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<document ID-DOI="10.3989/ajbm.2209" ID-GBIF-Dataset="c4363035-d9cd-4166-a623-4e29d85a4be9" ID-Zenodo-Dep="6326308" approvalRequired="1" approvalRequired_for_matCits="1" checkinTime="1646336223289" checkinUser="carolina" docAuthor="Madrid, Anales del Jardín Botánico de, 2009 &amp; Knapp, Sandra" docDate="2009" docId="03D387ADB3038A15FF2DFC5DFB36F9B3" docLanguage="en" docName="AnJardinBotMadrid.66.1.65-84.pdf" docOrigin="Anales del Jardín Botánico de Madrid 66 (1)" docTitle="Solanum aquartia var. luxurians Alain, Phytologia" docType="treatment" docVersion="7" lastPageNumber="81" masterDocId="FFEAFFD5B3138A05FFBAFF9BFF99FF80" masterDocTitle="Synopsis and lectotypification of Solanum (Solanaceae) species endemic in the West Indies" masterLastPageNumber="84" masterPageNumber="65" pageNumber="81" updateTime="1682088994004" updateUser="ExternalLinkService">
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<mods:titleInfo>
<mods:title>Synopsis and lectotypification of Solanum (Solanaceae) species endemic in the West Indies</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Madrid, Anales del Jardín Botánico de</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>2009</mods:namePart>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Knapp, Sandra</mods:namePart>
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<mods:title>Anales del Jardín Botánico de Madrid</mods:title>
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<mods:date>2009</mods:date>
<mods:detail type="pubDate">
<mods:number>2009-07-07</mods:number>
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<mods:detail type="volume">
<mods:number>66</mods:number>
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<mods:start>65</mods:start>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.6329391" ID-GBIF-Taxon="193860282" ID-Zenodo-Dep="6329391" LSID="urn:lsid:plazi:treatment:03D387ADB3038A15FF2DFC5DFB36F9B3" httpUri="http://treatment.plazi.org/id/03D387ADB3038A15FF2DFC5DFB36F9B3" lastPageNumber="81" pageId="16" pageNumber="81">
<subSubSection pageId="16" pageNumber="81" type="nomenclature">
<paragraph blockId="16.[151,753,207,1100]" pageId="16" pageNumber="81">
<taxonomicName ID-CoL="4Y35Z" authority="(O.E. Schulz)" authorityName="Alain, Phytologia" baseAuthorityName="O. E. Schulz" box="[151,752,966,992]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="variety" species="aquartia" variety="luxurians">
<emphasis box="[151,347,966,992]" italics="true" pageId="16" pageNumber="81">Solanum aquartia</emphasis>
Dunal var.
<emphasis box="[483,587,966,992]" italics="true" pageId="16" pageNumber="81">luxurians</emphasis>
(O.E. Schulz)
</taxonomicName>
<bibCitation box="[180,533,999,1025]" pageId="16" pageNumber="81">Alain, Phytologia 70: 150. 1991</bibCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="81" type="multiple">
<paragraph blockId="16.[151,753,207,1100]" pageId="16" pageNumber="81">
Type:
<taxonomicName authority="O. E. Schulz ex O. Shmidt." authorityName="O. Shmidt." class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="variety" species="aculeatum" variety="luxurians">
<emphasis box="[253,474,1040,1066]" italics="true" pageId="16" pageNumber="81">Solanum aculeatum</emphasis>
(Jacq.) O.E.Schulz var.
<emphasis box="[151,255,1074,1100]" italics="true" pageId="16" pageNumber="81">luxurians</emphasis>
O.E. Schulz ex O. Shmidt.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="81" type="distribution">
<paragraph blockId="16.[151,753,1123,1981]" pageId="16" pageNumber="81">
<emphasis box="[180,323,1123,1149]" italics="true" pageId="16" pageNumber="81">Distribution.</emphasis>
In forests and scrub on limestone dry hills on Cuba, Hispaniola and Jamaica, from sea level to 400 m elevation.
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="81" type="materials_examined">
<paragraph blockId="16.[151,753,1123,1981]" pageId="16" pageNumber="81">
<emphasis box="[180,461,1224,1249]" italics="true" pageId="16" pageNumber="81">Representative specimen.</emphasis>
Jamaica, Saint Catherine Parish,
<emphasis box="[236,392,1256,1282]" italics="true" pageId="16" pageNumber="81">
<taxonomicName authorityName="Gistel" authorityYear="1848" box="[236,317,1257,1282]" class="Malacostraca" family="Latreilliidae" genus="Proctor" kingdom="Animalia" order="Decapoda" pageId="16" pageNumber="81" phylum="Arthropoda" rank="genus">Proctor</taxonomicName>
32768
</emphasis>
(BM).
</paragraph>
</subSubSection>
<subSubSection pageId="16" pageNumber="81" type="discussion">
<paragraph blockId="16.[151,753,1123,1981]" pageId="16" pageNumber="81">
There has long been confusion over the correct name for this species.
<bibRefCitation author="Dunal, M. - F." box="[418,590,1323,1349]" journalOrPublisher="Montpellier" pageId="16" pageNumber="81" refId="ref13588" refString="Dunal, M. - F. 1813. Histoire naturelle, medicale et economique des Solanum et des genres qui ont ete confundus avec eux. Montpellier." title="Histoire naturelle, medicale et economique des Solanum et des genres qui ont ete confundus avec eux" type="book" year="1813">Dunals (1813)</bibRefCitation>
re-naming of Jacquins
<taxonomicName box="[258,451,1356,1382]" class="Magnoliopsida" family="Solanaceae" genus="Aquartia" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="species" species="aculeata">
<emphasis box="[258,451,1356,1382]" italics="true" pageId="16" pageNumber="81">Aquartia aculeata</emphasis>
</taxonomicName>
as
<taxonomicName box="[484,677,1356,1382]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="species" species="aquartia">
<emphasis box="[484,677,1356,1382]" italics="true" pageId="16" pageNumber="81">Solanum aquartia</emphasis>
</taxonomicName>
was illegitimate, but the name was used for many years. By the time
<bibRefCitation author="Schulz, O. E." box="[249,405,1422,1448]" editor="I. Urban" journalOrPublisher="Symbolae Antillanae" pageId="16" pageNumber="81" pagination="140 - 279" part="6" refId="ref14448" refString="Schulz, O. E. 1909. Solanacearum genera nonnulla, in I. Urban (ed.) Symbolae Antillanae 6: 140 - 279." title="Solanacearum genera nonnulla" type="journal article" year="1909">Schulz (1909)</bibRefCitation>
made the correct combination
<taxonomicName box="[151,367,1456,1482]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="species" species="aculeatum">
<emphasis box="[151,367,1456,1482]" italics="true" pageId="16" pageNumber="81">Solanum aculeatum</emphasis>
</taxonomicName>
, the name
<taxonomicName authority="St." authorityName="St." box="[490,744,1456,1482]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="species" species="aculeatum">
<emphasis box="[490,707,1456,1482]" italics="true" pageId="16" pageNumber="81">Solanum aculeatum</emphasis>
St.
</taxonomicName>
- Lag. had been validly (but illegitimately) published as a replacement for
<taxonomicName authority="Dunal" authorityName="Dunal" box="[379,680,1522,1548]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="species" species="aculeiger">
<emphasis box="[379,596,1522,1548]" italics="true" pageId="16" pageNumber="81">Solanum aculeiger</emphasis>
Dunal
</taxonomicName>
(now recognised as a synonym of the Brazilian species
<taxonomicName authority="Sendtn." authorityName="Sendtn." class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="species" species="thomasiifolium">
<emphasis italics="true" pageId="16" pageNumber="81">S. thomasiifolium</emphasis>
Sendtn.
</taxonomicName>
, see
<taxonomicName authority="Source" authorityName="Source" box="[511,748,1589,1615]" class="Magnoliopsida" family="Solanaceae" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="family">Solanaceae Source</taxonomicName>
, http://www.solanaceaesource.org). The oldest available name for this species is
<taxonomicName box="[468,624,1656,1682]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="species" species="tetramerum">
<emphasis box="[468,624,1656,1682]" italics="true" pageId="16" pageNumber="81">S. tetramerum</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph blockId="16.[151,753,1123,1981]" lastBlockId="16.[805,1406,207,1587]" pageId="16" pageNumber="81">
No specimen of
<taxonomicName box="[377,575,1689,1715]" class="Magnoliopsida" family="Solanaceae" genus="Aquartia" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="species" species="aculeata">
<emphasis box="[377,575,1689,1715]" italics="true" pageId="16" pageNumber="81">Aquartia aculeata</emphasis>
</taxonomicName>
attributable to Jacquin has been found, despite intensive searches in many herbaria (principally BM, LINN and W) where such a sheet, if it had ever existed, might have been preserved. No original material was cited in the protologue (Jacquin, 1760), so any type selected must be a neotype. The illustration of
<taxonomicName box="[354,473,1889,1915]" class="Magnoliopsida" family="Solanaceae" genus="Aquartia" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="species" species="aculeata">
<emphasis box="[354,473,1889,1915]" italics="true" pageId="16" pageNumber="81">A. aculeata</emphasis>
</taxonomicName>
in Jacquins first illustrated edition of the
<emphasis italics="true" pageId="16" pageNumber="81">Selectarum Stirpium Americanarum Historia</emphasis>
(Jaqcuin, 1763), a copper engraving taken from original drawings done by Jacquin in the field (see
<bibRefCitation author="Wiltshear, F. G." box="[805,980,240,266]" journalOrPublisher="Journal of Botany" pageId="16" pageNumber="81" pagination="140" part="1913" refId="ref14572" refString="Wiltshear, F. G. 1913. LII. Jacquin's ' Selectarum Stirpium Historia Iconibus Pictus'. Journal of Botany 1913: 140." title="LII. Jacquin's ' Selectarum Stirpium Historia Iconibus Pictus'" type="journal article" year="1913">Wiltshear, 1913</bibRefCitation>
) is very accurate, and I have chosen it here as the neotype of the species (
<figureCitation box="[1183,1261,274,300]" captionStart="Fig" captionStartId="15.[181,215,1854,1875]" captionTargetBox="[234,1383,205,1820]" captionTargetPageId="15" captionText="Fig. 5. a, lectotype of Solanum schulzianum Urb., Eggers 2208 (K000196492); b, lectotype of S. selleanum Urb. &amp; Ekman, Ekman H.3127 (S04-2981). Reproduced with permission of the Swedish Museum of Natural History; c, lectotype of Aquartia aculeata Jacq., Jacquin,Select.Stirp.Amer. t. 12, 1763) (= S. tetramerum Dunal). Reproduced with permission of the Natural History Museum Botany Library; d, neotype of S. inclusum Griseb.,Linden 1986 (BM000886483) (= S. tetramerum Dunal). Reproduced with permission of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.6326320" httpUri="https://zenodo.org/record/6326320/files/figure.png" pageId="16" pageNumber="81">Fig. 5c</figureCitation>
).
</paragraph>
<paragraph blockId="16.[805,1406,207,1587]" pageId="16" pageNumber="81">
<taxonomicName authorityName="Griseb., Linden" authorityYear="1986" box="[833,1041,308,334]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="species" species="inclusum">
<emphasis box="[833,1041,308,334]" italics="true" pageId="16" pageNumber="81">Solanum inclusum</emphasis>
</taxonomicName>
was described without specific collection or locality but said to be widespread across the Caribbean and in northern South America (
<bibRefCitation author="Grisebach, A." journalOrPublisher="Lovell, Reeve &amp; Co., London" pageId="16" pageNumber="81" part="4" refId="ref13800" refString="Grisebach, A. 1861. Flora of the British West Indian islands, parts 4 - 5. Lovell, Reeve &amp; Co., London." title="Flora of the British West Indian islands" type="book" year="1861">Grisebach, 1861</bibRefCitation>
). Various subsequent authors have suggested that Grisebachs
<taxonomicName authorityName="Griseb., Linden" authorityYear="1986" box="[1079,1208,444,470]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="species" species="inclusum">
<emphasis box="[1079,1208,444,470]" italics="true" pageId="16" pageNumber="81">S. inclusum</emphasis>
</taxonomicName>
included both
<taxonomicName class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="species" species="tetramerum">
<emphasis italics="true" pageId="16" pageNumber="81">S. tetramerum</emphasis>
</taxonomicName>
and
<taxonomicName box="[1000,1160,477,503]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="species" species="polygamum">
<emphasis box="[1000,1160,477,503]" italics="true" pageId="16" pageNumber="81">S. polygamum</emphasis>
</taxonomicName>
. No specimens have been identified at GOET that can be directly linked to Grisebachs naming of
<taxonomicName authorityName="Griseb., Linden" authorityYear="1986" box="[1062,1188,545,571]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="species" species="inclusum">
<emphasis box="[1062,1188,545,571]" italics="true" pageId="16" pageNumber="81">S. inclusum</emphasis>
</taxonomicName>
. In his Cuban catalogue
<bibRefCitation author="Grisebach, A." box="[881,1093,579,605]" journalOrPublisher="W. Engelmann, Leipzig" pageId="16" pageNumber="81" refId="ref13828" refString="Grisebach, A. 1866. Catalogus Plantarum Cubensium. W. Engelmann, Leipzig." title="Catalogus Plantarum Cubensium" type="book" year="1866">Grisebach (1866)</bibRefCitation>
cited specimens as
<taxonomicName authorityName="Griseb., Linden" authorityYear="1986" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="species" species="inclusum">
<emphasis italics="true" pageId="16" pageNumber="81">S. inclusum</emphasis>
</taxonomicName>
that are now identified as
<taxonomicName box="[1185,1342,613,639]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="species" species="tetramerum">
<emphasis box="[1185,1342,613,639]" italics="true" pageId="16" pageNumber="81">S. tetramerum</emphasis>
</taxonomicName>
(
<emphasis italics="true" pageId="16" pageNumber="81">Linden 1986</emphasis>
,
<emphasis box="[921,976,647,673]" italics="true" pageId="16" pageNumber="81">2000</emphasis>
). Specimens indicated as type material of
<taxonomicName authorityName="Griseb., Linden" authorityYear="1986" box="[859,985,680,706]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="species" species="inclusum">
<emphasis box="[859,985,680,706]" italics="true" pageId="16" pageNumber="81">S. inclusum</emphasis>
</taxonomicName>
in some herbaria (i.e.,
<emphasis box="[1238,1378,680,706]" italics="true" pageId="16" pageNumber="81">Wright 1350</emphasis>
in GH) are in fact
<taxonomicName box="[981,1087,714,740]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="species" species="torvum">
<emphasis box="[981,1087,714,740]" italics="true" pageId="16" pageNumber="81">S. torvum</emphasis>
</taxonomicName>
. Since
<bibRefCitation author="Grisebach, A." box="[1165,1362,714,740]" journalOrPublisher="Lovell, Reeve &amp; Co., London" pageId="16" pageNumber="81" part="4" refId="ref13800" refString="Grisebach, A. 1861. Flora of the British West Indian islands, parts 4 - 5. Lovell, Reeve &amp; Co., London." title="Flora of the British West Indian islands" type="book" year="1861">Grisebach (1861)</bibRefCitation>
did not indicate any specimens or specific localities for his name, these sheets cannot be considered type material. I neotypify Grisebachs
<taxonomicName authorityName="Griseb., Linden" authorityYear="1986" box="[1131,1344,816,842]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="species" species="inclusum">
<emphasis box="[1131,1344,816,842]" italics="true" pageId="16" pageNumber="81">Solanum inclusum</emphasis>
</taxonomicName>
here with the BM duplicate (
<figureCitation box="[1072,1153,850,876]" captionStart="Fig" captionStartId="15.[181,215,1854,1875]" captionTargetBox="[234,1383,205,1820]" captionTargetPageId="15" captionText="Fig. 5. a, lectotype of Solanum schulzianum Urb., Eggers 2208 (K000196492); b, lectotype of S. selleanum Urb. &amp; Ekman, Ekman H.3127 (S04-2981). Reproduced with permission of the Swedish Museum of Natural History; c, lectotype of Aquartia aculeata Jacq., Jacquin,Select.Stirp.Amer. t. 12, 1763) (= S. tetramerum Dunal). Reproduced with permission of the Natural History Museum Botany Library; d, neotype of S. inclusum Griseb.,Linden 1986 (BM000886483) (= S. tetramerum Dunal). Reproduced with permission of the Natural History Museum, London." figureDoi="http://doi.org/10.5281/zenodo.6326320" httpUri="https://zenodo.org/record/6326320/files/figure.png" pageId="16" pageNumber="81">Fig. 5d</figureCitation>
) of
<emphasis box="[1197,1338,850,876]" italics="true" pageId="16" pageNumber="81">Linden 1986</emphasis>
in order to stabilise usage (synonymy) of this ambiguous name.
</paragraph>
<paragraph blockId="16.[805,1406,207,1587]" pageId="16" pageNumber="81">
No duplicates of
<emphasis box="[1058,1247,951,977]" italics="true" pageId="16" pageNumber="81">Ekman H.7289</emphasis>
, the type of
<emphasis box="[805,1023,985,1011]" italics="true" pageId="16" pageNumber="81">
<taxonomicName box="[805,902,985,1011]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="genus">Solanum</taxonomicName>
miragonae
</emphasis>
, are extant at B, and the only sheet of this collection encountered is at S, so this is the clear choice as the lectotype (
<figureCitation box="[1141,1223,1053,1079]" captionStart="Fig" captionStartId="17.[181,214,1827,1848]" captionTargetBox="[234,1383,205,1804]" captionTargetPageId="17" captionText="Fig. 6. a, lectotype of Solanum miragoanae Urb.,Ekman 7289 (S04-2952) (= S.tetramerum). Reproduced with permission of the Swedish Museum of Natural History; b, lectotype of S. tetramerum Dunal var. fuertesii O.E. Schulz, Fuertes 284 (NY00111372) (= S. tetramerum Dunal). Reproduced with permission of the New York Botanical Garden; c, lectotype of S. tetramerum var. luxurians O.E. Schulz, Ekman H.9979 (S04-2827) (= S. tetramerum Dunal). Reproduced with permission of the Swedish Museum of Natural History; d, lectotype of S. lobulatum O.E. Schulz, Smith &amp; Smith 1575 (K000196493) (= S. urens Dunal). Reproduced with permission of the Royal Botanic Gardens,Kew." figureDoi="http://doi.org/10.5281/zenodo.6326322" httpUri="https://zenodo.org/record/6326322/files/figure.png" pageId="16" pageNumber="81">Fig. 6a</figureCitation>
). It may be that the S sheet is the holotype, as
<emphasis box="[1163,1308,1087,1113]" italics="true" pageId="16" pageNumber="81">S. miragonae</emphasis>
was described in a Swedish journal along with other taxa whose types are in S, but as there is no evidence of this, I prefer to lectotypify the name here.
</paragraph>
<paragraph blockId="16.[805,1406,207,1587]" pageId="16" pageNumber="81">
<taxonomicName box="[833,1197,1222,1248]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="variety" species="aculeatum" variety="fuertesii">
<emphasis box="[833,1051,1222,1248]" italics="true" pageId="16" pageNumber="81">Solanum aculeatum</emphasis>
var.
<emphasis box="[1107,1197,1222,1248]" italics="true" pageId="16" pageNumber="81">fuertesii</emphasis>
</taxonomicName>
was described using several collections:
<emphasis box="[1070,1258,1256,1282]" italics="true" pageId="16" pageNumber="81">Fuertes 384, 386</emphasis>
and
<emphasis box="[1317,1359,1256,1282]" italics="true" pageId="16" pageNumber="81">396</emphasis>
. Of these,
<emphasis box="[891,1040,1290,1316]" italics="true" pageId="16" pageNumber="81">Fuertes 384</emphasis>
has been located in several herbaria, the NY duplicate is here selected as the lectotype (
<figureCitation box="[892,976,1358,1384]" captionStart="Fig" captionStartId="17.[181,214,1827,1848]" captionTargetBox="[234,1383,205,1804]" captionTargetPageId="17" captionText="Fig. 6. a, lectotype of Solanum miragoanae Urb.,Ekman 7289 (S04-2952) (= S.tetramerum). Reproduced with permission of the Swedish Museum of Natural History; b, lectotype of S. tetramerum Dunal var. fuertesii O.E. Schulz, Fuertes 284 (NY00111372) (= S. tetramerum Dunal). Reproduced with permission of the New York Botanical Garden; c, lectotype of S. tetramerum var. luxurians O.E. Schulz, Ekman H.9979 (S04-2827) (= S. tetramerum Dunal). Reproduced with permission of the Swedish Museum of Natural History; d, lectotype of S. lobulatum O.E. Schulz, Smith &amp; Smith 1575 (K000196493) (= S. urens Dunal). Reproduced with permission of the Royal Botanic Gardens,Kew." figureDoi="http://doi.org/10.5281/zenodo.6326322" httpUri="https://zenodo.org/record/6326322/files/figure.png" pageId="16" pageNumber="81">Fig. 6b</figureCitation>
). In describing
<taxonomicName box="[1154,1341,1358,1384]" class="Magnoliopsida" family="Solanaceae" genus="Solanum" kingdom="Plantae" order="Solanales" pageId="16" pageNumber="81" phylum="Tracheophyta" rank="variety" species="aculeatum" variety="luxurians">
variety
<emphasis box="[1237,1341,1358,1384]" italics="true" pageId="16" pageNumber="81">luxurians</emphasis>
</taxonomicName>
, with larger, more repand leaves, several syntypes were cited:
<emphasis box="[848,1021,1425,1451]" italics="true" pageId="16" pageNumber="81">Ekman H.2318</emphasis>
,
<emphasis box="[1036,1398,1425,1451]" italics="true" pageId="16" pageNumber="81">Ekman H.6732, Ekman H.8548</emphasis>
,
<emphasis box="[805,976,1459,1485]" italics="true" pageId="16" pageNumber="81">Ekman H.9979</emphasis>
and
<emphasis box="[1033,1205,1459,1485]" italics="true" pageId="16" pageNumber="81">Ekman H.8773</emphasis>
. The duplicate of
<emphasis box="[805,977,1493,1519]" italics="true" pageId="16" pageNumber="81">Ekman H.9979</emphasis>
in S (
<figureCitation box="[1045,1125,1493,1519]" captionStart="Fig" captionStartId="17.[181,214,1827,1848]" captionTargetBox="[234,1383,205,1804]" captionTargetPageId="17" captionText="Fig. 6. a, lectotype of Solanum miragoanae Urb.,Ekman 7289 (S04-2952) (= S.tetramerum). Reproduced with permission of the Swedish Museum of Natural History; b, lectotype of S. tetramerum Dunal var. fuertesii O.E. Schulz, Fuertes 284 (NY00111372) (= S. tetramerum Dunal). Reproduced with permission of the New York Botanical Garden; c, lectotype of S. tetramerum var. luxurians O.E. Schulz, Ekman H.9979 (S04-2827) (= S. tetramerum Dunal). Reproduced with permission of the Swedish Museum of Natural History; d, lectotype of S. lobulatum O.E. Schulz, Smith &amp; Smith 1575 (K000196493) (= S. urens Dunal). Reproduced with permission of the Royal Botanic Gardens,Kew." figureDoi="http://doi.org/10.5281/zenodo.6326322" httpUri="https://zenodo.org/record/6326322/files/figure.png" pageId="16" pageNumber="81">Fig. 6c</figureCitation>
) has an annotation label in Schulzs hand with the varietal name
<emphasis box="[1246,1350,1527,1553]" italics="true" pageId="16" pageNumber="81">luxurians</emphasis>
, and is the logical choice for a lectotype.
</paragraph>
</subSubSection>
</treatment>
</document>