284 lines
34 KiB
XML
284 lines
34 KiB
XML
<document id="AE94839DED14F13C8124BADDC0EC47F5" ID-DOI="10.11646/zootaxa.3691.1.8" ID-GBIF-Dataset="366bc68a-ca62-459a-a01c-660fd1dd8bd6" ID-ISSN="1175-5326" ID-Zenodo-Dep="217303" ID-ZooBank="A3C253A5-AD10-4FCB-A053-26B6A956EC4D" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460650271193" checkinUser="plazi" docAuthor="Lujan, Nathan K., Steele, Sarah & Velasquez, Miquel" docDate="2013" docId="9A4F8785FFB3FF96FF2DF884FB3BE723" docLanguage="en" docName="zt03691p198.pdf" docOrigin="Zootaxa 3691 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Panaqolus albivermis Lujan, Steele & Velasquez, 2013, new species" docType="treatment" docVersion="5" lastPageNumber="197" masterDocId="6676FFFDFFB2FF93FFBAFF8AFFC6E47F" masterDocTitle="A new distinctively banded species of Panaqolus (Siluriformes: Loricariidae) from the western Amazon Basin in Peru" masterLastPageNumber="198" masterPageNumber="192" pageNumber="193" updateTime="1698337804190" updateUser="plazi">
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<mods:titleInfo id="33B2FDDD4BCFF1630DE1EA74CD8F6BA4">
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<mods:title id="DA20DA5D5D63B0234578BEF534FEC9CD">A new distinctively banded species of Panaqolus (Siluriformes: Loricariidae) from the western Amazon Basin in Peru</mods:title>
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<mods:name id="E76D5C8A58878389806854EF08F02E00" type="personal">
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<mods:namePart id="6995D214B52587ADA02F4194119D0858">Lujan, Nathan K.</mods:namePart>
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<mods:roleTerm id="6A43B30AC6443BCBF19DB79166347E77">Author</mods:roleTerm>
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<mods:namePart id="A3B8780E960DE1ED0FAA82F646B451BD">Steele, Sarah</mods:namePart>
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<mods:name id="B079706893661AB22053EE883CD9866D" type="personal">
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<mods:roleTerm id="E568D7976F9F748CAA3648095184C206">Author</mods:roleTerm>
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<mods:namePart id="8A5F1DF5613FE6438EE3E9C441FDFFDF">Velasquez, Miquel</mods:namePart>
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<mods:typeOfResource id="A31AE8C80137F5324C3D4710B1383983">text</mods:typeOfResource>
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<mods:title id="9576561762F43719AEE8439C5FA4EAB9">Zootaxa</mods:title>
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<mods:part id="E04760BA4277EFFBD0BAD557AB6751C8">
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<mods:date id="0813EA937155AF581D74F83A2EDC5FD0">2013</mods:date>
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<mods:detail id="95B3E0CD0368DB7643809B543D73940D" type="volume">
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<mods:number id="391D687BD71CAA492498394C4F8E04E0">3691</mods:number>
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<mods:identifier id="4BEF8A8A827E28CBBCA6548A049D6712" type="DOI">10.11646/zootaxa.3691.1.8</mods:identifier>
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<treatment id="9A4F8785FFB3FF96FF2DF884FB3BE723" ID-DOI="http://doi.org/10.5281/zenodo.6162737" ID-GBIF-Taxon="119572597" ID-Zenodo-Dep="6162737" LSID="urn:lsid:plazi:treatment:9A4F8785FFB3FF96FF2DF884FB3BE723" httpUri="http://treatment.plazi.org/id/9A4F8785FFB3FF96FF2DF884FB3BE723" lastPageId="5" lastPageNumber="197" pageId="1" pageNumber="193">
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<subSubSection id="5AFC6518FFB3FF92FF2DF884FE9BE336" pageId="1" pageNumber="193" type="nomenclature">
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<paragraph id="12593693FFB3FF92FF2DF884FDFCE357" blockId="1.[151,570,1806,1865]" box="[151,570,1806,1832]" pageId="1" pageNumber="193">
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<heading id="491181FFFFB3FF92FF2DF884FDFCE357" bold="true" box="[151,570,1806,1832]" fontSize="11" level="1" pageId="1" pageNumber="193" reason="1">
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<emphasis id="2092EA81FFB3FF92FF2DF884FDFCE357" bold="true" box="[151,570,1806,1832]" pageId="1" pageNumber="193">
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<taxonomicName id="D5E64D10FFB3FF92FF2DF884FE5AE357" ID-CoL="4C9CJ" box="[151,412,1806,1832]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="1" pageNumber="193" phylum="Chordata" rank="species" species="albivermis" status="sp. nov.">
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<emphasis id="2092EA81FFB3FF92FF2DF884FE5AE357" bold="true" box="[151,412,1806,1832]" italics="true" pageId="1" pageNumber="193">Panaqolus albivermis</emphasis>
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</taxonomicName>
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,
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<taxonomicNameLabel id="3BA157FAFFB3FF92FE11F884FDFCE357" box="[427,570,1806,1832]" pageId="1" pageNumber="193" rank="species">new species</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="12593693FFB3FF92FF2DF8BBFE9BE336" blockId="1.[151,570,1806,1865]" box="[151,349,1841,1865]" pageId="1" pageNumber="193">
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<figureCitation id="8ADD2A16FFB3FF92FF2DF8BBFF30E336" box="[151,246,1841,1865]" captionStart="FIGURE 1" captionStartId="2.[151,250,1946,1969]" captionTargetBox="[151,1436,501,1925]" captionTargetId="figure@2.[151,1436,497,1925]" captionTargetPageId="2" captionText="FIGURE 1. Holotype of Panaqolus albivermis n. sp., 95.8 mm SL, Peru, Padre Abad Province, Irazola District, San Alejandro River, Ucayali River drainage, 195 masl, 8 ° 55 ' 02 '' N, 75 ° 12 ' 26 '' W, M. Velasquez, 13 June 2009 (photo by NKL)." httpUri="https://zenodo.org/record/217304/files/figure.png" pageId="1" pageNumber="193">Figure 1</figureCitation>
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.
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<tableCitation id="5F640328FFB3FF92FEB9F8BBFE91E336" box="[259,343,1841,1865]" captionStart="TABLE 1" captionStartId="4.[151,242,151,175]" captionText="TABLE 1. Selected morphometric characters for Panaqolus albivermis n. sp. Interlandmarks (ILM) are the two points between which measurements were taken (from Armbruster, 2003; * = too small to be measured precisely)." pageId="1" pageNumber="193">Table 1</tableCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="5AFC6518FFB3FF91FF2DF8F0FD85E4AB" lastPageId="2" lastPageNumber="194" pageId="1" pageNumber="193" type="materials_examined">
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<paragraph id="12593693FFB3FF92FF2DF8F0FBDEE3CA" blockId="1.[151,1436,1913,1973]" pageId="1" pageNumber="193">
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<emphasis id="2092EA81FFB3FF92FF2DF8F0FECDE3ED" bold="true" box="[151,267,1914,1938]" pageId="1" pageNumber="193">
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<typeStatus id="CD5D8831FFB3FF92FF2DF8F0FEC0E3ED" box="[151,262,1914,1938]" pageId="1" pageNumber="193" type="holotype">Holotype</typeStatus>
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.
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</emphasis>
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MUSM 35879,
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<quantity id="D51E9B76FFB3FF92FE78F8F3FDE1E3EE" box="[450,551,1913,1937]" metricMagnitude="-2" metricUnit="m" metricValue="9.58" pageId="1" pageNumber="193" unit="mm" value="95.8">95.8 mm</quantity>
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SL,
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<collectingCountry id="6AF17603FFB3FF92FDE6F8F0FD54E3EE" box="[604,658,1914,1937]" name="Peru" pageId="1" pageNumber="193">Peru</collectingCountry>
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, Padre Abad Province, Irazola District, San Alejandro River, Ucayali River drainage, 195 masl,
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<geoCoordinate id="77D25054FFB3FF92FE04F817FDF3E3CA" box="[446,565,1949,1973]" direction="north" orientation="latitude" pageId="1" pageNumber="193" precision="15" value="8.917223">8°55'02''N</geoCoordinate>
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,
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<geoCoordinate id="77D25054FFB3FF92FDFBF817FD0CE3CA" box="[577,714,1949,1973]" direction="west" orientation="longitude" pageId="1" pageNumber="193" precision="15" value="-75.20722">75°12'26''W</geoCoordinate>
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, M. Velasquez,
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<date id="66581053FFB3FF92FCC1F817FBD2E3CA" box="[891,1044,1949,1973]" pageId="1" pageNumber="193" value="2009-06-13">13 June 2009</date>
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.
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</paragraph>
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<paragraph id="12593693FFB0FF91FF7DFF12FD85E4AB" blockId="2.[151,1437,151,463]" pageId="2" pageNumber="194">
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<emphasis id="2092EA81FFB0FF91FF7DFF12FE80E4CF" bold="true" box="[199,326,152,176]" pageId="2" pageNumber="194">
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<typeStatus id="CD5D8831FFB0FF91FF7DFF12FE87E4CF" box="[199,321,152,176]" pageId="2" pageNumber="194" type="paratype">Paratypes</typeStatus>
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.
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</emphasis>
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Three specimens, collected with
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<typeStatus id="CD5D8831FFB0FF91FD7DFF1DFCEAE4D0" box="[711,812,151,175]" pageId="2" pageNumber="194" type="holotype">holotype</typeStatus>
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. ANSP 182815, 1:
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<quantity id="D51E9B76FFB0FF91FBB6FF1DFBB4E4D0" box="[1036,1138,151,175]" metricMagnitude="-2" metricUnit="m" metricValue="7.87" pageId="2" pageNumber="194" unit="mm" value="78.7">78.7 mm</quantity>
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SL; AUM 58781, 1:
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<quantity id="D51E9B76FFB0FF91FAD1FF1DFF05E4AB" metricMagnitude="-2" metricUnit="m" metricValue="7.09" pageId="2" pageNumber="194" unit="mm" value="70.9">70.9 mm</quantity>
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SL; ROM 93945, 1:
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<quantity id="D51E9B76FFB0FF91FE0AFF36FDD2E4AB" box="[432,532,188,212]" metricMagnitude="-2" metricUnit="m" metricValue="4.55" pageId="2" pageNumber="194" unit="mm" value="45.5">45.5 mm</quantity>
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SL.
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</paragraph>
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</subSubSection>
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<subSubSection id="5AFC6518FFB0FF91FF7DFF55FAE5E5B0" pageId="2" pageNumber="194" type="diagnosis">
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||
<paragraph id="12593693FFB0FF91FF7DFF55FAE5E5B0" blockId="2.[151,1437,151,463]" pageId="2" pageNumber="194">
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<emphasis id="2092EA81FFB0FF91FF7DFF55FE87E487" bold="true" box="[199,321,223,248]" pageId="2" pageNumber="194">Diagnosis.</emphasis>
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<taxonomicName id="D5E64D10FFB0FF91FEFDFF6AFDFAE487" box="[327,572,224,248]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="194" phylum="Chordata" rank="species" species="albivermis">
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<emphasis id="2092EA81FFB0FF91FEFDFF6AFDFAE487" box="[327,572,224,248]" italics="true" pageId="2" pageNumber="194">Panaqolus albivermis</emphasis>
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</taxonomicName>
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can be diagnosed from all other
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<taxonomicName id="D5E64D10FFB0FF91FC12FF6AFBE6E487" box="[936,1056,224,248]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="194" phylum="Chordata" rank="genus">
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<emphasis id="2092EA81FFB0FF91FC12FF6AFBE6E487" box="[936,1056,224,248]" italics="true" pageId="2" pageNumber="194">Panaqolus</emphasis>
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</taxonomicName>
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except
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<taxonomicName id="D5E64D10FFB0FF91FBCFFF6BFB23E487" box="[1141,1253,225,248]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="194" phylum="Chordata" rank="species" species="maccus">
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<emphasis id="2092EA81FFB0FF91FBCFFF6BFB23E487" box="[1141,1253,225,248]" italics="true" pageId="2" pageNumber="194">P. maccus</emphasis>
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</taxonomicName>
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by having head, body, and fins with widely separated small white to yellow spots, vermiculations, and/or thin oblique bands on a black base (vs. exclusively small white to yellow spots on a black base in
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<taxonomicName id="D5E64D10FFB0FF91FBABFEA3FB10E53F" box="[1041,1238,296,320]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="194" phylum="Chordata" rank="species" species="albomaculatus">
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<emphasis id="2092EA81FFB0FF91FBABFEA3FB10E53F" box="[1041,1238,296,320]" italics="true" pageId="2" pageNumber="194">P. albomaculatus</emphasis>
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</taxonomicName>
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, generally broad oblique bands of alternating light to dark brown in
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<taxonomicName id="D5E64D10FFB0FF91FD76FEC7FC80E51B" box="[716,838,332,356]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="194" phylum="Chordata" rank="species" species="changae">
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<emphasis id="2092EA81FFB0FF91FD76FEC7FC80E51B" box="[716,838,332,356]" italics="true" pageId="2" pageNumber="194">P. changae</emphasis>
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</taxonomicName>
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,
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<emphasis id="2092EA81FFB0FF91FCE9FEC7FBABE51B" box="[851,1133,333,356]" italics="true" pageId="2" pageNumber="194">
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<taxonomicName id="D5E64D10FFB0FF91FCE9FEC7FC0EE51B" box="[851,968,333,356]" class="Actinopterygii" family="Loricariidae" genus="Panaque" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="194" phylum="Chordata" rank="species" species="gnomus">P. gnomus</taxonomicName>
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,
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<taxonomicName id="D5E64D10FFB0FF91FC68FEC7FBABE51B" box="[978,1133,333,356]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="194" phylum="Chordata" rank="species" species="purusiensis">P. purusiensis</taxonomicName>
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</emphasis>
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, and a uniformly dark gray to black body color in
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<emphasis id="2092EA81FFB0FF91FE1AFEFBFDC2E5F7" box="[416,516,368,392]" italics="true" pageId="2" pageNumber="194">P. d e n t e x</emphasis>
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,
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<emphasis id="2092EA81FFB0FF91FDA2FEFBFDA1E5F7" box="[536,615,368,392]" italics="true" pageId="2" pageNumber="194">P. k o k o</emphasis>
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, and
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<taxonomicName id="D5E64D10FFB0FF91FD14FEFBFCFAE5F7" box="[686,828,369,392]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="194" phylum="Chordata" rank="species" species="nocturnus">
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<emphasis id="2092EA81FFB0FF91FD14FEFBFCFAE5F7" box="[686,828,369,392]" italics="true" pageId="2" pageNumber="194">P. nocturnus</emphasis>
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</taxonomicName>
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);
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<taxonomicName id="D5E64D10FFB0FF91FCEDFEFBFC2DE5F7" box="[855,1003,368,392]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="194" phylum="Chordata" rank="species" species="albivermis">
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<emphasis id="2092EA81FFB0FF91FCEDFEFBFC2DE5F7" box="[855,1003,368,392]" italics="true" pageId="2" pageNumber="194">P. albivermis</emphasis>
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</taxonomicName>
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can be diagnosed from
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<taxonomicName id="D5E64D10FFB0FF91FABEFEFBFAB0E5F7" box="[1284,1398,369,392]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="194" phylum="Chordata" rank="species" species="maccus">
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<emphasis id="2092EA81FFB0FF91FABEFEFBFAB0E5F7" box="[1284,1398,369,392]" italics="true" pageId="2" pageNumber="194">P. maccus</emphasis>
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</taxonomicName>
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by having a black base color (vs. brown), by having parallel dentary tooth rows (vs. acute intermandibular tooth row angle), and by having a larger known adult body size (
|
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<quantity id="D51E9B76FFB0FF91FD4DFE3DFC9AE5B0" box="[759,860,439,463]" metricMagnitude="-2" metricUnit="m" metricValue="9.58" pageId="2" pageNumber="194" unit="mm" value="95.8">95.8 mm</quantity>
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SL vs. 84.8; Schaefer & Stewart, 1993).
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</paragraph>
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</subSubSection>
|
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<caption id="4699661BFFB0FF91FF2DF810FAC0E3AF" httpUri="https://zenodo.org/record/217304/files/figure.png" pageId="2" pageNumber="194" targetBox="[151,1436,501,1925]" targetPageId="2">
|
||
<paragraph id="12593693FFB0FF91FF2DF810FAC0E3AF" blockId="2.[151,1436,1946,2000]" pageId="2" pageNumber="194">
|
||
<emphasis id="2092EA81FFB0FF91FF2DF810FED4E3CF" bold="true" box="[151,274,1946,1969]" pageId="2" pageNumber="194">FIGURE 1.</emphasis>
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Holotype of
|
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<taxonomicName id="D5E64D10FFB0FF91FE2DF811FDB3E3CE" box="[407,629,1947,1969]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="194" phylum="Chordata" rank="species" species="albivermis" status="sp. nov.">
|
||
<emphasis id="2092EA81FFB0FF91FE2DF811FDB3E3CE" box="[407,629,1947,1969]" italics="true" pageId="2" pageNumber="194">Panaqolus albivermis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2092EA81FFB0FF91FDC1F811FD75E3CF" bold="true" box="[635,691,1947,1968]" pageId="2" pageNumber="194">
|
||
<taxonomicNameLabel id="3BA157FAFFB0FF91FDC1F811FD75E3CF" box="[635,691,1947,1968]" pageId="2" pageNumber="194" rank="species">n. sp.</taxonomicNameLabel>
|
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</emphasis>
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, 95.8 mm SL, Peru, Padre Abad Province, Irazola District, San Alejandro River, Ucayali River drainage, 195 masl, 8°55'02''N, 75°12'26''W, M. Velasquez, 13 June 2009 (photo by NKL).
|
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</paragraph>
|
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</caption>
|
||
<subSubSection id="5AFC6518FFB1FF96FF7DFF1DFE62E6B3" lastPageId="5" lastPageNumber="197" pageId="3" pageNumber="195" type="description">
|
||
<paragraph id="12593693FFB1FF90FF7DFF1DFF15E586" blockId="3.[151,1437,151,685]" pageId="3" pageNumber="195">
|
||
<emphasis id="2092EA81FFB1FF90FF7DFF1DFE9EE4D0" bold="true" box="[199,344,151,176]" pageId="3" pageNumber="195">Description.</emphasis>
|
||
Morphometrics in
|
||
<tableCitation id="5F640328FFB1FF90FD82FF1DFD49E4D0" box="[568,655,151,175]" captionStart="TABLE 1" captionStartId="4.[151,242,151,175]" captionText="TABLE 1. Selected morphometric characters for Panaqolus albivermis n. sp. Interlandmarks (ILM) are the two points between which measurements were taken (from Armbruster, 2003; * = too small to be measured precisely)." pageId="3" pageNumber="195">Table 1</tableCitation>
|
||
. Largest specimen
|
||
<quantity id="D51E9B76FFB1FF90FCCAFF1DFC11E4D0" box="[880,983,151,175]" metricMagnitude="-2" metricUnit="m" metricValue="9.58" pageId="3" pageNumber="195" unit="mm" value="95.8">95.8 mm</quantity>
|
||
SL. Body depth increasing from tip of snout to greatest body depth at origin of dorsal fin, decreasing to posterior margin of adipose fin with slight increase at base of caudal fin. Dorsal profile slightly convex between snout and dorsal-fin origin. Ventral profile flat from snout to anus, slightly concave between anus and caudal fin. Ossified dermal plates with small odontodes covering snout and body flanks. Abdomen naked, lacking plates except for area between opercular openings. Cheek plates bearing hypertrophied, distally-hooked odontodes (mean 21, range 15-38,
|
||
<typeStatus id="CD5D8831FFB1FF90FB38FEC6FB22E51B" box="[1154,1252,332,356]" pageId="3" pageNumber="195" type="holotype">holotype</typeStatus>
|
||
38) evertible to approximately 90o from sagittal plane in dorsal view; longest odontodes extending to posterior exposed margin of opercle. Orbit positioned dorsolaterally on head with opening sloped ventrolaterally at approximately 45o from sagittal plane in anterior view. Snout rounded. Slight ridge from anterolateral corner of naris to dorsal margin of orbit.
|
||
</paragraph>
|
||
<paragraph id="12593693FFB1FF90FF7DFD8FFCB2E6D2" blockId="3.[151,1437,151,685]" pageId="3" pageNumber="195">
|
||
Oral disk occupying majority of ventral surface of head anterior of cleithrum. Premaxillary teeth 4-5 on left ramus (
|
||
<typeStatus id="CD5D8831FFB1FF90FF50FDA3FE88E63E" box="[234,334,553,577]" pageId="3" pageNumber="195" type="holotype">holotype</typeStatus>
|
||
= 5); dentary teeth 4-6 on left ramus (
|
||
<typeStatus id="CD5D8831FFB1FF90FD4DFDA3FC9DE63E" box="[759,859,553,577]" pageId="3" pageNumber="195" type="holotype">holotype</typeStatus>
|
||
= 4). All teeth with thick, strong shafts, and spoonshaped, paired cusps bent inward slightly; teeth slightly larger on dentary, teeth in middle of each row slightly larger than marginal teeth. Maxillary barbel short and attached to lower lip along most of length; ventral surface of labial disk with hemispherical papillae decreasing in size distally.
|
||
</paragraph>
|
||
<caption id="4699661BFFB1FF90FF2DF84CFB43E3A2" box="[151,1157,1990,2013]" httpUri="https://zenodo.org/record/217305/files/figure.png" pageId="3" pageNumber="195" targetBox="[166,1422,751,1955]" targetPageId="3">
|
||
<paragraph id="12593693FFB1FF90FF2DF84CFB43E3A2" blockId="3.[151,1157,1990,2013]" box="[151,1157,1990,2013]" pageId="3" pageNumber="195">
|
||
<emphasis id="2092EA81FFB1FF90FF2DF84CFED5E3A3" bold="true" box="[151,275,1990,2013]" pageId="3" pageNumber="195">FIGURE 2.</emphasis>
|
||
Map of northern Peru showing the type locality (open circle) of
|
||
<taxonomicName id="D5E64D10FFB1FF90FC18F84DFBB9E3A2" box="[930,1151,1991,2013]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="195" phylum="Chordata" rank="species" species="albivermis">
|
||
<emphasis id="2092EA81FFB1FF90FC18F84DFBB9E3A2" box="[930,1151,1991,2013]" italics="true" pageId="3" pageNumber="195">Panaqolus albivermis</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="4699661BFFB6FF97FF2DFF1DFAD4E4B0" pageId="4" pageNumber="196">
|
||
<paragraph id="12593693FFB6FF97FF2DFF1DFAD4E4B0" blockId="4.[151,1436,151,207]" pageId="4" pageNumber="196">
|
||
<emphasis id="2092EA81FFB6FF97FF2DFF1DFECBE4D0" bold="true" box="[151,269,151,175]" pageId="4" pageNumber="196">TABLE 1.</emphasis>
|
||
Selected morphometric characters for
|
||
<taxonomicName id="D5E64D10FFB6FF97FD0AFF12FC5DE4D0" box="[688,923,152,175]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="196" phylum="Chordata" rank="species" species="albivermis" status="sp. nov.">
|
||
<emphasis id="2092EA81FFB6FF97FD0AFF12FC5DE4D0" box="[688,923,152,175]" italics="true" pageId="4" pageNumber="196">Panaqolus albivermis</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="2092EA81FFB6FF97FC18FF1DFC19E4D0" bold="true" box="[930,991,151,175]" pageId="4" pageNumber="196">
|
||
<taxonomicNameLabel id="3BA157FAFFB6FF97FC18FF1DFC19E4D0" box="[930,991,151,175]" pageId="4" pageNumber="196" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
Interlandmarks (ILM) are the two points between which measurements were taken (from Armbruster, 2003; * = too small to be measured precisely).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="12593693FFB6FF97FD54FF60FBDCE57F" blockId="4.[159,1402,234,1992]" box="[750,1050,234,256]" pageId="4" pageNumber="196">
|
||
<typeStatus id="CD5D8831FFB6FF97FD54FF60FC8AE57F" box="[750,844,234,256]" pageId="4" pageNumber="196" type="holotype">Holotype</typeStatus>
|
||
<typeStatus id="CD5D8831FFB6FF97FC0DFF61FBDCE57F" box="[951,1050,235,256]" pageId="4" pageNumber="196" type="paratype">Paratypes</typeStatus>
|
||
</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FE9FFABCE556" blockId="4.[159,1402,234,1992]" box="[159,1402,275,298]" pageId="4" pageNumber="196">ILM Measurement MUSM 35879 ANSP 182815 AUM 58781 ROM 93945</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FEB7FE4CE503" blockId="4.[159,1402,234,1992]" pageId="4" pageNumber="196">1–20 Standard length 95.8 78.7 69.9 45.5 % of SL:</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FE1AFAE1E5DA" blockId="4.[159,1402,234,1992]" box="[159,1319,400,422]" pageId="4" pageNumber="196">1–10 Predorsal length 45.2 42.8 46.2 44.0</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FE30FAEEE5B0" blockId="4.[159,1402,234,1992]" box="[159,1320,441,463]" pageId="4" pageNumber="196">1–7 Head length 36.1 35.3 37.9 36.4</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FE69FADDE587" blockId="4.[159,1402,234,1992]" box="[159,1307,482,504]" pageId="4" pageNumber="196">7–10 Head-dorsal length 9.0 8.6 8.9 8.1</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FD86FAEEE65E" blockId="4.[159,1402,234,1992]" box="[159,1320,523,545]" pageId="4" pageNumber="196">8–9 Cleithral width 30.8 32.4 31.4 29.7</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FDBFFAEEE635" blockId="4.[159,1402,234,1992]" box="[159,1320,565,587]" pageId="4" pageNumber="196">1–12 Head-pectoral length 23.1 24.1 25.2 23.7</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FDD5FAEEE60B" blockId="4.[159,1402,234,1992]" box="[159,1320,606,628]" pageId="4" pageNumber="196">12–13 Thorax length 21.5 27.3 23.9 25.5</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FD02FAEEE6E2" blockId="4.[159,1402,234,1992]" box="[159,1320,648,670]" pageId="4" pageNumber="196">12–29 Pectoral-spine length 35.2 33.9 32.1 33.4</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FD38FAE1E6B8" blockId="4.[159,1402,234,1992]" box="[159,1319,689,711]" pageId="4" pageNumber="196">13–14 Abdominal length 24.3 24.9 23.4 21.6</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FD51FAEEE68F" blockId="4.[159,1402,234,1992]" box="[159,1320,730,752]" pageId="4" pageNumber="196">13–30 Pelvic-spine length 30.7 30.7 26.8 30.4</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FC8EFAEEE766" blockId="4.[159,1402,234,1992]" box="[159,1320,771,793]" pageId="4" pageNumber="196">13-13' Pelvic girdle width 19.9 20.5 18.5 21.0</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FCA7FAEEE73D" blockId="4.[159,1402,234,1992]" box="[159,1320,813,835]" pageId="4" pageNumber="196">14–15 Postanal length 30.9 31.9 29.0 30.1</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FCDDFAE1E713" blockId="4.[159,1402,234,1992]" box="[159,1319,854,876]" pageId="4" pageNumber="196">14–31 Anal-fin spine length 13.8 14.2 14.9 11.0</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FC0AFAEEE7EA" blockId="4.[159,1402,234,1992]" box="[159,1320,896,918]" pageId="4" pageNumber="196">10–12 Dorsal-pectoral depth 29.5 28.5 30.9 30.0</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FC20FAE1E7C0" blockId="4.[159,1402,234,1992]" box="[159,1319,937,959]" pageId="4" pageNumber="196">10–11 Dorsal spine length 33.0 34.3 33.6 36.0</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FC59FAE1E797" blockId="4.[159,1402,234,1992]" box="[159,1319,978,1000]" pageId="4" pageNumber="196">10–13 Dorsal-pelvic depth 23.6 22.0 22.3 20.4</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FC71FAE1E06E" blockId="4.[159,1402,234,1992]" box="[159,1319,1019,1041]" pageId="4" pageNumber="196">10–16 Dorsal-fin base length 28.3 28.3 26.9 24.5</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FBAFFAE1E045" blockId="4.[159,1402,234,1992]" box="[159,1319,1061,1083]" pageId="4" pageNumber="196">16–17 Dorsal-adipose distance 14.7 13.9 14.1 11.8</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FBC5FAE1E01B" blockId="4.[159,1402,234,1992]" box="[159,1319,1102,1124]" pageId="4" pageNumber="196">17–18 Adipose-spine length 11.3 12.0 10.9 11.4</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FBF2FAE1E0F2" blockId="4.[159,1402,234,1992]" box="[159,1319,1144,1166]" pageId="4" pageNumber="196">17–19 Adipose-upper caudal distance 17.6 11.6 17.4 18.9</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FB28FADDE0C8" blockId="4.[159,1402,234,1992]" box="[159,1307,1185,1207]" pageId="4" pageNumber="196">17-18' Adipose height 10.7 9.7 11.2 9.9</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FB40FAEEE09F" blockId="4.[159,1402,234,1992]" box="[159,1320,1226,1248]" pageId="4" pageNumber="196">15–19 Caudal peduncle depth 14.9 13.3 14.4 12.2</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FB7EFAEEE176" blockId="4.[159,1402,234,1992]" box="[159,1320,1267,1290]" pageId="4" pageNumber="196">20-32 Caudal peduncle-middle caudal ray 16.4 12.0 16.0 17.5</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FA94FAEEE14C" blockId="4.[159,1402,234,1992]" box="[159,1320,1309,1331]" pageId="4" pageNumber="196">20-33 Caudal peduncle-dorsal caudal spine 31.4 33.7 28.4 20.4</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FACCFAEEE123" blockId="4.[159,1402,234,1992]" box="[159,1320,1350,1372]" pageId="4" pageNumber="196">15–17 Adipose-lower caudal depth 22.8 25.2 23.1 22.6</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FAFAFAEEE1F9" blockId="4.[159,1402,234,1992]" box="[159,1320,1392,1414]" pageId="4" pageNumber="196">14–17 Adipose-anal depth 17.3 16.4 17.9 15.8</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FA13FAE1E1D0" blockId="4.[159,1402,234,1992]" box="[159,1319,1433,1455]" pageId="4" pageNumber="196">14–16 Dorsal-anal depth 16.6 15.7 17.2 12.9</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25FA48FE48E27D" blockId="4.[159,1402,234,1992]" pageId="4" pageNumber="196">13–16 Pelvic-dorsal depth 27.7 25.0 28.4 24.4 % of HL:</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25F99FFAEEE254" blockId="4.[159,1402,234,1992]" box="[159,1320,1557,1579]" pageId="4" pageNumber="196">5–7 Head-eye length 29.3 34.1 32.1 53.3</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25F9B4FAEEE22B" blockId="4.[159,1402,234,1992]" box="[159,1320,1598,1620]" pageId="4" pageNumber="196">4–5 Orbit diameter 17.1 18.1 17.6 20.5</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25F9E2FAEEE201" blockId="4.[159,1402,234,1992]" box="[159,1320,1640,1662]" pageId="4" pageNumber="196">1–4 Snout length 66.1 62.1 60.3 59.0</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25F918FAE1E2D8" blockId="4.[159,1402,234,1992]" box="[159,1319,1681,1703]" pageId="4" pageNumber="196">2–3 Internares width 13.5 13.8 13.8 15.8</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25F930FAE1E2AF" blockId="4.[159,1402,234,1992]" box="[159,1319,1722,1744]" pageId="4" pageNumber="196">5–6 Interorbital width 40.8 39.4 38.7 41.0</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25F96EFAEEE286" blockId="4.[159,1402,234,1992]" box="[159,1320,1763,1785]" pageId="4" pageNumber="196">7–12 Head depth 68.3 70.4 63.3 68.3</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25F884FAEEE35C" blockId="4.[159,1402,234,1992]" box="[159,1320,1805,1827]" pageId="4" pageNumber="196">1–24 Mouth length 47.5 45.6 40.5 44.2</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25F8BDFAEEE333" blockId="4.[159,1402,234,1992]" box="[159,1320,1846,1868]" pageId="4" pageNumber="196">21–22 Mouth width 40.5 36.0 40.7 37.7</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25F8EAFADDE30A" blockId="4.[159,1402,234,1992]" box="[159,1307,1888,1910]" pageId="4" pageNumber="196">22–23 Barbel length 9.6 10.1 11.7 8.1</paragraph>
|
||
<paragraph id="12593693FFB6FF97FF25F803FACEE3E0" blockId="4.[159,1402,234,1992]" box="[159,1288,1929,1951]" pageId="4" pageNumber="196">25–26 Dentary tooth cup length 18.9 16.6 10.5 *</paragraph>
|
||
<paragraph id="12593693FFB6FF96FF25F839FA50E51B" blockId="4.[159,1402,234,1992]" lastBlockId="5.[151,1437,151,860]" lastPageId="5" lastPageNumber="197" pageId="4" pageNumber="196">
|
||
27–28 Premaxillary tooth cup length 11.9 11.4 9.4 * Dorsal fin II,7(3)*, I,8(1); dorsal-fin spinelet small,
|
||
<emphasis id="2092EA81FFB7FF96FCF9FF13FC97E4CF" box="[835,849,153,176]" italics="true" pageId="5" pageNumber="197">V</emphasis>
|
||
-shaped; (dorsal-fin lock present and functional); posteriormost dorsal-fin ray free from body. Pectoral fin I,5(2)*, I,6(2); adpressed pectoral-fin spine reaching approximately to anus, spine having hypertrophied odontodes increasing in length distally. Pelvic fin I,5; pelvic-fin spine extending to or past insertion of anal fin when adpressed. Anal fin I,4; second unbranched ray longest. Adipose-fin spine slightly curved distally, adnate to caudal peduncle via fleshy membrane with concave or convex posterior margin. Caudal fin I,14,I; caudal fin asymmetrically emarginate with ventral lobe longer than dorsal lobe.
|
||
</paragraph>
|
||
<paragraph id="12593693FFB7FF96FF7DFEE5FE57E668" blockId="5.[151,1437,151,860]" pageId="5" pageNumber="197">Body broadest at cleithrum; posterior margin of exposed posterior process of cleithrum tapering to a point. Lateral median plates 24 or 25*, middorsal plates 23 or 24*, midventral plates 22–25*; anteriormost midventral plates strongly bent forming slight lateral ridge between verticals through pectoral- and dorsal-fin origins. Caudal peduncle plate rows five. One azygous preadipose plate; two predorsal plate rows not including nuchal plate; five interdorsal plate rows.</paragraph>
|
||
<paragraph id="12593693FFB7FF96FF7DFDA9FE62E6B3" blockId="5.[151,1437,151,860]" pageId="5" pageNumber="197">
|
||
<emphasis id="2092EA81FFB7FF96FF7DFDA9FED6E643" bold="true" box="[199,272,547,572]" pageId="5" pageNumber="197">Color.</emphasis>
|
||
Head, body, and fins with widely separated small white to yellow spots, vermiculations, and/or thin oblique bands on a black base. Several specimens have only thin widely-spaced, white to yellow oblique bands and lack spots entirely. Numbers of bands increase, and width of bands decrease, with body size, with juveniles having relatively few broad bands and adults having more numerous, thin bands and bands that are broken into vermiculations or spots.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5AFC6518FFB7FF96FF7DFD5DFD99E76B" pageId="5" pageNumber="197" type="distribution">
|
||
<paragraph id="12593693FFB7FF96FF7DFD5DFD99E76B" blockId="5.[151,1437,151,860]" pageId="5" pageNumber="197">
|
||
<emphasis id="2092EA81FFB7FF96FF7DFD5DFE31E68F" bold="true" box="[199,503,727,752]" pageId="5" pageNumber="197">Distribution and habitat.</emphasis>
|
||
Known only from the San Alejandro River, a tributary of Aguaytia River in the upper Ucayali watershed in central
|
||
<collectingCountry id="6AF17603FFB7FF96FD9EFD77FD9DE76B" box="[548,603,765,788]" name="Peru" pageId="5" pageNumber="197">Peru</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="5AFC6518FFB7FF96FF7DFCAAFB3BE723" pageId="5" pageNumber="197" type="etymology">
|
||
<paragraph id="12593693FFB7FF96FF7DFCAAFB3BE723" blockId="5.[151,1437,151,860]" pageId="5" pageNumber="197">
|
||
<emphasis id="2092EA81FFB7FF96FF7DFCAAFE88E747" bold="true" box="[199,334,800,824]" pageId="5" pageNumber="197">Etymology.</emphasis>
|
||
The specific epithet
|
||
<taxonomicName id="D5E64D10FFB7FF96FDFAFCAAFD70E747" box="[576,694,800,824]" class="Actinopterygii" family="Loricariidae" genus="Panaqolus" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="197" phylum="Chordata" rank="species" species="albivermis">
|
||
<emphasis id="2092EA81FFB7FF96FDFAFCAAFD70E747" box="[576,694,800,824]" italics="true" pageId="5" pageNumber="197">albivermis</emphasis>
|
||
</taxonomicName>
|
||
is a combination of the Latin
|
||
<emphasis id="2092EA81FFB7FF96FBAEFCAAFB96E747" box="[1044,1104,800,824]" italics="true" pageId="5" pageNumber="197">albus</emphasis>
|
||
, meaning white, and
|
||
<emphasis id="2092EA81FFB7FF96FAF0FCABFA53E747" box="[1354,1429,801,824]" italics="true" pageId="5" pageNumber="197">vermis</emphasis>
|
||
, meaning worm, and is in reference to this species’ variable but distinctive white to yellow markings.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |