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<document id="E82957C7439424B37B79DA7E9455D3A6" ID-DOI="10.11646/zootaxa.3918.1.5" ID-GBIF-Dataset="d557df68-c126-4906-b52d-a429d87914b7" ID-ISSN="1175-5326" ID-Zenodo-Dep="234081" ID-ZooBank="392562D1-8772-4F8D-885C-57CAA5B670C1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460980574430" checkinUser="plazi" docAuthor="Murányi, Dávid, Li, Weihai, Jeon, Mi Jeong, Hwang, Jeong Mi &amp; Seo, Hong Yul" docDate="2015" docId="9A3687BC5D2E560DFF768B99FB43B613" docLanguage="en" docName="zt03918p127.pdf" docOrigin="Zootaxa 3918 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Neoperla quadrata Wu &amp; Claassen 1934" docType="treatment" docVersion="8" lastPageNumber="122" masterDocId="660FFFC45D275604FFE18B0EFF92B050" masterDocTitle="Korean species of the genus Neoperla Needham, 1905 (Plecoptera: Perlidae) *" masterLastPageNumber="127" masterPageNumber="113" pageNumber="122" updateTime="1698560147113" updateUser="plazi">
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<mods:title id="8E5F829CFB54D5C72DCA523B3493EAE8">Korean species of the genus Neoperla Needham, 1905 (Plecoptera: Perlidae) *</mods:title>
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<mods:namePart id="63393919DA91B0F61D442B8586ED5ADD">Murányi, Dávid</mods:namePart>
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<mods:namePart id="4A63FA26886D23A15D9302F8303B2CCD">Li, Weihai</mods:namePart>
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<mods:namePart id="1A86B83E79D7827A1F6FE8212E39A753">Jeon, Mi Jeong</mods:namePart>
</mods:name>
<mods:name id="574985E1A3D1E79DFE0ABD070ABCE3F3" type="personal">
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<mods:namePart id="CC0909C4EDFE07D8860810010C8A5541">Hwang, Jeong Mi</mods:namePart>
</mods:name>
<mods:name id="39A06C435BA09A0A1CDB0C67F8AA17BD" type="personal">
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<mods:namePart id="E27E47294ECB83AAFD10734A92D482AE">Seo, Hong Yul</mods:namePart>
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<mods:date id="9C12F530BCB9A26CB1CCEF8A33D95FE3">2015</mods:date>
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<mods:number id="E7C3F55F7973510654E1D5EA09A0583D">3918</mods:number>
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<treatment id="9A3687BC5D2E560DFF768B99FB43B613" ID-DOI="http://doi.org/10.5281/zenodo.6095856" ID-GBIF-Taxon="119594423" ID-Zenodo-Dep="6095856" LSID="urn:lsid:plazi:treatment:9A3687BC5D2E560DFF768B99FB43B613" httpUri="http://treatment.plazi.org/id/9A3687BC5D2E560DFF768B99FB43B613" lastPageNumber="122" pageId="9" pageNumber="122">
<subSubSection id="5A8565215D2E560DFF768B99FE10B083" pageId="9" pageNumber="122" type="nomenclature">
<paragraph id="122036AA5D2E560DFF768B99FD1FB0E2" blockId="9.[151,653,151,211]" box="[151,653,151,178]" pageId="9" pageNumber="122">
<heading id="496881C65D2E560DFF768B99FD1FB0E2" bold="true" box="[151,653,151,178]" fontSize="11" level="1" pageId="9" pageNumber="122" reason="1">
<taxonomicName id="D59F4D295D2E560DFF768B99FD1FB0E2" ID-CoL="46PRV" authority="Wu &amp; Claassen, 1934" authorityName="Wu &amp; Claassen" authorityYear="1934" box="[151,653,151,178]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="122" phylum="Arthropoda" rank="species" species="quadrata">
<emphasis id="20EBEAB85D2E560DFF768B99FD1FB0E2" bold="true" box="[151,653,151,178]" pageId="9" pageNumber="122">
<emphasis id="20EBEAB85D2E560DFF768B99FEE8B0E1" bold="true" box="[151,378,151,177]" italics="true" pageId="9" pageNumber="122">Neoperla quadrata</emphasis>
<bibRefCitation id="760E4B5B5D2E560DFE608B96FD1FB0E2" author="Wu" box="[385,653,152,178]" pageId="9" pageNumber="122" refString="Wu, C. F. &amp; Claassen, P. W. (1934) Aquatic Insects of China. Article XVIII. New species of Chinese stoneflies (Order Plecoptera). Peking Natural History Bulletin, 9 (2), 111 - 129." type="journal article" year="1934">Wu &amp; Claassen, 1934</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="122036AA5D2E560DFF768BB4FE10B083" blockId="9.[151,653,151,211]" box="[151,386,186,211]" pageId="9" pageNumber="122">
(
<figureCitation id="8AA42A2F5D2E560DFF7E8BB4FEB5B083" box="[159,295,186,211]" captionStart="FIGURES 34 37" captionStartId="10.[151,264,1906,1929]" captionTargetBox="[151,1436,196,1885]" captionTargetId="figure@10.[151,1436,193,1885]" captionTargetPageId="10" captionText="FIGURES 34 37. Holotype of Neoperla quadrata Wu &amp; Claassen, 1934; China, Sichuan Province, Lucheng — 34: aedeagal sac, lateral view; 35: aedeagal sac, ventral view; 36: aedeagal sac, dorsal view; 37: aedeagus, lateral view—not to scale." httpUri="https://zenodo.org/record/234087/files/figure.png" pageId="9" pageNumber="122">Figs. 3437</figureCitation>
,
<figureCitation id="8AA42A2F5D2E560DFED38BB5FEEBB083" box="[306,377,187,211]" captionStart="FIGURES 41 42" captionStartId="12.[151,264,1508,1531]" captionTargetBox="[154,1426,453,1486]" captionTargetId="figure@12.[154,1427,453,1486]" captionTargetPageId="12" captionText="FIGURES 41 42. Distribution of the Neoperla Needham, 1905 species known from Korea — 41: known occurrences in North and South Korea; 42: known range of N. quadrata Wu &amp; Claassen, 1934 and N. ussurica Sivec &amp; Zhiltzova, 1996." httpUri="https://zenodo.org/record/234089/files/figure.png" pageId="9" pageNumber="122">4142</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5A8565215D2E560DFF768BF0FDE5B162" pageId="9" pageNumber="122" type="materials_examined">
<paragraph id="122036AA5D2E560DFF768BF0FDE5B162" blockId="9.[151,1436,253,459]" pageId="9" pageNumber="122">
<typeStatus id="CD2488085D2E560DFF768BF0FF5BB143" box="[151,201,254,275]" pageId="9" pageNumber="122">Type</typeStatus>
locality:
<collectingCountry id="6A88763A5D2E560DFED98BF3FEE5B143" box="[312,375,253,275]" name="China" pageId="9" pageNumber="122">China</collectingCountry>
, Szechuan, Tatsienlu,
<quantity id="D5679B4F5D2E560DFD8F8BF3FC86B143" box="[622,788,253,275]" metricMagnitude="3" metricUnit="m" metricValue="2.5908" metricValueMax="2.7432" metricValueMin="2.4384" pageId="9" pageNumber="122" unit="ft" value="8500.0" valueMax="9000.0" valueMin="8000.0">80009000 feet</quantity>
(Sichuan Province, Garzê Tibetan Autonomous Prefecture, Kangding County, Lucheng, N30° E102°).
</paragraph>
</subSubSection>
<subSubSection id="5A8565215D2E560DFF768A34FCCAB2B3" pageId="9" pageNumber="122" type="reference_group">
<paragraph id="122036AA5D2E560DFF768A34FEBAB19B" blockId="9.[151,1436,253,459]" pageId="9" pageNumber="122">
<treatmentCitationGroup id="328F11845D2E560DFF768A34FEBAB19B" pageId="9" pageNumber="122">
<taxonomicName id="D59F4D295D2E560DFF768A34FDDFB100" ID-CoL="46PRV" authority="Wu &amp; Claassen, 1934" authorityName="Wu &amp; Claassen" authorityYear="1934" box="[151,589,314,336]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="122" phylum="Arthropoda" rank="species" species="quadrata">
<emphasis id="20EBEAB85D2E560DFF768A34FECEB100" box="[151,348,314,336]" italics="true" pageId="9" pageNumber="122">Neoperla quadrata</emphasis>
<bibRefCitation id="760E4B5B5D2E560DFE868A35FDDFB100" author="Wu" box="[359,589,314,336]" pageId="9" pageNumber="122" refString="Wu, C. F. &amp; Claassen, P. W. (1934) Aquatic Insects of China. Article XVIII. New species of Chinese stoneflies (Order Plecoptera). Peking Natural History Bulletin, 9 (2), 111 - 129." type="journal article" year="1934">Wu &amp; Claassen, 1934</bibRefCitation>
</taxonomicName>
<treatmentCitation id="933E10BB5D2E560DFD898A34FC15B100" author="Wu" box="[616,903,314,336]" page="124" pageId="9" pageNumber="122" year="1934">
<bibRefCitation id="760E4B5B5D2E560DFD898A34FCC2B100" author="Wu" box="[616,848,314,336]" pageId="9" pageNumber="122" refString="Wu, C. F. &amp; Claassen, P. W. (1934) Aquatic Insects of China. Article XVIII. New species of Chinese stoneflies (Order Plecoptera). Peking Natural History Bulletin, 9 (2), 111 - 129." type="journal article" year="1934">Wu &amp; Claassen 1934</bibRefCitation>
: 124
</treatmentCitation>
. (original description of the male);
<treatmentCitation id="933E10BB5D2E560DFAE48A35FA0EB100" author="Wu" box="[1285,1436,314,336]" page="117" pageId="9" pageNumber="122" year="1938">
<bibRefCitation id="760E4B5B5D2E560DFAE48A35FAFBB100" author="Wu" box="[1285,1385,314,336]" pageId="9" pageNumber="122" refString="Wu, C. F. (1938) Plecopterorum sinensium: A monograph of stoneflies of China (Order Plecoptera). Yenching University, Beijing, 225 pp. [plates I - XLVII]" type="book" year="1938">Wu 1938</bibRefCitation>
: 117
</treatmentCitation>
(monography, with further details of the aedeagus);
<treatmentCitation id="933E10BB5D2E560DFD368A57FC33B13F" author="Claassen" box="[727,929,345,367]" page="165" pageId="9" pageNumber="122" year="1940">
<bibRefCitation id="760E4B5B5D2E560DFD368A57FCFCB13F" author="Claassen" box="[727,878,345,367]" pageId="9" pageNumber="122" refString="Claassen, P. W. (1940) A Catalogue of the Plecoptera of the World. Memoirs of the Cornell University Agricultural Experiment Station, Ithaca, 232, 1 - 235." type="journal article" year="1940">Claassen 1940</bibRefCitation>
: 165
</treatmentCitation>
. (catalog);
<treatmentCitation id="933E10BB5D2E560DFBF28A57FB73B13F" author="Kimmins" box="[1043,1249,345,367]" page="195" pageId="9" pageNumber="122" year="1950">
<bibRefCitation id="760E4B5B5D2E560DFBF28A57FB3CB13F" author="Kimmins" box="[1043,1198,345,367]" pageId="9" pageNumber="122" refString="Kimmins, D. E. (1950) Some Assamese Plecoptera, with descriptions of new species of Nemouridae. The Annals and Magazine of Natural History, 3 (13), 194 - 209. http: // dx. doi. org / 10.1080 / 00222935008654707" type="journal article" year="1950">Kimmins 1950</bibRefCitation>
: 195
</treatmentCitation>
. (first record from
<collectingCountry id="6A88763A5D2E560DFF268A76FF6CB1DE" box="[199,254,376,398]" name="India" pageId="9" pageNumber="122">India</collectingCountry>
, Meghalaya State);
<treatmentCitation id="933E10BB5D2E560DFE328A76FDE8B1DE" author="Illies" box="[467,634,376,398]" page="276" pageId="9" pageNumber="122" year="1966">
<bibRefCitation id="760E4B5B5D2E560DFE328A76FDD4B1DE" author="Illies" box="[467,582,376,398]" pageId="9" pageNumber="122" refString="Illies, J. (1966) Katalog der rezenten Plecoptera. Das Tierreich, Berlin, 82, 1 - 632." type="journal article" year="1966">Illies 1966</bibRefCitation>
: 276
</treatmentCitation>
(catalog);
<treatmentCitation id="933E10BB5D2E560DFD108A76FC26B1DE" author="Zwick" box="[753,948,376,398]" page="153" pageId="9" pageNumber="122" year="1973">
<bibRefCitation id="760E4B5B5D2E560DFD108A76FCECB1DE" author="Zwick" box="[753,894,376,398]" pageId="9" pageNumber="122" refString="Zwick, P. (1973 a) On the stoneflies from Korea (Insecta, Plecoptera). Fragmenta Faunistica, 19 (8), 149 - 157. http: // dx. doi. org / 10.3161 / 00159301 FF 1973.19.8.149" type="journal article" year="1973" yearSuffix="a">Zwick 1973a</bibRefCitation>
: 153
</treatmentCitation>
. (first records from
<collectingCountry id="6A88763A5D2E560DFB6E8A76FA87B1DE" box="[1167,1301,376,398]" name="North Korea" pageId="9" pageNumber="122">North Korea</collectingCountry>
, notes about variability of males);
<treatmentCitation id="933E10BB5D2E560DFE408A99FD1BB1FC" author="Yoon" box="[417,649,406,428]" page="125" pageId="9" pageNumber="122" year="1985">
<bibRefCitation id="760E4B5B5D2E560DFE408A99FDC4B1FC" author="Yoon" box="[417,598,406,428]" pageId="9" pageNumber="122" refString="Yoon, I. B. &amp; Aw, S. J. (1985) A taxonomic study on the stonefly (Plecoptera) nymphs of Korea (I). Suborder Holognatha and Systellognatha. Entomological Research Bulletin, 11, 111 - 139." type="journal article" year="1985">Yoon &amp; Aw 1985</bibRefCitation>
: 125
</treatmentCitation>
. (first records from
<collectingCountry id="6A88763A5D2E560DFCB38A98FC41B1FC" box="[850,979,406,428]" name="South Korea" pageId="9" pageNumber="122">South Korea</collectingCountry>
, description of the larva);
<bibRefCitation id="760E4B5B5D2E560DFB3C8A98FA0EB1FC" author="DeWalt" box="[1245,1436,406,429]" pageId="9" pageNumber="122" refString="DeWalt, R. E., Maehr, M. D., Neu-Becker, U. &amp; Stueber, G. (2014) Plecoptera Species File Online. Version 5.0 / 5.0. Available from: http: // Plecoptera. Species. File. org (accessed 16 November 2014)" type="journal article" year="2014">
DeWalt
<emphasis id="20EBEAB85D2E560DFAD18A96FAF6B1FC" box="[1328,1380,406,429]" italics="true" pageId="9" pageNumber="122">et al.</emphasis>
2014
</bibRefCitation>
(catalog).
</treatmentCitationGroup>
</paragraph>
<paragraph id="122036AA5D2E560DFF768AFCFC14B203" blockId="9.[151,1437,498,1603]" pageId="9" pageNumber="122">
<emphasis id="20EBEAB85D2E560DFF768AFCFEC9B25B" bold="true" box="[151,347,498,523]" pageId="9" pageNumber="122">Literature data.</emphasis>
<bibRefCitation id="760E4B5B5D2E560DFE898AFCFD8AB25B" author="Zwick" box="[360,536,498,523]" pageId="9" pageNumber="122" refString="Zwick, P. (1973 a) On the stoneflies from Korea (Insecta, Plecoptera). Fragmenta Faunistica, 19 (8), 149 - 157. http: // dx. doi. org / 10.3161 / 00159301 FF 1973.19.8.149" type="journal article" year="1973" yearSuffix="a">Zwick (1973a)</bibRefCitation>
:
<collectingCountry id="6A88763A5D2E560DFDC68AFCFD2EB25A" box="[551,700,498,523]" name="North Korea" pageId="9" pageNumber="122">North Korea</collectingCountry>
, Mjohjang-san Mts. (North Pyongan Province, Hyangsan-gun, Myohyang Mts),
<date id="6621106A5D2E560DFE848918FE7EB27E" box="[357,492,534,558]" pageId="9" pageNumber="122" value="1959-08-05">5.VIII.1959</date>
, leg. B. Pisarski, J. Proszyński:
<specimenCount id="0499FD235D2E560DFC958919FC08B27F" box="[884,922,535,559]" pageId="9" pageNumber="122" type="generic">1♂</specimenCount>
;
<collectingCountry id="6A88763A5D2E560DFC4D8918FBD3B27E" box="[940,1089,534,558]" name="North Korea" pageId="9" pageNumber="122">North Korea</collectingCountry>
, Jongsǒng (North Hamgyong Province, Kyongsong County),
<date id="6621106A5D2E560DFE1B8934FDEEB203" box="[506,636,570,595]" pageId="9" pageNumber="122" value="1954-06-01">01.VI.1954</date>
, leg. H.R. Bullock:
<specimenCount id="0499FD235D2E560DFCBA8935FC13B203" box="[859,897,571,595]" pageId="9" pageNumber="122" type="generic">1♂</specimenCount>
.
</paragraph>
<paragraph id="122036AA5D2E560DFF768951FCCAB2B3" blockId="9.[151,1437,498,1603]" pageId="9" pageNumber="122">
<bibRefCitation id="760E4B5B5D2E560DFF768951FEECB227" author="Yoon" box="[151,382,606,631]" pageId="9" pageNumber="122" refString="Yoon, I. B. &amp; Aw, S. J. (1985) A taxonomic study on the stonefly (Plecoptera) nymphs of Korea (I). Suborder Holognatha and Systellognatha. Entomological Research Bulletin, 11, 111 - 139." type="journal article" year="1985">Yoon &amp; Aw (1985)</bibRefCitation>
:
<collectingCountry id="6A88763A5D2E560DFE6C8950FDB2B226" box="[397,544,606,631]" name="South Korea" pageId="9" pageNumber="122">South Korea</collectingCountry>
, Kangwon-do, Wonsǒng, Chiaksan (Gangwon Province, Wonju City, Chiak Mts),
<date id="6621106A5D2E560DFF3E898DFE46B2CB" box="[223,468,642,667]" pageId="9" pageNumber="122" value="1975-07-31" valueMax="1975-08-01" valueMin="1975-07-31">31.VII.01.VIII.1975</date>
: larvae;
<collectingCountry id="6A88763A5D2E560DFDDC898CFD40B2CA" box="[573,722,642,667]" name="South Korea" pageId="9" pageNumber="122">South Korea</collectingCountry>
, Kyǒnggi-do, Kapyǒng, Chojongchǒn (Gyeonggi Province, Gapyeong County),
<date id="6621106A5D2E560DFE7689A9FD9EB2EF" box="[407,524,679,703]" pageId="9" pageNumber="122" value="1973-05-27">27.V.1973</date>
,
<date id="6621106A5D2E560DFDC789A9FD34B2EF" box="[550,678,679,703]" pageId="9" pageNumber="122" value="1983-04-29">29.IV.1983</date>
: larvae;
<collectingCountry id="6A88763A5D2E560DFCFE89A8FC2FB2EE" box="[799,957,678,703]" name="South Korea" pageId="9" pageNumber="122">South Korea</collectingCountry>
, Chungchǒngbuk-do, Okchǒn (North Chungcheong Province, Okcheon County),
<date id="6621106A5D2E560DFD6089C5FC93B2B3" box="[641,769,715,739]" pageId="9" pageNumber="122" value="1972-04-09">09.IV.1972</date>
: larvae.
</paragraph>
</subSubSection>
<subSubSection id="5A8565215D2E560DFF2689E0FC16B43F" pageId="9" pageNumber="122" type="materials_examined">
<paragraph id="122036AA5D2E560DFF2689E0FE27B31F" blockId="9.[151,1437,498,1603]" pageId="9" pageNumber="122">
<emphasis id="20EBEAB85D2E560DFF2689E0FE2BB357" bold="true" box="[199,441,750,775]" pageId="9" pageNumber="122">Material examined.</emphasis>
<collectingCountry id="6A88763A5D2E560DFE2B89E1FDB5B357" box="[458,551,751,775]" name="China" pageId="9" pageNumber="122">CHINA</collectingCountry>
: Szechuan, Tatsienlu,
<quantity id="D5679B4F5D2E560DFCDD89E1FC66B357" box="[828,1012,750,775]" metricMagnitude="3" metricUnit="m" metricValue="2.5908" metricValueMax="2.7432" metricValueMin="2.4384" pageId="9" pageNumber="122" unit="ft" value="8500.0" valueMax="9000.0" valueMin="8000.0">80009000 feet</quantity>
(Sichuan Province, Garzê Tibetan Autonomous Prefecture, Kangding County, Lucheng, N30° E102°),
<date id="6621106A5D2E560DFC41881CFBA7B37B" box="[928,1077,786,811]" pageId="9" pageNumber="122" value="1930-08-16">16.VIII.1930</date>
, leg. D.C. Graham:
<typeStatus id="CD2488085D2E560DFAFD881CFAECB37B" box="[1308,1406,786,811]" pageId="9" pageNumber="122" type="holotype">holotype</typeStatus>
♂ and
<typeStatus id="CD2488085D2E560DFF278839FEB5B31F" box="[198,295,823,847]" pageId="9" pageNumber="122" type="paratype">paratype</typeStatus>
♂ (
<collectionCode id="748EAE6F5D2E560DFEB38839FE3BB31F" LSID="urn:lsid:biocol.org:col:34871" box="[338,425,823,847]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="122">USNM</collectionCode>
).
</paragraph>
<paragraph id="122036AA5D2E560DFF268854FC16B43F" blockId="9.[151,1437,498,1603]" pageId="9" pageNumber="122">
<emphasis id="20EBEAB85D2E560DFF268854FD8FB323" bold="true" box="[199,541,858,883]" pageId="9" pageNumber="122">Complementary description.</emphasis>
Aedeagus (
<figureCitation id="8AA42A2F5D2E560DFD408854FCB9B323" box="[673,811,858,883]" captionStart="FIGURES 34 37" captionStartId="10.[151,264,1906,1929]" captionTargetBox="[151,1436,196,1885]" captionTargetId="figure@10.[151,1436,193,1885]" captionTargetPageId="10" captionText="FIGURES 34 37. Holotype of Neoperla quadrata Wu &amp; Claassen, 1934; China, Sichuan Province, Lucheng — 34: aedeagal sac, lateral view; 35: aedeagal sac, ventral view; 36: aedeagal sac, dorsal view; 37: aedeagus, lateral view—not to scale." httpUri="https://zenodo.org/record/234087/files/figure.png" pageId="9" pageNumber="122">Figs. 3437</figureCitation>
): The aedeagal tube plump, with a triangular dorsobasal sclerite, a ventrobasal hump and two large, spherical ventroapical lobes. The ventroapical lobes close set but not touching each other ventrally, and bearing large spines on most of their surface, while anteriorly without spines. Tube mostly bald other than spines on ventroapical lobes. The everted aedeagal sac curved dorsad, hardly longer than the tube. Sac covered with fine spinules from the basal ⅓ up to the apex, and bearing three fields of larger spines: a basodorsal, triangular field of moderately large spines, a longitudinal ventral row of large spines that reach to the apex from the basal ⅓, and a large, spherical dorsal patch of strong spines on the bulging subapical section. The apex is slightly separated, straight and without spines.
</paragraph>
</subSubSection>
<subSubSection id="5A8565215D2E560DFF268F74FC31B587" pageId="9" pageNumber="122" type="discussion">
<paragraph id="122036AA5D2E560DFF268F74FC31B587" blockId="9.[151,1437,498,1603]" pageId="9" pageNumber="122">
<emphasis id="20EBEAB85D2E560DFF268F74FEA9B4C3" bold="true" box="[199,315,1146,1171]" pageId="9" pageNumber="122">Remarks.</emphasis>
Habitus, terminalia and uneverted aedeagus of this species are similar to several other Asian
<taxonomicName id="D59F4D295D2E560DFF768F90FF6DB4E6" box="[151,255,1182,1206]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="20EBEAB85D2E560DFF768F90FF6DB4E6" box="[151,255,1182,1206]" italics="true" pageId="9" pageNumber="122">Neoperla</emphasis>
</taxonomicName>
. Details of the aedeagal sac have not been previously studied. The aedeagus of the
<typeStatus id="CD2488085D2E560DFB428F90FA97B4E7" box="[1187,1285,1182,1207]" pageId="9" pageNumber="122" type="holotype">holotype</typeStatus>
and
<typeStatus id="CD2488085D2E560DFADD8F91FA0EB4E7" box="[1340,1436,1183,1207]" pageId="9" pageNumber="122" type="paratype">paratype</typeStatus>
are not everted. The
<typeStatus id="CD2488085D2E560DFE6F8FCDFE7CB48B" box="[398,494,1219,1243]" pageId="9" pageNumber="122" type="paratype">paratype</typeStatus>
is not conspecific with the
<typeStatus id="CD2488085D2E560DFCDD8FCCFC0CB48B" box="[828,926,1218,1243]" pageId="9" pageNumber="122" type="holotype">holotype</typeStatus>
and most probably belongs to an unnamed species; thus, our description refers only to the
<typeStatus id="CD2488085D2E560DFD328FE8FCABB4AF" box="[723,825,1254,1279]" pageId="9" pageNumber="122" type="holotype">holotype</typeStatus>
. It is closely related to certain other species of the
<emphasis id="20EBEAB85D2E560DFF768E05FE9FB572" box="[151,269,1291,1314]" italics="true" pageId="9" pageNumber="122">montivaga</emphasis>
group,
<taxonomicName id="D59F4D295D2E560DFE918E05FE5BB573" box="[368,457,1291,1315]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="122" phylum="Arthropoda" rank="species" species="lushana">
<emphasis id="20EBEAB85D2E560DFE918E05FE5BB573" box="[368,457,1291,1315]" italics="true" pageId="9" pageNumber="122">lushana</emphasis>
</taxonomicName>
subgroup, especially to
<taxonomicName id="D59F4D295D2E560DFD158E05FC6BB573" authority="Wu, 1935" authorityName="Wu" authorityYear="1935" box="[756,1017,1290,1315]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="122" phylum="Arthropoda" rank="species" species="lushana">
<emphasis id="20EBEAB85D2E560DFD158E05FCE7B573" box="[756,885,1291,1315]" italics="true" pageId="9" pageNumber="122">N. lushana</emphasis>
<bibRefCitation id="760E4B5B5D2E560DFC638E05FC6BB573" author="Wu" box="[898,1017,1290,1315]" pageId="9" pageNumber="122" refString="Wu, C. F. (1935) Aquatic Insects of China. Article XXI. New species of stoneflies from East and South China (Order Plecoptera). Peking Natural History Bulletin, 9 (3), 227 - 243." type="journal article" year="1935">Wu, 1935</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="D59F4D295D2E560DFBEB8E05FAADB573" authority="Banks, 1937" authorityName="Banks" authorityYear="1937" box="[1034,1343,1290,1315]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="122" phylum="Arthropoda" rank="species" species="signatalis">
<emphasis id="20EBEAB85D2E560DFBEB8E05FB0DB573" box="[1034,1183,1291,1315]" italics="true" pageId="9" pageNumber="122">N. signatalis</emphasis>
<bibRefCitation id="760E4B5B5D2E560DFB4D8E04FAADB573" author="Banks" box="[1196,1343,1290,1315]" pageId="9" pageNumber="122" refString="Banks, N. (1937) Neuropteroid insects from Formosa. Philippine Journal of Science, 62, 255 - 291. [plates I - III]" type="journal article" year="1937">Banks, 1937</bibRefCitation>
</taxonomicName>
and
<taxonomicName id="D59F4D295D2E560DFA638E05FE46B517" authority="Li &amp; Li, 2013" authorityName="Li &amp; Li" authorityYear="2013" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="122" phylum="Arthropoda" rank="species" species="xuoansongae">
<emphasis id="20EBEAB85D2E560DFA638E05FEB9B516" italics="true" pageId="9" pageNumber="122">N. xuoansongae</emphasis>
<bibRefCitation id="760E4B5B5D2E560DFED28E20FE46B517" author="Li" box="[307,468,1326,1351]" pageId="9" pageNumber="122" refString="Li, W. H., Liang, H. Y. &amp; Li, W. L. (2013) Review of Neoperla (Plecoptera: Perlidae) from Zhejiang Province, China. Zootaxa, 3652 (3), 353 - 369. http: // dx. doi. org / 10.11646 / zootaxa. 3652.3.4" type="journal article" year="2013">Li &amp; Li, 2013</bibRefCitation>
</taxonomicName>
(in
<bibRefCitation id="760E4B5B5D2E560DFDE48E20FD35B517" author="Li" box="[517,679,1326,1351]" pageId="9" pageNumber="122" refString="Li, W. H., Liang, H. Y. &amp; Li, W. L. (2013) Review of Neoperla (Plecoptera: Perlidae) from Zhejiang Province, China. Zootaxa, 3652 (3), 353 - 369. http: // dx. doi. org / 10.11646 / zootaxa. 3652.3.4" type="journal article" year="2013">
Li
<emphasis id="20EBEAB85D2E560DFDC68E21FDF2B516" box="[551,608,1326,1350]" italics="true" pageId="9" pageNumber="122">et al.</emphasis>
2013
</bibRefCitation>
).
<taxonomicName id="D59F4D295D2E560DFD5A8E21FCD6B516" box="[699,836,1326,1350]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="122" phylum="Arthropoda" rank="species" species="quadrata">
<emphasis id="20EBEAB85D2E560DFD5A8E21FCD6B516" box="[699,836,1326,1350]" italics="true" pageId="9" pageNumber="122">N. quadrata</emphasis>
</taxonomicName>
differs from all those species by the combination of ventral row and spherical dorsal patch of strong spines on the aedeagal sac. Literature data from
<collectingCountry id="6A88763A5D2E560DFB1C8E5DFAD3B53A" box="[1277,1345,1363,1386]" name="South Korea" pageId="9" pageNumber="122">Korea</collectingCountry>
(
<bibRefCitation id="760E4B5B5D2E560DFAB28E5CFF4CB5DF" author="Zwick" pageId="9" pageNumber="122" refString="Zwick, P. (1973 a) On the stoneflies from Korea (Insecta, Plecoptera). Fragmenta Faunistica, 19 (8), 149 - 157. http: // dx. doi. org / 10.3161 / 00159301 FF 1973.19.8.149" type="journal article" year="1973" yearSuffix="a">Zwick 1973a</bibRefCitation>
,
<bibRefCitation id="760E4B5B5D2E560DFF0A8E79FE25B5DF" author="Yoon" box="[235,439,1398,1423]" pageId="9" pageNumber="122" refString="Yoon, I. B. &amp; Aw, S. J. (1985) A taxonomic study on the stonefly (Plecoptera) nymphs of Korea (I). Suborder Holognatha and Systellognatha. Entomological Research Bulletin, 11, 111 - 139." type="journal article" year="1985">Yoon &amp; Aw 1985</bibRefCitation>
) and
<collectingCountry id="6A88763A5D2E560DFE188E78FDA1B5DF" box="[505,563,1398,1423]" name="India" pageId="9" pageNumber="122">India</collectingCountry>
(
<bibRefCitation id="760E4B5B5D2E560DFDA28E78FD7DB5DF" author="Kimmins" box="[579,751,1398,1423]" pageId="9" pageNumber="122" refString="Kimmins, D. E. (1950) Some Assamese Plecoptera, with descriptions of new species of Nemouridae. The Annals and Magazine of Natural History, 3 (13), 194 - 209. http: // dx. doi. org / 10.1080 / 00222935008654707" type="journal article" year="1950">Kimmins 1950</bibRefCitation>
) are in need of validation. Unfortunately, the North Korean male specimens deposited in the Polish Academy of Sciences was not available for study (Tomas Huflejt pers. com.). Therefore, all records from
<collectingCountry id="6A88763A5D2E560DFDFD8EB0FD3AB586" box="[540,680,1470,1495]" name="South Korea" pageId="9" pageNumber="122">South Korea</collectingCountry>
were based on larvae.
</paragraph>
</subSubSection>
<subSubSection id="5A8565215D2E560DFF268EECFB43B613" pageId="9" pageNumber="122" type="distribution">
<paragraph id="122036AA5D2E560DFF268EECFB43B613" blockId="9.[151,1437,498,1603]" pageId="9" pageNumber="122">
<emphasis id="20EBEAB85D2E560DFF268EECFE64B5AB" bold="true" box="[199,502,1506,1531]" pageId="9" pageNumber="122">Distribution and ecology.</emphasis>
The species was described from Sichuan Province of
<collectingCountry id="6A88763A5D2E560DFB848EECFB39B5AB" box="[1125,1195,1506,1531]" name="China" pageId="9" pageNumber="122">China</collectingCountry>
, later it was reported from Meghalaya State of
<collectingCountry id="6A88763A5D2E560DFE5D8D08FE67B64F" box="[444,501,1542,1567]" name="India" pageId="9" pageNumber="122">India</collectingCountry>
and from both North and
<collectingCountry id="6A88763A5D2E560DFCC58D08FC24B64E" box="[804,950,1542,1567]" name="South Korea" pageId="9" pageNumber="122">South Korea</collectingCountry>
. If the Indian and Korean data are correct, this species can be regarded as one of the most widely distributed Asian
<taxonomicName id="D59F4D295D2E560DFC228D25FBBEB613" box="[963,1068,1579,1603]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="9" pageNumber="122" phylum="Arthropoda" rank="genus">
<emphasis id="20EBEAB85D2E560DFC228D25FBBEB613" box="[963,1068,1579,1603]" italics="true" pageId="9" pageNumber="122">Neoperla</emphasis>
</taxonomicName>
(
<figureCitation id="8AA42A2F5D2E560DFBDA8D24FB57B613" box="[1083,1221,1578,1603]" captionStart="FIGURES 41 42" captionStartId="12.[151,264,1508,1531]" captionTargetBox="[154,1426,453,1486]" captionTargetId="figure@12.[154,1427,453,1486]" captionTargetPageId="12" captionText="FIGURES 41 42. Distribution of the Neoperla Needham, 1905 species known from Korea — 41: known occurrences in North and South Korea; 42: known range of N. quadrata Wu &amp; Claassen, 1934 and N. ussurica Sivec &amp; Zhiltzova, 1996." httpUri="https://zenodo.org/record/234089/files/figure.png" pageId="9" pageNumber="122">Figs. 4142</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>