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<document id="77550A551AD2F1E89E72AD4A3CE1C19E" ID-DOI="10.5281/zenodo.281561" ID-GBIF-Dataset="5630905c-8f0c-456f-bbbb-7371b3cf12e7" ID-ISSN="1175-5326" ID-Zenodo-Dep="281561" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460492706813" checkinUser="plazi" docAuthor="Komai, Tomoyuki &amp; Ng, Peter K. L." docDate="2012" docId="03ED87FEFFD6292B64F5F876380DD645" docLanguage="en" docName="zt03352p050.pdf" docOrigin="Zootaxa 3352" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Coralliocryptus Komai &amp; Ng, 2012, n. gen." docType="treatment" docVersion="8" lastPageNumber="42" masterDocId="FFD4FF86FFD629296462FFFC3C2AD41A" masterDocTitle="A new genus and new species of leucosiid crab (Crustacea: Decapoda: Brachyura) from coral reefs in the Indo-West Pacific" masterLastPageNumber="50" masterPageNumber="40" pageNumber="40" updateTime="1698310356395" updateUser="plazi">
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<mods:title id="E3E435666CFDF825485B293B9DA587B7">A new genus and new species of leucosiid crab (Crustacea: Decapoda: Brachyura) from coral reefs in the Indo-West Pacific</mods:title>
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<mods:namePart id="A28B8D2B7548B98F056040556A93D8D6">Komai, Tomoyuki</mods:namePart>
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<mods:namePart id="8CC3938CE4E15DAB91889545C84F309F">Ng, Peter K. L.</mods:namePart>
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<mods:date id="D135A1D7B9BDB9CECA89D6977F7F75CA">2012</mods:date>
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<treatment id="03ED87FEFFD6292B64F5F876380DD645" ID-DOI="http://doi.org/10.5281/zenodo.6174787" ID-GBIF-Taxon="119519909" ID-Zenodo-Dep="6174787" LSID="urn:lsid:plazi:treatment:03ED87FEFFD6292B64F5F876380DD645" httpUri="http://treatment.plazi.org/id/03ED87FEFFD6292B64F5F876380DD645" lastPageId="2" lastPageNumber="42" pageId="0" pageNumber="40">
<subSubSection id="C35E6563FFD6292964F5F8763D81D3B9" box="[151,427,1929,1956]" pageId="0" pageNumber="40" type="nomenclature">
<paragraph id="8BFB36E8FFD6292964F5F8763D81D3B9" blockId="0.[151,604,1860,1956]" box="[151,427,1929,1956]" pageId="0" pageNumber="40">
<heading id="D0B38184FFD6292964F5F8763D81D3B9" bold="true" box="[151,427,1929,1956]" fontSize="11" level="1" pageId="0" pageNumber="40" reason="1">
<emphasis id="B930EAFAFFD6292964F5F8763D81D3B9" bold="true" box="[151,427,1929,1956]" pageId="0" pageNumber="40">
<taxonomicName id="4C444D6BFFD6292964F5F8763D79D3BE" ID-CoL="7P4Q4" box="[151,339,1930,1956]" class="Malacostraca" family="Leucosiidae" genus="Coralliocryptus" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="40" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B930EAFAFFD6292964F5F8763D79D3BE" bold="true" box="[151,339,1930,1956]" italics="true" pageId="0" pageNumber="40">Coralliocryptus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2035781FFD62929653BF8753D81D3B9" box="[345,427,1929,1955]" pageId="0" pageNumber="40" rank="genus">n. gen.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35E6563FFD6292964F5F82D3F23D3FD" box="[151,777,2000,2023]" pageId="0" pageNumber="40" type="materials_examined">
<paragraph id="8BFB36E8FFD6292964F5F82D3F23D3FD" blockId="0.[151,777,2000,2023]" box="[151,777,2000,2023]" pageId="0" pageNumber="40">
<emphasis id="B930EAFAFFD6292964F5F82D3D31D3FD" bold="true" box="[151,283,2000,2023]" pageId="0" pageNumber="40">
<typeStatus id="54FF884AFFD6292964F5F82D3CE0D3FC" box="[151,202,2001,2022]" pageId="0" pageNumber="40">Type</typeStatus>
species
</emphasis>
.
<taxonomicName id="4C444D6BFFD62929654BF82D3E1BD3FD" box="[297,561,2001,2023]" class="Malacostraca" family="Leucosiidae" genus="Coralliocryptus" kingdom="Animalia" order="Decapoda" pageId="0" pageNumber="40" phylum="Arthropoda" rank="species" species="caementa" status="sp. nov.">
<emphasis id="B930EAFAFFD62929654BF82D3E1BD3FD" box="[297,561,2001,2023]" italics="true" pageId="0" pageNumber="40">Coralliocryptus caementa</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2035781FFD629296655F82D3E5AD3FC" box="[567,624,2001,2022]" pageId="0" pageNumber="40" rank="species">
<emphasis id="B930EAFAFFD629296655F82D3E5AD3FC" bold="true" box="[567,624,2001,2022]" pageId="0" pageNumber="40">n. sp.</emphasis>
</taxonomicNameLabel>
, by monotypy.
</paragraph>
</subSubSection>
<subSubSection id="C35E6563FFD7292864F5FF6B3ED9D7D2" pageId="1" pageNumber="41" type="diagnosis">
<paragraph id="8BFB36E8FFD7292864F5FF6B3ED9D7D2" blockId="1.[151,1437,151,2013]" pageId="1" pageNumber="41">
<emphasis id="B930EAFAFFD7292864F5FF6B3D20D4AA" bold="true" box="[151,266,151,176]" pageId="1" pageNumber="41">Diagnosis</emphasis>
. Carapace distinctly wider than long, subhexagonal in general outline; dorsal surface covered with tubercles of various sizes, often stalked, but without distinct keels; frontal region slightly produced, bilobed by shallow median sulcus; cardiac region with low subconical tubercle; intestinal region with large elevation; epibranchial region strongly expanded laterally, divided into 2 lobes; pterygostomial region with prominent subconical tubercle; posterolateral margin with 1 tubercle; posterior margin with distinct tubercle at each lateral angle. Orbital hiatus effectively closed with dorsolateral angle of antennular fossa fusing with inner suborbital angle. Ocular peduncle short. Antennular basal article and flagellum folding slightly obliquely in fossa, concealed by operculiform basal article when closed. Basal antennal article positioned at lateral half of ventral floor of antennular fossa, passing under orbital margin in distal part, with distal articles and antennal flagellum inside orbit. Buccal cavern reaching beyond anterodorsal ends of afferent channel. Exopod of first and second maxilliped with flagellum. Third maxilliped with basis completely fused with ischium; merus distinctly shorter than ischium measured along mesial margin; exopod with rounded proximolateral angle. Chelipeds moderately long; merus with row of prominent stalked-tubercles on anterior and posterior margins; carpus without crests; palm gently inflated; fingers elongated, distinctly longer than palm, each with row of prominent, sharp teeth on opposable (cutting) margin. Ambulatory legs similar in shape and ornamentation; meri weakly compressed, with row of mushroom-shaped tubercles on extensor and flexor margins; carpi without crests; dactylo-propodal locking mechanism well developed. Thoracic sternum covered with numerous granules of various sizes; each suture separating sternites interrupted medially; medial suture absent; female sterno-abdominal cavity deeply excavated, well defined, completely covered by abdomen. Male abdomen narrowly triangular, consisting of free somites 1, 2 and 6, and functionally fused somites 35 (trace of sutures still evident on outer surface) plus telson. Female abdomen relatively narrow, consisting of 6 free somites plus telson, with first somite concealed beneath carapace; telson distinctly wider than long. G1 slender, compressed laterally, slightly sinuous in lateral view, terminating in subacute tip. G2 short, less than one-third length of G1, with foliaceus distal part.
</paragraph>
</subSubSection>
<subSubSection id="C35E6563FFD7292864ABFC2F3DAED00A" pageId="1" pageNumber="41" type="distribution">
<paragraph id="8BFB36E8FFD7292864ABFC2F3DAED00A" blockId="1.[151,1437,151,2013]" pageId="1" pageNumber="41">
<emphasis id="B930EAFAFFD7292864ABFC2F3D71D7F6" bold="true" box="[201,347,979,1004]" pageId="1" pageNumber="41">Distribution</emphasis>
. Widely distributed in the Indo-West Pacific, from the Comoro Is. to the Western Pacific; shallow coral reefs,
<quantity id="4CBC9B0DFFD729286578FC0B3DAAD00A" box="[282,384,1015,1040]" metricMagnitude="1" metricUnit="m" metricValue="2.15" metricValueMax="3.0" metricValueMin="1.3" pageId="1" pageNumber="41" unit="m" value="21.5" valueMax="30.0" valueMin="13.0">
13
<specimenCount id="9D42FD61FFD729286521FC0B3DAAD00A" box="[323,384,1015,1040]" pageId="1" pageNumber="41" type="generic">30 m</specimenCount>
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.
</paragraph>
</subSubSection>
<subSubSection id="C35E6563FFD7292864ABFBE039A6D3C6" pageId="1" pageNumber="41" type="discussion">
<paragraph id="8BFB36E8FFD7292864ABFBE03D08D2FA" blockId="1.[151,1437,151,2013]" pageId="1" pageNumber="41">
<emphasis id="B930EAFAFFD7292864ABFBE03D1CD02E" bold="true" box="[201,310,1052,1076]" pageId="1" pageNumber="41">Remarks</emphasis>
. With regards to the highly eroded carapace,
<taxonomicName id="4C444D6BFFD729286755FBE03FC2D02E" box="[823,1000,1052,1076]" class="Malacostraca" family="Leucosiidae" genus="Coralliocryptus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B930EAFAFFD729286755FBE03FC2D02E" box="[823,1000,1052,1076]" italics="true" pageId="1" pageNumber="41">Coralliocryptus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2035781FFD729286792FBE03815D02E" box="[1008,1087,1052,1076]" pageId="1" pageNumber="41" rank="genus">
<emphasis id="B930EAFAFFD729286792FBE03815D02E" bold="true" box="[1008,1087,1052,1076]" pageId="1" pageNumber="41">n. gen.</emphasis>
</taxonomicNameLabel>
superficially resembles genera like
<taxonomicName id="4C444D6BFFD7292864A8FBBC3D7AD042" box="[202,336,1088,1112]" class="Malacostraca" family="Leucosiidae" genus="Oreophorus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAFAFFD7292864A8FBBC3D7AD042" box="[202,336,1088,1112]" italics="true" pageId="1" pageNumber="41">Oreophorus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C444D6BFFD729286502FBBC3DEBD042" box="[352,449,1088,1112]" class="Malacostraca" family="Leucosiidae" genus="Oreotlos" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAFAFFD729286502FBBC3DEBD042" box="[352,449,1088,1112]" italics="true" pageId="1" pageNumber="41">Oreotlos</emphasis>
</taxonomicName>
and
<taxonomicName id="4C444D6BFFD72928659EFBBC3E05D042" box="[508,559,1088,1112]" class="Malacostraca" family="Leucosiidae" genus="Alox" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAFAFFD72928659EFBBC3E05D042" box="[508,559,1088,1112]" italics="true" pageId="1" pageNumber="41">Alox</emphasis>
</taxonomicName>
, but many characters argue against a close relationship. Firstly, the cheliped fingers of
<taxonomicName id="4C444D6BFFD729286568FB983D91D066" box="[266,443,1124,1148]" class="Malacostraca" family="Leucosiidae" genus="Coralliocryptus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B930EAFAFFD729286568FB983D91D066" box="[266,443,1124,1148]" italics="true" pageId="1" pageNumber="41">Coralliocryptus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2035781FFD7292865A0FB983E3AD066" box="[450,528,1124,1148]" pageId="1" pageNumber="41" rank="genus">
<emphasis id="B930EAFAFFD7292865A0FB983E3AD066" bold="true" box="[450,528,1124,1148]" pageId="1" pageNumber="41">n. gen.</emphasis>
</taxonomicNameLabel>
are very slender and elongated with the cutting margins armed with strong spiniform teeth. In members of the latter three genera, the fingers are usually stout and stocky, and while the cutting margins may have sharp teeth, they are never spiniform (e.g., see
<bibRefCitation id="EFD54B19FFD7292867E7FB513811D0DE" author="Tan" box="[901,1083,1196,1221]" pageId="1" pageNumber="41" refString="Tan, C. G. S. &amp; Ng, P. K. L. (1995) A revision of the Indo-Pacific genus Oreophorus Ruppell, 1830 (Crustacea: Decapoda: Brachyura: Leucosiidae). In: B. Richer De Forges (ed.) Les fonds meubles des lagons de Nouvelle-Caledonie (Sedimentologie, benthos). Etudes &amp; Theses, 2, 101 - 189. ORSTOM, Paris." type="journal article" year="1995">Tan &amp; Ng 1995</bibRefCitation>
:
<figureCitation id="137F2A6DFFD729286025FB5038BCD0DE" box="[1095,1174,1196,1220]" captionStart="FIGURE 2" captionStartId="5.[151,250,1977,2000]" captionTargetBox="[186,1406,193,1956]" captionTargetId="figure@5.[186,1406,193,1958]" captionTargetPageId="5" captionText="FIGURE 2. Coralliocryptus caementa n. gen., n. sp., non-type, male (3.8 x 6.4 mm) (CBM-ZC 10876), Vanuatu, entire animal. A, dorsal view; B, anterior (frontal) view; C, ventral view." httpUri="https://zenodo.org/record/281563/files/figure.png" pageId="1" pageNumber="41">Figs. 2</figureCitation>
E; 8C; 12C; 16B; 17D;
<bibRefCitation id="EFD54B19FFD7292864F5FB2D3D44D0F2" author="Naruse" box="[151,366,1232,1257]" pageId="1" pageNumber="41" refString="Naruse, T. &amp; Ng, P. K. L. (2006) Two new species of leucosiid crabs (Decapoda: Brachyura) from the Ryukyu Islands, Japan. Crustacean Research, 35, 108 - 116." type="journal article" year="2006">Naruse &amp; Ng 2006</bibRefCitation>
:
<figureCitation id="137F2A6DFFD72928651AFB2C3D93D0F2" box="[376,441,1232,1256]" captionStart="FIGURE 2" captionStartId="5.[151,250,1977,2000]" captionTargetBox="[186,1406,193,1956]" captionTargetId="figure@5.[186,1406,193,1958]" captionTargetPageId="5" captionText="FIGURE 2. Coralliocryptus caementa n. gen., n. sp., non-type, male (3.8 x 6.4 mm) (CBM-ZC 10876), Vanuatu, entire animal. A, dorsal view; B, anterior (frontal) view; C, ventral view." httpUri="https://zenodo.org/record/281563/files/figure.png" pageId="1" pageNumber="41">Fig. 2</figureCitation>
C;
<bibRefCitation id="EFD54B19FFD7292865BBFB2C3EB2D0F2" author="Galil" box="[473,664,1232,1257]" pageId="1" pageNumber="41" refString="Galil, B. S. &amp; Ng, P. K. L. (2007) Leucosiid crabs from Panglao, Philippines, with descriptions of three new species (Crustacea: Decapoda: Brachyura). Raffles Bulletin of Zoology, Supplement 16, 79 - 94." type="book chapter" year="2007">Galil &amp; Ng 2007</bibRefCitation>
:
<figureCitation id="137F2A6DFFD7292866C0FB2C3EF4D0F2" box="[674,734,1232,1256]" captionStart="FIGURE 1" captionStartId="4.[151,250,1985,2008]" captionTargetBox="[255,1325,195,1959]" captionTargetId="figure@4.[255,1326,195,1960]" captionTargetPageId="4" captionText="FIGURE 1. Coralliocryptus caementa n. gen., n. sp., holotype, female (5.4 x 9.1 mm) (CBM-ZC 10875), Okinawa, entire animal. A, dorsal view; B, anterior (frontal) view; C, ventral view." httpUri="https://zenodo.org/record/281562/files/figure.png" pageId="1" pageNumber="41">Fig. 1</figureCitation>
A-C; 2009:
<figureCitation id="137F2A6DFFD72928670AFB2C3F83D0F2" box="[872,937,1232,1256]" captionStart="FIGURE 2" captionStartId="5.[151,250,1977,2000]" captionTargetBox="[186,1406,193,1956]" captionTargetId="figure@5.[186,1406,193,1958]" captionTargetPageId="5" captionText="FIGURE 2. Coralliocryptus caementa n. gen., n. sp., non-type, male (3.8 x 6.4 mm) (CBM-ZC 10876), Vanuatu, entire animal. A, dorsal view; B, anterior (frontal) view; C, ventral view." httpUri="https://zenodo.org/record/281563/files/figure.png" pageId="1" pageNumber="41">Fig. 2</figureCitation>
C;
<bibRefCitation id="EFD54B19FFD7292867ABFB2D387CD0F2" author="Huang" box="[969,1110,1232,1257]" pageId="1" pageNumber="41" refString="Huang, J. - F. (2010) On a new species, Oreophorus holthuisi sp. nov. (Decapoda, Brachyura, Leucosiidae), from the Indian Ocean. In Fransen, C. H. J. M., De Grave, S. &amp; Ng, P. K. L. (editors), Studies on Malacostraca: Lipke Bijdeley Holthuis Memorial Volume. Crustaceana Monographs, 14, 333 - 342." type="journal article" year="2010">Huang 2010</bibRefCitation>
:
<figureCitation id="137F2A6DFFD729286002FB2C3888D0F2" box="[1120,1186,1232,1256]" captionStart="FIGURE 2" captionStartId="5.[151,250,1977,2000]" captionTargetBox="[186,1406,193,1956]" captionTargetId="figure@5.[186,1406,193,1958]" captionTargetPageId="5" captionText="FIGURE 2. Coralliocryptus caementa n. gen., n. sp., non-type, male (3.8 x 6.4 mm) (CBM-ZC 10876), Vanuatu, entire animal. A, dorsal view; B, anterior (frontal) view; C, ventral view." httpUri="https://zenodo.org/record/281563/files/figure.png" pageId="1" pageNumber="41">Fig. 2</figureCitation>
D; Ng
<emphasis id="B930EAFAFFD72928608EFB2D390ED0F2" box="[1260,1316,1232,1256]" italics="true" pageId="1" pageNumber="41">et al.</emphasis>
2009:
<figureCitation id="137F2A6DFFD729286112FB2C3C8FD116" captionStart="FIGURE 3" captionStartId="6.[151,250,1818,1841]" captionTargetBox="[228,1360,188,1777]" captionTargetId="figure@6.[213,1382,161,1803]" captionTargetPageId="6" captionText="FIGURE 3. Coralliocryptus caementa n. gen., n. sp., holotype, female (5.4 x 9.1 mm) (CBM-ZC 10875), Okinawa. A, schematic drawing of carapace, dorsal view, showing outline and position of tubercles; B, anterior part of carapace, left side, dorsal view; C, lateral part of carapace, dorsal view; D, frontal part of carapace, right side, anterior (frontal) view, showing epistome, antennular fossa and antenna; E, right orbit, anterior (frontal) view; F, dorsal roof of orbit, posterodorsal view; G, left third maxilliped, outer view; H, anterior part of sterno-abdominal cavity, ventral view; I, abdomen and telson, ventral view (tubercles on somites 3 6 and telson, and setae omitted). Scale bars: 2 mm for A; 1 mm for B, C, I; 0.5 mm for D G. Abbreviations: A 1 B, basal segment of antennule; S 1, abdominal somite 1." httpUri="https://zenodo.org/record/281564/files/figure.png" pageId="1" pageNumber="41">Fig. 3</figureCitation>
A). Most significantly, the female abdomen of
<taxonomicName id="4C444D6BFFD7292866D0FB083F49D116" box="[690,867,1268,1292]" class="Malacostraca" family="Leucosiidae" genus="Coralliocryptus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B930EAFAFFD7292866D0FB083F49D116" box="[690,867,1268,1292]" italics="true" pageId="1" pageNumber="41">Coralliocryptus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2035781FFD729286709FB083F93D116" box="[875,953,1268,1292]" pageId="1" pageNumber="41" rank="genus">
<emphasis id="B930EAFAFFD729286709FB083F93D116" bold="true" box="[875,953,1268,1292]" pageId="1" pageNumber="41">n. gen.</emphasis>
</taxonomicNameLabel>
is unique in having all the somites and telson free, whereas all other known leucosiids have most of the somites fused to form a domed plate (e.g., see
<bibRefCitation id="EFD54B19FFD729286134FAE53CD6D14E" author="Tan" pageId="1" pageNumber="41" refString="Tan, C. G. S. &amp; Ng, P. K. L. (1995) A revision of the Indo-Pacific genus Oreophorus Ruppell, 1830 (Crustacea: Decapoda: Brachyura: Leucosiidae). In: B. Richer De Forges (ed.) Les fonds meubles des lagons de Nouvelle-Caledonie (Sedimentologie, benthos). Etudes &amp; Theses, 2, 101 - 189. ORSTOM, Paris." type="journal article" year="1995">Tan &amp; Ng 1995</bibRefCitation>
: Figs. 8F; 16D). However,
<taxonomicName id="4C444D6BFFD72928664DFAC03ECAD14E" box="[559,736,1340,1364]" class="Malacostraca" family="Leucosiidae" genus="Coralliocryptus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B930EAFAFFD72928664DFAC03ECAD14E" box="[559,736,1340,1364]" italics="true" pageId="1" pageNumber="41">Coralliocryptus</emphasis>
</taxonomicName>
still has a deeply excavated sterno-abdominal cavity in which the eggs are retained and are completely covered by the abdomen, as in other leucosiid taxa. As such,
<taxonomicName id="4C444D6BFFD729286160FA9C3C92D186" class="Malacostraca" family="Leucosiidae" genus="Coralliocryptus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B930EAFAFFD729286160FA9C3C92D186" italics="true" pageId="1" pageNumber="41">Coralliocryptus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2035781FFD7292864A3FA783D38D186" box="[193,274,1412,1436]" pageId="1" pageNumber="41" rank="genus">
<emphasis id="B930EAFAFFD7292864A3FA783D38D186" bold="true" box="[193,274,1412,1436]" pageId="1" pageNumber="41">n. gen.</emphasis>
</taxonomicNameLabel>
should be included in the
<taxonomicName id="4C444D6BFFD72928662BFA783EFAD186" box="[585,720,1412,1436]" class="Malacostraca" family="Leucosiidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="41" phylum="Arthropoda" rank="family">Leucosiidae</taxonomicName>
as defined at present rather than in
<taxonomicName id="4C444D6BFFD729286016FA7839B2D186" authority="Alcock, 1896" authorityName="Alcock" authorityYear="1896" box="[1140,1432,1412,1436]" class="Malacostraca" family="Iphiculidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="41" phylum="Arthropoda" rank="family">Iphiculidae Alcock, 1896</taxonomicName>
, another family placed in the Leucosioidea, in spite of the similarity in the armature of the fingers of the chelae and the freely articulated female abdominal somites between the present new taxon and the members of
<taxonomicName id="4C444D6BFFD729286175FA3039B3D1FE" box="[1303,1433,1484,1508]" class="Malacostraca" family="Iphiculidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="41" phylum="Arthropoda" rank="family">Iphiculidae</taxonomicName>
. The primary character that defines the
<taxonomicName id="4C444D6BFFD729286625FA0C3EEFD212" box="[583,709,1520,1544]" class="Malacostraca" family="Iphiculidae" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="41" phylum="Arthropoda" rank="family">Iphiculidae</taxonomicName>
is the relatively narrow female abdomen in which all the somites are free, the sterno-abdominal cavity is relatively shallow and the eggs are never completely covered by the abdomen, with the egg mass protruding from the sides of the abdomen (see Ng
<emphasis id="B930EAFAFFD729286782F9C53832D24A" box="[992,1048,1592,1616]" italics="true" pageId="1" pageNumber="41">et al.</emphasis>
2008). It is worth mentioning that all iphiculids have the same kind of chela as that of
<taxonomicName id="4C444D6BFFD729286697F9A03F8CD26E" box="[757,934,1628,1652]" class="Malacostraca" family="Leucosiidae" genus="Coralliocryptus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B930EAFAFFD729286697F9A03F8CD26E" box="[757,934,1628,1652]" italics="true" pageId="1" pageNumber="41">Coralliocryptus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2035781FFD7292867D2F9A0382BD26E" box="[944,1025,1628,1652]" pageId="1" pageNumber="41" rank="genus">
<emphasis id="B930EAFAFFD7292867D2F9A0382BD26E" bold="true" box="[944,1025,1628,1652]" pageId="1" pageNumber="41">n. gen.</emphasis>
</taxonomicNameLabel>
, with slender fingers and spiniform teeth along the cutting margins. This unusual chela, however, is not unique to iphiculids, and is also seen in various leucosiids, including some species of
<taxonomicName id="4C444D6BFFD729286627F9593F3CD2A6" authority="Leach, 1817" authorityName="Leach" authorityYear="1817" box="[581,790,1700,1724]" class="Malacostraca" family="Leucosiidae" genus="Myra" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAFAFFD729286627F9593EA8D2A6" box="[581,642,1701,1724]" italics="true" pageId="1" pageNumber="41">Myra</emphasis>
Leach, 1817
</taxonomicName>
(see
<bibRefCitation id="EFD54B19FFD729286737F9583FE4D2A6" author="Galil" box="[853,974,1700,1724]" pageId="1" pageNumber="41" refString="Galil, B. S. (2001) A revision of Myra Leach, 1817 (Crustacea: Decapoda: Leucosioidea). Zoologische Mededelingen, 75, 409 - 446." type="journal article" year="2001">Galil 2001</bibRefCitation>
) and
<taxonomicName id="4C444D6BFFD72928606DF95838C0D2A6" authority="Bell, 1855" authorityName="Bell" authorityYear="1855" box="[1039,1258,1700,1724]" class="Malacostraca" family="Leucosiidae" genus="Nursilia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAFAFFD72928606DF9583841D2A6" box="[1039,1131,1700,1724]" italics="true" pageId="1" pageNumber="41">Nursilia</emphasis>
Bell, 1855
</taxonomicName>
(see
<bibRefCitation id="EFD54B19FFD72928614AF95839BDD2A6" author="Ihle" box="[1320,1431,1700,1724]" pageId="1" pageNumber="41" refString="Ihle, J. E. W. (1918) Die Decapoda Brachyura der Siboga-Expedition, 3. Oxystomata: Calappidae, Leucosiidae, Raninidae. Siboga Expeditie Monogrphie, 39 b 2, 159 - 322." type="journal article" year="1918">Ihle 1918</bibRefCitation>
;
<bibRefCitation id="EFD54B19FFD7292864F5F9343D3FD2FA" author="Chen" box="[151,277,1736,1760]" pageId="1" pageNumber="41" refString="Chen, H. (1982) On the genus Nursilia (Crustacea, Decapoda: Leucosiidae) of Chinese waters. Oceanologia et Limnologia Sinica, 13, 267 - 272, 2 pls." type="journal article" year="1982">Chen 1982</bibRefCitation>
).
</paragraph>
<paragraph id="8BFB36E8FFD7292864ABF91139A6D3C6" blockId="1.[151,1437,151,2013]" pageId="1" pageNumber="41">
In most species of
<taxonomicName id="4C444D6BFFD7292865F8F9103E35D31E" box="[410,543,1772,1796]" class="Malacostraca" family="Leucosiidae" genus="Oreophorus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAFAFFD7292865F8F9103E35D31E" box="[410,543,1772,1796]" italics="true" pageId="1" pageNumber="41">Oreophorus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C444D6BFFD72928664FF9103EA7D31E" box="[557,653,1772,1796]" class="Malacostraca" family="Leucosiidae" genus="Oreotlos" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAFAFFD72928664FF9103EA7D31E" box="[557,653,1772,1796]" italics="true" pageId="1" pageNumber="41">Oreotlos</emphasis>
</taxonomicName>
and
<taxonomicName id="4C444D6BFFD7292866A6F9103EDDD31E" box="[708,759,1772,1796]" class="Malacostraca" family="Leucosiidae" genus="Alox" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAFAFFD7292866A6F9103EDDD31E" box="[708,759,1772,1796]" italics="true" pageId="1" pageNumber="41">Alox</emphasis>
</taxonomicName>
, the basal antennal article is tightly lodged in the orbital hiatus, fusing with it (e.g., see
<bibRefCitation id="EFD54B19FFD7292865A8F8ED3E51D332" author="Tan" box="[458,635,1808,1833]" pageId="1" pageNumber="41" refString="Tan, C. G. S. &amp; Ng, P. K. L. (1995) A revision of the Indo-Pacific genus Oreophorus Ruppell, 1830 (Crustacea: Decapoda: Brachyura: Leucosiidae). In: B. Richer De Forges (ed.) Les fonds meubles des lagons de Nouvelle-Caledonie (Sedimentologie, benthos). Etudes &amp; Theses, 2, 101 - 189. ORSTOM, Paris." type="journal article" year="1995">Tan &amp; Ng 1995</bibRefCitation>
:
<figureCitation id="137F2A6DFFD7292866E4F8EC3EF8D332" box="[646,722,1808,1832]" captionStart="FIGURE 5" captionStartId="8.[151,250,1457,1480]" captionTargetBox="[169,1407,213,1432]" captionTargetId="figure@8.[151,1436,193,1437]" captionTargetPageId="8" captionText="FIGURE 5. Coralliocryptus caementa n. gen., n. sp., holotype, female (5.4 x 9.1 mm) (CBM-ZC 10875), Okinawa. A, right cheliped, ventral view; B, same, dorsal view (tubercles on merus omitted); C, same, chela, extensor (anterior) view; D, right first ambulatory leg (second pereopod), ventral view; E, same, dactylus, ventral view. Scale bars: 1 mm for A D; 0.5 mm for E." httpUri="https://zenodo.org/record/281566/files/figure.png" pageId="1" pageNumber="41">Figs. 5</figureCitation>
K). In
<taxonomicName id="4C444D6BFFD729286779F8EC3FE0D332" box="[795,970,1808,1832]" class="Malacostraca" family="Leucosiidae" genus="Coralliocryptus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B930EAFAFFD729286779F8EC3FE0D332" box="[795,970,1808,1832]" italics="true" pageId="1" pageNumber="41">Coralliocryptus</emphasis>
</taxonomicName>
, however, while the basal antennal article is still in the same position, there is no real orbital hiatus because the orbital margin is effectively closed, with the dorsolateral angle of the antennular fossa and inner suborbital angle fused together (
<figureCitation id="137F2A6DFFD729286053F8AB3858D375" box="[1073,1138,1879,1903]" captionStart="FIGURE 3" captionStartId="6.[151,250,1818,1841]" captionTargetBox="[228,1360,188,1777]" captionTargetId="figure@6.[213,1382,161,1803]" captionTargetPageId="6" captionText="FIGURE 3. Coralliocryptus caementa n. gen., n. sp., holotype, female (5.4 x 9.1 mm) (CBM-ZC 10875), Okinawa. A, schematic drawing of carapace, dorsal view, showing outline and position of tubercles; B, anterior part of carapace, left side, dorsal view; C, lateral part of carapace, dorsal view; D, frontal part of carapace, right side, anterior (frontal) view, showing epistome, antennular fossa and antenna; E, right orbit, anterior (frontal) view; F, dorsal roof of orbit, posterodorsal view; G, left third maxilliped, outer view; H, anterior part of sterno-abdominal cavity, ventral view; I, abdomen and telson, ventral view (tubercles on somites 3 6 and telson, and setae omitted). Scale bars: 2 mm for A; 1 mm for B, C, I; 0.5 mm for D G. Abbreviations: A 1 B, basal segment of antennule; S 1, abdominal somite 1." httpUri="https://zenodo.org/record/281564/files/figure.png" pageId="1" pageNumber="41">Fig. 3</figureCitation>
E). The basal antennal article of
<taxonomicName id="4C444D6BFFD7292864BEF8803DA1D38E" box="[220,395,1916,1940]" class="Malacostraca" family="Leucosiidae" genus="Coralliocryptus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B930EAFAFFD7292864BEF8803DA1D38E" box="[220,395,1916,1940]" italics="true" pageId="1" pageNumber="41">Coralliocryptus</emphasis>
</taxonomicName>
is therefore, actually underneath the orbital margin (
<figureCitation id="137F2A6DFFD7292867B0F8803839D38E" box="[978,1043,1916,1940]" captionStart="FIGURE 5" captionStartId="8.[151,250,1457,1480]" captionTargetBox="[169,1407,213,1432]" captionTargetId="figure@8.[151,1436,193,1437]" captionTargetPageId="8" captionText="FIGURE 5. Coralliocryptus caementa n. gen., n. sp., holotype, female (5.4 x 9.1 mm) (CBM-ZC 10875), Okinawa. A, right cheliped, ventral view; B, same, dorsal view (tubercles on merus omitted); C, same, chela, extensor (anterior) view; D, right first ambulatory leg (second pereopod), ventral view; E, same, dactylus, ventral view. Scale bars: 1 mm for A D; 0.5 mm for E." httpUri="https://zenodo.org/record/281566/files/figure.png" pageId="1" pageNumber="41">Fig. 5</figureCitation>
D). This is a rare feature in leucosiids but is not well studied and may be of higher systematic value. It is noteworthy that at least one
<taxonomicName id="4C444D6BFFD7292860BCF85C3914D3A2" box="[1246,1342,1952,1976]" class="Malacostraca" family="Leucosiidae" genus="Oreotlos" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAFAFFD7292860BCF85C3914D3A2" box="[1246,1342,1952,1976]" italics="true" pageId="1" pageNumber="41">Oreotlos</emphasis>
</taxonomicName>
species,
<taxonomicName id="4C444D6BFFD7292864F5F8393DFED3C6" authority="Tan &amp; Ng, 1995" authorityName="Tan &amp; Ng" authorityYear="1995" box="[151,468,1988,2013]" class="Malacostraca" family="Leucosiidae" genus="Oreotlos" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="41" phylum="Arthropoda" rank="species" species="heuretos">
<emphasis id="B930EAFAFFD7292864F5F8393D3DD3C6" box="[151,279,1988,2012]" italics="true" pageId="1" pageNumber="41">O. heuretos</emphasis>
<bibRefCitation id="EFD54B19FFD72928657CF8393DFED3C6" author="Tan" box="[286,468,1988,2013]" pageId="1" pageNumber="41" refString="Tan, C. G. S. &amp; Ng, P. K. L. (1995) A revision of the Indo-Pacific genus Oreophorus Ruppell, 1830 (Crustacea: Decapoda: Brachyura: Leucosiidae). In: B. Richer De Forges (ed.) Les fonds meubles des lagons de Nouvelle-Caledonie (Sedimentologie, benthos). Etudes &amp; Theses, 2, 101 - 189. ORSTOM, Paris." type="journal article" year="1995">Tan &amp; Ng, 1995</bibRefCitation>
</taxonomicName>
, has an orbital structure similar to that of
<taxonomicName id="4C444D6BFFD7292867C2F8383865D3C6" box="[928,1103,1988,2012]" class="Malacostraca" family="Leucosiidae" genus="Coralliocryptus" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="41" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B930EAFAFFD7292867C2F8383865D3C6" box="[928,1103,1988,2012]" italics="true" pageId="1" pageNumber="41">Coralliocryptus</emphasis>
</taxonomicName>
(H. Komatsu, pers. comm.).
</paragraph>
</subSubSection>
<subSubSection id="C35E6563FFD4292B64ABFF643DFAD4ED" pageId="2" pageNumber="42" type="etymology">
<paragraph id="8BFB36E8FFD4292B64ABFF643DFAD4ED" blockId="2.[151,1436,151,608]" pageId="2" pageNumber="42">
<emphasis id="B930EAFAFFD4292B64ABFF643D62D4AA" bold="true" box="[201,328,152,176]" pageId="2" pageNumber="42">Etymology</emphasis>
. From the combination of the Latinized Greek
<emphasis id="B930EAFAFFD4292B6707FF643FF8D4AA" box="[869,978,152,176]" italics="true" pageId="2" pageNumber="42">corallium</emphasis>
(= coral) and
<emphasis id="B930EAFAFFD4292B6015FF6538E0D4AA" box="[1143,1226,153,176]" italics="true" pageId="2" pageNumber="42">cryptus</emphasis>
(hidden), in reference to the new taxon cryptic habitat and mimic habit in shallow coral reefs, having prevented its discovery until recently. Gender masculine.
</paragraph>
</subSubSection>
<subSubSection id="C35E6563FFD4292B64ABFEFF380DD645" pageId="2" pageNumber="42" type="materials_examined">
<paragraph id="8BFB36E8FFD4292B64ABFEFF380DD645" blockId="2.[151,1436,151,608]" pageId="2" pageNumber="42">
<emphasis id="B930EAFAFFD4292B64ABFEFF3DF9D506" bold="true" box="[201,467,259,284]" pageId="2" pageNumber="42">Comparative material</emphasis>
.
<taxonomicName id="4C444D6BFFD4292B6581FEF83F25D506" authority="Ihle, 1918" authorityName="Ihle" authorityYear="1918" box="[483,783,260,284]" class="Malacostraca" family="Leucosiidae" genus="Alox" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="42" phylum="Arthropoda" rank="species" species="ornatum">
<emphasis id="B930EAFAFFD4292B6581FEF83E57D506" box="[483,637,260,284]" italics="true" pageId="2" pageNumber="42">Alox ornatum</emphasis>
(
<bibRefCitation id="EFD54B19FFD4292B66ECFEF83F2DD506" author="Ihle" box="[654,775,260,284]" pageId="2" pageNumber="42" refString="Ihle, J. E. W. (1918) Die Decapoda Brachyura der Siboga-Expedition, 3. Oxystomata: Calappidae, Leucosiidae, Raninidae. Siboga Expeditie Monogrphie, 39 b 2, 159 - 322." type="journal article" year="1918">Ihle, 1918</bibRefCitation>
)
</taxonomicName>
:
<specimenCount id="9D42FD61FFD4292B6779FEF83F4DD506" box="[795,871,260,284]" pageId="2" pageNumber="42" type="male">1 male</specimenCount>
(CBM-ZC 6885), N of Yakushima I., Ohsumi Is, southwestern
<collectingCountry id="F3537678FFD4292B6556FED43D5DD525" box="[308,375,296,319]" name="Japan" pageId="2" pageNumber="42">Japan</collectingCountry>
,
<geoCoordinate id="EE70502FFFD4292B65E6FEDB3E21D525" box="[388,523,295,319]" direction="north" orientation="latitude" pageId="2" pageNumber="42" precision="9" value="30.466667">30°28.00N</geoCoordinate>
,
<geoCoordinate id="EE70502FFFD4292B667AFEDB3E80D525" box="[536,682,295,319]" direction="east" orientation="longitude" pageId="2" pageNumber="42" precision="9" value="130.57666">130°34.60E</geoCoordinate>
,
<quantity id="4CBC9B0DFFD4292B66D5FEDB3EDED525" box="[695,756,295,319]" metricMagnitude="1" metricUnit="m" metricValue="9.6" pageId="2" pageNumber="42" unit="m" value="96.0">96 m</quantity>
, TRV
<emphasis id="B930EAFAFFD4292B6723FED43FCDD55A" box="[833,999,296,320]" italics="true" pageId="2" pageNumber="42">Toyoshio-maru</emphasis>
, 1997-05 cruise, stn 11,
<date id="FFFA1028FFD4292B616BFEDB39B2D525" box="[1289,1432,295,319]" pageId="2" pageNumber="42" value="1997-06-03">3 June 1997</date>
, dredge, coll. T. Komai.
<taxonomicName id="4C444D6BFFD4292B65CEFEB03F34D57E" authority="Stimpson, 1858" authorityName="Stimpson" authorityYear="1858" box="[428,798,332,356]" class="Malacostraca" family="Leucosiidae" genus="Alox" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="42" phylum="Arthropoda" rank="species" species="rugosum">
<emphasis id="B930EAFAFFD4292B65CEFEB03E66D57E" box="[428,588,332,356]" italics="true" pageId="2" pageNumber="42">Alox rugosum</emphasis>
(Stimpson, 1858)
</taxonomicName>
:
<specimenCount id="9D42FD61FFD4292B674FFEB03F51D57E" box="[813,891,332,356]" pageId="2" pageNumber="42" type="male">1 male</specimenCount>
(CBM-ZC 3999),
<collectingCountry id="F3537678FFD4292B6034FEB038E1D57E" box="[1110,1227,332,356]" name="Singapore" pageId="2" pageNumber="42">Singapore</collectingCountry>
,
<quantity id="4CBC9B0DFFD4292B60BBFEB0396BD57E" box="[1241,1345,332,356]" metricMagnitude="1" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="2" pageNumber="42" unit="m" value="17.5" valueMax="20.0" valueMin="15.0">1520 m</quantity>
,
<date id="FFFA1028FFD4292B6132FEB03CFBD59D" pageId="2" pageNumber="42" value="1997-05-09">9 May 1997</date>
, dredge, coll. T. Komai;
<specimenCount id="9D42FD61FFD4292B658AFE933E63D59D" box="[488,585,367,391]" pageId="2" pageNumber="42" type="female">1 female</specimenCount>
(CBM-ZC 9635), Red Beach, Kin, Okinawa, Ryukyu Archipelago,
<quantity id="4CBC9B0DFFD4292B6125FE93395ED59D" box="[1351,1396,367,391]" metricMagnitude="0" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="42" unit="m" value="4.0">4 m</quantity>
,
<date id="FFFA1028FFD4292B61E2FE933D19D5B6" pageId="2" pageNumber="42" value="2009-01-23">23 January 2009</date>
,
<collectionCode id="ED55AE2DFFD4292B6521FE683D8AD5B6" box="[323,416,404,428]" pageId="2" pageNumber="42">SCUBA</collectionCode>
diving, coll. Yusuke Yamada.
<taxonomicName id="4C444D6BFFD4292B676EFE6838ADD5B6" authority="Naruse &amp; Ng, 2006: 1" authorityName="Naruse &amp; Ng" authorityPageNumber="1" authorityYear="2006" box="[780,1159,404,429]" class="Malacostraca" family="Leucosiidae" genus="Alox" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="42" phylum="Arthropoda" rank="species" species="uru">
<emphasis id="B930EAFAFFD4292B676EFE683F58D5B6" box="[780,882,404,428]" italics="true" pageId="2" pageNumber="42">Alox uru</emphasis>
<bibRefCitation id="EFD54B19FFD4292B671FFE693840D5B6" author="Naruse" box="[893,1130,404,429]" pageId="2" pageNumber="42" refString="Naruse, T. &amp; Ng, P. K. L. (2006) Two new species of leucosiid crabs (Decapoda: Brachyura) from the Ryukyu Islands, Japan. Crustacean Research, 35, 108 - 116." type="journal article" year="2006">Naruse &amp; Ng, 2006</bibRefCitation>
:
<specimenCount id="9D42FD61FFD4292B601BFE6838ADD5B6" box="[1145,1159,404,428]" pageId="2" pageNumber="42" type="male">1</specimenCount>
</taxonomicName>
male (CBM-ZC 7080), Uehara beach, Iriomote I., Yaeyama Is., Ryukyu Archipelago, subtidal,
<date id="FFFA1028FFD4292B67DAFE4B3817D5D5" box="[952,1085,439,463]" pageId="2" pageNumber="42" value="2001-07-08">8 July 2001</date>
, dip net, coll. T. Komai.
<taxonomicName id="4C444D6BFFD4292B6130FE443DF8D5EE" authority="Borradaile, 1903" authorityName="Borradaile" authorityYear="1903" class="Malacostraca" family="Leucosiidae" genus="Oreotlos" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="42" phylum="Arthropoda" rank="species" species="latus">
<emphasis id="B930EAFAFFD4292B6130FE443CDCD5EE" italics="true" pageId="2" pageNumber="42">Oreotlos latus</emphasis>
(Borradaile, 1903)
</taxonomicName>
:
<specimenCount id="9D42FD61FFD4292B65BFFE203E03D5EE" box="[477,553,476,500]" pageId="2" pageNumber="42" type="male">1 male</specimenCount>
(CBM-ZC 6906), off Muko-jima I., Ogasawara Is,
<geoCoordinate id="EE70502FFFD4292B6011FE2038D0D5EE" box="[1139,1274,476,500]" direction="north" orientation="latitude" pageId="2" pageNumber="42" precision="9" value="27.795334">27°47.72N</geoCoordinate>
,
<geoCoordinate id="EE70502FFFD4292B6165FE2039B2D5EE" box="[1287,1432,476,500]" direction="east" orientation="longitude" pageId="2" pageNumber="42" precision="9" value="142.0495">142°02.97E</geoCoordinate>
,
<quantity id="4CBC9B0DFFD4292B64F5FE033CD1D60D" box="[151,251,511,535]" metricMagnitude="1" metricUnit="m" metricValue="6.9" metricValueMax="7.0" metricValueMin="6.8" pageId="2" pageNumber="42" unit="m" value="69.0" valueMax="70.0" valueMin="68.0">6870 m</quantity>
, TRV
<emphasis id="B930EAFAFFD4292B6521FDFC3DF7D602" box="[323,477,512,536]" italics="true" pageId="2" pageNumber="42">Shinyo-maru</emphasis>
, 1997 cruise, stn 2,
<date id="FFFA1028FFD4292B66D9FE033F53D60D" box="[699,889,511,535]" pageId="2" pageNumber="42" value="1997-10-15">15 October 1997</date>
, dredge, coll. T. Komai. Comparative material of
<taxonomicName id="4C444D6BFFD4292B64F5FDD83D37D626" box="[151,285,548,572]" class="Malacostraca" family="Leucosiidae" genus="Oreophorus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAFAFFD4292B64F5FDD83D37D626" box="[151,285,548,572]" italics="true" pageId="2" pageNumber="42">Oreophorus</emphasis>
</taxonomicName>
,
<taxonomicName id="4C444D6BFFD4292B654EFDD83DA7D626" box="[300,397,548,572]" class="Malacostraca" family="Leucosiidae" genus="Oreotlos" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAFAFFD4292B654EFDD83DA7D626" box="[300,397,548,572]" italics="true" pageId="2" pageNumber="42">Oreotlos</emphasis>
</taxonomicName>
,
<taxonomicName id="4C444D6BFFD4292B65F9FDD83DE5D626" box="[411,463,548,572]" class="Malacostraca" family="Leucosiidae" genus="Alox" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAFAFFD4292B65F9FDD83DE5D626" box="[411,463,548,572]" italics="true" pageId="2" pageNumber="42">Alox</emphasis>
</taxonomicName>
,
<taxonomicName id="4C444D6BFFD4292B65BFFDD83E6FD626" box="[477,581,548,572]" class="Malacostraca" family="Iphiculidae" genus="Iphiculus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAFAFFD4292B65BFFDD83E6FD626" box="[477,581,548,572]" italics="true" pageId="2" pageNumber="42">Iphiculus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C444D6BFFD4292B661FFDD83F24D626" box="[637,782,548,572]" class="Malacostraca" family="Iphiculidae" genus="Pariphiculus" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="42" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAFAFFD4292B661FFDD83F24D626" box="[637,782,548,572]" italics="true" pageId="2" pageNumber="42">Pariphiculus</emphasis>
</taxonomicName>
species in the
<collectionCode id="ED55AE2DFFD4292B67D7FDD83FC1D626" box="[949,1003,548,572]" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="2" pageNumber="42">ZRC</collectionCode>
recorded in
<bibRefCitation id="EFD54B19FFD4292B6018FDD93915D626" author="Tan" box="[1146,1343,548,573]" pageId="2" pageNumber="42" refString="Tan, C. G. S. &amp; Ng, P. K. L. (1995) A revision of the Indo-Pacific genus Oreophorus Ruppell, 1830 (Crustacea: Decapoda: Brachyura: Leucosiidae). In: B. Richer De Forges (ed.) Les fonds meubles des lagons de Nouvelle-Caledonie (Sedimentologie, benthos). Etudes &amp; Theses, 2, 101 - 189. ORSTOM, Paris." type="journal article" year="1995">Tan &amp; Ng (1995)</bibRefCitation>
,
<bibRefCitation id="EFD54B19FFD4292B612EFDD93D00D645" author="Naruse" pageId="2" pageNumber="42" refString="Naruse, T. &amp; Ng, P. K. L. (2006) Two new species of leucosiid crabs (Decapoda: Brachyura) from the Ryukyu Islands, Japan. Crustacean Research, 35, 108 - 116." type="journal article" year="2006">Naruse &amp; Ng (2006)</bibRefCitation>
, Galil &amp; Ng (2007, 2009), and Ng
<emphasis id="B930EAFAFFD4292B66D0FDB53EC0D67A" box="[690,746,584,608]" italics="true" pageId="2" pageNumber="42">et al.</emphasis>
(2009) are not relisted here.
</paragraph>
</subSubSection>
</treatment>
</document>