plazi/treatments-xml
2
0
Fork 0
Code Issues Pull requests Releases Activity
treatments-xml/data/03/AB/39/03AB390CC061FFAE7DEBFD88361CFCD7.xml
maintenance bc2564723d added 03
2024-06-21 12:22:17 +02:00

329 lines
38 KiB
XML
Raw Blame History

This file contains ambiguous Unicode characters

This file contains Unicode characters that might be confused with other characters. If you think that this is intentional, you can safely ignore this warning. Use the Escape button to reveal them.

<document id="2C8FC095D61B8A3BF1B629A1BE632ACB" ID-DOI="10.5281/zenodo.190501" ID-GBIF-Dataset="1f0e8267-5d03-4f65-82e7-5ab451be4a39" ID-ISSN="1175-5326" ID-Zenodo-Dep="190501" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461114725693" checkinUser="plazi" docAuthor="Paulson, Dennis R." docDate="2009" docId="03AB390CC061FFAE7DEBFD88361CFCD7" docLanguage="en" docName="zt02239p068.pdf" docOrigin="Zootaxa 2239" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Leptobasis guanacaste Paulson, 2009, n. sp." docType="treatment" docVersion="9" lastPageNumber="66" masterDocId="FF924174C060FFAA7D7CFFC93304FFBD" masterDocTitle="A new species of Leptobasis from Costa Rica (Odonata: Coenagrionidae)" masterLastPageNumber="68" masterPageNumber="62" pageNumber="63" updateTime="1698594834445" updateUser="plazi">
<mods:mods id="5B36DC92F0881C3E54C7E95B06A070BB" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="DE052B13C4AA0B6A805BD531E9B22E1E">
<mods:title id="064D46F7BD215947C9730A160A5417F0">A new species of Leptobasis from Costa Rica (Odonata: Coenagrionidae)</mods:title>
</mods:titleInfo>
<mods:name id="9B261C3991596D9971EEF19BCF19FCE2" type="personal">
<mods:role id="26B964A2B632704E1F70486EEE1B8244">
<mods:roleTerm id="74C0FE1E2CB94B280D220B21909B53F6">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="2368B59DAD15ADCE823370477681DB37">Paulson, Dennis R.</mods:namePart>
</mods:name>
<mods:typeOfResource id="62FADB93F9345333C5FC3518E2C27854">text</mods:typeOfResource>
<mods:relatedItem id="6405B98A57E1F391223A2831630CEBA2" type="host">
<mods:titleInfo id="7ED9CA56D652C25782C60C1EB659123B">
<mods:title id="D29C68E3F35213D2E9AFA1C62F365B61">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="F54383A5C492CA6F3DB0E8E9EFE86398">
<mods:date id="5136C5257BCF77DCF04D6B3747B8F2BE">2009</mods:date>
<mods:detail id="F8728C396ED2B261982DB4C94A1132BD" type="volume">
<mods:number id="086300D96A7BFB3CE36CF2F5713554A8">2239</mods:number>
</mods:detail>
<mods:extent id="34D98CA32653B6E434FB10C1BFBD024F" unit="page">
<mods:start id="B3BF008323642468D74C8E805D29EAA6">62</mods:start>
<mods:end id="8E84BB0C1E4AF558EDE6C04A1B1D6F83">68</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="5162357681BD23F5368F1561AA37FB95">journal article</mods:classification>
<mods:identifier id="2BE27759F409E9903A3B88BB867CCF99" type="DOI">10.5281/zenodo.190501</mods:identifier>
<mods:identifier id="D951138A2F31434BC4E4613ACCCA2625" type="GBIF-Dataset">1f0e8267-5d03-4f65-82e7-5ab451be4a39</mods:identifier>
<mods:identifier id="130D7ABAD0BA47B15AF2F64F3BE9659B" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="E1D3A2BAB34AABDCE9EC44099D03B818" type="Zenodo-Dep">190501</mods:identifier>
</mods:mods>
<treatment id="03AB390CC061FFAE7DEBFD88361CFCD7" ID-DOI="http://doi.org/10.5281/zenodo.3503106" ID-GBIF-Taxon="119624572" ID-Zenodo-Dep="3503106" LSID="urn:lsid:plazi:treatment:03AB390CC061FFAE7DEBFD88361CFCD7" httpUri="http://treatment.plazi.org/id/03AB390CC061FFAE7DEBFD88361CFCD7" lastPageId="4" lastPageNumber="66" pageId="1" pageNumber="63">
<subSubSection id="C318DB91C061FFAB7DEBFD8832F5FDE7" box="[151,497,576,603]" pageId="1" pageNumber="63" type="nomenclature">
<paragraph id="8BBD881AC061FFAB7DEBFD8832F5FDE7" blockId="1.[151,497,576,603]" box="[151,497,576,603]" pageId="1" pageNumber="63">
<heading id="D0F53F76C061FFAB7DEBFD8832F5FDE7" bold="true" box="[151,497,576,603]" fontSize="11" level="1" pageId="1" pageNumber="63" reason="1">
<emphasis id="B9765408C061FFAB7DEBFD8832F5FDE7" bold="true" box="[151,497,576,603]" pageId="1" pageNumber="63">
<taxonomicName id="4C02F399C061FFAB7DEBFD8832ACFDE6" ID-CoL="3TJZL" box="[151,424,577,603]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="63" phylum="Arthropoda" rank="species" species="guanacaste" status="sp. nov.">
<emphasis id="B9765408C061FFAB7DEBFD8832ACFDE6" bold="true" box="[151,424,577,603]" italics="true" pageId="1" pageNumber="63">Leptobasis guanacaste</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A245E973C061FFAB7CD3FD8932F5FDE7" box="[431,497,576,602]" pageId="1" pageNumber="63" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C318DB91C061FFAB7DEBFD433098FC31" pageId="1" pageNumber="63" type="materials_examined">
<paragraph id="8BBD881AC061FFAB7DEBFD433098FC31" blockId="1.[151,1437,650,2030]" pageId="1" pageNumber="63">
<emphasis id="B9765408C061FFAB7DEBFD433247FD19" bold="true" box="[151,323,650,676]" pageId="1" pageNumber="63">
<typeStatus id="54B936B8C061FFAB7DEBFD4333D6FD19" box="[151,210,650,676]" pageId="1" pageNumber="63">Type</typeStatus>
material
</emphasis>
.
<emphasis id="B9765408C061FFAB7C28FD4332C2FD19" bold="true" box="[340,454,650,676]" pageId="1" pageNumber="63">
<typeStatus id="54B936B8C061FFAB7C28FD4332C2FD19" box="[340,454,650,676]" pageId="1" pageNumber="63" type="holotype">Holotype</typeStatus>
</emphasis>
: 3 from Hacienda Taboga,
<taxonomicName id="4C02F399C061FFAB7E70FD433097FD19" box="[780,915,650,676]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="1" pageNumber="67" phylum="Arthropoda" rank="species" species="guanacaste">Guanacaste</taxonomicName>
Province,
<collectingCountry id="F315C88AC061FFAB796DFD433795FD19" box="[1041,1169,650,676]" name="Costa Rica" pageId="1" pageNumber="63">Costa Rica</collectingCountry>
(
<geoCoordinate id="EE36EEDDC061FFAB79DCFD43360AFD19" box="[1184,1294,650,676]" direction="north" orientation="latitude" pageId="1" pageNumber="63" precision="555" value="10.33">10.33° N</geoCoordinate>
,
<geoCoordinate id="EE36EEDDC061FFAB7866FD43368AFD19" box="[1306,1422,650,676]" direction="west" orientation="longitude" pageId="1" pageNumber="63" precision="555" value="-85.2">85.20° W</geoCoordinate>
), elevation 4 meters,
<date id="FFBCAEDAC061FFAB7CF6FD783146FD76" box="[394,578,689,715]" pageId="1" pageNumber="63" value="1967-08-02">2 August 1967</date>
, leg. D. R. Paulson (
<collectionCode id="ED1310DFC061FFAB7E3FFD783092FD76" box="[835,918,689,715]" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="1" pageNumber="63">FSCA</collectionCode>
).
<emphasis id="B9765408C061FFAB7ED7FD79371DFD77" bold="true" box="[939,1049,688,714]" pageId="1" pageNumber="63">Allotype</emphasis>
: Ƥ same data but
<date id="FFBCAEDAC061FFAB7981FD783699FD76" box="[1277,1437,689,715]" pageId="1" pageNumber="63" value="1966-07-14">14 July 1966</date>
(
<collectionCode id="ED1310DFC061FFAB7DE3FD1133EBFD4F" box="[159,239,728,754]" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="1" pageNumber="63">FSCA</collectionCode>
).
<emphasis id="B9765408C061FFAB7C7EFD1E327BFD4C" bold="true" box="[258,383,727,753]" pageId="1" pageNumber="63">
<typeStatus id="54B936B8C061FFAB7C7EFD1E327BFD4C" box="[258,383,727,753]" pageId="1" pageNumber="63" type="paratype">Paratypes</typeStatus>
</emphasis>
(28 3, 39 ƤƤ). Location as
<typeStatus id="54B936B8C061FFAB7F99FD11304FFD4F" box="[741,843,728,754]" pageId="1" pageNumber="63" type="holotype">holotype</typeStatus>
and allotype, from following dates:
<date id="FFBCAEDAC061FFAB7989FD113679FD4F" box="[1269,1405,728,754]" pageId="1" pageNumber="63" value="1966-07-14">14 Jul 1966</date>
(2 3, 5 ƤƤ),
<date id="FFBCAEDAC061FFAB7C52FD3732BCFCA5" box="[302,440,766,792]" pageId="1" pageNumber="63" value="1966-07-16">16 Jul 1966</date>
(
<date id="FFBCAEDAC061FFAB7CB5FD37313DFCA5" box="[457,569,766,792]" pageId="1" pageNumber="63" value="1915-08-03">8 3, 15</date>
ƤƤ),
<date id="FFBCAEDAC061FFAB7FFAFD373014FCA5" box="[646,784,766,792]" pageId="1" pageNumber="63" value="1966-07-24">24 Jul 1966</date>
(2 ƤƤ),
<date id="FFBCAEDAC061FFAB7E01FD37370DFCA5" box="[893,1033,766,792]" pageId="1" pageNumber="63" value="1966-08-09">9 Aug 1966</date>
(1 3, 2 ƤƤ),
<date id="FFBCAEDAC061FFAB79CEFD373649FCA5" box="[1202,1357,766,792]" pageId="1" pageNumber="63" value="1966-08-24">24 Aug 1966</date>
(1 Ƥ),
<date id="FFBCAEDAC061FFAB7DEBFCEC3234FC82" box="[151,304,805,831]" pageId="1" pageNumber="63" value="1966-08-26">26 Aug 1966</date>
(1 3),
<date id="FFBCAEDAC061FFAB7CFAFCEC311CFC82" box="[390,536,805,831]" pageId="1" pageNumber="63" value="1966-10-25">25 Oct 1966</date>
(2 3, 2 ƤƤ),
<date id="FFBCAEDAC061FFAB7FA8FCEC3062FC82" box="[724,870,805,831]" pageId="1" pageNumber="63" value="1966-10-26">26 Oct 1966</date>
(1 3),
<date id="FFBCAEDAC061FFAB7EC7FCEC3751FC82" box="[955,1109,805,831]" pageId="1" pageNumber="63" value="1966-11-17">17 Nov 1966</date>
(1 Ƥ),
<date id="FFBCAEDAC061FFAB79D7FCEC3630FC82" box="[1195,1332,805,831]" pageId="1" pageNumber="63" value="1967-07-14">14 Jul 1967</date>
(
<date id="FFBCAEDAC061FFAB7838FCEC33B2FCDB" pageId="1" pageNumber="63" value="1910-11-03">11 3, 10</date>
ƤƤ),
<date id="FFBCAEDAC061FFAB7C7AFC85329DFCDB" box="[262,409,844,870]" pageId="1" pageNumber="63" value="1967-08-02">2 Aug 1967</date>
(1 3, 1 ƤƤ),
<date id="FFBCAEDAC061FFAB7F37FC8531D9FCDB" box="[587,733,844,870]" pageId="1" pageNumber="63" value="1967-08-09">9 Aug 1967</date>
(1 3) (3 and Ƥ
<typeStatus id="54B936B8C061FFAB7ECCFC85371EFCDB" box="[944,1050,844,870]" pageId="1" pageNumber="63" type="paratype">paratype</typeStatus>
to each of
<collectionCode id="ED1310DFC061FFAB79DAFC8537F2FCDB" box="[1190,1270,844,870]" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="1" pageNumber="63">FSCA</collectionCode>
, InBIO,
<collectionCode id="ED1310DFC061FFAB781EFC85369CFCDB" box="[1378,1432,844,870]" pageId="1" pageNumber="63">KJT</collectionCode>
,
<collectionCode id="ED1310DFC061FFAB7DEBFCBB33F0FC31" box="[151,244,882,908]" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="1" pageNumber="63">NMNH</collectionCode>
,
<collectionCode id="ED1310DFC061FFAB7D83FCBB3245FC31" box="[255,321,882,908]" pageId="1" pageNumber="63">RWG</collectionCode>
,
<collectionCode id="ED1310DFC061FFAB7C31FCBB3297FC31" box="[333,403,882,908]" pageId="1" pageNumber="63">TWD</collectionCode>
, and
<collectionCode id="ED1310DFC061FFAB7CACFCBB312BFC31" box="[464,559,882,908]" httpUri="http://grbio.org/cool/ra64-3eva" name="University of Michigan, Museum of Zoology" pageId="1" pageNumber="63">UMMZ</collectionCode>
; remainder in DRP collection).
</paragraph>
</subSubSection>
<subSubSection id="C318DB91C061FFAE7DBAFC5132F0FDB3" lastPageId="4" lastPageNumber="66" pageId="1" pageNumber="63" type="description">
<paragraph id="8BBD881AC061FFAB7DBAFC513691FBBD" blockId="1.[151,1437,650,2030]" pageId="1" pageNumber="63">
<emphasis id="B9765408C061FFAB7DBAFC513253FC0F" bold="true" box="[198,343,920,946]" pageId="1" pageNumber="63">Description</emphasis>
.
<emphasis id="B9765408C061FFAB7C17FC513126FC0F" bold="true" box="[363,546,920,946]" pageId="1" pageNumber="63">
Male
<typeStatus id="54B936B8C061FFAB7CCAFC513126FC0F" box="[438,546,920,946]" pageId="1" pageNumber="63" type="holotype">holotype</typeStatus>
</emphasis>
.
<emphasis id="B9765408C061FFAB7F49FC51317CFC0F" bold="true" box="[565,632,920,946]" pageId="1" pageNumber="63">Head</emphasis>
. Glossy black above, including antennae; labrum very pale blue; postocular spots represented by narrow pale blue triangles extending forward along inner edge of each eye. Mouthparts cream-colored. Rear of head black in center, pale blue adjacent to eyes. Frons in profile rounded.
</paragraph>
<paragraph id="8BBD881AC061FFAB7DBAFBC43187FA3E" blockId="1.[151,1437,650,2030]" pageId="1" pageNumber="63">
<emphasis id="B9765408C061FFAB7DBAFBC43227FB9A" bold="true" box="[198,291,1037,1063]" pageId="1" pageNumber="63">Thorax</emphasis>
. Prothorax black above, pale orange on sides, orange spot in middle of anterior lobe. Posterior lobe rounded and projecting caudally. Mesostigmal plates small and mostly orange, more or less triangular and projecting upward from thorax (as in
<figureCitation id="1339949FC061FFAB7FEAFB9331DBFBC9" box="[662,735,1114,1140]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,276,1900,1924]" captionTargetBox="[222,1357,449,1899]" captionTargetId="figure@2.[217,1369,438,1974]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Leptobasis guanacaste. 1 – 4. S 10, male paratype. (1) lateral view; (2) mediodorsal view; (3) medioventral view; (4) posterior view. 5 – 6. Genital ligula, male paratype. (5) lateral view; (6) entolateral view. 7. Metafemur, male paratype, view of flexor surface. 8 – 9. Posterior lobe of pronotum and mesostigmal plates, dorsal view. (8) male paratype; (9) female allotype. 10. S 8 - 10, female allotype, lateral view." httpUri="https://zenodo.org/record/190502/files/figure.png" pageId="1" pageNumber="63">Fig. 8</figureCitation>
). Pterothorax orange, paler below and with black median stripe and narrow black humeral stripes with irregular median borders. Legs same shade of orange, tinged brown at distal ends of segments; spines black. Spurs on metafemur distinctive, one longer and appressed distally at 1/3 length, then row of 7 (left) or 6 (right) much smaller and slightly curved spines at 5/8–7/8 length (as in
<figureCitation id="1339949FC061FFAB7D99FB3C3235FAB2" box="[229,305,1269,1295]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,276,1900,1924]" captionTargetBox="[222,1357,449,1899]" captionTargetId="figure@2.[217,1369,438,1974]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Leptobasis guanacaste. 1 – 4. S 10, male paratype. (1) lateral view; (2) mediodorsal view; (3) medioventral view; (4) posterior view. 5 – 6. Genital ligula, male paratype. (5) lateral view; (6) entolateral view. 7. Metafemur, male paratype, view of flexor surface. 8 – 9. Posterior lobe of pronotum and mesostigmal plates, dorsal view. (8) male paratype; (9) female allotype. 10. S 8 - 10, female allotype, lateral view." httpUri="https://zenodo.org/record/190502/files/figure.png" pageId="1" pageNumber="63">Fig. 7</figureCitation>
); metatibial spurs considerably shorter than intervening spaces; tarsal claws with vestigial supplementary tooth. Wings hyaline with brown veins; pt brown, a parallelogram with all sides subequal, covering slightly less than one cell (as in
<figureCitation id="1339949FC061FFAB7FF7FA8B31E6FAE1" box="[651,738,1346,1372]" captionStart="FIGURE 11" captionStartId="3.[151,254,836,860]" captionTargetBox="[240,1339,241,782]" captionTargetId="figure@3.[218,1369,197,811]" captionTargetPageId="3" captionText="FIGURE 11. Wings, male paratype of Leptobasis guanacaste." httpUri="https://zenodo.org/record/190503/files/figure.png" pageId="1" pageNumber="63">Fig. 11</figureCitation>
). Px
<quantity id="4CFA25FFC061FFAB7E5DFA8B3054FAE1" box="[801,848,1346,1372]" metricMagnitude="-1" metricUnit="m" metricValue="1.778" pageId="1" pageNumber="63" unit="in" value="7.0">7 in</quantity>
right FW,
<quantity id="4CFA25FFC061FFAB7EAEFA8B3705FAE1" box="[978,1025,1346,1372]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="1" pageNumber="63" unit="in" value="8.0">8 in</quantity>
other three wings. CuA extends 6 cells beyond quadrangle in all four wings.
</paragraph>
<paragraph id="8BBD881AC061FFAB7DBAFA46322BF92F" blockId="1.[151,1437,650,2030]" pageId="1" pageNumber="63">
<emphasis id="B9765408C061FFAB7DBAFA463244FA14" bold="true" box="[198,320,1423,1449]" pageId="1" pageNumber="63">Abdomen</emphasis>
. S1-2 light orange, S3-7 black above and orange below, S7 with very narrow orange apical ring; S8-10 and appendages orange. Cerci in lateral view slightly more than one-third length of S10, quadrate, in posterior view a small black cornified spine on ventral surface projecting ventrally and laterally. Paraprocts two-thirds length of cerci, broad at base but narrowing abruptly to a short dorsolateral tubercle with pointed, cornified spine at end. Appendages of
<typeStatus id="54B936B8C061FFAB7F10F9E331D0F9F9" box="[620,724,1578,1604]" pageId="1" pageNumber="63" type="paratype">paratype</typeStatus>
in
<figureCitation id="1339949FC061FFAB7F80F9E33075F9F9" box="[764,881,1578,1604]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,276,1900,1924]" captionTargetBox="[222,1357,449,1899]" captionTargetId="figure@2.[217,1369,438,1974]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Leptobasis guanacaste. 1 – 4. S 10, male paratype. (1) lateral view; (2) mediodorsal view; (3) medioventral view; (4) posterior view. 5 – 6. Genital ligula, male paratype. (5) lateral view; (6) entolateral view. 7. Metafemur, male paratype, view of flexor surface. 8 – 9. Posterior lobe of pronotum and mesostigmal plates, dorsal view. (8) male paratype; (9) female allotype. 10. S 8 - 10, female allotype, lateral view." httpUri="https://zenodo.org/record/190502/files/figure.png" pageId="1" pageNumber="63">Figs. 1–4</figureCitation>
. Genital ligula with two long basally directed slightly curved chitinized filaments on internal surface of terminal segment (as in
<figureCitation id="1339949FC061FFAB79F6F99837FAF9D6" box="[1162,1278,1617,1643]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,276,1900,1924]" captionTargetBox="[222,1357,449,1899]" captionTargetId="figure@2.[217,1369,438,1974]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Leptobasis guanacaste. 1 – 4. S 10, male paratype. (1) lateral view; (2) mediodorsal view; (3) medioventral view; (4) posterior view. 5 – 6. Genital ligula, male paratype. (5) lateral view; (6) entolateral view. 7. Metafemur, male paratype, view of flexor surface. 8 – 9. Posterior lobe of pronotum and mesostigmal plates, dorsal view. (8) male paratype; (9) female allotype. 10. S 8 - 10, female allotype, lateral view." httpUri="https://zenodo.org/record/190502/files/figure.png" pageId="1" pageNumber="63">Figs. 5–6</figureCitation>
) but with no internal fold.
</paragraph>
<paragraph id="8BBD881AC061FFAB7DBAF95737FDF905" blockId="1.[151,1437,650,2030]" box="[198,1273,1694,1720]" pageId="1" pageNumber="63">
<emphasis id="B9765408C061FFAB7DBAF9573279F905" bold="true" box="[198,381,1694,1720]" pageId="1" pageNumber="63">Measurements</emphasis>
. Total length
<quantity id="4CFA25FFC061FFAB7F5CF9573171F905" box="[544,629,1694,1720]" metricMagnitude="-2" metricUnit="m" metricValue="3.2" pageId="1" pageNumber="63" unit="mm" value="32.0">32 mm</quantity>
; abdomen length
<quantity id="4CFA25FFC061FFAB7E38F957309EF905" box="[836,922,1694,1720]" metricMagnitude="-2" metricUnit="m" metricValue="2.7" pageId="1" pageNumber="63" unit="mm" value="27.0">27 mm</quantity>
; FW
<quantity id="4CFA25FFC061FFAB7EA5F9573741F905" box="[985,1093,1694,1720]" metricMagnitude="-2" metricUnit="m" metricValue="1.58" pageId="1" pageNumber="63" unit="mm" value="15.8">15.8 mm</quantity>
; HW 15.0 mm.
</paragraph>
<paragraph id="8BBD881AC061FFAB7DBAF90C316FF962" blockId="1.[151,1437,650,2030]" box="[198,619,1733,1759]" pageId="1" pageNumber="63">
<emphasis id="B9765408C061FFAB7DBAF90C3288F962" bold="true" box="[198,396,1733,1759]" pageId="1" pageNumber="63">Female allotype</emphasis>
(not fully mature).
</paragraph>
<paragraph id="8BBD881AC061FFAB7DBAF92230D7F8BB" blockId="1.[151,1437,650,2030]" box="[198,979,1771,1798]" pageId="1" pageNumber="63">
<emphasis id="B9765408C061FFAB7DBAF922320DF8B8" bold="true" box="[198,265,1771,1797]" pageId="1" pageNumber="63">Head</emphasis>
. Colored exactly as male, but duller, brown instead of black.
</paragraph>
<paragraph id="8BBD881AC061FFA87DBAF8DB31F6FF42" blockId="1.[151,1437,650,2030]" lastBlockId="2.[151,1436,152,448]" lastPageId="2" lastPageNumber="64" pageId="1" pageNumber="63">
<emphasis id="B9765408C061FFAB7DBAF8DB3227F891" bold="true" box="[198,291,1810,1836]" pageId="1" pageNumber="63">Thorax</emphasis>
. Anterior lobe of prothorax pale orange, brown laterally; median lobe orange-brown dorsally, shading to pale orange laterally; posterior lobe darker brown, posterior margin very slightly concave rather than convex as in male. Pterothorax pale orange, with brown median stripe and only hint of brownish humeral stripe along mesopleural suture. Mesostigmal plates pale orange, rounded and projecting upward from thorax (
<figureCitation id="1339949FC061FFAB7DDCF86433EFF87A" box="[160,235,1965,1991]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,276,1900,1924]" captionTargetBox="[222,1357,449,1899]" captionTargetId="figure@2.[217,1369,438,1974]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Leptobasis guanacaste. 1 – 4. S 10, male paratype. (1) lateral view; (2) mediodorsal view; (3) medioventral view; (4) posterior view. 5 – 6. Genital ligula, male paratype. (5) lateral view; (6) entolateral view. 7. Metafemur, male paratype, view of flexor surface. 8 – 9. Posterior lobe of pronotum and mesostigmal plates, dorsal view. (8) male paratype; (9) female allotype. 10. S 8 - 10, female allotype, lateral view." httpUri="https://zenodo.org/record/190502/files/figure.png" pageId="1" pageNumber="63">Fig. 9</figureCitation>
). Legs pale orange with slightly indicated darker bands. Spurs on metafemur not as modified as in male; proximalmost spine at midlength, moderately long, then followed immediately by 5 (left) or 4 (right) spines, distalmost slightly longest; metatibial spurs about same length as intervening spaces (notably different from male). Wings as male, but pt pale tan; 9 Px in both FW, 7 Px in both HW. CuA extends 5 cells beyond quadrangle in both FW, 6 cells beyond in both HW.
</paragraph>
<paragraph id="8BBD881AC062FFA87DBAFEC232C4FE27" blockId="2.[151,1436,152,448]" pageId="2" pageNumber="64">
<emphasis id="B9765408C062FFA87DBAFEC23244FE98" bold="true" box="[198,320,267,293]" pageId="2" pageNumber="64">Abdomen</emphasis>
. S1-2 orange with brown at posterior margin. S3-7 black above, orange below; proximal 1/6 of S3 orange; S8-10 orange, somewhat obscured with brown wash laterally. No vulvar spine on S8. Ovipositor valves curved, extending slightly beyond cerci on S10 (
<figureCitation id="1339949FC062FFA87E5AFE90307AFECE" box="[806,894,345,371]" captionStart="FIGURES 1 – 10" captionStartId="2.[151,276,1900,1924]" captionTargetBox="[222,1357,449,1899]" captionTargetId="figure@2.[217,1369,438,1974]" captionTargetPageId="2" captionText="FIGURES 1 – 10. Leptobasis guanacaste. 1 – 4. S 10, male paratype. (1) lateral view; (2) mediodorsal view; (3) medioventral view; (4) posterior view. 5 – 6. Genital ligula, male paratype. (5) lateral view; (6) entolateral view. 7. Metafemur, male paratype, view of flexor surface. 8 – 9. Posterior lobe of pronotum and mesostigmal plates, dorsal view. (8) male paratype; (9) female allotype. 10. S 8 - 10, female allotype, lateral view." httpUri="https://zenodo.org/record/190502/files/figure.png" pageId="2" pageNumber="64">Fig. 10</figureCitation>
). Outer valve of ovipositor with single row of about 23 tiny black teeth.
</paragraph>
<paragraph id="8BBD881AC062FFA87DBAFE6F360BFE7D" blockId="2.[151,1436,152,448]" box="[198,1295,422,448]" pageId="2" pageNumber="64">
<emphasis id="B9765408C062FFA87DBAFE6F3279FE7D" bold="true" box="[198,381,422,448]" pageId="2" pageNumber="64">Measurements</emphasis>
. Total length
<quantity id="4CFA25FFC062FFA87F5CFE6F318FFE7D" box="[544,651,422,448]" metricMagnitude="-2" metricUnit="m" metricValue="3.05" pageId="2" pageNumber="64" unit="mm" value="30.5">30.5 mm</quantity>
; abdomen length
<quantity id="4CFA25FFC062FFA87E26FE6F30B4FE7D" box="[858,944,422,448]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" pageId="2" pageNumber="64" unit="mm" value="25.0">25 mm</quantity>
; FW 18.0 mm; HW 17.0 mm.
</paragraph>
<caption id="DF7DD892C062FFA87DEBF8A530E7F851" httpUri="https://zenodo.org/record/190502/files/figure.png" pageId="2" pageNumber="64" targetBox="[222,1357,449,1899]" targetPageId="2">
<paragraph id="8BBD881AC062FFA87DEBF8A530E7F851" blockId="2.[151,1437,1900,2029]" pageId="2" pageNumber="64">
<emphasis id="B9765408C062FFA87DEBF8A5325AF839" bold="true" box="[151,350,1900,1924]" pageId="2" pageNumber="64">FIGURES 1–10.</emphasis>
<emphasis id="B9765408C062FFA87C1AF8A4317CF839" box="[358,632,1901,1924]" italics="true" pageId="2" pageNumber="64">
<taxonomicName id="4C02F399C062FFA87C1AF8A43170F839" box="[358,628,1901,1924]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="2" pageNumber="64" phylum="Arthropoda" rank="species" species="guanacaste">Leptobasis guanacaste</taxonomicName>
.
</emphasis>
<emphasis id="B9765408C062FFA87F03F8A531B6F839" bold="true" box="[639,690,1900,1924]" pageId="2" pageNumber="64">1–4.</emphasis>
S10, male paratype. (1) lateral view; (2) mediodorsal view; (3) medioventral view; (4) posterior view.
<emphasis id="B9765408C062FFA87F2CF8593185F815" bold="true" box="[592,641,1936,1960]" pageId="2" pageNumber="64">5–6.</emphasis>
Genital ligula, male paratype. (5) lateral view; (6) entolateral view.
<emphasis id="B9765408C062FFA878FBF8593699F815" bold="true" box="[1415,1437,1936,1960]" pageId="2" pageNumber="64">7.</emphasis>
Metafemur, male paratype, view of flexor surface.
<emphasis id="B9765408C062FFA87FCBF87A31E1F876" bold="true" box="[695,741,1971,1995]" pageId="2" pageNumber="64">8–9.</emphasis>
Posterior lobe of pronotum and mesostigmal plates, dorsal view. (8) male paratype; (9) female allotype.
<emphasis id="B9765408C062FFA87F45F81C315EF850" bold="true" box="[569,602,2005,2029]" pageId="2" pageNumber="64">10.</emphasis>
S8-10, female allotype, lateral view.
</paragraph>
</caption>
<caption id="DF7DD892C063FFA97DEBFC8D3035FCE6" box="[151,817,836,860]" httpUri="https://zenodo.org/record/190503/files/figure.png" pageId="3" pageNumber="65" targetBox="[240,1339,241,782]" targetPageId="3">
<paragraph id="8BBD881AC063FFA97DEBFC8D3035FCE6" blockId="3.[151,817,836,860]" box="[151,817,836,860]" pageId="3" pageNumber="65">
<emphasis id="B9765408C063FFA97DEBFC8D3220FCE1" bold="true" box="[151,292,836,860]" pageId="3" pageNumber="65">FIGURE 11.</emphasis>
Wings, male paratype of
<taxonomicName id="4C02F399C063FFA97F4BFC8C302EFCE1" box="[567,810,837,860]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="65" phylum="Arthropoda" rank="species" species="guanacaste">
<emphasis id="B9765408C063FFA97F4BFC8C302EFCE1" box="[567,810,837,860]" italics="true" pageId="3" pageNumber="65">Leptobasis guanacaste</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<caption id="DF7DD892C063FFA97DEBF9B932CAF917" httpUri="https://zenodo.org/record/190504/files/figure.png" pageId="3" pageNumber="65" targetBox="[217,1369,914,1624]" targetPageId="3">
<paragraph id="8BBD881AC063FFA97DEBF9B932CAF917" blockId="3.[151,1436,1648,1706]" pageId="3" pageNumber="65">
<emphasis id="B9765408C063FFA97DEBF9B9322CF935" bold="true" box="[151,296,1648,1672]" pageId="3" pageNumber="65">FIGURE 12.</emphasis>
Male (above) and female (below) of
<taxonomicName id="4C02F399C063FFA97FBFF9B830BFF935" box="[707,955,1649,1672]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="65" phylum="Arthropoda" rank="species" species="guanacaste">
<emphasis id="B9765408C063FFA97FBFF9B830BFF935" box="[707,955,1649,1672]" italics="true" pageId="3" pageNumber="65">Leptobasis guanacaste</emphasis>
</taxonomicName>
, at Hacienda Taboga,
<taxonomicName id="4C02F399C063FFA979CDF9B9362AF93A" box="[1201,1326,1648,1671]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="3" pageNumber="67" phylum="Arthropoda" rank="species" species="guanacaste">Guanacaste</taxonomicName>
Province, Costa Rica (photos by DRP).
</paragraph>
</caption>
<paragraph id="8BBD881AC063FFA97DBAF9153039F8D6" blockId="3.[151,1436,1756,2014]" pageId="3" pageNumber="65">
<emphasis id="B9765408C063FFA97DBAF91532E2F94B" bold="true" box="[198,486,1756,1782]" pageId="3" pageNumber="65">
Variation in
<typeStatus id="54B936B8C063FFA97C16F91532E2F94B" box="[362,486,1756,1782]" pageId="3" pageNumber="65" type="paratype">paratypes</typeStatus>
</emphasis>
. Mature females have top of head and front of thorax glossy black, with blue labrum, postocular spots, and narrow antehumeral stripe. Abdomen entirely black above, pale orange-brown below, paler color extending higher on sides of S8-10. Metatibial spurs variable in length and distance apart in both sexes. See Coloration in Life below for more details.
</paragraph>
<paragraph id="8BBD881AC063FFAE7DBAF8BE3626FF0F" blockId="3.[151,1436,1756,2014]" lastBlockId="4.[151,1437,152,874]" lastPageId="4" lastPageNumber="66" pageId="3" pageNumber="65">
<emphasis id="B9765408C063FFA97DBAF8BE3279F82C" bold="true" box="[198,381,1911,1937]" pageId="3" pageNumber="65">Measurements</emphasis>
. Ten
<typeStatus id="54B936B8C063FFA97CBAF8BE312FF82C" box="[454,555,1911,1937]" pageId="3" pageNumber="65" type="paratype">paratype</typeStatus>
3 total length 31.5–35.0 mm (mean
<quantity id="4CFA25FFC063FFA97975F8BE377DF82C" box="[1033,1145,1911,1937]" metricMagnitude="-2" metricUnit="m" metricValue="3.32" pageId="3" pageNumber="65" unit="mm" value="33.2">33.2 mm</quantity>
), abdomen length
<quantity id="4CFA25FFC063FFA97826F8BE33FBF805" metricMagnitude="-2" metricUnit="m" metricValue="2.81" metricValueMax="2.95" metricValueMin="2.67" pageId="3" pageNumber="65" unit="mm" value="28.1" valueMax="29.5" valueMin="26.7">26.7– 29.5 mm</quantity>
(mean
<quantity id="4CFA25FFC063FFA97C2BF85732C7F805" box="[343,451,1950,1976]" metricMagnitude="-2" metricUnit="m" metricValue="2.82" pageId="3" pageNumber="65" unit="mm" value="28.2">28.2 mm</quantity>
), FW length 15.8–17.0 mm (mean
<quantity id="4CFA25FFC063FFA97E21F85730CEF805" box="[861,970,1950,1976]" metricMagnitude="-2" metricUnit="m" metricValue="1.63" pageId="3" pageNumber="65" unit="mm" value="16.3">16.3 mm</quantity>
), HW length 14.7–16.0 mm (mean
<quantity id="4CFA25FFC063FFA97815F85733CEF863" metricMagnitude="-2" metricUnit="m" metricValue="1.55" pageId="3" pageNumber="65" unit="mm" value="15.5">15.5 mm</quantity>
). Ten
<typeStatus id="54B936B8C063FFA97C6FF80D327EF863" box="[275,378,1988,2014]" pageId="3" pageNumber="65" type="paratype">paratype</typeStatus>
ƤƤ total length 28.0–
<quantity id="4CFA25FFC063FFA97FF7F80D31F1F863" box="[651,757,1988,2014]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" pageId="3" pageNumber="65" unit="mm" value="32.5">32.5 mm</quantity>
(mean
<quantity id="4CFA25FFC063FFA97E2CF80D30BBF863" box="[848,959,1988,2014]" metricMagnitude="-2" metricUnit="m" metricValue="3.02" pageId="3" pageNumber="65" unit="mm" value="30.2">30.2 mm</quantity>
), abdomen length 23.8–27.0 mm (mean
<quantity id="4CFA25FFC064FFAE7DEBFF513207FF0F" box="[151,259,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="2.4899999999999998" pageId="4" pageNumber="66" unit="mm" value="24.9">24.9 mm</quantity>
), FW length 17.3–19.0 mm (mean
<quantity id="4CFA25FFC064FFAE7FE1FF51300EFF0F" box="[669,778,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="1.7899999999999998" pageId="4" pageNumber="66" unit="mm" value="17.9">17.9 mm</quantity>
), HW length
<quantity id="4CFA25FFC064FFAE7EDAFF513755FF0F" box="[934,1105,152,178]" metricMagnitude="-2" metricUnit="m" metricValue="1.73" metricValueMax="1.83" metricValueMin="1.63" pageId="4" pageNumber="66" unit="mm" value="17.3" valueMax="18.3" valueMin="16.3">16.3–18.3 mm</quantity>
(mean 17.0 mm).
</paragraph>
<paragraph id="8BBD881AC064FFAE7DBAFF7732F0FDB3" blockId="4.[151,1437,152,874]" pageId="4" pageNumber="66">
<emphasis id="B9765408C064FFAE7DBAFF7732A2FF65" bold="true" box="[198,422,190,216]" pageId="4" pageNumber="66">Coloration in life</emphasis>
. (
<figureCitation id="1339949FC064FFAE7CBFFF773124FF65" box="[451,544,190,216]" captionStart="FIGURE 12" captionStartId="3.[151,255,1648,1672]" captionTargetBox="[217,1369,914,1624]" captionTargetId="figure@3.[217,1369,914,1624]" captionTargetPageId="3" captionText="FIGURE 12. Male (above) and female (below) of Leptobasis guanacaste, at Hacienda Taboga, Guanacaste Province, Costa Rica (photos by DRP)." httpUri="https://zenodo.org/record/190504/files/figure.png" pageId="4" pageNumber="66">Fig. 12</figureCitation>
). Mature male eyes black on upper third, greenish on lower two-thirds; postocular spots blue-green; prothoracic spot, legs, and face yellow. Thorax, S1-2, and S8-10 orange. Mature female quite unlike male except for similar eye color. Head black, face and postocular spots bluish; thorax black above, greenish on sides. Abdomen black with some dull orange markings ventrolaterally on S8-10. Substantial color changes with age, as in other
<taxonomicName id="4C02F399C064FFAE7FBDFE903044FECF" box="[705,832,345,370]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B9765408C064FFAE7FBDFE903044FECF" box="[705,832,345,370]" italics="true" pageId="4" pageNumber="66">Leptobasis</emphasis>
</taxonomicName>
species. Post-tenerals of both sexes mostly orange, with S4-7 quickly becoming blackish; looking much like
<taxonomicName id="4C02F399C064FFAE7E15FE4930F8FE24" box="[873,1020,384,409]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="vacillans">
<emphasis id="B9765408C064FFAE7E15FE4930F8FE24" box="[873,1020,384,409]" italics="true" pageId="4" pageNumber="66">L. vacillans</emphasis>
</taxonomicName>
at this stage. Apparently quickly develop to immature stage, in which female colored somewhat like mature male, but face and postocular spots orange; thorax orange with black median stripe, legs orange; abdomen black, with orange markings low on S8-9 and S10 entirely orange.
</paragraph>
</subSubSection>
<subSubSection id="C318DB91C064FFAE7DBAFDD3361CFCD7" pageId="4" pageNumber="66" type="diagnosis">
<paragraph id="8BBD881AC064FFAE7DBAFDD3361CFCD7" blockId="4.[151,1437,152,874]" pageId="4" pageNumber="66">
<emphasis id="B9765408C064FFAE7DBAFDD33242FD89" bold="true" box="[198,326,538,564]" pageId="4" pageNumber="66">Diagnosis.</emphasis>
<taxonomicName id="4C02F399C064FFAE7C32FDD3315DFD8E" box="[334,601,538,563]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="guanacaste">
<emphasis id="B9765408C064FFAE7C32FDD3315DFD8E" box="[334,601,538,563]" italics="true" pageId="4" pageNumber="66">Leptobasis guanacaste</emphasis>
</taxonomicName>
is easily distinguished from all other named species of its genus by its color pattern; no other coenagrionid in its range looks like the mature male. It is the only
<taxonomicName id="4C02F399C064FFAE79CCFD88362BFDE7" box="[1200,1327,577,602]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B9765408C064FFAE79CCFD88362BFDE7" box="[1200,1327,577,602]" italics="true" pageId="4" pageNumber="66">Leptobasis</emphasis>
</taxonomicName>
in which mature individuals lack the typical coenagrionid pattern of median and humeral thoracic stripes. The orange thorax with black median stripe and black abdomen with orange tip are found in males of no other
<taxonomicName id="4C02F399C064FFAE7869FD463690FD15" box="[1301,1428,655,680]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B9765408C064FFAE7869FD463690FD15" box="[1301,1428,655,680]" italics="true" pageId="4" pageNumber="66">Leptobasis</emphasis>
</taxonomicName>
. Some other
<taxonomicName id="4C02F399C064FFAE7C5BFD7C32A3FD73" box="[295,423,693,718]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="66" phylum="Arthropoda" rank="genus">
<emphasis id="B9765408C064FFAE7C5BFD7C32A3FD73" box="[295,423,693,718]" italics="true" pageId="4" pageNumber="66">Leptobasis</emphasis>
</taxonomicName>
have orange immatures, but they lack the black median thoracic stripe that develops during maturation in
<taxonomicName id="4C02F399C064FFAE7CE2FD15314FFD48" box="[414,587,732,757]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="guanacaste">
<emphasis id="B9765408C064FFAE7CE2FD15314FFD48" box="[414,587,732,757]" italics="true" pageId="4" pageNumber="66">L. guanacaste</emphasis>
</taxonomicName>
. In addition, male
<taxonomicName id="4C02F399C064FFAE7E44FD1530E2FD48" box="[824,998,732,757]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="guanacaste">
<emphasis id="B9765408C064FFAE7E44FD1530E2FD48" box="[824,998,732,757]" italics="true" pageId="4" pageNumber="66">L. guanacaste</emphasis>
</taxonomicName>
has a unique combination of short, angled, and spined cerci and even shorter paraprocts. It shares with
<taxonomicName id="4C02F399C064FFAE7EAEFCCB3771FCA6" box="[978,1141,770,795]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="candelaria">
<emphasis id="B9765408C064FFAE7EAEFCCB3771FCA6" box="[978,1141,770,795]" italics="true" pageId="4" pageNumber="66">L. candelaria</emphasis>
</taxonomicName>
distinctive metafemoral spines and genital ligula and a relatively short ovipositor. In addition, female
<taxonomicName id="4C02F399C064FFAE7947FCE037E6FCFF" box="[1083,1250,809,834]" class="Insecta" family="Coenagrionidae" genus="Leptobasis" kingdom="Animalia" order="Odonata" pageId="4" pageNumber="66" phylum="Arthropoda" rank="species" species="guanacaste">
<emphasis id="B9765408C064FFAE7947FCE037E6FCFF" box="[1083,1250,809,834]" italics="true" pageId="4" pageNumber="66">L. guanacaste</emphasis>
</taxonomicName>
is unique in its genus in the slightly concave posterior border of the prothorax; this is concave in all other species.
</paragraph>
</subSubSection>
</treatment>
</document>
Reference in a new issue View git blame Copy permalink