277 lines
51 KiB
XML
277 lines
51 KiB
XML
<document id="29CF365494CEB82F042E6C3A54448D3F" ID-CLB-Dataset="31254" ID-DOI="10.11646/zootaxa.4362.1.3" ID-GBIF-Dataset="e7983c67-0ce5-4b7f-9373-8782b86e059b" ID-ISSN="1175-5326" ID-Zenodo-Dep="1076176" ID-ZooBank="61A41EC1-2557-429D-B47C-595D12F2D8E3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1512393169915" checkinUser="plazi" docAuthor="Polotow, Daniele & Griswold, Charles" docDate="2017" docId="03D38783FF89FF88E6B36B10FD05601C" docLanguage="en" docName="zootaxa.4362.1.3.pdf" docOrigin="Zootaxa 4362 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Devendra Lehtinen 1967" docType="treatment" docVersion="8" lastPageNumber="65" masterDocId="FFEAFFFBFF84FF86E6246E6DFFC96633" masterDocTitle="Cleaning old cabinets: revealing the taxonomy of Sri Lankan wolf spiders (Araneae, Udubidae and Zoropsidae)" masterLastPageNumber="74" masterPageNumber="51" pageNumber="64" updateTime="1698506327670" updateUser="ExternalLinkService">
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<mods:title id="CF0F47F9885BB55F1C47911FB58EC241">Cleaning old cabinets: revealing the taxonomy of Sri Lankan wolf spiders (Araneae, Udubidae and Zoropsidae)</mods:title>
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<mods:namePart id="16005714B077BC21D550B0583162CABE">Polotow, Daniele</mods:namePart>
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<mods:namePart id="6FFEF55A3441FB9C395774F00E5DD76E">Griswold, Charles</mods:namePart>
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<mods:title id="219C32DC7025908B13F21E5020B2235B">Zootaxa</mods:title>
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<mods:date id="AF9FDF8842EA34FA8B63CA07AD233798">2017</mods:date>
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<treatment id="03D38783FF89FF88E6B36B10FD05601C" ID-DOI="http://doi.org/10.5281/zenodo.5999802" ID-GBIF-Taxon="137584765" ID-Zenodo-Dep="5999802" LSID="urn:lsid:plazi:treatment:03D38783FF89FF88E6B36B10FD05601C" httpUri="http://treatment.plazi.org/id/03D38783FF89FF88E6B36B10FD05601C" lastPageId="14" lastPageNumber="65" pageId="13" pageNumber="64">
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<subSubSection id="C360651EFF89FF8BE6B36B10FE0463A4" box="[151,461,1405,1431]" pageId="13" pageNumber="64" type="nomenclature">
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<paragraph id="8BC53695FF89FF8BE6B36B10FE0463A4" blockId="13.[151,461,1405,1431]" box="[151,461,1405,1431]" pageId="13" pageNumber="64">
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<heading id="D08D81F9FF89FF8BE6B36B10FE0463A4" bold="true" box="[151,461,1405,1431]" fontSize="11" level="1" pageId="13" pageNumber="64" reason="1">
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<taxonomicName id="4C7A4D16FF89FF8BE6B36B10FE0463A4" ID-CoL="43VW" authority="Lehtinen, 1967" authorityName="Lehtinen" authorityYear="1967" box="[151,461,1405,1431]" class="Arachnida" family="Zoropsidae" genus="Devendra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="64" phylum="Arthropoda" rank="genus">
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<emphasis id="B90EEA87FF89FF8BE6B36B10FE0463A4" bold="true" box="[151,461,1405,1431]" pageId="13" pageNumber="64">
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<emphasis id="B90EEA87FF89FF8BE6B36B10FEC463A4" bold="true" box="[151,269,1405,1431]" italics="true" pageId="13" pageNumber="64">Devendra</emphasis>
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<bibRefCitation id="EFEB4B64FF89FF8BE7376B10FE0463A4" author="Lehtinen" box="[275,461,1405,1431]" pageId="13" pageNumber="64" refString="LEhtInEn, P. T. (1967) ClAssIFIcAtIOn OF thE crIbEllAtE spIdErs And sOmE AllIEd FAmIlIEs, wIth nOtEs On thE EVOlutIOn OF thE subOrdEr ArAnEOmOrphA. Annales Zoologici Fennici, 4, 199 - 468." type="journal article" year="1967">Lehtinen, 1967</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C360651EFF89FF8BE6B36BA9FB1D6052" pageId="13" pageNumber="64" type="reference_group">
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<paragraph id="8BC53695FF89FF8BE6B36BA9FBCC63CB" blockId="13.[151,1437,1476,1528]" pageId="13" pageNumber="64">
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<treatmentCitationGroup id="AB6A11BBFF89FF8BE6B36BA9FBCC63CB" pageId="13" pageNumber="64">
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<treatmentCitation id="0ADB1084FF89FF8BE6B36BA9FE0563E9" author="LEhtInEn" box="[151,460,1476,1498]" page="228" pageId="13" pageNumber="64" year="1967">
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<taxonomicName id="4C7A4D16FF89FF8BE6B36BA9FE5763E9" ID-CoL="43VW" authority="LEhtInEn, 1967" authorityName="Lehtinen" authorityYear="1967" box="[151,414,1476,1498]" class="Arachnida" family="Zoropsidae" genus="Devendra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="64" phylum="Arthropoda" rank="genus">
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<emphasis id="B90EEA87FF89FF8BE6B36BA9FF3263E9" box="[151,251,1476,1498]" italics="true" pageId="13" pageNumber="64">Devendra</emphasis>
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<bibRefCitation id="EFEB4B64FF89FF8BE7256BA9FE5763E9" author="LEhtInEn" box="[257,414,1476,1498]" pageId="13" pageNumber="64" refString="LEhtInEn, P. T. (1967) ClAssIFIcAtIOn OF thE crIbEllAtE spIdErs And sOmE AllIEd FAmIlIEs, wIth nOtEs On thE EVOlutIOn OF thE subOrdEr ArAnEOmOrphA. Annales Zoologici Fennici, 4, 199 - 468." type="journal article" year="1967">LEhtInEn, 1967</bibRefCitation>
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</taxonomicName>
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: 228
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</treatmentCitation>
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(TypE spEcIEs by OrIgInAl dEsIgnAtIOn:
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<taxonomicName id="4C7A4D16FF89FF8BE57C6BA9FBAB63E9" box="[856,1122,1476,1498]" class="Arachnida" family="Zorocratidae" genus="Campostichomma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="64" phylum="Arthropoda" rank="species" species="pardale">
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<emphasis id="B90EEA87FF89FF8BE57C6BA9FBAB63E9" box="[856,1122,1476,1498]" italics="true" pageId="13" pageNumber="64">Campostichomma pardale</emphasis>
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</taxonomicName>
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SImOn, 1898).
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<treatmentCitation id="0ADB1084FF89FF8BE2DA6BA9FF1F63CB" author="GrIswOld" page="7" pageId="13" pageNumber="64" year="1993">
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<bibRefCitation id="EFEB4B64FF89FF8BE2DA6BA9FA5163E9" author="GrIswOld" box="[1278,1432,1476,1498]" pageId="13" pageNumber="64" refString="GrIswOld, C. E. (1993) InVEstIgAtIOns IntO thE phylOgEny OF thE lycOsOId spIdErs And thEIr kIn (ArAchnIdA: ArAnEAE: LycOsOIdEA). Smithsonian Contributions to Zoology, 539, 1 - 39. https: // dOI. Org / 10.5479 / sI. 00810282.539" type="journal article" year="1993">GrIswOld 1993</bibRefCitation>
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: 7
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</treatmentCitation>
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, FIgs 37–42;
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<treatmentCitation id="0ADB1084FF89FF8BE77D6B8EFDB463CB" author="RAVEn" box="[345,637,1506,1528]" page="356" pageId="13" pageNumber="64" year="2005">
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<bibRefCitation id="EFEB4B64FF89FF8BE77D6B8EFD8263CB" author="RAVEn" box="[345,587,1506,1528]" pageId="13" pageNumber="64" refString="RAVEn, R. J. & StumkAt, K. S. (2005) REVIsIOns OF AustrAlIAn grOund-huntIng spIdErs: II. ZOrOpsIdAE (LycOsOIdEA: ArAnEAE). Memoirs of the Queensland Museum, 50, 347 - 423." type="journal article" year="2005">RAVEn & StumkAt 2005</bibRefCitation>
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: 356
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||
</treatmentCitation>
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.
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<treatmentCitation id="0ADB1084FF89FF8BE4A36B8FFC4D63CB" author="POlOtOw" box="[647,900,1506,1528]" page="152" pageId="13" pageNumber="64" year="2015">
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<bibRefCitation id="EFEB4B64FF89FF8BE4A36B8FFC9863CB" author="POlOtOw" box="[647,849,1506,1528]" pageId="13" pageNumber="64" refString="POlOtOw, D., CArmIchEAl, A. & GrIswOld, C. E. (2015) TOtAl EVIdEncE AnAlysIs OF thE phylOgEnEtIc rElAtIOnshIps OF LycOsOIdEA spIdErs (ArAnEAE, EntElEgynAE). Invertebrate Systematics, 29, 124 - 163. https: // dOI. Org / 10.1071 / IS 14041" type="journal article" year="2015">
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POlOtOw
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<emphasis id="B90EEA87FF89FF8BE4C66B8EFCDC63CB" box="[738,789,1506,1528]" italics="true" pageId="13" pageNumber="64">et al.</emphasis>
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2015
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</bibRefCitation>
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: 152
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||
</treatmentCitation>
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; WSC 2017.
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||
</treatmentCitationGroup>
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</paragraph>
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<paragraph id="8BC53695FF89FF8BE6B36848FB1D6052" blockId="13.[151,1437,1573,2030]" pageId="13" pageNumber="64">
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||
<emphasis id="B90EEA87FF89FF8BE6B36848FF06600D" bold="true" box="[151,207,1573,1598]" pageId="13" pageNumber="64">Note</emphasis>
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. The genus name
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<taxonomicName id="4C7A4D16FF89FF8BE7806848FDDB600E" ID-CoL="43VW" authorityName="Lehtinen" authorityYear="1967" box="[420,530,1573,1597]" class="Arachnida" family="Zoropsidae" genus="Devendra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="64" phylum="Arthropoda" rank="genus">
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<emphasis id="B90EEA87FF89FF8BE7806848FDDB600E" box="[420,530,1573,1597]" italics="true" pageId="13" pageNumber="64">Devendra</emphasis>
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</taxonomicName>
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is masculine in the source language, Sanskrit, and a masculine a-stem noun in Latin (H. D. Cameron, in litt; WSC 2017), hence
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<taxonomicName id="4C7A4D16FF89FF8BE4E56824FC5A6052" ID-CoL="355V6" baseAuthorityName="Simon" baseAuthorityYear="1898" box="[705,915,1609,1633]" class="Arachnida" family="Zoropsidae" genus="Devendra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="64" phylum="Arthropoda" rank="species" species="pardalis">
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<emphasis id="B90EEA87FF89FF8BE4E56824FC5A6052" box="[705,915,1609,1633]" italics="true" pageId="13" pageNumber="64">Devendra pardalis</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C7A4D16FF89FF8BE5856827FBD56052" ID-CoL="355V7" baseAuthorityName="Simon" baseAuthorityYear="1898" box="[929,1052,1609,1633]" class="Arachnida" family="Zoropsidae" genus="Devendra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="64" phylum="Arthropoda" rank="species" species="pumilus">
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<emphasis id="B90EEA87FF89FF8BE5856827FBD56052" box="[929,1052,1609,1633]" italics="true" pageId="13" pageNumber="64">D. pumilus</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C7A4D16FF89FF8BE2766827FB076052" ID-CoL="355V9" baseAuthorityName="Simon" baseAuthorityYear="1898" box="[1106,1230,1610,1633]" class="Arachnida" family="Zoropsidae" genus="Devendra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="64" phylum="Arthropoda" rank="species" species="seriatus">
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<emphasis id="B90EEA87FF89FF8BE2766827FB076052" box="[1106,1230,1610,1633]" italics="true" pageId="13" pageNumber="64">D. seriatus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C360651EFF89FF88E6E36800FE5362DB" lastPageId="14" lastPageNumber="65" pageId="13" pageNumber="64" type="diagnosis">
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<paragraph id="8BC53695FF89FF8BE6E36800FD5261F9" blockId="13.[151,1437,1573,2030]" pageId="13" pageNumber="64">
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<emphasis id="B90EEA87FF89FF8BE6E36800FE8860B5" bold="true" box="[199,321,1645,1670]" pageId="13" pageNumber="64">Diagnosis.</emphasis>
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<taxonomicName id="4C7A4D16FF89FF8BE76D6800FE7E60B6" authorityName="Lehtinen" authorityYear="1967" box="[329,439,1645,1669]" class="Arachnida" family="Zoropsidae" genus="Devendra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="64" phylum="Arthropoda" rank="genus">
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<emphasis id="B90EEA87FF89FF8BE76D6800FE7E60B6" box="[329,439,1645,1669]" italics="true" pageId="13" pageNumber="64">Devendra</emphasis>
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</taxonomicName>
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are cribellate, RTA clade spiders with the PER usually straight, no claw tufts, leg tibiae of males with a visible, subcuticular transverse basal suture or crack (like in
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<figureCitation id="13412A10FF89FF8BE22668FCFB936099" box="[1026,1114,1681,1706]" captionStart="FIGURE 3" captionStartId="7.[151,250,1732,1754]" captionTargetBox="[151,1436,193,1711]" captionTargetId="figure@7.[151,1436,193,1711]" captionTargetPageId="7" captionText="FIGURE 3. Campostichomma manicatum KArsch, 1892. (A–D) mAlE lEFt pAlp; (A) VEntrAl VIEw, (B) rEtrOlAtErAl VIEw, (C) prOlAtErAl VIEw; (D) rIght pAlp, rEtrOlAtErAl VIEw, ExpAndEd; (E) mAlE lEg IV; (F–G) EpIgynum; (F) VEntrAl VIEw; (G) dOrsAl VIEw.AbbrEVIAtIOns:LO, lOckIng lObEs;TC, tIbIAl crAck." httpUri="https://zenodo.org/record/1076188/files/figure.png" pageId="13" pageNumber="64">Fig. 3E</figureCitation>
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). Males of
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<taxonomicName id="4C7A4D16FF89FF8BE2FF68FCFA83609A" authorityName="Lehtinen" authorityYear="1967" box="[1243,1354,1681,1705]" class="Arachnida" family="Zoropsidae" genus="Devendra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="64" phylum="Arthropoda" rank="genus">
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<emphasis id="B90EEA87FF89FF8BE2FF68FCFA83609A" box="[1243,1354,1681,1705]" italics="true" pageId="13" pageNumber="64">Devendra</emphasis>
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</taxonomicName>
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can be distinguished from other zoropsid genera by characters of the male palp, as the presence of a S-shaped median apophysis (
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<figureCitation id="13412A10FF89FF8BE73368B4FE4B60C1" box="[279,386,1753,1778]" captionStart="FIGURE 11" captionStartId="16.[151,250,1906,1928]" captionTargetBox="[163,1412,935,1884]" captionTargetId="figure@16.[151,1436,911,1884]" captionTargetPageId="16" captionText="FIGURE 11. Devendra pumilus (SImOn, 1898), mAlE. (A–C) lEFt pAlp; (A) VEntrAl VIEw; (B) rEtrOlAtErAl VIEw; (C) prOlAtErAl VIEw.AbbrEVIAtIOns: C, cOnductOr; E, EmbOlus; MA, mEdIAn ApOphysIs; RTA, rEtrOlAtErAl tIbIAl ApOphysIs; ST, subtEgulum; T, tEgulum; TI, tIbIA." httpUri="https://zenodo.org/record/1076216/files/figure.png" pageId="13" pageNumber="64">Figs 11A</figureCitation>
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,
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<figureCitation id="13412A10FF89FF8BE7AB68B7FE0A60C1" box="[399,451,1754,1778]" captionStart="FIGURE 12" captionStartId="18.[151,250,1893,1915]" captionTargetBox="[168,1414,388,1849]" captionTargetId="figure@18.[151,1436,375,1871]" captionTargetPageId="18" captionText="FIGURE 12. Devendra seriatus (SImOn, 1898). (A–C) lEFt mAlE pAlp; (A) VEntrAl VIEw; (B) rEtrOlAtErAl VIEw; (C) prOlAtErAl VIEw; (D–F) FEmAlE EpIgynum; (D) VEntrAl VIEw; (E) dOrsAl VIEw; (F) rIght spErmAthEcA,dOrsAl, clEArEd, dEtAIl.AbbrEVIAtIOns: C, cOnductOr; CD, cOpulAtOry ducts; E, EmbOlus; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; LP, lAtErAl prOcEss OF EpIgynum; MA, mEdIAn ApOphysIs; ML, mEdIAn lObE OF EpIgynum; RTA, rEtrOlAtErAl tIbIAl ApOphysIs; S, spErmAthEcAE; T, tEgulum; TI, tIbIA." httpUri="https://zenodo.org/record/1076218/files/figure.png" pageId="13" pageNumber="64">12A</figureCitation>
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), by the presence of a ventral tibial apophysis and the RTA positioned distant from the tip of tibia (
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<figureCitation id="13412A10FF89FF8BE7396890FEB16125" box="[285,376,1789,1814]" captionStart="FIGURE 9" captionStartId="15.[151,250,1972,1994]" captionTargetBox="[151,1436,663,1949]" captionTargetId="figure@15.[151,1436,663,1949]" captionTargetPageId="15" captionText="FIGURE 9. Devendra spp. (A–C) D. pardalis (SImOn, 1898), FEmAlE; (A) hAbItus, dOrsAl VIEw; (B) AntErIOr VIEw OF OculAr ArEA;(C)EpIgynum, VEntrAl VIEw;(D–E) D. pumilus (SImOn, 1898), mAlE;(D)hAbItus,dOrsAl VIEw;(E)rIght pAlp,VEntrAl VIEw." httpUri="https://zenodo.org/record/1076208/files/figure.png" pageId="13" pageNumber="64">Figs 9E</figureCitation>
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,
|
||
<figureCitation id="13412A10FF89FF8BE7A16893FE106125" box="[389,473,1790,1814]" captionStart="FIGURE 12" captionStartId="18.[151,250,1893,1915]" captionTargetBox="[168,1414,388,1849]" captionTargetId="figure@18.[151,1436,375,1871]" captionTargetPageId="18" captionText="FIGURE 12. Devendra seriatus (SImOn, 1898). (A–C) lEFt mAlE pAlp; (A) VEntrAl VIEw; (B) rEtrOlAtErAl VIEw; (C) prOlAtErAl VIEw; (D–F) FEmAlE EpIgynum; (D) VEntrAl VIEw; (E) dOrsAl VIEw; (F) rIght spErmAthEcA,dOrsAl, clEArEd, dEtAIl.AbbrEVIAtIOns: C, cOnductOr; CD, cOpulAtOry ducts; E, EmbOlus; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; LP, lAtErAl prOcEss OF EpIgynum; MA, mEdIAn ApOphysIs; ML, mEdIAn lObE OF EpIgynum; RTA, rEtrOlAtErAl tIbIAl ApOphysIs; S, spErmAthEcAE; T, tEgulum; TI, tIbIA." httpUri="https://zenodo.org/record/1076218/files/figure.png" pageId="13" pageNumber="64">12A, B</figureCitation>
|
||
); in at least
|
||
<taxonomicName id="4C7A4D16FF89FF8BE4416893FD286126" baseAuthorityName="Simon" baseAuthorityYear="1898" box="[613,737,1789,1813]" class="Arachnida" family="Zoropsidae" genus="Devendra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="13" pageNumber="64" phylum="Arthropoda" rank="species" species="pumilus">
|
||
<emphasis id="B90EEA87FF89FF8BE4416893FDB06126" box="[613,633,1790,1813]" italics="true" pageId="13" pageNumber="64">D</emphasis>
|
||
.
|
||
<emphasis id="B90EEA87FF89FF8BE4AC6890FD286126" box="[648,737,1789,1813]" italics="true" pageId="13" pageNumber="64">pumilus</emphasis>
|
||
</taxonomicName>
|
||
this process is surmounted by a spine (
|
||
<figureCitation id="13412A10FF89FF8BE2846890FB326125" box="[1184,1275,1789,1814]" captionStart="FIGURE 9" captionStartId="15.[151,250,1972,1994]" captionTargetBox="[151,1436,663,1949]" captionTargetId="figure@15.[151,1436,663,1949]" captionTargetPageId="15" captionText="FIGURE 9. Devendra spp. (A–C) D. pardalis (SImOn, 1898), FEmAlE; (A) hAbItus, dOrsAl VIEw; (B) AntErIOr VIEw OF OculAr ArEA;(C)EpIgynum, VEntrAl VIEw;(D–E) D. pumilus (SImOn, 1898), mAlE;(D)hAbItus,dOrsAl VIEw;(E)rIght pAlp,VEntrAl VIEw." httpUri="https://zenodo.org/record/1076208/files/figure.png" pageId="13" pageNumber="64">Figs 9E</figureCitation>
|
||
,
|
||
<figureCitation id="13412A10FF89FF8BE3236893FA926125" box="[1287,1371,1790,1814]" captionStart="FIGURE 11" captionStartId="16.[151,250,1906,1928]" captionTargetBox="[163,1412,935,1884]" captionTargetId="figure@16.[151,1436,911,1884]" captionTargetPageId="16" captionText="FIGURE 11. Devendra pumilus (SImOn, 1898), mAlE. (A–C) lEFt pAlp; (A) VEntrAl VIEw; (B) rEtrOlAtErAl VIEw; (C) prOlAtErAl VIEw.AbbrEVIAtIOns: C, cOnductOr; E, EmbOlus; MA, mEdIAn ApOphysIs; RTA, rEtrOlAtErAl tIbIAl ApOphysIs; ST, subtEgulum; T, tEgulum; TI, tIbIA." httpUri="https://zenodo.org/record/1076216/files/figure.png" pageId="13" pageNumber="64">11A, B</figureCitation>
|
||
). The females have a simple epigynum with large teeth on the lateral lobes (
|
||
<figureCitation id="13412A10FF89FF8BE5E3694CFBEC6109" box="[967,1061,1825,1850]" captionStart="FIGURE 9" captionStartId="15.[151,250,1972,1994]" captionTargetBox="[151,1436,663,1949]" captionTargetId="figure@15.[151,1436,663,1949]" captionTargetPageId="15" captionText="FIGURE 9. Devendra spp. (A–C) D. pardalis (SImOn, 1898), FEmAlE; (A) hAbItus, dOrsAl VIEw; (B) AntErIOr VIEw OF OculAr ArEA;(C)EpIgynum, VEntrAl VIEw;(D–E) D. pumilus (SImOn, 1898), mAlE;(D)hAbItus,dOrsAl VIEw;(E)rIght pAlp,VEntrAl VIEw." httpUri="https://zenodo.org/record/1076208/files/figure.png" pageId="13" pageNumber="64">Figs 9C</figureCitation>
|
||
,
|
||
<figureCitation id="13412A10FF89FF8BE210694FFBAE6109" box="[1076,1127,1826,1850]" captionStart="FIGURE 10" captionStartId="16.[151,250,641,663]" captionTargetBox="[163,1413,196,618]" captionTargetId="figure@16.[151,1436,193,620]" captionTargetPageId="16" captionText="FIGURE 10. Devendra pardalis (SImOn, 1898), FEmAlE. (A–C) EpIgynum; (A) VEntrAl VIEw; (B) dOrsAl VIEw; (C) rIght spErmAthEcA, dOrsAl, clEArEd, dEtAIl.AbbrEVIAtIOns: CD, cOpulAtOry ducts; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; LP, lAtErAl prOcEss OF EpIgynum; ML, mEdIAnlObE OF EpIgynum; S, spErmAthEcAE." httpUri="https://zenodo.org/record/1076212/files/figure.png" pageId="13" pageNumber="64">10A</figureCitation>
|
||
,
|
||
<figureCitation id="13412A10FF89FF8BE251694FFB616109" box="[1141,1192,1826,1850]" captionStart="FIGURE 12" captionStartId="18.[151,250,1893,1915]" captionTargetBox="[168,1414,388,1849]" captionTargetId="figure@18.[151,1436,375,1871]" captionTargetPageId="18" captionText="FIGURE 12. Devendra seriatus (SImOn, 1898). (A–C) lEFt mAlE pAlp; (A) VEntrAl VIEw; (B) rEtrOlAtErAl VIEw; (C) prOlAtErAl VIEw; (D–F) FEmAlE EpIgynum; (D) VEntrAl VIEw; (E) dOrsAl VIEw; (F) rIght spErmAthEcA,dOrsAl, clEArEd, dEtAIl.AbbrEVIAtIOns: C, cOnductOr; CD, cOpulAtOry ducts; E, EmbOlus; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; LP, lAtErAl prOcEss OF EpIgynum; MA, mEdIAn ApOphysIs; ML, mEdIAn lObE OF EpIgynum; RTA, rEtrOlAtErAl tIbIAl ApOphysIs; S, spErmAthEcAE; T, tEgulum; TI, tIbIA." httpUri="https://zenodo.org/record/1076218/files/figure.png" pageId="13" pageNumber="64">12D</figureCitation>
|
||
,
|
||
<figureCitation id="13412A10FF89FF8BE292694FFAC56109" box="[1206,1292,1826,1850]" captionStart="FIGURE 13" captionStartId="19.[151,250,1685,1707]" captionTargetBox="[159,1414,872,1660]" captionTargetId="figure@19.[151,1436,842,1674]" captionTargetPageId="19" captionText="FIGURE 13. Devendra spp. (A–C) D. saama sp. nov., FEmAlE hOlOtypE (MNHN#18302), EpIgynum; (A) VEntrAl VIEw; (B) dOrsAl VIEw; (C) rIght spErmAthEcA, dOrsAl, clEArEd, dEtAIl; (D–F) D. amaiti sp. nov., FEmAlE hOlOtypE (MNHN#18337), EpIgynum;(D) VEntrAl VIEw;(E)dOrsAlVIEw;(F)rIght spErmAthEcA,dOrsAl,clEArEd,dEtAIl.AbbrEVIAtIOns:CD, cOpulAtOry duct; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; LP, lAtErAl prOcEss OF EpIgynum; ML, mEdIAn lObE OF EpIgynum; S, spErmAthEcAE." httpUri="https://zenodo.org/record/1076222/files/figure.png" pageId="13" pageNumber="64">13A, D</figureCitation>
|
||
) and can be distinguished by the shape of the epigynal structures, as the presence of a smooth and slightly sclerotized epigynal plate, with a small lateral projection. Internally, the vulva is simple with the head of the spermathecae very small, sessile or on a tubercle, the copulatory ducts are short, the spermathecae is large and the fertilization ducts are elongated (
|
||
<figureCitation id="13412A10FF89FF8BE73669DCFE5761F9" box="[274,414,1969,1994]" captionStart="FIGURE 10" captionStartId="16.[151,250,641,663]" captionTargetBox="[163,1413,196,618]" captionTargetId="figure@16.[151,1436,193,620]" captionTargetPageId="16" captionText="FIGURE 10. Devendra pardalis (SImOn, 1898), FEmAlE. (A–C) EpIgynum; (A) VEntrAl VIEw; (B) dOrsAl VIEw; (C) rIght spErmAthEcA, dOrsAl, clEArEd, dEtAIl.AbbrEVIAtIOns: CD, cOpulAtOry ducts; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; LP, lAtErAl prOcEss OF EpIgynum; ML, mEdIAnlObE OF EpIgynum; S, spErmAthEcAE." httpUri="https://zenodo.org/record/1076212/files/figure.png" pageId="13" pageNumber="64">Figs 10B, C</figureCitation>
|
||
.
|
||
<figureCitation id="13412A10FF89FF8BE78D69DFFE3D61FA" box="[425,500,1970,1994]" captionStart="FIGURE 12" captionStartId="18.[151,250,1893,1915]" captionTargetBox="[168,1414,388,1849]" captionTargetId="figure@18.[151,1436,375,1871]" captionTargetPageId="18" captionText="FIGURE 12. Devendra seriatus (SImOn, 1898). (A–C) lEFt mAlE pAlp; (A) VEntrAl VIEw; (B) rEtrOlAtErAl VIEw; (C) prOlAtErAl VIEw; (D–F) FEmAlE EpIgynum; (D) VEntrAl VIEw; (E) dOrsAl VIEw; (F) rIght spErmAthEcA,dOrsAl, clEArEd, dEtAIl.AbbrEVIAtIOns: C, cOnductOr; CD, cOpulAtOry ducts; E, EmbOlus; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; LP, lAtErAl prOcEss OF EpIgynum; MA, mEdIAn ApOphysIs; ML, mEdIAn lObE OF EpIgynum; RTA, rEtrOlAtErAl tIbIAl ApOphysIs; S, spErmAthEcAE; T, tEgulum; TI, tIbIA." httpUri="https://zenodo.org/record/1076218/files/figure.png" pageId="13" pageNumber="64">12E, F</figureCitation>
|
||
,
|
||
<figureCitation id="13412A10FF89FF8BE7DB69DFFD4661F9" box="[511,655,1970,1994]" captionStart="FIGURE 13" captionStartId="19.[151,250,1685,1707]" captionTargetBox="[159,1414,872,1660]" captionTargetId="figure@19.[151,1436,842,1674]" captionTargetPageId="19" captionText="FIGURE 13. Devendra spp. (A–C) D. saama sp. nov., FEmAlE hOlOtypE (MNHN#18302), EpIgynum; (A) VEntrAl VIEw; (B) dOrsAl VIEw; (C) rIght spErmAthEcA, dOrsAl, clEArEd, dEtAIl; (D–F) D. amaiti sp. nov., FEmAlE hOlOtypE (MNHN#18337), EpIgynum;(D) VEntrAl VIEw;(E)dOrsAlVIEw;(F)rIght spErmAthEcA,dOrsAl,clEArEd,dEtAIl.AbbrEVIAtIOns:CD, cOpulAtOry duct; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; LP, lAtErAl prOcEss OF EpIgynum; ML, mEdIAn lObE OF EpIgynum; S, spErmAthEcAE." httpUri="https://zenodo.org/record/1076222/files/figure.png" pageId="13" pageNumber="64">13B, C, E, F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BC53695FF89FF88E6E369B8FE5362DB" blockId="13.[151,1437,1573,2030]" lastBlockId="14.[151,1437,151,1583]" lastPageId="14" lastPageNumber="65" pageId="13" pageNumber="64">
|
||
<emphasis id="B90EEA87FF89FF8BE6E369B8FE9061DD" bold="true" box="[199,345,2005,2030]" pageId="13" pageNumber="64">DesCription.</emphasis>
|
||
Medium sized-spiders, total length 3.90–7.10; sexual dimorphism slight (
|
||
<figureCitation id="13412A10FF89FF8BE29969B8FA8B61DD" box="[1213,1346,2005,2030]" captionStart="FIGURE 9" captionStartId="15.[151,250,1972,1994]" captionTargetBox="[151,1436,663,1949]" captionTargetId="figure@15.[151,1436,663,1949]" captionTargetPageId="15" captionText="FIGURE 9. Devendra spp. (A–C) D. pardalis (SImOn, 1898), FEmAlE; (A) hAbItus, dOrsAl VIEw; (B) AntErIOr VIEw OF OculAr ArEA;(C)EpIgynum, VEntrAl VIEw;(D–E) D. pumilus (SImOn, 1898), mAlE;(D)hAbItus,dOrsAl VIEw;(E)rIght pAlp,VEntrAl VIEw." httpUri="https://zenodo.org/record/1076208/files/figure.png" pageId="13" pageNumber="64">Fig. 9A, D</figureCitation>
|
||
), males smaller with relatively longer legs. Carapace with darker band along each side of
|
||
<emphasis id="B90EEA87FF8AFF88E2516EF4FAE86683" box="[1141,1313,152,176]" italics="true" pageId="14" pageNumber="65">pars cephalica</emphasis>
|
||
, each eye surrounded by black (
|
||
<figureCitation id="13412A10FF8AFF88E7BC6ED1FE3A66E7" box="[408,499,188,213]" captionStart="FIGURE 9" captionStartId="15.[151,250,1972,1994]" captionTargetBox="[151,1436,663,1949]" captionTargetId="figure@15.[151,1436,663,1949]" captionTargetPageId="15" captionText="FIGURE 9. Devendra spp. (A–C) D. pardalis (SImOn, 1898), FEmAlE; (A) hAbItus, dOrsAl VIEw; (B) AntErIOr VIEw OF OculAr ArEA;(C)EpIgynum, VEntrAl VIEw;(D–E) D. pumilus (SImOn, 1898), mAlE;(D)hAbItus,dOrsAl VIEw;(E)rIght pAlp,VEntrAl VIEw." httpUri="https://zenodo.org/record/1076208/files/figure.png" pageId="14" pageNumber="65">Fig. 9B</figureCitation>
|
||
), legs with faint annuli, especially on femora. Abdomen grey with anterodorsal cardiac mark, posteriorly with 2–3 paired chevrons (
|
||
<figureCitation id="13412A10FF8AFF88E4D36EB2FCB366CB" box="[759,890,223,248]" captionStart="FIGURE 9" captionStartId="15.[151,250,1972,1994]" captionTargetBox="[151,1436,663,1949]" captionTargetId="figure@15.[151,1436,663,1949]" captionTargetPageId="15" captionText="FIGURE 9. Devendra spp. (A–C) D. pardalis (SImOn, 1898), FEmAlE; (A) hAbItus, dOrsAl VIEw; (B) AntErIOr VIEw OF OculAr ArEA;(C)EpIgynum, VEntrAl VIEw;(D–E) D. pumilus (SImOn, 1898), mAlE;(D)hAbItus,dOrsAl VIEw;(E)rIght pAlp,VEntrAl VIEw." httpUri="https://zenodo.org/record/1076208/files/figure.png" pageId="14" pageNumber="65">Fig. 9A, D</figureCitation>
|
||
). Carapace pear-shaped in dorsal view, domed; fovea linear, deep (
|
||
<figureCitation id="13412A10FF8AFF88E7576F69FE3A672F" box="[371,499,260,285]" captionStart="FIGURE 9" captionStartId="15.[151,250,1972,1994]" captionTargetBox="[151,1436,663,1949]" captionTargetId="figure@15.[151,1436,663,1949]" captionTargetPageId="15" captionText="FIGURE 9. Devendra spp. (A–C) D. pardalis (SImOn, 1898), FEmAlE; (A) hAbItus, dOrsAl VIEw; (B) AntErIOr VIEw OF OculAr ArEA;(C)EpIgynum, VEntrAl VIEw;(D–E) D. pumilus (SImOn, 1898), mAlE;(D)hAbItus,dOrsAl VIEw;(E)rIght pAlp,VEntrAl VIEw." httpUri="https://zenodo.org/record/1076208/files/figure.png" pageId="14" pageNumber="65">Fig. 9A, D</figureCitation>
|
||
). Ocular area broad, ocular quadrangle trapezoidal, widest behind; eight eyes in 2 rows, anterior row slightly recurved, posterior row nearly straight (
|
||
<figureCitation id="13412A10FF8AFF88E5BA6F4AFC336773" box="[926,1018,295,320]" captionStart="FIGURE 9" captionStartId="15.[151,250,1972,1994]" captionTargetBox="[151,1436,663,1949]" captionTargetId="figure@15.[151,1436,663,1949]" captionTargetPageId="15" captionText="FIGURE 9. Devendra spp. (A–C) D. pardalis (SImOn, 1898), FEmAlE; (A) hAbItus, dOrsAl VIEw; (B) AntErIOr VIEw OF OculAr ArEA;(C)EpIgynum, VEntrAl VIEw;(D–E) D. pumilus (SImOn, 1898), mAlE;(D)hAbItus,dOrsAl VIEw;(E)rIght pAlp,VEntrAl VIEw." httpUri="https://zenodo.org/record/1076208/files/figure.png" pageId="14" pageNumber="65">Fig. 9B</figureCitation>
|
||
). Clypeus low (
|
||
<figureCitation id="13412A10FF8AFF88E2966F4AFAC76773" box="[1202,1294,295,320]" captionStart="FIGURE 9" captionStartId="15.[151,250,1972,1994]" captionTargetBox="[151,1436,663,1949]" captionTargetId="figure@15.[151,1436,663,1949]" captionTargetPageId="15" captionText="FIGURE 9. Devendra spp. (A–C) D. pardalis (SImOn, 1898), FEmAlE; (A) hAbItus, dOrsAl VIEw; (B) AntErIOr VIEw OF OculAr ArEA;(C)EpIgynum, VEntrAl VIEw;(D–E) D. pumilus (SImOn, 1898), mAlE;(D)hAbItus,dOrsAl VIEw;(E)rIght pAlp,VEntrAl VIEw." httpUri="https://zenodo.org/record/1076208/files/figure.png" pageId="14" pageNumber="65">Fig. 9B</figureCitation>
|
||
). Chelicerae stout with large boss, especially in female, pro- and retromargins of fang furrow with three and four stout teeth, respectively. Sternum broadly oval, bluntly curved or with small point posteriorly; labium with shallow basal notch; palpal coxae slightly converging anteriorly, serrula in a single row along outer margin. Legs moderately long; integument finely wrinkled (
|
||
<bibRefCitation id="EFEB4B64FF8AFF88E43C6FDAFCD567E3" author="Griswold" box="[536,796,439,464]" pageId="14" pageNumber="65" refString="GrIswOld, C. E. (1993) InVEstIgAtIOns IntO thE phylOgEny OF thE lycOsOId spIdErs And thEIr kIn (ArAchnIdA: ArAnEAE: LycOsOIdEA). Smithsonian Contributions to Zoology, 539, 1 - 39. https: // dOI. Org / 10.5479 / sI. 00810282.539" type="journal article" year="1993">Griswold 1993: fig. 77</bibRefCitation>
|
||
), with plumose setae (
|
||
<emphasis id="B90EEA87FF8AFF88E23C6FD4FB9F67E3" box="[1048,1110,441,464]" italics="true" pageId="14" pageNumber="65">sensu</emphasis>
|
||
<bibRefCitation id="EFEB4B64FF8AFF88E2796FDAFAC867E3" author="Lehtinen" box="[1117,1281,439,464]" pageId="14" pageNumber="65" refString="LEhtInEn, P. T. (1975) NOtEs On thE phylOgEnEtIc clAssIFIcAtIOn OF ArAnEAE. Proceedings of the 6 th Arachnological Congress, 1975, 26 - 29." type="journal article" year="1975">Lehtinen 1975</bibRefCitation>
|
||
) and feathery scales (
|
||
<bibRefCitation id="EFEB4B64FF8AFF88E6CD6FB1FDEF67C7" author="Griswold" box="[233,550,476,501]" pageId="14" pageNumber="65" refString="GrIswOld, C. E. (1993) InVEstIgAtIOns IntO thE phylOgEny OF thE lycOsOId spIdErs And thEIr kIn (ArAchnIdA: ArAnEAE: LycOsOIdEA). Smithsonian Contributions to Zoology, 539, 1 - 39. https: // dOI. Org / 10.5479 / sI. 00810282.539" type="journal article" year="1993">Griswold 1993: figs. 75, 77</bibRefCitation>
|
||
); trochanters notched. Femora of legs and palpi with dorsal and lateral spines, patellae III and IV with lateral spines, tibiae I and II with 4 pairs of ventral spines and an additional proventral spine just behind apical pair, metatarsi I and II with 3 pairs of ventral spines, males usually with, females without, lateral spines on tibiae and metatarsi I and II; tibiae and metatarsi III and IV with dorsal, lateral, and ventral spines; male with sub-basal crack on leg tibiae; palpal femora with dorsal and lateral spines, tibiae and tarsi of females with spines, tibiae and tarsi of males with or without spines, cymbium usually lacking spines. Preening combs and scopulae absent. Superior tarsal claws pectinate, with inferior tarsal claws simple; claw tufts absent. Trichobothria on tibiae with a basal dorsolateral group and pro and retrolateral rows, retrolateral row extending to apex, metatarsi with dorsal, irregular row or dorsal row plus dorso-basal group, tarsi with dorso-basal row that divides into 2–3 rows apically; palpal tibiae with pro- and retrolateral rows, apparently absent from tarsi; trichobothrial base with transversely-ridged hood (
|
||
<bibRefCitation id="EFEB4B64FF8AFF88E7EA6D29FD25656F" author="Griswold" box="[462,748,836,861]" pageId="14" pageNumber="65" refString="GrIswOld, C. E. (1993) InVEstIgAtIOns IntO thE phylOgEny OF thE lycOsOId spIdErs And thEIr kIn (ArAchnIdA: ArAnEAE: LycOsOIdEA). Smithsonian Contributions to Zoology, 539, 1 - 39. https: // dOI. Org / 10.5479 / sI. 00810282.539" type="journal article" year="1993">Griswold 1993: fig. 73</bibRefCitation>
|
||
). Tarsal organ median, capsulate, orifice keyhole-shaped (
|
||
<bibRefCitation id="EFEB4B64FF8AFF88E6BA6D0AFE6B65B3" author="Griswold" box="[158,418,871,896]" pageId="14" pageNumber="65" refString="GrIswOld, C. E. (1993) InVEstIgAtIOns IntO thE phylOgEny OF thE lycOsOId spIdErs And thEIr kIn (ArAchnIdA: ArAnEAE: LycOsOIdEA). Smithsonian Contributions to Zoology, 539, 1 - 39. https: // dOI. Org / 10.5479 / sI. 00810282.539" type="journal article" year="1993">Griswold 1993: fig. 72</bibRefCitation>
|
||
), with proximal seam. Male palpal tibia about as long as cymbium, tegulum simple; firmly attached embolus (E) arising on retrolateral side of tegulum and tapering to apex; median apophysis (MA) concave, attached in middle of tegulum (
|
||
<figureCitation id="13412A10FF8AFF88E7D36DC2FD9765FB" box="[503,606,943,968]" captionStart="FIGURE 11" captionStartId="16.[151,250,1906,1928]" captionTargetBox="[163,1412,935,1884]" captionTargetId="figure@16.[151,1436,911,1884]" captionTargetPageId="16" captionText="FIGURE 11. Devendra pumilus (SImOn, 1898), mAlE. (A–C) lEFt pAlp; (A) VEntrAl VIEw; (B) rEtrOlAtErAl VIEw; (C) prOlAtErAl VIEw.AbbrEVIAtIOns: C, cOnductOr; E, EmbOlus; MA, mEdIAn ApOphysIs; RTA, rEtrOlAtErAl tIbIAl ApOphysIs; ST, subtEgulum; T, tEgulum; TI, tIbIA." httpUri="https://zenodo.org/record/1076216/files/figure.png" pageId="14" pageNumber="65">Fig. 11A</figureCitation>
|
||
); conductor (C) hyaline, originating in the top of tegulum, fan-shaped and opposing but not embracing apex of embolus (
|
||
<figureCitation id="13412A10FF8AFF88E4F06DB9FCAF65DF" box="[724,870,980,1005]" captionStart="FIGURE 11" captionStartId="16.[151,250,1906,1928]" captionTargetBox="[163,1412,935,1884]" captionTargetId="figure@16.[151,1436,911,1884]" captionTargetPageId="16" captionText="FIGURE 11. Devendra pumilus (SImOn, 1898), mAlE. (A–C) lEFt pAlp; (A) VEntrAl VIEw; (B) rEtrOlAtErAl VIEw; (C) prOlAtErAl VIEw.AbbrEVIAtIOns: C, cOnductOr; E, EmbOlus; MA, mEdIAn ApOphysIs; RTA, rEtrOlAtErAl tIbIAl ApOphysIs; ST, subtEgulum; T, tEgulum; TI, tIbIA." httpUri="https://zenodo.org/record/1076216/files/figure.png" pageId="14" pageNumber="65">Figs 11A–C</figureCitation>
|
||
,
|
||
<figureCitation id="13412A10FF8AFF88E55E6DB9FC0665DF" box="[890,975,980,1004]" captionStart="FIGURE 12" captionStartId="18.[151,250,1893,1915]" captionTargetBox="[168,1414,388,1849]" captionTargetId="figure@18.[151,1436,375,1871]" captionTargetPageId="18" captionText="FIGURE 12. Devendra seriatus (SImOn, 1898). (A–C) lEFt mAlE pAlp; (A) VEntrAl VIEw; (B) rEtrOlAtErAl VIEw; (C) prOlAtErAl VIEw; (D–F) FEmAlE EpIgynum; (D) VEntrAl VIEw; (E) dOrsAl VIEw; (F) rIght spErmAthEcA,dOrsAl, clEArEd, dEtAIl.AbbrEVIAtIOns: C, cOnductOr; CD, cOpulAtOry ducts; E, EmbOlus; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; LP, lAtErAl prOcEss OF EpIgynum; MA, mEdIAn ApOphysIs; ML, mEdIAn lObE OF EpIgynum; RTA, rEtrOlAtErAl tIbIAl ApOphysIs; S, spErmAthEcAE; T, tEgulum; TI, tIbIA." httpUri="https://zenodo.org/record/1076218/files/figure.png" pageId="14" pageNumber="65">12A–C</figureCitation>
|
||
); fundus in subtegulum, reservoir and ejaculatory duct simple, without loops or switchbacks, spiralling around outer margin of tegulum. Abdomen oval, without scuta (
|
||
<figureCitation id="13412A10FF8AFF88E7636A71FE066207" box="[327,463,1052,1077]" captionStart="FIGURE 9" captionStartId="15.[151,250,1972,1994]" captionTargetBox="[151,1436,663,1949]" captionTargetId="figure@15.[151,1436,663,1949]" captionTargetPageId="15" captionText="FIGURE 9. Devendra spp. (A–C) D. pardalis (SImOn, 1898), FEmAlE; (A) hAbItus, dOrsAl VIEw; (B) AntErIOr VIEw OF OculAr ArEA;(C)EpIgynum, VEntrAl VIEw;(D–E) D. pumilus (SImOn, 1898), mAlE;(D)hAbItus,dOrsAl VIEw;(E)rIght pAlp,VEntrAl VIEw." httpUri="https://zenodo.org/record/1076208/files/figure.png" pageId="14" pageNumber="65">Fig. 9A, D</figureCitation>
|
||
). Respiratory system comprising two anterior book lungs and a posterior tracheal spiracle. Epigynum divided by longitudinal epigynal folds into median lobe and lateral lobes; lateral lobes with small teeth (
|
||
<figureCitation id="13412A10FF8AFF88E70A6A09FE56624F" box="[302,415,1124,1149]" captionStart="FIGURE 10" captionStartId="16.[151,250,641,663]" captionTargetBox="[163,1413,196,618]" captionTargetId="figure@16.[151,1436,193,620]" captionTargetPageId="16" captionText="FIGURE 10. Devendra pardalis (SImOn, 1898), FEmAlE. (A–C) EpIgynum; (A) VEntrAl VIEw; (B) dOrsAl VIEw; (C) rIght spErmAthEcA, dOrsAl, clEArEd, dEtAIl.AbbrEVIAtIOns: CD, cOpulAtOry ducts; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; LP, lAtErAl prOcEss OF EpIgynum; ML, mEdIAnlObE OF EpIgynum; S, spErmAthEcAE." httpUri="https://zenodo.org/record/1076212/files/figure.png" pageId="14" pageNumber="65">Figs 10A</figureCitation>
|
||
,
|
||
<figureCitation id="13412A10FF8AFF88E7966A09FE2C624F" box="[434,485,1124,1148]" captionStart="FIGURE 12" captionStartId="18.[151,250,1893,1915]" captionTargetBox="[168,1414,388,1849]" captionTargetId="figure@18.[151,1436,375,1871]" captionTargetPageId="18" captionText="FIGURE 12. Devendra seriatus (SImOn, 1898). (A–C) lEFt mAlE pAlp; (A) VEntrAl VIEw; (B) rEtrOlAtErAl VIEw; (C) prOlAtErAl VIEw; (D–F) FEmAlE EpIgynum; (D) VEntrAl VIEw; (E) dOrsAl VIEw; (F) rIght spErmAthEcA,dOrsAl, clEArEd, dEtAIl.AbbrEVIAtIOns: C, cOnductOr; CD, cOpulAtOry ducts; E, EmbOlus; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; LP, lAtErAl prOcEss OF EpIgynum; MA, mEdIAn ApOphysIs; ML, mEdIAn lObE OF EpIgynum; RTA, rEtrOlAtErAl tIbIAl ApOphysIs; S, spErmAthEcAE; T, tEgulum; TI, tIbIA." httpUri="https://zenodo.org/record/1076218/files/figure.png" pageId="14" pageNumber="65">12D</figureCitation>
|
||
,
|
||
<figureCitation id="13412A10FF8AFF88E7D36A09FD9C624F" box="[503,597,1124,1148]" captionStart="FIGURE 13" captionStartId="19.[151,250,1685,1707]" captionTargetBox="[159,1414,872,1660]" captionTargetId="figure@19.[151,1436,842,1674]" captionTargetPageId="19" captionText="FIGURE 13. Devendra spp. (A–C) D. saama sp. nov., FEmAlE hOlOtypE (MNHN#18302), EpIgynum; (A) VEntrAl VIEw; (B) dOrsAl VIEw; (C) rIght spErmAthEcA, dOrsAl, clEArEd, dEtAIl; (D–F) D. amaiti sp. nov., FEmAlE hOlOtypE (MNHN#18337), EpIgynum;(D) VEntrAl VIEw;(E)dOrsAlVIEw;(F)rIght spErmAthEcA,dOrsAl,clEArEd,dEtAIl.AbbrEVIAtIOns:CD, cOpulAtOry duct; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; LP, lAtErAl prOcEss OF EpIgynum; ML, mEdIAn lObE OF EpIgynum; S, spErmAthEcAE." httpUri="https://zenodo.org/record/1076222/files/figure.png" pageId="14" pageNumber="65">13A, D</figureCitation>
|
||
). Vulva with elongated copulatory duct, oval spermathecae and small fertilization ducts (
|
||
<figureCitation id="13412A10FF8AFF88E7566AEAFDCA6293" box="[370,515,1159,1184]" captionStart="FIGURE 10" captionStartId="16.[151,250,641,663]" captionTargetBox="[163,1413,196,618]" captionTargetId="figure@16.[151,1436,193,620]" captionTargetPageId="16" captionText="FIGURE 10. Devendra pardalis (SImOn, 1898), FEmAlE. (A–C) EpIgynum; (A) VEntrAl VIEw; (B) dOrsAl VIEw; (C) rIght spErmAthEcA, dOrsAl, clEArEd, dEtAIl.AbbrEVIAtIOns: CD, cOpulAtOry ducts; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; LP, lAtErAl prOcEss OF EpIgynum; ML, mEdIAnlObE OF EpIgynum; S, spErmAthEcAE." httpUri="https://zenodo.org/record/1076212/files/figure.png" pageId="14" pageNumber="65">Figs 10B, C</figureCitation>
|
||
,
|
||
<figureCitation id="13412A10FF8AFF88E4376AE5FDAB6293" box="[531,610,1160,1184]" captionStart="FIGURE 12" captionStartId="18.[151,250,1893,1915]" captionTargetBox="[168,1414,388,1849]" captionTargetId="figure@18.[151,1436,375,1871]" captionTargetPageId="18" captionText="FIGURE 12. Devendra seriatus (SImOn, 1898). (A–C) lEFt mAlE pAlp; (A) VEntrAl VIEw; (B) rEtrOlAtErAl VIEw; (C) prOlAtErAl VIEw; (D–F) FEmAlE EpIgynum; (D) VEntrAl VIEw; (E) dOrsAl VIEw; (F) rIght spErmAthEcA,dOrsAl, clEArEd, dEtAIl.AbbrEVIAtIOns: C, cOnductOr; CD, cOpulAtOry ducts; E, EmbOlus; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; LP, lAtErAl prOcEss OF EpIgynum; MA, mEdIAn ApOphysIs; ML, mEdIAn lObE OF EpIgynum; RTA, rEtrOlAtErAl tIbIAl ApOphysIs; S, spErmAthEcAE; T, tEgulum; TI, tIbIA." httpUri="https://zenodo.org/record/1076218/files/figure.png" pageId="14" pageNumber="65">12E, F</figureCitation>
|
||
,
|
||
<figureCitation id="13412A10FF8AFF88E4546AE5FCC26293" box="[624,779,1160,1184]" captionStart="FIGURE 13" captionStartId="19.[151,250,1685,1707]" captionTargetBox="[159,1414,872,1660]" captionTargetId="figure@19.[151,1436,842,1674]" captionTargetPageId="19" captionText="FIGURE 13. Devendra spp. (A–C) D. saama sp. nov., FEmAlE hOlOtypE (MNHN#18302), EpIgynum; (A) VEntrAl VIEw; (B) dOrsAl VIEw; (C) rIght spErmAthEcA, dOrsAl, clEArEd, dEtAIl; (D–F) D. amaiti sp. nov., FEmAlE hOlOtypE (MNHN#18337), EpIgynum;(D) VEntrAl VIEw;(E)dOrsAlVIEw;(F)rIght spErmAthEcA,dOrsAl,clEArEd,dEtAIl.AbbrEVIAtIOns:CD, cOpulAtOry duct; FD, FErtIlIZAtIOn duct; LL, lAtErAl lObE OF EpIgynum; LP, lAtErAl prOcEss OF EpIgynum; ML, mEdIAn lObE OF EpIgynum; S, spErmAthEcAE." httpUri="https://zenodo.org/record/1076222/files/figure.png" pageId="14" pageNumber="65">13B, C, E, F</figureCitation>
|
||
). Colulus a fleshy triangular lobe set with several setae. Cribellum absent. Six spinnerets, anterior (ALS) and posterior laterals (PLS) two-segmented, posterior medians (PMS) one-segmented.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C360651EFF8AFF88E6E36A99FCAB6038" pageId="14" pageNumber="65" type="description">
|
||
<paragraph id="8BC53695FF8AFF88E6E36A99FDAA6303" blockId="14.[151,1437,151,1583]" pageId="14" pageNumber="65">
|
||
<emphasis id="B90EEA87FF8AFF88E6E36A99FEAF633E" bold="true" box="[199,358,1268,1293]" pageId="14" pageNumber="65">Composition.</emphasis>
|
||
Five species:
|
||
<taxonomicName id="4C7A4D16FF8AFF88E4236A98FCF8633F" authority="(Simon, 1898)" baseAuthorityName="Simon" baseAuthorityYear="1898" box="[519,817,1268,1293]" class="Arachnida" family="Zoropsidae" genus="Devendra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="65" phylum="Arthropoda" rank="species" species="pardalis">
|
||
<emphasis id="B90EEA87FF8AFF88E4236A98FD4C633F" box="[519,645,1268,1292]" italics="true" pageId="14" pageNumber="65">D. pardalis</emphasis>
|
||
(Simon, 1898)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7A4D16FF8AFF88E5196A98FBAD633F" authority="(Simon, 1898)" baseAuthorityName="Simon" baseAuthorityYear="1898" box="[829,1124,1268,1293]" class="Arachnida" family="Zoropsidae" genus="Devendra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="65" phylum="Arthropoda" rank="species" species="pumilus">
|
||
<emphasis id="B90EEA87FF8AFF88E5196A98FC70633F" box="[829,953,1268,1292]" italics="true" pageId="14" pageNumber="65">D. pumilus</emphasis>
|
||
(Simon, 1898)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7A4D16FF8AFF88E2546A98FA5E633F" authority="(Simon, 1898)" baseAuthorityName="Simon" baseAuthorityYear="1898" box="[1136,1431,1268,1293]" class="Arachnida" family="Zoropsidae" genus="Devendra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="65" phylum="Arthropoda" rank="species" species="seriatus">
|
||
<emphasis id="B90EEA87FF8AFF88E2546A98FB25633F" box="[1136,1260,1269,1292]" italics="true" pageId="14" pageNumber="65">D. seriatus</emphasis>
|
||
(Simon, 1898)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C7A4D16FF8AFF88E6B36B74FECB6303" authority="Polotow & Griswold, 2017" authorityName="Polotow & Griswold" authorityYear="2017" box="[151,258,1305,1328]" class="Arachnida" family="Zoropsidae" genus="Devendra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="65" phylum="Arthropoda" rank="species" species="saama" status="sp. nov.">
|
||
<emphasis id="B90EEA87FF8AFF88E6B36B74FECB6303" box="[151,258,1305,1328]" italics="true" pageId="14" pageNumber="65">D. saama</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23D57FCFF8AFF88E72D6B7AFEAB6303" box="[265,354,1303,1328]" pageId="14" pageNumber="65" rank="species">
|
||
<emphasis id="B90EEA87FF8AFF88E72D6B7AFEAB6303" bold="true" box="[265,354,1303,1328]" pageId="14" pageNumber="65">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
and
|
||
<taxonomicName id="4C7A4D16FF8AFF88E7BD6B74FDCB631C" authority="Polotow & Griswold, 2017" authorityName="Polotow & Griswold" authorityYear="2017" box="[409,514,1304,1328]" class="Arachnida" family="Zoropsidae" genus="Devendra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="65" phylum="Arthropoda" rank="species" species="amaiti" status="sp. nov.">
|
||
<emphasis id="B90EEA87FF8AFF88E7BD6B74FDCB631C" box="[409,514,1304,1328]" italics="true" pageId="14" pageNumber="65">D. amaiti</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A23D57FCFF8AFF88E42E6B7AFDAA6303" box="[522,611,1303,1328]" pageId="14" pageNumber="65" rank="species">
|
||
<emphasis id="B90EEA87FF8AFF88E42E6B7AFDAA6303" bold="true" box="[522,611,1303,1328]" pageId="14" pageNumber="65">sp. nov.</emphasis>
|
||
</taxonomicNameLabel>
|
||
</paragraph>
|
||
<paragraph id="8BC53695FF8AFF88E6E36B53FCAB6038" blockId="14.[151,1437,151,1583]" pageId="14" pageNumber="65">
|
||
<emphasis id="B90EEA87FF8AFF88E6E36B53FE4F6364" bold="true" box="[199,390,1342,1367]" pageId="14" pageNumber="65">Natural history.</emphasis>
|
||
All known specimens were collected in the late 19th century near Kandy and Nuwara Eliya, which are located in the highlands of southern
|
||
<collectingCountry id="F36D7605FF8AFF88E4976B0FFCEB6348" box="[691,802,1378,1403]" name="Sri Lanka" pageId="14" pageNumber="65">Sri Lanka</collectingCountry>
|
||
at above
|
||
<quantity id="4C829B70FF8AFF88E5B06B0EFC146349" box="[916,989,1379,1403]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="14" pageNumber="65" unit="m" value="800.0">800 m</quantity>
|
||
elevation (
|
||
<figureCitation id="13412A10FF8AFF88E27A6B0FFB7A6348" box="[1118,1203,1378,1403]" captionStart="FIGURE14" captionStartId="20.[151,250,1476,1498]" captionTargetBox="[150,1436,807,1454]" captionTargetId="figure@20.[150,1436,807,1454]" captionTargetPageId="20" captionText="FIGURE14. (A) PArtIAl wOrld mAp (ExcEptNEArctIc AndNEOtrOpIcAl rEAlms),shOwIng thE lOcAtIOnOF thE IslAnd cOuntry OF SrI LAnkA In SOuth AsIA. (B) ClOsE up OF SrI LAnkA, wIth cOllEctIOn lOcAlItIEs OF Campostichomma And Devendra spEcIEs: whItE squArE, Devendra pardalis (SImOn, 1898), D. pumilus (SImOn, 1898) And D. saama sp. nov.; whItE cIrclE, D seriatus (SImOn, 1898) And Devendra amaiti sp. nov.; blAck squArE, Compostichomma harasbedda sp. nov.; blAck cIrclE, Compostichomma alawala sp. nov.; blAck trIAnglE, Compostichomma mudduk sp. nov. ThE cOllEctIOn dAtA OF Compostichomma manicatum KArsch, 1892 dO nOt prOVIdE A spEcIFIclOcAlIty In SrI LAnkA, thus thE spEcIEs Is nOt shOwn In thIs mAp." httpUri="https://zenodo.org/record/1076226/files/figure.png" pageId="14" pageNumber="65">Fig. 14</figureCitation>
|
||
). Today, this area in
|
||
<collectingRegion id="49BEF877FF8AFF88E6B36BEBFE9C63AC" box="[151,341,1414,1439]" country="Sri Lanka" name="Central" pageId="14" pageNumber="65">Central Province</collectingRegion>
|
||
is largely cultivated for tea plantations but there are remnants of montane forest, especially near Nuwara Eliya and at Horton Plains National Park, at elevations from 1800 to more than
|
||
<quantity id="4C829B70FF8AFF88E2836BC6FACB63F1" box="[1191,1282,1451,1475]" metricMagnitude="3" metricUnit="m" metricValue="2.2" pageId="14" pageNumber="65" unit="m" value="2200.0">2200 m</quantity>
|
||
. Such a high species richness (five) in one small area suggests that
|
||
<taxonomicName id="4C7A4D16FF8AFF88E52B6BA2FCB463D4" authorityName="Lehtinen" authorityYear="1967" box="[783,893,1487,1511]" class="Arachnida" family="Zoropsidae" genus="Devendra" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="14" pageNumber="65" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90EEA87FF8AFF88E52B6BA2FCB463D4" box="[783,893,1487,1511]" italics="true" pageId="14" pageNumber="65">Devendra</emphasis>
|
||
</taxonomicName>
|
||
species may be short-range endemics and that more species remain to be discovered in other montane regions.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C360651EFF8AFF88E6E3687BFD05601C" box="[199,716,1558,1583]" pageId="14" pageNumber="65" type="distribution">
|
||
<paragraph id="8BC53695FF8AFF88E6E3687BFD05601C" blockId="14.[151,1437,151,1583]" box="[199,716,1558,1583]" pageId="14" pageNumber="65">
|
||
<emphasis id="B90EEA87FF8AFF88E6E3687BFEA9601C" bold="true" box="[199,352,1558,1583]" pageId="14" pageNumber="65">Distribution.</emphasis>
|
||
Endemic to
|
||
<collectingCountry id="F36D7605FF8AFF88E7CA687BFD95601C" box="[494,604,1558,1583]" name="Sri Lanka" pageId="14" pageNumber="65">Sri Lanka</collectingCountry>
|
||
(
|
||
<figureCitation id="13412A10FF8AFF88E44F687BFD76601C" box="[619,703,1558,1583]" captionStart="FIGURE14" captionStartId="20.[151,250,1476,1498]" captionTargetBox="[150,1436,807,1454]" captionTargetId="figure@20.[150,1436,807,1454]" captionTargetPageId="20" captionText="FIGURE14. (A) PArtIAl wOrld mAp (ExcEptNEArctIc AndNEOtrOpIcAl rEAlms),shOwIng thE lOcAtIOnOF thE IslAnd cOuntry OF SrI LAnkA In SOuth AsIA. (B) ClOsE up OF SrI LAnkA, wIth cOllEctIOn lOcAlItIEs OF Campostichomma And Devendra spEcIEs: whItE squArE, Devendra pardalis (SImOn, 1898), D. pumilus (SImOn, 1898) And D. saama sp. nov.; whItE cIrclE, D seriatus (SImOn, 1898) And Devendra amaiti sp. nov.; blAck squArE, Compostichomma harasbedda sp. nov.; blAck cIrclE, Compostichomma alawala sp. nov.; blAck trIAnglE, Compostichomma mudduk sp. nov. ThE cOllEctIOn dAtA OF Compostichomma manicatum KArsch, 1892 dO nOt prOVIdE A spEcIFIclOcAlIty In SrI LAnkA, thus thE spEcIEs Is nOt shOwn In thIs mAp." httpUri="https://zenodo.org/record/1076226/files/figure.png" pageId="14" pageNumber="65">Fig. 14</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |