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<document id="8E61A009CE91AFC953E20EACF9CF0611" ID-CLB-Dataset="2382" ID-DOI="10.11646/zootaxa.5093.3.6" ID-GBIF-Dataset="ceddcc67-ae4d-4e17-bb2c-40f024d5c68f" ID-ISSN="1175-5326" ID-Zenodo-Dep="5909992" ID-ZooBank="E50B2E66-1852-40D8-AED7-3E412DCD6717" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1643276840499" checkinUser="plazi" docAuthor="Bayarsaikhan, Ulziijargal, Bucsek, Karol, Cha, Yeong-Bin, Le, Xuanvi, Kwon, Hyung Wook &amp; Bae, Yang-Seop" docDate="2022" docId="03AB87ECA052B51239FE16AF64C1FF21" docLanguage="en" docName="zootaxa.5093.3.6.pdf" docOrigin="Zootaxa 5093 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Halone arcuata Bayarsaikhan &amp; Bucsek &amp; Cha &amp; Le &amp; Kwon &amp; Bae 2022, sp. nov." docType="treatment" docVersion="7" lastPageNumber="388" masterDocId="FF92FF94A053B5113969120C673DFFD8" masterDocTitle="A new Halone Walker (Lepidoptera, Erebidae, Arctiinae, Lithosiini) from Vietnam with establishing of a new combination" masterLastPageNumber="391" masterPageNumber="385" pageNumber="386" updateTime="1699304535711" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="EC78BCC0C6399F6FC8C2C1A18766DA98">A new Halone Walker (Lepidoptera, Erebidae, Arctiinae, Lithosiini) from Vietnam with establishing of a new combination</mods:title>
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<mods:namePart id="1557ED38846819653DB17A124640601E">Bayarsaikhan, Ulziijargal</mods:namePart>
<mods:affiliation id="1ED85ACFFC4B1B8199395B25979F8089">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea.</mods:affiliation>
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<mods:namePart id="6D9F66F199767814D2CBDDA4D805A113">Bucsek, Karol</mods:namePart>
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<mods:affiliation id="65ECDE1AEFD89F872ED3B637B63F18A5">Slovak Academy of Sciences, Institute of Zoology, Dúbravská cesta 9, 845 06 Bratislava, Slovak Republic. &amp; monosyntaxis @ gmail. com; https: // orcid. org / 0000 - 0003 - 1799 - 8762</mods:affiliation>
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<mods:namePart id="5F6AAED286624045B83274E4EED3D7BF">Cha, Yeong-Bin</mods:namePart>
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<mods:affiliation id="83D32C557A71907ED8F5F6A63DF8FF7D">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; cyb 0201 @ naver. com; https: // orcid. org / 0000 - 0003 - 2635 - 5734</mods:affiliation>
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<mods:namePart id="063F41904BF576363E2B2DC5B60BEB78">Le, Xuanvi</mods:namePart>
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<mods:affiliation id="03DB157FD309FE73F5DB5D1B7BE4D3B5">Division of Pest &amp; Natural Enemy Identification, Plant Protection Research Institute, Ducthang-Bac Tuliem, Hanoi, Vietnam. &amp; lexuanvippri @ gmail. com; https: // orcid. org / 0000 - 0003 - 0409 - 3577</mods:affiliation>
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<mods:namePart id="494D265088C6F525E4114C5443ABBAC5">Kwon, Hyung Wook</mods:namePart>
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<mods:affiliation id="068AA3CD096BCD413E0A9448EE2CAD31">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Convergence Research Center for Insect Vectors, Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, Songdo ˗ dong, Incheon, 22012, Republic of Korea. &amp; hwkwon @ inu. ac. kr; https: // orcid. org / 0000 - 0001 - 9340 - 7974</mods:affiliation>
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<mods:namePart id="22E5FC79EECE71D70034E8A909CA5E25">Bae, Yang-Seop</mods:namePart>
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<mods:affiliation id="AB8B0920913B518C6614718410A01A98">Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Bio ˗ Resource and Environmental Center, Incheon National University, 119 Academy ˗ ro, Yeonsu ˗ gu, Incheon, 22012, Republic of Korea. &amp; Convergence Research Center for Insect Vectors, Division of Life Sciences, College of Life Sciences and Bioengineering, Incheon National University, Songdo ˗ dong, Incheon, 22012, Republic of Korea. &amp; baeys @ inu. ac. kr; https: // orcid. org / 0000 - 0001 - 7356 - 5633</mods:affiliation>
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<mods:date id="2F2050E823F47E967891D54F3B22838E">2022</mods:date>
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<treatment id="03AB87ECA052B51239FE16AF64C1FF21" ID-DOI="http://doi.org/10.5281/zenodo.5910002" ID-GBIF-Taxon="192662375" ID-Zenodo-Dep="5910002" LSID="urn:lsid:plazi:treatment:03AB87ECA052B51239FE16AF64C1FF21" httpUri="http://treatment.plazi.org/id/03AB87ECA052B51239FE16AF64C1FF21" lastPageId="3" lastPageNumber="388" pageId="1" pageNumber="386">
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<emphasis id="B976EAE8A052B51039FE16AF65F0FB66" bold="true" box="[151,717,1187,1214]" pageId="1" pageNumber="386">
<taxonomicName id="4C024D79A052B51039FE16AF655EFB66" authority="Bayarsaikhan &amp; Bae" authorityName="Bayarsaikhan &amp; Bucsek &amp; Cha &amp; Le &amp; Kwon &amp; Bae" authorityYear="2022" box="[151,611,1187,1214]" class="Insecta" family="Arctiidae" genus="Halone" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="386" phylum="Arthropoda" rank="species" species="arcuata" status="sp. nov.">
<emphasis id="B976EAE8A052B51039FE16AF6668FB66" bold="true" box="[151,341,1187,1214]" italics="true" pageId="1" pageNumber="386">Halone arcuata</emphasis>
Bayarsaikhan &amp; Bae
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,
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(
<figureCitation id="13392A7FA052B51039F616C467D0FB3A" box="[159,237,1224,1250]" captionStart="FIGURES 14" captionStartId="2.[151,264,1938,1962]" captionTargetBox="[164,1423,669,1915]" captionTargetId="figure-226@2.[164,1423,669,1915]" captionTargetPageId="2" captionText="FIGURES 14. Adults of new and compared species of Halone. 1a. H.arcuata sp. nov., male, Vietnam, holotype (INU˗10303V). 1b. ditto, female, Vietnam, paratype (INU˗10429V). 2. H. viktorai, male, Malaysia, holotype. 3. H. solitus, male, Malaysia, holotype. 4a. H. hollowayi, male, Cambodia (paratype, INU˗1568C). 4b. ditto, female, Cambodia (holotype, INU˗10033C)." figureDoi="http://doi.org/10.5281/zenodo.5909996" httpUri="https://zenodo.org/record/5909996/files/figure.png" pageId="1" pageNumber="386">Figs. 1</figureCitation>
,
<figureCitation id="13392A7FA052B510399216C46636FB3B" box="[251,267,1224,1251]" captionStart="FIGURES 58" captionStartId="3.[151,264,1611,1635]" captionTargetBox="[162,1428,282,1578]" captionTargetId="figure-52@3.[151,1436,273,1587]" captionTargetPageId="3" captionText="FIGURES 58. Male genitalia of new and compared species of Halone. 5a. H. arcuata sp. nov., holotype (INU˗10303V). 5b. ditto, paratype (INU˗10429V). 6. H. viktorai, holotype (MalS25). 7. H. solitus, holotype (MalS26). 8. H. hollowayi, paratype (INU˗1568C) (scale bar for 5a and 5b)." figureDoi="http://doi.org/10.5281/zenodo.5910000" httpUri="https://zenodo.org/record/5910000/files/figure.png" pageId="1" pageNumber="386">5</figureCitation>
,
<figureCitation id="13392A7FA052B510387E16C4661BFB3A" box="[279,294,1224,1250]" captionStart="FIGURES 910" captionStartId="4.[151,264,1918,1942]" captionTargetBox="[165,1420,1002,1881]" captionTargetId="figure-354@4.[151,1436,993,1895]" captionTargetPageId="4" captionText="FIGURES 910. Female genitalia of new and compared species of Halone. 9. H. arcuata sp. nov., Vietnam, paratype (INU˗10429V). 10. H. hollowayi, Cambodia, holotype (INU˗10033C)." figureDoi="http://doi.org/10.5281/zenodo.5910004" httpUri="https://zenodo.org/record/5910004/files/figure.png" pageId="1" pageNumber="386">9</figureCitation>
)
</paragraph>
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<emphasis id="B976EAE8A052B51039FE171C6673FAF2" bold="true" box="[151,334,1296,1322]" pageId="1" pageNumber="386">Type materials.</emphasis>
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<emphasis id="B976EAE8A052B510383D171C66F7FAF2" bold="true" box="[340,458,1296,1322]" pageId="1" pageNumber="386">
<typeStatus id="54B98858A052B510383D171C66F8FAF2" box="[340,453,1296,1322]" pageId="1" pageNumber="386" type="holotype">Holotype</typeStatus>
:
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<specimenCount id="9D04FD73A052B51038B9171D66DAFAF3" box="[464,487,1297,1323]" pageId="1" pageNumber="386" type="male"></specimenCount>
,
<collectorName id="26F7532CA052B5103898171C656EFAF2" box="[497,595,1296,1322]" pageId="1" pageNumber="386">Vietnam</collectorName>
,
<collectorName id="26F7532CA052B5103B35171D640BFAF3" box="[604,822,1296,1323]" pageId="1" pageNumber="386">Prov. Ninhbinh, NP</collectorName>
.
<collectorName id="26F7532CA052B5103A56171C64FEFAF2" box="[831,963,1296,1322]" pageId="1" pageNumber="386">Cucphuong</collectorName>
,
<date id="FFBC103AA052B5103AA4171C630FFAF3" box="[973,1074,1296,1323]" pageId="1" pageNumber="386" value="2005-05-01">
<collectingDate id="EFF8E9D2A052B5103AA4171C630FFAF3" box="[973,1074,1296,1323]" pageId="1" pageNumber="386" value="2005-05-01">1.V.2005</collectingDate>
</date>
(
<collectorName id="26F7532CA052B5103D56171C63ABFAF3" box="[1087,1174,1296,1323]" pageId="1" pageNumber="386">YS Bae</collectorName>
&amp;
<collectionCode id="ED13AE3FA052B5103DDF171D63E7FAF3" box="[1206,1242,1297,1323]" country="Germany" name="Jura Museum, Eichstatt" pageId="1" pageNumber="386">JM</collectionCode>
<collectorName id="26F7532CA052B5103D89171C622BFAF3" box="[1248,1302,1296,1323]" pageId="1" pageNumber="386">Kim</collectorName>
),
<collectorName id="26F7532CA052B5103C4E171C67FCFA97" pageId="1" pageNumber="386">Gen. Slide No.</collectorName>
INU˗10303
<collectionCode id="ED13AE3FA052B510382017386660FA96" box="[329,349,1332,1358]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="386" type="Museum">V</collectionCode>
, deposited in
<collectionCode id="ED13AE3FA052B510389317396504FA97" box="[506,569,1333,1359]" country="South Africa" httpUri="http://grbio.org/cool/6j3y-rabv" name="ARC-Plant Protection Research Institute, National Collection of Fungi: Culture Collection" pageId="1" pageNumber="386">PPRI</collectionCode>
</materialsCitation>
.
<materialsCitation id="3B6A3CA7A052B5103B2C173864F7FA96" ID-GBIF-Occurrence="3458072302" box="[581,970,1332,1359]" country="Vietnam" location="VIETNAM" pageId="1" pageNumber="386" specimenCount="3" specimenCount-female="2" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B976EAE8A052B5103B2C17386580FA96" bold="true" box="[581,701,1332,1358]" pageId="1" pageNumber="386">
<typeStatus id="54B98858A052B5103B2C17386580FA96" box="[581,701,1332,1358]" pageId="1" pageNumber="386" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="9D04FD73A052B5103BA6173865C6FA97" box="[719,763,1332,1359]" pageId="1" pageNumber="386" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D04FD73A052B5103A611738640CFA97" box="[776,817,1332,1359]" pageId="1" pageNumber="386" type="female">2 ♀</specimenCount>
)
<collectingCountry id="F315766AA052B5103A2D173864F7FA96" box="[836,970,1332,1358]" name="Vietnam" pageId="1" pageNumber="386">VIETNAM</collectingCountry>
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:
<specimenCount id="9D04FD73A052B5103ABC1738633FFA97" box="[981,1026,1332,1359]" pageId="1" pageNumber="386" type="male">1 ♂</specimenCount>
,
<specimenCount id="9D04FD73A052B5103D6617386307FA97" box="[1039,1082,1332,1359]" pageId="1" pageNumber="386" type="female">2 ♀</specimenCount>
,
<materialsCitation id="3B6A3CA7A052B5103D20173963BEFAAB" ID-GBIF-Occurrence="3458072305" collectingDate="2005-05-01" collectionCode="V" collectorName="Gen. Slide No." country="Vietnam" pageId="1" pageNumber="386" specimenCount="1" typeStatus="paratype">
same data as for the holotype,
<collectorName id="26F7532CA052B51039FE17546603FAAB" box="[151,318,1368,1395]" pageId="1" pageNumber="386">Gen. Slide No.</collectorName>
INU˗10156
<collectionCode id="ED13AE3FA052B51038A1175466E1FAAB" box="[456,476,1368,1395]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="386" type="Museum">V</collectionCode>
(female), 10429
<collectionCode id="ED13AE3FA052B5103BE41754659CFAAB" box="[653,673,1368,1395]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="386" type="Museum">V</collectionCode>
(female), 10431
<collectionCode id="ED13AE3FA052B5103A3D17546454FAAB" box="[852,873,1368,1395]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="386" type="Museum">V</collectionCode>
(male), deposited in INU
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3186571A052B51039AE177063F9F86A" pageId="1" pageNumber="386" type="diagnosis">
<paragraph id="8BBD36FAA052B51039AE177063F9F86A" blockId="1.[151,1437,1296,2042]" pageId="1" pageNumber="386">
<emphasis id="B976EAE8A052B51039AE1770667FFA4E" bold="true" box="[199,322,1404,1430]" pageId="1" pageNumber="386">Diagnosis.</emphasis>
<taxonomicName id="4C024D79A052B5103822177066C0FA4E" authority="Bayarsaikhan &amp; Bucsek &amp; Cha &amp; Le &amp; Kwon &amp; Bae, 2022" authorityName="Bayarsaikhan &amp; Bucsek &amp; Cha &amp; Le &amp; Kwon &amp; Bae" authorityYear="2022" box="[331,509,1404,1430]" class="Insecta" family="Arctiidae" genus="Halone" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="386" phylum="Arthropoda" rank="species" species="arcuata" status="sp. nov.">
<emphasis id="B976EAE8A052B5103822177066C0FA4E" box="[331,509,1404,1430]" italics="true" pageId="1" pageNumber="386">Halone arcuata</emphasis>
</taxonomicName>
<emphasis id="B976EAE8A052B5103B6E1770655EFA4E" bold="true" box="[519,611,1404,1430]" pageId="1" pageNumber="386">
<taxonomicNameLabel id="A2455793A052B5103B6E1770655EFA4E" box="[519,611,1404,1430]" pageId="1" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from other congeners by its dark brown longitudinal streak (the antemedial and the disco-cellular spots are almost connected) in the discal area of the forewing. The male genitalia of the new species is similar to those of
<taxonomicName id="4C024D79A052B5103B0B17C86485FA06" authority="Bucsek, 2014" authorityName="Bucsek" authorityYear="2014" box="[610,952,1476,1502]" class="Insecta" family="Arctiidae" genus="Halone" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="386" phylum="Arthropoda" rank="species" species="victorai">
<emphasis id="B976EAE8A052B5103B0B17C8642EFA06" box="[610,787,1476,1502]" italics="true" pageId="1" pageNumber="386">Halone victorai</emphasis>
<bibRefCitation id="EF934B0BA052B5103A7217C86485FA06" author="Bucsek, K." box="[795,952,1476,1502]" pageId="1" pageNumber="386" refId="ref3377" refString="Bucsek, K. (2014) Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula - Malaysia. Supplementum. Institute of Zoology SAS, Bratislava, 42 pp." type="book" year="2014">Bucsek, 2014</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C024D79A052B5103AAD17C963EFFA06" authority="Bucsek, 2014" authorityName="Bucsek" authorityYear="2014" box="[964,1234,1476,1502]" class="Insecta" family="Arctiidae" genus="Halone" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="386" phylum="Arthropoda" rank="species" species="solitus">
<emphasis id="B976EAE8A052B5103AAD17C9630DFA06" box="[964,1072,1476,1502]" italics="true" pageId="1" pageNumber="386">H. solitus</emphasis>
<bibRefCitation id="EF934B0BA052B5103D5117C863EFFA06" author="Bucsek, K." box="[1080,1234,1476,1502]" pageId="1" pageNumber="386" refId="ref3377" refString="Bucsek, K. (2014) Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula - Malaysia. Supplementum. Institute of Zoology SAS, Bratislava, 42 pp." type="book" year="2014">Bucsek, 2014</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="4C024D79A052B5103C6317C966F8F9DA" authority="Bayarsaikhan &amp; Bae, 2019" authorityName="Bayarsaikhan &amp; Bae" authorityYear="2019" class="Insecta" family="Arctiidae" genus="Halone" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="386" phylum="Arthropoda" rank="species" species="hollowayi">
<emphasis id="B976EAE8A052B5103C6317C962A1FA06" box="[1290,1436,1476,1502]" italics="true" pageId="1" pageNumber="386">H. hollowayi</emphasis>
Bayarsaikhan &amp;
<bibRefCitation id="EF934B0BA052B510383D17E566F8F9DA" author="Bae, Y. S. &amp; Bayarsaikhan, U." box="[340,453,1512,1539]" pageId="1" pageNumber="386" pagination="395 - 400" refId="ref3183" refString="Bae, Y. S. &amp; Bayarsaikhan, U. (2019) New and newly recorded species of the genus Halone Walker, 1854 (Lepidoptera, Erebidae, Arctiinae, Lithosiini) from Cambodia, Laos and Vietnam. Zootaxa, 4586 (2), 395 - 400. https: // doi. org / 10.11646 / zootaxa. 4586.2.13" type="journal article" year="2019">Bae, 2019</bibRefCitation>
</taxonomicName>
due to the irregularly projected tegumen, the well-developed saccular process as length as elongated, membranous cucculus. However,
<taxonomicName id="4C024D79A052B5103BCA14016427F9FE" authority="Bayarsaikhan &amp; Bucsek &amp; Cha &amp; Le &amp; Kwon &amp; Bae, 2022" authorityName="Bayarsaikhan &amp; Bucsek &amp; Cha &amp; Le &amp; Kwon &amp; Bae" authorityYear="2022" box="[675,794,1549,1574]" class="Insecta" family="Arctiidae" genus="Halone" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="386" phylum="Arthropoda" rank="species" species="arcuata" status="sp. nov.">
<emphasis id="B976EAE8A052B5103BCA14016427F9FE" box="[675,794,1549,1574]" italics="true" pageId="1" pageNumber="386">H. arcuata</emphasis>
</taxonomicName>
<emphasis id="B976EAE8A052B5103A761400644AF9FE" bold="true" box="[799,887,1548,1574]" pageId="1" pageNumber="386">
<taxonomicNameLabel id="A2455793A052B5103A761400644AF9FE" box="[799,887,1548,1574]" pageId="1" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
differs from the allied species in having a roundly arch-shaped projection of the tegumen and a dentate, strongly arcuate and sclerotized saccular process, with an acuminate apex, while in
<taxonomicName id="4C024D79A052B51038DE1459650DF9B6" authorityName="Bucsek" authorityYear="2014" box="[439,560,1620,1646]" class="Insecta" family="Arctiidae" genus="Halone" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="386" phylum="Arthropoda" rank="species" species="viktorai">
<emphasis id="B976EAE8A052B51038DE1459650DF9B6" box="[439,560,1620,1646]" italics="true" pageId="1" pageNumber="386">H. viktorai</emphasis>
</taxonomicName>
(illustrated by
<bibRefCitation id="EF934B0BA052B5103BB21458644DF9B6" author="Bucsek, K." box="[731,880,1620,1646]" pageId="1" pageNumber="386" refId="ref3377" refString="Bucsek, K. (2014) Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula - Malaysia. Supplementum. Institute of Zoology SAS, Bratislava, 42 pp." type="book" year="2014">Bucsek 2014</bibRefCitation>
: gen. MalS25; present paper fig. 6), the subapical projection of the tegumen is slender, almost straight and spine-shaped, and the saccular process is slender, weakly tapering, entire, with a rounded apex. In
<taxonomicName id="4C024D79A052B5103B36149165F6F96E" authorityName="Bucsek" authorityYear="2014" box="[607,715,1692,1718]" class="Insecta" family="Arctiidae" genus="Halone" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="386" phylum="Arthropoda" rank="species" species="solitus">
<emphasis id="B976EAE8A052B5103B36149165F6F96E" box="[607,715,1692,1718]" italics="true" pageId="1" pageNumber="386">H. solitus</emphasis>
</taxonomicName>
(illustrated by
<bibRefCitation id="EF934B0BA052B5103A1C14906336F96E" author="Bucsek, K." box="[885,1035,1692,1718]" pageId="1" pageNumber="386" refId="ref3377" refString="Bucsek, K. (2014) Erebidae, Arctiinae (Lithosiini, Arctiini) of Malay Peninsula - Malaysia. Supplementum. Institute of Zoology SAS, Bratislava, 42 pp." type="book" year="2014">Bucsek 2014</bibRefCitation>
: gen. MalS26; present paper fig. 7), the apical projection of the tegumen is broad and rectangular, and the slender saccular process is entire, slender and apically rounded. In
<taxonomicName id="4C024D79A052B51038E914E9652CF926" authorityName="Bayarsaikhan &amp; Bae" authorityYear="2019" box="[384,529,1764,1790]" class="Insecta" family="Arctiidae" genus="Halone" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="386" phylum="Arthropoda" rank="species" species="hollowayi">
<emphasis id="B976EAE8A052B51038E914E9652CF926" box="[384,529,1764,1790]" italics="true" pageId="1" pageNumber="386">H. hollowayi</emphasis>
</taxonomicName>
(illustrated by
<bibRefCitation id="EF934B0BA052B5103BD414E964D4F926" author="Bae, Y. S. &amp; Bayarsaikhan, U." box="[701,1001,1764,1791]" pageId="1" pageNumber="386" pagination="395 - 400" refId="ref3183" refString="Bae, Y. S. &amp; Bayarsaikhan, U. (2019) New and newly recorded species of the genus Halone Walker, 1854 (Lepidoptera, Erebidae, Arctiinae, Lithosiini) from Cambodia, Laos and Vietnam. Zootaxa, 4586 (2), 395 - 400. https: // doi. org / 10.11646 / zootaxa. 4586.2.13" type="journal article" year="2019">Bae &amp; Bayarsaikhan 2019</bibRefCitation>
: fig. 5; present paper fig. 8), the apical projection of the tegumen is strongly tapering, curved, the saccular process is more stout than in
<taxonomicName id="4C024D79A052B5103D9E1505624CF8FA" authority="Bayarsaikhan &amp; Bucsek &amp; Cha &amp; Le &amp; Kwon &amp; Bae, 2022" authorityName="Bayarsaikhan &amp; Bucsek &amp; Cha &amp; Le &amp; Kwon &amp; Bae" authorityYear="2022" box="[1271,1393,1801,1826]" class="Insecta" family="Arctiidae" genus="Halone" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="386" phylum="Arthropoda" rank="species" species="arcuata" status="sp. nov.">
<emphasis id="B976EAE8A052B5103D9E1505624CF8FA" box="[1271,1393,1801,1826]" italics="true" pageId="1" pageNumber="386">H. arcuata</emphasis>
</taxonomicName>
<emphasis id="B976EAE8A052B5103C12150467F5F89E" bold="true" pageId="1" pageNumber="386">
<taxonomicNameLabel id="A2455793A052B5103C12150467F5F89E" pageId="1" pageNumber="386" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, entire and has an irregularly rectangular apex. In the female genitalia, the following features clearly separate the new species from all the other known females of genus
<taxonomicName id="4C024D79A052B5103A5A155C64B8F8B2" authorityName="Walker" authorityYear="1854" box="[819,901,1872,1898]" class="Insecta" family="Arctiidae" genus="Halone" kingdom="Animalia" order="Lepidoptera" pageId="1" pageNumber="386" phylum="Arthropoda" rank="genus">
<emphasis id="B976EAE8A052B5103A5A155C64B8F8B2" box="[819,901,1872,1898]" italics="true" pageId="1" pageNumber="386">Halone</emphasis>
</taxonomicName>
: (1) the postvaginalis plate is sclerotized, “bow tie”-shaped and densely serrulate; (2) the anterior section of the antrum is broadly rounded, sclerotized, with weakly waved edge; (3) the lamella antevaginalis is horizontally elongate, arch-shaped and sclerotized.
</paragraph>
</subSubSection>
<subSubSection id="C3186571A052B51339AE15B0632DFD5D" lastPageId="2" lastPageNumber="387" pageId="1" pageNumber="386" type="description">
<paragraph id="8BBD36FAA052B51339AE15B0632DFD5D" blockId="1.[151,1437,1296,2042]" lastBlockId="2.[151,1437,151,645]" lastPageId="2" lastPageNumber="387" pageId="1" pageNumber="386">
<emphasis id="B976EAE8A052B51039AE15B0666CF80E" bold="true" box="[199,337,1980,2006]" pageId="1" pageNumber="386">Description</emphasis>
<emphasis id="B976EAE8A052B510383115B066A8F80E" box="[344,405,1980,2006]" italics="true" pageId="1" pageNumber="386">Adult</emphasis>
(
<figureCitation id="13392A7FA052B51038CA15B0651BF80F" box="[419,550,1980,2007]" captionStart="FIGURES 14" captionStartId="2.[151,264,1938,1962]" captionTargetBox="[164,1423,669,1915]" captionTargetId="figure-226@2.[164,1423,669,1915]" captionTargetPageId="2" captionText="FIGURES 14. Adults of new and compared species of Halone. 1a. H.arcuata sp. nov., male, Vietnam, holotype (INU˗10303V). 1b. ditto, female, Vietnam, paratype (INU˗10429V). 2. H. viktorai, male, Malaysia, holotype. 3. H. solitus, male, Malaysia, holotype. 4a. H. hollowayi, male, Cambodia (paratype, INU˗1568C). 4b. ditto, female, Cambodia (holotype, INU˗10033C)." figureDoi="http://doi.org/10.5281/zenodo.5909996" httpUri="https://zenodo.org/record/5909996/files/figure.png" pageId="1" pageNumber="386">Figs. 1a, 1b</figureCitation>
). Length of forewing
<quantity id="4CFA9B1FA052B5103A7115B0644DF80F" box="[792,880,1980,2007]" metricMagnitude="-3" metricUnit="m" metricValue="6.5" metricValueMax="7.0" metricValueMin="6.0" pageId="1" pageNumber="386" unit="mm" value="6.5" valueMax="7.0" valueMin="6.0">6˗7 mm</quantity>
in males (n=2), 7˗7.5 mm in females (n=2). Head, patagium, tegula and thorax sordid white, mixed with brown scales. Forewing ground color sordid white, with pale brown sprinkles; medial discal area with a short, longitudinal dark brown streak (antemedial and discocellular patches almost connected); cilia sordid white. Hindwing pale brown; cilia gray. Abdomen fuscous in both sexes, tinged with pale yellow in females.
<emphasis id="B976EAE8A051B5133B4112D365F1FF21" box="[552,716,223,249]" italics="true" pageId="2" pageNumber="387">Male genitalia</emphasis>
(
<figureCitation id="13392A7FA051B5133BB212D3645FFF21" box="[731,866,222,249]" captionStart="FIGURES 58" captionStartId="3.[151,264,1611,1635]" captionTargetBox="[162,1428,282,1578]" captionTargetId="figure-52@3.[151,1436,273,1587]" captionTargetPageId="3" captionText="FIGURES 58. Male genitalia of new and compared species of Halone. 5a. H. arcuata sp. nov., holotype (INU˗10303V). 5b. ditto, paratype (INU˗10429V). 6. H. viktorai, holotype (MalS25). 7. H. solitus, holotype (MalS26). 8. H. hollowayi, paratype (INU˗1568C) (scale bar for 5a and 5b)." figureDoi="http://doi.org/10.5281/zenodo.5910000" httpUri="https://zenodo.org/record/5910000/files/figure.png" pageId="2" pageNumber="387">Figs. 5a, 5b</figureCitation>
). Uncus stout, acuminate, with a stout and apically rounded ventral process. Tegumen weakly sclerotized, strongly shouldered, with roundly arch-shaped apical projection. Anal tube membranous, scaphium weakly trapezoid, with weakly sclerotized edge. Distal section of valva membranous, sparsely setose, narrow, apically rounded, almost equal in length to tegumen. Saccular process strongly arcuate, sclerotized, with dentate proximal half and with acuminate apex, almost equal in length to valva. Saccus broadly “U”˗shaped. Aedeagus stout, weakly sclerotized; vesical scobinated, with seven slender spines and elongate, sclerotized tube.
<emphasis id="B976EAE8A051B51338AA13BB65BFFE09" box="[451,642,439,465]" italics="true" pageId="2" pageNumber="387">Female genitalia</emphasis>
(
<figureCitation id="13392A7FA051B5133BF813BB65E8FE09" box="[657,725,439,465]" captionStart="FIGURES 910" captionStartId="4.[151,264,1918,1942]" captionTargetBox="[165,1420,1002,1881]" captionTargetId="figure-354@4.[151,1436,993,1895]" captionTargetPageId="4" captionText="FIGURES 910. Female genitalia of new and compared species of Halone. 9. H. arcuata sp. nov., Vietnam, paratype (INU˗10429V). 10. H. hollowayi, Cambodia, holotype (INU˗10033C)." figureDoi="http://doi.org/10.5281/zenodo.5910004" httpUri="https://zenodo.org/record/5910004/files/figure.png" pageId="2" pageNumber="387">Fig. 9</figureCitation>
). Papillae anales weakly covered with setose. Postvaginal plate sclerotized, “bow tie”˗shaped, densely serrulate.Antrum sclerotized, broadly rounded, with slightly waved posterior margin with a shallow incision medially. Apophysis posterioris slender, about 2 times longer than thiker apophysis anterioris. Lamella antevaginalis strongly sclerotized, horizontally elongated, arch-shaped. Right section of ductus bursae heavily sclerotized and tubular, while left section broad, membranous, with rectangular sclerotized plate. Corpus bursae globular, membranous, with a strongly dentate signum medially.
</paragraph>
</subSubSection>
<caption id="DF7D6672A051B51339FE159E6241F82A" ID-DOI="http://doi.org/10.5281/zenodo.5909996" ID-Zenodo-Dep="5909996" httpUri="https://zenodo.org/record/5909996/files/figure.png" pageId="2" pageNumber="387" startId="2.[151,264,1938,1962]" targetBox="[164,1423,669,1915]" targetPageId="2">
<paragraph id="8BBD36FAA051B51339FE159E6241F82A" blockId="2.[151,1437,1938,2034]" pageId="2" pageNumber="387">
<emphasis id="B976EAE8A051B51339FE159E6605F872" bold="true" box="[151,312,1938,1962]" pageId="2" pageNumber="387">FIGURES 14.</emphasis>
Adults of new and compared species of
<taxonomicName id="4C024D79A051B5133BAC159E642DF872" authorityName="Walker" authorityYear="1854" box="[709,784,1938,1962]" class="Insecta" family="Arctiidae" genus="Halone" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="387" phylum="Arthropoda" rank="genus">
<emphasis id="B976EAE8A051B5133BAC159E642DF872" box="[709,784,1938,1962]" italics="true" pageId="2" pageNumber="387">Halone</emphasis>
</taxonomicName>
. 1a.
<taxonomicName id="4C024D79A051B5133A55159F649AF872" authority="Bayarsaikhan &amp; Bucsek &amp; Cha &amp; Le &amp; Kwon &amp; Bae, 2022" authorityName="Bayarsaikhan &amp; Bucsek &amp; Cha &amp; Le &amp; Kwon &amp; Bae" authorityYear="2022" box="[828,935,1939,1962]" class="Insecta" family="Arctiidae" genus="Halone" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="387" phylum="Arthropoda" rank="species" species="arcuata" status="sp. nov.">
<emphasis id="B976EAE8A051B5133A55159F649AF872" box="[828,935,1939,1962]" italics="true" pageId="2" pageNumber="387">H. arcuata</emphasis>
</taxonomicName>
<emphasis id="B976EAE8A051B5133AC5159E64C7F872" bold="true" box="[940,1018,1938,1962]" pageId="2" pageNumber="387">
<taxonomicNameLabel id="A2455793A051B5133AC5159E64C7F872" box="[940,1018,1938,1962]" pageId="2" pageNumber="387" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, male, Vietnam, holotype (INU˗10303V). 1b.
<emphasis id="B976EAE8A051B51339A915BA67C9F816" box="[192,244,1974,1998]" italics="true" pageId="2" pageNumber="387">ditto,</emphasis>
female, Vietnam, paratype (INU˗10429V). 2.
<emphasis id="B976EAE8A051B5133BB115BB6473F816" box="[728,846,1974,1998]" italics="true" pageId="2" pageNumber="387">
<taxonomicName id="4C024D79A051B5133BB115BB6477F816" authorityName="Bucsek" authorityYear="2014" box="[728,842,1974,1998]" class="Insecta" family="Arctiidae" genus="Halone" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="387" phylum="Arthropoda" rank="species" species="viktorai">H. viktorai</taxonomicName>
,
</emphasis>
male, Malaysia, holotype. 3.
<taxonomicName id="4C024D79A051B5133DEF15BB63D7F816" authorityName="Bucsek" authorityYear="2014" box="[1158,1258,1974,1998]" class="Insecta" family="Arctiidae" genus="Halone" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="387" phylum="Arthropoda" rank="species" species="solitus">
<emphasis id="B976EAE8A051B5133DEF15BB63D7F816" box="[1158,1258,1974,1998]" italics="true" pageId="2" pageNumber="387">H. solitus</emphasis>
</taxonomicName>
, male, Malaysia, holotype. 4a.
<taxonomicName id="4C024D79A051B513384815D76699F82A" authorityName="Bayarsaikhan &amp; Bae" authorityYear="2019" box="[289,420,2010,2034]" class="Insecta" family="Arctiidae" genus="Halone" kingdom="Animalia" order="Lepidoptera" pageId="2" pageNumber="387" phylum="Arthropoda" rank="species" species="hollowayi">
<emphasis id="B976EAE8A051B513384815D76699F82A" box="[289,420,2010,2034]" italics="true" pageId="2" pageNumber="387">H. hollowayi</emphasis>
</taxonomicName>
, male, Cambodia (paratype, INU˗1568C). 4b.
<emphasis id="B976EAE8A051B5133A1415D66496F82A" box="[893,939,2010,2034]" italics="true" pageId="2" pageNumber="387">ditto</emphasis>
, female, Cambodia (holotype, INU˗10033C).
</paragraph>
</caption>
<subSubSection id="C3186571A050B51239AE129A65AFFF69" box="[199,658,150,177]" pageId="3" pageNumber="388" type="distribution">
<paragraph id="8BBD36FAA050B51239AE129A65AFFF69" blockId="3.[151,1437,150,249]" box="[199,658,150,177]" pageId="3" pageNumber="388">
<emphasis id="B976EAE8A050B51239AE129A665CFF68" bold="true" box="[199,353,150,176]" pageId="3" pageNumber="388">Distribution.</emphasis>
<collectingCountry id="F315766AA050B512380E129B66FBFF69" box="[359,454,151,177]" name="Vietnam" pageId="3" pageNumber="388">Vietnam</collectingCountry>
(Ninhbinh Prov.).
</paragraph>
</subSubSection>
<subSubSection id="C3186571A050B51239AE12B764C1FF21" pageId="3" pageNumber="388" type="etymology">
<paragraph id="8BBD36FAA050B51239AE12B764C1FF21" blockId="3.[151,1437,150,249]" pageId="3" pageNumber="388">
<emphasis id="B976EAE8A050B51239AE12B76673FF0D" bold="true" box="[199,334,187,213]" pageId="3" pageNumber="388">Etymology.</emphasis>
The specific epithet is derived from the Latin “
<emphasis id="B976EAE8A050B5123A0612B764EDFF0C" box="[879,976,187,212]" italics="true" pageId="3" pageNumber="388">arcuatus</emphasis>
” meaning “arched”, refers to the arched saccular process and arch˗shaped projection of tegumen in the male genitalia.
</paragraph>
</subSubSection>
</treatment>
</document>