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<mods:title id="39C792F54F7B2101FD58C6CDD929BE0E">A taxonomic study on the nominate subgenus Macrolycus Waterhouse, 1878 from China (Coleoptera, Lycidae)</mods:title>
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Key to the species of
<taxonomicName id="4C7A4D4BFF92FF98CAF9FA82037BFACF" authorityName="Waterhouse" authorityYear="1878" box="[413,555,1294,1321]" class="Insecta" family="Lycidae" genus="Macrolycus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="364" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B90EEADAFF92FF98CAF9FA82037BFACF" bold="true" box="[413,555,1294,1321]" italics="true" pageId="6" pageNumber="364">Macrolycus</emphasis>
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<taxonomicNameLabel id="A23D57A1FF92FF98C956FA830323FACF" box="[562,627,1295,1321]" pageId="6" pageNumber="364" sensu="stricto">s. str.</taxonomicNameLabel>
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<paragraph id="8BC536C8FF92FF98CBF3FABF0074FAAB" blockId="6.[151,627,1294,1357]" box="[151,292,1331,1357]" pageId="6" pageNumber="364">(males only)</paragraph>
<paragraph id="8BC536C8FF92FF98CBF3FAF404CCFA6B" blockId="6.[151,1437,1400,2037]" box="[151,1436,1400,1422]" pageId="6" pageNumber="364">
1. Aedeagus with strongly developed parameres, at least 1/3 length of phallus (e.g.
<figureCitation id="13412A4DFF92FF98C88DFAF40565FA6B" box="[1001,1077,1400,1422]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[216,1373,185,1909]" captionTargetId="figure-25@5.[205,1383,181,1919]" captionTargetPageId="5" captionText="FIGURE 3.Aedeagi of Macrolycus species, dorsal views:AC. M. oreophilus Kazantsev, 2002; DF. M. gansuensis Kazantsev, 2002; GI. M. bowringi Waterhouse, 1878; JL. M. subapicis sp. nov. (holotype). A, D, G, J: dorsal views; B, E, H, K: ventral views; C, F, I, L: lateral views. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821200" httpUri="https://zenodo.org/record/10821200/files/figure.png" pageId="6" pageNumber="364">Fig. 3G</figureCitation>
;
<bibRefCitation id="EFEB4B39FF92FF98CF5BFAF40594FA6B" author="Li, Y. &amp; Pang, H. &amp; Bocak, L." box="[1087,1220,1400,1422]" pageId="6" pageNumber="364" pagination="44 - 61" refId="ref5696" refString="Li, Y., Pang, H. &amp; Bocak, L. (2012) New species of Macrolycus Waterhouse, 1878 from China and Laos, with a checklist of the genus (Coleoptera: Lycidae). Zootaxa, 3232, 44 - 61. https: // doi. org / 10.11646 / zootaxa. 3232.1.2" type="journal article" year="2012">
Li
<emphasis id="B90EEADAFF92FF98CF3EFAF405DBFA6B" box="[1114,1163,1400,1421]" italics="true" pageId="6" pageNumber="364">et al.</emphasis>
2012
</bibRefCitation>
: fig. 37).......... 2
</paragraph>
<paragraph id="8BC536C8FF92FF98CBF3FA1904CCFA21" blockId="6.[151,1437,1400,2037]" pageId="6" pageNumber="364">
- Aedeagus with moderately developed parameres, less than 1/4 length of phallus (e.g.
<figureCitation id="13412A4DFF92FF98CF77FA1905C8FA4C" box="[1043,1176,1429,1451]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[216,1373,185,1909]" captionTargetId="figure-25@5.[205,1383,181,1919]" captionTargetPageId="5" captionText="FIGURE 3.Aedeagi of Macrolycus species, dorsal views:AC. M. oreophilus Kazantsev, 2002; DF. M. gansuensis Kazantsev, 2002; GI. M. bowringi Waterhouse, 1878; JL. M. subapicis sp. nov. (holotype). A, D, G, J: dorsal views; B, E, H, K: ventral views; C, F, I, L: lateral views. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821200" httpUri="https://zenodo.org/record/10821200/files/figure.png" pageId="6" pageNumber="364">Figs 3B, E, K</figureCitation>
;
<bibRefCitation id="EFEB4B39FF92FF98CFC6FA190410FA4C" author="Kazantsev, S. V." box="[1186,1344,1429,1450]" pageId="6" pageNumber="364" pagination="99 - 109" refId="ref5410" refString="Kazantsev, S. V. (2001) Macrolycus Waterhouse, 1878 (Coleoptera: Lycidae) of continental China. Elytron, 14 (2000), 99 - 109." type="journal article" year="2001">Kazantsev 2001</bibRefCitation>
: figs 2, 5; 2002: fig. 2, 6;
<bibRefCitation id="EFEB4B39FF92FF98CA12FA3E00ABFA21" author="Li, Y. &amp; Pang, H. &amp; Bocak, L." box="[374,507,1458,1480]" pageId="6" pageNumber="364" pagination="44 - 61" refId="ref5696" refString="Li, Y., Pang, H. &amp; Bocak, L. (2012) New species of Macrolycus Waterhouse, 1878 from China and Laos, with a checklist of the genus (Coleoptera: Lycidae). Zootaxa, 3232, 44 - 61. https: // doi. org / 10.11646 / zootaxa. 3232.1.2" type="journal article" year="2012">
Li
<emphasis id="B90EEADAFF92FF98CAF5FA3F0092FA21" box="[401,450,1458,1480]" italics="true" pageId="6" pageNumber="364">et al.</emphasis>
2012
</bibRefCitation>
: fig. 34)..................................................................... 3
</paragraph>
<paragraph id="8BC536C8FF92FF98CBF3FA5C04CCF9E4" blockId="6.[151,1437,1400,2037]" pageId="6" pageNumber="364">
2. Phallus present with a ridge along the whole length of lateral side (
<bibRefCitation id="EFEB4B39FF92FF98C807FA5C02BAFA03" author="Li, Y. &amp; Pang, H. &amp; Bocak, L." box="[867,1002,1488,1510]" pageId="6" pageNumber="364" pagination="44 - 61" refId="ref5696" refString="Li, Y., Pang, H. &amp; Bocak, L. (2012) New species of Macrolycus Waterhouse, 1878 from China and Laos, with a checklist of the genus (Coleoptera: Lycidae). Zootaxa, 3232, 44 - 61. https: // doi. org / 10.11646 / zootaxa. 3232.1.2" type="journal article" year="2012">
Li
<emphasis id="B90EEADAFF92FF98C8E4FA5C02E1FA03" box="[896,945,1488,1509]" italics="true" pageId="6" pageNumber="364">et al.</emphasis>
2012
</bibRefCitation>
: fig. 36).....................................................................................................
<taxonomicName id="4C7A4D4BFF92FF98CF22FA6104CCF9E4" authority="Li, Bocak &amp; Pang, 2012" authorityName="Li, Bocak &amp; Pang" authorityYear="2012" box="[1094,1436,1516,1539]" class="Insecta" family="Lycidae" genus="Macrolycus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="364" phylum="Arthropoda" rank="species" species="luteus">
<emphasis id="B90EEADAFF92FF98CF22FA6104CCF9E4" bold="true" box="[1094,1436,1516,1539]" pageId="6" pageNumber="364">
<emphasis id="B90EEADAFF92FF98CF22FA6105F4F9E4" bold="true" box="[1094,1188,1516,1538]" italics="true" pageId="6" pageNumber="364">M. luteus</emphasis>
<bibRefCitation id="EFEB4B39FF92FF98CFCEFA6104CCF9E4" author="Li, Y. &amp; Pang, H. &amp; Bocak, L." box="[1194,1436,1517,1539]" pageId="6" pageNumber="364" pagination="44 - 61" refId="ref5696" refString="Li, Y., Pang, H. &amp; Bocak, L. (2012) New species of Macrolycus Waterhouse, 1878 from China and Laos, with a checklist of the genus (Coleoptera: Lycidae). Zootaxa, 3232, 44 - 61. https: // doi. org / 10.11646 / zootaxa. 3232.1.2" type="journal article" year="2012">Li, Bocak &amp; Pang, 2012</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="8BC536C8FF92FF98CBF3F98704CCF9F9" blockId="6.[151,1437,1400,2037]" box="[151,1436,1546,1568]" pageId="6" pageNumber="364">
- Phallus smooth and absent with ridge on lateral side (
<figureCitation id="13412A4DFF92FF98C987F9860274F9F9" box="[739,804,1546,1568]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[216,1373,185,1909]" captionTargetId="figure-25@5.[205,1383,181,1919]" captionTargetPageId="5" captionText="FIGURE 3.Aedeagi of Macrolycus species, dorsal views:AC. M. oreophilus Kazantsev, 2002; DF. M. gansuensis Kazantsev, 2002; GI. M. bowringi Waterhouse, 1878; JL. M. subapicis sp. nov. (holotype). A, D, G, J: dorsal views; B, E, H, K: ventral views; C, F, I, L: lateral views. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821200" httpUri="https://zenodo.org/record/10821200/files/figure.png" pageId="6" pageNumber="364">Fig. 3I</figureCitation>
).........................
<taxonomicName id="4C7A4D4BFF92FF98CF02F98704CCF9F9" authority="Waterhouse, 1878" authorityName="Waterhouse" authorityYear="1878" box="[1126,1436,1546,1568]" class="Insecta" family="Lycidae" genus="Macrolycus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="364" phylum="Arthropoda" rank="species" species="bowringi">
<emphasis id="B90EEADAFF92FF98CF02F98704CCF9F9" bold="true" box="[1126,1436,1546,1568]" pageId="6" pageNumber="364">
<emphasis id="B90EEADAFF92FF98CF02F98705B0F9C6" bold="true" box="[1126,1248,1546,1568]" italics="true" pageId="6" pageNumber="364">M. bowringi</emphasis>
<bibRefCitation id="EFEB4B39FF92FF98CF82F98604CCF9F9" author="Waterhouse, C. O." box="[1254,1436,1546,1567]" pageId="6" pageNumber="364" pagination="95 - 118" refId="ref6154" refString="Waterhouse, C. O. (1878) On the different forms occurring in the Coleopterous family Lycidae, with descriptions of new genera and species. Transactions of the Entomological Society of London, 26, 95 - 118. https: // doi. org / 10.1111 / j. 1365 - 2311.1878. tb 01944. x" type="journal article" year="1878">Waterhouse, 1878</bibRefCitation>
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</paragraph>
<paragraph id="8BC536C8FF92FF98CBF3F9A404CCF991" blockId="6.[151,1437,1400,2037]" pageId="6" pageNumber="364">
3. Parameres shell-shaped, with lateral margins nearly straight or feebly protuberant in lateral view....
<taxonomicName id="4C7A4D4BFF92FF98CFD6F9A40016F9BC" authority="Pic, 1935 Parameres" authorityName="Pic" authorityYear="1935" class="Insecta" family="Lycidae" genus="Macrolycus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="364" phylum="Arthropoda" rank="species" species="jeanvoinei">
<emphasis id="B90EEADAFF92FF98CFD6F9A404CCF9DA" bold="true" box="[1202,1436,1575,1597]" pageId="6" pageNumber="364">
<emphasis id="B90EEADAFF92FF98CFD6F9A4046BF9DA" bold="true" box="[1202,1339,1575,1597]" italics="true" pageId="6" pageNumber="364">M. jeanvoinei</emphasis>
<bibRefCitation id="EFEB4B39FF92FF98CE24F9AB04CCF9DA" author="Pic, M." box="[1344,1436,1575,1597]" pageId="6" pageNumber="364" pagination="109 - 115" refId="ref6127" refString="Pic, M. (1935) Contribution al'etude des coleopteres malacodermes. Annales de la Societe Entomologique de France, 104, 109 - 115." type="journal article" year="1935">Pic, 1935</bibRefCitation>
</emphasis>
Parameres
</taxonomicName>
spoon-shaped, with lateral margins widely emarginate in lateral view (e.g.,
<figureCitation id="13412A4DFF92FF98CF4FF9C905FEF9BC" box="[1067,1198,1605,1627]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[216,1373,185,1909]" captionTargetId="figure-25@5.[205,1383,181,1919]" captionTargetPageId="5" captionText="FIGURE 3.Aedeagi of Macrolycus species, dorsal views:AC. M. oreophilus Kazantsev, 2002; DF. M. gansuensis Kazantsev, 2002; GI. M. bowringi Waterhouse, 1878; JL. M. subapicis sp. nov. (holotype). A, D, G, J: dorsal views; B, E, H, K: ventral views; C, F, I, L: lateral views. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821200" httpUri="https://zenodo.org/record/10821200/files/figure.png" pageId="6" pageNumber="364">Figs 3C, F, L</figureCitation>
;
<bibRefCitation id="EFEB4B39FF92FF98CFDDF9C90408F9BC" author="Kazantsev, S. V." box="[1209,1368,1605,1626]" pageId="6" pageNumber="364" pagination="99 - 109" refId="ref5410" refString="Kazantsev, S. V. (2001) Macrolycus Waterhouse, 1878 (Coleoptera: Lycidae) of continental China. Elytron, 14 (2000), 99 - 109." type="journal article" year="2001">Kazantsev 2001</bibRefCitation>
: fig. 6; 2002: figs 3, 7;
<bibRefCitation id="EFEB4B39FF92FF98CA1DF9EE00AFF991" author="Li, Y. &amp; Pang, H. &amp; Bocak, L." box="[377,511,1634,1656]" pageId="6" pageNumber="364" pagination="44 - 61" refId="ref5696" refString="Li, Y., Pang, H. &amp; Bocak, L. (2012) New species of Macrolycus Waterhouse, 1878 from China and Laos, with a checklist of the genus (Coleoptera: Lycidae). Zootaxa, 3232, 44 - 61. https: // doi. org / 10.11646 / zootaxa. 3232.1.2" type="journal article" year="2012">
Li
<emphasis id="B90EEADAFF92FF98CAF1F9EF0095F991" box="[405,453,1634,1656]" italics="true" pageId="6" pageNumber="364">et al.</emphasis>
2012
</bibRefCitation>
: fig. 41)..................................................................... 4
</paragraph>
<paragraph id="8BC536C8FF92FF98CBF3F90C04CCF972" blockId="6.[151,1437,1400,2037]" box="[151,1436,1663,1686]" pageId="6" pageNumber="364">
4. Phallus of aedeagus with tetraschistic at apex (
<bibRefCitation id="EFEB4B39FF92FF98C9C5F90C0278F973" author="Li, Y. &amp; Pang, H. &amp; Bocak, L." box="[673,808,1664,1686]" pageId="6" pageNumber="364" pagination="44 - 61" refId="ref5696" refString="Li, Y., Pang, H. &amp; Bocak, L. (2012) New species of Macrolycus Waterhouse, 1878 from China and Laos, with a checklist of the genus (Coleoptera: Lycidae). Zootaxa, 3232, 44 - 61. https: // doi. org / 10.11646 / zootaxa. 3232.1.2" type="journal article" year="2012">
Li
<emphasis id="B90EEADAFF92FF98C9DAF90C03BFF973" box="[702,751,1664,1685]" italics="true" pageId="6" pageNumber="364">et al.</emphasis>
2012
</bibRefCitation>
: fig. 34)............
<taxonomicName id="4C7A4D4BFF92FF98CF6BF90C04CCF972" authority="Li, Bocak &amp; Pang, 2012" authorityName="Li, Bocak &amp; Pang" authorityYear="2012" box="[1039,1436,1663,1685]" class="Insecta" family="Lycidae" genus="Macrolycus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="364" phylum="Arthropoda" rank="species" species="quadrifidus">
<emphasis id="B90EEADAFF92FF98CF6BF90C04CCF972" bold="true" box="[1039,1436,1663,1685]" pageId="6" pageNumber="364">
<emphasis id="B90EEADAFF92FF98CF6BF90C05F4F973" bold="true" box="[1039,1188,1663,1685]" italics="true" pageId="6" pageNumber="364">M. quadrifidus</emphasis>
<bibRefCitation id="EFEB4B39FF92FF98CFCEF9F304CCF972" author="Li, Y. &amp; Pang, H. &amp; Bocak, L." box="[1194,1436,1663,1685]" pageId="6" pageNumber="364" pagination="44 - 61" refId="ref5696" refString="Li, Y., Pang, H. &amp; Bocak, L. (2012) New species of Macrolycus Waterhouse, 1878 from China and Laos, with a checklist of the genus (Coleoptera: Lycidae). Zootaxa, 3232, 44 - 61. https: // doi. org / 10.11646 / zootaxa. 3232.1.2" type="journal article" year="2012">Li, Bocak &amp; Pang, 2012</bibRefCitation>
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BC536C8FF92FF98CBF3F91104CCF936" blockId="6.[151,1437,1400,2037]" pageId="6" pageNumber="364">
- Phallus of aedeagus present with two small processes at apex (e.g.
<figureCitation id="13412A4DFF92FF98C800F91102BAF954" box="[868,1002,1693,1715]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[216,1373,185,1909]" captionTargetId="figure-25@5.[205,1383,181,1919]" captionTargetPageId="5" captionText="FIGURE 3.Aedeagi of Macrolycus species, dorsal views:AC. M. oreophilus Kazantsev, 2002; DF. M. gansuensis Kazantsev, 2002; GI. M. bowringi Waterhouse, 1878; JL. M. subapicis sp. nov. (holotype). A, D, G, J: dorsal views; B, E, H, K: ventral views; C, F, I, L: lateral views. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821200" httpUri="https://zenodo.org/record/10821200/files/figure.png" pageId="6" pageNumber="364">Figs 3B, E, K</figureCitation>
;
<bibRefCitation id="EFEB4B39FF92FF98C890F91105C2F954" author="Kazantsev, S. V." box="[1012,1170,1693,1714]" pageId="6" pageNumber="364" pagination="99 - 109" refId="ref5410" refString="Kazantsev, S. V. (2001) Macrolycus Waterhouse, 1878 (Coleoptera: Lycidae) of continental China. Elytron, 14 (2000), 99 - 109." type="journal article" year="2001">Kazantsev 2001</bibRefCitation>
: figs 2, 5; 2002: fig. 2, 6). ................................................................................................... 5
</paragraph>
<paragraph id="8BC536C8FF92FF98CBF3F95B04CCF8EC" blockId="6.[151,1437,1400,2037]" pageId="6" pageNumber="364">
5. Antennomere III triangular in male (
<figureCitation id="13412A4DFF92FF98C926F95403DDF90B" box="[578,653,1752,1774]" captionStart="FIGURE 2" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[215,1373,190,1908]" captionTargetId="figure-25@3.[204,1383,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Male habitus of Macrolycus species, dorsal views: A. M. oreophilus Kazantsev, 2002; B. M. gansuensis Kazantsev, 2002; C. M. bowringi Waterhouse, 1878; D. M. subapicis sp. nov. (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821196" httpUri="https://zenodo.org/record/10821196/files/figure.png" pageId="6" pageNumber="364">Fig. 2B</figureCitation>
;
<bibRefCitation id="EFEB4B39FF92FF98C9FCF9540266F90B" author="Kazantsev, S. V." box="[664,822,1752,1773]" pageId="6" pageNumber="364" pagination="253 - 263" refId="ref5443" refString="Kazantsev, S. V. (2002) New and little known species of Lycidae (Coleoptera) from China. Russian Entomological Journal, 11, 253 - 263." type="journal article" year="2002">Kazantsev 2002</bibRefCitation>
: fig. 1); aedeagus with apical part of phallus expanded distad in ventral view (
<figureCitation id="13412A4DFF92FF98CA02F97900E1F8EC" box="[358,433,1781,1803]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[216,1373,185,1909]" captionTargetId="figure-25@5.[205,1383,181,1919]" captionTargetPageId="5" captionText="FIGURE 3.Aedeagi of Macrolycus species, dorsal views:AC. M. oreophilus Kazantsev, 2002; DF. M. gansuensis Kazantsev, 2002; GI. M. bowringi Waterhouse, 1878; JL. M. subapicis sp. nov. (holotype). A, D, G, J: dorsal views; B, E, H, K: ventral views; C, F, I, L: lateral views. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821200" httpUri="https://zenodo.org/record/10821200/files/figure.png" pageId="6" pageNumber="364">Fig. 3E</figureCitation>
;
<bibRefCitation id="EFEB4B39FF92FF98CADEF9790308F8EC" author="Kazantsev, S. V." box="[442,600,1781,1802]" pageId="6" pageNumber="364" pagination="253 - 263" refId="ref5443" refString="Kazantsev, S. V. (2002) New and little known species of Lycidae (Coleoptera) from China. Russian Entomological Journal, 11, 253 - 263." type="journal article" year="2002">Kazantsev 2002</bibRefCitation>
: fig. 2).....................................
<taxonomicName id="4C7A4D4BFF92FF98CF06F97904CCF8EC" authority="Kazantsev, 2002" authorityName="Kazantsev" authorityYear="2002" box="[1122,1436,1781,1802]" class="Insecta" family="Lycidae" genus="Macrolycus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="364" phylum="Arthropoda" rank="species" species="gansuensis">
<emphasis id="B90EEADAFF92FF98CF06F97904CCF8EC" bold="true" box="[1122,1436,1781,1802]" pageId="6" pageNumber="364">
<emphasis id="B90EEADAFF92FF98CF06F97905A1F8EC" bold="true" box="[1122,1265,1781,1802]" italics="true" pageId="6" pageNumber="364">M. gansuensis</emphasis>
<bibRefCitation id="EFEB4B39FF92FF98CF93F97904CCF8EC" author="Kazantsev, S. V." box="[1271,1436,1781,1802]" pageId="6" pageNumber="364" pagination="253 - 263" refId="ref5443" refString="Kazantsev, S. V. (2002) New and little known species of Lycidae (Coleoptera) from China. Russian Entomological Journal, 11, 253 - 263." type="journal article" year="2002">Kazantsev, 2002</bibRefCitation>
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BC536C8FF92FF98CBF3F89F04CCF8A0" blockId="6.[151,1437,1400,2037]" pageId="6" pageNumber="364">
- Antennomere III lamellate in male (e.g.,
<figureCitation id="13412A4DFF92FF98C918F89E03BEF8CE" box="[636,750,1810,1832]" captionStart="FIGURE 2" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[215,1373,190,1908]" captionTargetId="figure-25@3.[204,1383,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Male habitus of Macrolycus species, dorsal views: A. M. oreophilus Kazantsev, 2002; B. M. gansuensis Kazantsev, 2002; C. M. bowringi Waterhouse, 1878; D. M. subapicis sp. nov. (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821196" httpUri="https://zenodo.org/record/10821196/files/figure.png" pageId="6" pageNumber="364">Figs 2A, D</figureCitation>
;
<bibRefCitation id="EFEB4B39FF92FF98C99FF89F02CCF8C1" author="Kazantsev, S. V." box="[763,924,1810,1832]" pageId="6" pageNumber="364" pagination="99 - 109" refId="ref5410" refString="Kazantsev, S. V. (2001) Macrolycus Waterhouse, 1878 (Coleoptera: Lycidae) of continental China. Elytron, 14 (2000), 99 - 109." type="journal article" year="2001">Kazantsev 2001</bibRefCitation>
: figs 1, 4); aedeagus with apical part of apical part progressively constricted distad in ventral view (e.g.
<figureCitation id="13412A4DFF92FF98C9BAF8BC021AF8A3" box="[734,842,1840,1862]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[216,1373,185,1909]" captionTargetId="figure-25@5.[205,1383,181,1919]" captionTargetPageId="5" captionText="FIGURE 3.Aedeagi of Macrolycus species, dorsal views:AC. M. oreophilus Kazantsev, 2002; DF. M. gansuensis Kazantsev, 2002; GI. M. bowringi Waterhouse, 1878; JL. M. subapicis sp. nov. (holotype). A, D, G, J: dorsal views; B, E, H, K: ventral views; C, F, I, L: lateral views. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821200" httpUri="https://zenodo.org/record/10821200/files/figure.png" pageId="6" pageNumber="364">Figs 3B, K</figureCitation>
;
<bibRefCitation id="EFEB4B39FF92FF98C830F8BC02A2F8A3" author="Kazantsev, S. V." box="[852,1010,1840,1861]" pageId="6" pageNumber="364" pagination="99 - 109" refId="ref5410" refString="Kazantsev, S. V. (2001) Macrolycus Waterhouse, 1878 (Coleoptera: Lycidae) of continental China. Elytron, 14 (2000), 99 - 109." type="journal article" year="2001">Kazantsev 2001</bibRefCitation>
: figs 2, 5, 2002: fig. 6)................ 6
</paragraph>
<paragraph id="8BC536C8FF92FF98CBF3F8C104CCF899" blockId="6.[151,1437,1400,2037]" pageId="6" pageNumber="364">
6. Lamella of antennomere III 0.8 times as long as joint itself (
<bibRefCitation id="EFEB4B39FF92FF98C84EF8C1029AF884" author="Kazantsev, S. V." box="[810,970,1869,1890]" pageId="6" pageNumber="364" pagination="99 - 109" refId="ref5410" refString="Kazantsev, S. V. (2001) Macrolycus Waterhouse, 1878 (Coleoptera: Lycidae) of continental China. Elytron, 14 (2000), 99 - 109." type="journal article" year="2001">Kazantsev 2001</bibRefCitation>
: fig. 1); subapical part of phallus feebly inflated laterally in ventral view, 2.5 times as wide as basal part (
<bibRefCitation id="EFEB4B39FF92FF98C865F8E602F0F899" author="Kazantsev, S. V." box="[769,928,1898,1919]" pageId="6" pageNumber="364" pagination="99 - 109" refId="ref5410" refString="Kazantsev, S. V. (2001) Macrolycus Waterhouse, 1878 (Coleoptera: Lycidae) of continental China. Elytron, 14 (2000), 99 - 109." type="journal article" year="2001">Kazantsev 2001</bibRefCitation>
: fig. 2)...........
<taxonomicName id="4C7A4D4BFF92FF98CF09F8E704CCF899" authority="Kazantsev, 2001" authorityName="Kazantsev" authorityYear="2001" box="[1133,1436,1898,1920]" class="Insecta" family="Lycidae" genus="Macrolycus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="364" phylum="Arthropoda" rank="species" species="yunnanus">
<emphasis id="B90EEADAFF92FF98CF09F8E704CCF899" bold="true" box="[1133,1436,1898,1920]" pageId="6" pageNumber="364">
<emphasis id="B90EEADAFF92FF98CF09F8E705A1F866" bold="true" box="[1133,1265,1899,1920]" italics="true" pageId="6" pageNumber="364">M. yunnanus</emphasis>
<bibRefCitation id="EFEB4B39FF92FF98CF93F8E604CCF899" author="Kazantsev, S. V." box="[1271,1436,1898,1919]" pageId="6" pageNumber="364" pagination="99 - 109" refId="ref5410" refString="Kazantsev, S. V. (2001) Macrolycus Waterhouse, 1878 (Coleoptera: Lycidae) of continental China. Elytron, 14 (2000), 99 - 109." type="journal article" year="2001">Kazantsev, 2001</bibRefCitation>
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BC536C8FF92FF98CBF3F80404CCF85C" blockId="6.[151,1437,1400,2037]" pageId="6" pageNumber="364">
- Lamella of antennomere III at least 1.1 times as long as joint itself (
<figureCitation id="13412A4DFF92FF98C8E7F80402A0F87B" box="[899,1008,1928,1950]" captionStart="FIGURE 2" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[215,1373,190,1908]" captionTargetId="figure-25@3.[204,1383,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Male habitus of Macrolycus species, dorsal views: A. M. oreophilus Kazantsev, 2002; B. M. gansuensis Kazantsev, 2002; C. M. bowringi Waterhouse, 1878; D. M. subapicis sp. nov. (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821196" httpUri="https://zenodo.org/record/10821196/files/figure.png" pageId="6" pageNumber="364">Fig. 2A, D</figureCitation>
); subapical part of phallus strongly inflated laterally in ventral view, nearly 3.0 times as wide as basal part (
<figureCitation id="13412A4DFF92FF98C827F82902FFF85C" box="[835,943,1957,1979]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[216,1373,185,1909]" captionTargetId="figure-25@5.[205,1383,181,1919]" captionTargetPageId="5" captionText="FIGURE 3.Aedeagi of Macrolycus species, dorsal views:AC. M. oreophilus Kazantsev, 2002; DF. M. gansuensis Kazantsev, 2002; GI. M. bowringi Waterhouse, 1878; JL. M. subapicis sp. nov. (holotype). A, D, G, J: dorsal views; B, E, H, K: ventral views; C, F, I, L: lateral views. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821200" httpUri="https://zenodo.org/record/10821200/files/figure.png" pageId="6" pageNumber="364">Figs 3B, K</figureCitation>
;
<bibRefCitation id="EFEB4B39FF92FF98C8DDF8290507F85C" author="Kazantsev, S. V." box="[953,1111,1957,1978]" pageId="6" pageNumber="364" pagination="99 - 109" refId="ref5410" refString="Kazantsev, S. V. (2001) Macrolycus Waterhouse, 1878 (Coleoptera: Lycidae) of continental China. Elytron, 14 (2000), 99 - 109." type="journal article" year="2001">Kazantsev 2001</bibRefCitation>
: fig. 5; 2002: fig. 6).......... 7
</paragraph>
<paragraph id="8BC536C8FF92FF98CBF3F84F04CCF812" blockId="6.[151,1437,1400,2037]" pageId="6" pageNumber="364">
7. Pronotum and elytra dark red; subapical part of phallus flat ventrally in lateral view (
<bibRefCitation id="EFEB4B39FF92FF98CF74F84E05E0F831" author="Kazantsev, S. V." box="[1040,1200,1986,2007]" pageId="6" pageNumber="364" pagination="99 - 109" refId="ref5410" refString="Kazantsev, S. V. (2001) Macrolycus Waterhouse, 1878 (Coleoptera: Lycidae) of continental China. Elytron, 14 (2000), 99 - 109." type="journal article" year="2001">Kazantsev 2001</bibRefCitation>
: fig. 6)......................................................................................
<taxonomicName id="4C7A4D4BFF92FF98CF34F86C04CCF812" authority="Kazantsev, 2001" authorityName="Kazantsev" authorityYear="2001" box="[1104,1436,2015,2037]" class="Insecta" family="Lycidae" genus="Macrolycus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="364" phylum="Arthropoda" rank="species" species="shaanxiensis">
<emphasis id="B90EEADAFF92FF98CF34F86C04CCF812" bold="true" box="[1104,1436,2015,2037]" pageId="6" pageNumber="364">
<emphasis id="B90EEADAFF92FF98CF34F86C05A1F813" bold="true" box="[1104,1265,2015,2037]" italics="true" pageId="6" pageNumber="364">M. shaanxiensis</emphasis>
<bibRefCitation id="EFEB4B39FF92FF98CF93F86C04CCF812" author="Kazantsev, S. V." box="[1271,1436,2015,2037]" pageId="6" pageNumber="364" pagination="99 - 109" refId="ref5410" refString="Kazantsev, S. V. (2001) Macrolycus Waterhouse, 1878 (Coleoptera: Lycidae) of continental China. Elytron, 14 (2000), 99 - 109." type="journal article" year="2001">Kazantsev, 2001</bibRefCitation>
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BC536C8FF93FF99CBF3FF1404CCFF2C" blockId="7.[151,1436,151,320]" pageId="7" pageNumber="365">
- Pronotum and elytra orange red or light red (e.g.,
<figureCitation id="13412A4DFF93FF99C9DEFF1B0255FF4B" box="[698,773,151,173]" captionStart="FIGURE 2" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[215,1373,190,1908]" captionTargetId="figure-25@3.[204,1383,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Male habitus of Macrolycus species, dorsal views: A. M. oreophilus Kazantsev, 2002; B. M. gansuensis Kazantsev, 2002; C. M. bowringi Waterhouse, 1878; D. M. subapicis sp. nov. (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821196" httpUri="https://zenodo.org/record/10821196/files/figure.png" pageId="7" pageNumber="365">Fig. 2D</figureCitation>
); subapical part of phallus inflated ventrally in lateral view (e.g.,
<figureCitation id="13412A4DFF93FF99CE17FF1B004EFF2C" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[216,1373,185,1909]" captionTargetId="figure-25@5.[205,1383,181,1919]" captionTargetPageId="5" captionText="FIGURE 3.Aedeagi of Macrolycus species, dorsal views:AC. M. oreophilus Kazantsev, 2002; DF. M. gansuensis Kazantsev, 2002; GI. M. bowringi Waterhouse, 1878; JL. M. subapicis sp. nov. (holotype). A, D, G, J: dorsal views; B, E, H, K: ventral views; C, F, I, L: lateral views. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821200" httpUri="https://zenodo.org/record/10821200/files/figure.png" pageId="7" pageNumber="365">Figs 3B, K</figureCitation>
;
<bibRefCitation id="EFEB4B39FF93FF99CA4CFF390096FF2C" author="Kazantsev, S. V." box="[296,454,181,202]" pageId="7" pageNumber="365" pagination="253 - 263" refId="ref5443" refString="Kazantsev, S. V. (2002) New and little known species of Lycidae (Coleoptera) from China. Russian Entomological Journal, 11, 253 - 263." type="journal article" year="2002">Kazantsev 2002</bibRefCitation>
: fig. 7)........................................................................... 8
</paragraph>
<paragraph id="8BC536C8FF93FF99CBF3FF5E04CCFEE3" blockId="7.[151,1436,151,320]" pageId="7" pageNumber="365">
8. Elytra strongly widened posteriorly, 3.1 times longer than humeral width (
<figureCitation id="13412A4DFF93FF99C8C8FF5E02AAFF01" box="[940,1018,210,232]" captionStart="FIGURE 2" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[215,1373,190,1908]" captionTargetId="figure-25@3.[204,1383,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Male habitus of Macrolycus species, dorsal views: A. M. oreophilus Kazantsev, 2002; B. M. gansuensis Kazantsev, 2002; C. M. bowringi Waterhouse, 1878; D. M. subapicis sp. nov. (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821196" httpUri="https://zenodo.org/record/10821196/files/figure.png" pageId="7" pageNumber="365">Fig. 2D</figureCitation>
); apical part of phallus abruptly constricted distad then parallel-sided in ventral view (
<figureCitation id="13412A4DFF93FF99C912FF630391FEE3" box="[630,705,239,261]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[216,1373,185,1909]" captionTargetId="figure-25@5.[205,1383,181,1919]" captionTargetPageId="5" captionText="FIGURE 3.Aedeagi of Macrolycus species, dorsal views:AC. M. oreophilus Kazantsev, 2002; DF. M. gansuensis Kazantsev, 2002; GI. M. bowringi Waterhouse, 1878; JL. M. subapicis sp. nov. (holotype). A, D, G, J: dorsal views; B, E, H, K: ventral views; C, F, I, L: lateral views. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821200" httpUri="https://zenodo.org/record/10821200/files/figure.png" pageId="7" pageNumber="365">Fig. 3K</figureCitation>
)...........................................
<emphasis id="B90EEADAFF93FF99CFA8FF7C04CCFEE3" bold="true" box="[1228,1436,239,261]" pageId="7" pageNumber="365">
<taxonomicName id="4C7A4D4BFF93FF99CFA8FF7C0419FEE3" authority="Du &amp; Yang &amp; Yang &amp; Liu, 2024" authorityName="Du &amp; Yang &amp; Yang &amp; Liu" authorityYear="2024" box="[1228,1353,239,261]" class="Insecta" family="Lycidae" genus="Macrolycus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="365" phylum="Arthropoda" rank="species" species="subapicis" status="sp. nov.">
<emphasis id="B90EEADAFF93FF99CFA8FF7C0419FEE3" bold="true" box="[1228,1353,239,261]" italics="true" pageId="7" pageNumber="365">M. subapicis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23D57A1FF93FF99CE34FF7C04CCFEE3" box="[1360,1436,240,261]" pageId="7" pageNumber="365" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BC536C8FF93FF99CBF3FE8104CCFED9" blockId="7.[151,1436,151,320]" pageId="7" pageNumber="365">
- Elytra feebly widened posteriorly, 4.3 times longer than humeral width (
<figureCitation id="13412A4DFF93FF99C8ECFE810284FEC4" box="[904,980,269,291]" captionStart="FIGURE 2" captionStartId="3.[152,255,1943,1967]" captionTargetBox="[215,1373,190,1908]" captionTargetId="figure-25@3.[204,1383,181,1919]" captionTargetPageId="3" captionText="FIGURE 2. Male habitus of Macrolycus species, dorsal views: A. M. oreophilus Kazantsev, 2002; B. M. gansuensis Kazantsev, 2002; C. M. bowringi Waterhouse, 1878; D. M. subapicis sp. nov. (holotype). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821196" httpUri="https://zenodo.org/record/10821196/files/figure.png" pageId="7" pageNumber="365">Fig. 2A</figureCitation>
); apical part of phallus progressively constricted distad in ventral view (
<figureCitation id="13412A4DFF93FF99CADAFEA6035AFED9" box="[446,522,298,320]" captionStart="FIGURE 3" captionStartId="5.[152,255,1943,1967]" captionTargetBox="[216,1373,185,1909]" captionTargetId="figure-25@5.[205,1383,181,1919]" captionTargetPageId="5" captionText="FIGURE 3.Aedeagi of Macrolycus species, dorsal views:AC. M. oreophilus Kazantsev, 2002; DF. M. gansuensis Kazantsev, 2002; GI. M. bowringi Waterhouse, 1878; JL. M. subapicis sp. nov. (holotype). A, D, G, J: dorsal views; B, E, H, K: ventral views; C, F, I, L: lateral views. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.10821200" httpUri="https://zenodo.org/record/10821200/files/figure.png" pageId="7" pageNumber="365">Fig. 3C</figureCitation>
;
<bibRefCitation id="EFEB4B39FF93FF99C970FEA703E2FED9" author="Kazantsev, S. V." box="[532,690,298,320]" pageId="7" pageNumber="365" pagination="253 - 263" refId="ref5443" refString="Kazantsev, S. V. (2002) New and little known species of Lycidae (Coleoptera) from China. Russian Entomological Journal, 11, 253 - 263." type="journal article" year="2002">Kazantsev 2002</bibRefCitation>
: fig. 6)..............................
<taxonomicName id="4C7A4D4BFF93FF99CF02FEA604CCFED9" authority="Kazantsev, 2002" authorityName="Kazantsev" authorityYear="2002" box="[1126,1436,298,320]" class="Insecta" family="Lycidae" genus="Macrolycus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="365" phylum="Arthropoda" rank="species" species="oreophilus">
<emphasis id="B90EEADAFF93FF99CF02FEA604CCFED9" bold="true" box="[1126,1436,298,320]" pageId="7" pageNumber="365">
<emphasis id="B90EEADAFF93FF99CF02FEA605A1FEA6" bold="true" box="[1126,1265,298,320]" italics="true" pageId="7" pageNumber="365">M. oreophilus</emphasis>
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</emphasis>
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</paragraph>
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</treatment>
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