treatments-xml/data/9A/36/87/9A3687BC5D225603FF768E43FD63B1A5.xml

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<mods:title id="B91D0075112EBD821C2DEB534BA2CB17">Korean species of the genus Neoperla Needham, 1905 (Plecoptera: Perlidae) *</mods:title>
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<mods:namePart id="EBF0B5D9D3758F5C1C3B111BEC637550">Murányi, Dávid</mods:namePart>
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<mods:namePart id="1E751E74184C460979C98001AE242B59">Li, Weihai</mods:namePart>
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<mods:namePart id="1E427F9844A22B6648100078F3547E7E">Jeon, Mi Jeong</mods:namePart>
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<mods:namePart id="A781D753110E2D180D39D05622578284">Hwang, Jeong Mi</mods:namePart>
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<mods:namePart id="4362CB94E74B457CDD682218209F93B7">Seo, Hong Yul</mods:namePart>
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<treatment id="9A3687BC5D225603FF768E43FD63B1A5" ID-DOI="http://doi.org/10.5281/zenodo.6095854" ID-GBIF-Taxon="119594426" ID-Zenodo-Dep="6095854" LSID="urn:lsid:plazi:treatment:9A3687BC5D225603FF768E43FD63B1A5" httpUri="http://treatment.plazi.org/id/9A3687BC5D225603FF768E43FD63B1A5" lastPageId="7" lastPageNumber="120" pageId="5" pageNumber="118">
<subSubSection id="5A8565215D225601FF768E43FE39B5D8" pageId="5" pageNumber="118" type="nomenclature">
<paragraph id="122036AA5D225601FF768E43FD15B537" blockId="5.[151,647,1357,1417]" box="[151,647,1357,1383]" pageId="5" pageNumber="118">
<heading id="496881C65D225601FF768E43FD15B537" bold="true" box="[151,647,1357,1383]" fontSize="11" level="1" pageId="5" pageNumber="118" reason="1">
<emphasis id="20EBEAB85D225601FF768E43FD15B537" bold="true" box="[151,647,1357,1383]" pageId="5" pageNumber="118">
<taxonomicName id="D59F4D295D225601FF768E43FDA5B537" ID-CoL="46PMG" authority="Muranyi &amp; Li" authorityName="Muranyi &amp; Li" box="[151,567,1357,1383]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="5" pageNumber="118" phylum="Arthropoda" rank="species" species="goguryeo" status="sp. nov.">
<emphasis id="20EBEAB85D225601FF768E43FEEFB537" bold="true" box="[151,381,1357,1383]" italics="true" pageId="5" pageNumber="118">Neoperla goguryeo</emphasis>
Murányi &amp; Li
</taxonomicName>
, 
<taxonomicNameLabel id="3BD857C35D225601FDA48E43FD15B537" box="[581,647,1357,1383]" pageId="5" pageNumber="118" rank="species">sp. n.</taxonomicNameLabel>
</emphasis>
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</paragraph>
<paragraph id="122036AA5D225601FF768E7EFE39B5D8" blockId="5.[151,647,1357,1417]" box="[151,427,1392,1417]" pageId="5" pageNumber="118">
(
<figureCitation id="8AA42A2F5D225601FF7E8E7EFEB5B5D8" box="[159,295,1392,1417]" captionStart-0="FIGURES 10 – 17" captionStart-1="FIGURES 18 – 23" captionStartId-0="5.[151,264,1137,1160]" captionStartId-1="7.[151,264,1639,1662]" captionTargetBox-0="[191,1368,246,1103]" captionTargetBox-1="[168,1409,566,1600]" captionTargetId-0="figure@5.[151,1436,193,1116]" captionTargetId-1="figure@7.[151,1436,555,1618]" captionTargetPageId-0="5" captionTargetPageId-1="7" captionText-0="FIGURES 10 – 17. Females of Korean Neoperla Needham, 1905 — 10, 12, 14, 16: vagina, dorsal view; 11, 13, 15, 17: vagina, lateral view— 10 – 11: Neoperla adamantea Murányi &amp; Li, sp. n., North Korea, Kangwon Province, Kumgang Mts; 12 – 13: Neoperla coreensis Ra, Kim, Kang &amp; Ham, 1994, North Korea, North Pyongan Province, Myohyang Mts; 14 – 15: Neoperla goguryeo Murányi &amp; Li, sp. n., North Korea, Kangwon Province, Kumgang Mts; 16 – 17: Neoperla goguryeo Murányi &amp; Li, sp. n., North Korea, North Pyongan Province, Myohyang Mts—scale 1 mm." captionText-1="FIGURES 18 – 23. Holotype male of Neoperla goguryeo Murányi &amp; Li, sp. n.; North Korea, Kangwon Province, Kumgang Mts — 18: head and pronotum, dorsal view; 19: terminalia, dorsal view; 20: terminalia, lateral view; 21: aedeagus, lateral view; 22: aedeagus, ventral view; 23: aedeagus, dorsal view—scales 1 mm." httpUri-0="https://zenodo.org/record/234084/files/figure.png" httpUri-1="https://zenodo.org/record/234085/files/figure.png" pageId="5" pageNumber="118">Figs. 14–23</figureCitation>
, 
<figureCitation id="8AA42A2F5D225601FED38E7EFEE9B5D8" box="[306,379,1392,1416]" captionStart="FIGURES 24 – 33" captionStartId="8.[151,264,1858,1881]" captionTargetBox="[151,1436,195,1836]" captionTargetId="figure@8.[151,1436,195,1837]" captionTargetPageId="8" captionText="FIGURES 24 – 33. Eggs of Korean Neoperla Needham, 1905 — 24, 29 – 30: egg, lateral view; 25, 32: micropyle, lateral view; 26: collar, caudal view; 27: lid, caudal view; 28: collar, oblique view; 31: lid, lateral view; 33: collar, lateral view— 24 – 28: Neoperla coreensis Ra, Kim, Kang &amp; Ham, 1994, North Korea, North Pyongan Province, Myohyang Mts; 29, 31 – 33: Neoperla goguryeo Murányi &amp; Li, sp. n., North Korea, Kangwon Province, Kumgang Mts; 30: Neoperla goguryeo Murányi &amp; Li, sp. n., North Korea, North Pyongan Province, Myohyang Mts—scales 0.1 mm." httpUri="https://zenodo.org/record/234086/files/figure.png" pageId="5" pageNumber="118">29–33</figureCitation>
, 
<figureCitation id="8AA42A2F5D225601FE678E7EFE31B5D8" box="[390,419,1392,1416]" captionStart="FIGURES 41 – 42" captionStartId="12.[151,264,1508,1531]" captionTargetBox="[154,1426,453,1486]" captionTargetId="figure@12.[154,1427,453,1486]" captionTargetPageId="12" captionText="FIGURES 41 – 42. Distribution of the Neoperla Needham, 1905 species known from Korea — 41: known occurrences in North and South Korea; 42: known range of N. quadrata Wu &amp; Claassen, 1934 and N. ussurica Sivec &amp; Zhiltzova, 1996." httpUri="https://zenodo.org/record/234089/files/figure.png" pageId="5" pageNumber="118">41</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="5A8565215D225601FF768EB6FDA5B6D4" pageId="5" pageNumber="118" type="diagnosis">
<paragraph id="122036AA5D225601FF768EB6FDA5B6D4" blockId="5.[151,1437,1464,2029]" pageId="5" pageNumber="118">
<emphasis id="20EBEAB85D225601FF768EB6FE83B581" bold="true" box="[151,273,1464,1489]" pageId="5" pageNumber="118">Diagnosis.</emphasis>
General colour pale, head with two dark patches, ocelli set far from each other. Male terminalia with large posterior process on T7, apically narrow and erect mesal sclerite on T8, T9 unmodified, T10 hemitergal process long. Aedeagal tube with a triangular dorsobasal sclerite, and with two large ventroapical lobes that bear large spines posteriorly. Aedeagal sac long and covered with fine spinules up to the apex, with a small dorsoapical lobe and three fields of strong spines at the level of the lobe. Vagina large and with numerous folds, inner lateral folds form a Y-shaped dorsal pattern.
</paragraph>
</subSubSection>
<subSubSection id="5A8565215D225602FF268D9EFC96B0A8" lastPageId="6" lastPageNumber="119" pageId="5" pageNumber="118" type="materials_examined">
<paragraph id="122036AA5D225602FF268D9EFC96B0A8" blockId="5.[151,1437,1464,2029]" lastBlockId="6.[151,1437,151,2013]" lastPageId="6" lastPageNumber="119" pageId="5" pageNumber="118">
<emphasis id="20EBEAB85D225601FF268D9EFEE4B6F9" bold="true" box="[199,374,1680,1705]" pageId="5" pageNumber="118">
<typeStatus id="CD2488085D225601FF268D9EFE90B6F9" box="[199,258,1680,1705]" pageId="5" pageNumber="118">Type</typeStatus>
material.
</emphasis>
<typeStatus id="CD2488085D225601FE9F8D9EFE74B6F9" box="[382,486,1680,1705]" pageId="5" pageNumber="118" type="holotype">Holotype</typeStatus>
male: 
<collectingCountry id="6A88763A5D225601FDD48D9EFC93B6F8" box="[565,769,1680,1704]" name="North Korea" pageId="5" pageNumber="118">NORTH KOREA</collectingCountry>
: Kangwon Province, Kosŏng-gun, Kumgang Mts, Onjongri, at light in mixed coniferous-locust tree forest by Hotel Go-song (locality No.322), 
<quantity id="D5679B4F5D225601FB7A8DBAFB7AB69C" box="[1179,1256,1716,1741]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="118" unit="m" value="250.0">250 m</quantity>
a.s.l., 
<geoCoordinate id="77AB506D5D225601FADA8DBBFA0FB69D" box="[1339,1437,1717,1741]" direction="north" orientation="latitude" pageId="5" pageNumber="118" precision="925" value="38.666668">N38°40’</geoCoordinate>
<geoCoordinate id="77AB506D5D225601FF768DD6FE97B6A1" box="[151,261,1752,1777]" direction="east" orientation="longitude" pageId="5" pageNumber="118" precision="925" value="128.25">E128°15’</geoCoordinate>
, 
<date id="6621106A5D225601FEF48DD6FE3BB6A1" box="[277,425,1752,1777]" pageId="5" pageNumber="118" value="1975-08-06">06.VIII.1975</date>
, leg. Jenő Papp, András Vojnits (
<collectionCode id="748EAE6F5D225601FCD48DD6FCEFB6A0" box="[821,893,1752,1776]" httpUri="http://grbio.org/cool/r4iq-k5jx" name="National Institute of Biological Resources" pageId="5" pageNumber="118">NIBR</collectionCode>
; terminalia and aedeagus cleared in KOH and stored in the same vial). 
<typeStatus id="CD2488085D225601FE4B8DF3FD9EB745" box="[426,524,1789,1813]" pageId="5" pageNumber="118" type="paratype">Paratype</typeStatus>
females: same locality, collecting method and collectors, 
<date id="6621106A5D225601FB748DF2FABAB745" box="[1173,1320,1788,1813]" pageId="5" pageNumber="118" value="1975-08-05">05.VIII.1975</date>
(No.319): 1♀ (
<collectionCode id="748EAE6F5D225601FF2E8C2EFE85B768" box="[207,279,1824,1848]" httpUri="http://grbio.org/cool/r4iq-k5jx" name="National Institute of Biological Resources" pageId="5" pageNumber="118">NIBR</collectionCode>
; eggs prepared for SEM, terminalia cleared in KOH and stored in the same vial), 1♀ (
<collectionCode id="748EAE6F5D225601FADC8C2EFA0BB768" LSID="urn:lsid:biocol.org:col:34991" box="[1341,1433,1824,1848]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="5" pageNumber="118">HNHM</collectionCode>
: PLP4349; terminalia cleared in KOH and stored in the same vial); North Pyongan Province, Hyangsan-gun, Myohyang Mts, at light on the balcony and garden of Hotel Myohyang-san (locality No. 829), 
<quantity id="D5679B4F5D225601FAE88C66FAC7B7D0" box="[1289,1365,1896,1921]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="118" unit="m" value="150.0">150 m</quantity>
a.s.l., 
<geoCoordinate id="77AB506D5D225601FF768C82FF65B7F4" box="[151,247,1932,1956]" direction="north" orientation="latitude" pageId="5" pageNumber="118" precision="925" value="40.0">N40°00’</geoCoordinate>
<geoCoordinate id="77AB506D5D225601FF1F8C82FEFEB7F4" box="[254,364,1932,1956]" direction="east" orientation="longitude" pageId="5" pageNumber="118" precision="925" value="126.25">E126°15’</geoCoordinate>
, 
<date id="6621106A5D225601FE998C82FD91B7F4" box="[376,515,1932,1956]" pageId="5" pageNumber="118" value="1982-07-18">18.VII.1982</date>
, leg. László Forró, László Ronkay: 1♀ (
<collectionCode id="748EAE6F5D225601FC278C82FB9CB7F4" box="[966,1038,1932,1956]" httpUri="http://grbio.org/cool/r4iq-k5jx" name="National Institute of Biological Resources" pageId="5" pageNumber="118">NIBR</collectionCode>
; eggs prepared for SEM, terminalia cleared in KOH and stored in the same vial); North Pyongan Province, Hyangsan-gun, Myohyang Mts, Hotel Myohyang-san, 
<quantity id="D5679B4F5D225601FEB18CDAFE08B7BC" box="[336,410,2004,2028]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="5" pageNumber="118" unit="m" value="150.0">150 m</quantity>
a.s.l., 
<geoCoordinate id="77AB506D5D225601FE098CDAFDDAB7BC" box="[488,584,2004,2028]" direction="north" orientation="latitude" pageId="5" pageNumber="118" precision="925" value="40.0">N40°00’</geoCoordinate>
<geoCoordinate id="77AB506D5D225601FDB28CDAFD53B7BC" box="[595,705,2004,2028]" direction="east" orientation="longitude" pageId="5" pageNumber="118" precision="925" value="126.25">E126°15’</geoCoordinate>
, 
<date id="6621106A5D225601FD2E8CDAFCF6B7BC" box="[719,868,2004,2028]" pageId="5" pageNumber="118" value="1989-08-12">12.VIII.1989</date>
, leg. Lajos Berczi: 1♀ (
<collectionCode id="748EAE6F5D225601FB9C8CDAFB57B7BC" box="[1149,1221,2004,2028]" httpUri="http://grbio.org/cool/r4iq-k5jx" name="National Institute of Biological Resources" pageId="5" pageNumber="118">NIBR</collectionCode>
; eggs prepared for SEM, terminalia cleared in KOH and stored in the same vial); North Pyongan Province, Hyangsan-gun, Myohyang Mts, N40° E126°, 
<date id="6621106A5D215602FE898BB2FDB5B084" box="[360,551,188,212]" pageId="6" pageNumber="119" value="1989-08-17" valueMax="1989-08-20" valueMin="1989-08-17">17–20.VIII.1989</date>
, leg. Konstantin Dobolyi, György Szollát: 1♀ (
<collectionCode id="748EAE6F5D215602FBDD8BB2FB16B084" box="[1084,1156,188,212]" httpUri="http://grbio.org/cool/r4iq-k5jx" name="National Institute of Biological Resources" pageId="6" pageNumber="119">NIBR</collectionCode>
; eggs prepared for SEM, terminalia cleared in KOH and stored in the same vial).
</paragraph>
</subSubSection>
<subSubSection id="5A8565215D215602FF268A0AFDF4B78C" pageId="6" pageNumber="119" type="description">
<paragraph id="122036AA5D215602FF268A0AFEBAB230" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="119">
<emphasis id="20EBEAB85D215602FF268A0AFECBB14D" bold="true" box="[199,345,260,285]" pageId="6" pageNumber="119">Description.</emphasis>
Medium sized species. Macropterous, forewing length of 
<typeStatus id="CD2488085D215602FBEB8A0AFBFEB14D" box="[1034,1132,260,285]" pageId="6" pageNumber="119" type="holotype">holotype</typeStatus>
male: 11.0 mm; 
<typeStatus id="CD2488085D215602FADD8A0BFA0EB14D" box="[1340,1436,261,285]" pageId="6" pageNumber="119" type="paratype">paratype</typeStatus>
females: 14.5–15.0 mm. General colour pale. Setation generally consists of soft hairs, strong setae are on ventral keels of femora and ventral surface of cercal segments. Head yellowish, with a triangular, dark brown patch between the ocelli and a smaller one anterior to the M-line (
<figureCitation id="8AA42A2F5D215602FCA28A61FC05B1D8" box="[835,919,367,392]" captionStart="FIGURES 18 – 23" captionStartId="7.[151,264,1639,1662]" captionTargetBox="[168,1409,566,1600]" captionTargetId="figure@7.[151,1436,555,1618]" captionTargetPageId="7" captionText="FIGURES 18 – 23. Holotype male of Neoperla goguryeo Murányi &amp; Li, sp. n.; North Korea, Kangwon Province, Kumgang Mts — 18: head and pronotum, dorsal view; 19: terminalia, dorsal view; 20: terminalia, lateral view; 21: aedeagus, lateral view; 22: aedeagus, ventral view; 23: aedeagus, dorsal view—scales 1 mm." httpUri="https://zenodo.org/record/234085/files/figure.png" pageId="6" pageNumber="119">Fig. 18</figureCitation>
). Tentorial callosities and M-line indistinct; a wrinkle is located between M-line and the lateral margins. Both eyes and ocelli are large and black, distance between ocelli about as wide as diameter of one ocellus. Antenna and palpi not much darker than head. Pronotum square, anterior edges distinctly angled; narrower than head with eyes; ground colour yellow, with a longitudinal brown stripe medially and dark transverse anterior and posterior lines; rugosities large but obscure. Meso- and metanotum brownish. Legs pale, knees and tarsi darker; tibiae slightly dilated. Wings hyaline, costa yellow, other veins brown.
</paragraph>
<paragraph id="122036AA5D215602FF268962FC3CB3D0" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="119">
Male terminalia (
<figureCitation id="8AA42A2F5D215602FE6F8962FD89B2D4" box="[398,539,620,645]" captionStart="FIGURES 18 – 23" captionStartId="7.[151,264,1639,1662]" captionTargetBox="[168,1409,566,1600]" captionTargetId="figure@7.[151,1436,555,1618]" captionTargetPageId="7" captionText="FIGURES 18 – 23. Holotype male of Neoperla goguryeo Murányi &amp; Li, sp. n.; North Korea, Kangwon Province, Kumgang Mts — 18: head and pronotum, dorsal view; 19: terminalia, dorsal view; 20: terminalia, lateral view; 21: aedeagus, lateral view; 22: aedeagus, ventral view; 23: aedeagus, dorsal view—scales 1 mm." httpUri="https://zenodo.org/record/234085/files/figure.png" pageId="6" pageNumber="119">Figs. 19–20</figureCitation>
): Sterna simple, S9 broadly truncate. Terga 2–6 simple, with entire antecosta. Posterior process of T7 as wide as half of the segment’s width, with a well separated rectangular patch that bears sensilla basiconica mixed with thin hairs, and a posterior, bold and well sclerotized arch. Tergum 8 with an erect, slightly backcurved mesal sclerite that bears a row of sensilla basiconica on its antero-apical keel; base of the sclerite is wide, the erect apex is about one seventh of segment’s width. Tergum 9 with medially interrupted antecosta, otherwise unmodified. Hemiterga of tergum 10 bent in dorsal, not raised in lateral view; hemitergal lobe with a ventral patch of sensilla basiconica meso-posteriorly. Hemitergal process long, points backwards and inwards, basally curved in dorsal view, tip downcurved in lateral view.
</paragraph>
<paragraph id="122036AA5D215602FF268882FAC9B42D" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="119">
Aedeagus (
<figureCitation id="8AA42A2F5D215602FEA48882FE5DB3F4" box="[325,463,908,933]" captionStart="FIGURES 18 – 23" captionStartId="7.[151,264,1639,1662]" captionTargetBox="[168,1409,566,1600]" captionTargetId="figure@7.[151,1436,555,1618]" captionTargetPageId="7" captionText="FIGURES 18 – 23. Holotype male of Neoperla goguryeo Murányi &amp; Li, sp. n.; North Korea, Kangwon Province, Kumgang Mts — 18: head and pronotum, dorsal view; 19: terminalia, dorsal view; 20: terminalia, lateral view; 21: aedeagus, lateral view; 22: aedeagus, ventral view; 23: aedeagus, dorsal view—scales 1 mm." httpUri="https://zenodo.org/record/234085/files/figure.png" pageId="6" pageNumber="119">Figs. 21–23</figureCitation>
): The aedeagal tube is plump, with a triangular dorsobasal sclerite, a ventrobasal hump and two large, spherical ventroapical lobes. Dorsal and dorsolateral surface of the lobes covered with fine spinules, less dense spinules forming a triangular patch posterior to the bald ventrobasal hump. The ventroapical lobes are touching each other ventrally, and bear large spines on their posterior surface, while anteriorly without spines. The everted aedeagal sac is straight, nearly 2× as long as the tube and bears a small, hemispherical lobe dorsoapically. Sac is covered throughout with fine spinules up to the apex, while two lateral, and a ventral field of strong spines occur at the level of the dorsoapical lobe. The separated, slightly downcurved apex of the sac without spines.
</paragraph>
<paragraph id="122036AA5D215602FF268F89FB7DB560" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="119">
Female terminalia (
<figureCitation id="8AA42A2F5D215602FE4D8F89FDA6B4F0" box="[428,564,1159,1184]" captionStart="FIGURES 10 – 17" captionStartId="5.[151,264,1137,1160]" captionTargetBox="[191,1368,246,1103]" captionTargetId="figure@5.[151,1436,193,1116]" captionTargetPageId="5" captionText="FIGURES 10 – 17. Females of Korean Neoperla Needham, 1905 — 10, 12, 14, 16: vagina, dorsal view; 11, 13, 15, 17: vagina, lateral view— 10 – 11: Neoperla adamantea Murányi &amp; Li, sp. n., North Korea, Kangwon Province, Kumgang Mts; 12 – 13: Neoperla coreensis Ra, Kim, Kang &amp; Ham, 1994, North Korea, North Pyongan Province, Myohyang Mts; 14 – 15: Neoperla goguryeo Murányi &amp; Li, sp. n., North Korea, Kangwon Province, Kumgang Mts; 16 – 17: Neoperla goguryeo Murányi &amp; Li, sp. n., North Korea, North Pyongan Province, Myohyang Mts—scale 1 mm." httpUri="https://zenodo.org/record/234084/files/figure.png" pageId="6" pageNumber="119">Figs 14–17</figureCitation>
): Sterna and terga simple, posterior edge of S8 forms a small, triangular subgenital plate. Vagina large, longer than wide and with many membranous folds. Inner sclerite large with two papillate lobes dorsally to anterior folds, basal portion forked. The inner pair of lateral folds bear a dark, well sclerotized crest that forms a Y-shaped pattern, with its posterior end covered by the apical, rather undulant posterior folds. Spermatheca small, its origin covered by the protruding lateral folds in dorsal view.
</paragraph>
<paragraph id="122036AA5D215602FF268E33FCCCB67D" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="119">
Egg (
<figureCitation id="8AA42A2F5D215602FEE58E32FE18B504" box="[260,394,1340,1365]" captionStart="FIGURES 24 – 33" captionStartId="8.[151,264,1858,1881]" captionTargetBox="[151,1436,195,1836]" captionTargetId="figure@8.[151,1436,195,1837]" captionTargetPageId="8" captionText="FIGURES 24 – 33. Eggs of Korean Neoperla Needham, 1905 — 24, 29 – 30: egg, lateral view; 25, 32: micropyle, lateral view; 26: collar, caudal view; 27: lid, caudal view; 28: collar, oblique view; 31: lid, lateral view; 33: collar, lateral view— 24 – 28: Neoperla coreensis Ra, Kim, Kang &amp; Ham, 1994, North Korea, North Pyongan Province, Myohyang Mts; 29, 31 – 33: Neoperla goguryeo Murányi &amp; Li, sp. n., North Korea, Kangwon Province, Kumgang Mts; 30: Neoperla goguryeo Murányi &amp; Li, sp. n., North Korea, North Pyongan Province, Myohyang Mts—scales 0.1 mm." httpUri="https://zenodo.org/record/234086/files/figure.png" pageId="6" pageNumber="119">Figs 29–33</figureCitation>
): Chorion dark brown, 
<quantity id="D5679B4F5D215602FD758E32FCAAB504" box="[660,824,1340,1364]" metricMagnitude="-4" metricUnit="m" metricValue="4.1" metricValueMax="4.4" metricValueMin="3.8" pageId="6" pageNumber="119" unit="mm" value="0.41" valueMax="0.44" valueMin="0.38">0.38–0.44 mm</quantity>
long and 
<quantity id="D5679B4F5D215602FC4A8E32FBC2B504" box="[939,1104,1340,1365]" metricMagnitude="-4" metricUnit="m" metricValue="2.2" metricValueMax="2.6" metricValueMin="1.7999999999999998" pageId="6" pageNumber="119" unit="mm" value="0.22" valueMax="0.26" valueMin="0.18">0.18–0.26 mm</quantity>
wide (N=25). Barrel shaped, cross section round. Hatching line inconspicuous. Micropyles placed in a transverse row closer to the opercular end, each located between striae. Collar sessile without distinct flange, surrounded by two to four rows of regular FCIs that have fine inner punctations; anchor not studied. Chorion covered with striae in groupings of four; outer striae of each group thickened and raised at both collar and lid end, but raised striae more conspicuous near collar. Striae set closely without sulci between them. Lid bears one or two distinct rows of FCIs at junction of striae, apical part covered with less distinct FCIs with fine inner punctations.
</paragraph>
<paragraph id="122036AA5D215602FF268D37FE15B600" blockId="6.[151,1437,151,2013]" box="[199,391,1591,1616]" pageId="6" pageNumber="119">Larva: unknown.</paragraph>
<paragraph id="122036AA5D215602FF268D52FDF4B78C" blockId="6.[151,1437,151,2013]" pageId="6" pageNumber="119">
<emphasis id="20EBEAB85D215602FF268D52FEAEB625" bold="true" box="[199,316,1628,1653]" pageId="6" pageNumber="119">Affinities.</emphasis>
This is a member of the 
<emphasis id="20EBEAB85D215602FD8E8D53FD77B624" box="[623,741,1629,1652]" italics="true" pageId="6" pageNumber="119">montivaga</emphasis>
group, 
<taxonomicName id="D59F4D295D215602FCA58D52FC0FB624" box="[836,925,1628,1652]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="119" phylum="Arthropoda" rank="species" species="lushana">
<emphasis id="20EBEAB85D215602FCA58D52FC0FB624" box="[836,925,1628,1652]" italics="true" pageId="6" pageNumber="119">lushana</emphasis>
</taxonomicName>
subgroup as recognized by 
<bibRefCitation id="760E4B5B5D215602FB0B8D52FF71B6C8" author="Zwick" pageId="6" pageNumber="119" refString="Zwick, P. &amp; Sivec, I. (1980) Beitrage zur Kenntnis der Plecoptera des Himalaja. Entomologica Basiliensia, 5, 59 - 138." type="journal article" year="1980">Zwick &amp; Sivec (1980)</bibRefCitation>
, and closely related to certain Chinese and Japanese species. Among the Japanese species, it is the more similar to 
<taxonomicName id="D59F4D295D215602FEEE8DABFDC0B6EC" authority="Pictet, 1841" authorityName="Pictet" authorityYear="1841" box="[271,594,1700,1725]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="119" phylum="Arthropoda" rank="species" species="geniculata">
<emphasis id="20EBEAB85D215602FEEE8DABFE3BB6EC" box="[271,425,1700,1724]" italics="true" pageId="6" pageNumber="119">N. geniculata</emphasis>
(
<bibRefCitation id="760E4B5B5D215602FE5A8DAAFDDBB6EC" author="Pictet" box="[443,585,1700,1725]" pageId="6" pageNumber="119" refString="Pictet, F. J. (1841) Histoire naturelle des insectes Nevropteres. Famille des Perlides. Kessmann-Bailliere, Geneve-Paris, 423 pp." type="book" year="1841">Pictet, 1841</bibRefCitation>
)
</taxonomicName>
, and to 
<taxonomicName id="D59F4D295D215602FD528DABFB9EB6EC" authority="Li &amp; Li, 2013" authorityName="Li &amp; Li" authorityYear="2013" box="[691,1036,1700,1725]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="119" phylum="Arthropoda" rank="species" species="xuansongae">
<emphasis id="20EBEAB85D215602FD528DABFCCFB6EC" box="[691,861,1701,1724]" italics="true" pageId="6" pageNumber="119">N. xuansongae</emphasis>
<bibRefCitation id="760E4B5B5D215602FC868DAAFB9EB6EC" author="Li" box="[871,1036,1700,1725]" pageId="6" pageNumber="119" refString="Li, W. H., Liang, H. Y. &amp; Li, W. L. (2013) Review of Neoperla (Plecoptera: Perlidae) from Zhejiang Province, China. Zootaxa, 3652 (3), 353 - 369. http: // dx. doi. org / 10.11646 / zootaxa. 3652.3.4" type="journal article" year="2013">Li &amp; Li, 2013</bibRefCitation>
</taxonomicName>
(in 
<bibRefCitation id="760E4B5B5D215602FBA18DAAFB71B6EC" author="Li" box="[1088,1251,1700,1725]" pageId="6" pageNumber="119" refString="Li, W. H., Liang, H. Y. &amp; Li, W. L. (2013) Review of Neoperla (Plecoptera: Perlidae) from Zhejiang Province, China. Zootaxa, 3652 (3), 353 - 369. http: // dx. doi. org / 10.11646 / zootaxa. 3652.3.4" type="journal article" year="2013">
Li 
<emphasis id="20EBEAB85D215602FB828DABFB0CB6EC" box="[1123,1182,1700,1724]" italics="true" pageId="6" pageNumber="119">et al.</emphasis>
2013
</bibRefCitation>
) among species described from 
<collectingCountry id="6A88763A5D215602FEAE8DC9FE04B6B0" box="[335,406,1735,1760]" name="China" pageId="6" pageNumber="119">China</collectingCountry>
. These species can be easily distinguished by aedeagal characters: 
<taxonomicName id="D59F4D295D215602FB4F8DC7FAACB6B0" box="[1198,1342,1737,1760]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="119" phylum="Arthropoda" rank="species" species="goguryeo">
<emphasis id="20EBEAB85D215602FB4F8DC7FAACB6B0" box="[1198,1342,1737,1760]" italics="true" pageId="6" pageNumber="119">N. goguryeo</emphasis>
</taxonomicName>
bears a dorsoapical lobe that is surrounded by two lateral and one dorsal field of strong spines, while the lobe is lacking on 
<taxonomicName id="D59F4D295D215602FF768C1FFEADB778" box="[151,319,1809,1832]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="119" phylum="Arthropoda" rank="species" species="xuansongae">
<emphasis id="20EBEAB85D215602FF768C1FFEADB778" box="[151,319,1809,1832]" italics="true" pageId="6" pageNumber="119">N. xuansongae</emphasis>
</taxonomicName>
and strong spines are lacking on 
<taxonomicName id="D59F4D295D215602FD5D8C1FFCC1B778" box="[700,851,1808,1832]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="119" phylum="Arthropoda" rank="species" species="geniculata">
<emphasis id="20EBEAB85D215602FD5D8C1FFCC1B778" box="[700,851,1808,1832]" italics="true" pageId="6" pageNumber="119">N. geniculata</emphasis>
</taxonomicName>
aedeagus. Development of the vagina is typical for the 
<taxonomicName id="D59F4D295D215602FF218C3AFE8BB71C" box="[192,281,1844,1868]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="119" phylum="Arthropoda" rank="species" species="lushana">
<emphasis id="20EBEAB85D215602FF218C3AFE8BB71C" box="[192,281,1844,1868]" italics="true" pageId="6" pageNumber="119">lushana</emphasis>
</taxonomicName>
subgroup, can be distinguished from those of the known females by the large, Y-shaped pattern formed by the inner pair of lateral folds. Egg is of a usual 
<taxonomicName id="D59F4D295D215602FD248C56FCBFB720" box="[709,813,1880,1904]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="20EBEAB85D215602FD248C56FCBFB720" box="[709,813,1880,1904]" italics="true" pageId="6" pageNumber="119">Neoperla</emphasis>
</taxonomicName>
<typeStatus id="CD2488085D215602FCD58C57FCF5B721" box="[820,871,1881,1905]" pageId="6" pageNumber="119">type</typeStatus>
, hardly distinguishable from several other species. The habitus, coloration and male terminalia of the new species are similar to many other Palaearctic 
<taxonomicName id="D59F4D295D215602FACD8C72FA06B7C4" box="[1324,1428,1916,1940]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="119" phylum="Arthropoda" rank="genus">
<emphasis id="20EBEAB85D215602FACD8C72FA06B7C4" box="[1324,1428,1916,1940]" italics="true" pageId="6" pageNumber="119">Neoperla</emphasis>
</taxonomicName>
. Habitus differences from the coexisting 
<taxonomicName id="D59F4D295D215602FDBD8CAFFD69B7E8" box="[604,763,1952,1976]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="119" phylum="Arthropoda" rank="species" species="adamantea">
<emphasis id="20EBEAB85D215602FDBD8CAFFD69B7E8" box="[604,763,1952,1976]" italics="true" pageId="6" pageNumber="119">N. adamantea</emphasis>
</taxonomicName>
as discussed above. The sympatric 
<taxonomicName id="D59F4D295D215602FB6E8CAFFA88B7E8" box="[1167,1306,1953,1976]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="6" pageNumber="119" phylum="Arthropoda" rank="species" species="coreensis">
<emphasis id="20EBEAB85D215602FB6E8CAFFA88B7E8" box="[1167,1306,1953,1976]" italics="true" pageId="6" pageNumber="119">N. coreensis</emphasis>
</taxonomicName>
has a more darkly marked head and pronotal pattern.
</paragraph>
</subSubSection>
<subSubSection id="5A8565215D205603FF268B99FDDCB1D8" pageId="7" pageNumber="120" type="distribution">
<paragraph id="122036AA5D205603FF268B99FDDCB1D8" blockId="7.[151,1436,151,501]" pageId="7" pageNumber="120">
<emphasis id="20EBEAB85D205603FF268B99FE6CB0E0" bold="true" box="[199,510,151,176]" pageId="7" pageNumber="120">Distribution and ecology.</emphasis>
The species was collected at light in the lower regions of the Kumgang and Myohyang Mountains (
<figureCitation id="8AA42A2F5D205603FE7F8BB2FE62B084" box="[414,496,188,213]" captionStart="FIGURES 41 – 42" captionStartId="12.[151,264,1508,1531]" captionTargetBox="[154,1426,453,1486]" captionTargetId="figure@12.[154,1427,453,1486]" captionTargetPageId="12" captionText="FIGURES 41 – 42. Distribution of the Neoperla Needham, 1905 species known from Korea — 41: known occurrences in North and South Korea; 42: known range of N. quadrata Wu &amp; Claassen, 1934 and N. ussurica Sivec &amp; Zhiltzova, 1996." httpUri="https://zenodo.org/record/234089/files/figure.png" pageId="7" pageNumber="120">Fig. 41</figureCitation>
). The Kumgang Mountains form the northern part of the Thebaek Ranges, while the Myohyang Mountains form the western chains of the Changbai (Baekdu) Ranges that separate the Korean Peninsula from the Asian continent. In the Kumgang Mountains the species co-occurs with 
<emphasis id="20EBEAB85D205603FB7D8A0BFAD2B14C" box="[1180,1344,260,284]" italics="true" pageId="7" pageNumber="120">
<taxonomicName id="D59F4D295D205603FB7D8A0BFAA9B14C" box="[1180,1339,260,284]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="120" phylum="Arthropoda" rank="species" species="adamantea">N. adamantea</taxonomicName>
,
</emphasis>
while it was collected together with 
<taxonomicName id="D59F4D295D205603FE0B8A27FDE8B16F" box="[490,634,296,320]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="120" phylum="Arthropoda" rank="species" species="coreensis">
<emphasis id="20EBEAB85D205603FE0B8A27FDE8B16F" box="[490,634,296,320]" italics="true" pageId="7" pageNumber="120">N. coreensis</emphasis>
</taxonomicName>
in the Myohyang Mountains. Other large perlids like 
<taxonomicName id="D59F4D295D205603FAF18A26FE7DB134" authority="McLachlan, 1872" authorityName="McLachlan" authorityYear="1872" class="Insecta" family="Perlidae" genus="Paragnetina" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="120" phylum="Arthropoda" rank="species" species="flavotincta">
<emphasis id="20EBEAB85D205603FAF18A26FE9DB134" italics="true" pageId="7" pageNumber="120">Paragnetina flavotincta</emphasis>
(
<bibRefCitation id="760E4B5B5D205603FEFF8A42FE75B134" author="McLachlan" box="[286,487,332,357]" pageId="7" pageNumber="120" refString="McLachlan, M. R. (1872) Materiaux pour une Faune Nevropterologique de l'Asie Septentrionale. Seconde partie. Non- Odonates. Annales de la Societe Entomologique de Belgique, 15, 47 - 77. [plates I-II]" type="journal article" year="1872">McLachlan, 1872</bibRefCitation>
)
</taxonomicName>
also frequently occurred with 
<taxonomicName id="D59F4D295D205603FCAD8A43FC4AB134" box="[844,984,333,356]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="120" phylum="Arthropoda" rank="species" species="goguryeo">
<emphasis id="20EBEAB85D205603FCAD8A43FC4AB134" box="[844,984,333,356]" italics="true" pageId="7" pageNumber="120">N. goguryeo</emphasis>
</taxonomicName>
at the light traps. This species seems to occur in large streams and small rivers.
</paragraph>
</subSubSection>
<subSubSection id="5A8565215D205603FF268A9AFD63B1A5" pageId="7" pageNumber="120" type="etymology">
<paragraph id="122036AA5D205603FF268A9AFD63B1A5" blockId="7.[151,1436,151,501]" pageId="7" pageNumber="120">
<emphasis id="20EBEAB85D205603FF268A9AFEDCB1FD" bold="true" box="[199,334,404,429]" pageId="7" pageNumber="120">Etymology.</emphasis>
The name 
<taxonomicName id="D59F4D295D205603FE288A9BFDA1B1FC" box="[457,563,405,428]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="120" phylum="Arthropoda" rank="species" species="goguryeo">
<emphasis id="20EBEAB85D205603FE288A9BFDA1B1FC" box="[457,563,405,428]" italics="true" pageId="7" pageNumber="120">goguryeo</emphasis>
</taxonomicName>
refers to 
<taxonomicName id="D59F4D295D205603FD7D8A9AFC9EB1FC" box="[668,780,404,428]" class="Insecta" family="Perlidae" genus="Neoperla" kingdom="Animalia" order="Plecoptera" pageId="7" pageNumber="126" phylum="Arthropoda" rank="species" species="goguryeo">Goguryeo</taxonomicName>
or Koguryo, one of the ancient Three Kingdoms of 
<collectingCountry id="6A88763A5D205603FAB08A9BFA0AB1FC" box="[1361,1432,405,428]" name="South Korea" pageId="7" pageNumber="120">Korea</collectingCountry>
, which existed from 37 BC to 668 AD (
<bibRefCitation id="760E4B5B5D205603FDB38AB6FD74B180" author="Csoma" box="[594,742,440,464]" pageId="7" pageNumber="120" refString="Csoma, M. (2013) Korea. Egy nemzet, ket orszag - a kozos gyokerektol. 2 nd edition, Napvilag Kiado Kft, Budapest, 281 pp." type="book" year="2013">Csoma 2013</bibRefCitation>
). The known distribution of the species delimits the center of this ancient kingdom. Used as a noun, gender neutral.
</paragraph>
</subSubSection>
</treatment>
</document>