215 lines
30 KiB
XML
215 lines
30 KiB
XML
<document id="A3A61DEC00B779CBC0ABECD34EDB1A81" ID-CLB-Dataset="26407" ID-DOI="10.11646/zootaxa.4623.2.8" ID-GBIF-Dataset="db2b3bd4-2e27-4918-8fff-226b303c65b9" ID-ISSN="1175-5326" ID-Zenodo-Dep="3255499" ID-ZooBank="EEDC9982-ED9C-40BD-8584-CDADA572DDE" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1561449672174" checkinUser="plazi" docAuthor="Bocek, Matej & Adamkova, Kristyna" docDate="2019" docId="03AF335EC3666356FF06F99AFB3BFCFC" docLanguage="en" docName="zootaxa.4623.2.8.pdf" docOrigin="Zootaxa 4623 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Schizotrichalus Kleine 1926" docType="treatment" docVersion="9" lastPageNumber="344" masterDocId="FF964B26C3646355FF91FF95FFD3FFEB" masterDocTitle="New species of Moluccan trichaline net-winged beetles, with remarks on the phylogenetic position and distribution of Schizotrichalus (Coleoptera: Lycidae: Metriorrhynchinae)" masterLastPageNumber="350" masterPageNumber="341" pageNumber="343" updateTime="1698736235721" updateUser="ExternalLinkService">
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<mods:title id="4A3FCD1BE449E9CC1C7340663A2EFA18">New species of Moluccan trichaline net-winged beetles, with remarks on the phylogenetic position and distribution of Schizotrichalus (Coleoptera: Lycidae: Metriorrhynchinae)</mods:title>
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<mods:namePart id="861009EDB5676A8D62EA0AC220CD6529">Bocek, Matej</mods:namePart>
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<mods:namePart id="51F88D8722D0027DDD7E6F9C7EECB18A">Adamkova, Kristyna</mods:namePart>
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<mods:title id="3BD09D92968F59513B52161D2633CC35">Zootaxa</mods:title>
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<mods:part id="AA57E663D7633AA66BECAC7A45C9EE1B">
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<mods:date id="E223CF01ED972F0E81ED3FB73FF1180A">2019</mods:date>
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<mods:number id="D07AB1A5C7DA693EF92CE4AE2D43783E">2019-06-25</mods:number>
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<mods:number id="74D238F312F2AAAD0B6C5F125DAC1ED8">4623</mods:number>
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<subSubSection id="C31CD1C3C3666357FF06F99AFE26F9C2" box="[151,501,1551,1578]" pageId="2" pageNumber="343" type="nomenclature">
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<paragraph id="8BB98248C3666357FF06F99AFE26F9C2" blockId="2.[151,501,1551,1578]" box="[151,501,1551,1578]" pageId="2" pageNumber="343">
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<heading id="D0F13524C3666357FF06F99AFE26F9C2" bold="true" box="[151,501,1551,1578]" fontSize="11" level="1" pageId="2" pageNumber="343" reason="1">
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<taxonomicName id="4C06F9CBC3666357FF06F99AFE26F9C2" authority="Kleine, 1926" authorityName="Kleine" authorityYear="1926" box="[151,501,1551,1578]" class="Insecta" family="Lycidae" genus="Schizotrichalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
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<emphasis id="B9725E5AC3666357FF06F99AFE26F9C2" bold="true" box="[151,501,1551,1578]" pageId="2" pageNumber="343">
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<emphasis id="B9725E5AC3666357FF06F99AFE81F9C1" bold="true" box="[151,338,1551,1578]" italics="true" pageId="2" pageNumber="343">Schizotrichalus</emphasis>
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<bibRefCitation id="EF97FFB9C3666357FECBF99AFE26F9C2" author="Kleine, R." box="[346,501,1551,1577]" pageId="2" pageNumber="343" pagination="91 - 195" refId="ref5945" refString="Kleine, R. (1926) Coleoptera, Lycidae. Nova Guinea. Resultats de l'expedition scientifique neerlandaise a la Nouvelle-Guinee, 15, 91 - 195." type="journal article" year="1926">Kleine, 1926</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<subSubSection id="C31CD1C3C3666357FF06F9C1FE24F987" box="[151,503,1619,1644]" pageId="2" pageNumber="343" type="reference_group">
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<paragraph id="8BB98248C3666357FF06F9C1FE24F987" blockId="2.[151,998,1619,1675]" box="[151,503,1619,1644]" pageId="2" pageNumber="343">
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<treatmentCitationGroup id="AB16A566C3666357FF06F9C1FE24F987" box="[151,503,1619,1644]" pageId="2" pageNumber="343">
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<taxonomicName id="4C06F9CBC3666357FF06F9C1FE20F987" authority="Kleine, 1926: 183" authorityName="Kleine" authorityPageNumber="183" authorityYear="1926" box="[151,499,1619,1644]" class="Insecta" family="Lycidae" genus="Schizotrichalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
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<emphasis id="B9725E5AC3666357FF06F9C1FEE0F987" box="[151,307,1620,1644]" italics="true" pageId="2" pageNumber="343">Schizotrichalus</emphasis>
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<treatmentCitation id="0AA7A459C3666357FEABF9C6FE20F987" author="Kleine, R." box="[314,499,1619,1644]" page="183" pageId="2" pageNumber="343" year="1926">
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<bibRefCitation id="EF97FFB9C3666357FEABF9C6FE20F987" author="Kleine, R." box="[314,499,1619,1644]" pageId="2" pageNumber="343" pagination="91 - 195" refId="ref5945" refString="Kleine, R. (1926) Coleoptera, Lycidae. Nova Guinea. Resultats de l'expedition scientifique neerlandaise a la Nouvelle-Guinee, 15, 91 - 195." type="journal article" year="1926">Kleine, 1926: 183</bibRefCitation>
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</treatmentCitation>
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</taxonomicName>
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.
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection id="C31CD1C3C3666357FF06F9E7FC35F960" box="[151,998,1650,1675]" pageId="2" pageNumber="343" type="materials_examined">
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<paragraph id="8BB98248C3666357FF06F9E7FC35F960" blockId="2.[151,998,1619,1675]" box="[151,998,1650,1675]" pageId="2" pageNumber="343">
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<emphasis id="B9725E5AC3666357FF06F9E7FEF0F960" bold="true" box="[151,291,1650,1675]" pageId="2" pageNumber="343">
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<typeStatus id="54BD3CEAC3666357FF06F9E7FF1FF961" box="[151,204,1650,1674]" pageId="2" pageNumber="343">Type</typeStatus>
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species.
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</emphasis>
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<taxonomicName id="4C06F9CBC3666357FEBBF9E7FD0DF961" authority="Waterhouse, 1879: 70" authorityName="Waterhouse" authorityPageNumber="70" authorityYear="1879" box="[298,734,1650,1675]" class="Insecta" family="Lycidae" genus="Trichalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="343" phylum="Arthropoda" rank="species" species="nigrescens">
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<emphasis id="B9725E5AC3666357FEBBF9E7FE28F961" box="[298,507,1650,1674]" italics="true" pageId="2" pageNumber="343">Trichalus nigrescens</emphasis>
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<bibRefCitation id="EF97FFB9C3666357FD90F9E7FD0DF961" author="Waterhouse, C. O." box="[513,734,1650,1675]" pageId="2" pageNumber="343" refId="ref7129" refString="Waterhouse, C. O. (1879) Illustration of the typical specimens of Coleoptera in the collection of the British Museum. Part I. - Lycidae. British Museum, London, 93 pp." type="book" year="1879">Waterhouse, 1879: 70</bibRefCitation>
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</taxonomicName>
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(by original designation).
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</paragraph>
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</subSubSection>
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<subSubSection id="C31CD1C3C3666357FF06F92DFC54F840" pageId="2" pageNumber="343" type="diagnosis">
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<paragraph id="8BB98248C3666357FF06F92DFC54F840" blockId="2.[151,1437,1720,2035]" pageId="2" pageNumber="343">
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<emphasis id="B9725E5AC3666357FF06F92DFEC1F938" bold="true" box="[151,274,1720,1747]" pageId="2" pageNumber="343">Diagnosis.</emphasis>
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<taxonomicName id="4C06F9CBC3666357FE8AF92CFE1BF938" authorityName="Kleine" authorityYear="1926" box="[283,456,1721,1747]" class="Insecta" family="Lycidae" genus="Schizotrichalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
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<emphasis id="B9725E5AC3666357FE8AF92CFE1BF938" box="[283,456,1721,1747]" italics="true" pageId="2" pageNumber="343">Schizotrichalus</emphasis>
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</taxonomicName>
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shares the similar shortened elytral primary costa 1 with other trichaline net-winged beetles (
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<emphasis id="B9725E5AC3666357FF67F94BFEC1F91C" box="[246,274,1758,1783]" italics="true" pageId="2" pageNumber="343">i.e</emphasis>
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.,
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<taxonomicName id="4C06F9CBC3666357FEB6F948FE66F91C" authorityName="Kleine" authorityYear="1926" box="[295,437,1757,1783]" class="Insecta" family="Lycidae" genus="Diatrichalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
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<emphasis id="B9725E5AC3666357FEB6F948FE66F91C" box="[295,437,1757,1783]" italics="true" pageId="2" pageNumber="343">Diatrichalus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C06F9CBC3666357FE55F948FD55F91C" authorityName="Pic" authorityYear="1921" box="[452,646,1757,1783]" class="Insecta" family="Lycidae" genus="Flabellotrichalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
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<emphasis id="B9725E5AC3666357FE55F948FD55F91C" box="[452,646,1757,1783]" italics="true" pageId="2" pageNumber="343">Flabellotrichalus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C06F9CBC3666357FD04F948FD2CF91C" authorityName="Waterhouse" authorityYear="1877" box="[661,767,1757,1783]" class="Insecta" family="Lycidae" genus="Trichalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
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<emphasis id="B9725E5AC3666357FD04F948FD2CF91C" box="[661,767,1757,1783]" italics="true" pageId="2" pageNumber="343">Trichalus</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C06F9CBC3666357FC9CF948FCABF91C" authorityName="Waterhouse" authorityYear="1879" box="[781,888,1757,1783]" class="Insecta" family="Lycidae" genus="Eniclases" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
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<emphasis id="B9725E5AC3666357FC9CF948FCABF91C" box="[781,888,1757,1783]" italics="true" pageId="2" pageNumber="343">Eniclases</emphasis>
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</taxonomicName>
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,
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<taxonomicName id="4C06F9CBC3666357FC19F948FC20F91C" authorityName="Bocak" authorityYear="1998" box="[904,1011,1757,1783]" class="Insecta" family="Lycidae" genus="Lobatang" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
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<emphasis id="B9725E5AC3666357FC19F948FC20F91C" box="[904,1011,1757,1783]" italics="true" pageId="2" pageNumber="343">Lobatang</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C06F9CBC3666357FBBAF948FB01F91C" authorityName="Pic" authorityYear="1921" box="[1067,1234,1757,1783]" class="Insecta" family="Lycidae" genus="Microtrichalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
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<emphasis id="B9725E5AC3666357FBBAF948FB01F91C" box="[1067,1234,1757,1783]" italics="true" pageId="2" pageNumber="343">Microtrichalus</emphasis>
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</taxonomicName>
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; general morphological description of all trichaline genera was published by
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<bibRefCitation id="EF97FFB9C3666357FCDBF894FC35F8F0" author="Bocak, L." box="[842,998,1793,1819]" pageId="2" pageNumber="343" pagination="315 - 351" refId="ref5210" refString="Bocak, L. (2002) Revision and phylogenetic analysis of Metriorrhynchinae. European Journal of Entomology, 99, 315 - 351. https: // doi. org / 10.14411 / eje. 2002.043" type="journal article" year="2002">Bocak (2002)</bibRefCitation>
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and
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<bibRefCitation id="EF97FFB9C3666357FB8EF894FAE5F8F0" author="Bocek, M. & Bocak, L." box="[1055,1334,1793,1819]" pageId="2" pageNumber="343" pagination="3963" refId="ref5383" refString="Bocek, M. & Bocak, L. (2017) The comparison of molecular and morphology-based phylogenies of trichaline net-winged beetles (Coleoptera: Lycidae: Metriorrhynchini) with description of a new subgenus. PeerJ, 5, e 3963. https: // doi. org / 10.7717 / peerj. 3963" type="journal article" year="2017">Bocek & Bocak (2017))</bibRefCitation>
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.
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<taxonomicName id="4C06F9CBC3666357FAD3F894FF20F8D4" authorityName="Kleine" authorityYear="1926" class="Insecta" family="Lycidae" genus="Schizotrichalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
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<emphasis id="B9725E5AC3666357FAD3F894FF20F8D4" italics="true" pageId="2" pageNumber="343">Schizotrichalus</emphasis>
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</taxonomicName>
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differs from other trichalines in having five well-developed pronotal areoles unlike other trichalines (
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<figureCitation id="133D9ECDC3666357FAFBF8B0FF16F88F" captionStart="FIGURES 2–16" captionStartId="5.[151,264,1674,1699]" captionTargetBox="[153,1429,647,1644]" captionTargetId="figure@5.[151,1436,669,1654]" captionTargetPageId="5" captionText="FIGURES 2–16. General appearance and diagnostic characters used for the species delimitation. (2–5) General appearance. (2) Schizotrichalus halmaherensis sp. nov., male; (3) ditto, female; (4) Eniclases kusyi sp. nov., male; (5) E. moluccanus, male. (6–9) Pronota (6) S. halmaherensis sp. nov., male; (7) ditto, female; (8) E. moluccanus, male; (9) E. kusyi sp. nov., male. (10–12) Male genitalia (10) S. halmaherensis sp. nov.; (11) E. kusyi sp. nov.; (12) E. moluccanus; (13–16) Basal antennomeres (13) S. halmaherensis sp. nov., male; (14) ditto, female; (15) E. kusyi sp. nov., male; (16) E. moluccanus, male. Scale bars (2–5) 2 mm; (6–16) 0.5 mm." httpUri="https://zenodo.org/record/3255503/files/figure.png" pageId="2" pageNumber="343">Figs 6–7</figureCitation>
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). Similar to
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<taxonomicName id="4C06F9CBC3666357FEC9F8DCFE10F888" authorityName="Waterhouse" authorityYear="1879" box="[344,451,1865,1891]" class="Insecta" family="Lycidae" genus="Eniclases" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
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||
<emphasis id="B9725E5AC3666357FEC9F8DCFE10F888" box="[344,451,1865,1891]" italics="true" pageId="2" pageNumber="343">Eniclases</emphasis>
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</taxonomicName>
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, the males of
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<taxonomicName id="4C06F9CBC3666357FDFDF8DCFCCAF888" authorityName="Kleine" authorityYear="1926" box="[620,793,1865,1891]" class="Insecta" family="Lycidae" genus="Schizotrichalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
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<emphasis id="B9725E5AC3666357FDFDF8DCFCCAF888" box="[620,793,1865,1891]" italics="true" pageId="2" pageNumber="343">Schizotrichalus</emphasis>
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</taxonomicName>
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have either acutely serrate or flabellate antennae. Male genitalia of
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<taxonomicName id="4C06F9CBC3666357FE8EF8F8FE1FF86C" authorityName="Kleine" authorityYear="1926" box="[287,460,1901,1927]" class="Insecta" family="Lycidae" genus="Schizotrichalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="343" phylum="Arthropoda" rank="genus">
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<emphasis id="B9725E5AC3666357FE8EF8F8FE1FF86C" box="[287,460,1901,1927]" italics="true" pageId="2" pageNumber="343">Schizotrichalus</emphasis>
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</taxonomicName>
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have a long, slender phallus with a cup-like apex and pigmented dorsal line (
|
||
<figureCitation id="133D9ECDC3666357FAABF8F8FA5CF86C" box="[1338,1423,1901,1927]" captionStart="FIGURES 2–16" captionStartId="5.[151,264,1674,1699]" captionTargetBox="[153,1429,647,1644]" captionTargetId="figure@5.[151,1436,669,1654]" captionTargetPageId="5" captionText="FIGURES 2–16. General appearance and diagnostic characters used for the species delimitation. (2–5) General appearance. (2) Schizotrichalus halmaherensis sp. nov., male; (3) ditto, female; (4) Eniclases kusyi sp. nov., male; (5) E. moluccanus, male. (6–9) Pronota (6) S. halmaherensis sp. nov., male; (7) ditto, female; (8) E. moluccanus, male; (9) E. kusyi sp. nov., male. (10–12) Male genitalia (10) S. halmaherensis sp. nov.; (11) E. kusyi sp. nov.; (12) E. moluccanus; (13–16) Basal antennomeres (13) S. halmaherensis sp. nov., male; (14) ditto, female; (15) E. kusyi sp. nov., male; (16) E. moluccanus, male. Scale bars (2–5) 2 mm; (6–16) 0.5 mm." httpUri="https://zenodo.org/record/3255503/files/figure.png" pageId="2" pageNumber="343">Fig. 10</figureCitation>
|
||
). The phallobase is small and without an extensive membranous sac.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31CD1C3C3666356FF56F821FC5DFE44" lastPageId="3" lastPageNumber="344" pageId="2" pageNumber="343" type="description">
|
||
<paragraph id="8BB98248C3666356FF56F821FC5DFE44" blockId="2.[151,1437,1720,2035]" lastBlockId="3.[151,1437,153,791]" lastPageId="3" lastPageNumber="344" pageId="2" pageNumber="343">
|
||
<emphasis id="B9725E5AC3666357FF56F821FEA6F824" bold="true" box="[199,373,1972,1999]" pageId="2" pageNumber="343">Redescription.</emphasis>
|
||
Body
|
||
<quantity id="4CFE2FADC3666357FE54F821FD88F824" box="[453,603,1972,1999]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" metricValueMax="11.5" metricValueMin="6.5" pageId="2" pageNumber="343" unit="mm" value="9.0" valueMax="11.5" valueMin="6.5">6.5–11.5 mm</quantity>
|
||
long, coloration variable from orange to brown and black; dark colored or pronotum and humeral part of elytra orange and apical part of elytra brown to black (
|
||
<figureCitation id="133D9ECDC3666357FB04F84CFB2DF818" box="[1173,1278,2009,2035]" captionStart="FIGURES 2–16" captionStartId="5.[151,264,1674,1699]" captionTargetBox="[153,1429,647,1644]" captionTargetId="figure@5.[151,1436,669,1654]" captionTargetPageId="5" captionText="FIGURES 2–16. General appearance and diagnostic characters used for the species delimitation. (2–5) General appearance. (2) Schizotrichalus halmaherensis sp. nov., male; (3) ditto, female; (4) Eniclases kusyi sp. nov., male; (5) E. moluccanus, male. (6–9) Pronota (6) S. halmaherensis sp. nov., male; (7) ditto, female; (8) E. moluccanus, male; (9) E. kusyi sp. nov., male. (10–12) Male genitalia (10) S. halmaherensis sp. nov.; (11) E. kusyi sp. nov.; (12) E. moluccanus; (13–16) Basal antennomeres (13) S. halmaherensis sp. nov., male; (14) ditto, female; (15) E. kusyi sp. nov., male; (16) E. moluccanus, male. Scale bars (2–5) 2 mm; (6–16) 0.5 mm." httpUri="https://zenodo.org/record/3255503/files/figure.png" pageId="2" pageNumber="343">Figs 2–3</figureCitation>
|
||
). Head small, mostly covered by pronotum; without rostrum, hypognathous; eyes hemispherically prominent. Mouthparts welldeveloped, maxillary palpi slender, four-segmented, apical palpomere widest, mandibles short and robust. Antennae compressed, acutely serrate or flabellate with long lamellae in males, always serrate in females. Pronotum trapezoidal, wider than long, with one median areola apically reaching two third of pronotal length and frontally attached to pronotal margin by robust carina, two lateral carinae well-developed, attached to frontal pronotal margin (
|
||
<figureCitation id="133D9ECDC3676356FAFAFEBCFF16FE83" captionStart="FIGURES 2–16" captionStartId="5.[151,264,1674,1699]" captionTargetBox="[153,1429,647,1644]" captionTargetId="figure@5.[151,1436,669,1654]" captionTargetPageId="5" captionText="FIGURES 2–16. General appearance and diagnostic characters used for the species delimitation. (2–5) General appearance. (2) Schizotrichalus halmaherensis sp. nov., male; (3) ditto, female; (4) Eniclases kusyi sp. nov., male; (5) E. moluccanus, male. (6–9) Pronota (6) S. halmaherensis sp. nov., male; (7) ditto, female; (8) E. moluccanus, male; (9) E. kusyi sp. nov., male. (10–12) Male genitalia (10) S. halmaherensis sp. nov.; (11) E. kusyi sp. nov.; (12) E. moluccanus; (13–16) Basal antennomeres (13) S. halmaherensis sp. nov., male; (14) ditto, female; (15) E. kusyi sp. nov., male; (16) E. moluccanus, male. Scale bars (2–5) 2 mm; (6–16) 0.5 mm." httpUri="https://zenodo.org/record/3255503/files/figure.png" pageId="3" pageNumber="344">Figs 6–7</figureCitation>
|
||
). Elytral primary costa 1 shortened, secondary costae well-developed with regular transverse costae. Legs compressed, moderately long. Male genitalia without parameres, phallus long, subtle, mostly membranous, only basally well-sclerotized, with dorsal strengthened pigmented keel (
|
||
<figureCitation id="133D9ECDC3676356FCBCFE00FC52FE44" box="[813,897,405,431]" captionStart="FIGURES 2–16" captionStartId="5.[151,264,1674,1699]" captionTargetBox="[153,1429,647,1644]" captionTargetId="figure@5.[151,1436,669,1654]" captionTargetPageId="5" captionText="FIGURES 2–16. General appearance and diagnostic characters used for the species delimitation. (2–5) General appearance. (2) Schizotrichalus halmaherensis sp. nov., male; (3) ditto, female; (4) Eniclases kusyi sp. nov., male; (5) E. moluccanus, male. (6–9) Pronota (6) S. halmaherensis sp. nov., male; (7) ditto, female; (8) E. moluccanus, male; (9) E. kusyi sp. nov., male. (10–12) Male genitalia (10) S. halmaherensis sp. nov.; (11) E. kusyi sp. nov.; (12) E. moluccanus; (13–16) Basal antennomeres (13) S. halmaherensis sp. nov., male; (14) ditto, female; (15) E. kusyi sp. nov., male; (16) E. moluccanus, male. Scale bars (2–5) 2 mm; (6–16) 0.5 mm." httpUri="https://zenodo.org/record/3255503/files/figure.png" pageId="3" pageNumber="344">Fig. 10</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31CD1C3C3676356FF56FE2DFD12FE1C" pageId="3" pageNumber="344" type="distribution">
|
||
<paragraph id="8BB98248C3676356FF56FE2DFD12FE1C" blockId="3.[151,1437,153,791]" pageId="3" pageNumber="344">
|
||
<emphasis id="B9725E5AC3676356FF56FE2DFDD1FE38" bold="true" box="[199,514,440,467]" pageId="3" pageNumber="344">Distribution and diversity.</emphasis>
|
||
Until now,
|
||
<taxonomicName id="4C06F9CBC3676356FD14FE2CFCE1FE38" authorityName="Kleine" authorityYear="1926" box="[645,818,441,467]" class="Insecta" family="Lycidae" genus="Schizotrichalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9725E5AC3676356FD14FE2CFCE1FE38" box="[645,818,441,467]" italics="true" pageId="3" pageNumber="344">Schizotrichalus</emphasis>
|
||
</taxonomicName>
|
||
has been recorded only from New
|
||
<collectingCountry id="F311C2D8C3676356FB2BFE2CFADCFE38" box="[1210,1295,441,467]" name="Guinea-Bissau" pageId="3" pageNumber="344">Guinea</collectingCountry>
|
||
. For the first time, we record
|
||
<taxonomicName id="4C06F9CBC3676356FEDCFE48FE29FE1C" authorityName="Kleine" authorityYear="1926" box="[333,506,477,503]" class="Insecta" family="Lycidae" genus="Schizotrichalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9725E5AC3676356FEDCFE48FE29FE1C" box="[333,506,477,503]" italics="true" pageId="3" pageNumber="344">Schizotrichalus</emphasis>
|
||
</taxonomicName>
|
||
from Halmahera.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C31CD1C3C3676356FF56FD94FB3BFCFC" pageId="3" pageNumber="344" type="discussion">
|
||
<paragraph id="8BB98248C3676356FF56FD94FB3BFCFC" blockId="3.[151,1437,153,791]" pageId="3" pageNumber="344">
|
||
<emphasis id="B9725E5AC3676356FF56FD94FEE3FDF0" bold="true" box="[199,304,513,539]" pageId="3" pageNumber="344">Remark.</emphasis>
|
||
The phylogenetic placement in trichalines and close relationships of
|
||
<taxonomicName id="4C06F9CBC3676356FBDBFD94FB24FDF0" authorityName="Kleine" authorityYear="1926" box="[1098,1271,513,539]" class="Insecta" family="Lycidae" genus="Schizotrichalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9725E5AC3676356FBDBFD94FB24FDF0" box="[1098,1271,513,539]" italics="true" pageId="3" pageNumber="344">Schizotrichalus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C06F9CBC3676356FAA0FD94FA4FFDF0" authorityName="Waterhouse" authorityYear="1879" box="[1329,1436,513,539]" class="Insecta" family="Lycidae" genus="Eniclases" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9725E5AC3676356FAA0FD94FA4FFDF0" box="[1329,1436,513,539]" italics="true" pageId="3" pageNumber="344">Eniclases</emphasis>
|
||
</taxonomicName>
|
||
have been supported by the shape of male genitalia. The specimens properly fixed for the isolation of DNA have not been available and no DNA-based validation of proposed relationships was possible (
|
||
<bibRefCitation id="EF97FFB9C3676356FBC2FDDCFA9AFD88" author="Bocek, M. & Bocak, L." box="[1107,1353,585,611]" pageId="3" pageNumber="344" pagination="3963" refId="ref5383" refString="Bocek, M. & Bocak, L. (2017) The comparison of molecular and morphology-based phylogenies of trichaline net-winged beetles (Coleoptera: Lycidae: Metriorrhynchini) with description of a new subgenus. PeerJ, 5, e 3963. https: // doi. org / 10.7717 / peerj. 3963" type="journal article" year="2017">Bocek & Bocak 2017</bibRefCitation>
|
||
, 2019). The current analysis confirms the sister relationships between
|
||
<taxonomicName id="4C06F9CBC3676356FCC6FDF8FC11FD6C" authorityName="Waterhouse" authorityYear="1879" box="[855,962,621,647]" class="Insecta" family="Lycidae" genus="Eniclases" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9725E5AC3676356FCC6FDF8FC11FD6C" box="[855,962,621,647]" italics="true" pageId="3" pageNumber="344">Eniclases</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C06F9CBC3676356FC6AFDF8FB7BFD6C" authorityName="Kleine" authorityYear="1926" box="[1019,1192,621,647]" class="Insecta" family="Lycidae" genus="Schizotrichalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9725E5AC3676356FC6AFDF8FB7BFD6C" box="[1019,1192,621,647]" italics="true" pageId="3" pageNumber="344">Schizotrichalus</emphasis>
|
||
</taxonomicName>
|
||
. Morphological traits are sometimes insufficient to resolve genus-level phylogeny and the presence of multiple pronotal areolae could be considered as a plesiomorphic state of character. Based on this trait,
|
||
<taxonomicName id="4C06F9CBC3676356FC16FD20FBE7FD24" authorityName="Kleine" authorityYear="1926" box="[903,1076,693,719]" class="Insecta" family="Lycidae" genus="Schizotrichalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9725E5AC3676356FC16FD20FBE7FD24" box="[903,1076,693,719]" italics="true" pageId="3" pageNumber="344">Schizotrichalus</emphasis>
|
||
</taxonomicName>
|
||
could be a sister to other trichalines, which share the characteristic pronotum with a single areola (
|
||
<bibRefCitation id="EF97FFB9C3676356FCECFD4CFBD9FD18" author="Bocak, L." box="[893,1034,729,755]" pageId="3" pageNumber="344" pagination="315 - 351" refId="ref5210" refString="Bocak, L. (2002) Revision and phylogenetic analysis of Metriorrhynchinae. European Journal of Entomology, 99, 315 - 351. https: // doi. org / 10.14411 / eje. 2002.043" type="journal article" year="2002">Bocak 2002</bibRefCitation>
|
||
). The combination of molecular and morphological data suggests the placement of
|
||
<taxonomicName id="4C06F9CBC3676356FD0EFD68FC9FFCFC" authorityName="Kleine" authorityYear="1926" box="[671,844,765,791]" class="Insecta" family="Lycidae" genus="Schizotrichalus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9725E5AC3676356FD0EFD68FC9FFCFC" box="[671,844,765,791]" italics="true" pageId="3" pageNumber="344">Schizotrichalus</emphasis>
|
||
</taxonomicName>
|
||
in a close relationships to
|
||
<taxonomicName id="4C06F9CBC3676356FBE7FD68FB32FCFC" authorityName="Waterhouse" authorityYear="1879" box="[1142,1249,765,791]" class="Insecta" family="Lycidae" genus="Eniclases" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="344" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9725E5AC3676356FBE7FD68FB32FCFC" box="[1142,1249,765,791]" italics="true" pageId="3" pageNumber="344">Eniclases</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |