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<document id="2CD799900F05BC2695DE162D67523444" ID-CLB-Dataset="28065" ID-DOI="10.11646/zootaxa.4504.4.5" ID-GBIF-Dataset="343a6a67-e8bf-467c-9818-de9621bbe955" ID-ISSN="1175-5326" ID-Zenodo-Dep="2606614" ID-ZooBank="08E1A79F-BAA4-4EAB-9636-FE44BF7EB7F3" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553585132792" checkinUser="plazi" docAuthor="Jäger, Peter" docDate="2018" docId="03AB87D4E76E6F6312F9FD4AC8D37DD0" docLanguage="en" docName="zootaxa.4504.4.5.pdf" docOrigin="Zootaxa 4504 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Systaria Simon 1897, spec. nov." docType="treatment" docVersion="8" lastPageNumber="525" masterDocId="FF92FFACE76F6F62126EFFC4C9517912" masterDocTitle="On the genus Systaria (Araneae: Clubionidae) in Southeast Asia: new species from caves and forests" masterLastPageNumber="544" masterPageNumber="524" pageNumber="525" updateTime="1698712839155" updateUser="ExternalLinkService">
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<mods:title id="EED95C25D219E69AA4A188923F863610">On the genus Systaria (Araneae: Clubionidae) in Southeast Asia: new species from caves and forests</mods:title>
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<treatment id="03AB87D4E76E6F6312F9FD4AC8D37DD0" ID-DOI="http://doi.org/10.5281/zenodo.5988000" ID-GBIF-Taxon="154495195" ID-Zenodo-Dep="5988000" LSID="urn:lsid:plazi:treatment:03AB87D4E76E6F6312F9FD4AC8D37DD0" httpUri="http://treatment.plazi.org/id/03AB87D4E76E6F6312F9FD4AC8D37DD0" lastPageNumber="525" pageId="1" pageNumber="525">
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<paragraph id="8BBD36C2E76E6F6312F9FD4AC8B97BBA" blockId="1.[151,488,654,680]" box="[151,488,654,680]" pageId="1" pageNumber="525">
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<emphasis id="B976EAD0E76E6F6312F9FD4AC8B97BBA" bold="true" box="[151,488,654,680]" pageId="1" pageNumber="525">
Genus
<taxonomicName id="4C024D41E76E6F631280FD4AC8B97BBA" authority="Simon 1897" authorityName="Simon" authorityYear="1897" box="[238,488,654,680]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="525" phylum="Arthropoda" rank="genus">
<emphasis id="B976EAD0E76E6F631280FD4AC81E7BBA" bold="true" box="[238,335,654,680]" italics="true" pageId="1" pageNumber="525">Systaria</emphasis>
<bibRefCitation id="EF934B33E76E6F631338FD4AC8B97BBA" author="Simon, E." box="[342,488,654,680]" pageId="1" pageNumber="525" refId="ref12442" refString="Simon, E. (1897) Histoire naturelle des araignees. Uol. 2. Roret, Paris, 192 pp." type="book" year="1897">Simon 1897</bibRefCitation>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36C2E76E6F6312F9FD11C8D37DD0" blockId="1.[151,1437,725,1218]" pageId="1" pageNumber="525">
<emphasis id="B976EAD0E76E6F6312F9FD11C9877BFC" bold="true" box="[151,214,725,750]" pageId="1" pageNumber="525">Note.</emphasis>
The terminology of the female internal duct system was used ambiguously in previous literature. Copulatory duct (part, in which embolus is inserted; insemination duct sensu
<bibRefCitation id="EF934B33E76E6F6311E8FD3DCDAE7A00" author="Dankittipakul, P. &amp; Singtripop, T." box="[902,1279,761,786]" pageId="1" pageNumber="525" pagination="16 - 32" refId="ref11494" refString="Dankittipakul, P. &amp; Singtripop, T. (2011) Seven new species of Systaria Simon, 1897 from southeast Asia (Araneae, Clubionidae, Systariinae). Zootaxa, 2905, 16 - 32." type="journal article" year="2011">Dankittipakul &amp; Singtripop 2011</bibRefCitation>
) connects the copulatory opening with the part of the duct system where the bursae branches off. The part between that point and the spermathecae I call connecting tube according to the terminology used by
<bibRefCitation id="EF934B33E76E6F631659FC85CDA47A48" author="Jin, C. &amp; Fu, L. &amp; Yin, X. C. &amp; Zhang, F." box="[1079,1269,833,858]" pageId="1" pageNumber="525" pagination="30 - 55" refId="ref12173" refString="Jin, C., Fu, L., Yin, X. C. &amp; Zhang, F. (2016) Four new species of the genus Otacilia Thorell, 1897 from Hunan Province, China (Araneae, Phrurolithidae). ZooKeys, 620, 30 - 55." type="journal article" year="2016">
Jin
<emphasis id="B976EAD0E76E6F63160DFC86CDCE7A48" box="[1123,1183,833,858]" italics="true" pageId="1" pageNumber="525">et al.</emphasis>
(2016)
</bibRefCitation>
for the genus
<emphasis id="B976EAD0E76E6F6312F9FCA1C9A27A6C" box="[151,243,869,894]" italics="true" pageId="1" pageNumber="525">Otacilia</emphasis>
Thorell 1897 (
<taxonomicName id="4C024D41E76E6F6313F0FCA1CB147A6C" box="[414,581,869,894]" class="Arachnida" family="Phrurolithidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="525" phylum="Arthropoda" rank="family">Phrurolithidae</taxonomicName>
). Both ducts might be short and hardly discernible. As in
<bibRefCitation id="EF934B33E76E6F6316B4FCA2CCC67A6C" author="Jager, P." box="[1242,1431,869,894]" pageId="1" pageNumber="525" pagination="87 - 114" refId="ref11664" refString="Jager, P. (2005) New large-sized cave-dwelling Heteropoda species from Asia, with notes on their relationships (Araneae: Sparassidae: Heteropodinae). Revue Suisse de Zoologie, 112, 87 - 114. https: // doi. org / 10.5962 / bhl. part. 80288" type="journal article" year="2005">Jäger (2005: 88)</bibRefCitation>
, slit sense organs close to the epigyne are illustrated as descriptive characters. Same is true for sclerotised spheres in the epigyne (see Jäger 2012). These were interpreted as remnants of fused epigynal furrows. Parts of these furrows can be seen in
<taxonomicName id="4C024D41E76E6F63132AFC16CB537AF8" authority="Jäger, 2018" authorityName="Jäger" authorityYear="2018" box="[324,514,977,1002]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="525" phylum="Arthropoda" rank="species" species="lannops" status="sp. nov.">
<emphasis id="B976EAD0E76E6F63132AFC16CB537AF8" box="[324,514,977,1002]" italics="true" pageId="1" pageNumber="525">Systaria lannops</emphasis>
</taxonomicName>
<emphasis id="B976EAD0E76E6F631064FC15CB2E7AF8" bold="true" box="[522,639,977,1002]" pageId="1" pageNumber="525">
<taxonomicNameLabel id="A24557ABE76E6F631064FC15CB2E7AF8" box="[522,639,977,1002]" pageId="1" pageNumber="525" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C024D41E76E6F6310D7FC16CA7E7AFB" authority="Jäger, 2018" authorityName="Jäger" authorityYear="2018" box="[697,815,978,1001]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="525" phylum="Arthropoda" rank="species" species="procera" status="sp. nov.">
<emphasis id="B976EAD0E76E6F6310D7FC16CA7E7AFB" box="[697,815,978,1001]" italics="true" pageId="1" pageNumber="525">S. procera</emphasis>
</taxonomicName>
<emphasis id="B976EAD0E76E6F631156FC15CAFC7AF8" bold="true" box="[824,941,977,1002]" pageId="1" pageNumber="525">
<taxonomicNameLabel id="A24557ABE76E6F631156FC15CAFC7AF8" box="[824,941,977,1002]" pageId="1" pageNumber="525" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
In
<taxonomicName id="4C024D41E76E6F6311B8FC16CD377AF8" authority="Jäger, 2018" authorityName="Jäger" authorityYear="2018" box="[982,1126,977,1002]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="525" phylum="Arthropoda" rank="species" species="longinqua" status="sp. nov.">
<emphasis id="B976EAD0E76E6F6311B8FC16CD377AF8" box="[982,1126,977,1002]" italics="true" pageId="1" pageNumber="525">S. longinqua</emphasis>
</taxonomicName>
<emphasis id="B976EAD0E76E6F631600FC15CDB27AF8" bold="true" box="[1134,1251,977,1002]" pageId="1" pageNumber="525">
<taxonomicNameLabel id="A24557ABE76E6F631600FC15CDB27AF8" box="[1134,1251,977,1002]" pageId="1" pageNumber="525" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
and
<taxonomicName id="4C024D41E76E6F631773FC16CCCD7AFB" authority="Jäger, 2018" authorityName="Jäger" authorityYear="2018" box="[1309,1436,978,1001]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="525" phylum="Arthropoda" rank="species" species="princesa" status="sp. nov.">
<emphasis id="B976EAD0E76E6F631773FC16CCCD7AFB" box="[1309,1436,978,1001]" italics="true" pageId="1" pageNumber="525">S. princesa</emphasis>
</taxonomicName>
<emphasis id="B976EAD0E76E6F6312F9FC31C85B7D1C" bold="true" box="[151,266,1013,1038]" pageId="1" pageNumber="525">
<taxonomicNameLabel id="A24557ABE76E6F6312F9FC31C85B7D1C" box="[151,266,1013,1038]" pageId="1" pageNumber="525" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
such furrows are absent and generally more spheres are present then (see also discussion). The sternum of all
<taxonomicName id="4C024D41E76E6F6312B1FBDEC86A7D23" authorityName="Simon" authorityYear="1897" box="[223,315,1050,1073]" class="Arachnida" family="Clubionidae" genus="Systaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="525" phylum="Arthropoda" rank="genus">
<emphasis id="B976EAD0E76E6F6312B1FBDEC86A7D23" box="[223,315,1050,1073]" italics="true" pageId="1" pageNumber="525">Systaria</emphasis>
</taxonomicName>
species treated in this paper exhibit 4 pairs of indistinct humps, the posterior one almost fused. Between the humps 3 pairs of slit sensilla on each side are present, each consisting of 1 large and 1 smaller slit sensillum (
<figureCitation id="13392A47E76E6F63137BFBA5C83D7D68" box="[277,364,1121,1146]" captionStart="FIGURES 3339" captionStartId="16.[151,263,1879,1902]" captionTargetBox="[177,1393,321,1855]" captionTargetId="figure@16.[177,1413,298,1864]" captionTargetPageId="16" captionText="FIGURES 3339. Systaria princesa spec. nov. from Philippines, Palawan (3334, 37 holotype male from Tagusan cave, 35 36 male paratype from Puerto Princesa Underground River cave, 3839 female paratype from Tagusan Cave). 3336 Left palp (33, 35 ventral, 34, 36 retrolateral). 37 Sternum, ventral. 38 Epigyne, ventral. 39 Vulva, dorsal. For explanations of a, b see diagnosis. Abbreviation: CO—copulatory opening." httpUri="https://zenodo.org/record/2606654/files/figure.png" pageId="1" pageNumber="525">Fig. 35</figureCitation>
). The distal half of metatarsi and tarsi exhibit sparse scopula ventrally, spinnerets have long setae, posterior lateral spinnerets exhibit the distal segment distinctly longer than in remaining spinnerets (
<figureCitation id="13392A47E76E6F631752FB41CCDE7D8C" box="[1340,1423,1157,1182]" captionStart="FIGURES 1115" captionStartId="5.[151,263,1396,1419]" captionTargetBox="[151,1435,193,1383]" captionTargetId="figure@5.[151,1435,193,1383]" captionTargetPageId="5" captionText="FIGURES 1115. Systaria longinqua spec. nov., holotype female from N Laos, Tham Pathok. 11 Epigyne, ventral. 12 Vulva, dorsal. 13 Left chelicera, ventral. 14 Prosoma, dorsal. 15 Spinnerets, ventral (not all setae drawn). Abbreviation: COcopulatory opening." httpUri="https://zenodo.org/record/2606640/files/figure.png" pageId="1" pageNumber="525">Fig. 15</figureCitation>
). Claw tufts are dense.
</paragraph>
</subSubSection>
</treatment>
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