172 lines
24 KiB
XML
172 lines
24 KiB
XML
<document id="4A0819FCC5E70994EBC8B13B5C986974" ID-DOI="10.11646/zootaxa.4039.4.5" ID-GBIF-Dataset="5d48e0d6-4af9-422e-af66-81503dd42524" ID-ISSN="1175-5326" ID-Zenodo-Dep="241183" ID-ZooBank="C6FD1263-3CAB-47F3-8661-700612D0B59A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461333100469" checkinUser="plazi" docAuthor="Huang, Hao, Bartolozzi, Luca & Chen, Chang-Chin" docDate="2015" docId="0393D10F5A61755BD39FABB3F0F4FD45" docLanguage="en" docName="zt04039p565.pdf" docOrigin="Zootaxa 4039 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Echinoaesalus sabahensis Zelenka 1994" docType="treatment" docVersion="9" lastPageNumber="563" masterDocId="FFAAA9775A687551D308AE67F265FC59" masterDocTitle="Notes on the genus Echinoaesalus Zelenka, 1993 (Coleoptera: Lucanidae), with the description of a new species from the Malay Peninsula" masterLastPageNumber="565" masterPageNumber="553" pageNumber="562" updateTime="1698618809578" updateUser="plazi">
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<mods:title id="B78E607197BB57F588AAA6B8972529FE">Notes on the genus Echinoaesalus Zelenka, 1993 (Coleoptera: Lucanidae), with the description of a new species from the Malay Peninsula</mods:title>
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<mods:namePart id="7C24A9A5CD02E8D60FCAE088DC828206">Huang, Hao</mods:namePart>
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<mods:namePart id="9548ACA4997F36A248EB280F661B1F20">Bartolozzi, Luca</mods:namePart>
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<mods:namePart id="1E89CCBF9491A445D38F00844EA693AF">Chen, Chang-Chin</mods:namePart>
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<treatment id="0393D10F5A61755BD39FABB3F0F4FD45" ID-DOI="http://doi.org/10.5281/zenodo.6097388" ID-GBIF-Taxon="119644249" ID-Zenodo-Dep="6097388" LSID="urn:lsid:plazi:treatment:0393D10F5A61755BD39FABB3F0F4FD45" httpUri="http://treatment.plazi.org/id/0393D10F5A61755BD39FABB3F0F4FD45" lastPageId="10" lastPageNumber="563" pageId="9" pageNumber="562">
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<subSubSection id="C32033925A617558D39FABB3F079FA56" pageId="9" pageNumber="562" type="nomenclature">
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<paragraph id="8B8560195A617558D39FABB3F0ECF9B7" blockId="9.[151,649,1492,1552]" box="[151,649,1492,1518]" pageId="9" pageNumber="562">
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<heading id="D0CDD7755A617558D39FABB3F0ECF9B7" bold="true" box="[151,649,1492,1518]" fontSize="11" level="1" pageId="9" pageNumber="562" reason="1">
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<taxonomicName id="4C3A1B9A5A617558D39FABB3F0ECF9B7" ID-CoL="38G53" authority="Zelenka, 1994" authorityName="Zelenka" authorityYear="1994" box="[151,649,1492,1518]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="562" phylum="Arthropoda" rank="species" species="sabahensis">
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<emphasis id="B94EBC0B5A617558D39FABB3F0ECF9B7" bold="true" box="[151,649,1492,1518]" pageId="9" pageNumber="562">
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<emphasis id="B94EBC0B5A617558D39FABB3F3B1F9B7" bold="true" box="[151,468,1492,1518]" italics="true" pageId="9" pageNumber="562">Echinoaesalus sabahensis</emphasis>
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<bibRefCitation id="EFAB1DE85A617558D2D3ABB3F0ECF9B7" author="Zelenka" box="[475,649,1492,1518]" pageId="9" pageNumber="562" refString="Zelenka, W. (1994) Zwei neue Echinoaesalus - Arten aus Sudostasien (Coleoptera, Lucanidae). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 46, 56 - 61." type="journal article" year="1994">Zelenka, 1994</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8B8560195A617558D39FAB90F079FA56" blockId="9.[151,649,1492,1552]" box="[151,540,1527,1552]" pageId="9" pageNumber="562">
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(
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<figureCitation id="13017C9C5A617558D397AB90F28BFA56" box="[159,238,1527,1552]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1750,1773]" captionTargetBox="[183,1395,205,1699]" captionTargetId="figure@2.[183,1395,193,1713]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Antenna. 1, Echinoaesalus hidakai, L 12, western Malaysia; 2, E. borneoensis, L 11, Borneo; 3, E. timidus, L 19, western Malaysia; 4, E. sabahensis, L 14, Borneo; 5, E. gedeensis, holotype, Java; 6, E. cechovskyi, holotype, western Malaysia; 7, E. cechovskyi, paratype, Singapore. Arrow directed to end of third antennomere." httpUri="https://zenodo.org/record/241184/files/figure.png" pageId="9" pageNumber="562">Figs. 4</figureCitation>
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,
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<figureCitation id="13017C9C5A617558D3F2AB90F326FA56" box="[250,323,1527,1551]" captionStart="FIGURES 8 – 30" captionStartId="2.[151,264,1842,1865]" captionText="FIGURES 8 – 30. Mentum under same scale (scale bar 0.5 mm). 8, Echinoaesalus hidakai, L 12, western Malaysia; 9, E. borneoensis, L 01, Borneo; 10, E. hidakai, L 10, western Malaysia; 11, E. borneoensis, L 07, Borneo; 12, E. sabahensis, after Krikken (2008), Borneo; 13, E. sabahensis, L 14, Borneo; 14, E. cechovskyi, holotype, western Malaysia; 15, E. cechovskyi, paratype, Singapore; 16 – 18, E. timidus; 16, L 21; 17, L 22; 18, L 18; 19, E. javanus, after Krikken (2008), holotype, Java; 20, E. gedeensis, holotype, Java; 21, E. timidus, after Krikken (2008), holotype, probably male; 22 – 30, E. timidus; 22, L 08; 23, L 02; 24, L 17; 25, L 23; 26, L 09; 27, L 06; 28, L 20; 29, L 13; 30, L 29." pageId="9" pageNumber="562">12–13</figureCitation>
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,
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<figureCitation id="13017C9C5A617558D246AB90F309FA56" box="[334,364,1527,1551]" captionStart="FIGURES 35 – 52" captionStartId="3.[151,264,1853,1876]" captionTargetBox="[202,1395,193,1742]" captionTargetId="figure@3.[202,1396,193,1744]" captionTargetPageId="3" captionText="FIGURES 35 – 52. Protibia (left), mesotibia (middle), and metatibia (right), or just protibia. 35, Echinoaesalus hidakai, L 12; 36, E. hidakai, L 10; 37, E. borneoensis, L 11; 38, E. borneoensis, L 07; 39 – 48, E. timidus; 39, L 17; 40, L 13; 41, L 08; 42, L 02; 43, L 23; 44, L 21; 45, L 22; 46, L 18; 47, L 09; 48, L 06; 49, E. sabahensis, L 14; 50, E. cechovskyi, holotype; 51, E. cechovskyi, paratype; 52, E. gedeensis, holotype." httpUri="https://zenodo.org/record/241185/files/figure.png" pageId="9" pageNumber="562">49</figureCitation>
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,
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<figureCitation id="13017C9C5A617558D270AB90F3A4FA56" box="[376,449,1527,1552]" captionStart="FIGURES 57 – 76" captionStartId="4.[151,264,1567,1590]" captionTargetBox="[151,1436,203,1545]" captionTargetId="figure@4.[151,1436,193,1546]" captionTargetPageId="4" captionText="FIGURES 57 – 76. Prosternal process in ventral view. 57, Echinoaesalus hidakai, L 12; 58, E. hidakai, L 10; 59 – 62, E. borneoensis; 59, L 11; 60, L 01; 61, L 07; 62, L 16; 63, E. gedeensis, holotype; 64 – 67, E. timidus; 64, L 08; 65, L 02; 66, L 17; 67, L 21; 68, E. sabahensis, L 14; 69, E. cechovskyi, L 28; 70 – 73, E. timidus; 70, L 09; 71, L 06; 72, L 20; 73, L 13; 74, E. sabahensis, after Krikken 2008; 75, E. cechovskyi, paratype; 76, E. javanus, after Krikken (2008), holotype." httpUri="https://zenodo.org/record/241186/files/figure.png" pageId="9" pageNumber="562">68, 74</figureCitation>
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, 87, 99)
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</paragraph>
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</subSubSection>
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<subSubSection id="C32033925A617558D39FA858F74DFA01" box="[151,1320,1599,1624]" pageId="9" pageNumber="562" type="materials_examined">
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<paragraph id="8B8560195A617558D39FA858F74DFA01" blockId="9.[151,1436,1599,2022]" box="[151,1320,1599,1624]" pageId="9" pageNumber="562">
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<emphasis id="B94EBC0B5A617558D39FA858F29AFA01" bold="true" box="[151,255,1599,1624]" pageId="9" pageNumber="562">Material</emphasis>
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.
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<emphasis id="B94EBC0B5A617558D205A858F3C6FA01" bold="true" box="[269,419,1599,1624]" pageId="9" pageNumber="562">
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<collectingCountry id="F32D20895A617558D205A858F3C6FA01" box="[269,419,1599,1624]" name="Malaysia" pageId="9" pageNumber="562">MALAYSIA</collectingCountry>
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</emphasis>
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:
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<specimenCount id="9D3CAB905A617558D2BAA858F3B9FA0E" box="[434,476,1599,1623]" pageId="9" pageNumber="562" type="generic">1 ♂</specimenCount>
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(L14), BORNEO, Sabah, Crocker range, Gunung Emas,
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<date id="FF8446D95A617558D757A858F6DBFA0E" box="[1119,1214,1599,1623]" pageId="9" pageNumber="562" value="1998-06">VI.1998</date>
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, Kodada.
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</paragraph>
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</subSubSection>
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<subSubSection id="C32033925A617558D3CFA804F760FAB0" pageId="9" pageNumber="562" type="discussion">
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<paragraph id="8B8560195A617558D3CFA804F760FAB0" blockId="9.[151,1436,1599,2022]" pageId="9" pageNumber="562">
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<emphasis id="B94EBC0B5A617558D3CFA804F351FA25" bold="true" box="[199,308,1635,1660]" pageId="9" pageNumber="562">Remarks</emphasis>
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. The following male characters of
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<taxonomicName id="4C3A1B9A5A617558D1B4A803F133FA22" box="[700,854,1635,1659]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="562" phylum="Arthropoda" rank="species" species="sabahensis">
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<emphasis id="B94EBC0B5A617558D1B4A803F133FA22" box="[700,854,1635,1659]" italics="true" pageId="9" pageNumber="562">E. sabahensis</emphasis>
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</taxonomicName>
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have not been described before. Third antennomere (
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<figureCitation id="13017C9C5A617558D397A8E0F286FAC6" box="[159,227,1671,1696]" captionStart="FIGURES 1 – 7" captionStartId="2.[151,264,1750,1773]" captionTargetBox="[183,1395,205,1699]" captionTargetId="figure@2.[183,1395,193,1713]" captionTargetPageId="2" captionText="FIGURES 1 – 7. Antenna. 1, Echinoaesalus hidakai, L 12, western Malaysia; 2, E. borneoensis, L 11, Borneo; 3, E. timidus, L 19, western Malaysia; 4, E. sabahensis, L 14, Borneo; 5, E. gedeensis, holotype, Java; 6, E. cechovskyi, holotype, western Malaysia; 7, E. cechovskyi, paratype, Singapore. Arrow directed to end of third antennomere." httpUri="https://zenodo.org/record/241184/files/figure.png" pageId="9" pageNumber="562">Fig. 4</figureCitation>
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) more elongate than in
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<emphasis id="B94EBC0B5A617558D2E4A8EFF09DFAC6" box="[492,760,1671,1695]" italics="true" pageId="9" pageNumber="562">
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<taxonomicName id="4C3A1B9A5A617558D2E4A8EFF005FAC6" box="[492,608,1671,1695]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="562" phylum="Arthropoda" rank="species" species="timidus">E. timidus</taxonomicName>
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,
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<taxonomicName id="4C3A1B9A5A617558D164A8EFF09DFAC6" box="[620,760,1671,1695]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="562" phylum="Arthropoda" rank="species" species="gedeensis">E. gedeensis</taxonomicName>
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</emphasis>
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, and
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<taxonomicName id="4C3A1B9A5A617558D03DA8EFF1A8FAC6" box="[821,973,1671,1695]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="562" phylum="Arthropoda" rank="species" species="cechovskyi">
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<emphasis id="B94EBC0B5A617558D03DA8EFF1A8FAC6" box="[821,973,1671,1695]" italics="true" pageId="9" pageNumber="562">E. cechovskyi</emphasis>
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</taxonomicName>
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(described below). Labium, maxilla, and hindwing as in other species of the
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<taxonomicName id="4C3A1B9A5A617558D12BA8C8F0F0FA9F" box="[547,661,1710,1734]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="562" phylum="Arthropoda" rank="species" species="timidus">
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<emphasis id="B94EBC0B5A617558D12BA8C8F0F0FA9F" box="[547,661,1710,1734]" italics="true" pageId="9" pageNumber="562">E. timidus</emphasis>
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</taxonomicName>
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group. The 9th abdominal segment of male (Fig. 87) with a very long basal lobe which is at least three times as long as that of other species of the subgenus
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<emphasis id="B94EBC0B5A617558D768A8B5F69BFAB3" box="[1120,1278,1746,1770]" italics="true" pageId="9" pageNumber="562">Zelenkaesalus</emphasis>
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.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C32033925A61755BD3CFA892F0EEFCAE" lastPageId="10" lastPageNumber="563" pageId="9" pageNumber="562" type="description">
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<paragraph id="8B8560195A617558D3CFA892F7F9FBBF" blockId="9.[151,1436,1599,2022]" pageId="9" pageNumber="562">
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<emphasis id="B94EBC0B5A617558D3CFA892F311FB57" bold="true" box="[199,372,1781,1806]" pageId="9" pageNumber="562">Male genitalia</emphasis>
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(
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<figureCitation id="13017C9C5A617558D28DA892F3B9FB57" box="[389,476,1781,1806]" captionStart="FIGURES 90 – 102" captionStartId="5.[151,264,1797,1820]" captionTargetBox="[151,1436,519,1775]" captionTargetId="figure@5.[151,1436,519,1775]" captionTargetPageId="5" captionText="FIGURES 90 – 102. Aedeagus in dorsal, left lateral, ventral, and right lateral views under same scale (scale bar 1 mm). 90, Echinoaesalus hidakai, L 12, western Malaysia; 91, E. borneoensis, L 11, Borneo; 92, E. borneoensis, L 01, Borneo; 93 – 98, E. timidus; 93, L 08; 94, L 02; 95, L 17; 96, L 23; 97, L 21; 98, L 22; 99, E. sabahensis, L 14, Borneo; 100, E. timidus, L 18; 101, E. cechovskyi, holotype, western Malaysia; 102, E. gedeensis, holotype, Java. Arrows directed to ends of parameres." httpUri="https://zenodo.org/record/241187/files/figure.png" pageId="9" pageNumber="562">Fig. 99</figureCitation>
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). Since
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<taxonomicName id="4C3A1B9A5A617558D132A890F0CAFB57" box="[570,687,1782,1806]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="562" phylum="Arthropoda" rank="species" species="timidus">
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<emphasis id="B94EBC0B5A617558D132A890F0CAFB57" box="[570,687,1782,1806]" italics="true" pageId="9" pageNumber="562">E. timidus</emphasis>
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</taxonomicName>
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is known to be very variable in mentum and pronotal plate, it is rather difficult to identify the specimens of
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<taxonomicName id="4C3A1B9A5A617558D18AA97DF178FB6B" box="[642,797,1818,1842]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="562" phylum="Arthropoda" rank="species" species="sabahensis">
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<emphasis id="B94EBC0B5A617558D18AA97DF178FB6B" box="[642,797,1818,1842]" italics="true" pageId="9" pageNumber="562">E. sabahensis</emphasis>
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</taxonomicName>
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without a dissection. The aedeagus of
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<taxonomicName id="4C3A1B9A5A617558D7DCA97DF70AFB6B" box="[1236,1391,1818,1842]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="562" phylum="Arthropoda" rank="species" species="sabahensis">
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<emphasis id="B94EBC0B5A617558D7DCA97DF70AFB6B" box="[1236,1391,1818,1842]" italics="true" pageId="9" pageNumber="562">E. sabahensis</emphasis>
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</taxonomicName>
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has been illustrated as a rough hand drawing in the original description (
|
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<bibRefCitation id="EFAB1DE85A617558D0A7A95AF62AFB0F" author="Zelenka" box="[943,1103,1853,1878]" pageId="9" pageNumber="562" refString="Zelenka, W. (1994) Zwei neue Echinoaesalus - Arten aus Sudostasien (Coleoptera, Lucanidae). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 46, 56 - 61." type="journal article" year="1994">Zelenka 1994</bibRefCitation>
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), which shows the following characters: parameres half as long as the dorsal side of median lobe; dorsal side of median lobe enlarged from base to terminal end in width and broadly rounded at the end. The aedeagus of the specimen examined by us (
|
||
<figureCitation id="13017C9C5A617558D649A9E2F7F1FBC7" box="[1345,1428,1925,1950]" captionStart="FIGURES 90 – 102" captionStartId="5.[151,264,1797,1820]" captionTargetBox="[151,1436,519,1775]" captionTargetId="figure@5.[151,1436,519,1775]" captionTargetPageId="5" captionText="FIGURES 90 – 102. Aedeagus in dorsal, left lateral, ventral, and right lateral views under same scale (scale bar 1 mm). 90, Echinoaesalus hidakai, L 12, western Malaysia; 91, E. borneoensis, L 11, Borneo; 92, E. borneoensis, L 01, Borneo; 93 – 98, E. timidus; 93, L 08; 94, L 02; 95, L 17; 96, L 23; 97, L 21; 98, L 22; 99, E. sabahensis, L 14, Borneo; 100, E. timidus, L 18; 101, E. cechovskyi, holotype, western Malaysia; 102, E. gedeensis, holotype, Java. Arrows directed to ends of parameres." httpUri="https://zenodo.org/record/241187/files/figure.png" pageId="9" pageNumber="562">Fig. 99</figureCitation>
|
||
) fits the original figure (
|
||
<bibRefCitation id="EFAB1DE85A617558D2A4A9CEF028FB9B" author="Zelenka" box="[428,589,1961,1986]" pageId="9" pageNumber="562" refString="Zelenka, W. (1994) Zwei neue Echinoaesalus - Arten aus Sudostasien (Coleoptera, Lucanidae). Zeitschrift der Arbeitsgemeinschaft Osterreichischer Entomologen, 46, 56 - 61." type="journal article" year="1994">Zelenka 1994</bibRefCitation>
|
||
) very well. Beside this, the following characters are also noted: size of aedeagus similar to that of
|
||
<taxonomicName id="4C3A1B9A5A617558D2CDA9A8F034FBBF" box="[453,593,1998,2022]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="562" phylum="Arthropoda" rank="species" species="gedeensis">
|
||
<emphasis id="B94EBC0B5A617558D2CDA9A8F034FBBF" box="[453,593,1998,2022]" italics="true" pageId="9" pageNumber="562">E. gedeensis</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13017C9C5A617558D168A9AAF0DAFBBF" box="[608,703,1997,2022]" captionStart="FIGURES 90 – 102" captionStartId="5.[151,264,1797,1820]" captionTargetBox="[151,1436,519,1775]" captionTargetId="figure@5.[151,1436,519,1775]" captionTargetPageId="5" captionText="FIGURES 90 – 102. Aedeagus in dorsal, left lateral, ventral, and right lateral views under same scale (scale bar 1 mm). 90, Echinoaesalus hidakai, L 12, western Malaysia; 91, E. borneoensis, L 11, Borneo; 92, E. borneoensis, L 01, Borneo; 93 – 98, E. timidus; 93, L 08; 94, L 02; 95, L 17; 96, L 23; 97, L 21; 98, L 22; 99, E. sabahensis, L 14, Borneo; 100, E. timidus, L 18; 101, E. cechovskyi, holotype, western Malaysia; 102, E. gedeensis, holotype, Java. Arrows directed to ends of parameres." httpUri="https://zenodo.org/record/241187/files/figure.png" pageId="9" pageNumber="562">Fig. 102</figureCitation>
|
||
) and
|
||
<taxonomicName id="4C3A1B9A5A617558D1F5A9A8F1F0FBBF" box="[765,917,1998,2022]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="562" phylum="Arthropoda" rank="species" species="cechovskyi">
|
||
<emphasis id="B94EBC0B5A617558D1F5A9A8F1F0FBBF" box="[765,917,1998,2022]" italics="true" pageId="9" pageNumber="562">E. cechovskyi</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13017C9C5A617558D0ADA9AAF663FBBF" box="[933,1030,1997,2022]" captionStart="FIGURES 90 – 102" captionStartId="5.[151,264,1797,1820]" captionTargetBox="[151,1436,519,1775]" captionTargetId="figure@5.[151,1436,519,1775]" captionTargetPageId="5" captionText="FIGURES 90 – 102. Aedeagus in dorsal, left lateral, ventral, and right lateral views under same scale (scale bar 1 mm). 90, Echinoaesalus hidakai, L 12, western Malaysia; 91, E. borneoensis, L 11, Borneo; 92, E. borneoensis, L 01, Borneo; 93 – 98, E. timidus; 93, L 08; 94, L 02; 95, L 17; 96, L 23; 97, L 21; 98, L 22; 99, E. sabahensis, L 14, Borneo; 100, E. timidus, L 18; 101, E. cechovskyi, holotype, western Malaysia; 102, E. gedeensis, holotype, Java. Arrows directed to ends of parameres." httpUri="https://zenodo.org/record/241187/files/figure.png" pageId="9" pageNumber="562">Fig. 101</figureCitation>
|
||
), but markedly larger than that of
|
||
<emphasis id="B94EBC0B5A617558D68CA9A8F7F9FBBF" box="[1412,1436,1999,2022]" italics="true" pageId="9" pageNumber="562">E.</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8B8560195A62755BD39FAEFFF0EEFCAE" blockId="10.[151,1436,151,285]" pageId="10" pageNumber="563">
|
||
<taxonomicName id="4C3A1B9A5A62755BD39FAEFFF28FFCE9" box="[151,234,152,176]" class="Insecta" family="Lucanidae" genus="Echinoaesalus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="563" phylum="Arthropoda" rank="species" species="timidus">
|
||
<emphasis id="B94EBC0B5A62755BD39FAEFFF28FFCE9" box="[151,234,152,176]" italics="true" pageId="10" pageNumber="563">timidus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="13017C9C5A62755BD3F0AEF0F3DCFCE9" box="[248,441,151,176]" captionStart="FIGURES 90 – 102" captionStartId="5.[151,264,1797,1820]" captionTargetBox="[151,1436,519,1775]" captionTargetId="figure@5.[151,1436,519,1775]" captionTargetPageId="5" captionText="FIGURES 90 – 102. Aedeagus in dorsal, left lateral, ventral, and right lateral views under same scale (scale bar 1 mm). 90, Echinoaesalus hidakai, L 12, western Malaysia; 91, E. borneoensis, L 11, Borneo; 92, E. borneoensis, L 01, Borneo; 93 – 98, E. timidus; 93, L 08; 94, L 02; 95, L 17; 96, L 23; 97, L 21; 98, L 22; 99, E. sabahensis, L 14, Borneo; 100, E. timidus, L 18; 101, E. cechovskyi, holotype, western Malaysia; 102, E. gedeensis, holotype, Java. Arrows directed to ends of parameres." httpUri="https://zenodo.org/record/241187/files/figure.png" pageId="10" pageNumber="563">Figs. 93–98, 100</figureCitation>
|
||
); lateral sides of the median lobe broadly colorless and non-sclerotized, causing a ventral plate well defined; ventral plate of median lobe contracted at base and broadly rounded, markedly wider than in other species of the subgenus
|
||
<emphasis id="B94EBC0B5A62755BD2EEAE87F0E0FCA1" box="[486,645,224,248]" italics="true" pageId="10" pageNumber="563">Zelenkaesalus</emphasis>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32033925A62755BD3CFAF63F0F4FD45" box="[199,657,260,285]" pageId="10" pageNumber="563" type="distribution">
|
||
<paragraph id="8B8560195A62755BD3CFAF63F0F4FD45" blockId="10.[151,1436,151,285]" box="[199,657,260,285]" pageId="10" pageNumber="563">
|
||
<emphasis id="B94EBC0B5A62755BD3CFAF63F33CFD44" bold="true" box="[199,345,260,285]" pageId="10" pageNumber="563">Distribution</emphasis>
|
||
. Only known from Borneo.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |