719 lines
96 KiB
XML
719 lines
96 KiB
XML
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<mods:title id="E8B3E5DF9F0D5F0C242E7863F900D1D2">Taxonomic revision of the genus Oplognathus MacLeay, 1819 (Coleoptera: Scarabaeidae: Rutelinae: Rutelini)</mods:title>
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<mods:namePart id="89796A3A538EB8B7AC1DC53DD5745427">Carvalho, Tamara G.</mods:namePart>
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<mods:affiliation id="DB04E06D0BF707D2DD4BF12ABDA2ECC2">Programa de Pós-Graduação em Entomologia da UFRPE, Universidade Federal Rural de Pernambuco, Rua Manoel de Medeiros, s / n - Dois Irmãos, 52171 - 900, Recife, PE, Brazil.</mods:affiliation>
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<mods:namePart id="827AB32ACC4E08EFA844F88A0D030EEC">Seidel, Matthias</mods:namePart>
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<mods:affiliation id="CBDB7636A36CFED822DCFEB8ED30E9AE">Centrum für Naturkunde (CeNak), Universität Hamburg, Martin-Luther-King-Platz 3, 20146 Hamburg, Germany.</mods:affiliation>
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<mods:nameIdentifier id="125698164294323EE79EECCF4C85627C" type="email">matthias.seidel@uni-hamburg.de;matth.seidel@gmail.com</mods:nameIdentifier>
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<mods:affiliation id="00E19DA3F1ABA2D44FAC2EDDA3CE24D9">Programa de Pós-Graduação em Entomologia da UFRPE, Universidade Federal Rural de Pernambuco, Rua Manoel de Medeiros, s / n - Dois Irmãos, 52171 - 900, Recife, PE, Brazil.</mods:affiliation>
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<mods:title id="CCD5BB36B87ACEEC865FAF3F4EE061E2">European Journal of Taxonomy</mods:title>
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<mods:date id="0D09E797C757D43D99E11178D010274E">2021</mods:date>
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<taxonomicName id="4C7A4D11E15C453BFDA1FC65FBB9FBE9" authority="Ohaus, 1905" authorityName="Ohaus" authorityYear="1905" box="[554,1034,1017,1045]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="71" phylum="Arthropoda" rank="species" species="helmenreichi">
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<emphasis id="B90EEA80E15C453BFDA1FC65FCDDFBE8" bold="true" box="[554,878,1017,1044]" italics="true" pageId="9" pageNumber="71">Oplognathus helmenreichi</emphasis>
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<bibRefCitation id="EFEB4B63E15C453BFCFEFC66FBB9FBE9" author="Ohaus F." box="[885,1034,1018,1045]" pageId="9" pageNumber="71" pagination="283 - 329" refId="ref11320" refString="Ohaus F. 1905. Beitrage zur Kenntniss der amerikanischen Ruteliden. Stettiner Entomologische Zeitung 66: 283 - 329." type="journal article" year="1905">Ohaus, 1905</bibRefCitation>
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<paragraph id="8BC53692E15C453BFD9AFB81FB90FBCB" blockId="9.[529,1059,1017,1079]" box="[529,1059,1053,1079]" pageId="9" pageNumber="71">
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<figureCitation id="13412A17E15C453BFD9AFB81FD37FBCB" box="[529,644,1053,1079]" captionStart="Fig" captionStartId="7.[189,232,1885,1911]" captionTargetBox="[335,1250,321,1813]" captionTargetId="figure-17@7.[293,1296,265,1846]" captionTargetPageId="7" captionText="Fig. 1. a–b. Oplognathus kirbii MacLeay, 1819. a. Neotype, ♂ (NHMUK). b. Female specimen (MZUSP). c. O. helmenreichi Ohaus, 1905, holotype, ♂ (ZMHB). d–e. O. helmenreichi var.maculicollis Ohaus, 1914. d. Syntype, ♂ (ZMHB). e. Syntype, ♀ (ZMHB). f–g. O. bahianus Ohaus, 1912. f. Syntype, ♂ (ZMHB). g. Female specimen (EPGC). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514422" httpUri="https://zenodo.org/record/5514422/files/figure.png" pageId="9" pageNumber="71">Figs 1c–e</figureCitation>
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,
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<figureCitation id="13412A17E15C453BFD1BFB81FD7DFBCB" box="[656,718,1053,1079]" captionStart="Fig" captionStartId="8.[189,232,1920,1946]" captionTargetBox="[233,1372,1482,1865]" captionTargetId="figure-407@8.[197,1400,1458,1881]" captionTargetPageId="8" captionText="Fig. 2. Labels. a. Oplognathus bahianus Ohaus, 1912, syntype, ♂. b. O. helmenreichi Ohaus, 1905, holotype, ♂. c–d. O. helmenreichi var. maculicollis Ohaus, 1914.c. Syntype ♂. d. Syntype ♀. e. O. kirbii MacLeay, 1819, neotype, ♂." figureDoi="http://doi.org/10.5281/zenodo.5514424" httpUri="https://zenodo.org/record/5514424/files/figure.png" pageId="9" pageNumber="71">2b–d</figureCitation>
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,
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<figureCitation id="13412A17E15C453BFD51FB81FCA6FBCB" box="[730,789,1053,1079]" captionStart="Fig" captionStartId="11.[189,232,1918,1944]" captionTargetBox="[232,1370,378,1807]" captionTargetId="figure-16@11.[189,1398,315,1828]" captionTargetPageId="11" captionText="Fig. 3. Head and pronotum details. a. Oplognathus kirbii MacLeay, 1819, male specimen (ZMHB). b. O. bahianus Ohaus, 1912, syntype, ♂ (ZMHB). c. O. helmenreichiOhaus, 1905, holotype, ♂ (ZMHB). d. O. helmenreichi var. maculicollis Ohaus, 1914, syntype, ♂ (ZMHB). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514426" httpUri="https://zenodo.org/record/5514426/files/figure.png" pageId="9" pageNumber="71">3c–d</figureCitation>
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,
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<figureCitation id="13412A17E15C453BFCAAFB81FCD3FBCB" box="[801,864,1053,1079]" captionStart="Fig" captionStartId="12.[189,232,1918,1944]" captionTargetBox="[206,1384,870,1864]" captionTargetId="figure-172@12.[189,1398,850,1878]" captionTargetPageId="12" captionText="Fig. 4. Mouthparts. a–d. Oplognathus kirbii MacLeay, 1819 (ZMHB). e–h. O. bahianus Ohaus, 1912 (ZMHB). i–m. O. helmenreichi Ohaus, 1905 (ZMHB). a, e, i. Right mandible. b, f, j. Right maxilla. c, g, l. Detail of the last maxillary palpomere. d, h, m. Mentum. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514428" httpUri="https://zenodo.org/record/5514428/files/figure.png" pageId="9" pageNumber="71">4i–m</figureCitation>
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,
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<figureCitation id="13412A17E15C453BFCE7FB81FCCEFBCB" box="[876,893,1053,1079]" captionStart="Fig" captionStartId="15.[189,232,1921,1947]" captionTargetBox="[277,1269,617,1876]" captionTargetId="figure-111@15.[189,1398,583,1882]" captionTargetPageId="15" captionText="Fig. 6. Aedeagus in dorsal (a, e) and lateral view (b–c, d–f). a–c. Oplognathus helmenreichi Ohaus, 1905, holotype, ♂ (ZMHB). d–f. O. helmenreichi var. maculicollis Ohaus, 1914, syntype, ♂ (ZMHB). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514434" httpUri="https://zenodo.org/record/5514434/files/figure.png" pageId="9" pageNumber="71">6</figureCitation>
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,
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<figureCitation id="13412A17E15C453BFC01FB81FC16FBCB" box="[906,933,1053,1079]" captionStart="Fig" captionStartId="17.[189,232,1924,1950]" captionTargetBox="[270,1309,269,1863]" captionTargetId="figure-16@17.[243,1344,265,1886]" captionTargetPageId="17" captionText="Fig. 7. Variation in male genitalia. a–d. Oplognathus kirbii MacLeay, 1819 (a-b: CEMT; c-d: ZMHB). e–h. O. helmenreichi Ohaus, 1905 (ZMHB). i–m. O. bahianus Ohaus, 1912 (CERPE). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514436" httpUri="https://zenodo.org/record/5514436/files/figure.png" pageId="9" pageNumber="71">7e</figureCitation>
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–h,
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<figureCitation id="13412A17E15C453BFC5AFB81FBBBFBCB" box="[977,1032,1053,1079]" captionStart="Fig" captionStartId="18.[189,232,1925,1951]" captionTargetBox="[214,1361,1235,1854]" captionTargetId="figure-286@18.[189,1398,1189,1887]" captionTargetPageId="18" captionText="Fig. 8. Patterns of mesoventral process. a, d. Oplognathus kirbii MacLeay, 1819 (MZUSP). b, e. O. bahianus Ohaus, 1912 (ZMHB). c, f. O. helmenreichi Ohaus, 1905 (CERPE). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514440" httpUri="https://zenodo.org/record/5514440/files/figure.png" pageId="9" pageNumber="71">8c, f</figureCitation>
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, 9
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</paragraph>
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<treatmentCitation id="0ADB1083E15C453BFF36FBC2FD6DFB85" author="Ohaus F." box="[189,734,1118,1145]" page="322" pageId="9" pageNumber="71" year="1905">
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<taxonomicName id="4C7A4D11E15C453BFF36FBC2FD6DFB85" authority="Ohaus, 1905: 322" authorityName="Ohaus" authorityPageNumber="322" authorityYear="1905" box="[189,734,1118,1145]" class="Insecta" family="Rutelidae" genus="Hoplognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="71" phylum="Arthropoda" rank="species" species="helmenreichi">
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<emphasis id="B90EEA80E15C453BFF36FBC2FDB0FB84" box="[189,515,1118,1144]" italics="true" pageId="9" pageNumber="71">Hoplognathus helmenreichi</emphasis>
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<bibRefCitation id="EFEB4B63E15C453BFD81FBC2FD6DFB85" author="Ohaus F." box="[522,734,1118,1145]" pageId="9" pageNumber="71" pagination="283 - 329" refId="ref11320" refString="Ohaus F. 1905. Beitrage zur Kenntniss der amerikanischen Ruteliden. Stettiner Entomologische Zeitung 66: 283 - 329." type="journal article" year="1905">Ohaus, 1905: 322</bibRefCitation>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BC53692E15C453BFF36FB3CFD3CFB22" blockId="9.[189,1399,1184,1457]" pageId="9" pageNumber="71">
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<taxonomicName id="4C7A4D11E15C453BFF36FB3CFDB8FB46" authorityName="Ohaus" authorityYear="1905" box="[189,523,1184,1210]" class="Insecta" family="Rutelidae" genus="Hoplognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="71" phylum="Arthropoda" rank="species" species="helmenreichi">
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<emphasis id="B90EEA80E15C453BFF36FB3CFDB8FB46" box="[189,523,1184,1210]" italics="true" pageId="9" pageNumber="71">Hoplognathus helmenreichi</emphasis>
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</taxonomicName>
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–
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||
<treatmentCitation id="0ADB1083E15C453BFDB3FB3CFCADFB46" author="Ohaus F." box="[568,798,1184,1210]" page="301" pageId="9" pageNumber="71" year="1914">
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<bibRefCitation id="EFEB4B63E15C453BFDB3FB3CFCADFB46" author="Ohaus F." box="[568,798,1184,1210]" pageId="9" pageNumber="71" pagination="299 - 304" refId="ref11366" refString="Ohaus F. 1914 a. Neue Coleoptera lamellicornia aus Argentinien. Deutsche Entomologische Zeitschrift 1914: 299 - 304." type="journal article" year="1914">Ohaus 1914a: 301</bibRefCitation>
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</treatmentCitation>
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(species description); 1918: 12 (catalogue). —
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<treatmentCitation id="0ADB1083E15C453BFF61FB5FFE4FFB22" author="Blackwelder" box="[234,508,1219,1246]" page="235" pageId="9" pageNumber="71" year="1944">
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<bibRefCitation id="EFEB4B63E15C453BFF61FB5FFE4FFB22" author="Blackwelder R. E." box="[234,508,1219,1246]" pageId="9" pageNumber="71" pagination="189 - 341" refId="ref10657" refString="Blackwelder R. E. 1944. Checklist of the Coleopterous insects of Mexico, Central America, the West Indies and South America. Part 2. Bulletin of the United States National Museum 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 235</bibRefCitation>
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</treatmentCitation>
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(catalogue).
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BC53692E15C453BFF36FB7AFCFBFAD8" blockId="9.[189,1399,1184,1457]" pageId="9" pageNumber="71">
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<treatmentCitationGroup id="AB6A11BCE15C453BFF36FB7AFCFBFAD8" pageId="9" pageNumber="71">
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<taxonomicName id="4C7A4D11E15C453BFF36FB7AFDB7FAFC" baseAuthorityName="Ohaus" baseAuthorityYear="1905" box="[189,516,1254,1280]" class="Insecta" family="Rutelidae" genus="Hoplognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="71" phylum="Arthropoda" rank="species" species="helmreicheni">
|
||
<emphasis id="B90EEA80E15C453BFF36FB7AFDB7FAFC" box="[189,516,1254,1280]" italics="true" pageId="9" pageNumber="71">Hoplognathus helmreicheni</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0ADB1083E15C453BFDAFFB7AFC48FAFD" author="Ohaus" box="[548,1019,1254,1281]" page="302" pageId="9" pageNumber="71" year="1914">
|
||
<bibRefCitation id="EFEB4B63E15C453BFDAFFB7AFD48FAFC" author="Ohaus F." box="[548,763,1254,1280]" pageId="9" pageNumber="71" pagination="299 - 304" refId="ref11366" refString="Ohaus F. 1914 a. Neue Coleoptera lamellicornia aus Argentinien. Deutsche Entomologische Zeitschrift 1914: 299 - 304." type="journal article" year="1914">Ohaus 1914a: 302</bibRefCitation>
|
||
(species description)
|
||
</treatmentCitation>
|
||
; 1918: 12 (catalogue); 1934: 42 (catalogue). —
|
||
<treatmentCitation id="0ADB1083E15C453BFE28FA95FD06FAD8" author="Blackwelder" box="[419,693,1289,1316]" page="235" pageId="9" pageNumber="71" year="1944">
|
||
<bibRefCitation id="EFEB4B63E15C453BFE28FA95FD06FAD8" author="Blackwelder R. E." box="[419,693,1289,1316]" pageId="9" pageNumber="71" pagination="189 - 341" refId="ref10657" refString="Blackwelder R. E. 1944. Checklist of the Coleopterous insects of Mexico, Central America, the West Indies and South America. Part 2. Bulletin of the United States National Museum 185: 189 - 341. https: // doi. org / 10.5479 / si. 03629236.185.2" type="journal article" year="1944">Blackwelder 1944: 235</bibRefCitation>
|
||
</treatmentCitation>
|
||
(catalogue).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BC53692E15C453BFF36FAB1FC2CFA97" blockId="9.[189,1399,1184,1457]" pageId="9" pageNumber="71">
|
||
<treatmentCitationGroup id="AB6A11BCE15C453BFF36FAB1FC2CFA97" pageId="9" pageNumber="71">
|
||
<taxonomicName id="4C7A4D11E15C453BFF36FAB1FE47FABB" baseAuthorityName="Ohaus" baseAuthorityYear="1905" box="[189,500,1325,1351]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="71" phylum="Arthropoda" rank="species" species="helmreicheni">
|
||
<emphasis id="B90EEA80E15C453BFF36FAB1FE47FABB" box="[189,500,1325,1351]" italics="true" pageId="9" pageNumber="71">Oplognathus helmreicheni</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0ADB1083E15C453BFD9AFAB1FC2CFAB4" author="Machatschke" box="[529,927,1324,1352]" page="5" pageId="9" pageNumber="71" year="1972">
|
||
<bibRefCitation id="EFEB4B63E15C453BFD9AFAB1FCBFFABB" author="Machatschke J. W." box="[529,780,1324,1351]" pageId="9" pageNumber="71" pagination="1 - 361" refId="ref11259" refString="Machatschke J. W. 1972. Scarabaeoidea: Melolonthidae, Rutelinae. Coleopterorum Catalogus Supplementa 66: 1 - 361." type="journal article" year="1972">Machatschke 1972: 5</bibRefCitation>
|
||
(catalogue).
|
||
</treatmentCitation>
|
||
—
|
||
<treatmentCitation id="0ADB1083E15C453BFC40FAB1FA96FAB4" author="Krajcik" box="[971,1317,1325,1352]" page="90" pageId="9" pageNumber="71" year="2007">
|
||
<bibRefCitation id="EFEB4B63E15C453BFC40FAB1FB22FABB" author="Krajcik M." box="[971,1169,1325,1351]" pageId="9" pageNumber="71" refId="ref11075" refString="Krajcik M. 2007. Anima. X. Rutelinae of the world. Checklist of Scarabaeoidea of the world. 2. Rutelinae (Coleoptera: Scarabaeidae: Rutelinae). Plzen, Czech Republic." type="book" year="2007">Krajcik 2007: 90</bibRefCitation>
|
||
(catalogue).
|
||
</treatmentCitation>
|
||
<taxonomicName id="4C7A4D11E15C453BFF36FACCFE47FA96" authorityName="Ohaus" authorityYear="1905" box="[189,500,1360,1386]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="71" phylum="Arthropoda" rank="species" species="helmenreichi">
|
||
<emphasis id="B90EEA80E15C453BFF36FACCFE47FA96" box="[189,500,1360,1386]" italics="true" pageId="9" pageNumber="71">Oplognathus helmenreichi</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0ADB1083E15C453BFD9AFACCFC2CFA97" author="Machatschke" box="[529,927,1360,1387]" page="5" pageId="9" pageNumber="71" year="1972">
|
||
<bibRefCitation id="EFEB4B63E15C453BFD9AFACCFCBFFA97" author="Machatschke J. W." box="[529,780,1360,1387]" pageId="9" pageNumber="71" pagination="1 - 361" refId="ref11259" refString="Machatschke J. W. 1972. Scarabaeoidea: Melolonthidae, Rutelinae. Coleopterorum Catalogus Supplementa 66: 1 - 361." type="journal article" year="1972">Machatschke 1972: 5</bibRefCitation>
|
||
(catalogue).
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
<paragraph id="8BC53692E15C453BFF36FAEFFE30FA4D" blockId="9.[189,1399,1184,1457]" pageId="9" pageNumber="71">
|
||
<treatmentCitationGroup id="AB6A11BCE15C453BFF36FAEFFE30FA4D" pageId="9" pageNumber="71">
|
||
<taxonomicName id="4C7A4D11E15C453BFF36FAEFFD63FA71" authorityName="Ohaus" authorityYear="1914" box="[189,720,1395,1422]" class="Insecta" family="Rutelidae" genus="Hoplognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="71" phylum="Arthropoda" rank="variety" species="helmreicheni" variety="maculicollis">
|
||
<emphasis id="B90EEA80E15C453BFF36FAEFFDB7FA71" box="[189,516,1395,1421]" italics="true" pageId="9" pageNumber="71">Hoplognathus helmreicheni</emphasis>
|
||
var.
|
||
<emphasis id="B90EEA80E15C453BFDCAFAEFFD63FA71" box="[577,720,1395,1421]" italics="true" pageId="9" pageNumber="71">maculicollis</emphasis>
|
||
</taxonomicName>
|
||
–
|
||
<treatmentCitation id="0ADB1083E15C453BFD7AFAEFFE30FA4D" author="Ohaus" page="302" pageId="9" pageNumber="71" year="1914">
|
||
<bibRefCitation id="EFEB4B63E15C453BFD7AFAEFFC79FA71" author="Ohaus F." box="[753,970,1395,1421]" pageId="9" pageNumber="71" pagination="299 - 304" refId="ref11366" refString="Ohaus F. 1914 a. Neue Coleoptera lamellicornia aus Argentinien. Deutsche Entomologische Zeitschrift 1914: 299 - 304." type="journal article" year="1914">Ohaus 1914a: 302</bibRefCitation>
|
||
(description, infrasubspecific name, unavailable).
|
||
</treatmentCitation>
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3606519E15C453BFF36FA4BFBA3F908" pageId="9" pageNumber="71" type="diagnosis">
|
||
<paragraph id="8BC53692E15C453BFF36FA4BFE85FA0D" blockId="9.[189,310,1495,1521]" box="[189,310,1495,1521]" pageId="9" pageNumber="71">
|
||
<heading id="D08D81FEE15C453BFF36FA4BFE85FA0D" bold="true" box="[189,310,1495,1521]" fontSize="11" level="3" pageId="9" pageNumber="71" reason="3">
|
||
<emphasis id="B90EEA80E15C453BFF36FA4BFE85FA0D" bold="true" box="[189,310,1495,1521]" pageId="9" pageNumber="71">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC53692E15C453BFF36F99AFBA3F908" blockId="9.[187,1399,1542,1780]" pageId="9" pageNumber="71">
|
||
Elytra reddish brown to orange yellow; male with clypeus moderately trilobate, lobes distinctly rounded, sides parallel (
|
||
<figureCitation id="13412A17E15C453BFEECF9B5FE6EF9B8" box="[359,477,1577,1604]" captionStart="Fig" captionStartId="7.[189,232,1885,1911]" captionTargetBox="[335,1250,321,1813]" captionTargetId="figure-17@7.[293,1296,265,1846]" captionTargetPageId="7" captionText="Fig. 1. a–b. Oplognathus kirbii MacLeay, 1819. a. Neotype, ♂ (NHMUK). b. Female specimen (MZUSP). c. O. helmenreichi Ohaus, 1905, holotype, ♂ (ZMHB). d–e. O. helmenreichi var.maculicollis Ohaus, 1914. d. Syntype, ♂ (ZMHB). e. Syntype, ♀ (ZMHB). f–g. O. bahianus Ohaus, 1912. f. Syntype, ♂ (ZMHB). g. Female specimen (EPGC). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514422" httpUri="https://zenodo.org/record/5514422/files/figure.png" pageId="9" pageNumber="71">Figs 1c–e</figureCitation>
|
||
,
|
||
<figureCitation id="13412A17E15C453BFE61F9B5FD94F9B8" box="[490,551,1577,1604]" captionStart="Fig" captionStartId="11.[189,232,1918,1944]" captionTargetBox="[232,1370,378,1807]" captionTargetId="figure-16@11.[189,1398,315,1828]" captionTargetPageId="11" captionText="Fig. 3. Head and pronotum details. a. Oplognathus kirbii MacLeay, 1819, male specimen (ZMHB). b. O. bahianus Ohaus, 1912, syntype, ♂ (ZMHB). c. O. helmenreichiOhaus, 1905, holotype, ♂ (ZMHB). d. O. helmenreichi var. maculicollis Ohaus, 1914, syntype, ♂ (ZMHB). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514426" httpUri="https://zenodo.org/record/5514426/files/figure.png" pageId="9" pageNumber="71">3c–d</figureCitation>
|
||
); antennal club 1.8× longer than antennomeres 2–7 combined (
|
||
<figureCitation id="13412A17E15C453BFA98F9B5FF7CF994" captionStart="Fig" captionStartId="11.[189,232,1918,1944]" captionTargetBox="[232,1370,378,1807]" captionTargetId="figure-16@11.[189,1398,315,1828]" captionTargetPageId="11" captionText="Fig. 3. Head and pronotum details. a. Oplognathus kirbii MacLeay, 1819, male specimen (ZMHB). b. O. bahianus Ohaus, 1912, syntype, ♂ (ZMHB). c. O. helmenreichiOhaus, 1905, holotype, ♂ (ZMHB). d. O. helmenreichi var. maculicollis Ohaus, 1914, syntype, ♂ (ZMHB). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514426" httpUri="https://zenodo.org/record/5514426/files/figure.png" pageId="9" pageNumber="71">Fig. 3c– d</figureCitation>
|
||
); females with club length 1.5× longer than antennomeres 2–7 combined; last maxillary palpomere cylindrical in both sexes (
|
||
<figureCitation id="13412A17E15C453BFE67F9ECFD88F977" box="[492,571,1648,1675]" captionStart="Fig" captionStartId="12.[189,232,1918,1944]" captionTargetBox="[206,1384,870,1864]" captionTargetId="figure-172@12.[189,1398,850,1878]" captionTargetPageId="12" captionText="Fig. 4. Mouthparts. a–d. Oplognathus kirbii MacLeay, 1819 (ZMHB). e–h. O. bahianus Ohaus, 1912 (ZMHB). i–m. O. helmenreichi Ohaus, 1905 (ZMHB). a, e, i. Right mandible. b, f, j. Right maxilla. c, g, l. Detail of the last maxillary palpomere. d, h, m. Mentum. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514428" httpUri="https://zenodo.org/record/5514428/files/figure.png" pageId="9" pageNumber="71">Fig. 4l</figureCitation>
|
||
); mentum longer than wide, anterior margin slightly sinuous, posterior angles rounded (
|
||
<figureCitation id="13412A17E15C453BFE0BF90FFE52F951" box="[384,481,1683,1709]" captionStart="Fig" captionStartId="12.[189,232,1918,1944]" captionTargetBox="[206,1384,870,1864]" captionTargetId="figure-172@12.[189,1398,850,1878]" captionTargetPageId="12" captionText="Fig. 4. Mouthparts. a–d. Oplognathus kirbii MacLeay, 1819 (ZMHB). e–h. O. bahianus Ohaus, 1912 (ZMHB). i–m. O. helmenreichi Ohaus, 1905 (ZMHB). a, e, i. Right mandible. b, f, j. Right maxilla. c, g, l. Detail of the last maxillary palpomere. d, h, m. Mentum. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514428" httpUri="https://zenodo.org/record/5514428/files/figure.png" pageId="9" pageNumber="71">
|
||
Fig.
|
||
<quantity id="4C829B77E15C453BFE3CF90FFE52F951" box="[439,481,1683,1709]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="9" pageNumber="71" unit="m" value="4.0">4m</quantity>
|
||
</figureCitation>
|
||
). Elytral striae distinctly marked; elytral epipleuron of female slightly dilated; mesoventral process long, apex rounded, never reaching anterior coxae (
|
||
<figureCitation id="13412A17E15C453BFB8AF92AFBD8F92D" box="[1025,1131,1718,1745]" captionStart="Fig" captionStartId="18.[189,232,1925,1951]" captionTargetBox="[214,1361,1235,1854]" captionTargetId="figure-286@18.[189,1398,1189,1887]" captionTargetPageId="18" captionText="Fig. 8. Patterns of mesoventral process. a, d. Oplognathus kirbii MacLeay, 1819 (MZUSP). b, e. O. bahianus Ohaus, 1912 (ZMHB). c, f. O. helmenreichi Ohaus, 1905 (CERPE). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514440" httpUri="https://zenodo.org/record/5514440/files/figure.png" pageId="9" pageNumber="71">Fig. 8c, f</figureCitation>
|
||
); apex of the mesotibia with 7–13 spinules; apex of metatibia with 14–22 spinules in both sexes.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3606519E15C4538FF36F887FAFBFCF9" lastPageId="10" lastPageNumber="72" pageId="9" pageNumber="71" type="materials_examined">
|
||
<paragraph id="8BC53692E15C453BFF36F887FDF4F8CA" blockId="9.[189,583,1819,1846]" box="[189,583,1819,1846]" pageId="9" pageNumber="71">
|
||
<emphasis id="B90EEA80E15C453BFF36F887FE19F8C9" bold="true" box="[189,426,1819,1845]" pageId="9" pageNumber="71">Material examined</emphasis>
|
||
(
|
||
<figureCitation id="13412A17E15C453BFE31F887FDA1F8C9" box="[442,530,1819,1845]" captionStart="Fig" captionStartId="7.[189,232,1885,1911]" captionTargetBox="[335,1250,321,1813]" captionTargetId="figure-17@7.[293,1296,265,1846]" captionTargetPageId="7" captionText="Fig. 1. a–b. Oplognathus kirbii MacLeay, 1819. a. Neotype, ♂ (NHMUK). b. Female specimen (MZUSP). c. O. helmenreichi Ohaus, 1905, holotype, ♂ (ZMHB). d–e. O. helmenreichi var.maculicollis Ohaus, 1914. d. Syntype, ♂ (ZMHB). e. Syntype, ♀ (ZMHB). f–g. O. bahianus Ohaus, 1912. f. Syntype, ♂ (ZMHB). g. Female specimen (EPGC). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514422" httpUri="https://zenodo.org/record/5514422/files/figure.png" pageId="9" pageNumber="71">Figs 1c</figureCitation>
|
||
,
|
||
<figureCitation id="13412A17E15C453BFD94F887FD8DF8CA" box="[543,574,1819,1846]" captionStart="Fig" captionStartId="8.[189,232,1920,1946]" captionTargetBox="[233,1372,1482,1865]" captionTargetId="figure-407@8.[197,1400,1458,1881]" captionTargetPageId="8" captionText="Fig. 2. Labels. a. Oplognathus bahianus Ohaus, 1912, syntype, ♂. b. O. helmenreichi Ohaus, 1905, holotype, ♂. c–d. O. helmenreichi var. maculicollis Ohaus, 1914.c. Syntype ♂. d. Syntype ♀. e. O. kirbii MacLeay, 1819, neotype, ♂." figureDoi="http://doi.org/10.5281/zenodo.5514424" httpUri="https://zenodo.org/record/5514424/files/figure.png" pageId="9" pageNumber="71">2b</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BC53692E15C453BFF61F8D6FB66F832" blockId="9.[189,1399,1866,1998]" pageId="9" pageNumber="71">
|
||
<materialsCitation id="3B123CCFE15C453BFF61F8D6FB66F832" ID-GBIF-Occurrence="3382543319" collectionCode="ZMHB" country="Argentina" location="Typus" pageId="9" pageNumber="71" specimenCount="1" specimenCount-male="1" stateProvince="Ciudad Autonoma de Buenos Aires" typeStatus="holotype">
|
||
<emphasis id="B90EEA80E15C453BFF61F8D6FEEFF898" bold="true" box="[234,348,1866,1892]" pageId="9" pageNumber="71">
|
||
<typeStatus id="54C18830E15C453BFF61F8D6FEEFF898" box="[234,348,1866,1892]" pageId="9" pageNumber="71" type="holotype">Holotype</typeStatus>
|
||
</emphasis>
|
||
<collectingCountry id="F36D7602E15C453BFF36F8F1FED6F87B" box="[189,357,1901,1927]" name="Argentina" pageId="9" pageNumber="71">ARGENTINE</collectingCountry>
|
||
•
|
||
<specimenCount id="9D7CFD1BE15C453BFE05F8F1FE15F87B" box="[398,422,1901,1927]" pageId="9" pageNumber="71" type="male">♂</specimenCount>
|
||
,
|
||
<typeStatus id="54C18830E15C453BFE30F8F1FD91F87B" box="[443,546,1901,1927]" pageId="9" pageNumber="71" type="holotype">holotype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C7A4D11E15C453BFDD2F8F1FBFBF87B" authority="Ohaus, 1905" authorityName="Ohaus" authorityYear="1905" box="[601,1096,1900,1927]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="71" phylum="Arthropoda" rank="species" species="helmenreichi">
|
||
<emphasis id="B90EEA80E15C453BFDD2F8F1FC2AF87B" box="[601,921,1901,1927]" italics="true" pageId="9" pageNumber="71">Oplognathus helmenreichi</emphasis>
|
||
<bibRefCitation id="EFEB4B63E15C453BFC23F8F1FBFBF87B" author="Ohaus F." box="[936,1096,1900,1927]" pageId="9" pageNumber="71" pagination="283 - 329" refId="ref11320" refString="Ohaus F. 1905. Beitrage zur Kenntniss der amerikanischen Ruteliden. Stettiner Entomologische Zeitung 66: 283 - 329." type="journal article" year="1905">Ohaus, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
; “
|
||
<collectingRegion id="49BEF870E15C453BFBECF8F1FAA2F87B" box="[1127,1297,1901,1927]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="9" pageNumber="71">Buenos Aires</collectingRegion>
|
||
” [white typewritten label]; “
|
||
<taxonomicName id="4C7A4D11E15C453BFE3BF80CFD7FF856" authorityName="Ohaus" authorityYear="1905" box="[432,716,1936,1962]" class="Insecta" family="Rutelidae" genus="Pelidnota" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="71" phylum="Arthropoda" rank="species" species="helmenreichi">
|
||
<emphasis id="B90EEA80E15C453BFE3BF80CFD7FF856" box="[432,716,1936,1962]" italics="true" pageId="9" pageNumber="71">Pelidnota Helmenreichi</emphasis>
|
||
</taxonomicName>
|
||
<collectingRegion id="49BEF870E15C453BFD5DF80CFCC9F856" box="[726,890,1936,1962]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="9" pageNumber="71">Buenos Aires</collectingRegion>
|
||
” [white handwritten label]; “
|
||
<location id="8EA56049E15C453BFB57F80CFA9EF856" LSID="urn:lsid:plazi:treatment:03D38784E15C453CFDA1FC65FBDFF998:8EA56049E15C453BFB57F80CFA9EF856" box="[1244,1325,1936,1962]" country="Argentina" name="Typus" pageId="9" pageNumber="71" stateProvince="Ciudad Autonoma de Buenos Aires">
|
||
<typeStatus id="54C18830E15C453BFB57F80CFA9EF856" box="[1244,1325,1936,1962]" pageId="9" pageNumber="71">Typus</typeStatus>
|
||
</location>
|
||
!” [red typewritten label]; “
|
||
<taxonomicName id="4C7A4D11E15C453BFE22F82FFCF8F831" authority="Ohaus" authorityName="Ohaus" authorityYear="1905" box="[425,843,1971,1997]" class="Insecta" family="Rutelidae" genus="Hoplognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="71" phylum="Arthropoda" rank="species" species="helmenreichi">
|
||
<emphasis id="B90EEA80E15C453BFE22F82FFD45F831" box="[425,758,1971,1997]" italics="true" pageId="9" pageNumber="71">Hoplognathus Helmenreichi</emphasis>
|
||
Ohaus
|
||
</taxonomicName>
|
||
” [red handwritten label];
|
||
<collectionCode id="ED6BAE57E15C453BFBF3F828FB62F832" box="[1144,1233,1972,1998]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="9" pageNumber="71">ZMHB</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BC53692E15F4538FF61FE94FBC4FDE5" blockId="10.[189,1400,264,537]" pageId="10" pageNumber="72">
|
||
<materialsCitation id="3B123CCFE15F4538FF61FE94FEF8FE95" ID-GBIF-Occurrence="3382543322" pageId="10" pageNumber="72" specimenCount="1" specimenCount-male="1" typeStatus="syntype">
|
||
<emphasis id="B90EEA80E15F4538FF61FE94FEE9FEDE" bold="true" box="[234,346,264,290]" pageId="10" pageNumber="72">
|
||
<typeStatus id="54C18830E15F4538FF61FE94FEE9FEDE" box="[234,346,264,290]" pageId="10" pageNumber="72" type="syntype">Syntypes</typeStatus>
|
||
</emphasis>
|
||
COUNTRY UNKNOWN •
|
||
<specimenCount id="9D7CFD1BE15F4538FD9AFEB0FD9AFEBA" box="[529,553,300,326]" pageId="10" pageNumber="72" type="male">♂</specimenCount>
|
||
, syntype of
|
||
<taxonomicName id="4C7A4D11E15F4538FD4CFEB0FAC5FEBA" authority="Ohaus, 1914" authorityName="Ohaus" authorityYear="1914" box="[711,1398,300,326]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="72" phylum="Arthropoda" rank="variety" species="helmenreichi" variety="maculicollis">
|
||
<emphasis id="B90EEA80E15F4538FD4CFEB0FC4CFEBA" box="[711,1023,300,326]" italics="true" pageId="10" pageNumber="72">Oplognathus helmenreichi</emphasis>
|
||
var.
|
||
<emphasis id="B90EEA80E15F4538FBCFFEB0FB61FEBA" box="[1092,1234,300,326]" italics="true" pageId="10" pageNumber="72">maculicollis</emphasis>
|
||
Ohaus, 1914
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13412A17E15F4538FF4EFED3FEAFFE95" box="[197,284,335,361]" captionStart="Fig" captionStartId="7.[189,232,1885,1911]" captionTargetBox="[335,1250,321,1813]" captionTargetId="figure-17@7.[293,1296,265,1846]" captionTargetPageId="7" captionText="Fig. 1. a–b. Oplognathus kirbii MacLeay, 1819. a. Neotype, ♂ (NHMUK). b. Female specimen (MZUSP). c. O. helmenreichi Ohaus, 1905, holotype, ♂ (ZMHB). d–e. O. helmenreichi var.maculicollis Ohaus, 1914. d. Syntype, ♂ (ZMHB). e. Syntype, ♀ (ZMHB). f–g. O. bahianus Ohaus, 1912. f. Syntype, ♂ (ZMHB). g. Female specimen (EPGC). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514422" httpUri="https://zenodo.org/record/5514422/files/figure.png" pageId="10" pageNumber="72">Figs 1d</figureCitation>
|
||
,
|
||
<figureCitation id="13412A17E15F4538FEADFED3FEF1FE95" box="[294,322,335,361]" captionStart="Fig" captionStartId="8.[189,232,1920,1946]" captionTargetBox="[233,1372,1482,1865]" captionTargetId="figure-407@8.[197,1400,1458,1881]" captionTargetPageId="8" captionText="Fig. 2. Labels. a. Oplognathus bahianus Ohaus, 1912, syntype, ♂. b. O. helmenreichi Ohaus, 1905, holotype, ♂. c–d. O. helmenreichi var. maculicollis Ohaus, 1914.c. Syntype ♂. d. Syntype ♀. e. O. kirbii MacLeay, 1819, neotype, ♂." figureDoi="http://doi.org/10.5281/zenodo.5514424" httpUri="https://zenodo.org/record/5514424/files/figure.png" pageId="10" pageNumber="72">2c</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B123CCFE15F4538FED5FED3FC20FE4C" ID-GBIF-Occurrence="3382543308" collectionCode="ZMHB" pageId="10" pageNumber="72" specimenCode="MFNB 2017" specimenCount="1" typeStatus="typus">
|
||
“
|
||
<typeStatus id="54C18830E15F4538FEE2FED3FE09FE95" box="[361,442,335,361]" pageId="10" pageNumber="72">Typus</typeStatus>
|
||
!” [red typewritten label]; “Helmr.” [white handwritten label]; “
|
||
<emphasis id="B90EEA80E15F4538FBF5FECCFA8FFE95" box="[1150,1340,335,361]" italics="true" pageId="10" pageNumber="72">H. Helmenreichi</emphasis>
|
||
Ohs.
|
||
<taxonomicName id="4C7A4D11E15F4538FF36FEEFFE12FE70" authority="Ohs." authorityName="Ohaus" authorityYear="1914" box="[189,417,370,397]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="72" phylum="Arthropoda" rank="variety" species="helmenreichi" variety="maculicollis">
|
||
v.
|
||
<emphasis id="B90EEA80E15F4538FF5DFEEEFED7FE70" box="[214,356,370,396]" italics="true" pageId="10" pageNumber="72">maculicollis</emphasis>
|
||
Ohs.
|
||
</taxonomicName>
|
||
” [red handwritten label]; “SYNTYPE
|
||
<taxonomicName id="4C7A4D11E15F4538FCDDFEEEFEFEFE53" authority="Ohaus, 1914" authorityName="Ohaus" authorityYear="1914" class="Insecta" family="Rutelidae" genus="Hoplognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="72" phylum="Arthropoda" rank="variety" species="helmenreicheni" variety="maculicollis">
|
||
<emphasis id="B90EEA80E15F4538FCDDFEEEFB01FE70" box="[854,1202,370,396]" italics="true" pageId="10" pageNumber="72">Hoplognathus helmenreicheni</emphasis>
|
||
var.
|
||
<emphasis id="B90EEA80E15F4538FB6DFEEEFAC7FE70" box="[1254,1396,370,396]" italics="true" pageId="10" pageNumber="72">maculicollis</emphasis>
|
||
Ohaus, 1914
|
||
</taxonomicName>
|
||
labelled by
|
||
<specimenCode id="DBDC9EE9E15F4538FE5EFE0AFDDAFE53" box="[469,617,405,432]" pageId="10" pageNumber="72">MFNB 2017</specimenCode>
|
||
” [red typed label];
|
||
<collectionCode id="ED6BAE57E15F4538FCB4FE0AFC20FE4C" box="[831,915,406,432]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="10" pageNumber="72">ZMHB</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3B123CCFE15F4538FC2CFE0AFBC4FDE5" ID-GBIF-Occurrence="3382543323" collectionCode="ZMHB" location="Oplognathus" municipality="Oplognathus" pageId="10" pageNumber="72" specimenCode="MFNB 2017" specimenCount="1" specimenCount-female="1" typeStatus="syntype">
|
||
<specimenCount id="9D7CFD1BE15F4538FC2CFE0AFC0EFE4C" box="[935,957,406,432]" pageId="10" pageNumber="72" type="female">♀</specimenCount>
|
||
,
|
||
<typeStatus id="54C18830E15F4538FC42FE0AFB90FE4C" box="[969,1059,406,432]" pageId="10" pageNumber="72" type="syntype">syntype</typeStatus>
|
||
of
|
||
<taxonomicName id="4C7A4D11E15F4538FBCEFE09FDA9FE2F" authority="Ohaus, 1914" authorityName="Ohaus" authorityYear="1914" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="72" phylum="Arthropoda" rank="variety" species="helmenreichi" variety="maculicollis">
|
||
<emphasis id="B90EEA80E15F4538FBCEFE09FAC5FE53" box="[1093,1398,405,431]" italics="true" pageId="10" pageNumber="72">
|
||
<collectingMunicipality id="6BA1ACE8E15F4538FBCEFE09FB68FE53" box="[1093,1243,405,431]" pageId="10" pageNumber="72">Oplognathus</collectingMunicipality>
|
||
helmenreichi
|
||
</emphasis>
|
||
var.
|
||
<emphasis id="B90EEA80E15F4538FF7BFE24FECDFE2E" box="[240,382,440,466]" italics="true" pageId="10" pageNumber="72">maculicollis</emphasis>
|
||
Ohaus, 1914
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13412A17E15F4538FDA1FE24FD30FE2F" box="[554,643,440,467]" captionStart="Fig" captionStartId="7.[189,232,1885,1911]" captionTargetBox="[335,1250,321,1813]" captionTargetId="figure-17@7.[293,1296,265,1846]" captionTargetPageId="7" captionText="Fig. 1. a–b. Oplognathus kirbii MacLeay, 1819. a. Neotype, ♂ (NHMUK). b. Female specimen (MZUSP). c. O. helmenreichi Ohaus, 1905, holotype, ♂ (ZMHB). d–e. O. helmenreichi var.maculicollis Ohaus, 1914. d. Syntype, ♂ (ZMHB). e. Syntype, ♀ (ZMHB). f–g. O. bahianus Ohaus, 1912. f. Syntype, ♂ (ZMHB). g. Female specimen (EPGC). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514422" httpUri="https://zenodo.org/record/5514422/files/figure.png" pageId="10" pageNumber="72">Figs 1e</figureCitation>
|
||
,
|
||
<figureCitation id="13412A17E15F4538FD1AFE24FD01FE2F" box="[657,690,440,467]" captionStart="Fig" captionStartId="8.[189,232,1920,1946]" captionTargetBox="[233,1372,1482,1865]" captionTargetId="figure-407@8.[197,1400,1458,1881]" captionTargetPageId="8" captionText="Fig. 2. Labels. a. Oplognathus bahianus Ohaus, 1912, syntype, ♂. b. O. helmenreichi Ohaus, 1905, holotype, ♂. c–d. O. helmenreichi var. maculicollis Ohaus, 1914.c. Syntype ♂. d. Syntype ♀. e. O. kirbii MacLeay, 1819, neotype, ♂." figureDoi="http://doi.org/10.5281/zenodo.5514424" httpUri="https://zenodo.org/record/5514424/files/figure.png" pageId="10" pageNumber="72">2d</figureCitation>
|
||
); “♀” [white typewritten label with black border]; “Cotype” [red typewritten label]; “Helmr.” [white handwritten label]; “SYNTYPE
|
||
<taxonomicName id="4C7A4D11E15F4538FB92FE40FDA5FDE5" authority="Ohaus, 1914" authorityName="Ohaus" authorityYear="1914" class="Insecta" family="Rutelidae" genus="Hoplognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="72" phylum="Arthropoda" rank="variety" species="helmenreicheni" variety="maculicollis">
|
||
<emphasis id="B90EEA80E15F4538FB92FE40FACBFE0A" box="[1049,1400,476,502]" italics="true" pageId="10" pageNumber="72">Hoplognathus helmenreicheni</emphasis>
|
||
var.
|
||
<emphasis id="B90EEA80E15F4538FF64FE63FECEFDE5" box="[239,381,511,537]" italics="true" pageId="10" pageNumber="72">maculicollis</emphasis>
|
||
Ohaus, 1914
|
||
</taxonomicName>
|
||
labelled by
|
||
<specimenCode id="DBDC9EE9E15F4538FD2EFE63FC88FDE5" box="[677,827,511,537]" pageId="10" pageNumber="72">MFNB 2017</specimenCode>
|
||
” [red typed label];
|
||
<collectionCode id="ED6BAE57E15F4538FB90FE63FBC0FDE5" box="[1051,1139,511,537]" collectionName="Germany, Berlin, Museum fuer Naturkunde der Humboldt-Universitaet" pageId="10" pageNumber="72">ZMHB</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BC53692E15F4538FF61FDDCFD13FDA7" blockId="10.[189,1398,576,673]" box="[234,672,576,603]" pageId="10" pageNumber="72">
|
||
<emphasis id="B90EEA80E15F4538FF61FDDCFE52FDA6" bold="true" box="[234,481,576,602]" pageId="10" pageNumber="72">Additional material</emphasis>
|
||
(
|
||
<specimenCount id="9D7CFD1BE15F4538FE7AFDDCFD2BFDA7" box="[497,664,576,603]" pageId="10" pageNumber="72" type="generic">six specimens</specimenCount>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BC53692E15F4538FF36FDF8FE91FD5D" blockId="10.[189,1398,576,673]" pageId="10" pageNumber="72">
|
||
<materialsCitation id="3B123CCFE15F4538FF36FDF8FE91FD5D" ID-GBIF-Occurrence="3382543309" collectionCode="NHMW" pageId="10" pageNumber="72" specimenCount="1" specimenCount-female="1">
|
||
COUNTRY UNKNOWN • “Ohaus determin. 1913
|
||
<emphasis id="B90EEA80E15F4538FC9DFDFFFC4EFD81" box="[790,1021,611,637]" italics="true" pageId="10" pageNumber="72">Hopl. Helmenreichi</emphasis>
|
||
<taxonomicName id="4C7A4D11E15F4538FB88FDF8FB59FD81" authority="Ohs." authorityName="Ohaus" authorityYear="1914" box="[1027,1258,611,638]" class="Insecta" family="Rutelidae" genus="Hoplognathus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="72" phylum="Arthropoda" rank="variety" species="helmenreicheni" variety="maculicollis">
|
||
v.
|
||
<emphasis id="B90EEA80E15F4538FB97FDFFFB18FD81" box="[1052,1195,611,637]" italics="true" pageId="10" pageNumber="72">maculicollis</emphasis>
|
||
Ohs.
|
||
</taxonomicName>
|
||
Cotype
|
||
<specimenCount id="9D7CFD1BE15F4538FAC1FDF8FAECFD82" box="[1354,1375,612,638]" pageId="10" pageNumber="72" type="female">♀</specimenCount>
|
||
”;
|
||
<collectionCode id="ED6BAE57E15F4538FF36FD1BFEADFD5D" box="[189,286,647,673]" collectionName="Austria, Wien, Naturhistorisches Museum Wien" pageId="10" pageNumber="72">NHMW</collectionCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8BC53692E15F4538FF36FD54FAFBFCF9" blockId="10.[189,1399,712,774]" pageId="10" pageNumber="72">
|
||
<materialsCitation id="3B123CCFE15F4538FF36FD54FDD1FCF9" ID-GBIF-Occurrence="3382543317" collectingDate="2003-12-23" collectionCode="UFMG-ICO" collectorName="M. F. Vasconcelos" country="Brazil" location="Estacao Ecologica de Acaua" municipality="Leme do Prado" pageId="10" pageNumber="72" specimenCount="4" specimenCount-male="4" stateProvince="Minas Gerais">
|
||
<collectingCountry id="F36D7602E15F4538FF36FD54FE94FD1E" box="[189,295,712,738]" name="Brazil" pageId="10" pageNumber="72">BRAZIL</collectingCountry>
|
||
–
|
||
<emphasis id="B90EEA80E15F4538FEDBFD54FE4DFD1E" bold="true" box="[336,510,712,738]" pageId="10" pageNumber="72">
|
||
<collectingRegion id="49BEF870E15F4538FEDBFD54FE4DFD1E" box="[336,510,712,738]" country="Brazil" name="Minas Gerais" pageId="10" pageNumber="72">Minas Gerais</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D7CFD1BE15F4538FDA8FD54FDDDFD1F" box="[547,622,712,739]" pageId="10" pageNumber="72" type="male">4 ♂♂</specimenCount>
|
||
;
|
||
<collectingMunicipality id="6BA1ACE8E15F4538FD0BFD55FCF2FD1E" box="[640,833,712,739]" pageId="10" pageNumber="72">Leme do Prado</collectingMunicipality>
|
||
,
|
||
<location id="8EA56049E15F4538FCD8FD55FB07FD1E" LSID="urn:lsid:plazi:treatment:03D38784E15C453CFDA1FC65FBDFF998:8EA56049E15F4538FCD8FD55FB07FD1E" box="[851,1204,712,739]" country="Brazil" municipality="Leme do Prado" name="Estacao Ecologica de Acaua" pageId="10" pageNumber="72" stateProvince="Minas Gerais">Estação Ecológica de Acauã</location>
|
||
;
|
||
<date id="FFC41052E15F4538FB4EFD54FAC1FD1E" box="[1221,1394,712,739]" pageId="10" pageNumber="72" value="2003-12-23">
|
||
<collectingDate id="EF80E9BAE15F4538FB4EFD54FAC1FD1E" box="[1221,1394,712,739]" pageId="10" pageNumber="72" value="2003-12-23">23 Dec. 2003</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="268F5344E15F4538FF36FD70FE38FCF9" box="[189,395,747,774]" pageId="10" pageNumber="72">M.F. Vasconcelos</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED6BAE57E15F4538FE46FD77FDD1FCF9" box="[461,610,747,773]" pageId="10" pageNumber="72">UFMG-ICO</collectionCode>
|
||
</materialsCitation>
|
||
. –
|
||
<materialsCitation id="3B123CCFE15F4538FD08FD76FAF0FCF9" ID-GBIF-Occurrence="3382543305" box="[643,1347,746,774]" collectingDate="2010-08-03" collectionCode="CERPE" collectorName="Oliveira" country="Brazil" location="Curitiba" pageId="10" pageNumber="72" specimenCount="1" specimenCount-male="1" stateProvince="Parana">
|
||
<emphasis id="B90EEA80E15F4538FD08FD76FD6DFCF9" bold="true" box="[643,734,746,773]" pageId="10" pageNumber="72">
|
||
<collectingRegion id="49BEF870E15F4538FD08FD76FD6DFCF9" box="[643,734,746,773]" country="Brazil" name="Parana" pageId="10" pageNumber="72">Paraná</collectingRegion>
|
||
</emphasis>
|
||
•
|
||
<specimenCount id="9D7CFD1BE15F4538FD7CFD77FC95FCFA" box="[759,806,747,774]" pageId="10" pageNumber="72" type="male">1 ♂</specimenCount>
|
||
;
|
||
<location id="8EA56049E15F4538FCB8FD77FC24FCF9" LSID="urn:lsid:plazi:treatment:03D38784E15C453CFDA1FC65FBDFF998:8EA56049E15F4538FCB8FD77FC24FCF9" box="[819,919,747,773]" country="Brazil" name="Curitiba" pageId="10" pageNumber="72" stateProvince="Parana">Curitiba</location>
|
||
;
|
||
<date id="FFC41052E15F4538FC28FD77FB85FCF9" box="[931,1078,747,774]" pageId="10" pageNumber="72" value="2010-08-03">
|
||
<collectingDate id="EF80E9BAE15F4538FC28FD77FB85FCF9" box="[931,1078,747,774]" pageId="10" pageNumber="72" value="2010-08-03">3 Aug. 2010</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="268F5344E15F4538FBC9FD77FB11FCF9" box="[1090,1186,747,773]" pageId="10" pageNumber="72">Oliveira</collectorName>
|
||
leg.;
|
||
<collectionCode id="ED6BAE57E15F4538FB6FFD77FAF0FCF9" box="[1252,1347,747,773]" pageId="10" pageNumber="72">CERPE</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3606519E15F453FFF36FCB0FCEEFEBA" lastPageId="13" lastPageNumber="75" pageId="10" pageNumber="72" type="description">
|
||
<paragraph id="8BC53692E15F4538FF36FCB0FED8FCBA" blockId="10.[189,363,812,838]" box="[189,363,812,838]" pageId="10" pageNumber="72">
|
||
<heading id="D08D81FEE15F4538FF36FCB0FED8FCBA" bold="true" box="[189,363,812,838]" fontSize="11" level="3" pageId="10" pageNumber="72" reason="3">
|
||
<emphasis id="B90EEA80E15F4538FF36FCB0FED8FCBA" bold="true" box="[189,363,812,838]" pageId="10" pageNumber="72">Redescription</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC53692E15F4538FF61FCC7FCB8FC8A" blockId="10.[189,910,859,921]" box="[234,779,859,886]" pageId="10" pageNumber="72">
|
||
<emphasis id="B90EEA80E15F4538FF61FCC7FE2EFC89" bold="true" box="[234,413,859,885]" pageId="10" pageNumber="72">
|
||
Male
|
||
<typeStatus id="54C18830E15F4538FEBAFCC7FE2EFC89" box="[305,413,859,885]" pageId="10" pageNumber="72" type="holotype">holotype</typeStatus>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="13412A17E15F4538FE27FCC7FDB6FC89" box="[428,517,859,885]" captionStart="Fig" captionStartId="7.[189,232,1885,1911]" captionTargetBox="[335,1250,321,1813]" captionTargetId="figure-17@7.[293,1296,265,1846]" captionTargetPageId="7" captionText="Fig. 1. a–b. Oplognathus kirbii MacLeay, 1819. a. Neotype, ♂ (NHMUK). b. Female specimen (MZUSP). c. O. helmenreichi Ohaus, 1905, holotype, ♂ (ZMHB). d–e. O. helmenreichi var.maculicollis Ohaus, 1914. d. Syntype, ♂ (ZMHB). e. Syntype, ♀ (ZMHB). f–g. O. bahianus Ohaus, 1912. f. Syntype, ♂ (ZMHB). g. Female specimen (EPGC). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514422" httpUri="https://zenodo.org/record/5514422/files/figure.png" pageId="10" pageNumber="72">Figs 1c</figureCitation>
|
||
,
|
||
<figureCitation id="13412A17E15F4538FD99FCC7FD83FC89" box="[530,560,859,885]" captionStart="Fig" captionStartId="11.[189,232,1918,1944]" captionTargetBox="[232,1370,378,1807]" captionTargetId="figure-16@11.[189,1398,315,1828]" captionTargetPageId="11" captionText="Fig. 3. Head and pronotum details. a. Oplognathus kirbii MacLeay, 1819, male specimen (ZMHB). b. O. bahianus Ohaus, 1912, syntype, ♂ (ZMHB). c. O. helmenreichiOhaus, 1905, holotype, ♂ (ZMHB). d. O. helmenreichi var. maculicollis Ohaus, 1914, syntype, ♂ (ZMHB). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514426" httpUri="https://zenodo.org/record/5514426/files/figure.png" pageId="10" pageNumber="72">3c</figureCitation>
|
||
,
|
||
<figureCitation id="13412A17E15F4538FDB7FCC7FDC8FC89" box="[572,635,859,885]" captionStart="Fig" captionStartId="12.[189,232,1918,1944]" captionTargetBox="[206,1384,870,1864]" captionTargetId="figure-172@12.[189,1398,850,1878]" captionTargetPageId="12" captionText="Fig. 4. Mouthparts. a–d. Oplognathus kirbii MacLeay, 1819 (ZMHB). e–h. O. bahianus Ohaus, 1912 (ZMHB). i–m. O. helmenreichi Ohaus, 1905 (ZMHB). a, e, i. Right mandible. b, f, j. Right maxilla. c, g, l. Detail of the last maxillary palpomere. d, h, m. Mentum. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514428" httpUri="https://zenodo.org/record/5514428/files/figure.png" pageId="10" pageNumber="72">4i–m</figureCitation>
|
||
,
|
||
<figureCitation id="13412A17E15F4538FD0CFCC7FD72FC89" box="[647,705,859,885]" captionStart="Fig" captionStartId="15.[189,232,1921,1947]" captionTargetBox="[277,1269,617,1876]" captionTargetId="figure-111@15.[189,1398,583,1882]" captionTargetPageId="15" captionText="Fig. 6. Aedeagus in dorsal (a, e) and lateral view (b–c, d–f). a–c. Oplognathus helmenreichi Ohaus, 1905, holotype, ♂ (ZMHB). d–f. O. helmenreichi var. maculicollis Ohaus, 1914, syntype, ♂ (ZMHB). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514434" httpUri="https://zenodo.org/record/5514434/files/figure.png" pageId="10" pageNumber="72">6a–c</figureCitation>
|
||
,
|
||
<figureCitation id="13412A17E15F4538FD46FCC7FCB7FC8A" box="[717,772,859,886]" captionStart="Fig" captionStartId="17.[189,232,1924,1950]" captionTargetBox="[270,1309,269,1863]" captionTargetId="figure-16@17.[243,1344,265,1886]" captionTargetPageId="17" captionText="Fig. 7. Variation in male genitalia. a–d. Oplognathus kirbii MacLeay, 1819 (a-b: CEMT; c-d: ZMHB). e–h. O. helmenreichi Ohaus, 1905 (ZMHB). i–m. O. bahianus Ohaus, 1912 (CERPE). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514436" httpUri="https://zenodo.org/record/5514436/files/figure.png" pageId="10" pageNumber="72">7e–f</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BC53692E15F4538FF36FCE3FC3DFC65" blockId="10.[189,910,859,921]" box="[189,910,894,921]" pageId="10" pageNumber="72">
|
||
SIZE. Total length:
|
||
<quantity id="4C829B77E15F4538FE13FCE2FDB7FC65" box="[408,516,894,921]" metricMagnitude="-2" metricUnit="m" metricValue="1.76" pageId="10" pageNumber="72" unit="mm" value="17.6">17.6 mm</quantity>
|
||
; width across prothorax:
|
||
<quantity id="4C829B77E15F4538FCA7FCE3FC3AFC65" box="[812,905,895,921]" metricMagnitude="-3" metricUnit="m" metricValue="9.1" pageId="10" pageNumber="72" unit="mm" value="9.1">9.1 mm</quantity>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BC53692E15F4538FF36FC5CFDB5FC01" blockId="10.[187,1399,960,1022]" pageId="10" pageNumber="72">COLOUR. General aspect metallic dark green and reddish brown; head, pronotum, legs and venter metallic green, elytra reddish brown.</paragraph>
|
||
<paragraph id="8BC53692E15F4538FF36FBB9FAB6FAC9" blockId="10.[188,1399,1061,1334]" pageId="10" pageNumber="72">
|
||
HEAD. Clypeus with surface rugose, densely punctate, larges punctures, gradually coalescent from disc to margins; apex trilobate, lobes rounded, sides parallels with the slightly converging apex. Frontoclypeal suture obsolete, sinuous, forming an angle on disc. Clypeus and frons subequal in length. Frons subquadrangular, moderately punctate, more densely posteriorly, with medium to large punctures, coalescent in the margins (
|
||
<figureCitation id="13412A17E15F4538FE7FFB2DFDFAFB30" box="[500,585,1201,1228]" captionStart="Fig" captionStartId="11.[189,232,1918,1944]" captionTargetBox="[232,1370,378,1807]" captionTargetId="figure-16@11.[189,1398,315,1828]" captionTargetPageId="11" captionText="Fig. 3. Head and pronotum details. a. Oplognathus kirbii MacLeay, 1819, male specimen (ZMHB). b. O. bahianus Ohaus, 1912, syntype, ♂ (ZMHB). c. O. helmenreichiOhaus, 1905, holotype, ♂ (ZMHB). d. O. helmenreichi var. maculicollis Ohaus, 1914, syntype, ♂ (ZMHB). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514426" httpUri="https://zenodo.org/record/5514426/files/figure.png" pageId="10" pageNumber="72">Fig. 3c</figureCitation>
|
||
). Maxillary palpi with last palpomere cylindrical, sensorial area oval, occupying two thirds of dorsal area (
|
||
<figureCitation id="13412A17E15F4538FD08FB49FD6BFB0C" box="[643,728,1237,1264]" captionStart="Fig" captionStartId="12.[189,232,1918,1944]" captionTargetBox="[206,1384,870,1864]" captionTargetId="figure-172@12.[189,1398,850,1878]" captionTargetPageId="12" captionText="Fig. 4. Mouthparts. a–d. Oplognathus kirbii MacLeay, 1819 (ZMHB). e–h. O. bahianus Ohaus, 1912 (ZMHB). i–m. O. helmenreichi Ohaus, 1905 (ZMHB). a, e, i. Right mandible. b, f, j. Right maxilla. c, g, l. Detail of the last maxillary palpomere. d, h, m. Mentum. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514428" httpUri="https://zenodo.org/record/5514428/files/figure.png" pageId="10" pageNumber="72">Fig. 4l</figureCitation>
|
||
). Mentum near 1.2× longer than wide, anterior margin weakly sinuous, almost straight, posterior angles rounded (
|
||
<figureCitation id="13412A17E15F4538FC00FB64FC43FAEE" box="[907,1008,1272,1298]" captionStart="Fig" captionStartId="12.[189,232,1918,1944]" captionTargetBox="[206,1384,870,1864]" captionTargetId="figure-172@12.[189,1398,850,1878]" captionTargetPageId="12" captionText="Fig. 4. Mouthparts. a–d. Oplognathus kirbii MacLeay, 1819 (ZMHB). e–h. O. bahianus Ohaus, 1912 (ZMHB). i–m. O. helmenreichi Ohaus, 1905 (ZMHB). a, e, i. Right mandible. b, f, j. Right maxilla. c, g, l. Detail of the last maxillary palpomere. d, h, m. Mentum. Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514428" httpUri="https://zenodo.org/record/5514428/files/figure.png" pageId="10" pageNumber="72">
|
||
Fig.
|
||
<quantity id="4C829B77E15F4538FC4DFB64FC43FAEE" box="[966,1008,1272,1298]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="10" pageNumber="72" unit="m" value="4.0">4m</quantity>
|
||
</figureCitation>
|
||
); surface sparsely punctate with scattered setae, punctures large. Antennal club 1.8× longer than antennomeres 2–7 combined.
|
||
</paragraph>
|
||
<paragraph id="8BC53692E15F4538FF36FAC1FB17F906" blockId="10.[188,1400,1372,1786]" pageId="10" pageNumber="72">
|
||
THORAX. Pronotum with surface moderate to densely punctate, increasing density towards sides; anterior angles acute; sides slightly truncate (
|
||
<figureCitation id="13412A17E15F4538FD5BFA1CFC96FA66" box="[720,805,1408,1434]" captionStart="Fig" captionStartId="11.[189,232,1918,1944]" captionTargetBox="[232,1370,378,1807]" captionTargetId="figure-16@11.[189,1398,315,1828]" captionTargetPageId="11" captionText="Fig. 3. Head and pronotum details. a. Oplognathus kirbii MacLeay, 1819, male specimen (ZMHB). b. O. bahianus Ohaus, 1912, syntype, ♂ (ZMHB). c. O. helmenreichiOhaus, 1905, holotype, ♂ (ZMHB). d. O. helmenreichi var. maculicollis Ohaus, 1914, syntype, ♂ (ZMHB). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514426" httpUri="https://zenodo.org/record/5514426/files/figure.png" pageId="10" pageNumber="72">Fig. 3c</figureCitation>
|
||
). Propleuron slightly concave, surface rugose, and moderately setose. Scutellar plate moderately punctate, punctures bigger on disc. Elytral striae distinctly marked; first interstriae with puncture thin, dense. Epipleuron flattened, sparsely setose, minute setae. Mesoventral process long with rounded apex, never reaching procoxal base (
|
||
<figureCitation id="13412A17E15F4538FBBCFA75FB10F9F8" box="[1079,1187,1513,1540]" captionStart="Fig" captionStartId="18.[189,232,1925,1951]" captionTargetBox="[214,1361,1235,1854]" captionTargetId="figure-286@18.[189,1398,1189,1887]" captionTargetPageId="18" captionText="Fig. 8. Patterns of mesoventral process. a, d. Oplognathus kirbii MacLeay, 1819 (MZUSP). b, e. O. bahianus Ohaus, 1912 (ZMHB). c, f. O. helmenreichi Ohaus, 1905 (CERPE). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514440" httpUri="https://zenodo.org/record/5514440/files/figure.png" pageId="10" pageNumber="72">Fig. 8c, f</figureCitation>
|
||
); surface punctate and densely setose. Metasternum densely punctate and densely setose. Procoxa densely punctate and sparsely setose. Profemur sparsely punctate and sparsely setose. Mesofemur moderately punctate and densely setose. Metafemur flattened, broad, sparsely punctate, and with sparse posterior setae. Protibia with surface densely punctate, bigger punctures, C-shaped punctures on disc; apex with 10 distinct setae. Mesotibia with two transverse carinae: proximal carina marked by four spinules, distal carina marked by five spinules, surface moderately punctate and sparsely setose, short setae; apex with 11 spinules. Metatibia similar to mesotibia, but more robust and with 19 apical spinules.
|
||
</paragraph>
|
||
<paragraph id="8BC53692E15F4538FF36F8BDFE00F835" blockId="10.[189,1399,1825,1993]" pageId="10" pageNumber="72">
|
||
ABDOMEN. Pygidium subtrapezoidal, apex slightly rounded, surface rugose, minute setae sparsely disposed on margins. Ventrites with surface sparsely setose, fine and short setae. Parameres similar in length, irregularly V-shaped bifurcated, bifurcation base rounded; apical lobes well defined, left lobe smaller than right lobe; left lateral expanded in lateral view, parabolic expansion; right expansion obsolete (
|
||
<figureCitation id="13412A17E15F4538FEA4F832FE15F835" box="[303,422,1966,1993]" captionStart="Fig" captionStartId="15.[189,232,1921,1947]" captionTargetBox="[277,1269,617,1876]" captionTargetId="figure-111@15.[189,1398,583,1882]" captionTargetPageId="15" captionText="Fig. 6. Aedeagus in dorsal (a, e) and lateral view (b–c, d–f). a–c. Oplognathus helmenreichi Ohaus, 1905, holotype, ♂ (ZMHB). d–f. O. helmenreichi var. maculicollis Ohaus, 1914, syntype, ♂ (ZMHB). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514434" httpUri="https://zenodo.org/record/5514434/files/figure.png" pageId="10" pageNumber="72">Figs 6a–c</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<caption id="DF05661AE15E4539FF36F8E2FB52F823" ID-DOI="http://doi.org/10.5281/zenodo.5514426" ID-Zenodo-Dep="5514426" httpUri="https://zenodo.org/record/5514426/files/figure.png" pageId="11" pageNumber="73" startId="11.[189,232,1918,1944]" targetBox="[232,1370,378,1807]" targetPageId="11">
|
||
<paragraph id="8BC53692E15E4539FF36F8E2FB52F823" blockId="11.[189,1399,1918,2015]" pageId="11" pageNumber="73">
|
||
<emphasis id="B90EEA80E15E4539FF36F8E2FEBDF864" bold="true" box="[189,270,1918,1944]" pageId="11" pageNumber="73">Fig. 3.</emphasis>
|
||
Head and pronotum details.
|
||
<emphasis id="B90EEA80E15E4539FDF9F8E2FD32F864" bold="true" box="[626,641,1918,1944]" pageId="11" pageNumber="73">a</emphasis>
|
||
.
|
||
<taxonomicName id="4C7A4D11E15E4539FD19F8E2FB89F864" authority="MacLeay, 1819" authorityName="MacLeay" authorityYear="1819" box="[658,1082,1918,1945]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="73" phylum="Arthropoda" rank="species" species="kirbii">
|
||
<emphasis id="B90EEA80E15E4539FD19F8E2FCC7F864" box="[658,884,1918,1944]" italics="true" pageId="11" pageNumber="73">Oplognathus kirbii</emphasis>
|
||
<bibRefCitation id="EFEB4B63E15E4539FCF4F8E3FB89F864" author="MacLeay WS" box="[895,1082,1918,1945]" pageId="11" pageNumber="73" refId="ref11281" refString="MacLeay WS. 1819. Horae entomologicae: or, Essays on the Annulose Animals. R. and A. Taylor, London. https: // doi. org / 10.5962 / bhl. title. 65809" type="book" year="1819">MacLeay, 1819</bibRefCitation>
|
||
</taxonomicName>
|
||
, male specimen (ZMHB).
|
||
<emphasis id="B90EEA80E15E4539FF36F83DFF7EF847" bold="true" box="[189,205,1953,1979]" pageId="11" pageNumber="73">b</emphasis>
|
||
.
|
||
<taxonomicName id="4C7A4D11E15E4539FF51F83EFDB3F847" authority="Ohaus, 1912" authorityName="Ohaus" authorityYear="1912" box="[218,512,1953,1980]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="73" phylum="Arthropoda" rank="species" species="bahianus">
|
||
<emphasis id="B90EEA80E15E4539FF51F83EFED4F847" box="[218,359,1953,1980]" italics="true" pageId="11" pageNumber="73">O. bahianus</emphasis>
|
||
<bibRefCitation id="EFEB4B63E15E4539FEE7F83DFDB3F847" author="Ohaus F." box="[364,512,1953,1979]" pageId="11" pageNumber="73" pagination="650" refId="ref11341" refString="Ohaus F. 1912. Hoplognathus bahianus n. sp. (Col. Rutel.). Deutsche Entomologische Zeitschrift 1912: 650." type="journal article" year="1912">Ohaus, 1912</bibRefCitation>
|
||
</taxonomicName>
|
||
, syntype, ♂ (ZMHB).
|
||
<emphasis id="B90EEA80E15E4539FC89F83DFCBCF847" bold="true" box="[770,783,1953,1979]" pageId="11" pageNumber="73">c</emphasis>
|
||
.
|
||
<taxonomicName id="4C7A4D11E15E4539FC90F83EFBDFF840" authority="Ohaus, 1905" authorityName="Ohaus" authorityYear="1905" box="[795,1132,1953,1980]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="73" phylum="Arthropoda" rank="species" species="helmenreichi">
|
||
<emphasis id="B90EEA80E15E4539FC90F83EFC66F847" box="[795,981,1953,1980]" italics="true" pageId="11" pageNumber="73">O. helmenreichi</emphasis>
|
||
<bibRefCitation id="EFEB4B63E15E4539FC53F83DFBDFF840" author="Ohaus F." box="[984,1132,1953,1980]" pageId="11" pageNumber="73" pagination="283 - 329" refId="ref11320" refString="Ohaus F. 1905. Beitrage zur Kenntniss der amerikanischen Ruteliden. Stettiner Entomologische Zeitung 66: 283 - 329." type="journal article" year="1905">Ohaus, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
, holotype, ♂ (ZMHB).
|
||
<emphasis id="B90EEA80E15E4539FF36F859FF7EF823" bold="true" box="[189,205,1989,2015]" pageId="11" pageNumber="73">d</emphasis>
|
||
.
|
||
<taxonomicName id="4C7A4D11E15E4539FF57F859FD4CF823" authority="Ohaus, 1914" authorityName="Ohaus" authorityYear="1914" box="[220,767,1988,2015]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="73" phylum="Arthropoda" rank="variety" species="helmenreichi" variety="maculicollis">
|
||
<emphasis id="B90EEA80E15E4539FF57F859FE2BF822" box="[220,408,1988,2015]" italics="true" pageId="11" pageNumber="73">O. helmenreichi</emphasis>
|
||
var.
|
||
<emphasis id="B90EEA80E15E4539FE59F858FDD2F822" box="[466,609,1988,2014]" italics="true" pageId="11" pageNumber="73">maculicollis</emphasis>
|
||
Ohaus, 1914
|
||
</taxonomicName>
|
||
, syntype, ♂ (ZMHB). Scale bar = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC53692E159453EFF61FE95FE10FEDF" blockId="12.[188,1401,265,573]" box="[234,419,265,291]" pageId="12" pageNumber="74">
|
||
<emphasis id="B90EEA80E159453EFF61FE95FE10FEDF" bold="true" box="[234,419,265,291]" pageId="12" pageNumber="74">Male variation</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8BC53692E159453EFF36FEB0FDB6FDC1" blockId="12.[188,1401,265,573]" pageId="12" pageNumber="74">
|
||
Size: total length: 17.2–19.0 mm; width across prothorax:
|
||
<quantity id="4C829B77E159453EFCFFFEB0FBB5FEBB" box="[884,1030,300,327]" metricMagnitude="-3" metricUnit="m" metricValue="8.3" metricValueMax="9.1" metricValueMin="7.5" pageId="12" pageNumber="74" unit="mm" value="8.3" valueMax="9.1" valueMin="7.5">7.5–9.1 mm</quantity>
|
||
. General aspect orange-brown; head, pronotum and scutellar plate may have yellowish maculae (
|
||
<figureCitation id="13412A17E159453EFC49FED3FB86FE96" box="[962,1077,335,362]" captionStart="Fig" captionStartId="7.[189,232,1885,1911]" captionTargetBox="[335,1250,321,1813]" captionTargetId="figure-17@7.[293,1296,265,1846]" captionTargetPageId="7" captionText="Fig. 1. a–b. Oplognathus kirbii MacLeay, 1819. a. Neotype, ♂ (NHMUK). b. Female specimen (MZUSP). c. O. helmenreichi Ohaus, 1905, holotype, ♂ (ZMHB). d–e. O. helmenreichi var.maculicollis Ohaus, 1914. d. Syntype, ♂ (ZMHB). e. Syntype, ♀ (ZMHB). f–g. O. bahianus Ohaus, 1912. f. Syntype, ♂ (ZMHB). g. Female specimen (EPGC). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514422" httpUri="https://zenodo.org/record/5514422/files/figure.png" pageId="12" pageNumber="74">Fig. 1d–e</figureCitation>
|
||
). Suture frontoclypeal vary in the degree of sinuosity. Pronotum can be more or less truncate at sides. Surface of mesofemur varies from moderately to densely punctate. Mesotibia with proximal and distal carina marked by 4–7 spinules; apex truncate with 7–13 spinules. Metatibia with 14–22 spinules in the apex, distal carina with 5–11 spinules, and proximal carina with 5–8. Parameres vary from parabolic curvature of the aedeagus to the truncate curvature, as well as, in the degree and shape of lateral expansion, it can be longer and more parabolic (
|
||
<figureCitation id="13412A17E159453EFEB0FDBEFE1CFDC0" box="[315,431,546,572]" captionStart="Fig" captionStartId="15.[189,232,1921,1947]" captionTargetBox="[277,1269,617,1876]" captionTargetId="figure-111@15.[189,1398,583,1882]" captionTargetPageId="15" captionText="Fig. 6. Aedeagus in dorsal (a, e) and lateral view (b–c, d–f). a–c. Oplognathus helmenreichi Ohaus, 1905, holotype, ♂ (ZMHB). d–f. O. helmenreichi var. maculicollis Ohaus, 1914, syntype, ♂ (ZMHB). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514434" httpUri="https://zenodo.org/record/5514434/files/figure.png" pageId="12" pageNumber="74">Figs 6d–f</figureCitation>
|
||
,
|
||
<figureCitation id="13412A17E159453EFE30FDBEFE4BFDC1" box="[443,504,546,573]" captionStart="Fig" captionStartId="17.[189,232,1924,1950]" captionTargetBox="[270,1309,269,1863]" captionTargetId="figure-16@17.[243,1344,265,1886]" captionTargetPageId="17" captionText="Fig. 7. Variation in male genitalia. a–d. Oplognathus kirbii MacLeay, 1819 (a-b: CEMT; c-d: ZMHB). e–h. O. helmenreichi Ohaus, 1905 (ZMHB). i–m. O. bahianus Ohaus, 1912 (CERPE). Scale bar = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5514436" httpUri="https://zenodo.org/record/5514436/files/figure.png" pageId="12" pageNumber="74">7e–h</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC53692E159453EFF61FDF8FE01FD82" blockId="12.[189,1399,612,744]" box="[234,434,612,638]" pageId="12" pageNumber="74">
|
||
<emphasis id="B90EEA80E159453EFF61FDF8FEF6FD82" bold="true" box="[234,325,612,638]" pageId="12" pageNumber="74">Female</emphasis>
|
||
(
|
||
<figureCitation id="13412A17E159453EFEDDFDF8FE1BFD82" box="[342,424,612,638]" captionStart="Fig" captionStartId="7.[189,232,1885,1911]" captionTargetBox="[335,1250,321,1813]" captionTargetId="figure-17@7.[293,1296,265,1846]" captionTargetPageId="7" captionText="Fig. 1. a–b. Oplognathus kirbii MacLeay, 1819. a. Neotype, ♂ (NHMUK). b. Female specimen (MZUSP). c. O. helmenreichi Ohaus, 1905, holotype, ♂ (ZMHB). d–e. O. helmenreichi var.maculicollis Ohaus, 1914. d. Syntype, ♂ (ZMHB). e. Syntype, ♀ (ZMHB). f–g. O. bahianus Ohaus, 1912. f. Syntype, ♂ (ZMHB). g. Female specimen (EPGC). Scale bar = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5514422" httpUri="https://zenodo.org/record/5514422/files/figure.png" pageId="12" pageNumber="74">Fig. 1e</figureCitation>
|
||
)
|
||
</paragraph>
|
||
<paragraph id="8BC53692E159453FFF36FD1BFCEEFEBA" blockId="12.[189,1399,612,744]" lastBlockId="13.[188,1399,265,327]" lastPageId="13" lastPageNumber="75" pageId="12" pageNumber="74">Similar to males in general aspects, differentiating in the following aspects: pronotal maculae more elongate; clypeus rounded; frontoclypeal suture distinct, with central curvature; antennal club 0.7× smaller than in males, 1.5× longer than antennomeres 2–7 combined. Pronotum with parallel sides and anterior angles more rounded. Elytral epipleuron slightly rounded, distinct depression above epipleuron. Protarsomere I as long as protarsomeres II–IV combined.</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF05661AE159453EFF36F8E2FB34F823" ID-DOI="http://doi.org/10.5281/zenodo.5514428" ID-Zenodo-Dep="5514428" httpUri="https://zenodo.org/record/5514428/files/figure.png" pageId="12" pageNumber="74" startId="12.[189,232,1918,1944]" targetBox="[206,1384,870,1864]" targetPageId="12">
|
||
<paragraph id="8BC53692E159453EFF36F8E2FB34F823" blockId="12.[189,1399,1918,2015]" pageId="12" pageNumber="74">
|
||
<emphasis id="B90EEA80E159453EFF36F8E2FEBEF864" bold="true" box="[189,269,1918,1944]" pageId="12" pageNumber="74">Fig. 4.</emphasis>
|
||
Mouthparts.
|
||
<emphasis id="B90EEA80E159453EFE25F8E2FE69F864" bold="true" box="[430,474,1918,1944]" pageId="12" pageNumber="74">a–d</emphasis>
|
||
.
|
||
<taxonomicName id="4C7A4D11E159453EFE61F8E2FC38F864" authority="MacLeay, 1819" authorityName="MacLeay" authorityYear="1819" box="[490,907,1918,1945]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="74" phylum="Arthropoda" rank="species" species="kirbii">
|
||
<emphasis id="B90EEA80E159453EFE61F8E2FD78F864" box="[490,715,1918,1944]" italics="true" pageId="12" pageNumber="74">Oplognathus kirbii</emphasis>
|
||
<bibRefCitation id="EFEB4B63E159453EFD58F8E3FC38F864" author="MacLeay WS" box="[723,907,1918,1945]" pageId="12" pageNumber="74" refId="ref11281" refString="MacLeay WS. 1819. Horae entomologicae: or, Essays on the Annulose Animals. R. and A. Taylor, London. https: // doi. org / 10.5962 / bhl. title. 65809" type="book" year="1819">MacLeay, 1819</bibRefCitation>
|
||
</taxonomicName>
|
||
(ZMHB).
|
||
<emphasis id="B90EEA80E159453EFB86F8E2FB8BF864" bold="true" box="[1037,1080,1918,1944]" pageId="12" pageNumber="74">e–h</emphasis>
|
||
.
|
||
<taxonomicName id="4C7A4D11E159453EFBCCF8E3FAC4F864" authority="Ohaus, 1912" authorityName="Ohaus" authorityYear="1912" box="[1095,1399,1918,1945]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="74" phylum="Arthropoda" rank="species" species="bahianus">
|
||
<emphasis id="B90EEA80E159453EFBCCF8E3FB6AF864" box="[1095,1241,1918,1945]" italics="true" pageId="12" pageNumber="74">O. bahianus</emphasis>
|
||
<bibRefCitation id="EFEB4B63E159453EFB6AF8E2FAC4F864" author="Ohaus F." box="[1249,1399,1918,1944]" pageId="12" pageNumber="74" pagination="650" refId="ref11341" refString="Ohaus F. 1912. Hoplognathus bahianus n. sp. (Col. Rutel.). Deutsche Entomologische Zeitschrift 1912: 650." type="journal article" year="1912">Ohaus, 1912</bibRefCitation>
|
||
</taxonomicName>
|
||
(ZMHB).
|
||
<emphasis id="B90EEA80E159453EFEBDF83DFED6F847" bold="true" box="[310,357,1953,1979]" pageId="12" pageNumber="74">i–m</emphasis>
|
||
.
|
||
<taxonomicName id="4C7A4D11E159453EFEFDF83EFD66F840" authority="Ohaus, 1905" authorityName="Ohaus" authorityYear="1905" box="[374,725,1953,1980]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="74" phylum="Arthropoda" rank="species" species="helmenreichi">
|
||
<emphasis id="B90EEA80E159453EFEFDF83EFD87F847" box="[374,564,1953,1980]" italics="true" pageId="12" pageNumber="74">O. helmenreichi</emphasis>
|
||
<bibRefCitation id="EFEB4B63E159453EFDB5F83DFD66F840" author="Ohaus F." box="[574,725,1953,1980]" pageId="12" pageNumber="74" pagination="283 - 329" refId="ref11320" refString="Ohaus F. 1905. Beitrage zur Kenntniss der amerikanischen Ruteliden. Stettiner Entomologische Zeitung 66: 283 - 329." type="journal article" year="1905">Ohaus, 1905</bibRefCitation>
|
||
</taxonomicName>
|
||
(ZMHB).
|
||
<emphasis id="B90EEA80E159453EFCD2F83DFC2FF847" bold="true" box="[857,924,1953,1979]" pageId="12" pageNumber="74">a, e, i</emphasis>
|
||
. Right mandible.
|
||
<emphasis id="B90EEA80E159453EFBF3F83DFB0FF847" bold="true" box="[1144,1212,1953,1979]" pageId="12" pageNumber="74">b, f, j</emphasis>
|
||
. Right maxilla.
|
||
<emphasis id="B90EEA80E159453EFF34F859FF4DF823" bold="true" box="[191,254,1989,2015]" pageId="12" pageNumber="74">c, g, l</emphasis>
|
||
. Detail of the last maxillary palpomere.
|
||
<emphasis id="B90EEA80E159453EFD52F859FC9CF823" bold="true" box="[729,815,1989,2015]" pageId="12" pageNumber="74">d, h, m</emphasis>
|
||
. Mentum. Scale bar = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3606519E158453CFF36FEF2FE89FE95" lastPageId="14" lastPageNumber="76" pageId="13" pageNumber="75" type="materials_examined">
|
||
<paragraph id="8BC53692E158453FFF36FEF2FEEEFE7B" blockId="13.[189,349,365,392]" box="[189,349,365,392]" pageId="13" pageNumber="75">
|
||
<heading id="D08D81FEE158453FFF36FEF2FEEEFE7B" bold="true" box="[189,349,365,392]" fontSize="11" level="3" pageId="13" pageNumber="75" reason="3">
|
||
<emphasis id="B90EEA80E158453FFF36FEF2FEEEFE7B" bold="true" box="[189,349,365,392]" pageId="13" pageNumber="75">Type locality</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC53692E158453CFF36FE00FE89FE95" blockId="13.[187,1399,412,509]" lastBlockId="14.[189,1399,265,361]" lastPageId="14" lastPageNumber="76" pageId="13" pageNumber="75">
|
||
<taxonomicName id="4C7A4D11E158453FFF36FE00FE46FE4A" authorityName="Ohaus" authorityYear="1905" box="[189,501,412,438]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="75" phylum="Arthropoda" rank="species" species="helmenreichi">
|
||
<emphasis id="B90EEA80E158453FFF36FE00FE46FE4A" box="[189,501,412,438]" italics="true" pageId="13" pageNumber="75">Oplognathus helmenreichi</emphasis>
|
||
</taxonomicName>
|
||
was originally described from
|
||
<collectingRegion id="49BEF870E158453FFCEDFE01FBB7FE4A" box="[870,1028,412,439]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="13" pageNumber="75">Buenos Aires</collectingRegion>
|
||
(
|
||
<collectingCountry id="F36D7602E158453FFB98FE00FB3FFE4B" box="[1043,1164,412,439]" name="Argentina" pageId="13" pageNumber="75">Argentina</collectingCountry>
|
||
) (
|
||
<bibRefCitation id="EFEB4B63E158453FFB2FFE00FA85FE4B" author="Ohaus F." box="[1188,1334,412,439]" pageId="13" pageNumber="75" pagination="283 - 329" refId="ref11320" refString="Ohaus F. 1905. Beitrage zur Kenntniss der amerikanischen Ruteliden. Stettiner Entomologische Zeitung 66: 283 - 329." type="journal article" year="1905">Ohaus 1905</bibRefCitation>
|
||
), and this locality was later contested by the author (
|
||
<bibRefCitation id="EFEB4B63E158453FFD47FE23FCD4FE25" author="Ohaus F." box="[716,871,447,473]" pageId="13" pageNumber="75" pagination="299 - 304" refId="ref11366" refString="Ohaus F. 1914 a. Neue Coleoptera lamellicornia aus Argentinien. Deutsche Entomologische Zeitschrift 1914: 299 - 304." type="journal article" year="1914">Ohaus 1914a</bibRefCitation>
|
||
) while describing the variety,
|
||
<taxonomicName id="4C7A4D11E158453FFB30FE5CFECCFE00" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="75" phylum="Arthropoda" rank="variety" species="helmenreichi" variety="macullicolis">
|
||
<emphasis id="B90EEA80E158453FFB30FE5CFAC7FE25" box="[1211,1396,447,474]" italics="true" pageId="13" pageNumber="75">O. helmenreichi</emphasis>
|
||
var.
|
||
<emphasis id="B90EEA80E158453FFF7BFE7EFECCFE00" box="[240,383,482,508]" italics="true" pageId="13" pageNumber="75">macullicolis</emphasis>
|
||
</taxonomicName>
|
||
. This species can actually be found in the north and northeast of Minas Gerais state in the Jequitinhonha River Valley (
|
||
<bibRefCitation id="EFEB4B63E15B453CFDB3FE95FD6AFEDF" author="Ohaus F." box="[568,729,265,291]" pageId="14" pageNumber="76" pagination="299 - 304" refId="ref11366" refString="Ohaus F. 1914 a. Neue Coleoptera lamellicornia aus Argentinien. Deutsche Entomologische Zeitschrift 1914: 299 - 304." type="journal article" year="1914">Ohaus 1914a</bibRefCitation>
|
||
). Accordingly, after the examination of three more males from the same region, we agree with
|
||
<bibRefCitation id="EFEB4B63E15B453CFDFAFEB0FCAEFEBA" author="Ohaus F." box="[625,797,300,326]" pageId="14" pageNumber="76" pagination="299 - 304" refId="ref11366" refString="Ohaus F. 1914 a. Neue Coleoptera lamellicornia aus Argentinien. Deutsche Entomologische Zeitschrift 1914: 299 - 304." type="journal article" year="1914">Ohaus (1914a)</bibRefCitation>
|
||
and disregard the occurrence of
|
||
<taxonomicName id="4C7A4D11E15B453CFB16FEB1FAEBFEBA" authorityName="Ohaus" authorityYear="1905" box="[1181,1368,300,327]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="76" phylum="Arthropoda" rank="species" species="helmenreichi">
|
||
<emphasis id="B90EEA80E15B453CFB16FEB1FAEBFEBA" box="[1181,1368,300,327]" italics="true" pageId="14" pageNumber="76">O. helmenreichi</emphasis>
|
||
</taxonomicName>
|
||
in
|
||
<collectingCountry id="F36D7602E15B453CFF36FED3FE85FE95" box="[189,310,335,361]" name="Argentina" pageId="14" pageNumber="76">Argentina</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF05661AE158453FFF36F83AFB46F818" ID-DOI="http://doi.org/10.5281/zenodo.5514432" ID-Zenodo-Dep="5514432" httpUri="https://zenodo.org/record/5514432/files/figure.png" pageId="13" pageNumber="75" startId="13.[189,232,1958,1984]" targetBox="[273,1281,606,1893]" targetPageId="13">
|
||
<paragraph id="8BC53692E158453FFF36F83AFB46F818" blockId="13.[189,1399,1958,2020]" pageId="13" pageNumber="75">
|
||
<emphasis id="B90EEA80E158453FFF36F83AFEBCF83C" bold="true" box="[189,271,1958,1984]" pageId="13" pageNumber="75">Fig. 5.</emphasis>
|
||
Aedeagus in dorsal (a, e) y lateral view (b–c, d, f).
|
||
<emphasis id="B90EEA80E158453FFC18F83AFC0DF83C" bold="true" box="[915,958,1958,1984]" pageId="13" pageNumber="75">a–c</emphasis>
|
||
.
|
||
<taxonomicName id="4C7A4D11E158453FFC5BF83AFAC4F83C" authority="MacLeay, 1819" authorityName="MacLeay" authorityYear="1819" box="[976,1399,1958,1985]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="75" phylum="Arthropoda" rank="species" species="kirbii">
|
||
<emphasis id="B90EEA80E158453FFC5BF83AFB01F83C" box="[976,1202,1958,1984]" italics="true" pageId="13" pageNumber="75">Oplognathus kirbii</emphasis>
|
||
<bibRefCitation id="EFEB4B63E158453FFB36F83BFAC4F83C" author="MacLeay WS" box="[1213,1399,1958,1985]" pageId="13" pageNumber="75" refId="ref11281" refString="MacLeay WS. 1819. Horae entomologicae: or, Essays on the Annulose Animals. R. and A. Taylor, London. https: // doi. org / 10.5962 / bhl. title. 65809" type="book" year="1819">MacLeay, 1819</bibRefCitation>
|
||
</taxonomicName>
|
||
(ZMHB).
|
||
<emphasis id="B90EEA80E158453FFEBFF855FED7F81F" bold="true" box="[308,356,1993,2019]" pageId="13" pageNumber="75">d–f.</emphasis>
|
||
<taxonomicName id="4C7A4D11E158453FFEE0F855FCA7F818" authority="Ohaus, 1912" authorityName="Ohaus" authorityYear="1912" box="[363,788,1993,2020]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="75" phylum="Arthropoda" rank="species" species="bahianus">
|
||
<emphasis id="B90EEA80E158453FFEE0F855FDC5F81F" box="[363,630,1993,2019]" italics="true" pageId="13" pageNumber="75">Oplognathus bahianus</emphasis>
|
||
<bibRefCitation id="EFEB4B63E158453FFDF6F855FCA7F818" author="Ohaus F." box="[637,788,1993,2020]" pageId="13" pageNumber="75" pagination="650" refId="ref11341" refString="Ohaus F. 1912. Hoplognathus bahianus n. sp. (Col. Rutel.). Deutsche Entomologische Zeitschrift 1912: 650." type="journal article" year="1912">Ohaus, 1912</bibRefCitation>
|
||
</taxonomicName>
|
||
, syntype, ♂ (ZMHB). Scale bar = 1 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C3606519E15B453CFF36FE0DFDA3FB66" pageId="14" pageNumber="76" type="discussion">
|
||
<paragraph id="8BC53692E15B453CFF36FE0DFE9CFE57" blockId="14.[189,303,401,427]" box="[189,303,401,427]" pageId="14" pageNumber="76">
|
||
<heading id="D08D81FEE15B453CFF36FE0DFE9CFE57" bold="true" box="[189,303,401,427]" fontSize="11" level="3" pageId="14" pageNumber="76" reason="3">
|
||
<emphasis id="B90EEA80E15B453CFF36FE0DFE9CFE57" bold="true" box="[189,303,401,427]" pageId="14" pageNumber="76">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC53692E15B453CFF36FE5CFDA3FB66" blockId="14.[188,1401,447,1178]" pageId="14" pageNumber="76">
|
||
We consider the type of
|
||
<taxonomicName id="4C7A4D11E15B453CFE78FE23FC87FE25" authorityName="Ohaus" authorityYear="1905" box="[499,820,447,473]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="76" phylum="Arthropoda" rank="species" species="helmenreichi">
|
||
<emphasis id="B90EEA80E15B453CFE78FE23FC87FE25" box="[499,820,447,473]" italics="true" pageId="14" pageNumber="76">Oplognathus helmenreichi</emphasis>
|
||
</taxonomicName>
|
||
a
|
||
<typeStatus id="54C18830E15B453CFCD3FE23FC72FE25" box="[856,961,447,473]" pageId="14" pageNumber="76" type="holotype">holotype</typeStatus>
|
||
since
|
||
<bibRefCitation id="EFEB4B63E15B453CFB9DFE23FB0EFE26" author="Ohaus F." box="[1046,1213,447,474]" pageId="14" pageNumber="76" pagination="283 - 329" refId="ref11320" refString="Ohaus F. 1905. Beitrage zur Kenntniss der amerikanischen Ruteliden. Stettiner Entomologische Zeitung 66: 283 - 329." type="journal article" year="1905">Ohaus (1905)</bibRefCitation>
|
||
mentions only a single specimen. For
|
||
<taxonomicName id="4C7A4D11E15B453CFE61FE7EFB7EFE01" authority="Ohaus (1914 a)" authorityName="Ohaus" authorityYear="1914" box="[490,1229,482,509]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="76" phylum="Arthropoda" rank="variety" species="helmreicheni" variety="maculicollis">
|
||
<emphasis id="B90EEA80E15B453CFE61FE7EFC9EFE00" box="[490,813,482,508]" italics="true" pageId="14" pageNumber="76">Oplognathus helmreicheni</emphasis>
|
||
var.
|
||
<emphasis id="B90EEA80E15B453CFCFEFE7EFBBBFE00" box="[885,1032,482,508]" italics="true" pageId="14" pageNumber="76">maculicollis</emphasis>
|
||
<bibRefCitation id="EFEB4B63E15B453CFB9EFE7EFB7EFE01" author="Ohaus F." box="[1045,1229,482,509]" pageId="14" pageNumber="76" pagination="299 - 304" refId="ref11366" refString="Ohaus F. 1914 a. Neue Coleoptera lamellicornia aus Argentinien. Deutsche Entomologische Zeitschrift 1914: 299 - 304." type="journal article" year="1914">Ohaus (1914a)</bibRefCitation>
|
||
</taxonomicName>
|
||
had multiple specimens available. ICZN (1999, Article 45.6.1.) states that a name “is infrasubspecific if its author expressly gave it infrasubspecific rank, or if the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity”. Furthermore, the Article 45.6.4. (ICZN 1999) clarifies that a name “is subspecific if first published before 1961 and its author expressly used one of the terms “variety” or “form” (including use of the terms “var.”, “forma”, “v.” and “f.”), unless its author also expressly gave it infrasubspecific rank, or the content of the work unambiguously reveals that the name was proposed for an infrasubspecific entity, in which case it is infrasubspecific”. In the time Ohaus described
|
||
<taxonomicName id="4C7A4D11E15B453CFD8BFD60FBA6FCEA" authorityName="Ohaus" authorityYear="1914" box="[512,1045,764,791]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="76" phylum="Arthropoda" rank="variety" species="helmenreichi" variety="maculicollis">
|
||
<emphasis id="B90EEA80E15B453CFD8BFD60FCF3FCEA" box="[512,832,764,790]" italics="true" pageId="14" pageNumber="76">Oplognathus helmenreichi</emphasis>
|
||
var.
|
||
<emphasis id="B90EEA80E15B453CFC09FD60FBA6FCEA" box="[898,1045,764,790]" italics="true" pageId="14" pageNumber="76">maculicollis</emphasis>
|
||
</taxonomicName>
|
||
he already clearly separated between species, subspecies and varieties as an article from the same year (see
|
||
<bibRefCitation id="EFEB4B63E15B453CFBF0FC83FAA8FCC6" author="Ohaus F." box="[1147,1307,799,826]" pageId="14" pageNumber="76" pagination="138 - 156" refId="ref11387" refString="Ohaus F. 1914 b. XIV. Beitrag zur Kenntnis der Ruteliden. Stettiner Entomologische Zeitung 75: 138 - 156." type="journal article" year="1914">Ohaus 1914b</bibRefCitation>
|
||
) shows where he described new taxa on the species, subspecies and infrasubspecific levels. Therefore,
|
||
<taxonomicName id="4C7A4D11E15B453CFF36FCFAFD7AFC7C" authorityName="Ohaus" authorityYear="1914" box="[189,713,870,896]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="76" phylum="Arthropoda" rank="variety" species="helmenreichi" variety="maculicollis">
|
||
<emphasis id="B90EEA80E15B453CFF36FCFAFE48FC7C" box="[189,507,870,896]" italics="true" pageId="14" pageNumber="76">Oplognathus helmenreichi</emphasis>
|
||
var.
|
||
<emphasis id="B90EEA80E15B453CFDBDFCFAFD7AFC7C" box="[566,713,870,896]" italics="true" pageId="14" pageNumber="76">maculicollis</emphasis>
|
||
</taxonomicName>
|
||
is an unavailable name. Although the pair of
|
||
<typeStatus id="54C18830E15B453CFB66FCFAFAE5FC7C" box="[1261,1366,870,896]" pageId="14" pageNumber="76" type="syntype">syntypes</typeStatus>
|
||
of
|
||
<taxonomicName id="4C7A4D11E15B453CFF36FC15FDFFFC5F" authorityName="Ohaus" authorityYear="1914" box="[189,588,905,931]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="76" phylum="Arthropoda" rank="variety" species="helmenreichi" variety="maculicollis">
|
||
<emphasis id="B90EEA80E15B453CFF36FC15FECEFC5F" box="[189,381,905,931]" italics="true" pageId="14" pageNumber="76">O. helmenreichi</emphasis>
|
||
var.
|
||
<emphasis id="B90EEA80E15B453CFE32FC15FDFFFC5F" box="[441,588,905,931]" italics="true" pageId="14" pageNumber="76">maculicollis</emphasis>
|
||
</taxonomicName>
|
||
examined have some distinct characteristics of the
|
||
<typeStatus id="54C18830E15B453CFB33FC15FA92FC5F" box="[1208,1313,905,931]" pageId="14" pageNumber="76" type="holotype">holotype</typeStatus>
|
||
– such as yellowish colour, smaller size, less dense punctures and the male genitalia almost parallel sided, the other specimens examined show many similarities both with the
|
||
<typeStatus id="54C18830E15B453CFC71FC4CFBD0FC16" box="[1018,1123,976,1002]" pageId="14" pageNumber="76" type="syntype">syntypes</typeStatus>
|
||
and with the
|
||
<typeStatus id="54C18830E15B453CFA83FC53FAC6FC15" box="[1288,1397,975,1001]" pageId="14" pageNumber="76" type="holotype">holotype</typeStatus>
|
||
. The variation of these characteristics was also observed within the other species of the genus. Despite these variations, mouthpart characters (like mentum shape, maxillary palpus shape) and mesoventral process remained constant in the
|
||
<specimenCount id="9D7CFD1BE15B453CFDD6FBA5FC93FBAF" box="[605,800,1081,1107]" pageId="14" pageNumber="76" type="generic">eight specimens</specimenCount>
|
||
examined. Thus,
|
||
<taxonomicName id="4C7A4D11E15B453CFC74FBA5FEE3FB8A" authorityName="Ohaus" authorityYear="1914" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="76" phylum="Arthropoda" rank="variety" species="helmenreichi" variety="maculicollis">
|
||
<emphasis id="B90EEA80E15B453CFC74FBA5FA8CFBAF" box="[1023,1343,1081,1107]" italics="true" pageId="14" pageNumber="76">Oplognathus helmenreichi</emphasis>
|
||
var.
|
||
<emphasis id="B90EEA80E15B453CFF36FBC0FEE3FB8A" box="[189,336,1116,1142]" italics="true" pageId="14" pageNumber="76">maculicollis</emphasis>
|
||
</taxonomicName>
|
||
does not present a unique combination of character states that justify validating it to species or subspecies status.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3606519E15B453CFF36FB5CFBDFF998" pageId="14" pageNumber="76" type="distribution">
|
||
<paragraph id="8BC53692E15B453CFF36FB5CFEE5FB26" blockId="14.[189,342,1216,1242]" box="[189,342,1216,1242]" pageId="14" pageNumber="76">
|
||
<heading id="D08D81FEE15B453CFF36FB5CFEE5FB26" bold="true" box="[189,342,1216,1242]" fontSize="11" level="3" pageId="14" pageNumber="76" reason="3">
|
||
<emphasis id="B90EEA80E15B453CFF36FB5CFEE5FB26" bold="true" box="[189,342,1216,1242]" pageId="14" pageNumber="76">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BC53692E15B453CFF36FB73FBEAFAF5" blockId="14.[189,1113,1263,1290]" box="[189,1113,1263,1290]" pageId="14" pageNumber="76">
|
||
<emphasis id="B90EEA80E15B453CFF36FB73FEB9FAF5" bold="true" box="[189,266,1263,1289]" pageId="14" pageNumber="76">
|
||
<collectingCountry id="F36D7602E15B453CFF36FB73FEB9FAF5" box="[189,266,1263,1289]" name="Brazil" pageId="14" pageNumber="76">Brazil</collectingCountry>
|
||
</emphasis>
|
||
:
|
||
<collectingRegion id="49BEF870E15B453CFE92FB73FE04FAF5" box="[281,439,1263,1289]" country="Brazil" name="Minas Gerais" pageId="14" pageNumber="76">Minas Gerais</collectingRegion>
|
||
: Grão Mogol, Leme do Prado.
|
||
<collectingRegion id="49BEF870E15B453CFCADFB73FCC5FAF5" box="[806,886,1263,1289]" country="Brazil" name="Parana" pageId="14" pageNumber="76">Paraná</collectingRegion>
|
||
: Curitiba? (Fig. 9).
|
||
</paragraph>
|
||
<paragraph id="8BC53692E15B453CFF36FAADFBDFF998" blockId="14.[189,1400,1329,1637]" pageId="14" pageNumber="76">
|
||
No other specimen of
|
||
<taxonomicName id="4C7A4D11E15B453CFE47FAADFD39FAB7" authorityName="Ohaus" authorityYear="1905" box="[460,650,1329,1355]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="76" phylum="Arthropoda" rank="species" species="helmenreichi">
|
||
<emphasis id="B90EEA80E15B453CFE47FAADFD39FAB7" box="[460,650,1329,1355]" italics="true" pageId="14" pageNumber="76">O. helmenreichi</emphasis>
|
||
</taxonomicName>
|
||
was subsequently collected in
|
||
<collectingRegion id="49BEF870E15B453CFC76FAADFB2DFAB7" box="[1021,1182,1329,1355]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="14" pageNumber="76">Buenos Aires</collectingRegion>
|
||
, or anywhere else in
|
||
<collectingCountry id="F36D7602E15B453CFF57FAC8FEE2FA92" box="[220,337,1364,1390]" name="Argentina" pageId="14" pageNumber="76">Argentina</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFEB4B63E15B453CFEEBFAC8FDB2FA92" author="Ohaus F." box="[352,513,1364,1390]" pageId="14" pageNumber="76" pagination="299 - 304" refId="ref11366" refString="Ohaus F. 1914 a. Neue Coleoptera lamellicornia aus Argentinien. Deutsche Entomologische Zeitschrift 1914: 299 - 304." type="journal article" year="1914">Ohaus 1914a</bibRefCitation>
|
||
). One part of the material of
|
||
<taxonomicName id="4C7A4D11E15B453CFCDCFAC8FB6EFA92" authorityName="Ohaus" authorityYear="1914" box="[855,1245,1364,1390]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="76" phylum="Arthropoda" rank="variety" species="helmenreichi" variety="maculicollis">
|
||
<emphasis id="B90EEA80E15B453CFCDCFAC8FBA0FA92" box="[855,1043,1364,1390]" italics="true" pageId="14" pageNumber="76">O. helmenreichi</emphasis>
|
||
var.
|
||
<emphasis id="B90EEA80E15B453CFBC5FAC8FB6EFA92" box="[1102,1245,1364,1390]" italics="true" pageId="14" pageNumber="76">maculicollis</emphasis>
|
||
</taxonomicName>
|
||
was labelled “Helm., Brasilia, XXII, 1853” and the other part “Serra do Grão Mogor, 1846” [Municipality of Grão Mogol,
|
||
<collectingRegion id="49BEF870E15B453CFE90FA06FE0CFA49" box="[283,447,1434,1461]" country="Brazil" name="Minas Gerais" pageId="14" pageNumber="76">Minas Gerais</collectingRegion>
|
||
].
|
||
<bibRefCitation id="EFEB4B63E15B453CFE59FA06FDCCFA48" author="Ohaus F." box="[466,639,1434,1460]" pageId="14" pageNumber="76" pagination="299 - 304" refId="ref11366" refString="Ohaus F. 1914 a. Neue Coleoptera lamellicornia aus Argentinien. Deutsche Entomologische Zeitschrift 1914: 299 - 304." type="journal article" year="1914">Ohaus (1914a)</bibRefCitation>
|
||
then made it clear that the most probable locality of this species would be the north of
|
||
<collectingRegion id="49BEF870E15B453CFE45FA21FD00FA24" box="[462,691,1469,1496]" country="Brazil" name="Minas Gerais" pageId="14" pageNumber="76">Minas Gerais State</collectingRegion>
|
||
(Grão Mogol) and not
|
||
<collectingRegion id="49BEF870E15B453CFC5BFA22FBC1FA2B" box="[976,1138,1469,1496]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="14" pageNumber="76">Buenos Aires</collectingRegion>
|
||
, corresponding to the known distribution of the genus. Thus, we agree with
|
||
<bibRefCitation id="EFEB4B63E15B453CFCC5FA7DFC4DFA07" author="Ohaus F." box="[846,1022,1505,1531]" pageId="14" pageNumber="76" pagination="299 - 304" refId="ref11366" refString="Ohaus F. 1914 a. Neue Coleoptera lamellicornia aus Argentinien. Deutsche Entomologische Zeitschrift 1914: 299 - 304." type="journal article" year="1914">Ohaus (1914a)</bibRefCitation>
|
||
and disregard
|
||
<collectingRegion id="49BEF870E15B453CFB3FFA7DFAE7FA07" box="[1204,1364,1505,1531]" country="Argentina" name="Ciudad Autonoma de Buenos Aires" pageId="14" pageNumber="76">Buenos Aires</collectingRegion>
|
||
as a locality for this species, as well as for the genus
|
||
<emphasis id="B90EEA80E15B453CFCAFF998FC77F9E2" box="[804,964,1540,1566]" italics="true" pageId="14" pageNumber="76">
|
||
<taxonomicName id="4C7A4D11E15B453CFCAFF998FC0CF9E2" authorityName="MacLeay" authorityYear="1819" box="[804,959,1540,1566]" class="Insecta" family="Rutelidae" genus="Oplognathus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="76" phylum="Arthropoda" rank="genus">Oplognathus</taxonomicName>
|
||
.
|
||
</emphasis>
|
||
One male of the examined material is labelled as being from Curitiba,
|
||
<collectingRegion id="49BEF870E15B453CFDD0F9BBFD1DF9BD" box="[603,686,1575,1601]" country="Brazil" name="Parana" pageId="14" pageNumber="76">Paraná</collectingRegion>
|
||
, South of
|
||
<collectingCountry id="F36D7602E15B453CFCA2F9BBFCC7F9BD" box="[809,884,1575,1601]" name="Brazil" pageId="14" pageNumber="76">Brazil</collectingCountry>
|
||
, but the origin is doubtful, since this single specimen was found in a didactic collection of the Federal University of
|
||
<collectingRegion id="49BEF870E15B453CFB9FF9D6FBDBF998" box="[1044,1128,1610,1636]" country="Brazil" name="Parana" pageId="14" pageNumber="76">Paraná</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |