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<document id="5434ED773FA7BF843CD316675670BEC7" ID-DOI="10.5252/geodiversitas2019v41a5" ID-GBIF-Dataset="1dd366f3-9e6d-4dcf-a082-408a0ea91bc4" ID-ISSN="1638-9395" ID-Zenodo-Dep="3699932" ID-ZooBank="urn:lsid:zoobank.org:pub:33E8EF32-4BE4-4992-9917-8611CA0AD579" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1583514705026" checkinUser="valdenar" docAuthor="Sanders, Malcolm T., Merle, Didier &amp; Puillandre, Nicolas" docDate="2019" docId="03D387DF193BFFF0FF3BFA46FC0BF83F" docLanguage="en" docName="GEODIVERSITAS.41.5.247-265.pdf.imf" docOrigin="Geodiversitas 41 (5)" docStyle="DocumentStyle:F830B10FF475E64C1F1601E3B32DDC00.4:Geodiversitas.2018-.journal_article" docStyleId="F830B10FF475E64C1F1601E3B32DDC00" docStyleName="Geodiversitas.2018-.journal_article" docStyleVersion="4" docTitle="Aquitanobursa chipolana Sanders &amp; Merle &amp; Puillandre 2019, n. comb." docType="treatment" docVersion="5" lastPageNumber="258" masterDocId="FFEAFFA71936FFFDFF96FFF2FFC1FA0D" masterDocTitle="A review of fossil Bursidae and their use for phylogeny calibration" masterLastPageNumber="265" masterPageNumber="247" pageNumber="258" updateTime="1698725915702" updateUser="plazi" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="5D19C9A623810264DCC1619A5BF2F2F9">A review of fossil Bursidae and their use for phylogeny calibration</mods:title>
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<paragraph id="8BC536C9193BFFF0FF3BFA46FDC4FFE2" blockId="13.[173,735,1460,1520]" pageId="13" pageNumber="258">
<heading id="D08D81A5193BFFF0FF3BFA46FD1EFFC2" box="[173,735,1460,1487]" pageId="13" pageNumber="258">
<taxonomicName id="4C7A4D4A193BFFF0FF3BFA46FDB6FFC2" ID-CoL="7Q6LV" authority="(Schmelz, 1997)" authorityName="Sanders &amp; Merle &amp; Puillandre" authorityYear="2019" baseAuthorityName="Schmelz" baseAuthorityYear="1997" box="[173,631,1460,1487]" class="Gastropoda" family="Bursidae" genus="Aquitanobursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="258" phylum="Mollusca" rank="species" species="chipolana" status="comb. nov.">
<emphasis id="B90EEADB193BFFF0FF3BFA46FE7EFFC2" bold="true" box="[173,447,1460,1487]" italics="true" pageId="13" pageNumber="258">Aquitanobursa chipolana</emphasis>
(
<bibRefCitation id="EFEB4B38193BFFF0FE5BFA47FDAEFFC2" author="SCHMELZ G. W." box="[461,623,1461,1487]" pageId="13" pageNumber="258" pagination="105 - 108" refId="ref12815" refString="SCHMELZ G. W. 1997. - Notes on the fauna of the Chipola Formation - XLI. A reexamination of Bursa (Mollusca: Gastropoda) from the Chipola Formation, with the description of a new species. Tulane Studies in Geology and Paleontology 30: 105 - 108." type="journal article" year="1997">Schmelz, 1997</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A23D57A0193BFFF0FDE8FA47FD1EFFC2" box="[638,735,1461,1487]" pageId="13" pageNumber="258" rank="species">n. comb.</taxonomicNameLabel>
</heading>
(
<figureCitation id="13412A4C193BFFF0FE19FA27FE3CFFE2" box="[399,509,1493,1520]" captionStart="FIG" captionStartId="10.[132,142,1805,1823]" captionTargetBox="[236,1352,213,1781]" captionTargetId="figure@10.[700,1167,959,1334]" captionTargetPageId="10" captionText="FIG. 5. — Extinct Bursidae species:A,B, Marsupina judensis Beu,2010; NMB H 18308,holotype,late Miocene,Punta Judas, Pacific Costa Rica,reproduced from Beu (2010); C, D, “Bursa” landaui Harzhauser, 2009, NHMW 2007z0181/0029 (C) holotype and NHMW 2007z0181/0030 (D), both from the Aquitanian of Ras Tipuli, Lindi Bay,Tanzania,reproduced from Harzhauser (2009); E, Lampadopsis sangirana (Beu, 2005), RGM 456 230, holotype, late Pliocene of Kalibeng layers,Sangiran,central Java, reproduced from Beu (2005); F, G, Aquitanobursa chipolana (Schmelz, 1997) n. comb., USNM 647108, paratype, specimen illustrated by Vokes (1973: text-figs 2a, b), coral reef facies of Chipola Formation (late Early Miocene), loc. TU547, reproduced from Beu (2010); H, I, Marsupina freya (Olsson, 1932), PRI 2312, holotype, “Zorritos Miocene,” divide between Quebrada Conchudo Bravo and Quebrada Seca,Mancora,Peru;J, K, Olssonia chira (Olsson,1930) n.comb.,PRI 24257,holotype, Late Eocene/Oligocene of Chira Formation, Quercotilla, Peru; L-O, Olssonia yasila (Olsson, 1930) n. comb., PRI 24254 (L, M) and PRI 24255 (N, O), Middle Eocene Talara Formation, Yasila, Peru; P-S, Aquitanobursa morrisi (dArchiac &amp; Haime, 1853) n. comb., original representation of one of the syntypes by dArchiac &amp; Haime (1853) (P, R) and (NHMUK PI TG 27045) lectotype,from “Calcaire grossier jaune de la chaîne dHala” (Aquitanian?) Pakistan;T, Aquitanobursa amphitrites (Maury,1917) n. comb., PRI 28763, holotype, Cercado Fm (Late Miocene), Maurys bluff 3, Cercado de Mao, Dominican Republic, reproduced from Beu (2010). Scale bar: 2 cm." figureDoi="http://doi.org/10.5281/zenodo.3699942" httpUri="https://zenodo.org/record/3699942/files/figure.png" pageId="13" pageNumber="258">Fig. 5F, G</figureCitation>
)
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<treatmentCitation id="0ADB10D8193BFFF0FF12F9FCFDAFFC28" author="SCHMELZ G. W." box="[132,622,1549,1574]" page="105" pageId="13" pageNumber="258" year="1997">
<taxonomicName id="4C7A4D4A193BFFF0FF12F9FCFDAFFC28" ID-CoL="7QH99" authority="Schmelz, 1997: 105" authorityName="Schmelz" authorityPageNumber="105" authorityYear="1997" box="[132,622,1549,1574]" class="Gastropoda" family="Bursidae" genus="Bursa" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="258" phylum="Mollusca" rank="species" species="chipolana" subGenus="Bufonariella">
<emphasis id="B90EEADB193BFFF0FF12F9FCFF7BFC28" box="[132,186,1550,1573]" italics="true" pageId="13" pageNumber="258">Bursa</emphasis>
(
<emphasis id="B90EEADB193BFFF0FF5EF9FFFEFDFC28" box="[200,316,1549,1573]" italics="true" pageId="13" pageNumber="258">Bufonariella</emphasis>
)
<emphasis id="B90EEADB193BFFF0FEDCF9FFFE64FC28" box="[330,421,1549,1573]" italics="true" pageId="13" pageNumber="258">chipolana</emphasis>
<bibRefCitation id="EFEB4B38193BFFF0FE3CF9FCFDAFFC28" author="SCHMELZ G. W." box="[426,622,1549,1574]" pageId="13" pageNumber="258" pagination="105 - 108" refId="ref12815" refString="SCHMELZ G. W. 1997. - Notes on the fauna of the Chipola Formation - XLI. A reexamination of Bursa (Mollusca: Gastropoda) from the Chipola Formation, with the description of a new species. Tulane Studies in Geology and Paleontology 30: 105 - 108." type="journal article" year="1997">Schmelz, 1997: 105</bibRefCitation>
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, pl. 1, figs 1a-c; pl. 2, figs 1a-b, 2a-b. —
<treatmentCitation id="0ADB10D8193BFFF0FEE2F9DAFDDDFC4D" author="VOKES E. H." box="[372,540,1576,1600]" page="212" pageId="13" pageNumber="258" year="1997">
<bibRefCitation id="EFEB4B38193BFFF0FEE2F9DAFDDDFC4D" author="VOKES E. H." box="[372,540,1576,1600]" pageId="13" pageNumber="258" pagination="211 - 216" refId="ref13458" refString="VOKES E. H. 1997. - Notes on the fauna of the Chipola Formation - XLIII Additions to the molluscan fauna since 1947. Tulane Study in Geology and Paleontology 30: 211 - 216." type="journal article" year="1997">Vokes 1997: 212</bibRefCitation>
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.
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</paragraph>
<paragraph id="8BC536C9193BFFF0FF12F9AAFE58FC87" blockId="13.[132,774,1624,1675]" pageId="13" pageNumber="258">
<treatmentCitationGroup id="AB6A11E7193BFFF0FF12F9AAFE58FC87" pageId="13" pageNumber="258">
<taxonomicName id="4C7A4D4A193BFFF0FF12F9AAFE85FC7D" authority="" box="[132,324,1624,1648]" class="Gastropoda" family="Bursidae" genus="Bursa" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="258" phylum="Mollusca" rank="subGenus" subGenus="Bufonariella">
<emphasis id="B90EEADB193BFFF0FF12F9AAFF7BFC62" box="[132,186,1624,1647]" italics="true" pageId="13" pageNumber="258">Bursa</emphasis>
(
<emphasis id="B90EEADB193BFFF0FF5FF9AAFEFCFC7D" box="[201,317,1624,1648]" italics="true" pageId="13" pageNumber="258">Bufonariella</emphasis>
)
</taxonomicName>
<emphasis id="B90EEADB193BFFF0FEDDF9AAFE79FC7D" box="[331,440,1624,1648]" italics="true" pageId="13" pageNumber="258">pelouatensis</emphasis>
<treatmentCitation id="0ADB10D8193BFFF0FE44F9AAFDAFFC7D" author="VOKES E. H." box="[466,622,1624,1648]" page="97" pageId="13" pageNumber="258" year="1973">
<bibRefCitation id="EFEB4B38193BFFF0FE44F9AAFDAFFC7D" author="VOKES E. H." box="[466,622,1624,1648]" pageId="13" pageNumber="258" pagination="97 - 101" refId="ref13409" refString="VOKES E. H. 1973. - Notes on the fauna of the Chipola Formation- XIV. On the occurrence of Bursa (Mollusca: Gastropoda) with comments on the genus. Tulane Study in Geology and Paleontology 10 (2): 97 - 101." type="journal article" year="1973">Vokes 1973: 97</bibRefCitation>
</treatmentCitation>
, text-fig. 2 (not
<bibRefCitation id="EFEB4B38193BFFF0FF12F980FE4CFC87" author="COSSMANN M. &amp; PEYROT A." box="[132,397,1650,1675]" pageId="13" pageNumber="258" pagination="193 - 318" refId="ref10623" refString="COSSMANN M. &amp; PEYROT A. 1924. - Conchologie neogenique de l'Aquitaine (Suite). Actes de la Societe linneenne de Bordeaux 75 (3): 193 - 318. https: // biodiversitylibrary. org / page / 32207388" type="journal article" year="1924">Cossmann &amp; Peyrot, 1924</bibRefCitation>
).
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</paragraph>
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<treatmentCitationGroup id="AB6A11E7193BFFF0FF12F950FDB5FCB7" box="[132,628,1698,1722]" pageId="13" pageNumber="258">
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<emphasis id="B90EEADB193BFFF0FF12F950FEDAFCB7" box="[132,283,1698,1722]" italics="true" pageId="13" pageNumber="258">Bursa chipolana</emphasis>
</taxonomicName>
<treatmentCitation id="0ADB10D8193BFFF0FEA2F950FE7DFCB7" author="BEU A. G." box="[308,444,1698,1722]" page="44" pageId="13" pageNumber="258" year="2010">
<bibRefCitation id="EFEB4B38193BFFF0FEA2F950FE7DFCB7" author="BEU A. G." box="[308,444,1698,1722]" pageId="13" pageNumber="258" pagination="1 - 550" refId="ref10501" refString="BEU A. G. 2010. - Neogene tonnoidean gastropods of tropical and South America; a contribution to the Dominican Republic and Panama Paleontology projects and uplift of the Central American Isthmus. Bulletins of American Paleontology 377 - 378: 1 - 550." type="journal article" year="2010">Beu 2010: 44</bibRefCitation>
</treatmentCitation>
, pl. 1, figs 5-7, 10.
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</paragraph>
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<typeStatus id="54C1886B193BFFF0FF12F923FF77FCEB" box="[132,182,1745,1769]" pageId="13" pageNumber="258">TYPE</typeStatus>
LOCALITY. — Tulane University locality TU546, Chipola Formation (Burdigalian), Tenmile Creek,
<collectingRegion id="49BEF82B193BFFF0FD8AF91EFDA4FD09" box="[540,613,1772,1796]" country="United States of America" name="Florida" pageId="13" pageNumber="258">Florida</collectingRegion>
,
<collectingCountry id="F36D7659193BFFF0FDF8F91EFD35FD09" box="[622,756,1772,1796]" name="United States of America" pageId="13" pageNumber="258">United States</collectingCountry>
.
</paragraph>
<paragraph id="8BC536C9193BFFF0FF12F8E9FE0DFDE0" blockId="13.[132,775,1819,2029]" pageId="13" pageNumber="258">
TYPE MATERIAL. —
<taxonomicName id="4C7A4D4A193BFFF0FECCF8EEFD5BFD3E" authorityName="Schmelz" authorityYear="1997" box="[346,666,1819,1844]" class="Gastropoda" family="Bursidae" genus="Bursa" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="258" phylum="Mollusca" rank="species" species="chipolana" subGenus="Bufonariella">
<emphasis id="B90EEADB193BFFF0FECCF8EEFE55FD3E" box="[346,404,1820,1843]" italics="true" pageId="13" pageNumber="258">Bursa</emphasis>
(
<emphasis id="B90EEADB193BFFF0FE30F8E9FDE4FD3E" box="[422,549,1819,1843]" italics="true" pageId="13" pageNumber="258">Bufonariella</emphasis>
)
<emphasis id="B90EEADB193BFFF0FDA1F8E9FD5BFD3E" box="[567,666,1819,1843]" italics="true" pageId="13" pageNumber="258">chipolana</emphasis>
</taxonomicName>
,
<typeStatus id="54C1886B193BFFF0FD3CF8E9FCC6FD3E" box="[682,775,1819,1843]" pageId="13" pageNumber="258" type="holotype">holotype</typeStatus>
(UF 73199), figured
<typeStatus id="54C1886B193BFFF0FECAF8C4FE74FD43" box="[348,437,1846,1870]" pageId="13" pageNumber="258" type="paratype">paratype</typeStatus>
(USNM 647108), from TU547, west bank Chipola
<collectingRegion id="49BEF82B193BFFF0FED3F8A2FEBDFD65" box="[325,380,1872,1896]" country="Nigeria" name="Rivers" pageId="13" pageNumber="258">River</collectingRegion>
<quantity id="4C829B2C193BFFF0FE17F8A2FE03FD65" box="[385,450,1872,1896]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="13" pageNumber="258" unit="m" value="600.0">600 m</quantity>
upstream from Fourmile Creek, Calhoun Co. (specimen figured by
<bibRefCitation id="EFEB4B38193BFFF0FE64F899FDADFD8E" author="VOKES E. H." box="[498,620,1899,1923]" pageId="13" pageNumber="258" pagination="97 - 101" refId="ref13409" refString="VOKES E. H. 1973. - Notes on the fauna of the Chipola Formation- XIV. On the occurrence of Bursa (Mollusca: Gastropoda) with comments on the genus. Tulane Study in Geology and Paleontology 10 (2): 97 - 101." type="journal article" year="1973">Vokes 1973</bibRefCitation>
: text-figs 2a-b, refigured in good quality by
<bibRefCitation id="EFEB4B38193BFFF0FE3CF874FDCFFD90" author="BEU A. G." box="[426,526,1925,1949]" pageId="13" pageNumber="258" pagination="1 - 550" refId="ref10501" refString="BEU A. G. 2010. - Neogene tonnoidean gastropods of tropical and South America; a contribution to the Dominican Republic and Panama Paleontology projects and uplift of the Central American Isthmus. Bulletins of American Paleontology 377 - 378: 1 - 550." type="journal article" year="2010">Beu 2010</bibRefCitation>
: pl. 1, figs 6, 7);
<materialsCitation id="3B123C94193BFFF0FD29F877FE06FDE0" ID-GBIF-Occurrence="2574964551" location="Calhoun Co" municipality="Tenmile Creek" pageId="13" pageNumber="258" specimenCount="1" stateProvince="Rivers" typeStatus="paratype">
figured
<typeStatus id="54C1886B193BFFF0FF12F852FF1CFDB5" box="[132,221,1952,1976]" pageId="13" pageNumber="258" type="paratype">paratype</typeStatus>
(UF 73200), from TU951,
<collectingMunicipality id="6BA1ACB3193BFFF0FD93F852FD63FDB5" box="[517,674,1952,1976]" pageId="13" pageNumber="258">Tenmile Creek</collectingMunicipality>
,
<location id="8EA56012193BFFF0FD3BF852FF66FDDF" LSID="urn:lsid:plazi:treatment:03D387DF193BFFF0FF3BFA46FC0BF83F:8EA56012193BFFF0FD3BF852FF66FDDF" municipality="Tenmile Creek" name="Calhoun Co" pageId="13" pageNumber="258">Calhoun Co</location>
.; nine further
<typeStatus id="54C1886B193BFFF0FEA7F849FE4EFDDE" box="[305,399,1979,2003]" pageId="13" pageNumber="258" type="paratype">paratypes</typeStatus>
from localities on or near the
<location id="8EA56012193BFFF0FD20F848FCC6FDDF" LSID="urn:lsid:plazi:treatment:03D387DF193BFFF0FF3BFA46FC0BF83F:8EA56012193BFFF0FD20F848FCC6FDDF" box="[694,775,1978,2002]" name="Chipola" pageId="13" pageNumber="258" stateProvince="Rivers">Chipola</location>
<collectingRegion id="49BEF82B193BFFF0FF12F827FF7AFDE0" box="[132,187,2005,2029]" country="Nigeria" name="Rivers" pageId="13" pageNumber="258">River</collectingRegion>
listed by
<bibRefCitation id="EFEB4B38193BFFF0FEB7F827FE06FDE0" author="SCHMELZ G. W." box="[289,455,2005,2029]" pageId="13" pageNumber="258" pagination="105 - 108" refId="ref12815" refString="SCHMELZ G. W. 1997. - Notes on the fauna of the Chipola Formation - XLI. A reexamination of Bursa (Mollusca: Gastropoda) from the Chipola Formation, with the description of a new species. Tulane Studies in Geology and Paleontology 30: 105 - 108." type="journal article" year="1997">Schmelz (1997)</bibRefCitation>
</materialsCitation>
.
</paragraph>
<paragraph id="8BC536C9193BFFF0FCBBFF2AFC76FB4D" blockId="13.[813,1456,216,320]" pageId="13" pageNumber="258">
GEOGRAPHIC AND STRATIGRAPHIC OCCURRENCE. —
<taxonomicName id="4C7A4D4A193BFFF0FAB3FF2BFC49FB06" authorityName="Sanders &amp; Merle &amp; Puillandre" authorityYear="2019" baseAuthorityName="Schmelz" baseAuthorityYear="1997" class="Gastropoda" family="Bursidae" genus="Aquitanobursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="258" phylum="Mollusca" rank="species" species="chipolana" status="comb. nov.">
<emphasis id="B90EEADB193BFFF0FAB3FF2BFC49FB06" italics="true" pageId="13" pageNumber="258">Aquitanobursa chipolana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D57A0193BFFF0FC19FF06FC29FB06" box="[911,1000,243,267]" pageId="13" pageNumber="258" rank="species">n. comb.</taxonomicNameLabel>
is recorded only from the Chipola Formation (Burdigalian) on and near the Chipola River, Calhoun Co.,
<collectingRegion id="49BEF82B193BFFF0FAF5FEFCFA6AFB2B" box="[1379,1451,270,294]" country="United States of America" name="Florida" pageId="13" pageNumber="258">Florida</collectingRegion>
,
<collectingCountry id="F36D7659193BFFF0FCBBFEDAFC73FB4D" box="[813,946,296,320]" name="United States of America" pageId="13" pageNumber="258">United States</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C3606542193BFFF0FCBBFE8AFC0BF83F" pageId="13" pageNumber="258" type="discussion">
<paragraph id="8BC536C9193BFFF0FCBBFE8AFC50FB9C" blockId="13.[813,1455,376,563]" box="[813,913,376,402]" pageId="13" pageNumber="258">REMARKS</paragraph>
<paragraph id="8BC536C9193BFFF0FCBBFE6AFC0BF83F" blockId="13.[813,1455,376,563]" pageId="13" pageNumber="258">
<taxonomicName id="4C7A4D4A193BFFF0FCBBFE6AFBFFFBBF" authorityName="Sanders &amp; Merle &amp; Puillandre" authorityYear="2019" baseAuthorityName="Schmelz" baseAuthorityYear="1997" box="[813,1086,408,434]" class="Gastropoda" family="Bursidae" genus="Aquitanobursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="258" phylum="Mollusca" rank="species" species="chipolana" status="comb. nov.">
<emphasis id="B90EEADB193BFFF0FCBBFE6AFBFFFBBF" box="[813,1086,408,434]" italics="true" pageId="13" pageNumber="258">Aquitanobursa chipolana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D57A0193BFFF0FBD2FE6BFB69FBBF" box="[1092,1192,408,435]" pageId="13" pageNumber="258" rank="species">n. comb.</taxonomicNameLabel>
seems to have a slightly greater number of primary cords than other species referred to
<taxonomicName id="4C7A4D4A193BFFF0FC24FE2AFB99FBFF" authority="Sanders &amp; Merle &amp; Puillandre, 2019" authorityName="Sanders &amp; Merle &amp; Puillandre" authorityYear="2019" box="[946,1112,472,498]" class="Gastropoda" family="Bursidae" genus="Aquitanobursa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Littorinimorpha" pageId="13" pageNumber="258" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B90EEADB193BFFF0FC24FE2AFB99FBFF" box="[946,1112,472,498]" italics="true" pageId="13" pageNumber="258">Aquitanobursa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D57A0193BFFF0FBF7FE2BFB72FBFE" box="[1121,1203,473,499]" pageId="13" pageNumber="258" rank="genus">n. gen.</taxonomicNameLabel>
This species possibly belongs in another genus, for which a new name needs to be created.
</paragraph>
</subSubSection>
</treatment>
</document>