treatments-xml/data/03/D3/87/03D387539E1F66504EA0FD156CE8FAF6.xml

<document id="E2C316E156291654234343073F59AE9A" ID-DOI="10.5281/zenodo.4531515" ID-ISSN="1942-1354" ID-Zenodo-Dep="4531515" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1612980280246" checkinUser="felipe" docAuthor="Stebnicka, Z. Teresa &amp; Skelley, Paul E." docDate="2009" docId="03D387539E1F66504EA0FD156CE8FAF6" docLanguage="en" docName="InsectaMundi.2009.0062.1-16.pdf" docOrigin="Insecta Mundi 2009 (62)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Haroldiataenius mariarum" docType="treatment" docVersion="6" lastPageNumber="5" masterDocId="FFEAFF2B9E1A66564E60FFB06948FFDB" masterDocTitle="A revision of the genus Haroldiataenius Chalumeau (Scarabaeidae: Aphodiinae: Eupariini)" masterLastPageNumber="16" masterPageNumber="1" pageNumber="4" updateTime="1698869595032" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="B6B83F85A7F9227C7F50ABF4BABEB8AC">A revision of the genus Haroldiataenius Chalumeau (Scarabaeidae: Aphodiinae: Eupariini)</mods:title>
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<mods:namePart id="4EF8B9E122D89E089A6880F040CD7117">Stebnicka, Z. Teresa</mods:namePart>
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<mods:namePart id="9B9A688658EADCAA2AE781245DC2AF74">Skelley, Paul E.</mods:namePart>
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<mods:title id="17E1FCA99CF0DC984C79F71629EAB412">Insecta Mundi</mods:title>
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<mods:date id="172241585F23A1326D15D82B1B29C4FF">2009</mods:date>
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<subSubSection id="C36065CE9E1F66534EA0FD156BD4FD60" box="[192,668,675,700]" pageId="5" pageNumber="4" type="nomenclature">
<paragraph id="8BC536459E1F66534EA0FD156BD4FD60" blockId="5.[192,668,675,731]" box="[192,668,675,700]" pageId="5" pageNumber="4">
<heading id="D08D81299E1F66534EA0FD156BD4FD60" bold="true" box="[192,668,675,700]" fontSize="10" level="2" pageId="5" pageNumber="4" reason="3">
<taxonomicName id="4C7A4DC69E1F66534EA0FD156BD4FD60" authority="(Bates)" baseAuthorityName="Bates" baseAuthorityYear="1887" box="[192,668,675,700]" class="Insecta" family="Aphodiidae" genus="Haroldiataenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="mariarum">
<emphasis id="B90EEA579E1F66534EA0FD156BD4FD60" bold="true" box="[192,668,675,700]" pageId="5" pageNumber="4">
<emphasis id="B90EEA579E1F66534EA0FD156B7EFD67" bold="true" box="[192,566,677,700]" italics="true" pageId="5" pageNumber="4">Haroldiataenius mariarum</emphasis>
(Bates)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36065CE9E1F66534EA0FD7368D2FD00" box="[192,410,707,731]" pageId="5" pageNumber="4" type="description">
<paragraph id="8BC536459E1F66534EA0FD7368D2FD00" blockId="5.[192,668,675,731]" box="[192,410,707,731]" pageId="5" pageNumber="4">
(
<figureCitation id="13412AC09E1F66534EA9FD736846FD00" box="[201,270,707,731]" captionStart="Figures 1-7" captionStartId="4.[192,289,1434,1456]" captionTargetBox="[192,1440,193,1402]" captionTargetId="figure-142@4.[190,1442,191,1404]" captionTargetPageId="4" captionText="Figures 1-7. Haroldiataenius spp. 1) H. mariarum, lateral habitus. 2) H. hintoni lateral habitus. 3) H. lucanus, mesotibia. 4) H. semipilosus, mesotibia. 5) H. saramari, lateral habitus. 6) H. lucanus, lateral habitus. 7) H. limbatus, lateral habitus." figureDoi="http://doi.org/10.5281/zenodo.4531517" httpUri="https://zenodo.org/record/4531517/files/figure.png" pageId="5" pageNumber="4">Fig. 1</figureCitation>
, 12, 16, 22)
</paragraph>
</subSubSection>
<subSubSection id="C36065CE9E1F66534EA0FCB46CC0FC80" pageId="5" pageNumber="4" type="reference_group">
<paragraph id="8BC536459E1F66534EA0FCB46AD0FCC0" blockId="5.[192,1416,771,860]" box="[192,920,771,795]" pageId="5" pageNumber="4">
<treatmentCitationGroup id="AB6A116B9E1F66534EA0FCB46AD0FCC0" box="[192,920,771,795]" pageId="5" pageNumber="4">
<treatmentCitation id="0ADB10549E1F66534EA0FCB46B3CFCC0" author="Bates, H. W." box="[192,628,771,795]" page="102" pageId="5" pageNumber="4" year="1887">
<taxonomicName id="4C7A4DC69E1F66534EA0FCB46B3CFCC0" authority="Bates 1887: 102" authorityName="Bates" authorityPageNumber="102" authorityYear="1887" box="[192,628,771,795]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="mariarum">
<emphasis id="B90EEA579E1F66534EA0FCB468F9FCC0" box="[192,433,772,795]" italics="true" pageId="5" pageNumber="4">Ataenius mariarum</emphasis>
<bibRefCitation id="EFEB4BB49E1F66534FD6FCB46B3CFCC0" author="Bates, H. W." box="[438,628,771,795]" pageId="5" pageNumber="4" pagination="25 - 160" refId="ref10671" refString="Bates, H. W. 1887. Insecta. Coleoptera. Pectinicornia and Lamellicornia. (Copridae, Aphodiidae, Orphnidae, Hybosoridae, Geotrupidae, Trogidae, Aclopidae, Chasmatopteridae, Melolonthidae). Biologia Centrali- Americana, Vol. 2, Part 2: 25 - 160." type="journal article" year="1887">Bates 1887: 102</bibRefCitation>
</taxonomicName>
</treatmentCitation>
– 
<treatmentCitation id="0ADB10549E1F66534CEDFCB36ADCFCC0" author="Cartwright, O. L." box="[653,916,771,795]" page="101" pageId="5" pageNumber="4" year="1964">
<bibRefCitation id="EFEB4BB49E1F66534CEDFCB36ADCFCC0" author="Cartwright, O. L." box="[653,916,771,795]" pageId="5" pageNumber="4" pagination="101 - 104" refId="ref10748" refString="Cartwright, O. L. 1964. Lectotype designations and new synonymy in the genus Ataenius. Coleopterists Bulletin 18: 101 - 104." type="journal article" year="1964">Cartwright 1964: 101</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BC536459E1F66534EA0FC936CC0FC80" blockId="5.[192,1416,771,860]" pageId="5" pageNumber="4">
<treatmentCitationGroup id="AB6A116B9E1F66534EA0FC936CC0FC80" pageId="5" pageNumber="4">
<taxonomicName id="4C7A4DC69E1F66534EA0FC936B46FCE0" baseAuthorityName="Bates" baseAuthorityYear="1887" box="[192,526,803,827]" class="Insecta" family="Aphodiidae" genus="Haroldiataenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="mariarum">
<emphasis id="B90EEA579E1F66534EA0FC936B46FCE0" box="[192,526,803,827]" italics="true" pageId="5" pageNumber="4">Haroldiataenius mariarum</emphasis>
</taxonomicName>
– 
<treatmentCitation id="0ADB10549E1F66534C45FC936A2DFCE0" author="Chalumeau, F." box="[549,869,803,827]" page="138" pageId="5" pageNumber="4" year="1981">
<bibRefCitation id="EFEB4BB49E1F66534C45FC936A2DFCE0" author="Chalumeau, F." box="[549,869,803,827]" pageId="5" pageNumber="4" pagination="137 - 142" refId="ref10805" refString="Chalumeau, F. 1981. Nouveaux genre et sous-genres d'Eupariini (Aphodiinae) d'Amerique tropicale. Nouvelle Revue d'Entomologie 11 (2): 137 - 142." type="journal article" year="1981">Chalumeau 1981: 138-139</bibRefCitation>
</treatmentCitation>
, fig. 1, 3; 
<treatmentCitation id="0ADB10549E1F66534DAFFC936DF6FCE0" author="Dellacasa, M." box="[975,1214,803,827]" page="339" pageId="5" pageNumber="4" year="1988">
<bibRefCitation id="EFEB4BB49E1F66534DAFFC936DF6FCE0" author="Dellacasa, M." box="[975,1214,803,827]" pageId="5" pageNumber="4" pagination="1 - 455" refId="ref10876" refString="Dellacasa, M. 1988. Contribution to a world-wide catalogue of Aegialiidae, Aphodiidae, Aulonocnemidae, Termitotrogidae (Coleoptera Scarabaeoidea). Memorie della Societa Entomologica Italiana [1987] 66: 1 - 455." type="journal article" year="1988">Dellacasa 1988: 339</bibRefCitation>
</treatmentCitation>
(catalogue). 
<treatmentCitation id="0ADB10549E1F66534EA0FCF36B8CFC80" author="Schmidt, A." box="[192,708,835,859]" page="105" pageId="5" pageNumber="4" year="1916">
<taxonomicName id="4C7A4DC69E1F66534EA0FCF36B8CFC80" authority="Schmidt 1916: 105" authorityName="Schmidt" authorityPageNumber="105" authorityYear="1916" box="[192,708,835,859]" class="Insecta" family="Aphodiidae" genus="Psammodius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="perpolitus">
<emphasis id="B90EEA579E1F66534EA0FCF36897FC80" box="[192,479,835,859]" italics="true" pageId="5" pageNumber="4">Psammodius perpolitus</emphasis>
<bibRefCitation id="EFEB4BB49E1F66534F84FCF36B8CFC80" author="Schmidt, A." box="[484,708,835,859]" pageId="5" pageNumber="4" pagination="95 - 116" refId="ref11168" refString="Schmidt, A. 1916. Namenanderungen und Beschreibung neuer Aphodiinen (Col.). Archiv fur Naturgeschichte, Abt. A, 82: 95 - 116." type="journal article" year="1916">Schmidt 1916: 105</bibRefCitation>
</taxonomicName>
</treatmentCitation>
– 
<treatmentCitation id="0ADB10549E1F66534CBEFCF36D54FC80" author="Chalumeau, F." box="[734,1052,835,859]" page="138" pageId="5" pageNumber="4" year="1981">
<bibRefCitation id="EFEB4BB49E1F66534CBEFCF36D54FC80" author="Chalumeau, F." box="[734,1052,835,859]" pageId="5" pageNumber="4" pagination="137 - 142" refId="ref10805" refString="Chalumeau, F. 1981. Nouveaux genre et sous-genres d'Eupariini (Aphodiinae) d'Amerique tropicale. Nouvelle Revue d'Entomologie 11 (2): 137 - 142." type="journal article" year="1981">Chalumeau 1981: 138-139</bibRefCitation>
</treatmentCitation>
(as synonym of 
<taxonomicName id="4C7A4DC69E1F66534AB9FCF46C30FC80" baseAuthorityName="Bates" baseAuthorityYear="1887" box="[1241,1400,836,859]" class="Insecta" family="Aphodiidae" genus="Haroldiataenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="mariarum">
<emphasis id="B90EEA579E1F66534AB9FCF46C30FC80" box="[1241,1400,836,859]" italics="true" pageId="5" pageNumber="4">A. mariarum</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C36065CE9E1F66534EA0FC336AD0FC20" pageId="5" pageNumber="4" type="diagnosis">
<paragraph id="8BC536459E1F66534EA0FC336AD0FC20" blockId="5.[192,1440,899,1019]" pageId="5" pageNumber="4">
<emphasis id="B90EEA579E1F66534EA0FC336801FC40" bold="true" box="[192,329,899,923]" pageId="5" pageNumber="4">Diagnosis</emphasis>
. Clypeal margin angulate each side of middle, surface strongly wrinkled. Pronotum with coarse punctures laterally, widely scattered, weakly defined. Pronotum with basal marginal line strong, broad laterally, narrowed medially. Lateral margins of pronotum and elytron with long fringe of setae. 
<taxonomicName id="4C7A4DC69E1F66534EA0FC536B47FC20" baseAuthorityName="Bates" baseAuthorityYear="1887" box="[192,527,995,1019]" class="Insecta" family="Aphodiidae" genus="Haroldiataenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="mariarum">
<emphasis id="B90EEA579E1F66534EA0FC536B47FC20" box="[192,527,995,1019]" italics="true" pageId="5" pageNumber="4">Haroldiataenius mariarum</emphasis>
</taxonomicName>
is found in 
<collectingCountry id="F36D76D59E1F66534CFAFC536BB9FC20" box="[666,753,995,1019]" name="Mexico" pageId="5" pageNumber="4">Mexico</collectingCountry>
and 
<collectingCountry id="F36D76D59E1F66534D4AFC546ADCFC20" box="[810,916,996,1019]" name="Panama" pageId="5" pageNumber="4">Panama</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C36065CE9E1F66534EA0FB936ADCFA60" pageId="5" pageNumber="4" type="description">
<paragraph id="8BC536459E1F66534EA0FB936ADCFA60" blockId="5.[192,1441,1059,1467]" pageId="5" pageNumber="4">
<emphasis id="B90EEA579E1F66534EA0FB93682CFBE0" bold="true" box="[192,356,1059,1083]" pageId="5" pageNumber="4">Description</emphasis>
. Length 4.5-5.0 mm. Body oval, strongly convex, castaneous, shining (
<figureCitation id="13412AC09E1F66534ADBFB936C48FBE0" box="[1211,1280,1059,1083]" captionStart="Figures 1-7" captionStartId="4.[192,289,1434,1456]" captionTargetBox="[192,1440,193,1402]" captionTargetId="figure-142@4.[190,1442,191,1404]" captionTargetPageId="4" captionText="Figures 1-7. Haroldiataenius spp. 1) H. mariarum, lateral habitus. 2) H. hintoni lateral habitus. 3) H. lucanus, mesotibia. 4) H. semipilosus, mesotibia. 5) H. saramari, lateral habitus. 6) H. lucanus, lateral habitus. 7) H. limbatus, lateral habitus." figureDoi="http://doi.org/10.5281/zenodo.4531517" httpUri="https://zenodo.org/record/4531517/files/figure.png" pageId="5" pageNumber="4">Fig. 1</figureCitation>
, 12). Clypeal margin subdentate each side of median emargination; surface coarsely wrinkled, frontal area smooth; genae small. Pronotum transverse, sides and base margined, setose; surface with extremely minute punctures throughout and moderate, shallow punctures scattered on sides. Scutellum relatively large, semioval. Elytral humeral lamella obtusely produced laterad; striae narrowly impressed, punctures fine; intervals flat to slightly convex, smooth; elytral margin with long pale setae. Ventral sclerites setose; meso-metasternal carina short with clump of setae; metasternum smooth; abdominal sternites 2-4 without fluting along sutures, all sternites with row of setae; disc of pygidium weakly eroded, punctate. All femora setigerous; posterior line of metafemur incomplete; meso- and metatibia strongly widened apically; mesotibia with traces of transverse ridges; metatibia with row of granules on outer side and row of very short setae apically, without accessory spine; basal tarsomere of metatarsus markedly shorter than upper tibial spur and subequal in length to following three tarsomeres combined. Epipharynx as in Fig. 16. Male genitalia with parameres narrowed, elongate (
<figureCitation id="13412AC09E1F66534D51FA136ACFFA60" box="[817,903,1443,1467]" captionStart="Figures 22-28" captionStartId="9.[193,290,1259,1281]" captionTargetBox="[239,1414,202,1227]" captionTargetId="figure-200@9.[190,1443,192,1243]" captionTargetPageId="9" captionText="Figures 22-28. Haroldiataenius spp., male genitalia. 22) H. mariarum. 23) H. limbatus. 24) H. hintoni. 25) H. lucanus. 26) H. griffini. 27) H. saramari. 28) H. semipilosus." figureDoi="http://doi.org/10.5281/zenodo.4531521" httpUri="https://zenodo.org/record/4531521/files/figure.png" pageId="5" pageNumber="4">Fig. 22</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C36065CE9E1F66534EA0FA536882F880" pageId="5" pageNumber="4" type="materials_examined">
<paragraph id="8BC536459E1F66534EA0FA536B9CF9C0" blockId="5.[192,1440,1507,1627]" pageId="5" pageNumber="4">
<emphasis id="B90EEA579E1F66534EA0FA536801FA20" bold="true" box="[192,329,1507,1531]" pageId="5" pageNumber="4">Type data</emphasis>
. 
<taxonomicName id="4C7A4DC69E1F66534F37FA546B02FA20" authorityName="Bates" authorityYear="1887" box="[343,586,1508,1531]" class="Insecta" family="Aphodiidae" genus="Ataenius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="mariarum">
<emphasis id="B90EEA579E1F66534F37FA546B02FA20" box="[343,586,1508,1531]" italics="true" pageId="5" pageNumber="4">Ataenius mariarum</emphasis>
</taxonomicName>
(examined, sex not determined) 
<typeStatus id="54C188E79E1F66534DB9FA536D01FA20" box="[985,1097,1507,1531]" pageId="5" pageNumber="4" type="lectotype">lectotype</typeStatus>
(Tres Marias Is., W 
<collectingCountry id="F36D76D59E1F66534B20FA536CD1FA20" box="[1344,1433,1507,1531]" name="Mexico" pageId="5" pageNumber="4">Mexico</collectingCountry>
) designated by 
<bibRefCitation id="EFEB4BB49E1F66534F0EF9B36B06F9C0" author="Cartwright, O. L." box="[366,590,1539,1563]" pageId="5" pageNumber="4" pagination="101 - 104" refId="ref10748" refString="Cartwright, O. L. 1964. Lectotype designations and new synonymy in the genus Ataenius. Coleopterists Bulletin 18: 101 - 104." type="journal article" year="1964">Cartwright (1964)</bibRefCitation>
, in 
<collectionCode id="ED6BAE809E1F66534C16F9B46B99F9C0" box="[630,721,1540,1563]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="5" pageNumber="4">NHML</collectionCode>
.
</paragraph>
<paragraph id="8BC536459E1F66534E90F9936B6CF980" blockId="5.[192,1440,1507,1627]" pageId="5" pageNumber="4">
<taxonomicName id="4C7A4DC69E1F66534E90F9936B46F9E0" authorityName="Schmidt" authorityYear="1916" box="[240,526,1571,1595]" class="Insecta" family="Aphodiidae" genus="Psammodius" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="perpolitus">
<emphasis id="B90EEA579E1F66534E90F9936B46F9E0" box="[240,526,1571,1595]" italics="true" pageId="5" pageNumber="4">Psammodius perpolitus</emphasis>
</taxonomicName>
(examined, sex not determined): 
<typeStatus id="54C188E79E1F66534DFDF9936D43F9E0" box="[925,1035,1571,1595]" pageId="5" pageNumber="4" type="lectotype">lectotype</typeStatus>
(
<collectingCountry id="F36D76D59E1F66534A79F9936DCAF9E0" box="[1049,1154,1571,1595]" name="Panama" pageId="5" pageNumber="4">Panama</collectingCountry>
, Vadona) designated by 
<bibRefCitation id="EFEB4BB49E1F66534EA0F9F368ECF980" author="Chalumeau, F." box="[192,420,1603,1627]" pageId="5" pageNumber="4" pagination="137 - 142" refId="ref10805" refString="Chalumeau, F. 1981. Nouveaux genre et sous-genres d'Eupariini (Aphodiinae) d'Amerique tropicale. Nouvelle Revue d'Entomologie 11 (2): 137 - 142." type="journal article" year="1981">Chalumeau (1981)</bibRefCitation>
, in 
<collectionCode id="ED6BAE809E1F66534FADF9F36B69F980" box="[461,545,1603,1627]" country="Sweden" httpUri="http://grbio.org/cool/rsbi-vwmq" name="Swedish Museum of Natural History, Entomology Collections" pageId="5" pageNumber="4">NHRS</collectionCode>
.
</paragraph>
<paragraph id="8BC536459E1F66534EA0F9336882F880" blockId="5.[192,1441,1667,1883]" pageId="5" pageNumber="4">
<emphasis id="B90EEA579E1F66534EA0F93368ACF940" bold="true" box="[192,484,1667,1691]" pageId="5" pageNumber="4">Specimens examined</emphasis>
(21). 
<materialsCitation id="3B123C189E1F66534C47F9336C69F940" box="[551,1313,1667,1692]" collectionCode="ISEA" collectorName="Tres Marias Is." country="Mexico" location="Mexico" pageId="5" pageNumber="4" specimenCount="1">
<emphasis id="B90EEA579E1F66534C47F9336BC2F940" bold="true" box="[551,650,1667,1691]" pageId="5" pageNumber="4">
<collectingCountry id="F36D76D59E1F66534C47F9336BC2F940" box="[551,650,1667,1691]" name="Mexico" pageId="5" pageNumber="4">Mexico</collectingCountry>
</emphasis>
– (1 ex from 
<typeStatus id="54C188E79E1F66534D46F9346A14F947" box="[806,860,1668,1692]" pageId="5" pageNumber="4">type</typeStatus>
series) 
<collectorName id="268F53939E1F66534DDAF9346D3EF940" box="[954,1142,1667,1691]" pageId="5" pageNumber="4">Tres Marias Is.</collectorName>
(
<collectionCode id="ED6BAE809E1F66534AE4F9336D83F940" box="[1156,1227,1667,1691]" pageId="5" pageNumber="4">ISEA</collectionCode>
); (6 ex)
</materialsCitation>
<materialsCitation id="3B123C189E1F66534B47F9336A2BF960" collectingDate="1956-06-28" collectionCode="VI, R, PESC" collectorName="E. Beer" country="Mexico" location="Acapulco" pageId="5" pageNumber="4" specimenCount="1" stateProvince="Guerrero">
<collectingRegion id="49BEF8A79E1F66534B47F9336CD4F940" box="[1319,1436,1667,1691]" country="Mexico" name="Guerrero" pageId="5" pageNumber="4">Guerrero</collectingRegion>
, 
<location id="8EA5609E9E1F66534EA0F913687BF960" LSID="urn:lsid:plazi:treatment:03D387539E1F66504EA0FD156CE8FAF6:8EA5609E9E1F66534EA0F913687BF960" box="[192,307,1699,1723]" country="Mexico" name="Acapulco" pageId="5" pageNumber="4" stateProvince="Guerrero">Acapulco</location>
, 
<date id="FFC410859E1F66534F5DF9136881F960" box="[317,457,1699,1723]" pageId="5" pageNumber="4" value="1956-06-28">
<collectingDate id="EF80E96D9E1F66534F5DF9136881F960" box="[317,457,1699,1723]" pageId="5" pageNumber="4" value="1956-06-28">
28. 
<collectionCode id="ED6BAE809E1F66534F01F91368CCF960" box="[353,388,1699,1723]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="5" pageNumber="4">VI</collectionCode>
.1956
</collectingDate>
</date>
. 
<collectionCode id="ED6BAE809E1F66534FB2F91468A1F960" box="[466,489,1700,1723]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="5" pageNumber="4">R</collectionCode>
. 
<collectorName id="268F53939E1F66534F93F9146B05F960" box="[499,589,1700,1723]" pageId="5" pageNumber="4">E. Beer</collectorName>
&amp; party (
<collectionCode id="ED6BAE809E1F66534CA0F9136A46F960" box="[704,782,1699,1723]" pageId="5" pageNumber="4">PESC</collectionCode>
); (2 ex)
</materialsCitation>
<materialsCitation id="3B123C189E1F66534D09F9136823F900" collectingDate="1938-08-20" collectionCode="FSCA" collectorName="L. Lipovsky" country="Mexico" location="Acapulco" pageId="5" pageNumber="4" specimenCount="1" stateProvince="Guerrero">
<collectingRegion id="49BEF8A79E1F66534D09F9136A96F960" box="[873,990,1699,1723]" country="Mexico" name="Guerrero" pageId="5" pageNumber="4">Guerrero</collectingRegion>
, 
<location id="8EA5609E9E1F66534D88F9136D13F960" LSID="urn:lsid:plazi:treatment:03D387539E1F66504EA0FD156CE8FAF6:8EA5609E9E1F66534D88F9136D13F960" box="[1000,1115,1699,1723]" country="Mexico" name="Acapulco" pageId="5" pageNumber="4" stateProvince="Guerrero">Acapulco</location>
, 
<date id="FFC410859E1F66534A06F9136C4EF960" box="[1126,1286,1699,1723]" pageId="5" pageNumber="4" value="1938-08-20">
<collectingDate id="EF80E96D9E1F66534A06F9136C4EF960" box="[1126,1286,1699,1723]" pageId="5" pageNumber="4" value="1938-08-20">20.VIII.1938</collectingDate>
</date>
, 
<collectorName id="268F53939E1F66534B70F9146CE8F960" box="[1296,1440,1699,1723]" pageId="5" pageNumber="4">L. Lipovsky</collectorName>
(
<collectionCode id="ED6BAE809E1F66534EA8F973685EF900" box="[200,278,1731,1755]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="5" pageNumber="4" type="Museum">FSCA</collectionCode>
); (2 ex)
</materialsCitation>
<materialsCitation id="3B123C189E1F66534F11F9736C7AF900" box="[369,1330,1731,1755]" collectingDate="1962-08-12" collectionCode="PESC" collectorName="U. Kans. Mex. Exped." country="Mexico" location="20 mi E Acapulco" pageId="5" pageNumber="4" specimenCount="1" stateProvince="Guerrero">
<collectingRegion id="49BEF8A79E1F66534F11F97368ADF900" box="[369,485,1731,1755]" country="Mexico" name="Guerrero" pageId="5" pageNumber="4">Guerrero</collectingRegion>
, 
<location id="8EA5609E9E1F66534F8FF9736B8DF900" LSID="urn:lsid:plazi:treatment:03D387539E1F66504EA0FD156CE8FAF6:8EA5609E9E1F66534F8FF9736B8DF900" box="[495,709,1731,1755]" country="Mexico" name="20 mi E Acapulco" pageId="5" pageNumber="4" stateProvince="Guerrero">
<quantity id="4C829BA09E1F66534F8FF9736B7BF900" box="[495,563,1731,1755]" metricMagnitude="4" metricUnit="m" metricValue="3.218688" pageId="5" pageNumber="4" unit="mi" value="20.0">20 mi</quantity>
E Acapulco
</location>
, 
<date id="FFC410859E1F66534CAFF9736A27F900" box="[719,879,1731,1755]" pageId="5" pageNumber="4" value="1962-08-12">
<collectingDate id="EF80E96D9E1F66534CAFF9736A27F900" box="[719,879,1731,1755]" pageId="5" pageNumber="4" value="1962-08-12">12.VIII.1962</collectingDate>
</date>
, 
<collectorName id="268F53939E1F66534D19F9746DCAF900" box="[889,1154,1731,1755]" pageId="5" pageNumber="4">U. Kans. Mex. Exped.</collectorName>
(
<collectionCode id="ED6BAE809E1F66534AEFF9736D94F900" box="[1167,1244,1731,1755]" pageId="5" pageNumber="4">PESC</collectionCode>
); (1 ex)
</materialsCitation>
<materialsCitation id="3B123C189E1F66534B58F9736A33F920" collectingDate="1953-09-17" collectingDateMax="1953-09-21" collectingDateMin="1953-09-17" collectionCode="FSCA" collectorName="B. Malkin" country="Mexico" location="San Blas" pageId="5" pageNumber="4" specimenCount="1" stateProvince="Nayarit">
<collectingRegion id="49BEF8A79E1F66534B58F9736CD4F900" box="[1336,1436,1731,1755]" country="Mexico" name="Nayarit" pageId="5" pageNumber="4">Nayarit</collectingRegion>
, 
<location id="8EA5609E9E1F66534EA0F9536866F920" LSID="urn:lsid:plazi:treatment:03D387539E1F66504EA0FD156CE8FAF6:8EA5609E9E1F66534EA0F9536866F920" box="[192,302,1763,1787]" country="Mexico" name="San Blas" pageId="5" pageNumber="4" stateProvince="Nayarit">San Blas</location>
, 
<date id="FFC410859E1F66534F58F95368AEF920" box="[312,486,1763,1787]" pageId="5" pageNumber="4" value="1953-09-17" valueMax="1953-09-21" valueMin="1953-09-17">
<collectingDate id="EF80E96D9E1F66534F58F95368AEF920" box="[312,486,1763,1787]" pageId="5" pageNumber="4" value="1953-09-17" valueMax="1953-09-21" valueMin="1953-09-17">17-21.IX.1953</collectingDate>
</date>
, at light, 
<collectorName id="268F53939E1F66534C35F9546B85F920" box="[597,717,1763,1787]" pageId="5" pageNumber="4">B. Malkin</collectorName>
(
<collectionCode id="ED6BAE809E1F66534CBBF9536A60F920" box="[731,808,1763,1787]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="5" pageNumber="4" type="Museum">FSCA</collectionCode>
); (7 ex)
</materialsCitation>
<materialsCitation id="3B123C189E1F66534DE1F9536889F8C0" collectingDate="1970-06-29" collectionCode="VI, PESC, ISEA" collectorName="P. M. Jump" country="Mexico" location="Salino Cruz" pageId="5" pageNumber="4" specimenCount="1" stateProvince="Oaxaca">
<collectingRegion id="49BEF8A79E1F66534DE1F9536A97F920" box="[897,991,1763,1787]" country="Mexico" name="Oaxaca" pageId="5" pageNumber="4">Oaxaca</collectingRegion>
, 
<location id="8EA5609E9E1F66534D88F9536D32F920" LSID="urn:lsid:plazi:treatment:03D387539E1F66504EA0FD156CE8FAF6:8EA5609E9E1F66534D88F9536D32F920" box="[1000,1146,1763,1787]" country="Mexico" name="Salino Cruz" pageId="5" pageNumber="4" stateProvince="Oaxaca">Salino Cruz</location>
, 
<date id="FFC410859E1F66534AE4F9536C44F920" box="[1156,1292,1763,1787]" pageId="5" pageNumber="4" value="1970-06-29">
<collectingDate id="EF80E96D9E1F66534AE4F9536C44F920" box="[1156,1292,1763,1787]" pageId="5" pageNumber="4" value="1970-06-29">
29. 
<collectionCode id="ED6BAE809E1F66534AC7F9536D81F920" box="[1191,1225,1763,1787]" country="Norway" name="Mykotektet, National Veterinary Institute" pageId="5" pageNumber="4">VI</collectionCode>
.1970
</collectingDate>
</date>
, 
<collectorName id="268F53939E1F66534B76F9546CE8F920" box="[1302,1440,1764,1787]" pageId="5" pageNumber="4">P. M. Jump</collectorName>
(
<collectionCode id="ED6BAE809E1F66534EA8F8B3685DF8C0" box="[200,277,1795,1819]" pageId="5" pageNumber="4">PESC</collectionCode>
, 
<collectionCode id="ED6BAE809E1F66534F42F8B36820F8C0" box="[290,360,1795,1819]" pageId="5" pageNumber="4">ISEA</collectionCode>
); (1 ex)
</materialsCitation>
<materialsCitation id="3B123C189E1F66534FA9F8B36DF6F8C0" box="[457,1214,1795,1819]" collectingDate="1972-07-02" collectionCode="FSCA" collectorName="K. Stephan" country="Mexico" location="Culiacan" pageId="5" pageNumber="4" specimenCount="1" stateProvince="Sinaloa">
<collectingRegion id="49BEF8A79E1F66534FA9F8B36B62F8C0" box="[457,554,1795,1819]" country="Mexico" name="Sinaloa" pageId="5" pageNumber="4">Sinaloa</collectingRegion>
, 
<location id="8EA5609E9E1F66534C56F8B36BE0F8C0" LSID="urn:lsid:plazi:treatment:03D387539E1F66504EA0FD156CE8FAF6:8EA5609E9E1F66534C56F8B36BE0F8C0" box="[566,680,1795,1819]" country="Mexico" name="Culiacan" pageId="5" pageNumber="4" stateProvince="Sinaloa">Culiacan</location>
, 
<date id="FFC410859E1F66534CD4F8B36A73F8C0" box="[692,827,1795,1819]" pageId="5" pageNumber="4" value="1972-07-02">
<collectingDate id="EF80E96D9E1F66534CD4F8B36A73F8C0" box="[692,827,1795,1819]" pageId="5" pageNumber="4" value="1972-07-02">2.VII.1972</collectingDate>
</date>
, 
<collectorName id="268F53939E1F66534D27F8B46A9BF8C0" box="[839,979,1795,1819]" pageId="5" pageNumber="4">K. Stephan</collectorName>
leg. (
<collectionCode id="ED6BAE809E1F66534A77F8B36D2DF8C0" box="[1047,1125,1795,1819]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="5" pageNumber="4" type="Museum">FSCA</collectionCode>
); (6 ex)
</materialsCitation>
<materialsCitation id="3B123C189E1F66534AA6F8B36D79F8E0" collectingDate="1965-08-10" collectionCode="FSCA" collectorName="G. H. Nelson" country="Mexico" location="1 mi S of Mazatlan" pageId="5" pageNumber="4" specimenCount="1" stateProvince="Sinaloa">
<collectingRegion id="49BEF8A79E1F66534AA6F8B36C6FF8C0" box="[1222,1319,1795,1819]" country="Mexico" name="Sinaloa" pageId="5" pageNumber="4">Sinaloa</collectingRegion>
, 
<location id="8EA5609E9E1F66534B53F8B3687FF8E0" LSID="urn:lsid:plazi:treatment:03D387539E1F66504EA0FD156CE8FAF6:8EA5609E9E1F66534B53F8B3687FF8E0" country="Mexico" name="1 mi S of Mazatlan" pageId="5" pageNumber="4" stateProvince="Sinaloa">
<quantity id="4C829BA09E1F66534B53F8B36C21F8C0" box="[1331,1385,1795,1819]" metricMagnitude="3" metricUnit="m" metricValue="1.609344" pageId="5" pageNumber="4" unit="mi" value="1.0">1 mi</quantity>
S of Mazatlan
</location>
, 
<date id="FFC410859E1F66534F21F8936897F8E0" box="[321,479,1827,1851]" pageId="5" pageNumber="4" value="1965-08-10">
<collectingDate id="EF80E96D9E1F66534F21F8936897F8E0" box="[321,479,1827,1851]" pageId="5" pageNumber="4" value="1965-08-10">10.VIII.1965</collectingDate>
</date>
, collection 
<collectorName id="268F53939E1F66534C00F8936BB4F8E0" box="[608,764,1827,1851]" pageId="5" pageNumber="4">G. H. Nelson</collectorName>
, at lite [sic] (
<collectionCode id="ED6BAE809E1F66534DF1F8936A96F8E0" box="[913,990,1827,1851]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="5" pageNumber="4" type="Museum">FSCA</collectionCode>
); (1 ex)
</materialsCitation>
<materialsCitation id="3B123C189E1F66534A57F893688EF880" collectingDate="1952-07-31" collectionCode="FSCA" collectorName="C. &amp; P. Vaurie" country="Mexico" location="Yavaros" pageId="5" pageNumber="4" specimenCount="1" stateProvince="Sonora">
<collectingRegion id="49BEF8A79E1F66534A57F8936DD8F8E0" box="[1079,1168,1827,1851]" country="Mexico" name="Sonora" pageId="5" pageNumber="4">Sonora</collectingRegion>
, 
<location id="8EA5609E9E1F66534AF9F8946DB7F8E0" LSID="urn:lsid:plazi:treatment:03D387539E1F66504EA0FD156CE8FAF6:8EA5609E9E1F66534AF9F8946DB7F8E0" box="[1177,1279,1828,1851]" country="Mexico" name="Yavaros" pageId="5" pageNumber="4" stateProvince="Sonora">Yavaros</location>
, 
<date id="FFC410859E1F66534B69F8936CD4F8E0" box="[1289,1436,1827,1851]" pageId="5" pageNumber="4" value="1952-07-31">
<collectingDate id="EF80E96D9E1F66534B69F8936CD4F8E0" box="[1289,1436,1827,1851]" pageId="5" pageNumber="4" value="1952-07-31">31.VII.1952</collectingDate>
</date>
, 
<collectorName id="268F53939E1F66534EA0F8F3682BF880" box="[192,355,1859,1883]" pageId="5" pageNumber="4">C.&amp;P. Vaurie</collectorName>
(
<collectionCode id="ED6BAE809E1F66534F10F8F368F6F880" box="[368,446,1859,1883]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="5" pageNumber="4" type="Museum">FSCA</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="DF0566CD9E1C66504EA0FC6D68FBFBCA" ID-DOI="http://doi.org/10.5281/zenodo.4531519" ID-Zenodo-Dep="4531519" httpUri="https://zenodo.org/record/4531519/files/figure.png" pageId="6" pageNumber="5" startId="6.[192,287,989,1011]" targetBox="[192,1443,196,958]" targetPageId="6">
<paragraph id="8BC536459E1C66504EA0FC6D68FBFBCA" blockId="6.[192,1440,989,1041]" pageId="6" pageNumber="5">
<emphasis id="B90EEA579E1C66504EA0FC6D6814FC28" bold="true" box="[192,348,989,1011]" pageId="6" pageNumber="5">Figures 8-11.</emphasis>
<taxonomicName id="4C7A4DC69E1C66504F01FC6E6B5EFC28" authorityName="Chalumeau" authorityYear="1981" box="[353,534,990,1011]" class="Insecta" family="Aphodiidae" genus="Haroldiataenius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="genus">
<emphasis id="B90EEA579E1C66504F01FC6E6B5EFC28" box="[353,534,990,1011]" italics="true" pageId="6" pageNumber="5">Haroldiataenius</emphasis>
</taxonomicName>
spp., lateral habitus. 
<emphasis id="B90EEA579E1C66504D63FC6D6A51FC28" bold="true" box="[771,793,989,1011]" pageId="6" pageNumber="5">8)</emphasis>
<taxonomicName id="4C7A4DC69E1C66504D7FFC6E6AF6FC28" authorityName="Stebnicka &amp; Skelley" authorityYear="2009" baseAuthorityName="Stebnicka &amp; Skelley" baseAuthorityYear="2009" box="[799,958,990,1011]" class="Insecta" family="Aphodiidae" genus="Haroldiataenius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="semipilosus">
<emphasis id="B90EEA579E1C66504D7FFC6E6AF6FC28" box="[799,958,990,1011]" italics="true" pageId="6" pageNumber="5">H. semipilosus</emphasis>
</taxonomicName>
. 
<emphasis id="B90EEA579E1C66504DAAFC6D6AA9FC28" bold="true" box="[970,993,989,1011]" pageId="6" pageNumber="5">9)</emphasis>
<taxonomicName id="4C7A4DC69E1C66504D86FC6E6D1CFC28" authorityName="Stebnicka &amp; Skelley" authorityYear="2009" baseAuthorityName="Stebnicka &amp; Skelley" baseAuthorityYear="2009" box="[998,1108,990,1011]" class="Insecta" family="Aphodiidae" genus="Haroldiataenius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="griffini">
<emphasis id="B90EEA579E1C66504D86FC6E6D1CFC28" box="[998,1108,990,1011]" italics="true" pageId="6" pageNumber="5">H. griffini</emphasis>
</taxonomicName>
. 
<emphasis id="B90EEA579E1C66504A00FC6D6DCCFC28" bold="true" box="[1120,1156,989,1011]" pageId="6" pageNumber="5">10)</emphasis>
<taxonomicName id="4C7A4DC69E1C66504AE9FC6E6C49FC28" authorityName="Stebnicka &amp; Skelley" authorityYear="2009" box="[1161,1281,990,1011]" class="Insecta" family="Aphodiidae" genus="Haroldiataenius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="buvexus">
<emphasis id="B90EEA579E1C66504AE9FC6E6C49FC28" box="[1161,1281,990,1011]" italics="true" pageId="6" pageNumber="5">H. buvexus</emphasis>
</taxonomicName>
, paratype. 
<emphasis id="B90EEA579E1C66504B1BFC6D6CE8FC28" bold="true" box="[1403,1440,989,1011]" pageId="6" pageNumber="5">11)</emphasis>
<taxonomicName id="4C7A4DC69E1C66504EA0FC4B680AFBCB" authorityName="Stebnicka &amp; Skelley" authorityYear="2009" baseAuthorityName="Stebnicka &amp; Skelley" baseAuthorityYear="2009" box="[192,322,1019,1040]" class="Insecta" family="Aphodiidae" genus="Haroldiataenius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="convexus">
<emphasis id="B90EEA579E1C66504EA0FC4B680AFBCB" box="[192,322,1019,1040]" italics="true" pageId="6" pageNumber="5">H. convexus</emphasis>
</taxonomicName>
, holotype.
</paragraph>
</caption>
<subSubSection id="C36065CE9E1C66504EA0FB856CE8FAF6" pageId="6" pageNumber="5" type="discussion">
<paragraph id="8BC536459E1C66504EA0FB856CE8FAF6" blockId="6.[192,1440,1077,1326]" pageId="6" pageNumber="5">
<emphasis id="B90EEA579E1C66504EA0FB856875FB96" bold="true" box="[192,317,1077,1101]" pageId="6" pageNumber="5">Remarks</emphasis>
. 
<taxonomicName id="4C7A4DC69E1C66504F2BFB856BD6FB96" baseAuthorityName="Bates" baseAuthorityYear="1887" box="[331,670,1077,1101]" class="Insecta" family="Aphodiidae" genus="Haroldiataenius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="mariarum">
<emphasis id="B90EEA579E1C66504F2BFB856BD6FB96" box="[331,670,1077,1101]" italics="true" pageId="6" pageNumber="5">Haroldiataenius mariarum</emphasis>
</taxonomicName>
is readily distinguished from other species of the genus by the long setae along the lateral margin of the pronotum and elytra, distinctly wrinkled clypeus, strongly dilated apex of meso- and metatibiae, and the basal meso- and metatarsomere being distinctly shorter than the long tibial spur. The species is similar in appearance to the psammodiine species 
<taxonomicName id="4C7A4DC69E1C66504AAEFB256B12FB16" authority="(Horn) (Verdu et al. 2006)" authorityName="Verdu et al." authorityYear="2006" baseAuthorityName="Horn" baseAuthorityYear="1871" class="Insecta" family="Aphodiidae" genus="Parapsammodius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="bidens">
<emphasis id="B90EEA579E1C66504AAEFB256859FB16" italics="true" pageId="6" pageNumber="5">Parapsammodius bidens</emphasis>
(Horn) (
<bibRefCitation id="EFEB4BB49E1C66504F19FB056B1BFB16" author="Verdu, J. R. &amp; Z. T. Stebnicka &amp; E. Galante" box="[377,595,1205,1229]" pageId="6" pageNumber="5" pagination="1 - 2" refId="ref11346" refString="Verdu, J. R., Z. T. Stebnicka, and E. Galante. 2006. A new Neotropical genus of the Eupariini- Psammodiini complex with comparative morphology of mouthparts structures and analysis of char- acters among related taxa (Coleoptera: Scarabaeidae: Aphodiinae). Acta Zoologica Cracoviensia 49 B (1 - 2): 55 - 72." type="journal article" year="2006">Verdú et al. 2006</bibRefCitation>
)
</taxonomicName>
, sharing with that species the strongly expanded metatibia with an external row of granules and similar epipharyngeal structures. In general, 
<taxonomicName id="4C7A4DC69E1C66504A24FB666DAEFB36" baseAuthorityName="Bates" baseAuthorityYear="1887" box="[1092,1254,1238,1261]" class="Insecta" family="Aphodiidae" genus="Haroldiataenius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="mariarum">
<emphasis id="B90EEA579E1C66504A24FB666DAEFB36" box="[1092,1254,1238,1261]" italics="true" pageId="6" pageNumber="5">H. mariarum</emphasis>
</taxonomicName>
fits the diagnosis given by 
<bibRefCitation id="EFEB4BB49E1C66504F2FFB456B65FAD6" author="Chalumeau, F." box="[335,557,1269,1293]" pageId="6" pageNumber="5" pagination="137 - 142" refId="ref10805" refString="Chalumeau, F. 1981. Nouveaux genre et sous-genres d'Eupariini (Aphodiinae) d'Amerique tropicale. Nouvelle Revue d'Entomologie 11 (2): 137 - 142." type="journal article" year="1981">Chalumeau (1981)</bibRefCitation>
for 
<emphasis id="B90EEA579E1C66504C39FB456B88FAD6" box="[601,704,1269,1293]" italics="true" pageId="6" pageNumber="5">Sayloria</emphasis>
, except that the lateral setae are much longer and the clypeus is distinctly wrinkled. These characters are variable among the species placed herein in 
<emphasis id="B90EEA579E1C66504AB2FAA56CE8FAF6" box="[1234,1440,1301,1325]" italics="true" pageId="6" pageNumber="5">
<taxonomicName id="4C7A4DC69E1C66504AB2FAA56CD4FAF6" authorityName="Chalumeau" authorityYear="1981" box="[1234,1436,1301,1325]" class="Insecta" family="Aphodiidae" genus="Haroldiataenius" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="genus">Haroldiataenius</taxonomicName>
.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>