plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/03/AB/87/03AB87BA2D1FA22DE41FF9887B18FC95.xml
maintenance bc2564723d added 03
2024-06-21 12:22:17 +02:00

468 lines
71 KiB
XML
Raw Blame History

This file contains invisible Unicode characters

This file contains invisible Unicode characters that are indistinguishable to humans but may be processed differently by a computer. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="1BA57AD5531000A2BAC6E6179DBF8571" ID-DOI="10.11646/zootaxa.1074.1.1" ID-ISSN="1175­5334" ID-Zenodo-Dep="5050779" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1625108368558" checkinUser="felipe" docAuthor="Ponder, W. F., Clark, S. A., Eberhard, S. &amp; Studdert, J. B." docDate="2005" docId="03AB87BA2D1FA22DE41FF9887B18FC95" docLanguage="en" docName="zt01074p066.pdf" docOrigin="Zootaxa 1074 (1)" docSource="https://biotaxa.org/Zootaxa/article/view/zootaxa.1074.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Pseudotricula conica Ponder &amp; Clark &amp; Eberhard &amp; Studdert 2005, n.sp." docType="treatment" docVersion="4" lastPageNumber="34" masterDocId="FF92FFC22D02A20CE517FFCC7E3DFF83" masterDocTitle="A radiation of hydrobiid snails in the caves and streams at Precipitous Bluff southwest Tasmania, Australia (Mollusca: Caenogastropoda: Rissooidea: Hydrobiidae s. l.)," masterLastPageNumber="66" masterPageNumber="1" pageNumber="30" updateTime="1698981223375" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="D9A9B555EBAB24C0CCE14E970271B2BE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="87D17CBAAA0299A305CA52AFAA53F19F">
<mods:title id="D60514008624E3114708F733A49DD7B6">A radiation of hydrobiid snails in the caves and streams at Precipitous Bluff southwest Tasmania, Australia (Mollusca: Caenogastropoda: Rissooidea: Hydrobiidae s. l.),</mods:title>
</mods:titleInfo>
<mods:name id="1667684E7072CE3BB322080DD9710219" type="personal">
<mods:role id="E1FF2599674E372DF78B26EDC61C4A21">
<mods:roleTerm id="625EEDBF18A124411C93163FEEEF1710">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F50E28341A06B25DD2B2D474A52CE28F">Ponder, W. F.</mods:namePart>
<mods:affiliation id="ECCEDE6F83C21516F7EDBE1954B8122C">Australian Museum, 6 College Street, Sydney, NSW, 2010, Australia. Email: winstonp @ austmus. gov. au Previously Australian Museum, now Department of Biological Sciences, Biodiversity and Systematics. University of Alabama, Box 870345, Tuscaloosa, AL 35487, United States of America Department of Conservation and Land Management, Science Division, PO Box 51, Wanneroo, WA 6065, Australia</mods:affiliation>
</mods:name>
<mods:name id="EBEA9FA863D27E0AFB2697498A939C50" type="personal">
<mods:role id="B23A749C0FBE16FED6950F6B4A528B60">
<mods:roleTerm id="408D8269D4F35A9B95CA0C52160A55F1">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="E28C19B0E8D6FFCABEEACC2741772F4E">Clark, S. A.</mods:namePart>
</mods:name>
<mods:name id="0767958FC2D15C1E8203B7C6C644E7BC" type="personal">
<mods:role id="D0C49C313C1199BD44B3915ED4C4B027">
<mods:roleTerm id="E6CD08F1E9B15DB3FB7B8586C984193B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6F5D0659278383B9C57587280579F20C">Eberhard, S.</mods:namePart>
</mods:name>
<mods:name id="FD94A47532C564B86A5148D563C83EC1" type="personal">
<mods:role id="EE66864F8562B5B1ADC01874AD505BFE">
<mods:roleTerm id="EDE317795B916CBEE6C3DC71778C87F3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="504D1906EEF7425DEC7373B251677649">Studdert, J. B.</mods:namePart>
<mods:affiliation id="813C3B5F7B9E553E3547C41BFE1B485A">Australian Museum, 6 College Street, Sydney, NSW, 2010, Australia. Email: winstonp @ austmus. gov. au Previously Australian Museum, now Department of Biological Sciences, Biodiversity and Systematics. University of Alabama, Box 870345, Tuscaloosa, AL 35487, United States of America Department of Conservation and Land Management, Science Division, PO Box 51, Wanneroo, WA 6065, Australia</mods:affiliation>
</mods:name>
<mods:typeOfResource id="0A61E8BD945D2F33F8EBD565C33C3F72">text</mods:typeOfResource>
<mods:relatedItem id="9AB45A4AA589EA15401D70777C78C2F4" type="host">
<mods:titleInfo id="06069029C6147BA31136DDA03CC1E0B2">
<mods:title id="6C62CAEE20603FC153508657C1DA41CF">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="979BB33A4D6FB0959527715DE482AAEE">
<mods:date id="2501728FF651DC64A02DCA687CBB82A0">2005</mods:date>
<mods:detail id="40199E8CF0CB9134A36DE8BBE6DD8532" type="pubDate">
<mods:number id="8AC9FE22D5B8668DCD319242737CC168">2005-11-01</mods:number>
</mods:detail>
<mods:detail id="4E63DE79C0A2C5EF5D958B25915035B7" type="volume">
<mods:number id="3873E3908176D2BE9766DBDF6D9DEA12">1074</mods:number>
</mods:detail>
<mods:detail id="B547FED00A719ACC78079A541CDB2E62" type="issue">
<mods:number id="717F7C1E4D9D48BFCA376C711A8ABA21">1</mods:number>
</mods:detail>
<mods:extent id="AB3CE42B0E7129F6FAF06D411479C5D5" unit="page">
<mods:start id="C2A1C8FB657A39AE12B897A282F7DB53">1</mods:start>
<mods:end id="E8CA6FF3D5208B8E59245BED84A405F9">66</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location id="E369B2EBD7E35FD4735F9036D1DA405B">
<mods:url id="9AAC5FFDC1A7E3CA41F47D1660F7790E">https://biotaxa.org/Zootaxa/article/view/zootaxa.1074.1.1</mods:url>
</mods:location>
<mods:classification id="8E1DD621A6ECA1465D36630A81ED4C89">journal article</mods:classification>
<mods:identifier id="5CA29AD1BE0C684FD2F876D3CCDF9E43" type="DOI">10.11646/zootaxa.1074.1.1</mods:identifier>
<mods:identifier id="42C92B3457DB0E3BB37EFD843CCB8F8C" type="ISSN">1175­5334</mods:identifier>
<mods:identifier id="D1FF59981E39B5CE0ECCE3083461EECF" type="Zenodo-Dep">5050779</mods:identifier>
<mods:identifier id="2A7D1E3BC08274353EC4E05CF8A41E28" type="ZooBank">8C36619A-8876-40C0-BA06-60AE4449DD49</mods:identifier>
</mods:mods>
<treatment id="03AB87BA2D1FA22DE41FF9887B18FC95" ID-DOI="http://doi.org/10.5281/zenodo.5053151" ID-Zenodo-Dep="5053151" LSID="urn:lsid:plazi:treatment:03AB87BA2D1FA22DE41FF9887B18FC95" httpUri="http://treatment.plazi.org/id/03AB87BA2D1FA22DE41FF9887B18FC95" lastPageId="33" lastPageNumber="34" pageId="29" pageNumber="30">
<subSubSection id="C31865272D1FA211E41FF9887C74F9DD" box="[264,585,1604,1630]" pageId="29" pageNumber="30" type="nomenclature">
<paragraph id="8BBD36AC2D1FA211E41FF9887C74F9DD" blockId="29.[264,1082,1604,1665]" box="[264,585,1604,1630]" pageId="29" pageNumber="30">
<heading id="D0F581C02D1FA211E41FF9887C74F9DD" bold="true" box="[264,585,1604,1630]" fontSize="11" level="1" pageId="29" pageNumber="30" reason="1">
<emphasis id="B976EABE2D1FA211E41FF9887C74F9DD" bold="true" box="[264,585,1604,1630]" pageId="29" pageNumber="30">
<taxonomicName id="4C024D2F2D1FA211E41FF9887C3AF9DD" authority="Ponder &amp; Clark &amp; Eberhard &amp; Studdert, 2005" authorityName="Ponder &amp; Clark &amp; Eberhard &amp; Studdert" authorityYear="2005" box="[264,519,1604,1630]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="29" pageNumber="30" phylum="Mollusca" rank="species" species="conica" status="sp. nov.">
<emphasis id="B976EABE2D1FA211E41FF9887C3AF9DD" bold="true" box="[264,519,1604,1630]" italics="true" pageId="29" pageNumber="30">Pseudotricula conica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24557C52D1FA211E719F9887C74F9DD" box="[526,585,1604,1630]" pageId="29" pageNumber="30" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C31865272D1FA22CE41FF9AB7DABF955" lastPageId="32" lastPageNumber="33" pageId="29" pageNumber="30" type="description">
<paragraph id="8BBD36AC2D1FA211E41FF9AB7A07F902" blockId="29.[264,1082,1604,1665]" box="[264,1082,1639,1665]" pageId="29" pageNumber="30">
<figureCitation id="13392A292D1FA211E41FF9AB7F99F902" box="[264,420,1639,1665]" captionStart="FIGURE 3" captionStartId="13.[264,357,1408,1429]" captionTargetBox="[319,1271,284,1386]" captionTargetId="figure-152@13.[311,1275,280,1392]" captionTargetPageId="13" captionText="FIGURE 3: Radulae of Pseudotricula species. A, B, Pseudotricula eberhardi, C.165053, Black Curtain Streamway, Cueva Blanca; C–H, Pseudotricula expandolabra, C, D, C.201495, Persephone Streamway, Bauhaus, E, G, H, C.201811, Persephone Pot, Bauhaus, F, C.201822, main streamway, Bauhaus; I, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; J–L, Pseudotricula conica, C.203676, paratype, Cane Toad Abuse Streamway, Damper Cave. A, C, G, K, central, lateral inner and outer marginal teeth; B, E, L, central teeth; D, outer marginal teeth; F, J, lateral and inner marginal teeth; H, lateral tooth; I, central and lateral teeth. Scales: A, B, E, H–J, L 10 m; C, F, G, K, 20 m; D, 5 m." figureDoi="http://doi.org/10.5281/zenodo.5050787" httpUri="https://zenodo.org/record/5050787/files/figure.png" pageId="29" pageNumber="30">Figures 3J–L</figureCitation>
;
<figureCitation id="13392A292D1FA211E4A7F9AB7FE9F902" box="[432,468,1639,1665]" captionStart="FIGURE 4" captionStartId="14.[264,357,1822,1843]" captionTargetBox="[406,1181,963,1757]" captionTargetId="figure-174@14.[378,1210,945,1795]" captionTargetPageId="14" captionText="FIGURE 4. Dorsal views of penes of Pseudotricula and Nanocochlea species. A,. Pseudotricula eberhardi; B, Pseudotricula expandolabra; C, Nanocochlea stylesae; D, Nanocochlea damperensis; E, Pseudotricula conica. Scales A, B, E, 0.2 mm; C, D, 0.1 mm. Abbreviations: ds, distal swelling; pd, penial duct." figureDoi="http://doi.org/10.5281/zenodo.5050791" httpUri="https://zenodo.org/record/5050791/files/figure.png" pageId="29" pageNumber="30">4E</figureCitation>
;
<figureCitation id="13392A292D1FA211E4F7F9AB7C1BF902" box="[480,550,1639,1665]" captionStart="FIGURE 6" captionStartId="16.[264,357,1641,1662]" captionTargetBox="[302,1285,329,1616]" captionTargetId="figure-146@16.[302,1285,329,1633]" captionTargetPageId="16" captionText="FIGURE 6. Protoconchs and teleoconch microsculpture of Pseudotricula and Nanocochlea species. A–D, Pseudotricula expandolabra, A, B, D, C.201493, paratype, main streamway, Bauhaus, C, C.201495, Persephone Streamway, Bauhaus; E–G, Pseudotricula conica, E, F, C.201464, Cane Toad Abuse Streamway, Damper Cave, G, Persephone Pot, Bauhaus; Honey and Cream Streamway, Damper Cave; H, Nanocochlea exigua, C.201475; I, J, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; K–M, Nanocochlea damperensis, K, L, C.203685, paratypes, first crossing, Damper Creek, M, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; N, O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, E, F, H, K–N, protoconch; C, D, G, I, J, O, microsculpture. Scales: A, C–E, J, K, M, O, 200 m; B, F–I, L, N, 100 m." figureDoi="http://doi.org/10.5281/zenodo.5050797" httpUri="https://zenodo.org/record/5050797/files/figure.png" pageId="29" pageNumber="30">6E–G</figureCitation>
;
<figureCitation id="13392A292D1FA211E724F9AB7C56F902" box="[563,619,1639,1665]" captionStart="FIGURE 7" captionStartId="19.[264,357,1755,1776]" captionTargetBox="[369,1242,439,1733]" captionTargetId="figure-114@19.[317,1269,433,1748]" captionTargetPageId="19" captionText="FIGURE 7. Opercula of Pseudotricula species. A, B, Pseudotricula expandolabra, C.201493, paratypes, main streamway, Bauhaus; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula arthurclarkei, C.203671, paratypes, Quetzalcoatl Conduit; F–I, Pseudotricula conica, F, G, C.203680, main streamway, near entrance, Damper Cave; H, I, C.201459, Screaming Stals streamway, Bauhaus; A, D, F, H, outer side; B, C, E, H, I, inner side; Scales: A–I, 200 m." figureDoi="http://doi.org/10.5281/zenodo.5050801" httpUri="https://zenodo.org/record/5050801/files/figure.png" pageId="29" pageNumber="30">7F–I</figureCitation>
;
<figureCitation id="13392A292D1FA211E76FF9AB7C82F902" box="[632,703,1639,1665]" captionStart="FIGURE 8" captionStartId="20.[264,357,1612,1633]" captionTargetBox="[341,1270,339,1585]" captionTargetId="figure-146@20.[299,1287,324,1608]" captionTargetPageId="20" captionText="FIGURE 8. Pallial cavities of male specimens of Pseudotricula and Nanocochlea species. A, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula conica, D, C.203680, main streamway near entrance, Damper Cave, E, Cane Toad Abuse Streamway, Damper Cave; F, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; G, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: A–D, F–H, 500 m, E, 1 mm. Abbreviations: a, anus; ct, ctenidium; ev, efferent vein; os, osphradium; pc, pericardium; pe, pallial edge; ppw, posterior pallial wall; r, rectum; rg, renal gland. The stippled area represents the hypobranchial gland." figureDoi="http://doi.org/10.5281/zenodo.5050803" httpUri="https://zenodo.org/record/5050803/files/figure.png" pageId="29" pageNumber="30">8D, E</figureCitation>
;
<figureCitation id="13392A292D1FA211E7DCF9AB7D2BF902" box="[715,790,1639,1665]" captionStart="FIGURE 9" captionStartId="21.[264,357,1723,1744]" captionTargetBox="[350,1309,380,1631]" captionTargetId="figure-97@21.[299,1342,370,1653]" captionTargetPageId="21" captionText="FIGURE 9. Head, penis and stomach of Pseudotricula species. A, D, Pseudotricula conica, C.203680, main streamway near entrance, Damper Cave; C, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, E, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, head and penis; C–E, stomach. Scales: A–E, 500 m. Abbreviations: dgo, opening from stomach to digestive gland; ey, eye; int, intestine; oes, oesophagus, p, penis; pg, penial duck; sn, snout; ss, style sack; st, stomach; tn, tentacle." figureDoi="http://doi.org/10.5281/zenodo.5050807" httpUri="https://zenodo.org/record/5050807/files/figure.png" pageId="29" pageNumber="30">9A, D</figureCitation>
;
<figureCitation id="13392A292D1FA211E634F9AB7D49F902" box="[803,884,1639,1665]" captionStart="FIGURE 10" captionStartId="22.[264,357,1296,1317]" captionTargetBox="[279,1315,341,1234]" captionTargetId="figure-191@22.[267,1319,329,1269]" captionTargetPageId="22" captionText="FIGURE 10. Female reproductive systems of Pseudotricula species. A ­ C, Pseudotricula expandolabra, A, B, C.201493, paratype, Persephone Streamway, Bauhaus, C, C.201815, Persephone Streamway, Bauhaus; D, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; E, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; F, G, Pseudotricula conica, F, C203680, main streamway near entrance, Damper Cave, G, Cane Toad Abuse Streamway, Damper Cave. Scale: 500 m. Abbreviations: ag, albumen gland; bc, bursa copulatrix; bd, bursal duct; cg, capsule gland; co, coiled oviduct; po, genital opening; pw, pallial wall; sr, seminal receptacle; vc, ventral channel." figureDoi="http://doi.org/10.5281/zenodo.5050809" httpUri="https://zenodo.org/record/5050809/files/figure.png" pageId="29" pageNumber="30">10F, G</figureCitation>
;
<figureCitation id="13392A292D1FA211E696F9AB7DE5F902" box="[897,984,1639,1665]" captionStart="FIGURE 12" captionStartId="28.[264,357,1479,1500]" captionTargetBox="[382,1170,289,1464]" captionTargetId="figure-168@28.[354,1233,269,1476]" captionTargetPageId="28" captionText="FIGURE 12. Shells of Pseudotricula species. A, Pseudotricula arthurclarkei, holotype, C.439656, Quetzalcoatl Conduit; B ­ D, Pseudotricula conica, B, holotype, Cane Toad Abuse Streamway, Damper Cave; C, C.203680, main streamway near entrance, Damper Cave; D, C.201813, Persephone Pot, Bauhaus. Scales: A–D, 500 m." figureDoi="http://doi.org/10.5281/zenodo.5050815" httpUri="https://zenodo.org/record/5050815/files/figure.png" pageId="29" pageNumber="30">12B–D</figureCitation>
;
<figureCitation id="13392A292D1FA211E6F3F9AB7A08F902" box="[996,1077,1639,1665]" captionStart="FIGURE 13" captionStartId="30.[264,357,1326,1347]" captionTargetBox="[393,1223,370,1252]" captionTargetId="figure-178@30.[348,1252,362,1266]" captionTargetPageId="30" captionText="FIGURE 13. Shells of Pseudotricula species. A ­ F, Pseudotricula conica, A, C.203694, Black Curtain streamway, Cueva Blanca, B, Screaming Stals streamway, Bauhaus; C, C.201813, Persephone Pot, Bauhaus, D, C.203464, Cane Toad Abuse Streamway, Damper Cave, E, C.201488, main streamway, Damper Cave, F, C.203692, Damper Cave. Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5050817" httpUri="https://zenodo.org/record/5050817/files/figure.png" pageId="29" pageNumber="30">13A–F</figureCitation>
.
</paragraph>
<paragraph id="8BBD36AC2D1FA211E41FF97B7F94F953" blockId="29.[264,1323,1719,1905]" box="[264,425,1719,1744]" pageId="29" pageNumber="30">
<emphasis id="B976EABE2D1FA211E41FF97B7F94F953" box="[264,425,1719,1744]" italics="true" pageId="29" pageNumber="30">Type material</emphasis>
</paragraph>
<paragraph id="8BBD36AC2D1FA211E42FF9137F9AF8A2" blockId="29.[264,1323,1719,1905]" pageId="29" pageNumber="30">
<materialsCitation id="3B6A3CF12D1FA211E42FF9137F9AF8A2" collectingDate="1994-03-28" collectionCode="AMS" location="Cane Toad Abuse Streamway Damper Cave" pageId="29" pageNumber="30" specimenCount="1" typeStatus="holotype">
<typeStatus id="54B9880E2D1FA211E42FF9137F94F97A" box="[312,425,1759,1785]" pageId="29" pageNumber="30" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED13AE692D1FA211E4A2F9137FC8F97A" box="[437,501,1759,1785]" pageId="29" pageNumber="30">AMS</collectionCode>
C.439649,
<location id="8EDD60772D1FA211E768F9137AB4F97A" LSID="urn:lsid:plazi:treatment:03AB87BA2D1FA22DE41FF9887B18FC95:8EDD60772D1FA211E768F9137AB4F97A" box="[639,1161,1759,1785]" name="Cane Toad Abuse Streamway Damper Cave" pageId="29" pageNumber="30">Cane Toad Abuse Streamway Damper Cave</location>
, stn PB1–2A,
<date id="FFBC106C2D1FA211E41FF8CB7F9FF8A2" box="[264,418,1799,1825]" pageId="29" pageNumber="30" value="1994-03-28">
<collectingDate id="EFF8E9842D1FA211E41FF8CB7F9FF8A2" box="[264,418,1799,1825]" pageId="29" pageNumber="30" value="1994-03-28">28 Mar 1994</collectingDate>
</date>
.
</materialsCitation>
</paragraph>
<paragraph id="8BBD36AC2D1FA211E42FF8E37C5CF8F2" blockId="29.[264,1323,1719,1905]" pageId="29" pageNumber="30">
<materialsCitation id="3B6A3CF12D1FA211E42FF8E37F7DF8F2" collectionCode="AMS" location="Damper Cave" pageId="29" pageNumber="30" specimenCount="1" typeStatus="paratype">
<typeStatus id="54B9880E2D1FA211E42FF8E37F93F8CA" box="[312,430,1839,1865]" pageId="29" pageNumber="30" type="paratype">Paratypes</typeStatus>
:
<collectionCode id="ED13AE692D1FA211E4AAF8E37FC1F8CA" box="[445,508,1839,1865]" pageId="29" pageNumber="30">AMS</collectionCode>
C.203676,
<location id="8EDD60772D1FA211E79AF8E37D0BF8CA" LSID="urn:lsid:plazi:treatment:03AB87BA2D1FA22DE41FF9887B18FC95:8EDD60772D1FA211E79AF8E37D0BF8CA" box="[653,822,1839,1865]" name="Damper Cave" pageId="29" pageNumber="30">Damper Cave</location>
, same locality, stn PB1–2A (10 dry, 20+ wet)
</materialsCitation>
; QVM, 9:20539 (5 wet).
</paragraph>
<caption id="DF7D66242D1CA212E41FFAE27CB7FA27" ID-DOI="http://doi.org/10.5281/zenodo.5050817" ID-Zenodo-Dep="5050817" httpUri="https://zenodo.org/record/5050817/files/figure.png" pageId="30" pageNumber="31" startId="30.[264,357,1326,1347]" targetBox="[393,1223,370,1252]" targetPageId="30">
<paragraph id="8BBD36AC2D1CA212E41FFAE27CB7FA27" blockId="30.[264,1323,1326,1444]" pageId="30" pageNumber="31">
<emphasis id="B976EABE2D1CA212E41FFAE27FB3FAC0" bold="true" box="[264,398,1326,1347]" pageId="30" pageNumber="31">FIGURE 13.</emphasis>
Shells of
<taxonomicName id="4C024D2F2D1CA212E4EBFAE37CC8FAC7" box="[508,757,1327,1348]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="undefined-a">
<emphasis id="B976EABE2D1CA212E4EBFAE37CBFFAC7" box="[508,642,1327,1348]" italics="true" pageId="30" pageNumber="31">Pseudotricula</emphasis>
species. A
</taxonomicName>
­ F,
<taxonomicName id="4C024D2F2D1CA212E639FAE37DC2FAC7" authorityName="Ponder &amp; Clark &amp; Eberhard &amp; Studdert" authorityYear="2005" box="[814,1023,1327,1348]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="30" pageNumber="31" phylum="Mollusca" rank="species" species="conica">
<emphasis id="B976EABE2D1CA212E639FAE37DC2FAC7" box="[814,1023,1327,1348]" italics="true" pageId="30" pageNumber="31">Pseudotricula conica</emphasis>
</taxonomicName>
, A, C.203694, Black Curtain streamway, Cueva Blanca, B, Screaming Stals streamway, Bauhaus; C, C.201813, Persephone Pot, Bauhaus, D, C.203464, Cane Toad Abuse Streamway, Damper Cave, E, C.201488, main streamway, Damper Cave, F, C.203692, Damper Cave. Scales: 1 mm.
</paragraph>
</caption>
<paragraph id="8BBD36AC2D1CA212E41FFA1B7C95FA73" blockId="30.[264,1324,1495,1881]" box="[264,680,1495,1520]" pageId="30" pageNumber="31">
<emphasis id="B976EABE2D1CA212E41FFA1B7C95FA73" box="[264,680,1495,1520]" italics="true" pageId="30" pageNumber="31">Other material examined (all AMS)</emphasis>
</paragraph>
<paragraph id="8BBD36AC2D1CA212E42FFA337FB9F8DA" blockId="30.[264,1324,1495,1881]" pageId="30" pageNumber="31">
<emphasis id="B976EABE2D1CA212E42FFA337FA3F99B" box="[312,414,1535,1560]" italics="true" pageId="30" pageNumber="31">Bauhaus</emphasis>
: C.201823, main streamway, stn PB6­1A, 14 dry); C.201824, same locality, stn PB6­1A (14 dry); C.201460, Screaming Stals streamway, stn 12 (2 dry, 17wet); C.201459, Screaming Stals streamway, stn 11 (12 dry, 20+ wet); C.201469, same locality, stn 13 (3 dry); C.439498, same locality, stn 12 (2 dry); C.201813, Persephone Pot, stn PB17­2a.2, middle and upper streamway (9 dry); C.201817, same locality, stn PB17­2.1, lower streamway (20+ dry, 20+ wet); C.439650, same locality, stn PB17­2a.1 (1dry) figured specimen; C.439496, Persephone Streamway, stn 8 (4 dry); C.439501, same locality, stn 9 (2 dry); C.439499, same locality, stn 9 (1 dry); C.439500, same locality, stn 10 (3 dry).
</paragraph>
<paragraph id="8BBD36AC2D1DA213E42FFED07D17FE7D" blockId="31.[264,1322,284,510]" pageId="31" pageNumber="32">
<emphasis id="B976EABE2D1DA213E42FFED07FE6FEB6" box="[312,475,284,309]" italics="true" pageId="31" pageNumber="32">Damper Cave</emphasis>
: C.203692, stn PB1­1B (1 dry, 3 wet); C.201488, main streamway, stn 6 (9 dry, 16 wet); C.203680, main streamway near entrance, stn PB1­1A (20+ dry, 20+ wet); C.439652, same locality, stn PB1­1A, (1dry), figured specimen; C.201464, Cane Toad Abuse streamway, stn 2 (11 dry, 20+ wet); C.201487, same locality, stn 2 (4 dry, 4 wet); C.201470, stn 2 (6 dry);
<emphasis id="B976EABE2D1DA213E720FE707CE2FE56" box="[567,735,444,469]" italics="true" pageId="31" pageNumber="32">Cueva Blanca</emphasis>
: C.166967, no stn no. (2 dry, 6 wet); C.203694, Black Curtain Streamway, stn PB4­1C (5 dry).
</paragraph>
<paragraph id="8BBD36AC2D1DA213E41FFDE57FB8FDC1" blockId="31.[264,1323,553,659]" box="[264,389,553,578]" pageId="31" pageNumber="32">
<emphasis id="B976EABE2D1DA213E41FFDE57FB8FDC1" box="[264,389,553,578]" italics="true" pageId="31" pageNumber="32">Etymology</emphasis>
</paragraph>
<paragraph id="8BBD36AC2D1DA213E42FFD9D7F59FD10" blockId="31.[264,1323,553,659]" pageId="31" pageNumber="32">
<taxonomicName id="4C024D2F2D1DA213E42FFD9D7FD5FDE8" authority="Latin" authorityName="Latin" box="[312,488,593,619]" class="Insecta" family="Carabidae" genus="Conicus" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Mollusca" rank="genus">
<emphasis id="B976EABE2D1DA213E42FFD9D7FA5FDE9" box="[312,408,593,618]" italics="true" pageId="31" pageNumber="32">Conicus</emphasis>
Latin
</taxonomicName>
adjective, cone­shaped; refers to the conical shape of the shell of this species.
</paragraph>
<paragraph id="8BBD36AC2D1DA213E41FFD737FAFFD5B" blockId="31.[264,1323,703,1169]" box="[264,402,703,728]" pageId="31" pageNumber="32">
<emphasis id="B976EABE2D1DA213E41FFD737FAFFD5B" box="[264,402,703,728]" italics="true" pageId="31" pageNumber="32">Description</emphasis>
</paragraph>
<paragraph id="8BBD36AC2D1DA213E42FFD2A7A63FBEA" blockId="31.[264,1323,703,1169]" pageId="31" pageNumber="32">
<emphasis id="B976EABE2D1DA213E42FFD2A7F4BFC83" bold="true" box="[312,374,742,768]" pageId="31" pageNumber="32">Shell</emphasis>
(
<figureCitation id="13392A292D1DA213E49EFD2B7C30FC82" box="[393,525,743,769]" captionStart="FIGURE 6" captionStartId="16.[264,357,1641,1662]" captionTargetBox="[302,1285,329,1616]" captionTargetId="figure-146@16.[302,1285,329,1633]" captionTargetPageId="16" captionText="FIGURE 6. Protoconchs and teleoconch microsculpture of Pseudotricula and Nanocochlea species. A–D, Pseudotricula expandolabra, A, B, D, C.201493, paratype, main streamway, Bauhaus, C, C.201495, Persephone Streamway, Bauhaus; E–G, Pseudotricula conica, E, F, C.201464, Cane Toad Abuse Streamway, Damper Cave, G, Persephone Pot, Bauhaus; Honey and Cream Streamway, Damper Cave; H, Nanocochlea exigua, C.201475; I, J, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; K–M, Nanocochlea damperensis, K, L, C.203685, paratypes, first crossing, Damper Creek, M, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; N, O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, E, F, H, K–N, protoconch; C, D, G, I, J, O, microsculpture. Scales: A, C–E, J, K, M, O, 200 m; B, F–I, L, N, 100 m." figureDoi="http://doi.org/10.5281/zenodo.5050797" httpUri="https://zenodo.org/record/5050797/files/figure.png" pageId="31" pageNumber="32">Figs 6E–G</figureCitation>
;
<figureCitation id="13392A292D1DA213E70AFD2B7C49FC82" box="[541,628,743,769]" captionStart="FIGURE 12" captionStartId="28.[264,357,1479,1500]" captionTargetBox="[382,1170,289,1464]" captionTargetId="figure-168@28.[354,1233,269,1476]" captionTargetPageId="28" captionText="FIGURE 12. Shells of Pseudotricula species. A, Pseudotricula arthurclarkei, holotype, C.439656, Quetzalcoatl Conduit; B ­ D, Pseudotricula conica, B, holotype, Cane Toad Abuse Streamway, Damper Cave; C, C.203680, main streamway near entrance, Damper Cave; D, C.201813, Persephone Pot, Bauhaus. Scales: A–D, 500 m." figureDoi="http://doi.org/10.5281/zenodo.5050815" httpUri="https://zenodo.org/record/5050815/files/figure.png" pageId="31" pageNumber="32">12B–D</figureCitation>
;
<figureCitation id="13392A292D1DA213E793FD2B7CEAFC82" box="[644,727,743,769]" captionStart="FIGURE 13" captionStartId="30.[264,357,1326,1347]" captionTargetBox="[393,1223,370,1252]" captionTargetId="figure-178@30.[348,1252,362,1266]" captionTargetPageId="30" captionText="FIGURE 13. Shells of Pseudotricula species. A ­ F, Pseudotricula conica, A, C.203694, Black Curtain streamway, Cueva Blanca, B, Screaming Stals streamway, Bauhaus; C, C.201813, Persephone Pot, Bauhaus, D, C.203464, Cane Toad Abuse Streamway, Damper Cave, E, C.201488, main streamway, Damper Cave, F, C.203692, Damper Cave. Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5050817" httpUri="https://zenodo.org/record/5050817/files/figure.png" pageId="31" pageNumber="32">13A–F</figureCitation>
). Length up to
<quantity id="4CFA9B492D1DA213E680FD2B7DC9FC82" box="[919,1012,743,769]" metricMagnitude="-3" metricUnit="m" metricValue="3.2" pageId="31" pageNumber="32" unit="mm" value="3.2">3.2 mm</quantity>
(usually less than
<quantity id="4CFA9B492D1DA213E1CBFD2B7B1BFC82" box="[1244,1318,743,769]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" pageId="31" pageNumber="32" unit="mm" value="3.0">3 mm</quantity>
, mean
<quantity id="4CFA9B492D1DA213E446FCC37F8EFCAA" box="[337,435,783,809]" metricMagnitude="-3" metricUnit="m" metricValue="2.3" pageId="31" pageNumber="32" unit="mm" value="2.3">2.3 mm</quantity>
); conical to broadly conical (SW/SL 0.58–0.75, mean 0.67, n = 60); spire low to moderate, straight to slightly convex in outline; last whorl evenly rounded, or angular to subangular in middle of whorl; whorls very slightly convex; suture simple. Protoconch microsculpture of small, distinct, closely spaced pits (
<figureCitation id="13392A292D1DA213E107FC4B7AB1FC22" box="[1040,1164,903,929]" captionStart="FIGURE 6" captionStartId="16.[264,357,1641,1662]" captionTargetBox="[302,1285,329,1616]" captionTargetId="figure-146@16.[302,1285,329,1633]" captionTargetPageId="16" captionText="FIGURE 6. Protoconchs and teleoconch microsculpture of Pseudotricula and Nanocochlea species. A–D, Pseudotricula expandolabra, A, B, D, C.201493, paratype, main streamway, Bauhaus, C, C.201495, Persephone Streamway, Bauhaus; E–G, Pseudotricula conica, E, F, C.201464, Cane Toad Abuse Streamway, Damper Cave, G, Persephone Pot, Bauhaus; Honey and Cream Streamway, Damper Cave; H, Nanocochlea exigua, C.201475; I, J, Nanocochlea stylesae, C.201476, paratypes, Malani Creek; K–M, Nanocochlea damperensis, K, L, C.203685, paratypes, first crossing, Damper Creek, M, C.201282, Near New River Lagoon, E side, Limestone Ck, where tagged track crosses; N, O, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, E, F, H, K–N, protoconch; C, D, G, I, J, O, microsculpture. Scales: A, C–E, J, K, M, O, 200 m; B, F–I, L, N, 100 m." figureDoi="http://doi.org/10.5281/zenodo.5050797" httpUri="https://zenodo.org/record/5050797/files/figure.png" pageId="31" pageNumber="32">Fig. 6E, F</figureCitation>
). Teleoconch up to 3.3 whorls in adult; aperture oval to pear­shaped; moderately large, shorter to longer than spire (AL/SL 0.40–0.60, mean 0.50, n = 60); outer lip prosocline, straight, with or without slight reflection; external varix absent; posterior notch absent; inner lip thin to moderately thickened and narrow to medium width, firmly adhering to narrowly separated from parietal wall. Shell colour white or yellowish­brown to pale brown.
</paragraph>
<paragraph id="8BBD36AC2D1DA213E42FFBBB7C5EFB12" blockId="31.[264,1323,703,1169]" box="[312,611,1143,1169]" pageId="31" pageNumber="32">
Dimensions. See
<tableCitation id="C68003172D1DA213E713FBBB7C60FB12" box="[516,605,1143,1169]" captionStart="TABLE 8" captionStartId="31.[264,343,1207,1228]" captionText="TABLE 8. Shell dimensions and teleoconch whorl counts of Pseudotricula conica n.sp." pageId="31" pageNumber="32">Table 8</tableCitation>
.
</paragraph>
<caption id="DF7D66242D1DA213E41FFB7B7A6EFB4F" box="[264,1107,1207,1229]" pageId="31" pageNumber="32" startId="31.[264,343,1207,1228]">
<paragraph id="8BBD36AC2D1DA213E41FFB7B7A6EFB4F" blockId="31.[264,1107,1207,1229]" box="[264,1107,1207,1229]" pageId="31" pageNumber="32">
<emphasis id="B976EABE2D1DA213E41FFB7B7F52FB4F" bold="true" box="[264,367,1207,1228]" pageId="31" pageNumber="32">TABLE 8.</emphasis>
Shell dimensions and teleoconch whorl counts of
<taxonomicName id="4C024D2F2D1DA213E644FB747A22FB4E" authority="Ponder &amp; Clark &amp; Eberhard &amp; Studdert, 2005" authorityName="Ponder &amp; Clark &amp; Eberhard &amp; Studdert" authorityYear="2005" box="[851,1055,1208,1229]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="31" pageNumber="32" phylum="Mollusca" rank="species" species="conica" status="sp. nov.">
<emphasis id="B976EABE2D1DA213E644FB747A22FB4E" box="[851,1055,1208,1229]" italics="true" pageId="31" pageNumber="32">Pseudotricula conica</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24557C52D1DA213E132FB7B7A6EFB4F" box="[1061,1107,1207,1228]" pageId="31" pageNumber="32" rank="species">n.sp.</taxonomicNameLabel>
</paragraph>
</caption>
<paragraph id="8BBD36AC2D1DA213E4ECFB267ADBF8EE" pageId="31" pageNumber="32">
<table id="F902C40C2D1D5DF3E407FB267ADAF8EE" box="[272,1255,1258,1901]" colsContinueFrom="27.[272,1267,1372,1616]" colsContinueIn="34.[272,1255,707,1027]" gridcols="8" gridrows="19" pageId="31" pageNumber="32">
<tr id="353234EE2D1D5DF3E407FB267ADAFB7C" box="[272,1255,1258,1279]" gridrow="0" pageId="31" pageNumber="32" rowspan-0="1">
<th id="76E35D922D1D5DF3E4ECFB267C18FB7C" box="[507,549,1258,1279]" gridcol="1" gridrow="0" pageId="31" pageNumber="32">
<emphasis id="B976EABE2D1DA213E4ECFB267C25FB7C" bold="true" box="[507,536,1258,1279]" pageId="31" pageNumber="32">SL</emphasis>
</th>
<th id="76E35D922D1D5DF3E767FB267CA6FB7C" box="[624,667,1258,1279]" gridcol="2" gridrow="0" pageId="31" pageNumber="32">
<emphasis id="B976EABE2D1DA213E766FB267CABFB7C" bold="true" box="[625,662,1258,1279]" pageId="31" pageNumber="32">SW</emphasis>
</th>
<th id="76E35D922D1D5DF3E7F1FB267D2DFB7C" box="[742,784,1258,1279]" gridcol="3" gridrow="0" pageId="31" pageNumber="32">
<emphasis id="B976EABE2D1DA213E7F1FB267D35FB7C" bold="true" box="[742,776,1258,1279]" pageId="31" pageNumber="32">AL</emphasis>
</th>
<th id="76E35D922D1D5DF3E64BFB267DBBFB7C" box="[860,902,1258,1279]" gridcol="4" gridrow="0" pageId="31" pageNumber="32">
<emphasis id="B976EABE2D1DA213E64BFB267DBEFB7C" bold="true" box="[860,899,1258,1279]" pageId="31" pageNumber="32">AW</emphasis>
</th>
<th id="76E35D922D1D5DF3E6C6FB267DC6FB7C" box="[977,1019,1258,1279]" gridcol="5" gridrow="0" pageId="31" pageNumber="32">
<emphasis id="B976EABE2D1DA213E6C6FB267DC4FB7C" bold="true" box="[977,1017,1258,1279]" pageId="31" pageNumber="32">BW</emphasis>
</th>
<th id="76E35D922D1D5DF3E150FB267A4CFB7C" box="[1095,1137,1258,1279]" gridcol="6" gridrow="0" pageId="31" pageNumber="32">
<emphasis id="B976EABE2D1DA213E150FB267A57FB7C" bold="true" box="[1095,1130,1258,1279]" pageId="31" pageNumber="32">CV</emphasis>
</th>
<th id="76E35D922D1D5DF3E1ABFB267ADAFB7C" box="[1212,1255,1258,1279]" gridcol="7" gridrow="0" pageId="31" pageNumber="32">
<emphasis id="B976EABE2D1DA213E1ABFB267AD9FB7C" bold="true" box="[1212,1252,1258,1279]" pageId="31" pageNumber="32">TW</emphasis>
</th>
</tr>
<tr id="353234EE2D1D5DF3E407FAC17ADAFAA1" box="[272,1255,1293,1314]" gridrow="1" pageId="31" pageNumber="32">
<th id="76E35D922D1D5DF3E407FAC17F83FAA1" box="[272,446,1293,1314]" gridcol="0" gridrow="1" pageId="31" pageNumber="32">Holotype</th>
<td id="76E35D922D1D5DF3E4ECFAC17C18FAA1" box="[507,549,1293,1314]" gridcol="1" gridrow="1" pageId="31" pageNumber="32">2.26</td>
<td id="76E35D922D1D5DF3E767FAC17CA6FAA1" box="[624,667,1293,1314]" gridcol="2" gridrow="1" pageId="31" pageNumber="32">1.43</td>
<td id="76E35D922D1D5DF3E7F1FAC17D2DFAA1" box="[742,784,1293,1314]" gridcol="3" gridrow="1" pageId="31" pageNumber="32">1.05</td>
<td id="76E35D922D1D5DF3E64BFAC17DBBFAA1" box="[860,902,1293,1314]" gridcol="4" gridrow="1" pageId="31" pageNumber="32">1.06</td>
<td id="76E35D922D1D5DF3E6C6FAC17DC6FAA1" box="[977,1019,1293,1314]" gridcol="5" gridrow="1" pageId="31" pageNumber="32">1.76</td>
<td id="76E35D922D1D5DF3E150FAC17A4CFAA1" box="[1095,1137,1293,1314]" gridcol="6" gridrow="1" pageId="31" pageNumber="32">0.11</td>
<td id="76E35D922D1D5DF3E1ABFAC17ADAFAA1" box="[1212,1255,1293,1314]" gridcol="7" gridrow="1" pageId="31" pageNumber="32">2.75</td>
</tr>
<tr id="353234EE2D1D5DF3E407FAE37ADAFAC7" box="[272,1255,1327,1348]" gridrow="2" pageId="31" pageNumber="32">
<th id="76E35D922D1D5DF3E407FAE37F83FAC7" box="[272,446,1327,1348]" gridcol="0" gridrow="2" pageId="31" pageNumber="32">Figured Specimen</th>
<td id="76E35D922D1D5DF3E4ECFAE37C18FAC7" box="[507,549,1327,1348]" gridcol="1" gridrow="2" pageId="31" pageNumber="32">2.07</td>
<td id="76E35D922D1D5DF3E767FAE37CA6FAC7" box="[624,667,1327,1348]" gridcol="2" gridrow="2" pageId="31" pageNumber="32">1.28</td>
<td id="76E35D922D1D5DF3E7F1FAE37D2DFAC7" box="[742,784,1327,1348]" gridcol="3" gridrow="2" pageId="31" pageNumber="32">1.01</td>
<td id="76E35D922D1D5DF3E64BFAE37DBBFAC7" box="[860,902,1327,1348]" gridcol="4" gridrow="2" pageId="31" pageNumber="32">0.89</td>
<td id="76E35D922D1D5DF3E6C6FAE37DC6FAC7" box="[977,1019,1327,1348]" gridcol="5" gridrow="2" pageId="31" pageNumber="32">1.58</td>
<td id="76E35D922D1D5DF3E150FAE37A4CFAC7" box="[1095,1137,1327,1348]" gridcol="6" gridrow="2" pageId="31" pageNumber="32">0.14</td>
<td id="76E35D922D1D5DF3E1ABFAE37ADAFAC7" box="[1212,1255,1327,1348]" gridcol="7" gridrow="2" pageId="31" pageNumber="32">2.60</td>
</tr>
<tr id="353234EE2D1D5DF3E407FA9E7ADAFAE4" box="[272,1255,1362,1383]" gridrow="3" pageId="31" pageNumber="32">
<th id="76E35D922D1D5DF3E407FA9E7F83FAE4" box="[272,446,1362,1383]" gridcol="0" gridrow="3" pageId="31" pageNumber="32">Figured specimen</th>
<td id="76E35D922D1D5DF3E4ECFA9E7C18FAE4" box="[507,549,1362,1383]" gridcol="1" gridrow="3" pageId="31" pageNumber="32">2.3</td>
<td id="76E35D922D1D5DF3E767FA9E7CA6FAE4" box="[624,667,1362,1383]" gridcol="2" gridrow="3" pageId="31" pageNumber="32">1.51</td>
<td id="76E35D922D1D5DF3E7F1FA9E7D2DFAE4" box="[742,784,1362,1383]" gridcol="3" gridrow="3" pageId="31" pageNumber="32">1.23</td>
<td id="76E35D922D1D5DF3E64BFA9E7DBBFAE4" box="[860,902,1362,1383]" gridcol="4" gridrow="3" pageId="31" pageNumber="32">1.06</td>
<td id="76E35D922D1D5DF3E6C6FA9E7DC6FAE4" box="[977,1019,1362,1383]" gridcol="5" gridrow="3" pageId="31" pageNumber="32">1.85</td>
<td id="76E35D922D1D5DF3E150FA9E7A4CFAE4" box="[1095,1137,1362,1383]" gridcol="6" gridrow="3" pageId="31" pageNumber="32">0.1</td>
<td id="76E35D922D1D5DF3E1ABFA9E7ADAFAE4" box="[1212,1255,1362,1383]" gridcol="7" gridrow="3" pageId="31" pageNumber="32">2.75</td>
</tr>
<tr id="353234EE2D1D5DF3E407FAB97ADAFA09" box="[272,1255,1397,1418]" gridrow="4" pageId="31" pageNumber="32" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76E35D922D1D5DF3E407FAB97F83FA09" box="[272,446,1397,1418]" gridcol="0" gridrow="4" pageId="31" pageNumber="32">C.201459(20)</th>
</tr>
<tr id="353234EE2D1D5DF3E407FA5B7ADAFA2F" box="[272,1255,1431,1452]" gridrow="5" pageId="31" pageNumber="32">
<th id="76E35D922D1D5DF3E407FA5B7F83FA2F" box="[272,446,1431,1452]" gridcol="0" gridrow="5" pageId="31" pageNumber="32">Minimum</th>
<td id="76E35D922D1D5DF3E4ECFA5B7C18FA2F" box="[507,549,1431,1452]" gridcol="1" gridrow="5" pageId="31" pageNumber="32">1.83</td>
<td id="76E35D922D1D5DF3E767FA5B7CA6FA2F" box="[624,667,1431,1452]" gridcol="2" gridrow="5" pageId="31" pageNumber="32">1.21</td>
<td id="76E35D922D1D5DF3E7F1FA5B7D2DFA2F" box="[742,784,1431,1452]" gridcol="3" gridrow="5" pageId="31" pageNumber="32">0.92</td>
<td id="76E35D922D1D5DF3E64BFA5B7DBBFA2F" box="[860,902,1431,1452]" gridcol="4" gridrow="5" pageId="31" pageNumber="32">0.87</td>
<td id="76E35D922D1D5DF3E6C6FA5B7DC6FA2F" box="[977,1019,1431,1452]" gridcol="5" gridrow="5" pageId="31" pageNumber="32">1.42</td>
<td id="76E35D922D1D5DF3E150FA5B7A4CFA2F" box="[1095,1137,1431,1452]" gridcol="6" gridrow="5" pageId="31" pageNumber="32">0.09</td>
<td id="76E35D922D1D5DF3E1ABFA5B7ADAFA2F" box="[1212,1255,1431,1452]" gridcol="7" gridrow="5" pageId="31" pageNumber="32">2.50</td>
</tr>
<tr id="353234EE2D1D5DF3E407FA767ADAFA4C" box="[272,1255,1466,1487]" gridrow="6" pageId="31" pageNumber="32">
<th id="76E35D922D1D5DF3E407FA767F83FA4C" box="[272,446,1466,1487]" gridcol="0" gridrow="6" pageId="31" pageNumber="32">Maximum</th>
<td id="76E35D922D1D5DF3E4ECFA767C18FA4C" box="[507,549,1466,1487]" gridcol="1" gridrow="6" pageId="31" pageNumber="32">2.52</td>
<td id="76E35D922D1D5DF3E767FA767CA6FA4C" box="[624,667,1466,1487]" gridcol="2" gridrow="6" pageId="31" pageNumber="32">1.63</td>
<td id="76E35D922D1D5DF3E7F1FA767D2DFA4C" box="[742,784,1466,1487]" gridcol="3" gridrow="6" pageId="31" pageNumber="32">1.20</td>
<td id="76E35D922D1D5DF3E64BFA767DBBFA4C" box="[860,902,1466,1487]" gridcol="4" gridrow="6" pageId="31" pageNumber="32">1.12</td>
<td id="76E35D922D1D5DF3E6C6FA767DC6FA4C" box="[977,1019,1466,1487]" gridcol="5" gridrow="6" pageId="31" pageNumber="32">1.94</td>
<td id="76E35D922D1D5DF3E150FA767A4CFA4C" box="[1095,1137,1466,1487]" gridcol="6" gridrow="6" pageId="31" pageNumber="32">0.18</td>
<td id="76E35D922D1D5DF3E1ABFA767ADAFA4C" box="[1212,1255,1466,1487]" gridcol="7" gridrow="6" pageId="31" pageNumber="32">3.00</td>
</tr>
<tr id="353234EE2D1D5DF3E407FA107ADAFA72" box="[272,1255,1500,1521]" gridrow="7" pageId="31" pageNumber="32">
<th id="76E35D922D1D5DF3E407FA107F83FA72" box="[272,446,1500,1521]" gridcol="0" gridrow="7" pageId="31" pageNumber="32">Mean</th>
<td id="76E35D922D1D5DF3E4ECFA107C18FA72" box="[507,549,1500,1521]" gridcol="1" gridrow="7" pageId="31" pageNumber="32">2.22</td>
<td id="76E35D922D1D5DF3E767FA107CA6FA72" box="[624,667,1500,1521]" gridcol="2" gridrow="7" pageId="31" pageNumber="32">1.46</td>
<td id="76E35D922D1D5DF3E7F1FA107D2DFA72" box="[742,784,1500,1521]" gridcol="3" gridrow="7" pageId="31" pageNumber="32">1.07</td>
<td id="76E35D922D1D5DF3E64BFA107DBBFA72" box="[860,902,1500,1521]" gridcol="4" gridrow="7" pageId="31" pageNumber="32">1.01</td>
<td id="76E35D922D1D5DF3E6C6FA107DC6FA72" box="[977,1019,1500,1521]" gridcol="5" gridrow="7" pageId="31" pageNumber="32">1.71</td>
<td id="76E35D922D1D5DF3E150FA107A4CFA72" box="[1095,1137,1500,1521]" gridcol="6" gridrow="7" pageId="31" pageNumber="32">0.13</td>
<td id="76E35D922D1D5DF3E1ABFA107ADAFA72" box="[1212,1255,1500,1521]" gridcol="7" gridrow="7" pageId="31" pageNumber="32">2.76</td>
</tr>
<tr id="353234EE2D1D5DF3E407FA337ADAF997" box="[272,1255,1535,1556]" gridrow="8" pageId="31" pageNumber="32">
<th id="76E35D922D1D5DF3E407FA337F83F997" box="[272,446,1535,1556]" gridcol="0" gridrow="8" pageId="31" pageNumber="32">Standard Dev.</th>
<td id="76E35D922D1D5DF3E4ECFA337C18F997" box="[507,549,1535,1556]" gridcol="1" gridrow="8" pageId="31" pageNumber="32">0.16</td>
<td id="76E35D922D1D5DF3E767FA337CA6F997" box="[624,667,1535,1556]" gridcol="2" gridrow="8" pageId="31" pageNumber="32">0.11</td>
<td id="76E35D922D1D5DF3E7F1FA337D2DF997" box="[742,784,1535,1556]" gridcol="3" gridrow="8" pageId="31" pageNumber="32">0.09</td>
<td id="76E35D922D1D5DF3E64BFA337DBBF997" box="[860,902,1535,1556]" gridcol="4" gridrow="8" pageId="31" pageNumber="32">0.07</td>
<td id="76E35D922D1D5DF3E6C6FA337DC6F997" box="[977,1019,1535,1556]" gridcol="5" gridrow="8" pageId="31" pageNumber="32">0.12</td>
<td id="76E35D922D1D5DF3E150FA337A4CF997" box="[1095,1137,1535,1556]" gridcol="6" gridrow="8" pageId="31" pageNumber="32">0.02</td>
<td id="76E35D922D1D5DF3E1ABFA337ADAF997" box="[1212,1255,1535,1556]" gridcol="7" gridrow="8" pageId="31" pageNumber="32">0.15</td>
</tr>
<tr id="353234EE2D1D5DF3E407F9ED7ADAF9B5" box="[272,1255,1569,1590]" gridrow="9" pageId="31" pageNumber="32">
<th id="76E35D922D1D5DF3E407F9ED7ADAF9B5" box="[272,1255,1569,1590]" colspan="8" colspanRight="7" gridcol="0" gridrow="9" pageId="31" pageNumber="32">C.201813(17)</th>
</tr>
<tr id="353234EE2D1D5DF3E407F9887ADAF9DA" box="[272,1255,1604,1625]" gridrow="10" pageId="31" pageNumber="32">
<th id="76E35D922D1D5DF3E407F9887F83F9DA" box="[272,446,1604,1625]" gridcol="0" gridrow="10" pageId="31" pageNumber="32">Minimum</th>
<td id="76E35D922D1D5DF3E4ECF9887C18F9DA" box="[507,549,1604,1625]" gridcol="1" gridrow="10" pageId="31" pageNumber="32">1.65</td>
<td id="76E35D922D1D5DF3E767F9887CA6F9DA" box="[624,667,1604,1625]" gridcol="2" gridrow="10" pageId="31" pageNumber="32">1.01</td>
<td id="76E35D922D1D5DF3E7F1F9887D2DF9DA" box="[742,784,1604,1625]" gridcol="3" gridrow="10" pageId="31" pageNumber="32">0.67</td>
<td id="76E35D922D1D5DF3E64BF9887DBBF9DA" box="[860,902,1604,1625]" gridcol="4" gridrow="10" pageId="31" pageNumber="32">0.57</td>
<td id="76E35D922D1D5DF3E6C6F9887DC6F9DA" box="[977,1019,1604,1625]" gridcol="5" gridrow="10" pageId="31" pageNumber="32">1.18</td>
<td id="76E35D922D1D5DF3E150F9887A4CF9DA" box="[1095,1137,1604,1625]" gridcol="6" gridrow="10" pageId="31" pageNumber="32">0.01</td>
<td id="76E35D922D1D5DF3E1ABF9887ADAF9DA" box="[1212,1255,1604,1625]" gridcol="7" gridrow="10" pageId="31" pageNumber="32">2.12</td>
</tr>
<tr id="353234EE2D1D5DF3E407F9AA7ADAF9F8" box="[272,1255,1638,1659]" gridrow="11" pageId="31" pageNumber="32">
<th id="76E35D922D1D5DF3E407F9AA7F83F9F8" box="[272,446,1638,1659]" gridcol="0" gridrow="11" pageId="31" pageNumber="32">Maximum</th>
<td id="76E35D922D1D5DF3E4ECF9AA7C18F9F8" box="[507,549,1638,1659]" gridcol="1" gridrow="11" pageId="31" pageNumber="32">2.44</td>
<td id="76E35D922D1D5DF3E767F9AA7CA6F9F8" box="[624,667,1638,1659]" gridcol="2" gridrow="11" pageId="31" pageNumber="32">1.47</td>
<td id="76E35D922D1D5DF3E7F1F9AA7D2DF9F8" box="[742,784,1638,1659]" gridcol="3" gridrow="11" pageId="31" pageNumber="32">1.12</td>
<td id="76E35D922D1D5DF3E64BF9AA7DBBF9F8" box="[860,902,1638,1659]" gridcol="4" gridrow="11" pageId="31" pageNumber="32">0.99</td>
<td id="76E35D922D1D5DF3E6C6F9AA7DC6F9F8" box="[977,1019,1638,1659]" gridcol="5" gridrow="11" pageId="31" pageNumber="32">1.75</td>
<td id="76E35D922D1D5DF3E150F9AA7A4CF9F8" box="[1095,1137,1638,1659]" gridcol="6" gridrow="11" pageId="31" pageNumber="32">0.22</td>
<td id="76E35D922D1D5DF3E1ABF9AA7ADAF9F8" box="[1212,1255,1638,1659]" gridcol="7" gridrow="11" pageId="31" pageNumber="32">3.30</td>
</tr>
<tr id="353234EE2D1D5DF3E407F9457ADAF91D" box="[272,1255,1673,1694]" gridrow="12" pageId="31" pageNumber="32">
<th id="76E35D922D1D5DF3E407F9457F83F91D" box="[272,446,1673,1694]" gridcol="0" gridrow="12" pageId="31" pageNumber="32">Mean</th>
<td id="76E35D922D1D5DF3E4ECF9457C18F91D" box="[507,549,1673,1694]" gridcol="1" gridrow="12" pageId="31" pageNumber="32">2.02</td>
<td id="76E35D922D1D5DF3E767F9457CA6F91D" box="[624,667,1673,1694]" gridcol="2" gridrow="12" pageId="31" pageNumber="32">1.29</td>
<td id="76E35D922D1D5DF3E7F1F9457D2DF91D" box="[742,784,1673,1694]" gridcol="3" gridrow="12" pageId="31" pageNumber="32">0.94</td>
<td id="76E35D922D1D5DF3E64BF9457DBBF91D" box="[860,902,1673,1694]" gridcol="4" gridrow="12" pageId="31" pageNumber="32">0.82</td>
<td id="76E35D922D1D5DF3E6C6F9457DC6F91D" box="[977,1019,1673,1694]" gridcol="5" gridrow="12" pageId="31" pageNumber="32">1.52</td>
<td id="76E35D922D1D5DF3E150F9457A4CF91D" box="[1095,1137,1673,1694]" gridcol="6" gridrow="12" pageId="31" pageNumber="32">0.11</td>
<td id="76E35D922D1D5DF3E1ABF9457ADAF91D" box="[1212,1255,1673,1694]" gridcol="7" gridrow="12" pageId="31" pageNumber="32">2.75</td>
</tr>
<tr id="353234EE2D1D5DF3E407F9677ADAF943" box="[272,1255,1707,1728]" gridrow="13" pageId="31" pageNumber="32">
<th id="76E35D922D1D5DF3E407F9677F83F943" box="[272,446,1707,1728]" gridcol="0" gridrow="13" pageId="31" pageNumber="32">Standard Dev.</th>
<td id="76E35D922D1D5DF3E4ECF9677C18F943" box="[507,549,1707,1728]" gridcol="1" gridrow="13" pageId="31" pageNumber="32">0.25</td>
<td id="76E35D922D1D5DF3E767F9677CA6F943" box="[624,667,1707,1728]" gridcol="2" gridrow="13" pageId="31" pageNumber="32">0.15</td>
<td id="76E35D922D1D5DF3E7F1F9677D2DF943" box="[742,784,1707,1728]" gridcol="3" gridrow="13" pageId="31" pageNumber="32">0.12</td>
<td id="76E35D922D1D5DF3E64BF9677DBBF943" box="[860,902,1707,1728]" gridcol="4" gridrow="13" pageId="31" pageNumber="32">0.14</td>
<td id="76E35D922D1D5DF3E6C6F9677DC6F943" box="[977,1019,1707,1728]" gridcol="5" gridrow="13" pageId="31" pageNumber="32">0.17</td>
<td id="76E35D922D1D5DF3E150F9677A4CF943" box="[1095,1137,1707,1728]" gridcol="6" gridrow="13" pageId="31" pageNumber="32">0.05</td>
<td id="76E35D922D1D5DF3E1ABF9677ADAF943" box="[1212,1255,1707,1728]" gridcol="7" gridrow="13" pageId="31" pageNumber="32">0.31</td>
</tr>
<tr id="353234EE2D1D5DF3E407F9027ADAF960" box="[272,1255,1742,1763]" gridrow="14" pageId="31" pageNumber="32" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1" rowspan-6="1" rowspan-7="1">
<th id="76E35D922D1D5DF3E407F9027F83F960" box="[272,446,1742,1763]" gridcol="0" gridrow="14" pageId="31" pageNumber="32">C.203680(20)</th>
</tr>
<tr id="353234EE2D1D5DF3E407F93C7ADAF886" box="[272,1255,1776,1797]" gridrow="15" pageId="31" pageNumber="32">
<th id="76E35D922D1D5DF3E407F93C7F83F886" box="[272,446,1776,1797]" gridcol="0" gridrow="15" pageId="31" pageNumber="32">Minimum</th>
<td id="76E35D922D1D5DF3E4ECF93C7C18F886" box="[507,549,1776,1797]" gridcol="1" gridrow="15" pageId="31" pageNumber="32">2.38</td>
<td id="76E35D922D1D5DF3E767F93C7CA6F886" box="[624,667,1776,1797]" gridcol="2" gridrow="15" pageId="31" pageNumber="32">1.66</td>
<td id="76E35D922D1D5DF3E7F1F93C7D2DF886" box="[742,784,1776,1797]" gridcol="3" gridrow="15" pageId="31" pageNumber="32">1.27</td>
<td id="76E35D922D1D5DF3E64BF93C7DBBF886" box="[860,902,1776,1797]" gridcol="4" gridrow="15" pageId="31" pageNumber="32">1.11</td>
<td id="76E35D922D1D5DF3E6C6F93C7DC6F886" box="[977,1019,1776,1797]" gridcol="5" gridrow="15" pageId="31" pageNumber="32">1.98</td>
<td id="76E35D922D1D5DF3E150F93C7A4CF886" box="[1095,1137,1776,1797]" gridcol="6" gridrow="15" pageId="31" pageNumber="32">0.08</td>
<td id="76E35D922D1D5DF3E1ABF93C7ADAF886" box="[1212,1255,1776,1797]" gridcol="7" gridrow="15" pageId="31" pageNumber="32">2.50</td>
</tr>
<tr id="353234EE2D1D5DF3E407F8DF7ADAF8AB" box="[272,1255,1811,1832]" gridrow="16" pageId="31" pageNumber="32">
<th id="76E35D922D1D5DF3E407F8DF7F83F8AB" box="[272,446,1811,1832]" gridcol="0" gridrow="16" pageId="31" pageNumber="32">Maximum</th>
<td id="76E35D922D1D5DF3E4ECF8DF7C18F8AB" box="[507,549,1811,1832]" gridcol="1" gridrow="16" pageId="31" pageNumber="32">3.23</td>
<td id="76E35D922D1D5DF3E767F8DF7CA6F8AB" box="[624,667,1811,1832]" gridcol="2" gridrow="16" pageId="31" pageNumber="32">2.07</td>
<td id="76E35D922D1D5DF3E7F1F8DF7D2DF8AB" box="[742,784,1811,1832]" gridcol="3" gridrow="16" pageId="31" pageNumber="32">1.57</td>
<td id="76E35D922D1D5DF3E64BF8DF7DBBF8AB" box="[860,902,1811,1832]" gridcol="4" gridrow="16" pageId="31" pageNumber="32">1.39</td>
<td id="76E35D922D1D5DF3E6C6F8DF7DC6F8AB" box="[977,1019,1811,1832]" gridcol="5" gridrow="16" pageId="31" pageNumber="32">2.45</td>
<td id="76E35D922D1D5DF3E150F8DF7A4CF8AB" box="[1095,1137,1811,1832]" gridcol="6" gridrow="16" pageId="31" pageNumber="32">0.16</td>
<td id="76E35D922D1D5DF3E1ABF8DF7ADAF8AB" box="[1212,1255,1811,1832]" gridcol="7" gridrow="16" pageId="31" pageNumber="32">3.10</td>
</tr>
<tr id="353234EE2D1D5DF3E407F8FA7ADAF8C8" box="[272,1255,1846,1867]" gridrow="17" pageId="31" pageNumber="32">
<th id="76E35D922D1D5DF3E407F8FA7F83F8C8" box="[272,446,1846,1867]" gridcol="0" gridrow="17" pageId="31" pageNumber="32">Mean</th>
<td id="76E35D922D1D5DF3E4ECF8FA7C18F8C8" box="[507,549,1846,1867]" gridcol="1" gridrow="17" pageId="31" pageNumber="32">2.61</td>
<td id="76E35D922D1D5DF3E767F8FA7CA6F8C8" box="[624,667,1846,1867]" gridcol="2" gridrow="17" pageId="31" pageNumber="32">1.84</td>
<td id="76E35D922D1D5DF3E7F1F8FA7D2DF8C8" box="[742,784,1846,1867]" gridcol="3" gridrow="17" pageId="31" pageNumber="32">1.41</td>
<td id="76E35D922D1D5DF3E64BF8FA7DBBF8C8" box="[860,902,1846,1867]" gridcol="4" gridrow="17" pageId="31" pageNumber="32">1.26</td>
<td id="76E35D922D1D5DF3E6C6F8FA7DC6F8C8" box="[977,1019,1846,1867]" gridcol="5" gridrow="17" pageId="31" pageNumber="32">2.15</td>
<td id="76E35D922D1D5DF3E150F8FA7A4CF8C8" box="[1095,1137,1846,1867]" gridcol="6" gridrow="17" pageId="31" pageNumber="32">0.12</td>
<td id="76E35D922D1D5DF3E1ABF8FA7ADAF8C8" box="[1212,1255,1846,1867]" gridcol="7" gridrow="17" pageId="31" pageNumber="32">2.66</td>
</tr>
<tr id="353234EE2D1D5DF3E407F8947ADAF8EE" box="[272,1255,1880,1901]" gridrow="18" pageId="31" pageNumber="32">
<th id="76E35D922D1D5DF3E407F8947F83F8EE" box="[272,446,1880,1901]" gridcol="0" gridrow="18" pageId="31" pageNumber="32">Standard Dev.</th>
<td id="76E35D922D1D5DF3E4ECF8947C18F8EE" box="[507,549,1880,1901]" gridcol="1" gridrow="18" pageId="31" pageNumber="32">0.20</td>
<td id="76E35D922D1D5DF3E767F8947CA6F8EE" box="[624,667,1880,1901]" gridcol="2" gridrow="18" pageId="31" pageNumber="32">0.12</td>
<td id="76E35D922D1D5DF3E7F1F8947D2DF8EE" box="[742,784,1880,1901]" gridcol="3" gridrow="18" pageId="31" pageNumber="32">0.09</td>
<td id="76E35D922D1D5DF3E64BF8947DBBF8EE" box="[860,902,1880,1901]" gridcol="4" gridrow="18" pageId="31" pageNumber="32">0.07</td>
<td id="76E35D922D1D5DF3E6C6F8947DC6F8EE" box="[977,1019,1880,1901]" gridcol="5" gridrow="18" pageId="31" pageNumber="32">0.13</td>
<td id="76E35D922D1D5DF3E150F8947A4CF8EE" box="[1095,1137,1880,1901]" gridcol="6" gridrow="18" pageId="31" pageNumber="32">0.02</td>
<td id="76E35D922D1D5DF3E1ABF8947ADAF8EE" box="[1212,1255,1880,1901]" gridcol="7" gridrow="18" pageId="31" pageNumber="32">0.16</td>
</tr>
</table>
</paragraph>
<paragraph id="8BBD36AC2D22A22CE42FFED77A6DFEB5" blockId="32.[264,1323,283,1550]" box="[312,1104,283,310]" pageId="32" pageNumber="33">
<emphasis id="B976EABE2D22A22CE42FFED77FFAFEB6" bold="true" box="[312,455,283,309]" pageId="32" pageNumber="33">Operculum</emphasis>
(
<figureCitation id="13392A292D22A22CE4C1FED07C78FEB5" box="[470,581,284,310]" captionStart="FIGURE 7" captionStartId="19.[264,357,1755,1776]" captionTargetBox="[369,1242,439,1733]" captionTargetId="figure-114@19.[317,1269,433,1748]" captionTargetPageId="19" captionText="FIGURE 7. Opercula of Pseudotricula species. A, B, Pseudotricula expandolabra, C.201493, paratypes, main streamway, Bauhaus; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula arthurclarkei, C.203671, paratypes, Quetzalcoatl Conduit; F–I, Pseudotricula conica, F, G, C.203680, main streamway, near entrance, Damper Cave; H, I, C.201459, Screaming Stals streamway, Bauhaus; A, D, F, H, outer side; B, C, E, H, I, inner side; Scales: A–I, 200 m." figureDoi="http://doi.org/10.5281/zenodo.5050801" httpUri="https://zenodo.org/record/5050801/files/figure.png" pageId="32" pageNumber="33">Fig. 7F–I</figureCitation>
). Yellowish; inner surface with white smear.
</paragraph>
<paragraph id="8BBD36AC2D22A22CE42FFE8F7FACFE55" blockId="32.[264,1323,283,1550]" pageId="32" pageNumber="33">
<emphasis id="B976EABE2D22A22CE42FFE8F7FDAFEDE" bold="true" box="[312,487,323,349]" pageId="32" pageNumber="33">Pallial cavity</emphasis>
(
<figureCitation id="13392A292D22A22CE714FE887CA1FEDD" box="[515,668,324,350]" captionStart="FIGURE 8" captionStartId="20.[264,357,1612,1633]" captionTargetBox="[341,1270,339,1585]" captionTargetId="figure-146@20.[299,1287,324,1608]" captionTargetPageId="20" captionText="FIGURE 8. Pallial cavities of male specimens of Pseudotricula and Nanocochlea species. A, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; C, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; D, E, Pseudotricula conica, D, C.203680, main streamway near entrance, Damper Cave, E, Cane Toad Abuse Streamway, Damper Cave; F, Nanocochlea exigua, C.203677, paratype, Cane Toad Abuse Streamway, Damper Cave; G, Nanocochlea damperensis, C.203685, paratype, first crossing, Damper Creek; H, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. Scales: A–D, F–H, 500 m, E, 1 mm. Abbreviations: a, anus; ct, ctenidium; ev, efferent vein; os, osphradium; pc, pericardium; pe, pallial edge; ppw, posterior pallial wall; r, rectum; rg, renal gland. The stippled area represents the hypobranchial gland." figureDoi="http://doi.org/10.5281/zenodo.5050803" httpUri="https://zenodo.org/record/5050803/files/figure.png" pageId="32" pageNumber="33">Fig. 8D, E</figureCitation>
). Ctenidium narrow; 10–14 very small filaments; osphradium between posterior end and middle of ctenidium; hypobranchial gland thick; renal organ extends forward ca. ⅓–½ into pallial cavity; pericardium more than ½ in pallial roof.
</paragraph>
<paragraph id="8BBD36AC2D22A22CE42FFE2F7C4DFD6D" blockId="32.[264,1323,283,1550]" pageId="32" pageNumber="33">
<emphasis id="B976EABE2D22A22CE42FFE2F7FAEFE7E" bold="true" box="[312,403,483,509]" pageId="32" pageNumber="33">Radula</emphasis>
(
<figureCitation id="13392A292D22A22CE4B2FE287C26FE7D" box="[421,539,484,510]" captionStart="FIGURE 3" captionStartId="13.[264,357,1408,1429]" captionTargetBox="[319,1271,284,1386]" captionTargetId="figure-152@13.[311,1275,280,1392]" captionTargetPageId="13" captionText="FIGURE 3: Radulae of Pseudotricula species. A, B, Pseudotricula eberhardi, C.165053, Black Curtain Streamway, Cueva Blanca; C–H, Pseudotricula expandolabra, C, D, C.201495, Persephone Streamway, Bauhaus, E, G, H, C.201811, Persephone Pot, Bauhaus, F, C.201822, main streamway, Bauhaus; I, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; J–L, Pseudotricula conica, C.203676, paratype, Cane Toad Abuse Streamway, Damper Cave. A, C, G, K, central, lateral inner and outer marginal teeth; B, E, L, central teeth; D, outer marginal teeth; F, J, lateral and inner marginal teeth; H, lateral tooth; I, central and lateral teeth. Scales: A, B, E, H–J, L 10 m; C, F, G, K, 20 m; D, 5 m." figureDoi="http://doi.org/10.5281/zenodo.5050787" httpUri="https://zenodo.org/record/5050787/files/figure.png" pageId="32" pageNumber="33">Fig. 3J–L</figureCitation>
).
<emphasis id="B976EABE2D22A22CE725FE287CF3FE7E" box="[562,718,484,509]" italics="true" pageId="32" pageNumber="33">Central teeth</emphasis>
: dorsal edge with moderate to deep indentation; 5 lateral cusps, median cusp of narrow to medium width, bluntly to sharply pointed, less than twice as long as adjacent cusps.
<emphasis id="B976EABE2D22A22CE7ECFDF87DA7FDCE" box="[763,922,564,589]" italics="true" pageId="32" pageNumber="33">Lateral teeth</emphasis>
: dorsal edge lightly convex to moderately indented; with 4–7 cusps on outer and 4–6 on inner side; median cusp of medium to narrow, sharp to blunt, less than twice as long as adjacent cusps; ratio of cutting edge to shaft about ⅓; basal projection bluntly pointed.
<emphasis id="B976EABE2D22A22CE6B7FD607A72FD46" box="[928,1103,684,709]" italics="true" pageId="32" pageNumber="33">Marginal teeth</emphasis>
: Inner with 21–32 cusps; outer with 22–32 cusps.
</paragraph>
<paragraph id="8BBD36AC2D22A22CE42FFD307F9FFCBD" blockId="32.[264,1323,283,1550]" pageId="32" pageNumber="33">
<emphasis id="B976EABE2D22A22CE42FFD307F9EFC95" bold="true" box="[312,419,764,790]" pageId="32" pageNumber="33">Stomach</emphasis>
(
<figureCitation id="13392A292D22A22CE4AEFD307C23FC95" box="[441,542,764,790]" captionStart="FIGURE 9" captionStartId="21.[264,357,1723,1744]" captionTargetBox="[350,1309,380,1631]" captionTargetId="figure-97@21.[299,1342,370,1653]" captionTargetPageId="21" captionText="FIGURE 9. Head, penis and stomach of Pseudotricula species. A, D, Pseudotricula conica, C.203680, main streamway near entrance, Damper Cave; C, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, E, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, head and penis; C–E, stomach. Scales: A–E, 500 m. Abbreviations: dgo, opening from stomach to digestive gland; ey, eye; int, intestine; oes, oesophagus, p, penis; pg, penial duck; sn, snout; ss, style sack; st, stomach; tn, tentacle." figureDoi="http://doi.org/10.5281/zenodo.5050807" httpUri="https://zenodo.org/record/5050807/files/figure.png" pageId="32" pageNumber="33">Fig. 9D</figureCitation>
). Stomach with posterior chamber and anterior chamber about equal in size.
</paragraph>
<paragraph id="8BBD36AC2D22A22CE42FFC807AF9FB85" blockId="32.[264,1323,283,1550]" pageId="32" pageNumber="33">
<emphasis id="B976EABE2D22A22CE42FFC807C05FCE5" bold="true" box="[312,568,844,870]" pageId="32" pageNumber="33">Male genital system.</emphasis>
Testis of 1.25 whorls; prostate gland oval to elongate pyriform; compressed in section. Pallial vas deferens straight. Penis (
<figureCitation id="13392A292D22A22CE6E0FCB87A64FC0D" box="[1015,1113,884,910]" captionStart="FIGURE 4" captionStartId="14.[264,357,1822,1843]" captionTargetBox="[406,1181,963,1757]" captionTargetId="figure-174@14.[378,1210,945,1795]" captionTargetPageId="14" captionText="FIGURE 4. Dorsal views of penes of Pseudotricula and Nanocochlea species. A,. Pseudotricula eberhardi; B, Pseudotricula expandolabra; C, Nanocochlea stylesae; D, Nanocochlea damperensis; E, Pseudotricula conica. Scales A, B, E, 0.2 mm; C, D, 0.1 mm. Abbreviations: ds, distal swelling; pd, penial duct." figureDoi="http://doi.org/10.5281/zenodo.5050791" httpUri="https://zenodo.org/record/5050791/files/figure.png" pageId="32" pageNumber="33">Fig. 4E</figureCitation>
;
<figureCitation id="13392A292D22A22CE17AFCB87AAFFC0D" box="[1133,1170,884,910]" captionStart="FIGURE 9" captionStartId="21.[264,357,1723,1744]" captionTargetBox="[350,1309,380,1631]" captionTargetId="figure-97@21.[299,1342,370,1653]" captionTargetPageId="21" captionText="FIGURE 9. Head, penis and stomach of Pseudotricula species. A, D, Pseudotricula conica, C.203680, main streamway near entrance, Damper Cave; C, Pseudotricula expandolabra, C.201493, paratype, Persephone Streamway, Bauhaus; B, E, Pseudotricula progenitor, C.201821, main streamway, Bauhaus. A, B, head and penis; C–E, stomach. Scales: A–E, 500 m. Abbreviations: dgo, opening from stomach to digestive gland; ey, eye; int, intestine; oes, oesophagus, p, penis; pg, penial duck; sn, snout; ss, style sack; st, stomach; tn, tentacle." figureDoi="http://doi.org/10.5281/zenodo.5050807" httpUri="https://zenodo.org/record/5050807/files/figure.png" pageId="32" pageNumber="33">9A</figureCitation>
) with weak swelling in mid­distal portion; distal end long, papilla­like; medial section parallel sided; of medium length; penial duct in medial section straight to undulating; base of penis moderately wide; with weak to moderate folds; penial duct straight to undulating.
</paragraph>
<paragraph id="8BBD36AC2D22A22CE42FFBD87F6CF98D" blockId="32.[264,1323,283,1550]" pageId="32" pageNumber="33">
<emphasis id="B976EABE2D22A22CE42FFBD87C6FFBAD" bold="true" box="[312,594,1044,1070]" pageId="32" pageNumber="33">Female genital system</emphasis>
(
<figureCitation id="13392A292D22A22CE771FBD87CCAFBAD" box="[614,759,1044,1070]" captionStart="FIGURE 10" captionStartId="22.[264,357,1296,1317]" captionTargetBox="[279,1315,341,1234]" captionTargetId="figure-191@22.[267,1319,329,1269]" captionTargetPageId="22" captionText="FIGURE 10. Female reproductive systems of Pseudotricula species. A ­ C, Pseudotricula expandolabra, A, B, C.201493, paratype, Persephone Streamway, Bauhaus, C, C.201815, Persephone Streamway, Bauhaus; D, Pseudotricula auriforma, C.201496, paratype, main streamway, Damper Cave; E, Pseudotricula arthurclarkei, C.203671, paratype, Quetzalcoatl Conduit; F, G, Pseudotricula conica, F, C203680, main streamway near entrance, Damper Cave, G, Cane Toad Abuse Streamway, Damper Cave. Scale: 500 m. Abbreviations: ag, albumen gland; bc, bursa copulatrix; bd, bursal duct; cg, capsule gland; co, coiled oviduct; po, genital opening; pw, pallial wall; sr, seminal receptacle; vc, ventral channel." figureDoi="http://doi.org/10.5281/zenodo.5050809" httpUri="https://zenodo.org/record/5050809/files/figure.png" pageId="32" pageNumber="33">Fig. 10F, G</figureCitation>
). Ovary of 0.6–1.0 whorls; oviduct extends to posterior edge of bursa copulatrix or slightly anterior or posterior to edge; straight distal to seminal receptacle; joins bursal duct in front of posterior pallial wall about half way between posterior pallial wall and capsule gland to about junction of albumen and capsule glands. Bursa copulatrix large, extending to posterior pallial wall or into pallial roof; elongately oval to pyriform; with bursal duct arising from middle of anterior edge of bursa or from antero­ventral edge; straight or with undulations. Seminal receptacle at middle of inner wall of bursa copulatrix or near mid ventral edge; ovoid to pyriform. About ½ to more than ½ of albumen gland in front of posterior pallial wall; capsule gland about same length as albumen gland to about ⅓ length of albumen gland; compressed oval in section; anterior end tapering to blunt; ventral channel simple, approximately parallel­sided throughout; vestibular area indistinct; genital opening overlapping anterior end of capsule gland.
</paragraph>
<paragraph id="8BBD36AC2D22A22CE41FF9887C18F9DE" blockId="32.[264,1323,1604,1750]" box="[264,549,1604,1629]" pageId="32" pageNumber="33">
<emphasis id="B976EABE2D22A22CE41FF9887C18F9DE" box="[264,549,1604,1629]" italics="true" pageId="32" pageNumber="33">Distribution and habitat</emphasis>
</paragraph>
<paragraph id="8BBD36AC2D22A22CE42FF9A07DABF955" blockId="32.[264,1323,1604,1750]" pageId="32" pageNumber="33">This is one of the commonest and widely distributed species in the caves. It appears to prefer low energy streams with mixed substrates, although a few specimens have been found in high energy habitats, notably in Cueva Blanca.</paragraph>
</subSubSection>
<subSubSection id="C31865272D22A22DE41FF8C07B18FC95" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33" type="discussion">
<paragraph id="8BBD36AC2D22A22DE41FF8C07A2BFDF5" blockId="32.[264,1323,1804,1870]" lastBlockId="33.[264,1325,284,790]" lastPageId="33" lastPageNumber="34" pageId="32" pageNumber="33">
<emphasis id="B976EABE2D22A22CE41FF8C07C09F8CE" box="[264,564,1804,1869]" italics="true" pageId="32" pageNumber="33">
Remarks
<taxonomicName id="4C024D2F2D22A22CE42FF8F87C09F8CE" authorityName="Ponder &amp; Clark &amp; Eberhard &amp; Studdert" authorityYear="2005" box="[312,564,1844,1869]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="32" pageNumber="33" phylum="Mollusca" rank="species" species="conica">Pseudotricula conica</taxonomicName>
</emphasis>
exhibits considerable variation in shell size and shape in what we have interpreted as a single species. In particular, samples from Cueva Blanca and Bauhaus caves are fairly consistently elongate (e.g.,
<figureCitation id="13392A292D23A22DE66EFE887DD8FEDD" box="[889,997,324,350]" captionStart="FIGURE 12" captionStartId="28.[264,357,1479,1500]" captionTargetBox="[382,1170,289,1464]" captionTargetId="figure-168@28.[354,1233,269,1476]" captionTargetPageId="28" captionText="FIGURE 12. Shells of Pseudotricula species. A, Pseudotricula arthurclarkei, holotype, C.439656, Quetzalcoatl Conduit; B ­ D, Pseudotricula conica, B, holotype, Cane Toad Abuse Streamway, Damper Cave; C, C.203680, main streamway near entrance, Damper Cave; D, C.201813, Persephone Pot, Bauhaus. Scales: A–D, 500 m." figureDoi="http://doi.org/10.5281/zenodo.5050815" httpUri="https://zenodo.org/record/5050815/files/figure.png" pageId="33" pageNumber="34">Fig. 12D</figureCitation>
;
<figureCitation id="13392A292D23A22DE6E4FE887A71FEDD" box="[1011,1100,324,350]" captionStart="FIGURE 13" captionStartId="30.[264,357,1326,1347]" captionTargetBox="[393,1223,370,1252]" captionTargetId="figure-178@30.[348,1252,362,1266]" captionTargetPageId="30" captionText="FIGURE 13. Shells of Pseudotricula species. A ­ F, Pseudotricula conica, A, C.203694, Black Curtain streamway, Cueva Blanca, B, Screaming Stals streamway, Bauhaus; C, C.201813, Persephone Pot, Bauhaus, D, C.203464, Cane Toad Abuse Streamway, Damper Cave, E, C.201488, main streamway, Damper Cave, F, C.203692, Damper Cave. Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5050817" httpUri="https://zenodo.org/record/5050817/files/figure.png" pageId="33" pageNumber="34">13A, C</figureCitation>
), while those from Damper Cave are usually broader (
<figureCitation id="13392A292D23A22DE7BDFEA07D07FE05" box="[682,826,364,390]" captionStart="FIGURE 12" captionStartId="28.[264,357,1479,1500]" captionTargetBox="[382,1170,289,1464]" captionTargetId="figure-168@28.[354,1233,269,1476]" captionTargetPageId="28" captionText="FIGURE 12. Shells of Pseudotricula species. A, Pseudotricula arthurclarkei, holotype, C.439656, Quetzalcoatl Conduit; B ­ D, Pseudotricula conica, B, holotype, Cane Toad Abuse Streamway, Damper Cave; C, C.203680, main streamway near entrance, Damper Cave; D, C.201813, Persephone Pot, Bauhaus. Scales: A–D, 500 m." figureDoi="http://doi.org/10.5281/zenodo.5050815" httpUri="https://zenodo.org/record/5050815/files/figure.png" pageId="33" pageNumber="34">Fig. 12B, C</figureCitation>
;
<figureCitation id="13392A292D23A22DE65FFEA07DA6FE05" box="[840,923,364,390]" captionStart="FIGURE 13" captionStartId="30.[264,357,1326,1347]" captionTargetBox="[393,1223,370,1252]" captionTargetId="figure-178@30.[348,1252,362,1266]" captionTargetPageId="30" captionText="FIGURE 13. Shells of Pseudotricula species. A ­ F, Pseudotricula conica, A, C.203694, Black Curtain streamway, Cueva Blanca, B, Screaming Stals streamway, Bauhaus; C, C.201813, Persephone Pot, Bauhaus, D, C.203464, Cane Toad Abuse Streamway, Damper Cave, E, C.201488, main streamway, Damper Cave, F, C.203692, Damper Cave. Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5050817" httpUri="https://zenodo.org/record/5050817/files/figure.png" pageId="33" pageNumber="34">13D–F</figureCitation>
). At least some material from the latter cave shows a range of shell morphology tall to broad with intermediate shell shapes (as for example in
<figureCitation id="13392A292D23A22DE4F3FE707C77FE55" box="[484,586,444,470]" captionStart="FIGURE 13" captionStartId="30.[264,357,1326,1347]" captionTargetBox="[393,1223,370,1252]" captionTargetId="figure-178@30.[348,1252,362,1266]" captionTargetPageId="30" captionText="FIGURE 13. Shells of Pseudotricula species. A ­ F, Pseudotricula conica, A, C.203694, Black Curtain streamway, Cueva Blanca, B, Screaming Stals streamway, Bauhaus; C, C.201813, Persephone Pot, Bauhaus, D, C.203464, Cane Toad Abuse Streamway, Damper Cave, E, C.201488, main streamway, Damper Cave, F, C.203692, Damper Cave. Scales: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.5050817" httpUri="https://zenodo.org/record/5050817/files/figure.png" pageId="33" pageNumber="34">Fig. 13B</figureCitation>
from Bauhaus). In addition, the anatomical and radular details of the broad and narrow forms are very similar. Because of this, and the existence of at least some apparent intermediates, we treat both forms as a single species pending more detailed studies. Specimens near the entrance to Damper cave (C.203680) are larger with rather dark­coloured shells than those from further inside the cave.
</paragraph>
<paragraph id="8BBD36AC2D23A22DE42FFD487B18FC95" blockId="33.[264,1325,284,790]" pageId="33" pageNumber="34">
<taxonomicName id="4C024D2F2D23A22DE42FFD487C0CFD1E" authorityName="Ponder &amp; Clark &amp; Eberhard &amp; Studdert" authorityYear="2005" box="[312,561,644,669]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="conica">
<emphasis id="B976EABE2D23A22DE42FFD487C0CFD1E" box="[312,561,644,669]" italics="true" pageId="33" pageNumber="34">Pseudotricula conica</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="4C024D2F2D23A22DE610FD487D82FD1E" authorityName="Ponder &amp; Clark &amp; Eberhard &amp; Studdert" authorityYear="2005" box="[775,959,644,669]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="arthurclarkei">
<emphasis id="B976EABE2D23A22DE610FD487D82FD1E" box="[775,959,644,669]" italics="true" pageId="33" pageNumber="34">P. arthurclarkei</emphasis>
</taxonomicName>
from Quetzalcoatl Conduit but differs in its smaller size (P&lt;0.001) although one lot from close to the entrance of Damper Cave (C.203680) is nearly as large as the former species.
<taxonomicName id="4C024D2F2D23A22DE6A6FD187A21FD6E" authorityName="Ponder &amp; Clark &amp; Eberhard &amp; Studdert" authorityYear="2005" box="[945,1052,724,749]" class="Gastropoda" family="Hydrobiidae" genus="Pseudotricula" kingdom="Animalia" order="Littorinimorpha" pageId="33" pageNumber="34" phylum="Mollusca" rank="species" species="conica">
<emphasis id="B976EABE2D23A22DE6A6FD187A21FD6E" box="[945,1052,724,749]" italics="true" pageId="33" pageNumber="34">P. conica</emphasis>
</taxonomicName>
also tends to have less convex whorls and has weak to obsolete opercular ridges. The two taxa are not sympatric.
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink