treatments-xml/data/D2/3C/87/D23C87A0FFBF2E0FFEF8253AFCD2BB40.xml
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<document id="0782839F95A5D7D786885C9387A17B47" ID-CLB-Dataset="198285" ID-DOI="10.1080/00222933.2022.2138600" ID-GBIF-Dataset="f3ea9905-e60b-4102-9404-65b3dcd1583c" ID-ISSN="1464-5262" ID-Zenodo-Dep="7380764" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669805273948" checkinUser="plazi" docAuthor="Tiwari, Shivam, Padate, Vinay P., Cubelio, Sherine S. &amp; Osawa, Masayuki" docDate="2022" docId="D23C87A0FFBF2E0FFEF8253AFCD2BB40" docLanguage="en" docName="JNATHIST.56.41-44.1819-1839.pdf" docOrigin="Journal of Natural History 56" docSource="http://dx.doi.org/10.1080/00222933.2022.2138600" docStyle="DocumentStyle:3C762F9783B768A90048B82ACD135A62.3:JNatHist.2017-.journal_article.0cover" docStyleId="3C762F9783B768A90048B82ACD135A62" docStyleName="JNatHist.2017-.journal_article.0cover" docStyleVersion="3" docTitle="Raymunida shraddhanandi Tiwari &amp; Padate &amp; Cubelio &amp; Osawa 2022, sp. nov." docType="treatment" docVersion="6" lastPageNumber="1829" masterDocId="2E05FFD8FFBC2E05FF442110FFC2BF11" masterDocTitle="Squat lobsters of the genera Raymunida Macpherson &amp; Machordom, 2000 and Munidopsis Whiteaves, 1874 (Decapoda: Anomura: Galatheoidea) from the Indian Exclusive Economic Zone, with descriptions of three new species" masterLastPageNumber="1839" masterPageNumber="1819" pageNumber="1822" updateTime="1699412247313" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C9384344F51BF8F066AC04E68E63408C">Squat lobsters of the genera Raymunida Macpherson &amp; Machordom, 2000 and Munidopsis Whiteaves, 1874 (Decapoda: Anomura: Galatheoidea) from the Indian Exclusive Economic Zone, with descriptions of three new species</mods:title>
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<mods:namePart id="BD82238A3B78E378251C0FD0C1A11AC0">Tiwari, Shivam</mods:namePart>
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<mods:namePart id="A4068E339C83A065A5D23D44AD1CDCAC">Padate, Vinay P.</mods:namePart>
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<mods:namePart id="3E6EAAA6B1B5FE96D285F6AC1027DAB4">Cubelio, Sherine S.</mods:namePart>
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<mods:namePart id="8B26AE8B27C6A5F67F8ACF1C925BC697">Osawa, Masayuki</mods:namePart>
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<mods:title id="892283659D4007AD69B4A891B9463841">Journal of Natural History</mods:title>
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<mods:part id="5471504C0ADCD55C1156F0A15A6974E4">
<mods:date id="A64D4BD1E514DBD14D3CCDD2F45FB429">2022</mods:date>
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<mods:number id="049A41293214AB48CD3CFE2A9F20638E">2022-11-29</mods:number>
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<mods:number id="3B5566DCCCAD0F8E3B9E9F4A334EBDA0">56</mods:number>
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<mods:start id="ECD5FDDC482558DF159BD361EA7B7B1B">1819</mods:start>
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<treatment id="D23C87A0FFBF2E0FFEF8253AFCD2BB40" ID-DOI="http://doi.org/10.5281/zenodo.7383872" ID-GBIF-Taxon="205278693" ID-Zenodo-Dep="7383872" LSID="urn:lsid:plazi:treatment:D23C87A0FFBF2E0FFEF8253AFCD2BB40" httpUri="http://treatment.plazi.org/id/D23C87A0FFBF2E0FFEF8253AFCD2BB40" lastPageId="10" lastPageNumber="1829" pageId="3" pageNumber="1822">
<subSubSection id="128F653DFFBF2E06FEF8253AFCAABB55" box="[444,872,1066,1092]" pageId="3" pageNumber="1822" type="nomenclature">
<paragraph id="5A2A36B6FFBF2E06FEF8253AFCAABB55" blockId="3.[394,922,1066,1130]" box="[444,872,1066,1092]" pageId="3" pageNumber="1822">
<heading id="016281DAFFBF2E06FEF8253AFCAABB55" box="[444,872,1066,1092]" centered="true" fontSize="11" level="2" pageId="3" pageNumber="1822" reason="2">
<emphasis id="68E1EAA4FFBF2E06FEF8253AFCAABB55" box="[444,872,1066,1092]" italics="true" pageId="3" pageNumber="1822">
<taxonomicName id="9D954D35FFBF2E06FEF8253AFCCBBB55" ID-CoL="dad1780c-eeba-4c30-9456-a8ba675ae5a7" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[444,777,1066,1092]" class="Malacostraca" family="Munididae" genus="Raymunida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="1822" phylum="Arthropoda" rank="species" species="shraddhanandi" status="sp. nov.">
<emphasis id="68E1EAA4FFBF2E06FEF8253AFCCBBB55" bold="true" box="[444,777,1066,1092]" italics="true" pageId="3" pageNumber="1822">Raymunida shraddhanandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFBF2E06FC54253AFCAABB55" box="[784,872,1066,1092]" pageId="3" pageNumber="1822" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="128F653DFFBF2E06FECE2540FBDDBB8F" pageId="3" pageNumber="1822" type="description">
<paragraph id="5A2A36B6FFBF2E06FECE2540FC58BB7B" blockId="3.[394,922,1066,1130]" box="[394,922,1104,1130]" pageId="3" pageNumber="1822">
(
<figureCitation id="C2AE2A33FFBF2E06FED52540FE3FBB7B" box="[401,509,1104,1130]" captionStart="Figure 2" captionStartId="4.[160,225,957,980]" captionTargetBox="[163,1152,142,925]" captionTargetId="figure-248@4.[163,1152,142,925]" captionTargetPageId="4" captionText="Figure 2. Habitus,dorsal view.(a) Raymunida shraddhanandi sp.nov., holotype ♂ (PCL 9.7 mm,CW 8.2 mm), IO/SS/ANO/00141;(b) R.shraddhanandi sp.nov.,paratype ♀ (PCL 5.8 mm,CW 4.9 mm),IO/SS/ANO/00140;(c) R. vittata Macpherson, 2009, ♀ (PCL 9.0 mm, CW 7.7 mm), IO/SS/ANO/00142. Scale bars: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7380768" httpUri="https://zenodo.org/record/7380768/files/figure.png" pageId="3" pageNumber="1822">Figures 2</figureCitation>
(a), 3(a3),
<figureCitation id="C2AE2A33FFBF2E06FDD02540FD36BB7B" box="[660,756,1104,1130]" captionStart="Figure 4" captionStartId="6.[160,225,1513,1536]" captionTargetBox="[164,1151,143,1481]" captionTargetId="figure-1@6.[163,1152,142,1482]" captionTargetPageId="6" captionText="Figure 4. Raymunida shraddhanandi sp. nov., holotype ♂ (PCL 9.7 mm, CW 8.2 mm), IO/SS/ANO/ 00141. (a) carapace (dorsal view); (b) carapace (left lateral view); (c) thoracic sternum (ventral view); (d) pleomeres 15 (dorsal view); (e) pleomeres 5 and 6 and telson (external view); (f) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (g) left Mxp3 (lateral view); (h) left P1 merus and carpus (dorsal view); (i) left P1 chela (dorsal view); (j) right P2 (lateral view); (k) right P3 (lateral view); (l), right P4 (lateral view); (m) right P5 dactylus (lateral view); (n) left G1 (internal view); (o) left G2 (external view). Scale bars: a, cl, o = 1.0 mm; b = 2.0 mm; m, n = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380774" httpUri="https://zenodo.org/record/7380774/files/figure.png" pageId="3" pageNumber="1822">Figure 4</figureCitation>
and
<figureCitation id="C2AE2A33FFBF2E06FC752540FC53BB7B" box="[817,913,1104,1130]" captionStart="Figure 5" captionStartId="7.[160,225,1379,1402]" captionTargetBox="[165,1150,143,1346]" captionTargetId="figure-56@7.[164,1151,142,1347]" captionTargetPageId="7" captionText="Figure 5. Raymunida shraddhanandi sp. nov., paratype ♀ (PCL 5.8 mm, CW 4.9 mm), IO/SS/ANO/00140. (a) carapace, pleon and cephalic appendages (dorsal view); (b) thoracic sternum (ventral view); (c) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (d) right Mxp3 (lateral view); (e) right P1 (dorsal view);(f) left P2 propodus and dactylus (lateral view); (g) left P2 merus and carpus (lateral view);(h) left P3 propodus and dactylus (lateral view); (i) left P3 merus and carpus (lateral view);(j) left P4 propodus and dactylus (lateral view); (k) left P4 merus and carpus (lateral view). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7380776" httpUri="https://zenodo.org/record/7380776/files/figure.png" pageId="3" pageNumber="1822">Figure 5</figureCitation>
)
</paragraph>
<paragraph id="5A2A36B6FFBF2E06FE612595FBDDBB8F" blockId="3.[293,1055,1157,1182]" box="[293,1055,1157,1182]" pageId="3" pageNumber="1822">
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urn:lsid:zoobank.org:act:
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<emphasis id="68E1EAA4FFBF2E06FFE425C0FE86BBFB" bold="true" box="[160,324,1232,1258]" italics="true" pageId="3" pageNumber="1822">Type material</emphasis>
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<paragraph id="5A2A36B6FFBF2E06FFE425E2FD92BA40" blockId="3.[160,1156,1232,1361]" pageId="3" pageNumber="1822">
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<typeStatus id="852E8814FFBF2E06FFE425E2FECFBA1D" box="[160,269,1266,1292]" pageId="3" pageNumber="1822" type="holotype">Holotype</typeStatus>
.
</emphasis>
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<collectingCountry id="22827626FFBF2E06FE6525E3FE9CBA1D" box="[289,350,1267,1292]" name="India" pageId="3" pageNumber="1822">India</collectingCountry>
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<specimenCount id="4C93FD3FFFBF2E06FE3725E4FE45BA1A" box="[371,391,1268,1291]" pageId="3" pageNumber="1822" type="male"></specimenCount>
(PCL
<quantity id="9D6D9B53FFBF2E06FE8325E3FDDCBA1D" box="[455,542,1267,1292]" metricMagnitude="-3" metricUnit="m" metricValue="9.7" pageId="3" pageNumber="1822" unit="mm" value="9.7">9.7 mm</quantity>
, CW
<quantity id="9D6D9B53FFBF2E06FD1025E3FD69BA1D" box="[596,683,1267,1292]" metricMagnitude="-3" metricUnit="m" metricValue="8.2" pageId="3" pageNumber="1822" unit="mm" value="8.2">8.2 mm</quantity>
);
<location id="5F4A606DFFBF2E06FDF825E3FC98BA1D" LSID="urn:lsid:plazi:treatment:D23C87A0FFBF2E0FFEF8253AFCD2BB40:5F4A606DFFBF2E06FDF825E3FC98BA1D" box="[700,858,1267,1292]" country="India" latitude="10.72" longLatPrecision="5486" longitude="92.7" municipality="Little Andaman Island" name="Andaman Sea" pageId="3" pageNumber="1822">Andaman Sea</location>
, off
<collectingMunicipality id="BA4EACCCFFBF2E06FCCD25E3FB43BA1D" box="[905,1153,1267,1292]" pageId="3" pageNumber="1822">Little Andaman Island</collectingMunicipality>
; FORVSS stn. 38806;
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,
<geoCoordinate id="3FA15071FFBF2E06FEA82406FDF1BA3E" box="[492,563,1302,1327]" degrees="92.7" direction="east" orientation="longitude" pageId="3" pageNumber="1822" precision="5555" value="92.7">92.7°E</geoCoordinate>
;
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<quantity id="9D6D9B53FFBF2E06FD7B2406FDB8BA3E" box="[575,634,1302,1327]" metricMagnitude="1" metricUnit="m" metricValue="5.3" pageId="3" pageNumber="1822" unit="m" value="53.0">53 m</quantity>
depth
</elevation>
;
<collectorName id="F7605360FFBF2E06FD932406FCD5BA3E" box="[727,791,1302,1327]" pageId="3" pageNumber="1822">Chain</collectorName>
dredge;
<date id="2E2B1076FFBF2E06FCC52406FBFBBA3E" box="[897,1081,1302,1327]" pageId="3" pageNumber="1822" value="2019-08-10">
<collectingDate id="3E6FE99EFFBF2E06FCC52406FBFBBA3E" box="[897,1081,1302,1327]" pageId="3" pageNumber="1822" value="2019-08-10">10 August 2019</collectingDate>
</date>
;
<collectorName id="F7605360FFBF2E06FB022406FECABA40" pageId="3" pageNumber="1822">Vinay P. Padate</collectorName>
leg.;
<collectionCode id="3C84AE73FFBF2E06FE01242EFEB9BA5E" box="[325,379,1342,1359]" pageId="3" pageNumber="1822">CMLRE</collectionCode>
<collectionCode id="3C84AE73FFBF2E06FEC62428FE58BA40" box="[386,410,1336,1361]" country="Portugal" name="Instituto de Oceanografia da Universidade de Lisboa" pageId="3" pageNumber="1822">IO</collectionCode>
/SS/ANO/00141.
</materialsCitation>
</paragraph>
<paragraph id="5A2A36B6FFBF2E06FFE42494FEEBBAD1" blockId="3.[160,1155,1412,1472]" pageId="3" pageNumber="1822">
<materialsCitation id="EAFD3CEBFFBF2E06FFE42494FEE7BAD1" ID-GBIF-Occurrence="3978782309" collectingDate="2019-08-10" collectionCode="CMLRE, IO" collectorName="Chain &amp; Vinay P. Padate" country="India" elevation="53" latitude="10.72" location="Andaman Sea" longLatPrecision="5486" longitude="92.7" municipality="Little Andaman Island" pageId="3" pageNumber="1822" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<emphasis id="68E1EAA4FFBF2E06FFE42494FED3BA8F" bold="true" box="[160,273,1412,1438]" italics="true" pageId="3" pageNumber="1822">
<typeStatus id="852E8814FFBF2E06FFE42494FECEBA8F" box="[160,268,1412,1438]" pageId="3" pageNumber="1822" type="paratype">Paratype</typeStatus>
.
</emphasis>
<emphasis id="68E1EAA4FFBF2E06FE652495FE9CBA8F" bold="true" box="[289,350,1413,1438]" pageId="3" pageNumber="1822">
<collectingCountry id="22827626FFBF2E06FE652495FE9CBA8F" box="[289,350,1413,1438]" name="India" pageId="3" pageNumber="1822">India</collectingCountry>
</emphasis>
<specimenCount id="4C93FD3FFFBF2E06FE372495FE59BA8D" box="[371,411,1413,1438]" pageId="3" pageNumber="1822" type="female">1 ♀</specimenCount>
(PCL
<quantity id="9D6D9B53FFBF2E06FE9F2495FDF0BA8F" box="[475,562,1413,1438]" metricMagnitude="-3" metricUnit="m" metricValue="5.8" pageId="3" pageNumber="1822" unit="mm" value="5.8">5.8 mm</quantity>
, CW
<quantity id="9D6D9B53FFBF2E06FD2D2495FD02BA8F" box="[617,704,1413,1438]" metricMagnitude="-3" metricUnit="m" metricValue="4.9" pageId="3" pageNumber="1822" unit="mm" value="4.9">4.9 mm</quantity>
); same data as for holotype;
<collectionCode id="3C84AE73FFBF2E06FB45249AFBF5BA8A" box="[1025,1079,1418,1435]" pageId="3" pageNumber="1822">CMLRE</collectionCode>
<collectionCode id="3C84AE73FFBF2E06FB7A2495FB97BA8F" box="[1086,1109,1413,1438]" country="Portugal" name="Instituto de Oceanografia da Universidade de Lisboa" pageId="3" pageNumber="1822">IO</collectionCode>
/SS/ ANO/00140
</materialsCitation>
.
</paragraph>
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<emphasis id="68E1EAA4FFBF2E06FFE424E2FEF4B91D" bold="true" box="[160,310,1522,1548]" italics="true" pageId="3" pageNumber="1822">Type locality</emphasis>
</paragraph>
<paragraph id="5A2A36B6FFBF2E06FFE42706FC36B93E" blockId="3.[160,1012,1522,1583]" box="[160,1012,1558,1583]" pageId="3" pageNumber="1822">
<materialsCitation id="EAFD3CEBFFBF2E06FFE42706FC33B93E" ID-GBIF-Occurrence="3978782306" box="[160,1009,1558,1583]" country="India" elevation="53" latitude="10.72" location="Andaman Sea" longLatPrecision="5486" longitude="92.7" municipality="Little Andaman Island" pageId="3" pageNumber="1822" specimenCount="1">
<collectingCountry id="22827626FFBF2E06FFE42706FF18B93E" box="[160,218,1558,1583]" name="India" pageId="3" pageNumber="1822">India</collectingCountry>
:
<location id="5F4A606DFFBF2E06FFA02706FE41B93E" LSID="urn:lsid:plazi:treatment:D23C87A0FFBF2E0FFEF8253AFCD2BB40:5F4A606DFFBF2E06FFA02706FE41B93E" box="[228,387,1558,1583]" country="India" latitude="10.72" longLatPrecision="5486" longitude="92.7" municipality="Little Andaman Island" name="Andaman Sea" pageId="3" pageNumber="1822">Andaman Sea</location>
, off
<collectingMunicipality id="BA4EACCCFFBF2E06FEF72706FD6EB93E" box="[435,684,1558,1583]" pageId="3" pageNumber="1822">Little Andaman Island</collectingMunicipality>
;
<geoCoordinate id="3FA15071FFBF2E06FDF22706FCCDB93E" box="[694,783,1558,1583]" degrees="10.72" direction="north" orientation="latitude" pageId="3" pageNumber="1822" precision="555" value="10.72">10.72°N</geoCoordinate>
,
<geoCoordinate id="3FA15071FFBF2E06FC5D2706FCA2B93E" box="[793,864,1558,1583]" degrees="92.7" direction="east" orientation="longitude" pageId="3" pageNumber="1822" precision="5555" value="92.7">92.7°E</geoCoordinate>
;
<elevation id="D1B8D185FFBF2E06FC2E2706FC33B93E" box="[874,1009,1558,1583]" metricMagnitude="1" metricUnit="m" metricValue="5.3" pageId="3" pageNumber="1822" unit="m" value="53.0">
<quantity id="9D6D9B53FFBF2E06FC2E2706FC61B93E" box="[874,931,1558,1583]" metricMagnitude="1" metricUnit="m" metricValue="5.3" pageId="3" pageNumber="1822" unit="m" value="53.0">53 m</quantity>
depth
</elevation>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="128F653DFFBF2E06FFE42771FDE3B9D1" pageId="3" pageNumber="1822" type="etymology">
<paragraph id="5A2A36B6FFBF2E06FFE42771FEDFB96A" blockId="3.[160,1156,1633,1728]" box="[160,285,1633,1659]" pageId="3" pageNumber="1822">
<emphasis id="68E1EAA4FFBF2E06FFE42771FEDFB96A" bold="true" box="[160,285,1633,1659]" italics="true" pageId="3" pageNumber="1822">Etymology</emphasis>
</paragraph>
<paragraph id="5A2A36B6FFBF2E06FFE42795FDE3B9D1" blockId="3.[160,1156,1633,1728]" pageId="3" pageNumber="1822">
The species is named in honour of the Swami Shraddhanand College, University of
<collectingRegion id="9851F854FFBF2E06FB072795FB42B98F" box="[1091,1152,1669,1694]" country="India" name="Delhi" pageId="3" pageNumber="1822">Delhi</collectingRegion>
, the alma mater of the first author.
</paragraph>
</subSubSection>
<caption id="0EEA663EFFB82E01FFE422ADFBE5BB1F" ID-DOI="http://doi.org/10.5281/zenodo.7380768" ID-Zenodo-Dep="7380768" httpUri="https://zenodo.org/record/7380768/files/figure.png" pageId="4" pageNumber="1823" startId="4.[160,225,957,980]" targetBox="[163,1152,142,925]" targetPageId="4">
<paragraph id="5A2A36B6FFB82E01FFE422ADFBE5BB1F" blockId="4.[159,1156,957,1038]" pageId="4" pageNumber="1823">
<emphasis id="68E1EAA4FFB82E01FFE422ADFF3BBCC5" bold="true" box="[160,249,957,980]" pageId="4" pageNumber="1823">Figure 2.</emphasis>
Habitus, dorsal view. (a)
<taxonomicName id="9D954D35FFB82E01FE9F22ADFD0FBCC5" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[475,717,957,980]" class="Malacostraca" family="Munididae" genus="Raymunida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="1823" phylum="Arthropoda" rank="species" species="shraddhanandi" status="sp. nov.">
<emphasis id="68E1EAA4FFB82E01FE9F22ADFD0FBCC5" box="[475,717,957,980]" italics="true" pageId="4" pageNumber="1823">Raymunida shraddhanandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFB82E01FD9522ADFCD9BCC5" box="[721,795,957,980]" pageId="4" pageNumber="1823" rank="species">sp. nov.</taxonomicNameLabel>
, holotype ♂ (PCL 9.7 mm, CW 8.2 mm), IO/SS/ANO/00141; (b)
<taxonomicName id="9D954D35FFB82E01FE2E22CAFDC5BCE0" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[362,519,986,1009]" class="Malacostraca" family="Munididae" genus="Raymunida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="1823" phylum="Arthropoda" rank="species" species="shraddhanandi" status="sp. nov.">
<emphasis id="68E1EAA4FFB82E01FE2E22CAFDC5BCE0" box="[362,519,986,1009]" italics="true" pageId="4" pageNumber="1823">R. shraddhanandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFB82E01FD4F22CAFD96BCE0" box="[523,596,986,1009]" pageId="4" pageNumber="1823" rank="species">sp. nov.</taxonomicNameLabel>
, paratype ♀ (PCL 5.8 mm, CW 4.9 mm), IO/SS/ANO/00140; (c)
<taxonomicName id="9D954D35FFB82E01FFDB22E7FE62BB1F" authority="Macpherson, 2009" authorityName="Macpherson" authorityYear="2009" box="[159,416,1015,1038]" class="Malacostraca" family="Munididae" genus="Raymunida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="1823" phylum="Arthropoda" rank="species" species="vittata">
<emphasis id="68E1EAA4FFB82E01FFDB22E7FF32BB1F" box="[159,240,1015,1038]" italics="true" pageId="4" pageNumber="1823">R. vittata</emphasis>
<bibRefCitation id="3E044B47FFB82E01FFB222E7FE62BB1F" author="Macpherson E" box="[246,416,1015,1038]" pageId="4" pageNumber="1823" pagination="431 - 451" refId="ref9854" refString="Macpherson E. 2009. New species of squat lobsters of the genera Munida and Raymunida (Crustacea, Decapoda, Galatheidae) from Vanuatu and New Caledonia. Zoosystema. 31 (3): 431 - 451. doi: 10. 5252 / z 2009 n 3 a 3." type="journal article" year="2009">Macpherson, 2009</bibRefCitation>
</taxonomicName>
, ♀ (PCL 9.0 mm, CW 7.7 mm), IO/SS/ANO/00142. Scale bars: 2.0 mm.
</paragraph>
</caption>
<subSubSection id="128F653DFFB82E01FFE4255AFCBAB9A0" pageId="4" pageNumber="1823" type="diagnosis">
<paragraph id="5A2A36B6FFB82E01FFE4255AFED6BB75" blockId="4.[159,1156,1098,1713]" box="[160,276,1098,1124]" pageId="4" pageNumber="1823">
<emphasis id="68E1EAA4FFB82E01FFE4255AFED6BB75" bold="true" box="[160,276,1098,1124]" italics="true" pageId="4" pageNumber="1823">Diagnosis</emphasis>
</paragraph>
<paragraph id="5A2A36B6FFB82E01FFE4257EFCBAB9A0" blockId="4.[159,1156,1098,1713]" pageId="4" pageNumber="1823">
Carapace (exclusive of rostrum) 1.2 times as long as wide; dorsal surface with sparse long iridescent setae, 5 pairs of epigastric spines, 01 anterior branchial spine, 1 parahepatic spine and 1 post-cervical spine on either side; frontal margin with short spine mesial to anterolateral spine; branchial margin with 4 spines (2 on each of anterior and posterior margins). Rostrum 0.4 times PCL, 2 times longer than supraocular spines. Anterior margin of thoracic sternite 4 as wide as sternite 3, sternites 57 smooth or with few short oblique striae on lateral portions. Pleonal tergites 2 and 3 each with or without row of interrupted, short striae between 2 transverse ridges. Antennular peduncle article 1 (excluding distal spines) not reaching distal corneal margin. Antennal peduncle article 1 with distomesial spine slightly overreaching distal margin of article 3, not reaching distal margin of antennular peduncle article 1; article 2 with distomesial spine reaching distal margin of article 3, unarmed on mesial margin; article 3 with short distomesial spine. Mxp3 merus with dorsodistal spine, ventral margin with 2 slender spines;carpus with 1 or 2 distal spines on ventral margin. P1 3.13.6 times PCL,with numerous long, simple setae; chela 4.6 (
<typeStatus id="852E8814FFB82E01FE982720FDFFB958" box="[476,573,1584,1609]" pageId="4" pageNumber="1823" type="paratype">paratype</typeStatus>
female) to 7.5 (
<typeStatus id="852E8814FFB82E01FDA32720FC8BB958" box="[743,841,1584,1609]" pageId="4" pageNumber="1823" type="holotype">holotype</typeStatus>
male) times as long as wide, with dorsolateral row of sparsely arranged spines; fingers each with distinct rounded crest along dorsal midline. P24 meri squamate on lateral surface, P4 mero-carpal articulation reaching level of lateral end of anterior cervical groove of carapace.
</paragraph>
</subSubSection>
<caption id="0EEA663EFFB92E00FFE42451FEE5BAFA" ID-DOI="http://doi.org/10.5281/zenodo.7380770" ID-Zenodo-Dep="7380770" httpUri="https://zenodo.org/record/7380770/files/figure.png" pageId="5" pageNumber="1824" startId="5.[160,225,1345,1368]" targetBox="[163,1152,142,1314]" targetPageId="5">
<paragraph id="5A2A36B6FFB92E00FFE42451FEE5BAFA" blockId="5.[160,1157,1345,1515]" pageId="5" pageNumber="1824">
<emphasis id="68E1EAA4FFB92E00FFE42451FF3EBA49" bold="true" box="[160,252,1345,1368]" pageId="5" pageNumber="1824">Figure 3.</emphasis>
Carapace, pleon and cephalic appendages, dorsal view. (a)
<taxonomicName id="9D954D35FFB92E00FC1E2451FBA2BA49" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[858,1120,1345,1368]" class="Malacostraca" family="Munididae" genus="Raymunida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="1824" phylum="Arthropoda" rank="species" species="shraddhanandi" status="sp. nov.">
<emphasis id="68E1EAA4FFB92E00FC1E2451FBA2BA49" box="[858,1120,1345,1368]" italics="true" pageId="5" pageNumber="1824">Raymunida shraddhanandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFB92E00FB2C2451FF0FBA67" pageId="5" pageNumber="1824" rank="species">sp. nov.</taxonomicNameLabel>
, holotype ♂ (PCL 9.7 mm, CW 8.2 mm), IO/SS/ANO/00141; (b)
<taxonomicName id="9D954D35FFB92E00FC6E244FFC10BA67" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[810,978,1375,1398]" class="Malacostraca" family="Munididae" genus="Raymunida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="1824" phylum="Arthropoda" rank="species" species="shraddhanandi" status="sp. nov.">
<emphasis id="68E1EAA4FFB92E00FC6E244FFC10BA67" box="[810,978,1375,1398]" italics="true" pageId="5" pageNumber="1824">R. shraddhanandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFB92E00FC93244FFBE7BA67" box="[983,1061,1375,1398]" pageId="5" pageNumber="1824" rank="species">sp. nov.</taxonomicNameLabel>
, paratype ♀ (PCL 5.8 mm, CW 4.9 mm), IO/SS/ANO/00140; (c)
<taxonomicName id="9D954D35FFB92E00FDE5246CFC71BA82" authority="Macpherson, 2009" authorityName="Macpherson" authorityYear="2009" box="[673,947,1404,1427]" class="Malacostraca" family="Munididae" genus="Raymunida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="1824" phylum="Arthropoda" rank="species" species="vittata">
<emphasis id="68E1EAA4FFB92E00FDE5246CFD3ABA82" box="[673,760,1404,1427]" italics="true" pageId="5" pageNumber="1824">R. vittata</emphasis>
<bibRefCitation id="3E044B47FFB92E00FDBB246CFC71BA82" author="Macpherson E" box="[767,947,1404,1427]" pageId="5" pageNumber="1824" pagination="431 - 451" refId="ref9854" refString="Macpherson E. 2009. New species of squat lobsters of the genera Munida and Raymunida (Crustacea, Decapoda, Galatheidae) from Vanuatu and New Caledonia. Zoosystema. 31 (3): 431 - 451. doi: 10. 5252 / z 2009 n 3 a 3." type="journal article" year="2009">Macpherson, 2009</bibRefCitation>
</taxonomicName>
, ♀ (PCL 9.0 mm, CW 7.7 mm), IO/SS/ANO/00142; (d)
<taxonomicName id="9D954D35FFB92E00FE972489FD53BAA1" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[467,657,1433,1456]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="1824" phylum="Arthropoda" rank="species" species="bengala" status="sp. nov.">
<emphasis id="68E1EAA4FFB92E00FE972489FD53BAA1" box="[467,657,1433,1456]" italics="true" pageId="5" pageNumber="1824">Munidopsis bengala</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFB92E00FDD32489FD27BAA1" box="[663,741,1433,1456]" pageId="5" pageNumber="1824" rank="species">sp. nov.</taxonomicNameLabel>
, holotype ♀ (PCL 8.0 mm, CW 7.7 mm), IO/ SS/ANO/00145; (e)
<taxonomicName id="9D954D35FFB92E00FE1924A7FE72BADF" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[349,432,1463,1486]" class="Malacostraca" family="Munidopsidae" genus="Munidopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="1824" phylum="Arthropoda" rank="species" species="kadal" status="sp. nov.">
<emphasis id="68E1EAA4FFB92E00FE1924A7FE72BADF" box="[349,432,1463,1486]" italics="true" pageId="5" pageNumber="1824">M. kadal</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFB92E00FEF324A7FDC4BADF" box="[439,518,1463,1486]" pageId="5" pageNumber="1824" rank="species">sp. nov.</taxonomicNameLabel>
, holotype ♂ (PCL 5.2 mm, CW 4.9 mm), IO/SS/ANO/00146. Scale bars: 2.0 mm.
</paragraph>
</caption>
<subSubSection id="128F653DFFB92E0CFFE42738FB90BADC" lastPageId="9" lastPageNumber="1828" pageId="5" pageNumber="1824" type="description">
<paragraph id="5A2A36B6FFB92E00FFE42738FE74B953" blockId="5.[160,1156,1576,1706]" box="[160,438,1576,1602]" pageId="5" pageNumber="1824">
<emphasis id="68E1EAA4FFB92E00FFE42738FE74B953" bold="true" box="[160,438,1576,1602]" italics="true" pageId="5" pageNumber="1824">
Description of
<typeStatus id="852E8814FFB92E00FE0A2738FE74B953" box="[334,438,1576,1602]" pageId="5" pageNumber="1824" type="holotype">holotype</typeStatus>
</emphasis>
</paragraph>
<paragraph id="5A2A36B6FFB92E0DFFE4275CFC58BE44" blockId="5.[160,1156,1576,1706]" lastBlockId="8.[160,1156,142,1727]" lastPageId="8" lastPageNumber="1827" pageId="5" pageNumber="1824">
Carapace (
<figureCitation id="C2AE2A33FFB92E00FE5E275CFE44B974" box="[282,390,1612,1637]" captionStart="Figure 2" captionStartId="4.[160,225,957,980]" captionTargetBox="[163,1152,142,925]" captionTargetId="figure-248@4.[163,1152,142,925]" captionTargetPageId="4" captionText="Figure 2. Habitus,dorsal view.(a) Raymunida shraddhanandi sp.nov., holotype ♂ (PCL 9.7 mm,CW 8.2 mm), IO/SS/ANO/00141;(b) R.shraddhanandi sp.nov.,paratype ♀ (PCL 5.8 mm,CW 4.9 mm),IO/SS/ANO/00140;(c) R. vittata Macpherson, 2009, ♀ (PCL 9.0 mm, CW 7.7 mm), IO/SS/ANO/00142. Scale bars: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7380768" httpUri="https://zenodo.org/record/7380768/files/figure.png" pageId="5" pageNumber="1824">Figures 2</figureCitation>
(a), 2(a), 2(a, 2)) (exclusive of rostrum) 1.2 times as long as wide, bearing a few long iridescent setae dorsally; transverse striae with dense short setae; gastric region with 6 transverse striae and 5 pairs of spines (epigastric spines); cardiac region with 2 complete transverse striae followed by 2 median striae and 1 pair of lateral striae; intestinal region with 3 transverse striae, first and third complete, second interrupted medially. One parahepatic and 1 postcervical spine present on each side; anterior branchial spines absent. Frontal margin moderately oblique, with 1 small spine between supraocular spine and anterolateral spine. Lateral margin gently convex. Anterolateral spine distinct, but falling far short of level of sinus between rostrum and supraocular spine, followed by small spine anterior to cervical groove. Anterior and posterior branchial margins each with 2 spines, posteriormost spine smallest. Rostrum spiniform, 0.4 times PCL, 2 times longer than supraocular spines, directed slightly downward. Supraocular spines horizontal, subparallel, overreaching distal margin of cornea.
</paragraph>
<caption id="0EEA663EFFBA2E03FFE424F9FC6DB9A1" ID-DOI="http://doi.org/10.5281/zenodo.7380774" ID-Zenodo-Dep="7380774" httpUri="https://zenodo.org/record/7380774/files/figure.png" pageId="6" pageNumber="1825" startId="6.[160,225,1513,1536]" targetBox="[164,1151,143,1481]" targetPageId="6">
<paragraph id="5A2A36B6FFBA2E03FFE424F9FC6DB9A1" blockId="6.[160,1156,1513,1712]" pageId="6" pageNumber="1825">
<emphasis id="68E1EAA4FFBA2E03FFE424F9FF3FB911" bold="true" box="[160,253,1513,1536]" pageId="6" pageNumber="1825">Figure 4.</emphasis>
<taxonomicName id="9D954D35FFBA2E03FE4324F9FDCCB911" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[263,526,1513,1536]" class="Malacostraca" family="Munididae" genus="Raymunida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="1825" phylum="Arthropoda" rank="species" species="shraddhanandi" status="sp. nov.">
<emphasis id="68E1EAA4FFBA2E03FE4324F9FDCCB911" box="[263,526,1513,1536]" italics="true" pageId="6" pageNumber="1825">Raymunida shraddhanandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFBA2E03FD5324F9FDABB911" box="[535,617,1513,1536]" pageId="6" pageNumber="1825" rank="species">sp. nov.</taxonomicNameLabel>
, holotype ♂ (PCL 9.7 mm, CW 8.2 mm), IO/SS/ANO/ 00141. (a) carapace (dorsal view); (b) carapace (left lateral view); (c) thoracic sternum (ventral view); (d) pleomeres 15 (dorsal view); (e) pleomeres 5 and 6 and telson (external view); (f) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (g) left Mxp3 (lateral view); (h) left P1 merus and carpus (dorsal view); (i) left P1 chela (dorsal view); (j) right P2 (lateral view); (k) right P3 (lateral view); (l), right P4 (lateral view); (m) right P5 dactylus (lateral view); (n) left G1 (internal view); (o) left G2 (external view). Scale bars: a, cl, o = 1.0 mm; b = 2.0 mm; m, n = 0.5 mm.
</paragraph>
</caption>
<caption id="0EEA663EFFBB2E02FFE42473FBA5B91C" ID-DOI="http://doi.org/10.5281/zenodo.7380776" ID-Zenodo-Dep="7380776" httpUri="https://zenodo.org/record/7380776/files/figure.png" pageId="7" pageNumber="1826" startId="7.[160,225,1379,1402]" targetBox="[165,1150,143,1346]" targetPageId="7">
<paragraph id="5A2A36B6FFBB2E02FFE42473FBA5B91C" blockId="7.[160,1157,1379,1549]" pageId="7" pageNumber="1826">
<emphasis id="68E1EAA4FFBB2E02FFE42473FF38BA6B" bold="true" box="[160,250,1379,1402]" pageId="7" pageNumber="1826">Figure 5.</emphasis>
<taxonomicName id="9D954D35FFBB2E02FE452473FE3CBA6B" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[257,510,1379,1402]" class="Malacostraca" family="Munididae" genus="Raymunida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="1826" phylum="Arthropoda" rank="species" species="shraddhanandi" status="sp. nov.">
<emphasis id="68E1EAA4FFBB2E02FE452473FE3CBA6B" box="[257,510,1379,1402]" italics="true" pageId="7" pageNumber="1826">Raymunida shraddhanandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFBB2E02FD402473FD93BA6B" box="[516,593,1379,1402]" pageId="7" pageNumber="1826" rank="species">sp. nov.</taxonomicNameLabel>
, paratype ♀ (PCL 5.8 mm, CW 4.9 mm), IO/SS/ANO/00140. (a) carapace, pleon and cephalic appendages (dorsal view); (b) thoracic sternum (ventral view); (c) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (d) right Mxp3 (lateral view); (e) right P1 (dorsal view); (f) left P2 propodus and dactylus (lateral view); (g) left P2 merus and carpus (lateral view); (h) left P3 propodus and dactylus (lateral view); (i) left P3 merus and carpus (lateral view); (j) left P4 propodus and dactylus (lateral view); (k) left P4 merus and carpus (lateral view). Scale bars: 1.0 mm.
</paragraph>
</caption>
<paragraph id="5A2A36B6FFB42E0DFF84204EFC88BD34" blockId="8.[160,1156,142,1727]" pageId="8" pageNumber="1827">
Thoracic sternite 3 (
<figureCitation id="C2AE2A33FFB42E0DFEE9204EFDCCBE66" box="[429,526,350,375]" captionStart="Figure 4" captionStartId="6.[160,225,1513,1536]" captionTargetBox="[164,1151,143,1481]" captionTargetId="figure-1@6.[163,1152,142,1482]" captionTargetPageId="6" captionText="Figure 4. Raymunida shraddhanandi sp. nov., holotype ♂ (PCL 9.7 mm, CW 8.2 mm), IO/SS/ANO/ 00141. (a) carapace (dorsal view); (b) carapace (left lateral view); (c) thoracic sternum (ventral view); (d) pleomeres 15 (dorsal view); (e) pleomeres 5 and 6 and telson (external view); (f) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (g) left Mxp3 (lateral view); (h) left P1 merus and carpus (dorsal view); (i) left P1 chela (dorsal view); (j) right P2 (lateral view); (k) right P3 (lateral view); (l), right P4 (lateral view); (m) right P5 dactylus (lateral view); (n) left G1 (internal view); (o) left G2 (external view). Scale bars: a, cl, o = 1.0 mm; b = 2.0 mm; m, n = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380774" httpUri="https://zenodo.org/record/7380774/files/figure.png" pageId="8" pageNumber="1827">Figure 4</figureCitation>
(c)) 3.3 times as wide as long, with 2 flattened lobes separated by shallow V-shaped notch on anterior margin, anterior margins of lobes faintly granulate, lateral angle with distinct acute projection. Sternite 4 with anterior margin as wide as sternite 3, with 2 transverse striae, anterior stria medially interrupted, bearing sparse setae, posterior stria uninterrupted. Sternites 5 and 6 with short oblique stria on each lateral portion. Sternite 7 with few short striae laterally.
</paragraph>
<paragraph id="5A2A36B6FFB42E0DFF84233EFC9EBDC3" blockId="8.[160,1156,142,1727]" pageId="8" pageNumber="1827">
Pleonal tergites (
<figureCitation id="C2AE2A33FFB42E0DFEC7233EFE21BD56" box="[387,483,558,583]" captionStart="Figure 4" captionStartId="6.[160,225,1513,1536]" captionTargetBox="[164,1151,143,1481]" captionTargetId="figure-1@6.[163,1152,142,1482]" captionTargetPageId="6" captionText="Figure 4. Raymunida shraddhanandi sp. nov., holotype ♂ (PCL 9.7 mm, CW 8.2 mm), IO/SS/ANO/ 00141. (a) carapace (dorsal view); (b) carapace (left lateral view); (c) thoracic sternum (ventral view); (d) pleomeres 15 (dorsal view); (e) pleomeres 5 and 6 and telson (external view); (f) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (g) left Mxp3 (lateral view); (h) left P1 merus and carpus (dorsal view); (i) left P1 chela (dorsal view); (j) right P2 (lateral view); (k) right P3 (lateral view); (l), right P4 (lateral view); (m) right P5 dactylus (lateral view); (n) left G1 (internal view); (o) left G2 (external view). Scale bars: a, cl, o = 1.0 mm; b = 2.0 mm; m, n = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380774" httpUri="https://zenodo.org/record/7380774/files/figure.png" pageId="8" pageNumber="1827">Figure 4</figureCitation>
(d)) with few moderately long iridescent setae and row of short plumose setae on transverse ridges; tergites 2 and 3 each with row of short, interrupted striae between 2 transverse ridges, tergite 4 without striae between 2 transverse ridges; tergites 5 and 6 with 2 medially interrupted ridges, those on tergite 5 longer and nearly straight, those on tergite 6 somewhat squamiform.
</paragraph>
<paragraph id="5A2A36B6FFB42E0DFF8423CCFDADBC2B" blockId="8.[160,1156,142,1727]" pageId="8" pageNumber="1827">
Eye (
<figureCitation id="C2AE2A33FFB42E0DFFB123CCFE8DBDE4" box="[245,335,732,757]" captionStart="Figure 4" captionStartId="6.[160,225,1513,1536]" captionTargetBox="[164,1151,143,1481]" captionTargetId="figure-1@6.[163,1152,142,1482]" captionTargetPageId="6" captionText="Figure 4. Raymunida shraddhanandi sp. nov., holotype ♂ (PCL 9.7 mm, CW 8.2 mm), IO/SS/ANO/ 00141. (a) carapace (dorsal view); (b) carapace (left lateral view); (c) thoracic sternum (ventral view); (d) pleomeres 15 (dorsal view); (e) pleomeres 5 and 6 and telson (external view); (f) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (g) left Mxp3 (lateral view); (h) left P1 merus and carpus (dorsal view); (i) left P1 chela (dorsal view); (j) right P2 (lateral view); (k) right P3 (lateral view); (l), right P4 (lateral view); (m) right P5 dactylus (lateral view); (n) left G1 (internal view); (o) left G2 (external view). Scale bars: a, cl, o = 1.0 mm; b = 2.0 mm; m, n = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380774" httpUri="https://zenodo.org/record/7380774/files/figure.png" pageId="8" pageNumber="1827">Figure 4</figureCitation>
(a)) moderately large, corneal diameter 0.3 times distance between mesial bases of anterolateral spines, eye lashes simple and relatively short, long stout seta on rounded dorsodistal margin of peduncle.
</paragraph>
<paragraph id="5A2A36B6FFB42E0DFF842254FD9DBCD4" blockId="8.[160,1156,142,1727]" pageId="8" pageNumber="1827">
Antennular peduncle article 1 (
<figureCitation id="C2AE2A33FFB42E0DFD712254FD5BBC4C" box="[565,665,836,861]" captionStart="Figure 4" captionStartId="6.[160,225,1513,1536]" captionTargetBox="[164,1151,143,1481]" captionTargetId="figure-1@6.[163,1152,142,1482]" captionTargetPageId="6" captionText="Figure 4. Raymunida shraddhanandi sp. nov., holotype ♂ (PCL 9.7 mm, CW 8.2 mm), IO/SS/ANO/ 00141. (a) carapace (dorsal view); (b) carapace (left lateral view); (c) thoracic sternum (ventral view); (d) pleomeres 15 (dorsal view); (e) pleomeres 5 and 6 and telson (external view); (f) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (g) left Mxp3 (lateral view); (h) left P1 merus and carpus (dorsal view); (i) left P1 chela (dorsal view); (j) right P2 (lateral view); (k) right P3 (lateral view); (l), right P4 (lateral view); (m) right P5 dactylus (lateral view); (n) left G1 (internal view); (o) left G2 (external view). Scale bars: a, cl, o = 1.0 mm; b = 2.0 mm; m, n = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380774" httpUri="https://zenodo.org/record/7380774/files/figure.png" pageId="8" pageNumber="1827">Figure 4</figureCitation>
(f)) (distal spines excluded) only reaching proximal margin of cornea; distomesial spine distinctly shorter than distolateral spine; 2 lateral spines, anterior spine distinctly overreaching tip of distolateral spine, posterior spine much shorter than anterior spine.
</paragraph>
<paragraph id="5A2A36B6FFB42E0DFF8422DEFE74BB63" blockId="8.[160,1156,142,1727]" pageId="8" pageNumber="1827">
Antennal peduncle article 1 (
<figureCitation id="C2AE2A33FFB42E0DFD5222DEFDBABCF6" box="[534,632,974,999]" captionStart="Figure 4" captionStartId="6.[160,225,1513,1536]" captionTargetBox="[164,1151,143,1481]" captionTargetId="figure-1@6.[163,1152,142,1482]" captionTargetPageId="6" captionText="Figure 4. Raymunida shraddhanandi sp. nov., holotype ♂ (PCL 9.7 mm, CW 8.2 mm), IO/SS/ANO/ 00141. (a) carapace (dorsal view); (b) carapace (left lateral view); (c) thoracic sternum (ventral view); (d) pleomeres 15 (dorsal view); (e) pleomeres 5 and 6 and telson (external view); (f) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (g) left Mxp3 (lateral view); (h) left P1 merus and carpus (dorsal view); (i) left P1 chela (dorsal view); (j) right P2 (lateral view); (k) right P3 (lateral view); (l), right P4 (lateral view); (m) right P5 dactylus (lateral view); (n) left G1 (internal view); (o) left G2 (external view). Scale bars: a, cl, o = 1.0 mm; b = 2.0 mm; m, n = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380774" httpUri="https://zenodo.org/record/7380774/files/figure.png" pageId="8" pageNumber="1827">Figure 4</figureCitation>
(f)) with long distomesial spine slightly overreaching distal margin of article 3, not reaching distal margin of antennular article 1; article 2 with distomesial spine reaching distal margin of article 3, distolateral spine nearly reaching distal margin of article 3, mesial margin unarmed; article 3 with short distomesial spine; article 4 unarmed.
</paragraph>
<paragraph id="5A2A36B6FFB42E0DFF84256CFDBDBA0E" blockId="8.[160,1156,142,1727]" pageId="8" pageNumber="1827">
Mxp3 (
<figureCitation id="C2AE2A33FFB42E0DFE48256CFEA5BB84" box="[268,359,1148,1173]" captionStart="Figure 4" captionStartId="6.[160,225,1513,1536]" captionTargetBox="[164,1151,143,1481]" captionTargetId="figure-1@6.[163,1152,142,1482]" captionTargetPageId="6" captionText="Figure 4. Raymunida shraddhanandi sp. nov., holotype ♂ (PCL 9.7 mm, CW 8.2 mm), IO/SS/ANO/ 00141. (a) carapace (dorsal view); (b) carapace (left lateral view); (c) thoracic sternum (ventral view); (d) pleomeres 15 (dorsal view); (e) pleomeres 5 and 6 and telson (external view); (f) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (g) left Mxp3 (lateral view); (h) left P1 merus and carpus (dorsal view); (i) left P1 chela (dorsal view); (j) right P2 (lateral view); (k) right P3 (lateral view); (l), right P4 (lateral view); (m) right P5 dactylus (lateral view); (n) left G1 (internal view); (o) left G2 (external view). Scale bars: a, cl, o = 1.0 mm; b = 2.0 mm; m, n = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380774" httpUri="https://zenodo.org/record/7380774/files/figure.png" pageId="8" pageNumber="1827">Figure 4</figureCitation>
(g)) ischium subequal in length to merus measured along dorsal margin, with strong spine each at dorsodistal and ventrodistal angles; crista dentata consisting of 33 denticles. Merus with small dorsodistal spine; ventral margin with 3 spines, second spine much smaller than other 2 slender main prominent spines. Carpus with 2 spines distally on ventral margin. Epipod present.
</paragraph>
<paragraph id="5A2A36B6FFB42E0CFF842439FDB5BE44" blockId="8.[160,1156,142,1727]" lastBlockId="9.[160,1156,142,930]" lastPageId="9" lastPageNumber="1828" pageId="8" pageNumber="1827">
P1 (
<figureCitation id="C2AE2A33FFB42E0DFFB42439FE94BA53" box="[240,342,1321,1346]" captionStart="Figure 4" captionStartId="6.[160,225,1513,1536]" captionTargetBox="[164,1151,143,1481]" captionTargetId="figure-1@6.[163,1152,142,1482]" captionTargetPageId="6" captionText="Figure 4. Raymunida shraddhanandi sp. nov., holotype ♂ (PCL 9.7 mm, CW 8.2 mm), IO/SS/ANO/ 00141. (a) carapace (dorsal view); (b) carapace (left lateral view); (c) thoracic sternum (ventral view); (d) pleomeres 15 (dorsal view); (e) pleomeres 5 and 6 and telson (external view); (f) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (g) left Mxp3 (lateral view); (h) left P1 merus and carpus (dorsal view); (i) left P1 chela (dorsal view); (j) right P2 (lateral view); (k) right P3 (lateral view); (l), right P4 (lateral view); (m) right P5 dactylus (lateral view); (n) left G1 (internal view); (o) left G2 (external view). Scale bars: a, cl, o = 1.0 mm; b = 2.0 mm; m, n = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380774" httpUri="https://zenodo.org/record/7380774/files/figure.png" pageId="8" pageNumber="1827">Figure 4</figureCitation>
(h,)) subequal from right to left, massive, 3.6 times PCL, moderately depressed, with numerous long, iridescent setae; surfaces of merus, carpus and palm of chela squamate. Merus 1.1 times PCL; dorsal and ventral surfaces each with 2 irregular rows of spines, distomesial spine strongest; mesial surface with 2 spines distal to mid length, distal spine much larger than proximal spine; lateral surface with 1 subdistal spine, ventrolateral margin with 1 small distal spine. Carpus with 2 irregular rows of spines on dorsal surface and with 1 spine each on ventral surface, mesial margin and lateral margin; mesial margin also with elongated spine on distal one-third. Chela 4.6 (left) or 4.8 (right) times as wide as long. Palm 1.3 times longer than carpus, 2.1 (right) or 2.3 (left) times as long as wide; dorsal surface with 2 irregular rows of spines and small spine at dactylar articulation; ventral surface with 1 mesial spine at mid length; mesial surface with longitudinal row of 4 spines; lateral margin with row of sparsely arranged spines extending to entire length of fixed finger. Fingers distally broken, but distinctly longer than palm at least, each with rounded longitudinal crest on dorsal surface; fixed finger curved on proximal half; dactylus with 3 spines on proximal one-third portion, 1 median spine and small, unevenly spaced spinules on distal portion of mesial margin; occlusal margins minutely dentate, dactylus with blunt teeth on proximal one-third fitting into large concavity on proximal half of fixed finger.
</paragraph>
<paragraph id="5A2A36B6FFB52E0CFF84204EFC67BC06" blockId="9.[160,1156,142,930]" pageId="9" pageNumber="1828">
P24 (
<figureCitation id="C2AE2A33FFB52E0CFE47204EFE99BE66" box="[259,347,350,375]" captionStart="Figure 4" captionStartId="6.[160,225,1513,1536]" captionTargetBox="[164,1151,143,1481]" captionTargetId="figure-1@6.[163,1152,142,1482]" captionTargetPageId="6" captionText="Figure 4. Raymunida shraddhanandi sp. nov., holotype ♂ (PCL 9.7 mm, CW 8.2 mm), IO/SS/ANO/ 00141. (a) carapace (dorsal view); (b) carapace (left lateral view); (c) thoracic sternum (ventral view); (d) pleomeres 15 (dorsal view); (e) pleomeres 5 and 6 and telson (external view); (f) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (g) left Mxp3 (lateral view); (h) left P1 merus and carpus (dorsal view); (i) left P1 chela (dorsal view); (j) right P2 (lateral view); (k) right P3 (lateral view); (l), right P4 (lateral view); (m) right P5 dactylus (lateral view); (n) left G1 (internal view); (o) left G2 (external view). Scale bars: a, cl, o = 1.0 mm; b = 2.0 mm; m, n = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380774" httpUri="https://zenodo.org/record/7380774/files/figure.png" pageId="9" pageNumber="1828">Figure 4</figureCitation>
(j)) somewhat compressed laterally, 2.1 (P2, P3) and 1.7 (P4) times PCL; P4 mero-carpal articulation reaching only level of first branchial spine (lateral end of anterior cervical groove of carapace) (
<figureCitation id="C2AE2A33FFB52E0CFE9A20B4FDF5BEAC" box="[478,567,420,445]" captionStart="Figure 2" captionStartId="4.[160,225,957,980]" captionTargetBox="[163,1152,142,925]" captionTargetId="figure-248@4.[163,1152,142,925]" captionTargetPageId="4" captionText="Figure 2. Habitus,dorsal view.(a) Raymunida shraddhanandi sp.nov., holotype ♂ (PCL 9.7 mm,CW 8.2 mm), IO/SS/ANO/00141;(b) R.shraddhanandi sp.nov.,paratype ♀ (PCL 5.8 mm,CW 4.9 mm),IO/SS/ANO/00140;(c) R. vittata Macpherson, 2009, ♀ (PCL 9.0 mm, CW 7.7 mm), IO/SS/ANO/00142. Scale bars: 2.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7380768" httpUri="https://zenodo.org/record/7380768/files/figure.png" pageId="9" pageNumber="1828">Figure 2</figureCitation>
(a)); meri 6.3 (P2), 4.5 (P3) and 3.5 (P4) times as long as wide, 0.9 (P2), 0.8 (P3) and 0.6 (P4) times PCL, 3.0 (P2), 2.6 (P3) and 2.2 (P4) times longer than carpi; propodi 7.5 (P2), 6.7 (P3) and 5.7 (P4) times as long as wide, 2.1 (P2), 2.0 (P3) and 1.9 (P4) times longer than carpi, 2.3 (P2), 2.2 (P3) and 2.0 (P4) times longer than dactyli. Meri covered with distinct squamiform ridges on lateral and mesial surfaces; dorsal margins each with row of 10 (P2), 6 (P3) and 3 (P4) spines randomly arranged and increasing in size distally; lateral surfaces each with dorsal row of 3 (P2), 9 (P3) and 8 (P4) spines; ventral margins with 4 (P2), 3 (P3) and 2 (P4) spines on distal one-third. Carpi with 5 (P2), 4 (P3) and 1 (P4) dorsal spines; ventral margins each with 1 distal spine. Propodi dorsally unarmed; ventral margins with 7 (P2), 4 (P3) and 4 (P4) slender corneous spines. Dactyli with 6 (P2), 5 (P3) and 4 (P4) slender corneous spines on ventral margins. Epipods present on P13.
</paragraph>
<paragraph id="5A2A36B6FFB52E0CFF842231FCAEBC2B" blockId="9.[160,1156,142,930]" box="[192,876,801,826]" pageId="9" pageNumber="1828">
Male with G1 (
<figureCitation id="C2AE2A33FFB52E0CFE202231FE01BC2B" box="[356,451,801,826]" captionStart="Figure 4" captionStartId="6.[160,225,1513,1536]" captionTargetBox="[164,1151,143,1481]" captionTargetId="figure-1@6.[163,1152,142,1482]" captionTargetPageId="6" captionText="Figure 4. Raymunida shraddhanandi sp. nov., holotype ♂ (PCL 9.7 mm, CW 8.2 mm), IO/SS/ANO/ 00141. (a) carapace (dorsal view); (b) carapace (left lateral view); (c) thoracic sternum (ventral view); (d) pleomeres 15 (dorsal view); (e) pleomeres 5 and 6 and telson (external view); (f) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (g) left Mxp3 (lateral view); (h) left P1 merus and carpus (dorsal view); (i) left P1 chela (dorsal view); (j) right P2 (lateral view); (k) right P3 (lateral view); (l), right P4 (lateral view); (m) right P5 dactylus (lateral view); (n) left G1 (internal view); (o) left G2 (external view). Scale bars: a, cl, o = 1.0 mm; b = 2.0 mm; m, n = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380774" httpUri="https://zenodo.org/record/7380774/files/figure.png" pageId="9" pageNumber="1828">Figure 4</figureCitation>
(n)) and G2 (
<figureCitation id="C2AE2A33FFB52E0CFD092231FD69BC2B" box="[589,683,801,826]" captionStart="Figure 4" captionStartId="6.[160,225,1513,1536]" captionTargetBox="[164,1151,143,1481]" captionTargetId="figure-1@6.[163,1152,142,1482]" captionTargetPageId="6" captionText="Figure 4. Raymunida shraddhanandi sp. nov., holotype ♂ (PCL 9.7 mm, CW 8.2 mm), IO/SS/ANO/ 00141. (a) carapace (dorsal view); (b) carapace (left lateral view); (c) thoracic sternum (ventral view); (d) pleomeres 15 (dorsal view); (e) pleomeres 5 and 6 and telson (external view); (f) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (g) left Mxp3 (lateral view); (h) left P1 merus and carpus (dorsal view); (i) left P1 chela (dorsal view); (j) right P2 (lateral view); (k) right P3 (lateral view); (l), right P4 (lateral view); (m) right P5 dactylus (lateral view); (n) left G1 (internal view); (o) left G2 (external view). Scale bars: a, cl, o = 1.0 mm; b = 2.0 mm; m, n = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380774" httpUri="https://zenodo.org/record/7380774/files/figure.png" pageId="9" pageNumber="1828">Figure 4</figureCitation>
(o)) as illustrated.
</paragraph>
<paragraph id="5A2A36B6FFB52E0CFF842254FB87BCB3" blockId="9.[160,1156,142,930]" pageId="9" pageNumber="1828">
Uropodal protopod (
<figureCitation id="C2AE2A33FFB52E0CFEEC2254FDC1BC4C" box="[424,515,836,861]" captionStart="Figure 4" captionStartId="6.[160,225,1513,1536]" captionTargetBox="[164,1151,143,1481]" captionTargetId="figure-1@6.[163,1152,142,1482]" captionTargetPageId="6" captionText="Figure 4. Raymunida shraddhanandi sp. nov., holotype ♂ (PCL 9.7 mm, CW 8.2 mm), IO/SS/ANO/ 00141. (a) carapace (dorsal view); (b) carapace (left lateral view); (c) thoracic sternum (ventral view); (d) pleomeres 15 (dorsal view); (e) pleomeres 5 and 6 and telson (external view); (f) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (g) left Mxp3 (lateral view); (h) left P1 merus and carpus (dorsal view); (i) left P1 chela (dorsal view); (j) right P2 (lateral view); (k) right P3 (lateral view); (l), right P4 (lateral view); (m) right P5 dactylus (lateral view); (n) left G1 (internal view); (o) left G2 (external view). Scale bars: a, cl, o = 1.0 mm; b = 2.0 mm; m, n = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.7380774" httpUri="https://zenodo.org/record/7380774/files/figure.png" pageId="9" pageNumber="1828">Figure 4</figureCitation>
(e)) with 1 distal spine and proximal produced portion on lateral margin; endopod broader than exopod, lateral and distal margins each with row of spines (lateral spines damaged); exopod also spinose on lateral and distal margins.
</paragraph>
<paragraph id="5A2A36B6FFB52E0CFFE422DEFEDBBCF9" blockId="9.[160,1156,974,1485]" box="[160,281,974,1000]" pageId="9" pageNumber="1828">
<emphasis id="68E1EAA4FFB52E0CFFE422DEFEDBBCF9" bold="true" box="[160,281,974,1000]" italics="true" pageId="9" pageNumber="1828">Variations</emphasis>
</paragraph>
<paragraph id="5A2A36B6FFB52E0CFFE422E1FB90BADC" blockId="9.[160,1156,974,1485]" pageId="9" pageNumber="1828">
The female
<typeStatus id="852E8814FFB52E0CFE6722E1FE47BB1B" box="[291,389,1009,1034]" pageId="9" pageNumber="1828" type="paratype">paratype</typeStatus>
differs from the male
<typeStatus id="852E8814FFB52E0CFD3B22E1FD23BB1B" box="[639,737,1009,1034]" pageId="9" pageNumber="1828" type="holotype">holotype</typeStatus>
in the following points, although the difference in the P1 represents sexual dimorphism. Carapace (
<figureCitation id="C2AE2A33FFB52E0CFC7B2504FC55BB3C" box="[831,919,1044,1069]" captionStart="Figure 5" captionStartId="7.[160,225,1379,1402]" captionTargetBox="[165,1150,143,1346]" captionTargetId="figure-56@7.[164,1151,142,1347]" captionTargetPageId="7" captionText="Figure 5. Raymunida shraddhanandi sp. nov., paratype ♀ (PCL 5.8 mm, CW 4.9 mm), IO/SS/ANO/00140. (a) carapace, pleon and cephalic appendages (dorsal view); (b) thoracic sternum (ventral view); (c) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (d) right Mxp3 (lateral view); (e) right P1 (dorsal view);(f) left P2 propodus and dactylus (lateral view); (g) left P2 merus and carpus (lateral view);(h) left P3 propodus and dactylus (lateral view); (i) left P3 merus and carpus (lateral view);(j) left P4 propodus and dactylus (lateral view); (k) left P4 merus and carpus (lateral view). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7380776" httpUri="https://zenodo.org/record/7380776/files/figure.png" pageId="9" pageNumber="1828">Figure 5</figureCitation>
(a)) bearing 1 anterior branchial spine on dorsal surface. Thoracic sternites 57 (
<figureCitation id="C2AE2A33FFB52E0CFC782527FC5FBB41" box="[828,925,1079,1104]" captionStart="Figure 5" captionStartId="7.[160,225,1379,1402]" captionTargetBox="[165,1150,143,1346]" captionTargetId="figure-56@7.[164,1151,142,1347]" captionTargetPageId="7" captionText="Figure 5. Raymunida shraddhanandi sp. nov., paratype ♀ (PCL 5.8 mm, CW 4.9 mm), IO/SS/ANO/00140. (a) carapace, pleon and cephalic appendages (dorsal view); (b) thoracic sternum (ventral view); (c) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (d) right Mxp3 (lateral view); (e) right P1 (dorsal view);(f) left P2 propodus and dactylus (lateral view); (g) left P2 merus and carpus (lateral view);(h) left P3 propodus and dactylus (lateral view); (i) left P3 merus and carpus (lateral view);(j) left P4 propodus and dactylus (lateral view); (k) left P4 merus and carpus (lateral view). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7380776" httpUri="https://zenodo.org/record/7380776/files/figure.png" pageId="9" pageNumber="1828">Figure 5</figureCitation>
(b)) smooth. Pleonal tergites 2 and 3 (
<figureCitation id="C2AE2A33FFB52E0CFE262549FE7FBB63" box="[354,445,1113,1138]" captionStart="Figure 5" captionStartId="7.[160,225,1379,1402]" captionTargetBox="[165,1150,143,1346]" captionTargetId="figure-56@7.[164,1151,142,1347]" captionTargetPageId="7" captionText="Figure 5. Raymunida shraddhanandi sp. nov., paratype ♀ (PCL 5.8 mm, CW 4.9 mm), IO/SS/ANO/00140. (a) carapace, pleon and cephalic appendages (dorsal view); (b) thoracic sternum (ventral view); (c) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (d) right Mxp3 (lateral view); (e) right P1 (dorsal view);(f) left P2 propodus and dactylus (lateral view); (g) left P2 merus and carpus (lateral view);(h) left P3 propodus and dactylus (lateral view); (i) left P3 merus and carpus (lateral view);(j) left P4 propodus and dactylus (lateral view); (k) left P4 merus and carpus (lateral view). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7380776" httpUri="https://zenodo.org/record/7380776/files/figure.png" pageId="9" pageNumber="1828">Figure 5</figureCitation>
(a)) without striae. Antennal peduncle article 2 with distomesial spine feebly overreaching distal margin of article 3 (
<figureCitation id="C2AE2A33FFB52E0CFDA6256CFCFEBB84" box="[738,828,1148,1173]" captionStart="Figure 5" captionStartId="7.[160,225,1379,1402]" captionTargetBox="[165,1150,143,1346]" captionTargetId="figure-56@7.[164,1151,142,1347]" captionTargetPageId="7" captionText="Figure 5. Raymunida shraddhanandi sp. nov., paratype ♀ (PCL 5.8 mm, CW 4.9 mm), IO/SS/ANO/00140. (a) carapace, pleon and cephalic appendages (dorsal view); (b) thoracic sternum (ventral view); (c) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (d) right Mxp3 (lateral view); (e) right P1 (dorsal view);(f) left P2 propodus and dactylus (lateral view); (g) left P2 merus and carpus (lateral view);(h) left P3 propodus and dactylus (lateral view); (i) left P3 merus and carpus (lateral view);(j) left P4 propodus and dactylus (lateral view); (k) left P4 merus and carpus (lateral view). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7380776" httpUri="https://zenodo.org/record/7380776/files/figure.png" pageId="9" pageNumber="1828">Figure 5</figureCitation>
(c)). Mxp3 (
<figureCitation id="C2AE2A33FFB52E0CFCFC256CFBD6BB84" box="[952,1044,1148,1173]" captionStart="Figure 5" captionStartId="7.[160,225,1379,1402]" captionTargetBox="[165,1150,143,1346]" captionTargetId="figure-56@7.[164,1151,142,1347]" captionTargetPageId="7" captionText="Figure 5. Raymunida shraddhanandi sp. nov., paratype ♀ (PCL 5.8 mm, CW 4.9 mm), IO/SS/ANO/00140. (a) carapace, pleon and cephalic appendages (dorsal view); (b) thoracic sternum (ventral view); (c) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (d) right Mxp3 (lateral view); (e) right P1 (dorsal view);(f) left P2 propodus and dactylus (lateral view); (g) left P2 merus and carpus (lateral view);(h) left P3 propodus and dactylus (lateral view); (i) left P3 merus and carpus (lateral view);(j) left P4 propodus and dactylus (lateral view); (k) left P4 merus and carpus (lateral view). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7380776" httpUri="https://zenodo.org/record/7380776/files/figure.png" pageId="9" pageNumber="1828">Figure 5</figureCitation>
(d)) merus with only 2 main spines on ventral margin, carpus with 1 distal spine on ventral margin. P1 slender, 3.1 times PCL, chela 7.5 times as long as wide; palm subequal in length to carpus, ventral surface with 2 irregular longitudinal rows of spines, lateral margin with row of spines extending to distal one-fourth of fixed finger; fingers about 1.9 times as long as palm, parallel to each other, occlusal margins minutely dentate, each with slightly larger acute teeth at regular intervals. P24 (
<figureCitation id="C2AE2A33FFB52E0CFD46245CFD9CBA74" box="[514,606,1356,1381]" captionStart="Figure 5" captionStartId="7.[160,225,1379,1402]" captionTargetBox="[165,1150,143,1346]" captionTargetId="figure-56@7.[164,1151,142,1347]" captionTargetPageId="7" captionText="Figure 5. Raymunida shraddhanandi sp. nov., paratype ♀ (PCL 5.8 mm, CW 4.9 mm), IO/SS/ANO/00140. (a) carapace, pleon and cephalic appendages (dorsal view); (b) thoracic sternum (ventral view); (c) right eye, antennular peduncle article 1 and antennal peduncle (ventral view); (d) right Mxp3 (lateral view); (e) right P1 (dorsal view);(f) left P2 propodus and dactylus (lateral view); (g) left P2 merus and carpus (lateral view);(h) left P3 propodus and dactylus (lateral view); (i) left P3 merus and carpus (lateral view);(j) left P4 propodus and dactylus (lateral view); (k) left P4 merus and carpus (lateral view). Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7380776" httpUri="https://zenodo.org/record/7380776/files/figure.png" pageId="9" pageNumber="1828">Figure 5</figureCitation>
(fi)) 1.9 (P2), 2.2 (P3) and 1.7 (P4) times PCL; meri with 4 (P2), 4 (P3) and 2 (P4) spines on dorsal margins, lateral surfaces with dorsal row of 6 (P2) and 6 (P4) spines, P4 ventral margin with only 1 distal spine; carpi with 4 (P2), 3 (P3) and 2 (P4) spines on dorsal margins; P2 propodus and dactylus each with 5 ventral spines.
</paragraph>
</subSubSection>
<subSubSection id="128F653DFFB52E0FFFE424E9FCD2BB40" lastPageId="10" lastPageNumber="1829" pageId="9" pageNumber="1828" type="discussion">
<paragraph id="5A2A36B6FFB52E0CFFE424E9FEC4B902" blockId="9.[160,1156,1529,1728]" box="[160,262,1529,1555]" pageId="9" pageNumber="1828">
<emphasis id="68E1EAA4FFB52E0CFFE424E9FEC4B902" bold="true" box="[160,262,1529,1555]" italics="true" pageId="9" pageNumber="1828">Remarks</emphasis>
</paragraph>
<paragraph id="5A2A36B6FFB52E0FFFE4270DFC23BFDB" blockId="9.[160,1156,1529,1728]" lastBlockId="10.[160,1156,142,271]" lastPageId="10" lastPageNumber="1829" pageId="9" pageNumber="1828">
Among the known congeners,
<taxonomicName id="9D954D35FFB52E0CFD4C270DFD09B927" authority="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa, 2022" authorityName="Tiwari &amp; Padate &amp; Cubelio &amp; Osawa" authorityYear="2022" box="[520,715,1565,1590]" class="Malacostraca" family="Munididae" genus="Raymunida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="1828" phylum="Arthropoda" rank="species" species="shraddhanandi" status="sp. nov.">
<emphasis id="68E1EAA4FFB52E0CFD4C270DFD09B927" box="[520,715,1565,1590]" italics="true" pageId="9" pageNumber="1828">R. shraddhanandi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="73D257DFFFB52E0CFD92270DFCF2B927" box="[726,816,1565,1590]" pageId="9" pageNumber="1828" rank="species">sp. nov.</taxonomicNameLabel>
is morphologically closest to
<taxonomicName id="9D954D35FFB52E0CFFE4272FFEFAB949" authorityName="Lin, Chan and Chu" authorityYear="2004" box="[160,312,1599,1624]" class="Malacostraca" family="Munididae" genus="Raymunida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="1828" phylum="Arthropoda" rank="species" species="formasanus">
<emphasis id="68E1EAA4FFB52E0CFFE4272FFEFAB949" box="[160,312,1599,1624]" italics="true" pageId="9" pageNumber="1828">R. formasanus</emphasis>
</taxonomicName>
(known from
<collectingCountry id="22827626FFB52E0CFE90272FFDE6B949" box="[468,548,1599,1624]" name="Taiwan" pageId="9" pageNumber="1828">Taiwan</collectingCountry>
and the south-eastern
<collectingCountry id="22827626FFB52E0CFC61272FFC4BB949" box="[805,905,1599,1624]" name="Australia" pageId="9" pageNumber="1828">Australia</collectingCountry>
,
<quantity id="9D6D9B53FFB52E0CFCD6272FFBCFB949" box="[914,1037,1599,1624]" metricMagnitude="2" metricUnit="m" metricValue="2.02" metricValueMax="3.0" metricValueMin="1.04" pageId="9" pageNumber="1828" unit="m" value="202.0" valueMax="300.0" valueMin="104.0">104300 m</quantity>
depths) in having the carapace with some long setae on the dorsal surface, the frontal carapace margin with a small spine mesial to the anterolateral spine, the antennal peduncle article 2 without a small spine on the mesial margin, the P1 bearing numerous long simple setae, the P1 fixed finger with a row of relatively sparse spines on the dorsolateral margin, and the P1 dactylus with a distinct longitudinal crest along the dorsal midline.
</paragraph>
<paragraph id="5A2A36B6FFB62E0FFF8421C4FDABBE1E" blockId="10.[160,1156,142,271]" pageId="10" pageNumber="1829">
However, the new species differs from
<taxonomicName id="9D954D35FFB62E0FFD3C21C4FCD0BFFC" authorityName="Lin, Chan and Chu" authorityYear="2004" box="[632,786,212,237]" class="Malacostraca" family="Munididae" genus="Raymunida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="1829" phylum="Arthropoda" rank="species" species="formasanus">
<emphasis id="68E1EAA4FFB62E0FFD3C21C4FCD0BFFC" box="[632,786,212,237]" italics="true" pageId="10" pageNumber="1829">R. formasanus</emphasis>
</taxonomicName>
) in the following characters (see
<bibRefCitation id="3E044B47FFB62E0FFFE421E6FE70BE1E" author="Ahyong ST &amp; Poore GCB" box="[160,434,246,271]" pageId="10" pageNumber="1829" pagination="1 - 76" refId="ref9330" refString="Ahyong ST, Poore GCB. 2004. Deep-water Galatheidae (Crustacea: Decapoda: Anomura) from southern and eastern Australia. Zootaxa. 472: 1 - 76. doi: 10.11646 / zootaxa. 472.1.1." type="journal article" year="2004">Ahyong and Poore 2004</bibRefCitation>
;
<bibRefCitation id="3E044B47FFB62E0FFEFA21E6FD9EBE1E" author="Lin CW &amp; Chan TY &amp; Chu KH" box="[446,604,246,271]" pageId="10" pageNumber="1829" pagination="149 - 156" refId="ref9746" refString="Lin CW, Chan TY, Chu KH. 2004. A new squat lobster of the genus Raymunida (Decapoda: Galatheidae) from Taiwan. J Crustac Biol. 24 (1): 149 - 156. doi: 10.1651 / C- 2432." type="journal article" year="2004">Lin et al. 2004</bibRefCitation>
).
</paragraph>
<paragraph id="5A2A36B6FFB62E0FFF84202CFD66BE66" blockId="10.[192,1156,316,722]" pageId="10" pageNumber="1829">
(1) The anterior branchial carapace region is unarmed or has only one spine, instead of three to five spines as in
<taxonomicName id="9D954D35FFB62E0FFD47204EFD5FBE66" authorityName="Lin, Chan and Chu" authorityYear="2004" box="[515,669,350,375]" class="Malacostraca" family="Munididae" genus="Raymunida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="1829" phylum="Arthropoda" rank="species" species="formasanus">
<emphasis id="68E1EAA4FFB62E0FFD47204EFD5FBE66" box="[515,669,350,375]" italics="true" pageId="10" pageNumber="1829">R. formasanus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="5A2A36B6FFB62E0FFF842091FBC2BECE" blockId="10.[192,1156,316,722]" pageId="10" pageNumber="1829">
(2) The distomesial spine of the antennal peduncle article 2 reaches only or feebly overreaches the distal margin of the article 3, while it clearly overreaches that margin in
<taxonomicName id="9D954D35FFB62E0FFE1920D6FE35BECE" authorityName="Lin, Chan and Chu" authorityYear="2004" box="[349,503,454,479]" class="Malacostraca" family="Munididae" genus="Raymunida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="1829" phylum="Arthropoda" rank="species" species="formasanus">
<emphasis id="68E1EAA4FFB62E0FFE1920D6FE35BECE" box="[349,503,454,479]" italics="true" pageId="10" pageNumber="1829">R. formasanus</emphasis>
</taxonomicName>
based on the re-examination of the
<typeStatus id="852E8814FFB62E0FFCD220D6FC3FBECE" box="[918,1021,454,479]" pageId="10" pageNumber="1829" type="holotype">holotype</typeStatus>
.
</paragraph>
<paragraph id="5A2A36B6FFB62E0FFF8420F9FE4BBD34" blockId="10.[192,1156,316,722]" pageId="10" pageNumber="1829">
(3) The antennal peduncle article 3 has a short distomesial spine, which is absent in
<taxonomicName id="9D954D35FFB62E0FFFAD231CFE41BD34" authorityName="Lin, Chan and Chu" authorityYear="2004" box="[233,387,524,549]" class="Malacostraca" family="Munididae" genus="Raymunida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="1829" phylum="Arthropoda" rank="species" species="formasanus">
<emphasis id="68E1EAA4FFB62E0FFFAD231CFE41BD34" box="[233,387,524,549]" italics="true" pageId="10" pageNumber="1829">R. formasanus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="5A2A36B6FFB62E0FFF84233EFE4BBD7B" blockId="10.[192,1156,316,722]" pageId="10" pageNumber="1829">
(4) The Mxp3 merus has a small but distinct dorsodistal spine, which is absent in
<taxonomicName id="9D954D35FFB62E0FFFAD2341FE41BD7B" authorityName="Lin, Chan and Chu" authorityYear="2004" box="[233,387,593,618]" class="Malacostraca" family="Munididae" genus="Raymunida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="1829" phylum="Arthropoda" rank="species" species="formasanus">
<emphasis id="68E1EAA4FFB62E0FFFAD2341FE41BD7B" box="[233,387,593,618]" italics="true" pageId="10" pageNumber="1829">R. formasanus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="5A2A36B6FFB62E0FFF842364FE64BDC3" blockId="10.[192,1156,316,722]" pageId="10" pageNumber="1829">
(5) The P4 mero-carpal articulation reaches only the lateral end of the anterior cervical groove of the carapace, whereas it overreaches the frontal margin of the carapace in
<taxonomicName id="9D954D35FFB62E0FFE4123A9FE5DBDC3" authorityName="Lin, Chan and Chu" authorityYear="2004" box="[261,415,697,722]" class="Malacostraca" family="Munididae" genus="Raymunida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="1829" phylum="Arthropoda" rank="species" species="formasanus">
<emphasis id="68E1EAA4FFB62E0FFE4123A9FE5DBDC3" box="[261,415,697,722]" italics="true" pageId="10" pageNumber="1829">R. formasanus</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="5A2A36B6FFB62E0FFFE423EEFB89BCF6" blockId="10.[159,1155,766,999]" pageId="10" pageNumber="1829">
<taxonomicName id="9D954D35FFB62E0FFFE423EEFE28BC06" authority="Southwell, 1906" authorityName="Southwell" authorityYear="1906" box="[160,490,766,791]" class="Malacostraca" family="Munididae" genus="Munida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="1829" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="68E1EAA4FFB62E0FFFE423EEFEFEBC06" box="[160,316,766,791]" italics="true" pageId="10" pageNumber="1829">Munida alcocki</emphasis>
<bibRefCitation id="3E044B47FFB62E0FFE0423EEFE28BC06" author="Southwell T" box="[320,490,766,791]" pageId="10" pageNumber="1829" pagination="211 - 224" refId="ref10412" refString="Southwell T. 1906. Report on Anomura collected by Professor Herdman, at Ceylon, 1902. Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Mannar. Supplementary Report. 5: 211 - 224." type="journal article" year="1906">Southwell,1906</bibRefCitation>
</taxonomicName>
was described from the Gulf of Mannar (Dutch Modragam Paar and Aripu Reef,off western coast of
<collectingCountry id="22827626FFB62E0FFD5D2231FDB9BC2B" box="[537,635,801,826]" name="Sri Lanka" pageId="10" pageNumber="1829">Sri Lanka</collectingCountry>
),in shallow water to a depth of
<specimenCount id="4C93FD3FFFB62E0FFC812231FBC2BC2B" box="[965,1024,801,826]" pageId="10" pageNumber="1829" type="generic">
<quantity id="9D6D9B53FFB62E0FFC812231FC3EBC2B" box="[965,1020,801,826]" metricMagnitude="1" metricUnit="m" metricValue="6.6" pageId="10" pageNumber="1829" unit="m" value="66.0">66 m</quantity>
.
</specimenCount>
The identity of the taxon is not clear, although it has been questionably included in the synonymy of
<taxonomicName id="9D954D35FFB62E0FFFDB2276FE69BC6E" baseAuthorityName="De Man" baseAuthorityYear="1902" box="[159,427,870,895]" class="Malacostraca" family="Munididae" genus="Raymunida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="1829" phylum="Arthropoda" rank="species" species="elegantissima">
<emphasis id="68E1EAA4FFB62E0FFFDB2276FE69BC6E" box="[159,427,870,895]" italics="true" pageId="10" pageNumber="1829">Raymunida elegantissima</emphasis>
</taxonomicName>
(see
<bibRefCitation id="3E044B47FFB62E0FFEA62276FD4EBC6E" author="Baba K &amp; Macpherson E &amp; Poore GCB &amp; Ahyong ST &amp; Bermudez A &amp; Cabezas P &amp; Lin CW &amp; Nizinski M &amp; Rodrigues C &amp; Schnabel KE" box="[482,652,870,895]" pageId="10" pageNumber="1829" pagination="1 - 220" refId="ref9673" refString="Baba K, Macpherson E, Poore GCB, Ahyong ST, Bermudez A, Cabezas P, Lin CW, Nizinski M, Rodrigues C, Schnabel KE. 2008. Catalogue of squat lobsters of the world (Crustacea: Decapoda: Anomura - Families Chirostylidae, Galatheidae and Kiwaidae). Zootaxa. 1905 (1): 1 - 220. doi: 10.11646 / zootaxa. 1905.1.1." type="journal article" year="2008">Baba et al. 2008</bibRefCitation>
).
<bibRefCitation id="3E044B47FFB62E0FFDDA2276FC91BC6E" author="Southwell T" box="[670,851,870,895]" pageId="10" pageNumber="1829" pagination="211 - 224" refId="ref10412" refString="Southwell T. 1906. Report on Anomura collected by Professor Herdman, at Ceylon, 1902. Report to the Government of Ceylon on the Pearl Oyster Fisheries of the Gulf of Mannar. Supplementary Report. 5: 211 - 224." type="journal article" year="1906">Southwell (1906)</bibRefCitation>
described
<taxonomicName id="9D954D35FFB62E0FFC8C2276FBA4BC6E" authorityName="Southwell" authorityYear="1906" box="[968,1126,870,895]" class="Malacostraca" family="Munididae" genus="Munida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="1829" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="68E1EAA4FFB62E0FFC8C2276FBA4BC6E" box="[968,1126,870,895]" italics="true" pageId="10" pageNumber="1829">Munida alcocki</emphasis>
</taxonomicName>
as having seven spines on the carapace lateral margin; the number agrees only with that of
<taxonomicName id="9D954D35FFB62E0FFFDB22BCFF3CBCD4" authorityName="Osawa and Safaie" authorityYear="2014" box="[159,254,940,965]" class="Malacostraca" family="Munididae" genus="Raymunida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="1829" phylum="Arthropoda" rank="species" species="iranica">
<emphasis id="68E1EAA4FFB62E0FFFDB22BCFF3CBCD4" box="[159,254,940,965]" italics="true" pageId="10" pageNumber="1829">R. iranica</emphasis>
</taxonomicName>
;in the other known
<taxonomicName id="9D954D35FFB62E0FFE9122BCFD62BCD4" box="[469,672,940,965]" pageId="10" pageNumber="1829">
<emphasis id="68E1EAA4FFB62E0FFE9122BCFD8EBCD4" box="[469,588,940,965]" italics="true" pageId="10" pageNumber="1829">Raymunida</emphasis>
species
</taxonomicName>
,only five or six spines are present (
<bibRefCitation id="3E044B47FFB62E0FFB4F22BCFED8BCF6" author="Osawa M &amp; Safaie M" pageId="10" pageNumber="1829" pagination="265 - 274" refId="ref10145" refString="Osawa M, Safaie M. 2014. Two squat lobster species (Crustacea: Decapoda: Anomura) from the Persian Gulf, with description of a new species of Raymunida Macpherson &amp; Machordom, 2000. Zootaxa. 3861 (3): 265 - 274. doi: 10.11646 / zootaxa. 3861.3.4." type="journal article" year="2014">Osawa and Safaie 2014</bibRefCitation>
). It is unlikely that
<taxonomicName id="9D954D35FFB62E0FFEA722DEFD43BCF6" authorityName="Southwell" authorityYear="1906" box="[483,641,974,999]" class="Malacostraca" family="Munididae" genus="Munida" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="1829" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="68E1EAA4FFB62E0FFEA722DEFD43BCF6" box="[483,641,974,999]" italics="true" pageId="10" pageNumber="1829">Munida alcocki</emphasis>
</taxonomicName>
is conspecific with the present new taxon.
</paragraph>
<paragraph id="5A2A36B6FFB62E0FFFE42504FE12BB3F" blockId="10.[160,784,1044,1105]" box="[160,464,1044,1070]" pageId="10" pageNumber="1829">
<emphasis id="68E1EAA4FFB62E0FFFE42504FE12BB3F" bold="true" box="[160,464,1044,1070]" italics="true" pageId="10" pageNumber="1829">Geographical distribution</emphasis>
</paragraph>
<paragraph id="5A2A36B6FFB62E0FFFE42528FCD2BB40" blockId="10.[160,784,1044,1105]" box="[160,784,1080,1105]" pageId="10" pageNumber="1829">
So far known only from the Andaman Sea,
<quantity id="9D6D9B53FFB62E0FFDC22528FD7CBB40" box="[646,702,1080,1105]" metricMagnitude="1" metricUnit="m" metricValue="5.3" pageId="10" pageNumber="1829" unit="m" value="53.0">53 m</quantity>
depth.
</paragraph>
</subSubSection>
</treatment>
</document>