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<document id="686866BFA3AF14C23F68E8B657C83C18" ID-CLB-Dataset="171013" ID-DOI="10.5852/ejt.2022.843.1947" ID-GBIF-Dataset="044b60a6-0e4e-44be-b75a-614b58434dcc" ID-ISSN="2118-9773" ID-Zenodo-Dep="7222308" ID-ZooBank="C03BE897-EFE2-4CCD-913A-723792CDF050" IM.bibliography_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1666126358981" checkinUser="felipe" docAuthor="Wood, Thomas J. &amp; Monfared, Alireza" docDate="2022" docId="03AB87A2FFB5510DDA43DFE2FD1BFD8B" docLanguage="en" docName="ejt-843_1-136.pdf" docOrigin="European Journal of Taxonomy 843" docSource="http://dx.doi.org/10.5852/ejt.2022.843.1947" docStyle="DocumentStyle:EF2B578F1D15862ADE45B0C07C620911.14:EJT.2018-.journal_article.type1" docStyleId="EF2B578F1D15862ADE45B0C07C620911" docStyleName="EJT.2018-.journal_article.type1" docStyleVersion="14" docTitle="Andrena (Notandrena) idigna Wood &amp; Monfared 2022, sp. nov." docType="treatment" docUuid="E4762290-100B-4C49-94F9-5DB4273D990B" docUuidSource="ZooBank" docVersion="10" lastPageNumber="37" masterDocId="FF92FFDAFF945129D84DD914FFB0FFB4" masterDocTitle="A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species" masterLastPageNumber="136" masterPageNumber="1" pageNumber="34" updateTime="1699408809500" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="2377EC6C90DEED24C4E844B5D20C2A0B">A revision of the Andrena (Hymenoptera: Andrenidae) fauna of Iran, with the description of 16 new species</mods:title>
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<mods:namePart id="85B99786F86E33ECE35BA15992208AEF">Wood, Thomas J.</mods:namePart>
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<mods:affiliation id="AD9B041F52DF0A36B3FD9A46494336F1">Laboratory of Zoology, University of Mons, 7000 Mons, Belgium.</mods:affiliation>
<mods:nameIdentifier id="BA6D915750C621F24212DEBCD87B1746" type="email">thomasjames.wood@umons.ac.be</mods:nameIdentifier>
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<mods:namePart id="80C996CF69E85423DA18089DC1C99ED1">Monfared, Alireza</mods:namePart>
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<mods:affiliation id="56D18A1243B9B65BEA889539188FDEBB">Department of Plant Protection, Faculty of Agriculture, Yasouj University, Yasouj, Iran.</mods:affiliation>
<mods:nameIdentifier id="C527256E09E94DA1F5B3D7B65869C4D4" type="email">amonfared@yu.ac.ir</mods:nameIdentifier>
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<mods:date id="0DA12200CE2E3195B57D11258E289DDA">2022</mods:date>
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<treatment id="03AB87A2FFB5510DDA43DFE2FD1BFD8B" ID-DOI="http://doi.org/10.5281/zenodo.7237391" ID-GBIF-Taxon="203125091" ID-Zenodo-Dep="7237391" LSID="urn:lsid:zoobank.org:act:E4762290-100B-4C49-94F9-5DB4273D990B" httpUri="http://treatment.plazi.org/id/03AB87A2FFB5510DDA43DFE2FD1BFD8B" lastPageId="36" lastPageNumber="37" pageId="33" pageNumber="34">
<subSubSection id="C318653FFFB55108DA43DFE2FB96F8A5" box="[526,1062,1782,1809]" pageId="33" pageNumber="34" type="nomenclature">
<paragraph id="8BBD36B4FFB55108DA43DFE2FB96F8A5" blockId="33.[374,1213,1782,1876]" box="[526,1062,1782,1809]" pageId="33" pageNumber="34">
<heading id="D0F581D8FFB55108DA43DFE2FB96F8A5" box="[526,1062,1782,1809]" centered="true" fontSize="11" level="2" pageId="33" pageNumber="34" reason="2">
<taxonomicName id="4C024D37FFB55108DA43DFE2FC74F8A4" authority="Wood" authorityName="Wood &amp; Monfared" authorityYear="2022" box="[526,964,1782,1809]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="idigna" status="sp. nov." subGenus="Notandrena">
<emphasis id="B976EAA6FFB55108DA43DFE2FCC8F8A5" bold="true" box="[526,888,1782,1809]" pageId="33" pageNumber="34">
<emphasis id="B976EAA6FFB55108DA43DFE2FDC8F8A5" bold="true" box="[526,632,1782,1809]" italics="true" pageId="33" pageNumber="34">Andrena</emphasis>
(
<emphasis id="B976EAA6FFB55108DAC4DFE2FCAAF8A5" bold="true" box="[649,794,1782,1809]" italics="true" pageId="33" pageNumber="34">Notandrena</emphasis>
)
<emphasis id="B976EAA6FFB55108DB66DFE2FCC8F8A5" bold="true" box="[811,888,1782,1809]" italics="true" pageId="33" pageNumber="34">idigna</emphasis>
</emphasis>
Wood
</taxonomicName>
<taxonomicNameLabel id="A24557DDFFB55108DB81DFE3FB96F8A5" box="[972,1062,1783,1809]" pageId="33" pageNumber="34" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
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urn:lsid:zoobank.org:act:
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</paragraph>
<paragraph id="8BBD36B4FFB55108DA95DE2EFCECF8E0" blockId="33.[374,1213,1782,1876]" box="[728,860,1850,1876]" pageId="33" pageNumber="34">
<figureCitation id="13392A31FFB55108DA95DE2EFCECF8E0" box="[728,860,1850,1876]" captionStart="Figs 7984" captionStartId="35.[189,241,1963,1989]" captionTargetBox="[188,1397,563,1923]" captionTargetId="figure-107@35.[188,1397,563,1923]" captionTargetPageId="35" captionText="Figs 7984. Andrena idigna Wood sp. nov., ♀ (OÖLM). 79. Profile. 80. Face. 81. Face, detail. 82. Dorsum. 83. Terga. 84. Terga, detail." figureDoi="http://doi.org/10.5281/zenodo.7222367" httpUri="https://zenodo.org/record/7222367/files/figure.png" pageId="33" pageNumber="34">Figs 7984</figureCitation>
</paragraph>
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<subSubSection id="C318653FFFB5510BD8F0DE6EFCB1FCAF" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34" type="diagnosis">
<paragraph id="8BBD36B4FFB55108D8F0DE6EFE86F820" blockId="33.[189,310,1914,1940]" box="[189,310,1914,1940]" pageId="33" pageNumber="34">
<heading id="D0F581D8FFB55108D8F0DE6EFE86F820" bold="true" box="[189,310,1914,1940]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
<emphasis id="B976EAA6FFB55108D8F0DE6EFE86F820" bold="true" box="[189,310,1914,1940]" pageId="33" pageNumber="34">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36B4FFB5510BD8F0DEBCFCB1FCAF" blockId="33.[188,1398,1960,2022]" lastBlockId="34.[187,1401,265,795]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">
Placement is tricky because of taxonomic confusion surrounding taxa previously placed in the subgenus
<taxonomicName id="4C024D37FFB55108D8F1DEDFFDB3F851" authority="Warncke, 1968" authorityName="Warncke" authorityYear="1968" box="[188,515,1995,2021]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="subGenus" subGenus="Carandrena">
<emphasis id="B976EAA6FFB55108D8F1DEDFFEFBF851" box="[188,331,1995,2021]" italics="true" pageId="33" pageNumber="34">Carandrena</emphasis>
<bibRefCitation id="EF934B45FFB55108D91CDEDFFDB3F851" author="Warncke" box="[337,515,1995,2021]" firstAuthor="Warncke" pageId="33" pageNumber="34" pagination="1 - 110" refId="ref70132" refString="Warncke K. 1968. Die Untergattungen der westpalaarktischen Bienengattung Andrena F. Memorias e Estudos do Museu Zoologico da Universidade de Coimbra 307: 1 - 110." type="journal article" year="1968">Warncke, 1968</bibRefCitation>
</taxonomicName>
, which is now a strict synonym of
<taxonomicName id="4C024D37FFB55108DBDEDEDFFB01F851" authority="Perez, 1890" authorityName="Perez" authorityYear="1890" box="[915,1201,1995,2021]" class="Insecta" family="Andrenidae" genus="Notandrena" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="128" phylum="Arthropoda" rank="genus">
<emphasis id="B976EAA6FFB55108DBDEDEDFFBAEF851" box="[915,1054,1995,2021]" italics="true" pageId="33" pageNumber="34">Notandrena</emphasis>
Pérez, 1890
</taxonomicName>
, but
<taxonomicName id="4C024D37FFB55108DCAADEDFFAC5F851" authorityName="Warncke" authorityYear="1968" box="[1255,1397,1995,2021]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="33" pageNumber="133" phylum="Arthropoda" rank="subGenus" subGenus="Carandrena">
<emphasis id="B976EAA6FFB55108DCAADEDFFAC5F851" box="[1255,1397,1995,2021]" italics="true" pageId="33" pageNumber="34">Carandrena</emphasis>
</taxonomicName>
sensu Warncke is polyphyletic (
<bibRefCitation id="EF934B45FFB6510BDA67D81DFCA5FE97" author="Pisanty" box="[554,789,265,292]" etAl="et al." firstAuthor="Pisanty" pageId="34" pageNumber="35" pagination="107151" refId="ref68841" refString="Pisanty G., Richter R., Martin T., Dettman J. &amp; Cardinal S. 2022 b. Molecular phylogeny, historical biogeography and revised classification of andrenine bees (Hymenoptera: Andrenidae). Molecular Phylogenetics and Evolution 170: e 107151. https: // doi. org / 10.1016 / j. ympev. 2021.107151" type="journal article" year="2022" yearSuffix="b">
Pisanty
<emphasis id="B976EAA6FFB6510BDACAD81EFD09FE97" box="[647,697,265,291]" italics="true" pageId="34" pageNumber="35">et al</emphasis>
. 2022b
</bibRefCitation>
).
<taxonomicName id="4C024D37FFB6510BDB6BD81DFC6AFE97" authority="Wood &amp; Monfared, 2022" authorityName="Wood &amp; Monfared" authorityYear="2022" box="[806,986,265,291]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="idigna" status="sp. nov.">
<emphasis id="B976EAA6FFB6510BDB6BD81DFC6AFE97" box="[806,986,265,291]" italics="true" pageId="34" pageNumber="35">Andrena idigna</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A24557DDFFB6510BDBAFD81EFB8BFE90" box="[994,1083,266,292]" pageId="34" pageNumber="35" rank="species">sp. nov.</taxonomicNameLabel>
has the general appearance of a
<taxonomicName id="4C024D37FFB6510BD8BDD839FECEFEF3" authorityName="Warncke" authorityYear="1968" box="[240,382,301,327]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="133" phylum="Arthropoda" rank="subGenus" subGenus="Carandrena">
<emphasis id="B976EAA6FFB6510BD8BDD839FECEFEF3" box="[240,382,301,327]" italics="true" pageId="34" pageNumber="35">Carandrena</emphasis>
</taxonomicName>
because of its broad head (
<figureCitation id="13392A31FFB6510BDAF0D839FCA3FEF3" box="[701,787,301,327]" captionStart="Figs 7984" captionStartId="35.[189,241,1963,1989]" captionTargetBox="[188,1397,563,1923]" captionTargetId="figure-107@35.[188,1397,563,1923]" captionTargetPageId="35" captionText="Figs 7984. Andrena idigna Wood sp. nov., ♀ (OÖLM). 79. Profile. 80. Face. 81. Face, detail. 82. Dorsum. 83. Terga. 84. Terga, detail." figureDoi="http://doi.org/10.5281/zenodo.7222367" httpUri="https://zenodo.org/record/7222367/files/figure.png" pageId="34" pageNumber="35">Fig. 81</figureCitation>
), tergal hairbands (
<figureCitation id="13392A31FFB6510BDBB9D839FBFAFEF3" box="[1012,1098,301,327]" captionStart="Figs 7984" captionStartId="35.[189,241,1963,1989]" captionTargetBox="[188,1397,563,1923]" captionTargetId="figure-107@35.[188,1397,563,1923]" captionTargetPageId="35" captionText="Figs 7984. Andrena idigna Wood sp. nov., ♀ (OÖLM). 79. Profile. 80. Face. 81. Face, detail. 82. Dorsum. 83. Terga. 84. Terga, detail." figureDoi="http://doi.org/10.5281/zenodo.7222367" httpUri="https://zenodo.org/record/7222367/files/figure.png" pageId="34" pageNumber="35">Fig. 84</figureCitation>
), and lack of any obvious derived features, one of the reasons why this former subgenus has been used as a waste basket for species that cannot be easily allocated elsewhere. Its integument lacks metallic reflections or red colouration on the terga, immediately separating it from almost all species in this group. It is therefore superficially similar to
<taxonomicName id="4C024D37FFB6510BD99FD8A9FD50FE6C" authority="Perez, 1895" authorityName="Perez" authorityYear="1895" box="[466,736,445,472]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="euzona">
<emphasis id="B976EAA6FFB6510BD99FD8A9FDF8FE63" box="[466,584,445,471]" italics="true" pageId="34" pageNumber="35">A. euzona</emphasis>
Pérez, 1895
</taxonomicName>
, which also has dark terga. However,
<taxonomicName id="4C024D37FFB6510BDCE0D8A9FA93FE63" authorityName="Perez" authorityYear="1895" box="[1197,1315,445,471]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="euzona">
<emphasis id="B976EAA6FFB6510BDCE0D8A9FA93FE63" box="[1197,1315,445,471]" italics="true" pageId="34" pageNumber="35">A. euzona</emphasis>
</taxonomicName>
has an extremely shiny scutum and is found outside the
<taxonomicName id="4C024D37FFB6510BDAB9D8F5FC32FE4F" authorityName="Warncke" authorityYear="1968" box="[756,898,481,507]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="133" phylum="Arthropoda" rank="subGenus" subGenus="Carandrena">
<emphasis id="B976EAA6FFB6510BDAB9D8F5FC32FE4F" box="[756,898,481,507]" italics="true" pageId="34" pageNumber="35">Carandrena</emphasis>
</taxonomicName>
+
<taxonomicName id="4C024D37FFB6510BDBD9D8F5FBAFFE4F" box="[916,1055,481,507]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="128" phylum="Arthropoda" rank="subGenus" subGenus="Notandrena">
<emphasis id="B976EAA6FFB6510BDBD9D8F5FBAFFE4F" box="[916,1055,481,507]" italics="true" pageId="34" pageNumber="35">Notandrena</emphasis>
</taxonomicName>
group, whereas
<taxonomicName id="4C024D37FFB6510BDC90D8F5FAF8FE4F" authorityName="Wood &amp; Monfared" authorityYear="2022" box="[1245,1352,481,507]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="idigna">
<emphasis id="B976EAA6FFB6510BDC90D8F5FAF8FE4F" box="[1245,1352,481,507]" italics="true" pageId="34" pageNumber="35">A. idigna</emphasis>
</taxonomicName>
has the scutum entirely and uniformly shagreened and dull. Though
<taxonomicName id="4C024D37FFB6510BDBF0DB11FB9AFDAB" authorityName="Wood &amp; Monfared" authorityYear="2022" box="[957,1066,517,543]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="idigna">
<emphasis id="B976EAA6FFB6510BDBF0DB11FB9AFDAB" box="[957,1066,517,543]" italics="true" pageId="34" pageNumber="35">A. idigna</emphasis>
</taxonomicName>
lacks a clear humeral angle on the pronotum (faint hint of this trait present but obscure), a character that is usually present in the
<taxonomicName id="4C024D37FFB6510BD8F2DB59FEF8FDD3" box="[191,328,589,615]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="128" phylum="Arthropoda" rank="subGenus" subGenus="Notandrena">
<emphasis id="B976EAA6FFB6510BD8F2DB59FEF8FDD3" box="[191,328,589,615]" italics="true" pageId="34" pageNumber="35">Notandrena</emphasis>
</taxonomicName>
, provisional placement in this subgenus is the most appropriate because of the broad head, shagreened scutum, shagreened and essentially impunctate terga, and tergal hairbands. Its dark terga, shagreened scutum, and unusual propodeal triangle, which is delineated laterally by fine carinae but with a narrow, longitudinal, and slightly impressed channel surrounded by irregular latitudinal rugae along its length, allow separation from all related taxa. This subgeneric placement should be reassessed following discovery of male specimens and DNA sampling.
</paragraph>
</subSubSection>
<subSubSection id="C318653FFFB6510BD8F0DA57FD2DFC05" pageId="34" pageNumber="35" type="etymology">
<paragraph id="8BBD36B4FFB6510BD8F0DA57FEF4FCE9" blockId="34.[189,324,835,861]" box="[189,324,835,861]" pageId="34" pageNumber="35">
<heading id="D0F581D8FFB6510BD8F0DA57FEF4FCE9" bold="true" box="[189,324,835,861]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="B976EAA6FFB6510BD8F0DA57FEF4FCE9" bold="true" box="[189,324,835,861]" pageId="34" pageNumber="35">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36B4FFB6510BD8F0DA67FD2DFC05" blockId="34.[189,1398,883,945]" pageId="34" pageNumber="35">
From the Sumerian
<taxonomicName id="4C024D37FFB6510BD9F9DA67FE4FFC39" authorityName="Wood &amp; Monfared" authorityYear="2022" box="[436,511,883,909]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="idigna">
<emphasis id="B976EAA6FFB6510BD9F9DA67FE4FFC39" box="[436,511,883,909]" italics="true" pageId="34" pageNumber="35">idigna</emphasis>
</taxonomicName>
or
<emphasis id="B976EAA6FFB6510BDA77DA67FD3DFC39" box="[570,653,883,909]" italics="true" pageId="34" pageNumber="35">idigina</emphasis>
, meaning running water, in reference to its locus typicus on the Karun River, the largest river in
<collectingCountry id="F3157624FFB6510BDA2BDA83FD29FC05" box="[614,665,919,945]" name="Iran" pageId="34" pageNumber="35">Iran</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C318653FFFB6510BD8F0DACDFBF7FBFD" pageId="34" pageNumber="35" type="materials_examined">
<paragraph id="8BBD36B4FFB6510BD8F0DACDFE1AFC47" blockId="34.[189,426,985,1011]" box="[189,426,985,1011]" pageId="34" pageNumber="35">
<heading id="D0F581D8FFB6510BD8F0DACDFE1AFC47" bold="true" box="[189,426,985,1011]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="B976EAA6FFB6510BD8F0DACDFE1AFC47" bold="true" box="[189,426,985,1011]" pageId="34" pageNumber="35">Material examined</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36B4FFB6510BD8A7DD1CFBF7FBFD" blockId="34.[189,1095,1032,1097]" pageId="34" pageNumber="35">
<materialsCitation id="3B6A3CE9FFB6510BD8A7DD1CFBF7FBFD" ID-GBIF-Occurrence="3944044015" collectingDate="1965-02-17" collectionCode="OOLM" collectorName="S. Tirgari" country="Iran" location="Ahwaz" pageId="34" pageNumber="35" specimenCount="1" specimenCount-female="1" stateProvince="Mollasani" typeStatus="holotype">
<emphasis id="B976EAA6FFB6510BD8A7DD1CFEECFB96" bold="true" box="[234,348,1032,1058]" pageId="34" pageNumber="35">
<typeStatus id="54B98816FFB6510BD8A7DD1CFEECFB96" box="[234,348,1032,1058]" pageId="34" pageNumber="35" type="holotype">Holotype</typeStatus>
</emphasis>
<collectingCountry id="F3157624FFB6510BD8F0DD38FEB5FBF2" box="[189,261,1068,1094]" name="Iran" pageId="34" pageNumber="35">IRAN</collectingCountry>
<specimenCount id="9D04FD3DFFB6510BD953DD39FE85FBF3" box="[286,309,1069,1095]" pageId="34" pageNumber="35" type="female"></specimenCount>
;
<collectingRegion id="49C6F856FFB6510BD90EDD38FE0BFBF2" box="[323,443,1068,1094]" pageId="34" pageNumber="35">Mollasani</collectingRegion>
,
<location id="8EDD606FFFB6510BD988DD38FDA8FBF2" LSID="urn:lsid:plazi:treatment:03AB87A2FFB5510DDA43DFE2FD1BFD8B:8EDD606FFFB6510BD988DD38FDA8FBF2" box="[453,536,1068,1094]" country="Iran" name="Ahwaz" pageId="34" pageNumber="35" stateProvince="Mollasani">Ahwaz</location>
[Ahvaz];
<date id="FFBC1074FFB6510BDAC2DD38FC9DFBF3" box="[655,813,1068,1095]" pageId="34" pageNumber="35" value="1965-02-17">
<collectingDate id="EFF8E99CFFB6510BDAC2DD38FC9DFBF3" box="[655,813,1068,1095]" pageId="34" pageNumber="35" value="1965-02-17">17 Feb. 1965</collectingDate>
</date>
;
<collectorName id="26F75362FFB6510BDB74DD38FC17FBF2" box="[825,935,1068,1094]" pageId="34" pageNumber="35">S. Tirgari</collectorName>
leg.;
<collectionCode id="ED13AE71FFB6510BDBA4DD39FBF3FBFD" box="[1001,1091,1069,1097]" collectionName="OOLM" pageId="34" pageNumber="35">OÖLM</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C318653FFFB6510DD8F0DD7AFE89FE7B" lastPageId="36" lastPageNumber="37" pageId="34" pageNumber="35" type="description">
<paragraph id="8BBD36B4FFB6510BD8F0DD7AFEFEFB3C" blockId="34.[189,334,1134,1160]" box="[189,334,1134,1160]" pageId="34" pageNumber="35">
<heading id="D0F581D8FFB6510BD8F0DD7AFEFEFB3C" bold="true" box="[189,334,1134,1160]" fontSize="11" level="3" pageId="34" pageNumber="35" reason="3">
<emphasis id="B976EAA6FFB6510BD8F0DD7AFEFEFB3C" bold="true" box="[189,334,1134,1160]" pageId="34" pageNumber="35">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36B4FFB6510BD8A7DD8AFEF5FB0C" blockId="34.[189,716,1182,1244]" box="[234,325,1182,1208]" pageId="34" pageNumber="35">
<emphasis id="B976EAA6FFB6510BD8A7DD8AFEF5FB0C" bold="true" box="[234,325,1182,1208]" pageId="34" pageNumber="35">Female</emphasis>
</paragraph>
<paragraph id="8BBD36B4FFB6510BD8F0DDD6FD7CFB68" blockId="34.[189,716,1182,1244]" box="[189,716,1218,1244]" pageId="34" pageNumber="35">
MEASUREMENTS. Body length
<quantity id="4CFA9B51FFB6510BDA5EDDD6FDE7FB68" box="[531,599,1218,1244]" metricMagnitude="-3" metricUnit="m" metricValue="9.0" pageId="34" pageNumber="35" unit="mm" value="9.0">9 mm</quantity>
(
<figureCitation id="13392A31FFB6510BDA2BDDD6FD0EFB68" box="[614,702,1218,1244]" captionStart="Figs 7984" captionStartId="35.[189,241,1963,1989]" captionTargetBox="[188,1397,563,1923]" captionTargetId="figure-107@35.[188,1397,563,1923]" captionTargetPageId="35" captionText="Figs 7984. Andrena idigna Wood sp. nov., ♀ (OÖLM). 79. Profile. 80. Face. 81. Face, detail. 82. Dorsum. 83. Terga. 84. Terga, detail." figureDoi="http://doi.org/10.5281/zenodo.7222367" httpUri="https://zenodo.org/record/7222367/files/figure.png" pageId="34" pageNumber="35">Fig. 79</figureCitation>
).
</paragraph>
<paragraph id="8BBD36B4FFB6510BD8F0DC11FB24F9AF" blockId="34.[189,1400,1284,1563]" pageId="34" pageNumber="35">
HEAD. 1.4 times as wide as long (
<figureCitation id="13392A31FFB6510BDA1DDC10FD1AFAAA" box="[592,682,1284,1310]" captionStart="Figs 7984" captionStartId="35.[189,241,1963,1989]" captionTargetBox="[188,1397,563,1923]" captionTargetId="figure-107@35.[188,1397,563,1923]" captionTargetPageId="35" captionText="Figs 7984. Andrena idigna Wood sp. nov., ♀ (OÖLM). 79. Profile. 80. Face. 81. Face, detail. 82. Dorsum. 83. Terga. 84. Terga, detail." figureDoi="http://doi.org/10.5281/zenodo.7222367" httpUri="https://zenodo.org/record/7222367/files/figure.png" pageId="34" pageNumber="35">Fig. 80</figureCitation>
). Clypeus dark, weakly domed, densely punctate, punctures separated by 0.52 puncture diameters (
<figureCitation id="13392A31FFB6510BDAD1DC3CFD43FAF6" box="[668,755,1320,1346]" captionStart="Figs 7984" captionStartId="35.[189,241,1963,1989]" captionTargetBox="[188,1397,563,1923]" captionTargetId="figure-107@35.[188,1397,563,1923]" captionTargetPageId="35" captionText="Figs 7984. Andrena idigna Wood sp. nov., ♀ (OÖLM). 79. Profile. 80. Face. 81. Face, detail. 82. Dorsum. 83. Terga. 84. Terga, detail." figureDoi="http://doi.org/10.5281/zenodo.7222367" httpUri="https://zenodo.org/record/7222367/files/figure.png" pageId="34" pageNumber="35">Fig. 81</figureCitation>
). Clypeus with fine and dense granular shagreen, dull to weakly shining centrally. Process of labrum trapezoidal, two times as wide as long, corners rounded. Face, gena, vertex, and scape with moderately long white hairs, not exceeding length of scape. Gena slightly exceeding width of compound eye; ocelloccipital distance subequal to diameter of lateral ocellus. Foveae dorsally broad, occupying ⅔
<superScript id="7C779BFCFFB6510BDA85DCA3FD66FA71" attach="left" box="[712,726,1463,1477]" fontSize="6" pageId="34" pageNumber="35">rd</superScript>
of space between compound eye and lateral ocellus, very slightly narrowed below at level of antennal insertions; filled with whitish hairs. Antennae dark basally, A512 lightened dark orange-red below; A3 exceeding A4+5, shorter than A4+5+6.
</paragraph>
<paragraph id="8BBD36B4FFB6510AD8F0DF57FE89FEF2" blockId="34.[188,1401,1602,2025]" lastBlockId="35.[189,1398,265,326]" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35">
MESOSOMA. Scutum and scutellum densely and uniformly granularly shagreened, dull; surface with irregular and obscure punctures, punctures separated by 23 puncture diameters, not well-differentiated from underlying surface (
<figureCitation id="13392A31FFB6510BD9BBDF9EFDE3F910" box="[502,595,1674,1700]" captionStart="Figs 7984" captionStartId="35.[189,241,1963,1989]" captionTargetBox="[188,1397,563,1923]" captionTargetId="figure-107@35.[188,1397,563,1923]" captionTargetPageId="35" captionText="Figs 7984. Andrena idigna Wood sp. nov., ♀ (OÖLM). 79. Profile. 80. Face. 81. Face, detail. 82. Dorsum. 83. Terga. 84. Terga, detail." figureDoi="http://doi.org/10.5281/zenodo.7222367" httpUri="https://zenodo.org/record/7222367/files/figure.png" pageId="34" pageNumber="35">Fig. 82</figureCitation>
). Pronotum without clear humeral angle, slightly angulate above, obscure. Mesepisternum and lateral and dorsolateral faces of propodeum with fine granular shagreen, dull; dorsolateral faces of propodeum with sparse network of fine, slightly raised rugae. Propodeal triangle declineated laterally with fine carinae, internal surface with narrow, longitudinal, and slightly impressed channel surrounded by irregular latitudinal rugae along its length. Remaining surface of propodeal triangle with fine granular shagreen, finer than on dorsolateral faces of propodeum; propodeal triangle thus defined by change in surface sculpture. Mesosoma with moderately long whitish to greyish hairs, not exceeding length of scape; propodeal corbiculae with weakly plumose hairs dorsally, inner surface with weakly plumose hairs. Legs dark, apical tarsal segments lightened dark orange; pubescence whitish. Scopae and flocculus white. Hind tarsal claws with strong inner tooth. Wings hyaline, stigma and venation orange, nervulus weakly antefurcal; first recurrent vein enters second submarginal cell at its middle.
</paragraph>
<paragraph id="8BBD36B4FFB7510DD8F0D87AFD50FEDE" blockId="35.[189,1399,366,498]" lastBlockId="36.[189,1399,265,362]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
METASOMA. Terga dark, apical margins of T24 slightly depressed, marginal areas of T14 lightened dark red to yellow-hyaline apically (
<figureCitation id="13392A31FFB7510ADA3ED885FD7EFE1F" box="[627,718,401,427]" captionStart="Figs 7984" captionStartId="35.[189,241,1963,1989]" captionTargetBox="[188,1397,563,1923]" captionTargetId="figure-107@35.[188,1397,563,1923]" captionTargetPageId="35" captionText="Figs 7984. Andrena idigna Wood sp. nov., ♀ (OÖLM). 79. Profile. 80. Face. 81. Face, detail. 82. Dorsum. 83. Terga. 84. Terga, detail." figureDoi="http://doi.org/10.5281/zenodo.7222367" httpUri="https://zenodo.org/record/7222367/files/figure.png" pageId="35" pageNumber="36">Fig. 83</figureCitation>
). Terga with uniform granular shagreen, weakly shining, essentially impunctate with only scattered and obscure hair-bearing punctures (
<figureCitation id="13392A31FFB7510ADC35D8A0FB61FE7A" box="[1144,1233,436,462]" captionStart="Figs 7984" captionStartId="35.[189,241,1963,1989]" captionTargetBox="[188,1397,563,1923]" captionTargetId="figure-107@35.[188,1397,563,1923]" captionTargetPageId="35" captionText="Figs 7984. Andrena idigna Wood sp. nov., ♀ (OÖLM). 79. Profile. 80. Face. 81. Face, detail. 82. Dorsum. 83. Terga. 84. Terga, detail." figureDoi="http://doi.org/10.5281/zenodo.7222367" httpUri="https://zenodo.org/record/7222367/files/figure.png" pageId="35" pageNumber="36">Fig. 84</figureCitation>
). Tergal discs with sparse whitish hairs, longest on T1, becoming progressively shorter apically. T14 with white apical hair fringes, weak and interrupted on T1, complete and dense on T24, obscuring underlying surface. Fringe of T5 and hairs flanking pygidial plate golden; pygidial plate broadly rounded, centrally densely punctate, with narrow impunctate rim.
</paragraph>
<caption id="DF7D663CFFB7510AD8F0DEBFFDDBF85D" ID-DOI="http://doi.org/10.5281/zenodo.7222367" ID-Zenodo-Dep="7222367" httpUri="https://zenodo.org/record/7222367/files/figure.png" pageId="35" pageNumber="36" startId="35.[189,241,1963,1989]" targetBox="[188,1397,563,1923]" targetPageId="35">
<paragraph id="8BBD36B4FFB7510AD8F0DEBFFDDBF85D" blockId="35.[189,1398,1963,2026]" pageId="35" pageNumber="36">
<emphasis id="B976EAA6FFB7510AD8F0DEBFFEFDF872" bold="true" box="[189,333,1963,1990]" pageId="35" pageNumber="36">Figs 7984.</emphasis>
<taxonomicName id="4C024D37FFB7510AD915DEB8FDD1F872" authority="Wood" authorityName="Wood &amp; Monfared" authorityYear="2022" box="[344,609,1964,1990]" class="Insecta" family="Andrenidae" genus="Andrena" kingdom="Animalia" order="Hymenoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="idigna" status="sp. nov.">
<emphasis id="B976EAA6FFB7510AD915DEB8FDA2F872" box="[344,530,1964,1990]" italics="true" pageId="35" pageNumber="36">Andrena idigna</emphasis>
Wood
</taxonomicName>
<taxonomicNameLabel id="A24557DDFFB7510ADA20DEB8FD7DF872" box="[621,717,1964,1990]" pageId="35" pageNumber="36" rank="species">sp. nov.</taxonomicNameLabel>
, ♀ (OÖLM).
<emphasis id="B976EAA6FFB7510ADB37DEB8FC27F872" bold="true" box="[890,919,1964,1990]" pageId="35" pageNumber="36">79</emphasis>
. Profile.
<emphasis id="B976EAA6FFB7510ADC46DEB8FB98F872" bold="true" box="[1035,1064,1964,1990]" pageId="35" pageNumber="36">80</emphasis>
. Face.
<emphasis id="B976EAA6FFB7510ADCC8DEB8FB12F872" bold="true" box="[1157,1186,1964,1990]" pageId="35" pageNumber="36">81</emphasis>
. Face, detail.
<emphasis id="B976EAA6FFB7510ADD1FDEB8FADFF872" bold="true" box="[1362,1391,1964,1990]" pageId="35" pageNumber="36">82</emphasis>
. Dorsum.
<emphasis id="B976EAA6FFB7510AD967DEDBFEF7F85D" bold="true" box="[298,327,1999,2025]" pageId="35" pageNumber="36">83</emphasis>
. Terga.
<emphasis id="B976EAA6FFB7510AD9EBDEDBFE73F85D" bold="true" box="[422,451,1999,2025]" pageId="35" pageNumber="36">84</emphasis>
. Terga, detail.
</paragraph>
</caption>
<paragraph id="8BBD36B4FFB0510DD8A7D886FE9AFE18" blockId="36.[189,313,402,463]" box="[234,298,402,428]" pageId="36" pageNumber="37">
<emphasis id="B976EAA6FFB0510DD8A7D886FE9AFE18" bold="true" box="[234,298,402,428]" pageId="36" pageNumber="37">Male</emphasis>
</paragraph>
<paragraph id="8BBD36B4FFB0510DD8F0D8A1FE89FE7B" blockId="36.[189,313,402,463]" box="[189,313,437,463]" pageId="36" pageNumber="37">Unknown.</paragraph>
</subSubSection>
<subSubSection id="C318653FFFB0510DD8F0D8E2FD1BFD8B" box="[189,683,502,576]" pageId="36" pageNumber="37" type="distribution">
<paragraph id="8BBD36B4FFB0510DD8F0D8E2FEE6FDA4" blockId="36.[189,342,502,528]" box="[189,342,502,528]" pageId="36" pageNumber="37">
<heading id="D0F581D8FFB0510DD8F0D8E2FEE6FDA4" bold="true" box="[189,342,502,528]" fontSize="11" level="3" pageId="36" pageNumber="37" reason="3">
<emphasis id="B976EAA6FFB0510DD8F0D8E2FEE6FDA4" bold="true" box="[189,342,502,528]" pageId="36" pageNumber="37">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BBD36B4FFB0510DD8F0DB31FD1BFD8B" blockId="36.[189,683,549,576]" box="[189,683,549,576]" pageId="36" pageNumber="37">
South-western
<collectingCountry id="F3157624FFB0510DD923DB32FE2DFDF4" box="[366,413,550,576]" name="Iran" pageId="36" pageNumber="37">Iran</collectingCountry>
(
<collectingRegion id="49C6F856FFB0510DD9E1DB31FD2FFD8B" box="[428,671,549,575]" country="Iran" name="Khuzestan" pageId="36" pageNumber="37">Khuzestan Province</collectingRegion>
).
</paragraph>
</subSubSection>
</treatment>
</document>