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<mods:title id="59B70A93C05F2D8C35CC6A6CBB56433B">Revision of the genus Porcellionides Miers, 1877 (Isopoda: Porcellionidae) in the Ibero-Balearic region</mods:title>
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<mods:namePart id="2A4F5A89369515E4EEAA7609580E8CA0">Cifuentes, Julio</mods:namePart>
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<mods:affiliation id="521D2FCC9C370A529790FE2EB3912CE1">Departamento de Biología (Zoología), Facultad de Ciencias, Universidad Autónoma de Madrid, 28049 Cantoblanco, Madrid, Spain.</mods:affiliation>
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<mods:namePart id="390365748B2B4C34D65CC3F2AEBB3C3C">Da Silva, Luís P.</mods:namePart>
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<mods:affiliation id="B9F4E65D1B0C56E59B1D26860388A809">CIBIO, Centro de Investigação em Biodiversidade e Recursos Genéticos, InBIO Laboratório Associado, Campus de Vairão, Universidade do Porto, 4485 - 661 Vairão, Portugal. &amp; BIOPOLIS Program in Genomics, Biodiversity and Land Planning, CIBIO, Campus de Vairão, 4485 - 661 Vairão, Portugal.</mods:affiliation>
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<subSubSection id="C3186558FF9EFFBF3464FD8FFBE4FDD2" box="[538,1049,536,563]" pageId="35" pageNumber="36" type="nomenclature">
<paragraph id="8BBD36D3FF9EFFBF3464FD8FFBE4FDD2" blockId="35.[538,1049,536,563]" box="[538,1049,536,563]" pageId="35" pageNumber="36">
<heading id="D0F581BFFF9EFFBF3464FD8FFBE4FDD2" box="[538,1049,536,563]" centered="true" fontSize="11" level="2" pageId="35" pageNumber="36" reason="2">
<taxonomicName id="4C024D50FF9EFFBF3464FD8FFBE4FDD2" ID-CoL="4LWK6" authority="(Dollfus, 1892)" baseAuthorityName="Dollfus" baseAuthorityYear="1892" box="[538,1049,536,563]" class="Malacostraca" family="Porcellionidae" genus="Porcellionides" kingdom="Animalia" order="Isopoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="rufocinctus">
<emphasis id="B976EAC1FF9EFFBF3464FD8FFCA0FDD3" bold="true" box="[538,861,536,563]" italics="true" pageId="35" pageNumber="36">Porcellionides rufocinctus</emphasis>
(
<bibRefCitation id="EF934B22FF9EFFBF3512FD8EFBECFDD2" author="Dollfus A." box="[876,1041,537,563]" pageId="35" pageNumber="36" pagination="161 - 190" refId="ref24087" refString="Dollfus A. 1892. Catalogue raisonne des isopodes terrestres de l'Espagne. Anales de la Sociedad espanola de Historia natural 21: 161 - 190." type="journal article" year="1892">Dollfus, 1892</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
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<paragraph id="8BBD36D3FF9EFFBF36C3FDCDFD16FD95" blockId="35.[189,747,602,628]" box="[189,747,602,628]" pageId="35" pageNumber="36">
<treatmentCitationGroup id="AB1211FDFF9EFFBF36C3FDCDFD16FD95" box="[189,747,602,628]" pageId="35" pageNumber="36">
<treatmentCitation id="0AA310C2FF9EFFBF36C3FDCDFD1BFD95" author="Dollfus A." box="[189,742,602,628]" page="184" pageId="35" pageNumber="36" year="1892">
<taxonomicName id="4C024D50FF9EFFBF36C3FDCDFD1BFD95" ID-CoL="42HK8" authority="Dollfus, 1892: 184" authorityName="Dollfus" authorityPageNumber="184" authorityYear="1892" box="[189,742,602,628]" class="Malacostraca" family="Porcellionidae" genus="Metoponorthus" kingdom="Animalia" order="Isopoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="rufocinctus">
<emphasis id="B976EAC1FF9EFFBF36C3FDCDFE02FD95" box="[189,511,602,628]" italics="true" pageId="35" pageNumber="36">Metoponorthus rufocinctus</emphasis>
<bibRefCitation id="EF934B22FF9EFFBF3479FDCDFD1BFD95" author="Dollfus A." box="[519,742,602,628]" pageId="35" pageNumber="36" pagination="161 - 190" refId="ref24087" refString="Dollfus A. 1892. Catalogue raisonne des isopodes terrestres de l'Espagne. Anales de la Sociedad espanola de Historia natural 21: 161 - 190." type="journal article" year="1892">Dollfus, 1892: 184</bibRefCitation>
</taxonomicName>
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.
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<subSubSection id="C3186558FF9EFFBF36C3FD0BFCB4F936" pageId="35" pageNumber="36" type="discussion">
<paragraph id="8BBD36D3FF9EFFBF36C3FD0BFECDFD57" blockId="35.[189,304,668,694]" box="[189,304,668,694]" pageId="35" pageNumber="36">
<heading id="D0F581BFFF9EFFBF36C3FD0BFECDFD57" bold="true" box="[189,304,668,694]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
<emphasis id="B976EAC1FF9EFFBF36C3FD0BFECDFD57" bold="true" box="[189,304,668,694]" pageId="35" pageNumber="36">Remarks</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36D3FF9EFFBF36C3FD5DFDD3FB6A" blockId="35.[188,1399,714,1163]" pageId="35" pageNumber="36">
<taxonomicName id="4C024D50FF9EFFBF36C3FD5DFE0BFD05" baseAuthorityName="Dollfus" baseAuthorityYear="1892" box="[189,502,714,740]" class="Malacostraca" family="Porcellionidae" genus="Porcellionides" kingdom="Animalia" order="Isopoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="rufocinctus">
<emphasis id="B976EAC1FF9EFFBF36C3FD5DFE0BFD05" box="[189,502,714,740]" italics="true" pageId="35" pageNumber="36">Porcellionides rufocinctus</emphasis>
</taxonomicName>
was described by
<bibRefCitation id="EF934B22FF9EFFBF34B1FD5DFC81FD05" author="Dollfus A." box="[719,892,714,740]" pageId="35" pageNumber="36" pagination="161 - 190" refId="ref24087" refString="Dollfus A. 1892. Catalogue raisonne des isopodes terrestres de l'Espagne. Anales de la Sociedad espanola de Historia natural 21: 161 - 190." type="journal article" year="1892">Dollfus (1892)</bibRefCitation>
in his work on terrestrial isopods of
<collectingCountry id="F3157643FF9EFFBF3352FD5DFA8EFD05" box="[1324,1395,714,740]" name="Spain" pageId="35" pageNumber="36">Spain</collectingCountry>
. The description was based on a male specimen deposited in the Museo Nacional de Ciencias de
<collectingRegion id="49C6F831FF9EFFBF335EFD7AFA8BFCE6" box="[1312,1398,749,775]" country="Spain" name="Madrid" pageId="35" pageNumber="36">Madrid</collectingRegion>
(MNCN), collected by Bolívar in Villa Rutis, officially known as Santa María de Rutis, belonging to the municipality of Culleredo in the province of Coruña. Later, the same author (1893) added another locality for the species, San Roque, situated close to the first location and within the municipality of Coruña in the same province. Within the MNCN collection, there is a specimen with the following data: “Coruña, Villa Rutis, 1893,
<specimenCount id="9D04FD5AFF9EFFBF3479FC0AFDA4FC56" box="[519,601,925,951]" count="1" pageId="35" pageNumber="36" type="male">1 male</specimenCount>
, Bolívar, MNCN 20.04/11518”. Although the date of this specimen does not match Dollfuss original description (1892), the remaining data is consistent. This specimen was stored with the code MNCN 20.04/7895, former number 86, alongside specimens of
<taxonomicName id="4C024D50FF9EFFBF3369FC73FE2DFBC3" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" class="Malacostraca" family="Oniscidae" genus="Oniscus" kingdom="Animalia" order="Isopoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="asellus">
<emphasis id="B976EAC1FF9EFFBF3369FC73FEF2FBC0" italics="true" pageId="35" pageNumber="36">Oniscus asellus</emphasis>
Linnaeus, 1758
</taxonomicName>
,
<taxonomicName id="4C024D50FF9EFFBF37A0FB90FC9DFBC0" authority="Brandt, 1833" authorityName="Brandt" authorityYear="1833" box="[478,864,1031,1057]" class="Malacostraca" family="Porcellionidae" genus="Porcellio" kingdom="Animalia" order="Isopoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="dilatatus">
<emphasis id="B976EAC1FF9EFFBF37A0FB90FD41FBC0" box="[478,700,1031,1057]" italics="true" pageId="35" pageNumber="36">Porcellio dilatatus</emphasis>
Brandt, 1833
</taxonomicName>
,
<taxonomicName id="4C024D50FF9EFFBF3510FB9FFB34FBC0" authority="Vandel, 1946" authorityName="Vandel" authorityYear="1946" box="[878,1225,1031,1057]" class="Malacostraca" family="Porcellionidae" genus="Porcellio" kingdom="Animalia" order="Isopoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="herminiensis">
<emphasis id="B976EAC1FF9EFFBF3510FB9FFBD8FBC0" box="[878,1061,1031,1057]" italics="true" pageId="35" pageNumber="36">P. herminiensis</emphasis>
<bibRefCitation id="EF934B22FF9EFFBF3255FB90FB34FBC0" author="Vandel A." box="[1067,1225,1031,1057]" pageId="35" pageNumber="36" pagination="135 - 427" refId="ref25660" refString="Vandel A. 1946. Crustaces isopodes terrestres (Oniscoidea) epiges et cavernicoles du Portugal. Anais da Faculdade de Ciencias do Porto 30: 135 - 427." type="journal article" year="1946">Vandel, 1946</bibRefCitation>
</taxonomicName>
, and
<taxonomicName id="4C024D50FF9EFFBF3374FB9FFE92FBA5" authority="Latreille, 1804" authorityName="Latreille" authorityYear="1804" class="Malacostraca" family="Porcellionidae" genus="Porcellio" kingdom="Animalia" order="Isopoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="scaber">
<emphasis id="B976EAC1FF9EFFBF3374FB9FFA8AFBC0" box="[1290,1399,1031,1057]" italics="true" pageId="35" pageNumber="36">P. scaber</emphasis>
Latreille, 1804
</taxonomicName>
. Currently, the specimen has undergone discoloration due to its extended preservation period in alcohol. However, we identified it as
<taxonomicName id="4C024D50FF9EFFBF3492FBDAFAD6FB89" authority="(Budde-Lund, 1885)" baseAuthorityName="Budde-Lund" baseAuthorityYear="1885" box="[748,1323,1101,1128]" class="Malacostraca" family="Porcellionidae" genus="Porcellionides" kingdom="Animalia" order="Isopoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="sexfasciatus">
<emphasis id="B976EAC1FF9EFFBF3492FBDAFBCDFB86" box="[748,1072,1101,1127]" italics="true" pageId="35" pageNumber="36">Porcellionides sexfasciatus</emphasis>
(
<bibRefCitation id="EF934B22FF9EFFBF323FFBDAFADEFB89" author="Budde-Lund G." box="[1089,1315,1101,1128]" pageId="35" pageNumber="36" refId="ref23578" refString="Budde-Lund G. 1885. Crustacea Isopoda Terrestria per Familias et Genera et Species Descripta. Nielsen &amp; Lydiche, Copenhagen. https: // doi. org / 10.5962 / bhl. title. 109769" type="book" year="1885">Budde-Lund, 1885</bibRefCitation>
)
</taxonomicName>
based on the exopod of the pleopod I.
</paragraph>
<paragraph id="8BBD36D3FF9EFFBF36C3FB25FA8AF993" blockId="35.[187,1399,1202,1651]" pageId="35" pageNumber="36">
The original description of
<taxonomicName id="4C024D50FF9EFFBF3784FB24FD66FB2D" baseAuthorityName="Dollfus" baseAuthorityYear="1892" box="[506,667,1202,1228]" class="Malacostraca" family="Porcellionidae" genus="Porcellionides" kingdom="Animalia" order="Isopoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="rufocinctus">
<emphasis id="B976EAC1FF9EFFBF3784FB24FD66FB2D" box="[506,667,1202,1228]" italics="true" pageId="35" pageNumber="36">P. rufocinctus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EF934B22FF9EFFBF34B9FB25FC8EFB2D" author="Dollfus A." box="[711,883,1202,1228]" pageId="35" pageNumber="36" pagination="161 - 190" refId="ref24087" refString="Dollfus A. 1892. Catalogue raisonne des isopodes terrestres de l'Espagne. Anales de la Sociedad espanola de Historia natural 21: 161 - 190." type="journal article" year="1892">Dollfus (1892)</bibRefCitation>
followed the common approach of the time, focusing on the coloration and morphology of the antennae, cephalon, pereon, pleon, telson, and uropods. In closely related species, these characters often prove insufficient for establishing specific identity. Absent are mentions of other now significant characters, such as pleopod and pereopod morphology, as well as certain integumentary characters like glandular fields or scale-setae. Nonetheless, Dollfus (1982) provided the following description: oval body, quite convex, pleon noticeably retracted (the body shape is exactly that of
<taxonomicName id="4C024D50FF9EFFBF345AFA12FCE4FA7E" baseAuthorityName="Scopoli" baseAuthorityYear="1763" box="[548,793,1413,1439]" class="Malacostraca" family="Philosciidae" genus="Philoscia" kingdom="Animalia" order="Isopoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="muscorum">
<emphasis id="B976EAC1FF9EFFBF345AFA12FCE4FA7E" box="[548,793,1413,1439]" italics="true" pageId="35" pageNumber="36">Philoscia muscorum</emphasis>
</taxonomicName>
); cephalon with a frontal line finely but distinctly bordered, lateral lobes very short and forehead almost flat; antennae reaching approximately half the body length, flagellum segments equal. Dollfus (1983) observed regarding the San Roque specimens that their coloration resembles that of
<taxonomicName id="4C024D50FF9EFFBF34F9FA78FCC8F9E8" baseAuthorityName="Budde-Lund" baseAuthorityYear="1885" box="[647,821,1519,1545]" class="Malacostraca" family="Porcellionidae" genus="Porcellionides" kingdom="Animalia" order="Isopoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="sexfasciatus">
<emphasis id="B976EAC1FF9EFFBF34F9FA78FCC8F9E8" box="[647,821,1519,1545]" italics="true" pageId="35" pageNumber="36">P. sexfasciatus</emphasis>
</taxonomicName>
but
<taxonomicName id="4C024D50FF9EFFBF3513FA78FBECF9E8" baseAuthorityName="Dollfus" baseAuthorityYear="1892" box="[877,1041,1519,1545]" class="Malacostraca" family="Porcellionidae" genus="Porcellionides" kingdom="Animalia" order="Isopoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="rufocinctus">
<emphasis id="B976EAC1FF9EFFBF3513FA78FBECF9E8" box="[877,1041,1519,1545]" italics="true" pageId="35" pageNumber="36">P. rufocinctus</emphasis>
</taxonomicName>
is cleary distinguished by the flat forehead and the much shorter pleotelson. All these characters could also correspond to
<taxonomicName id="4C024D50FF9EFFBF3285F984FA92F9CD" baseAuthorityName="Verhoeff" baseAuthorityYear="1901" box="[1275,1391,1554,1580]" class="Malacostraca" family="Porcellionidae" genus="Porcellionides" kingdom="Animalia" order="Isopoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="molleri">
<emphasis id="B976EAC1FF9EFFBF3285F984FA92F9CD" box="[1275,1391,1554,1580]" italics="true" pageId="35" pageNumber="36">P. molleri</emphasis>
</taxonomicName>
, as mentioned in the description provided above, as well as other species within this genus, and even certain specimens of
<taxonomicName id="4C024D50FF9EFFBF37CEF9CEFDA6F993" baseAuthorityName="Budde-Lund" baseAuthorityYear="1885" box="[432,603,1624,1650]" class="Malacostraca" family="Porcellionidae" genus="Porcellionides" kingdom="Animalia" order="Isopoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="sexfasciatus">
<emphasis id="B976EAC1FF9EFFBF37CEF9CEFDA6F993" box="[432,603,1624,1650]" italics="true" pageId="35" pageNumber="36">P. sexfasciatus</emphasis>
</taxonomicName>
. The latter aligns with our analysis of the specimen from Villa Rutis.
</paragraph>
<paragraph id="8BBD36D3FF9EFFBF36C3F90DFCB4F936" blockId="35.[189,1399,1690,1751]" pageId="35" pageNumber="36">
Therefore, at least for now, we classify this as species inquirenda, because it is not possible to definitively differentiate it from the other species of
<taxonomicName id="4C024D50FF9EFFBF34E8F92AFCBFF936" authorityName="Miers" authorityYear="1877" box="[662,834,1725,1751]" class="Malacostraca" family="Porcellionidae" genus="Porcellionides" kingdom="Animalia" order="Isopoda" pageId="35" pageNumber="36" phylum="Arthropoda" rank="genus">
<emphasis id="B976EAC1FF9EFFBF34E8F92AFCBFF936" box="[662,834,1725,1751]" italics="true" pageId="35" pageNumber="36">Porcellionides</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>