBCF909" bold="true" box="[151,317,1685,1710]" pageId="7" pageNumber="528">Type material</emphasis>
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.
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<typeStatus id="54C1885DFFA07779FEB4E8D5FE35F909" box="[332,436,1685,1710]" pageId="7" pageNumber="528" type="holotype">Holotype</typeStatus>
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labeled:
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<taxonomicName id="4C7A4D7CFFA07779FDE2E8D5FCAAF909" authorityName="Muona" authorityYear="2019" box="[538,811,1685,1710]" class="Insecta" family="Throscidae" genus="Pseudothroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="genus">
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<emphasis id="B90EEAEDFFA07779FDE2E8D5FCAAF909" box="[538,811,1685,1710]" italics="true" pageId="7" pageNumber="528">Pseudothroscus balticus</emphasis>
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</taxonomicName>
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n.sp/
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<typeStatus id="54C1885DFFA07779FC9BE8D6FC7DF909" box="[867,1020,1686,1710]" pageId="7" pageNumber="528" type="holotype">HOLOTYPE</typeStatus>
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/J.Muona des. 10-2015.
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</paragraph>
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<paragraph id="8BC536FFFFA07779FF3FE8FAFCC5F821" blockId="7.[151,1437,1685,1998]" pageId="7" pageNumber="528">
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Embedded in clear Baltic amber piece,
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<quantity id="4C829B1AFFA07779FD87E8FAFD4CF975" box="[639,717,1722,1746]" metricMagnitude="-2" metricUnit="m" metricValue="1.2" pageId="7" pageNumber="528" unit="mm" value="12.0">12 mm</quantity>
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<date id="FFC4103FFFA07779FD2CE8FBFD76F974" box="[724,759,1722,1747]" pageId="7" pageNumber="528">
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x
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<quantity id="4C829B1AFFA07779FD11E8FAFD76F974" box="[745,759,1722,1747]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" pageId="7" pageNumber="528" unit="mm" value="6.0">6</quantity>
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</date>
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mm, sex unknown. Stellate oak hairs, other plant matter and an unidentified psocopteran are present in the piece as well (
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<figureCitation id="13412A7AFFA07779FCCEE89DFC08F951" box="[822,905,1757,1782]" captionStart="FIGURE 22" captionStartId="18.[151,250,1887,1909]" captionTargetBox="[178,1417,1018,1876]" captionTargetId="figure@18.[178,1417,1018,1876]" captionTargetPageId="18" captionText="FIGURE 22. Pseudothroscus balticus n. sp., holotype." figureDoi="http://doi.org/10.5281/zenodo.3715818" httpUri="https://zenodo.org/record/3715818/files/figure.png" pageId="7" pageNumber="528">Fig. 22</figureCitation>
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).
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<typeStatus id="54C1885DFFA07779FC64E89FFC7FF950" box="[924,1022,1759,1783]" pageId="7" pageNumber="528" type="paratype">Paratype</typeStatus>
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of unknown sex embedded in a large clear piece of Baltic Amber,
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<quantity id="4C829B1AFFA07779FE21E942FDA9F8BD" box="[473,552,1794,1818]" metricMagnitude="-2" metricUnit="m" metricValue="2.8" pageId="7" pageNumber="528" unit="mm" value="28.0">28 mm</quantity>
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<date id="FFC4103FFFA07779FDD6E943FDDEF8BC" box="[558,607,1794,1819]" pageId="7" pageNumber="528" value="1959-10">
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x
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<quantity id="4C829B1AFFA07779FDBBE942FDDEF8BC" box="[579,607,1794,1819]" metricMagnitude="-2" metricUnit="m" metricValue="5.9" pageId="7" pageNumber="528" unit="mm" value="59.0">59</quantity>
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</date>
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mm. Some plant matter and two sciarid nematocerans are present as well. One of the dipterans is a perfectly preserved
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<taxonomicName id="4C7A4D7CFFA07779FD58E965FC9EF899" authorityName="Steffan" authorityYear="1969" box="[672,799,1829,1854]" class="Insecta" family="Sciaridae" genus="Lobosciara" higherTaxonomySource="GBIF" kingdom="Animalia" order="Diptera" pageId="7" pageNumber="528" phylum="Arthropoda" rank="genus">
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<emphasis id="B90EEAEDFFA07779FD58E965FC9EF899" box="[672,799,1829,1854]" italics="true" pageId="7" pageNumber="528">Lobosciara</emphasis>
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</taxonomicName>
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sp. male (Pekka Vilkamaa det.). Two further
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<typeStatus id="54C1885DFFA07779FAC9E967FA1DF898" box="[1329,1436,1831,1855]" pageId="7" pageNumber="528" type="paratype">paratypes</typeStatus>
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embedded together in rectangular Baltic Amber piece,
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<quantity id="4C829B1AFFA07779FCE4E90AFCEEF8C5" box="[796,879,1866,1890]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="7" pageNumber="528" unit="mm" value="27.0">27 mm</quantity>
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<date id="FFC4103FFFA07779FC82E90BFC31F8C5" box="[890,944,1866,1890]" pageId="7" pageNumber="528" value="2013-10">
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x
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<quantity id="4C829B1AFFA07779FC6CE90AFC31F8C5" box="[916,944,1866,1890]" metricMagnitude="-2" metricUnit="m" metricValue="1.3" pageId="7" pageNumber="528" unit="mm" value="13.0">13</quantity>
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</date>
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mm
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<date id="FFC4103FFFA07779FC0BE90BFB9BF8C4" box="[1011,1050,1866,1891]" pageId="7" pageNumber="528">
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x
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<quantity id="4C829B1AFFA07779FBF4E90AFB9BF8C4" box="[1036,1050,1866,1891]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="528" unit="mm" value="5.0">5</quantity>
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</date>
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mm, sex unknown. Age 50 My. Numerous hirsute plant remains are present as well (
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<figureCitation id="13412A7AFFA07779FD1BE92DFCB9F821" box="[739,824,1901,1926]" captionStart="FIGURE 22" captionStartId="18.[151,250,1887,1909]" captionTargetBox="[178,1417,1018,1876]" captionTargetId="figure@18.[178,1417,1018,1876]" captionTargetPageId="18" captionText="FIGURE 22. Pseudothroscus balticus n. sp., holotype." figureDoi="http://doi.org/10.5281/zenodo.3715818" httpUri="https://zenodo.org/record/3715818/files/figure.png" pageId="7" pageNumber="528">Fig. 22</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606574FFA07779FF3FE9D2FDACF869" pageId="7" pageNumber="528" type="diagnosis">
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<paragraph id="8BC536FFFFA07779FF3FE9D2FDACF869" blockId="7.[151,1437,1685,1998]" pageId="7" pageNumber="528">
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<emphasis id="B90EEAEDFFA07779FF3FE9D2FEBBF80C" bold="true" box="[199,314,1938,1963]" pageId="7" pageNumber="528">Diagnosis</emphasis>
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. Characterized by complete lateral pronotal carinae, large eyes, absence of tarsal grooves and small antennal fossae with delicate ridges.
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</paragraph>
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</subSubSection>
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<caption id="DF056677FFAF7776FF6FEA6FFCEBFBC3" ID-DOI="http://doi.org/10.5281/zenodo.3715782" ID-Zenodo-Dep="3715782" httpUri="https://zenodo.org/record/3715782/files/figure.png" pageId="8" pageNumber="529" startId="8.[151,250,1071,1093]" targetBox="[150,1436,193,1050]" targetPageId="8">
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<paragraph id="8BC536FFFFAF7776FF6FEA6FFCEBFBC3" blockId="8.[151,1436,1071,1124]" pageId="8" pageNumber="529">
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<emphasis id="B90EEAEDFFAF7776FF6FEA6FFE93FBE2" bold="true" box="[151,274,1071,1093]" pageId="8" pageNumber="529">FIGURE 5.</emphasis>
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<taxonomicName id="4C7A4D7CFFAF7776FEE0EA6FFD8FFBE2" authority="Muona, 2019" authorityName="Muona" authorityYear="2019" box="[280,526,1071,1093]" class="Insecta" family="Throscidae" genus="Pseudothroscus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="529" phylum="Arthropoda" rank="genus" status="sp. nov.">
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<emphasis id="B90EEAEDFFAF7776FEE0EA6FFD8FFBE2" box="[280,526,1071,1093]" italics="true" pageId="8" pageNumber="529">Pseudothroscus balticus</emphasis>
|
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</taxonomicName>
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<taxonomicNameLabel id="A23D5796FFAF7776FDECEA6FFDCDFBE2" box="[532,588,1071,1093]" pageId="8" pageNumber="529" rank="species">
|
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<emphasis id="B90EEAEDFFAF7776FDECEA6FFDC7FBE2" bold="true" box="[532,582,1071,1093]" pageId="8" pageNumber="529">n. sp</emphasis>
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.
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</taxonomicNameLabel>
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, ventral view. AC = antennal club; AF = antennal pit boarded by delicate ridge; L = labrum; LC = complete lateral pronotal ridge; MsT = simple mesotibia.
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</paragraph>
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</caption>
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<subSubSection id="C3606574FFAF7776FF3FEAD1FBC5FA9E" pageId="8" pageNumber="529" type="description">
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<paragraph id="8BC536FFFFAF7776FF3FEAD1FBC5FA9E" blockId="8.[151,1436,1169,1374]" pageId="8" pageNumber="529">
|
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<emphasis id="B90EEAEDFFAF7776FF3FEAD1FED0FB0D" bold="true" box="[199,337,1169,1194]" pageId="8" pageNumber="529">Description</emphasis>
|
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. Length 3.2 (HT) – 4.0 (PT) mm. Head and pronotum with relatively dense punctation, punctures mostly small, microsculpture weak (
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<figureCitation id="13412A7AFFAF7776FDD7EAF4FDFEFB6A" box="[559,639,1204,1229]" captionStart="FIGURE 4" captionStartId="7.[151,250,1482,1504]" captionTargetBox="[150,1436,802,1467]" captionTargetId="figure@7.[150,1436,802,1467]" captionTargetPageId="7" captionText="FIGURE 4. Pseudothroscus balticus n. sp., dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3715780" httpUri="https://zenodo.org/record/3715780/files/figure.png" pageId="8" pageNumber="529">Figs. 4</figureCitation>
|
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,
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<figureCitation id="13412A7AFFAF7776FD73EAF4FD1AFB6A" box="[651,667,1204,1229]" captionStart="FIGURE 5" captionStartId="8.[151,250,1071,1093]" captionTargetBox="[150,1436,193,1050]" captionTargetId="figure@8.[150,1436,193,1050]" captionTargetPageId="8" captionText="FIGURE 5. Pseudothroscus balticus n. sp., ventral view. AC = antennal club; AF = antennal pit boarded by delicate ridge; L = labrum; LC = complete lateral pronotal ridge; MsT = simple mesotibia." figureDoi="http://doi.org/10.5281/zenodo.3715782" httpUri="https://zenodo.org/record/3715782/files/figure.png" pageId="8" pageNumber="529">5</figureCitation>
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). Elytral striae deep, wide and strongly punctate, interstices twice as wide, very densely minutely punctate, convex (
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<figureCitation id="13412A7AFFAF7776FD3EEA99FC91FB56" box="[710,784,1241,1266]" captionStart="FIGURE 4" captionStartId="7.[151,250,1482,1504]" captionTargetBox="[150,1436,802,1467]" captionTargetId="figure@7.[150,1436,802,1467]" captionTargetPageId="7" captionText="FIGURE 4. Pseudothroscus balticus n. sp., dorsal view." figureDoi="http://doi.org/10.5281/zenodo.3715780" httpUri="https://zenodo.org/record/3715780/files/figure.png" pageId="8" pageNumber="529">Fig. 4</figureCitation>
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). Hypomera very densely punctate, presternum less so, metaventrite fairly sparsely punctate, punctures small. Abdomen mostly sparsely punctate, more densely towards apex, punctures large. Vestiture dense, on dorsum slightly erect, on abdomen not so.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3606574FFAF7776FF36EB05FD40FAF9" box="[206,705,1349,1374]" pageId="8" pageNumber="529" type="etymology">
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<paragraph id="8BC536FFFFAF7776FF36EB05FD40FAF9" blockId="8.[151,1436,1169,1374]" box="[206,705,1349,1374]" pageId="8" pageNumber="529">
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<emphasis id="B90EEAEDFFAF7776FF36EB05FECCFAF9" bold="true" box="[206,333,1349,1374]" pageId="8" pageNumber="529">Etymology</emphasis>
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. Known only from Baltic amber.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |