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<document id="20E6837E0E49322FBE7CB3AD5A4D70BF" ID-CLB-Dataset="24185" ID-DOI="10.11646/zootaxa.4731.1.5" ID-GBIF-Dataset="f24b4ebb-c309-431f-996d-c677b5582202" ID-ISSN="1175-5326" ID-Zenodo-Dep="3635652" ID-ZooBank="6C467606-BFA4-468D-ABBB-ECE3B2AE59C6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1580819012913" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2020" docId="03ED87EDFFC79D768FB8FCDBFD1EF874" docLanguage="en" docName="zootaxa.4731.1.5.pdf" docOrigin="Zootaxa 4731 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pachelpheus pachyacanthus Anker 2020, sp. nov." docType="treatment" docVersion="6" lastPageNumber="85" masterDocId="FFD4FF95FFC49D7C8F2FFFB8FF8CFF95" masterDocTitle="A remarkable burrow-dwelling alpheid shrimp, new genus and new species, from the tropical eastern Pacific (Malacostraca: Decapoda: Caridea)" masterLastPageNumber="88" masterPageNumber="75" pageNumber="78" updateTime="1698780339262" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="59D8438E55C80EDCAD775F9F8C469C52" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="51B81CC26AB9E74A0B58434F0B7FB53C">
<mods:title id="2377554ED561FBFF69149107E316FAEB">A remarkable burrow-dwelling alpheid shrimp, new genus and new species, from the tropical eastern Pacific (Malacostraca: Decapoda: Caridea)</mods:title>
</mods:titleInfo>
<mods:name id="46303C24F7379FE2DA29ECB749D281B6" type="personal">
<mods:role id="C7FEED23FC83232327E7226F26C445CE">
<mods:roleTerm id="30CFF1F99A019580ED43B0280A8A879B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F191D621C373BB872B240C686E9FA53A">Anker, Arthur</mods:namePart>
</mods:name>
<mods:typeOfResource id="B54D7C717B52A4B22A638F60F39A9B24">text</mods:typeOfResource>
<mods:relatedItem id="43118522F6F96A832E01287BE662D438" type="host">
<mods:titleInfo id="E0B95B5A9609C3058D4496948B91F7B4">
<mods:title id="4FFC918596DF3B4D4AB5AD408FA4F1DC">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="90560C1E054A20E022444DE381BA8A4C">
<mods:date id="899B1AAC1BF15B2756001F7C6DEA793A">2020</mods:date>
<mods:detail id="A4610709340CB843361EBC5B83771331" type="pubDate">
<mods:number id="1E8CD3496A607AF47E55013E327E50F6">2020-02-04</mods:number>
</mods:detail>
<mods:detail id="9B9DD6D053A0496A27B8657353CA2FAE" type="volume">
<mods:number id="099CAB6575A6F7273B92C804BCF42378">4731</mods:number>
</mods:detail>
<mods:detail id="74B9635EBE2806528902EA141A0E9CFD" type="issue">
<mods:number id="CCF8522DDAD7271FDC46E2E3055C70C8">1</mods:number>
</mods:detail>
<mods:extent id="F28BC8E52A0A6E287BF236261CF24551" unit="page">
<mods:start id="96AB0889F66B74E53D97D9B145360B0E">75</mods:start>
<mods:end id="A0165433D3491995788763975E99E808">88</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="CB7A7D6E3A761E4C7AB4CE37740BEEAF">journal article</mods:classification>
<mods:identifier id="31757D49FFE278F3E8EBBBF0CEF66DFE" type="CLB-Dataset">24185</mods:identifier>
<mods:identifier id="350AACFECA5AC075A0DAE8606CC8A719" type="DOI">10.11646/zootaxa.4731.1.5</mods:identifier>
<mods:identifier id="1E87F9398411C56148A4816B095B3B06" type="GBIF-Dataset">f24b4ebb-c309-431f-996d-c677b5582202</mods:identifier>
<mods:identifier id="5063FB5EA0E90EBAB4F5767750C1B126" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="947D3C472B90A5C2B3DA688DD23C0D85" type="Zenodo-Dep">3635652</mods:identifier>
<mods:identifier id="79C10464C5FB55FA97CBCA5CFD728C33" type="ZooBank">6C467606-BFA4-468D-ABBB-ECE3B2AE59C6</mods:identifier>
</mods:mods>
<treatment id="03ED87EDFFC79D768FB8FCDBFD1EF874" ID-GBIF-Taxon="161939307" LSID="urn:lsid:plazi:treatment:03ED87EDFFC79D768FB8FCDBFD1EF874" httpUri="http://treatment.plazi.org/id/03ED87EDFFC79D768FB8FCDBFD1EF874" lastPageId="10" lastPageNumber="85" pageId="3" pageNumber="78">
<subSubSection id="C35E6570FFC79D7F8FB8FCDBFDD9FCEB" box="[151,597,867,894]" pageId="3" pageNumber="78" type="nomenclature">
<paragraph id="8BFB36FBFFC79D7F8FB8FCDBFDD9FCEB" blockId="3.[151,597,867,930]" box="[151,597,867,894]" pageId="3" pageNumber="78">
<heading id="D0B38197FFC79D7F8FB8FCDBFDD9FCEB" bold="true" box="[151,597,867,894]" fontSize="11" level="1" pageId="3" pageNumber="78" reason="1">
<emphasis id="B930EAE9FFC79D7F8FB8FCDBFDD9FCEB" bold="true" box="[151,597,867,894]" pageId="3" pageNumber="78">
<taxonomicName id="4C444D78FFC79D7F8FB8FCDBFE7CFCEB" ID-CoL="86BB6" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[151,496,867,894]" class="Malacostraca" family="Alpheidae" genus="Pachelpheus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="78" phylum="Arthropoda" rank="species" species="pachyacanthus" status="sp. nov.">
<emphasis id="B930EAE9FFC79D7F8FB8FCDBFE7CFCEB" bold="true" box="[151,496,867,894]" italics="true" pageId="3" pageNumber="78">Pachelpheus pachyacanthus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2035792FFC79D7F8ED8FCDCFDD9FCEB" box="[503,597,868,894]" pageId="3" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35E6570FFC79D7F8FB8FC3FFE8CFC37" box="[151,256,903,930]" pageId="3" pageNumber="78" type="description">
<paragraph id="8BFB36FBFFC79D7F8FB8FC3FFE8CFC37" blockId="3.[151,597,867,930]" box="[151,256,903,930]" pageId="3" pageNumber="78">
<figureCitation id="137F2A7EFFC79D7F8FB8FC3FFE8CFC37" box="[151,256,903,930]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStartId-0="4.[151,250,1177,1202]" captionStartId-1="5.[151,250,1789,1814]" captionStartId-2="6.[151,250,1634,1659]" captionStartId-3="7.[151,250,1542,1567]" captionStartId-4="8.[151,250,1846,1871]" captionStartId-5="9.[151,250,1823,1848]" captionStartId-6="10.[151,250,1637,1662]" captionTargetBox-0="[178,1411,246,1150]" captionTargetBox-1="[185,1402,194,1754]" captionTargetBox-2="[177,1418,191,1585]" captionTargetBox-3="[156,1427,643,1510]" captionTargetBox-4="[163,1420,190,1804]" captionTargetBox-5="[151,1436,181,1798]" captionTargetBox-6="[152,1436,921,1612]" captionTargetId-0="figure@4.[164,1423,234,1150]" captionTargetId-1="figure@5.[164,1423,181,1764]" captionTargetId-2="figure@6.[151,1436,181,1614]" captionTargetId-3="figure@7.[151,1436,633,1517]" captionTargetId-4="figure@8.[151,1436,181,1821]" captionTargetId-5="figure@9.[151,1436,181,1798]" captionTargetId-6="figure@10.[152,1436,921,1612]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionTargetPageId-6="10" captionText-0="FIGURE 1. Pachelpheus pachyacanthus gen. et sp. nov., holotype, male (cl 3.5 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51917): habitus, lateral view." captionText-1="FIGURE 2. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918) [A–G, K, L]; holotype, male (cl 3.5 mm) from the same locality (FLMNH UF 51917) [H–J]: A—frontal margin, dorsal view; B—same, lateral view; C—fifth and sixth pleonites, lateral view; D—telson, dorsal view; E—same, detail of posterior margin, dorsal view; F—ventromesial carina on first article of antennular peduncle, lateral view; G—antennal scaphocerite, dorsal view; H—male first pleopod, mesial (posterior) view; I—male second pleopod, mesial (posterior) view; J—same, appendix masculina and appendix interna, lateral (anterior) view; K—uropod, dorsal view; L—same, detail of distolateral angle of exopod and spiniform setae, dorsal and slightly posterior view." captionText-2="FIGURE 3. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918): A—mandible, mesial view; B—maxillule, lateral view; C—maxilla, lateral view; D—first maxilliped, lateral view; E—same, endopod, mesial view; F—second maxilliped, lateral view; G—third maxilliped, lateral view; H—same, apex of ultimate article, lateral view." captionText-3="FIGURE 4. Pachelpheus pachyacanthus gen. et sp. nov., holotype, male (cl 3.5 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51917):A—right cheliped, lateral view; B—same, distal portion of merus, carpus and chela, mesial view; C—same, merus and carpus, mesial view; D—same, carpus and chela, lateral view." captionText-4="FIGURE 5. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918): A—second pereiopod, lateral view; B—same, chela, mesial view; C—third pereiopod, lateral view; D—same, distal portion of carpus, propodus and dactylus, lateral view; E—fourth pereiopod, lateral view; F—fifth pereiopod, lateral view; G—same, ischium and merus, dorsomesial view; H—same, ischium and merus, lateral view; I—same, distal portion of carpus, propodus and dactylus, mesial view." captionText-5="FIGURE 6. Pachelpheus pachyacanthus gen. et sp. nov., holotype, male (cl 3.5 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51917), living individual:A—dorsal view; B—lateral view. Photographs by the author." captionText-6="FIGURE 7. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918), living individual (chelipeds missing), lateral view. Photograph by the author." figureDoi-0="http://doi.org/10.5281/zenodo.3635654" figureDoi-1="http://doi.org/10.5281/zenodo.3635656" figureDoi-2="http://doi.org/10.5281/zenodo.3635658" figureDoi-3="http://doi.org/10.5281/zenodo.3635660" figureDoi-4="http://doi.org/10.5281/zenodo.3635662" figureDoi-5="http://doi.org/10.5281/zenodo.3635664" figureDoi-6="http://doi.org/10.5281/zenodo.3635666" httpUri-0="https://zenodo.org/record/3635654/files/figure.png" httpUri-1="https://zenodo.org/record/3635656/files/figure.png" httpUri-2="https://zenodo.org/record/3635658/files/figure.png" httpUri-3="https://zenodo.org/record/3635660/files/figure.png" httpUri-4="https://zenodo.org/record/3635662/files/figure.png" httpUri-5="https://zenodo.org/record/3635664/files/figure.png" httpUri-6="https://zenodo.org/record/3635666/files/figure.png" pageId="3" pageNumber="78">Figs. 1–7</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35E6570FFC79D7F8FB8FC72FE73FB0D" pageId="3" pageNumber="78" type="materials_examined">
<paragraph id="8BFB36FBFFC79D7F8FB8FC72FE73FB0D" blockId="3.[151,1437,969,2041]" pageId="3" pageNumber="78">
<emphasis id="B930EAE9FFC79D7F8FB8FC72FECCFC71" bold="true" box="[151,320,969,996]" pageId="3" pageNumber="78">Type material</emphasis>
.
<materialsCitation id="3B2C3CA6FFC79D7F8E7EFC72FE37FBE1" ID-GBIF-Occurrence="2562577884" collectingDate="2019-02-24" collectionCode="FLMNH" collectorName="de Coibita &amp; A. Anker &amp; P. P. G. Pachelle" country="Panama" county="Rancheria" elevation="1" latitude="7.636972" location="Pacific" longLatPrecision="1" longitude="-81.70491" municipality="Coiba Archipelago" pageId="3" pageNumber="78" specimenCode="UF 51917" specimenCount="1" typeStatus="holotype">
<typeStatus id="54FF8859FFC79D7F8E7EFC72FE32FC71" box="[337,446,970,996]" pageId="3" pageNumber="78" type="holotype">Holotype</typeStatus>
: male (cl
<quantity id="4CBC9B1EFFC79D7F8D1EFC71FD07FC70" box="[561,651,969,997]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="3" pageNumber="78" unit="mm" value="3.5">3.5 mm</quantity>
, tl
<quantity id="4CBC9B1EFFC79D7F8D9DFC72FC91FC70" box="[690,797,970,997]" metricMagnitude="-2" metricUnit="m" metricValue="1.23" pageId="3" pageNumber="78" unit="mm" value="12.3">12.3 mm</quantity>
),
<collectionCode id="ED55AE3EFFC79D7F8C1DFC73FC18FC70" box="[818,916,971,997]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="3" pageNumber="78" type="Museum">FLMNH</collectionCode>
<specimenCode id="DBE29E80FFC79D7F8CB2FC73FB98FC71" box="[925,1044,969,997]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="3" pageNumber="78" type="Museum">UF 51917</specimenCode>
,
<collectingCountry id="F353766BFFC79D7F8B0EFC73FBF1FC70" box="[1057,1149,971,997]" name="Panama" pageId="3" pageNumber="78">Panama</collectingCountry>
,
<location id="8E9B6020FFC79D7F8BA5FC72FB54FC71" LSID="urn:lsid:plazi:treatment:03ED87EDFFC79D768FB8FCDBFD1EF874:8E9B6020FFC79D7F8BA5FC72FB54FC71" box="[1162,1240,970,996]" country="Panama" county="Rancheria" latitude="7.636972" longLatPrecision="1" longitude="-81.70491" municipality="Coiba Archipelago" name="Pacific" pageId="3" pageNumber="78">Pacific</location>
coast,
<collectingMunicipality id="6B9FAC81FFC79D7F8A04FC72FE8BFB9D" pageId="3" pageNumber="78">Coiba Archipelago</collectingMunicipality>
, Isla
<collectingCounty id="629A4E77FFC79D7F8E6BFC56FE3AFB9D" box="[324,438,1006,1032]" pageId="3" pageNumber="78">Rancheria</collectingCounty>
(
<collectingCountry id="F353766BFFC79D7F8EE9FC56FE7CFB9D" box="[454,496,1006,1032]" name="Iceland" pageId="3" pageNumber="78">Isla</collectingCountry>
<collectorName id="26B1532DFFC79D7F8ED7FC56FDFEFB9C" box="[504,626,1006,1033]" pageId="3" pageNumber="78">de Coibita</collectorName>
), southern side near STRI boat “parking” area and large rocky outcrop,
<geoCoordinate id="EE70503CFFC79D7F8FB8FBAAFEABFBB9" box="[151,295,1042,1068]" degrees="7" direction="north" minutes="38" orientation="latitude" pageId="3" pageNumber="78" precision="1" seconds="13.1" value="7.636972">7°38’13.1”N</geoCoordinate>
<geoCoordinate id="EE70503CFFC79D7F8E00FBAAFE5FFBB9" box="[303,467,1042,1068]" degrees="81" direction="west" minutes="42" orientation="longitude" pageId="3" pageNumber="78" precision="1" seconds="17.7" value="-81.70491">81°42’17.7”W</geoCoordinate>
, depth:
<quantity id="4CBC9B1EFFC79D7F8D03FBA9FD07FBB8" box="[556,651,1041,1069]" metricMagnitude="0" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="3" pageNumber="78" unit="m" value="1.25" valueMax="1.5" valueMin="1.0">
<elevation id="0069D1C8FFC79D7F8D03FBA9FD07FBB8" box="[556,651,1041,1069]" metricMagnitude="0" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="3" pageNumber="78" unit="m" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 m</elevation>
</quantity>
, subtidal sand flat with some muddier patches, coral rubble and small colonies of living corals, suction (yabby) pump, in burrow (host not collected), leg.
<collectorName id="26B1532DFFC79D7F8B6FFB8EFB2BFBC5" box="[1088,1191,1078,1104]" pageId="3" pageNumber="78">A. Anker</collectorName>
&amp;
<collectorName id="26B1532DFFC79D7F8BE5FB8EFAF9FBC5" box="[1226,1397,1078,1104]" pageId="3" pageNumber="78">P.P.G. Pachelle</collectorName>
,
<date id="FFFA103BFFC79D7F8AAFFB8EFE84FBE1" pageId="3" pageNumber="78" value="2019-02-24">
<collectingDate id="EFBEE9D3FFC79D7F8AAFFB8EFE84FBE1" pageId="3" pageNumber="78" value="2019-02-24">24 Feb. 2019</collectingDate>
</date>
[fcn PAN-282]
</materialsCitation>
.
<materialsCitation id="3B2C3CA6FFC79D7F8EE9FBE3FE77FB0D" ID-GBIF-Occurrence="2562577883" collectingDate="2019-02-24" collectionCode="FLMNH" collectorName="de Coibita &amp; A. Anker &amp; P. P. G. Pachelle" country="Panama" county="Rancheria" elevation="1" latitude="7.636972" location="Pacific" longLatPrecision="1" longitude="-81.70491" municipality="Coiba Archipelago" pageId="3" pageNumber="78" specimenCode="UF 51918" specimenCount="1" typeStatus="paratype">
<typeStatus id="54FF8859FFC79D7F8EE9FBE3FDA0FBE0" box="[454,556,1115,1141]" pageId="3" pageNumber="78" type="paratype">Paratype</typeStatus>
: female (cl
<quantity id="4CBC9B1EFFC79D7F8D9DFBE2FC87FBE0" box="[690,779,1114,1141]" metricMagnitude="-3" metricUnit="m" metricValue="3.6" pageId="3" pageNumber="78" unit="mm" value="3.6">3.6 mm</quantity>
, tl
<quantity id="4CBC9B1EFFC79D7F8C00FBE2FC14FBE0" box="[815,920,1114,1141]" metricMagnitude="-2" metricUnit="m" metricValue="1.26" pageId="3" pageNumber="78" unit="mm" value="12.6">12.6 mm</quantity>
),
<collectionCode id="ED55AE3EFFC79D7F8C83FBE3FB82FBE0" box="[940,1038,1115,1141]" country="USA" httpUri="http://grbio.org/cool/24h4-51mc" name="Florida Museum of Natural History" pageId="3" pageNumber="78" type="Museum">FLMNH</collectionCode>
<specimenCode id="DBE29E80FFC79D7F8B39FBE3FB06FBE1" box="[1046,1162,1113,1141]" collectionCode="UF" country="USA" lsid="urn:lsid:biocol.org:col:34858" name="Florida Museum of Natural History- Zoology, Paleontology and Paleobotany" pageId="3" pageNumber="78" type="Museum">UF 51918</specimenCode>
, same collection data as for the holotype [fcn PAN-280]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C35E6570FFC79D758FE8FB19FABDF85C" lastPageId="9" lastPageNumber="84" pageId="3" pageNumber="78" type="description">
<paragraph id="8BFB36FBFFC79D7F8FE8FB19FDCBFABD" blockId="3.[151,1437,969,2041]" pageId="3" pageNumber="78">
<emphasis id="B930EAE9FFC79D7F8FE8FB19FEDDFB29" bold="true" box="[199,337,1185,1212]" pageId="3" pageNumber="78">Description</emphasis>
. Small-sized (cl
<quantity id="4CBC9B1EFFC79D7F8D24FB19FD1EFB28" box="[523,658,1185,1213]" metricMagnitude="-3" metricUnit="m" metricValue="3.55" metricValueMax="3.6" metricValueMin="3.5" pageId="3" pageNumber="78" unit="mm" value="3.55" valueMax="3.6" valueMin="3.5">3.5–3.6 mm</quantity>
) alpheid shrimp with moderately slender, somewhat compressed body (
<figureCitation id="137F2A7EFFC79D7F8F8FFB7EFF6FFB75" box="[160,227,1222,1248]" captionStart="FIGURE 1" captionStartId="4.[151,250,1177,1202]" captionTargetBox="[178,1411,246,1150]" captionTargetId="figure@4.[164,1423,234,1150]" captionTargetPageId="4" captionText="FIGURE 1. Pachelpheus pachyacanthus gen. et sp. nov., holotype, male (cl 3.5 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51917): habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3635654" httpUri="https://zenodo.org/record/3635654/files/figure.png" pageId="3" pageNumber="78">Fig. 1</figureCitation>
). Carapace smooth, not setose, unarmed; frontal margin with broadly rounded rostral projection, without well-defined rostrum, orbital teeth absent; pterygostomial angle rounded, slightly produced anteriorly (
<figureCitation id="137F2A7EFFC79D7F8A3AFB52FA02FA90" box="[1301,1422,1258,1285]" captionStart="FIGURE 2" captionStartId="5.[151,250,1789,1814]" captionTargetBox="[185,1402,194,1754]" captionTargetId="figure@5.[164,1423,181,1764]" captionTargetPageId="5" captionText="FIGURE 2. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918) [A–G, K, L]; holotype, male (cl 3.5 mm) from the same locality (FLMNH UF 51917) [H–J]: A—frontal margin, dorsal view; B—same, lateral view; C—fifth and sixth pleonites, lateral view; D—telson, dorsal view; E—same, detail of posterior margin, dorsal view; F—ventromesial carina on first article of antennular peduncle, lateral view; G—antennal scaphocerite, dorsal view; H—male first pleopod, mesial (posterior) view; I—male second pleopod, mesial (posterior) view; J—same, appendix masculina and appendix interna, lateral (anterior) view; K—uropod, dorsal view; L—same, detail of distolateral angle of exopod and spiniform setae, dorsal and slightly posterior view." figureDoi="http://doi.org/10.5281/zenodo.3635656" httpUri="https://zenodo.org/record/3635656/files/figure.png" pageId="3" pageNumber="78">Fig. 2A, B</figureCitation>
). Cardiac notch well developed (
<figureCitation id="137F2A7EFFC79D7F8EDBFAB6FDB9FABD" box="[500,565,1294,1320]" captionStart="FIGURE 1" captionStartId="4.[151,250,1177,1202]" captionTargetBox="[178,1411,246,1150]" captionTargetId="figure@4.[164,1423,234,1150]" captionTargetPageId="4" captionText="FIGURE 1. Pachelpheus pachyacanthus gen. et sp. nov., holotype, male (cl 3.5 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51917): habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3635654" httpUri="https://zenodo.org/record/3635654/files/figure.png" pageId="3" pageNumber="78">Fig. 1</figureCitation>
).
</paragraph>
<paragraph id="8BFB36FBFFC79D7F8FE8FA8AFF44FA2D" blockId="3.[151,1437,969,2041]" pageId="3" pageNumber="78">
Pleon with first to fifth pleura broadly rounded posteroventrally; second pleuron greatly expanded in females (
<figureCitation id="137F2A7EFFC79D7F8FB0FAEEFF63FAE5" box="[159,239,1366,1392]" captionStart="FIGURE 1" captionStartId="4.[151,250,1177,1202]" captionTargetBox="[178,1411,246,1150]" captionTargetId="figure@4.[164,1423,234,1150]" captionTargetPageId="4" captionText="FIGURE 1. Pachelpheus pachyacanthus gen. et sp. nov., holotype, male (cl 3.5 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51917): habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3635654" httpUri="https://zenodo.org/record/3635654/files/figure.png" pageId="3" pageNumber="78">Figs. 1</figureCitation>
,
<figureCitation id="137F2A7EFFC79D7F8FD0FAEFFE83FAE4" box="[255,271,1367,1393]" captionStart="FIGURE 7" captionStartId="10.[151,250,1637,1662]" captionTargetBox="[152,1436,921,1612]" captionTargetId="figure@10.[152,1436,921,1612]" captionTargetPageId="10" captionText="FIGURE 7. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918), living individual (chelipeds missing), lateral view. Photograph by the author." figureDoi="http://doi.org/10.5281/zenodo.3635666" httpUri="https://zenodo.org/record/3635666/files/figure.png" pageId="3" pageNumber="78">7</figureCitation>
). Sixth pleonite with slight dorsal constriction near proximal margin; posteroventral angle with small triangular articulated plate; posterolateral margin not produced posteriorly, blunt; preanal plate rounded (
<figureCitation id="137F2A7EFFC79D7F8A69FAC2FA19FA01" box="[1350,1429,1402,1428]" captionStart="FIGURE 1" captionStartId="4.[151,250,1177,1202]" captionTargetBox="[178,1411,246,1150]" captionTargetId="figure@4.[164,1423,234,1150]" captionTargetPageId="4" captionText="FIGURE 1. Pachelpheus pachyacanthus gen. et sp. nov., holotype, male (cl 3.5 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51917): habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3635654" httpUri="https://zenodo.org/record/3635654/files/figure.png" pageId="3" pageNumber="78">Figs. 1</figureCitation>
,
<figureCitation id="137F2A7EFFC79D7F8FB8FA26FF30FA2D" box="[151,188,1438,1464]" captionStart="FIGURE 2" captionStartId="5.[151,250,1789,1814]" captionTargetBox="[185,1402,194,1754]" captionTargetId="figure@5.[164,1423,181,1764]" captionTargetPageId="5" captionText="FIGURE 2. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918) [A–G, K, L]; holotype, male (cl 3.5 mm) from the same locality (FLMNH UF 51917) [H–J]: A—frontal margin, dorsal view; B—same, lateral view; C—fifth and sixth pleonites, lateral view; D—telson, dorsal view; E—same, detail of posterior margin, dorsal view; F—ventromesial carina on first article of antennular peduncle, lateral view; G—antennal scaphocerite, dorsal view; H—male first pleopod, mesial (posterior) view; I—male second pleopod, mesial (posterior) view; J—same, appendix masculina and appendix interna, lateral (anterior) view; K—uropod, dorsal view; L—same, detail of distolateral angle of exopod and spiniform setae, dorsal and slightly posterior view." figureDoi="http://doi.org/10.5281/zenodo.3635656" httpUri="https://zenodo.org/record/3635656/files/figure.png" pageId="3" pageNumber="78">2C</figureCitation>
).
</paragraph>
<paragraph id="8BFB36FBFFC79D7F8FE8FA7AFAE3F9F8" blockId="3.[151,1437,969,2041]" pageId="3" pageNumber="78">
Telson fairly broad, subrectangular, strongly tapering distally, about 1.9 times as long as maximal (proximal) width; dorsal surface with two pairs of small cuspidate setae inserted at small distance from lateral margin, first pair slightly posterior to telson half-length, second pair at about 0.7 of telson length; posterior margin about 0.4 of maximal telson width, broadly rounded, with two pairs of spiniform setae, mesial noticeably thicker and at least three times longer than lateral; three or four long plumose setae arising between mesial spiniform setae (
<figureCitation id="137F2A7EFFC79D7F8BC6F9EAFAEEF9F8" box="[1257,1378,1618,1645]" captionStart="FIGURE 2" captionStartId="5.[151,250,1789,1814]" captionTargetBox="[185,1402,194,1754]" captionTargetId="figure@5.[164,1423,181,1764]" captionTargetPageId="5" captionText="FIGURE 2. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918) [A–G, K, L]; holotype, male (cl 3.5 mm) from the same locality (FLMNH UF 51917) [H–J]: A—frontal margin, dorsal view; B—same, lateral view; C—fifth and sixth pleonites, lateral view; D—telson, dorsal view; E—same, detail of posterior margin, dorsal view; F—ventromesial carina on first article of antennular peduncle, lateral view; G—antennal scaphocerite, dorsal view; H—male first pleopod, mesial (posterior) view; I—male second pleopod, mesial (posterior) view; J—same, appendix masculina and appendix interna, lateral (anterior) view; K—uropod, dorsal view; L—same, detail of distolateral angle of exopod and spiniform setae, dorsal and slightly posterior view." figureDoi="http://doi.org/10.5281/zenodo.3635656" httpUri="https://zenodo.org/record/3635656/files/figure.png" pageId="3" pageNumber="78">Fig. 2D, E</figureCitation>
).
</paragraph>
<paragraph id="8BFB36FBFFC79D7F8FE8F9CFFBDDF921" blockId="3.[151,1437,969,2041]" pageId="3" pageNumber="78">
Eyes fully concealed in dorsal view, partly exposed in lateral view; cornea relatively small, well pigmented; anteromesial margin of eyestalk unarmed (
<figureCitation id="137F2A7EFFC79D7F8D5AF922FD63F920" box="[629,751,1690,1717]" captionStart="FIGURE 2" captionStartId="5.[151,250,1789,1814]" captionTargetBox="[185,1402,194,1754]" captionTargetId="figure@5.[164,1423,181,1764]" captionTargetPageId="5" captionText="FIGURE 2. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918) [A–G, K, L]; holotype, male (cl 3.5 mm) from the same locality (FLMNH UF 51917) [H–J]: A—frontal margin, dorsal view; B—same, lateral view; C—fifth and sixth pleonites, lateral view; D—telson, dorsal view; E—same, detail of posterior margin, dorsal view; F—ventromesial carina on first article of antennular peduncle, lateral view; G—antennal scaphocerite, dorsal view; H—male first pleopod, mesial (posterior) view; I—male second pleopod, mesial (posterior) view; J—same, appendix masculina and appendix interna, lateral (anterior) view; K—uropod, dorsal view; L—same, detail of distolateral angle of exopod and spiniform setae, dorsal and slightly posterior view." figureDoi="http://doi.org/10.5281/zenodo.3635656" httpUri="https://zenodo.org/record/3635656/files/figure.png" pageId="3" pageNumber="78">Fig. 2A, B</figureCitation>
). Epistomial sclerites unarmed.
</paragraph>
<paragraph id="8BFB36FBFFC79D7F8FE8F906FE1CF818" blockId="3.[151,1437,969,2041]" pageId="3" pageNumber="78">
Antennular peduncle stout; stylocerite somewhat appressed to first article, tip blunt or at most subacute, not reaching distal margin of first article; ventromesial carina with anteriorly directed tooth; second article squareshaped in dorsal view, somewhat elongate on mesial side; lateral antennular flagellum much thicker than mesial antennular flagellum, with fused portion consisting of four subdivisions, secondary ramus (= accessory flagellum) well discernable, at least six groups of aesthetascs present on last subdivisions of fused portion and on secondary ramus (
<figureCitation id="137F2A7EFFC79D7F8FC4F8CAFE0FF818" box="[235,387,1906,1933]" captionStart="FIGURE 2" captionStartId="5.[151,250,1789,1814]" captionTargetBox="[185,1402,194,1754]" captionTargetId="figure@5.[164,1423,181,1764]" captionTargetPageId="5" captionText="FIGURE 2. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918) [A–G, K, L]; holotype, male (cl 3.5 mm) from the same locality (FLMNH UF 51917) [H–J]: A—frontal margin, dorsal view; B—same, lateral view; C—fifth and sixth pleonites, lateral view; D—telson, dorsal view; E—same, detail of posterior margin, dorsal view; F—ventromesial carina on first article of antennular peduncle, lateral view; G—antennal scaphocerite, dorsal view; H—male first pleopod, mesial (posterior) view; I—male second pleopod, mesial (posterior) view; J—same, appendix masculina and appendix interna, lateral (anterior) view; K—uropod, dorsal view; L—same, detail of distolateral angle of exopod and spiniform setae, dorsal and slightly posterior view." figureDoi="http://doi.org/10.5281/zenodo.3635656" httpUri="https://zenodo.org/record/3635656/files/figure.png" pageId="3" pageNumber="78">Fig. 2A, B, F</figureCitation>
).
</paragraph>
<paragraph id="8BFB36FBFFC79D788FE8F82EFDB5FF42" blockId="3.[151,1437,969,2041]" lastBlockId="4.[151,1436,153,215]" lastPageId="4" lastPageNumber="79" pageId="3" pageNumber="78">
Antenna with basicerite very stout, its distoventral margin with large subacute tooth; scaphocerite relatively small, subrectangular, not reaching end of antennular peduncle, with straight lateral margin and rather narrow blade, latter not overreaching small distolateral tooth; carpocerite stout, cylindrical, reaching well beyond scaphocerite and end of antennular peduncle; flagellum long, longer than half of body, relatively thick, although thinner than lateral antennular flagellum (
<figureCitation id="137F2A7EFFC09D788EA0FF05FDA0FF42" box="[399,556,189,215]" captionStart="FIGURE 2" captionStartId="5.[151,250,1789,1814]" captionTargetBox="[185,1402,194,1754]" captionTargetId="figure@5.[164,1423,181,1764]" captionTargetPageId="5" captionText="FIGURE 2. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918) [A–G, K, L]; holotype, male (cl 3.5 mm) from the same locality (FLMNH UF 51917) [H–J]: A—frontal margin, dorsal view; B—same, lateral view; C—fifth and sixth pleonites, lateral view; D—telson, dorsal view; E—same, detail of posterior margin, dorsal view; F—ventromesial carina on first article of antennular peduncle, lateral view; G—antennal scaphocerite, dorsal view; H—male first pleopod, mesial (posterior) view; I—male second pleopod, mesial (posterior) view; J—same, appendix masculina and appendix interna, lateral (anterior) view; K—uropod, dorsal view; L—same, detail of distolateral angle of exopod and spiniform setae, dorsal and slightly posterior view." figureDoi="http://doi.org/10.5281/zenodo.3635656" httpUri="https://zenodo.org/record/3635656/files/figure.png" pageId="4" pageNumber="79">Fig. 2A, B, G</figureCitation>
).
</paragraph>
<caption id="DF3B6673FFC09D788FB8FB21FD2FFB43" ID-DOI="http://doi.org/10.5281/zenodo.3635654" ID-Zenodo-Dep="3635654" httpUri="https://zenodo.org/record/3635654/files/figure.png" pageId="4" pageNumber="79" startId="4.[151,250,1177,1202]" targetBox="[178,1411,246,1150]" targetPageId="4">
<paragraph id="8BFB36FBFFC09D788FB8FB21FD2FFB43" blockId="4.[151,1437,1177,1238]" pageId="4" pageNumber="79">
<emphasis id="B930EAE9FFC09D788FB8FB21FE9FFB27" bold="true" box="[151,275,1177,1202]" pageId="4" pageNumber="79">FIGURE 1.</emphasis>
<taxonomicName id="4C444D78FFC09D788E36FB22FDB5FB27" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[281,569,1178,1202]" class="Malacostraca" family="Alpheidae" genus="Pachelpheus" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="79" phylum="Arthropoda" rank="species" species="pachyacanthus" status="gen. et sp. nov.">
<emphasis id="B930EAE9FFC09D788E36FB22FDB5FB27" box="[281,569,1178,1202]" italics="true" pageId="4" pageNumber="79">Pachelpheus pachyacanthus</emphasis>
</taxonomicName>
<emphasis id="B930EAE9FFC09D788D10FB22FD50FB27" bold="true" box="[575,732,1178,1202]" pageId="4" pageNumber="79">
<taxonomicNameLabel id="A2035792FFC09D788D10FB22FD50FB27" box="[575,732,1178,1202]" pageId="4" pageNumber="79" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, male (cl 3.5 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51917): habitus, lateral view.
</paragraph>
</caption>
<paragraph id="8BFB36FBFFC09D788FE8FABCFD7CF95A" blockId="4.[151,1437,1284,2031]" pageId="4" pageNumber="79">
Mouthparts typical for family. Mandible with robust molar process; incisor process terminating in five cutting teeth, second-dorsal largest; palp with faint subdivision between narrow first (proximal) and expanded second (distal) article (
<figureCitation id="137F2A7EFFC09D788E34FAF4FEFAFAF2" box="[283,374,1356,1383]" captionStart="FIGURE 3" captionStartId="6.[151,250,1634,1659]" captionTargetBox="[177,1418,191,1585]" captionTargetId="figure@6.[151,1436,181,1614]" captionTargetPageId="6" captionText="FIGURE 3. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918): A—mandible, mesial view; B—maxillule, lateral view; C—maxilla, lateral view; D—first maxilliped, lateral view; E—same, endopod, mesial view; F—second maxilliped, lateral view; G—third maxilliped, lateral view; H—same, apex of ultimate article, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3635658" httpUri="https://zenodo.org/record/3635658/files/figure.png" pageId="4" pageNumber="79">Fig. 3A</figureCitation>
). Maxillule with endopod (palp) bilobed, dorsal lobe furnished with slender seta, ventral lobe somewhat square-shaped, without seta; dorsal endite somewhat square-shaped, ventral endite with thick setae (
<figureCitation id="137F2A7EFFC09D788A40FAC8FF37FA3A" captionStart="FIGURE 3" captionStartId="6.[151,250,1634,1659]" captionTargetBox="[177,1418,191,1585]" captionTargetId="figure@6.[151,1436,181,1614]" captionTargetPageId="6" captionText="FIGURE 3. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918): A—mandible, mesial view; B—maxillule, lateral view; C—maxilla, lateral view; D—first maxilliped, lateral view; E—same, endopod, mesial view; F—second maxilliped, lateral view; G—third maxilliped, lateral view; H—same, apex of ultimate article, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3635658" httpUri="https://zenodo.org/record/3635658/files/figure.png" pageId="4" pageNumber="79">Fig. 3B</figureCitation>
). Maxilla with narrow scaphognathite; endopod simple, without setae; dorsal endite with moderately deep cleft; ventral endite short, with some elongate setae (
<figureCitation id="137F2A7EFFC09D788D8EFA00FD75FA46" box="[673,761,1464,1491]" captionStart="FIGURE 3" captionStartId="6.[151,250,1634,1659]" captionTargetBox="[177,1418,191,1585]" captionTargetId="figure@6.[151,1436,181,1614]" captionTargetPageId="6" captionText="FIGURE 3. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918): A—mandible, mesial view; B—maxillule, lateral view; C—maxilla, lateral view; D—first maxilliped, lateral view; E—same, endopod, mesial view; F—second maxilliped, lateral view; G—third maxilliped, lateral view; H—same, apex of ultimate article, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3635658" httpUri="https://zenodo.org/record/3635658/files/figure.png" pageId="4" pageNumber="79">Fig. 3C</figureCitation>
). First maxilliped with dorsal and ventral endites fused; exopod with moderately expanded caridean lobe; endopod not clearly subdivided, furnished with row of setae mesially, with two long thick setae distally; epipod bilobed (
<figureCitation id="137F2A7EFFC09D788DFBF9B8FCC3F98E" box="[724,847,1536,1563]" captionStart="FIGURE 3" captionStartId="6.[151,250,1634,1659]" captionTargetBox="[177,1418,191,1585]" captionTargetId="figure@6.[151,1436,181,1614]" captionTargetPageId="6" captionText="FIGURE 3. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918): A—mandible, mesial view; B—maxillule, lateral view; C—maxilla, lateral view; D—first maxilliped, lateral view; E—same, endopod, mesial view; F—second maxilliped, lateral view; G—third maxilliped, lateral view; H—same, apex of ultimate article, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3635658" httpUri="https://zenodo.org/record/3635658/files/figure.png" pageId="4" pageNumber="79">Fig. 3D, E</figureCitation>
). Second maxilliped with typically shaped endopod, propodal and dactylar articles distinct, forming apical “head”; exopod thickened in its proximal half; epipod rather small (
<figureCitation id="137F2A7EFFC09D788FCCF9F0FEBBF9F6" box="[227,311,1608,1635]" captionStart="FIGURE 3" captionStartId="6.[151,250,1634,1659]" captionTargetBox="[177,1418,191,1585]" captionTargetId="figure@6.[151,1436,181,1614]" captionTargetPageId="6" captionText="FIGURE 3. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918): A—mandible, mesial view; B—maxillule, lateral view; C—maxilla, lateral view; D—first maxilliped, lateral view; E—same, endopod, mesial view; F—second maxilliped, lateral view; G—third maxilliped, lateral view; H—same, apex of ultimate article, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3635658" httpUri="https://zenodo.org/record/3635658/files/figure.png" pageId="4" pageNumber="79">Fig. 3F</figureCitation>
). Third maxilliped slender, pediform; coxa with lateral plate subrectangular, almost ear-shaped; antepenultimate article slender, not twisted or markedly flattened; penultimate article about four times as long as wide; ultimate article slightly tapering distally, with numerous rows of short serrulate setae, tip with two robust spiniform setae; arthrobranch present, rather small (
<figureCitation id="137F2A7EFFC09D788D48F90CFD68F95A" box="[615,740,1716,1743]" captionStart="FIGURE 3" captionStartId="6.[151,250,1634,1659]" captionTargetBox="[177,1418,191,1585]" captionTargetId="figure@6.[151,1436,181,1614]" captionTargetPageId="6" captionText="FIGURE 3. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918): A—mandible, mesial view; B—maxillule, lateral view; C—maxilla, lateral view; D—first maxilliped, lateral view; E—same, endopod, mesial view; F—second maxilliped, lateral view; G—third maxilliped, lateral view; H—same, apex of ultimate article, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3635658" httpUri="https://zenodo.org/record/3635658/files/figure.png" pageId="4" pageNumber="79">Fig. 3G, H</figureCitation>
).
</paragraph>
<paragraph id="8BFB36FBFFC09D788FE8F960FD1FF87A" blockId="4.[151,1437,1284,2031]" pageId="4" pageNumber="79">
Chelipeds moderately enlarged, equal in size, symmetrical in shape, carried extended with dactylus in ventrolateral position (
<figureCitation id="137F2A7EFFC09D788E60F944FE1DF882" box="[335,401,1788,1815]" captionStart="FIGURE 1" captionStartId="4.[151,250,1177,1202]" captionTargetBox="[178,1411,246,1150]" captionTargetId="figure@4.[164,1423,234,1150]" captionTargetPageId="4" captionText="FIGURE 1. Pachelpheus pachyacanthus gen. et sp. nov., holotype, male (cl 3.5 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51917): habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3635654" httpUri="https://zenodo.org/record/3635654/files/figure.png" pageId="4" pageNumber="79">Fig. 1</figureCitation>
); basis and ischium short, robust, unarmed; merus short, stout, flattened ventrally, unarmed distally and on margins, distodorsal margin forming blunt angle; carpus wide, short, cup-shaped, mesial surface without setal rows; palm slightly enlarged, somewhat swollen, smooth, subcylindrical in cross-section; fingers short, about 0.6 length of palm, slightly gaping when closed, tips crossing; cutting edge of pollex with large tooth occupying about 0.4–0.6 of pollex length, fitting into broad distal hiatus on opposing dactylar cutting edge; cutting edge of dactylus with large tooth occupying about 0.2–0.3 of dacylar length, fitting into broad proximal hiatus on opposing cutting edge of pollex (
<figureCitation id="137F2A7EFFC09D788D26F86CFDC0F87A" box="[521,588,2004,2031]" captionStart="FIGURE 4" captionStartId="7.[151,250,1542,1567]" captionTargetBox="[156,1427,643,1510]" captionTargetId="figure@7.[151,1436,633,1517]" captionTargetPageId="7" captionText="FIGURE 4. Pachelpheus pachyacanthus gen. et sp. nov., holotype, male (cl 3.5 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51917):A—right cheliped, lateral view; B—same, distal portion of merus, carpus and chela, mesial view; C—same, merus and carpus, mesial view; D—same, carpus and chela, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3635660" httpUri="https://zenodo.org/record/3635660/files/figure.png" pageId="4" pageNumber="79">Fig. 4</figureCitation>
A–D).
</paragraph>
<caption id="DF3B6673FFC19D798FB8F945FC73F87B" ID-DOI="http://doi.org/10.5281/zenodo.3635656" ID-Zenodo-Dep="3635656" httpUri="https://zenodo.org/record/3635656/files/figure.png" pageId="5" pageNumber="80" startId="5.[151,250,1789,1814]" targetBox="[185,1402,194,1754]" targetPageId="5">
<paragraph id="8BFB36FBFFC19D798FB8F945FC73F87B" blockId="5.[151,1437,1789,2030]" pageId="5" pageNumber="80">
<emphasis id="B930EAE9FFC19D798FB8F945FE9EF883" bold="true" box="[151,274,1789,1814]" pageId="5" pageNumber="80">FIGURE 2.</emphasis>
<taxonomicName id="4C444D78FFC19D798E38F946FDB9F883" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[279,565,1790,1814]" class="Malacostraca" family="Alpheidae" genus="Pachelpheus" kingdom="Animalia" order="Decapoda" pageId="5" pageNumber="80" phylum="Arthropoda" rank="species" species="pachyacanthus" status="gen. et sp. nov.">
<emphasis id="B930EAE9FFC19D798E38F946FDB9F883" box="[279,565,1790,1814]" italics="true" pageId="5" pageNumber="80">Pachelpheus pachyacanthus</emphasis>
</taxonomicName>
<emphasis id="B930EAE9FFC19D798D14F946FD58F883" bold="true" box="[571,724,1790,1814]" pageId="5" pageNumber="80">
<taxonomicNameLabel id="A2035792FFC19D798D14F946FD58F883" box="[571,724,1790,1814]" pageId="5" pageNumber="80" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918) [A–G, K, L]; holotype, male (cl 3.5 mm) from the same locality (FLMNH UF 51917) [H–J]: A—frontal margin, dorsal view; B—same, lateral view; C—fifth and sixth pleonites, lateral view; D—telson, dorsal view; E—same, detail of posterior margin, dorsal view; F—ventromesial carina on first article of antennular peduncle, lateral view; G—antennal scaphocerite, dorsal view; H—male first pleopod, mesial (posterior) view; I—male second pleopod, mesial (posterior) view; J—same, appendix masculina and appendix interna, lateral (anterior) view; K—uropod, dorsal view; L—same, detail of distolateral angle of exopod and spiniform setae, dorsal and slightly posterior view.
</paragraph>
</caption>
<caption id="DF3B6673FFC29D7A8FB8F9DAFDFBF972" ID-DOI="http://doi.org/10.5281/zenodo.3635658" ID-Zenodo-Dep="3635658" httpUri="https://zenodo.org/record/3635658/files/figure.png" pageId="6" pageNumber="81" startId="6.[151,250,1634,1659]" targetBox="[177,1418,191,1585]" targetPageId="6">
<paragraph id="8BFB36FBFFC29D7A8FB8F9DAFDFBF972" blockId="6.[151,1437,1634,1767]" pageId="6" pageNumber="81">
<emphasis id="B930EAE9FFC29D7A8FB8F9DAFE9EF9EE" bold="true" box="[151,274,1634,1659]" pageId="6" pageNumber="81">FIGURE 3.</emphasis>
<taxonomicName id="4C444D78FFC29D7A8E38F9DAFDB9F9EF" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[279,565,1634,1658]" class="Malacostraca" family="Alpheidae" genus="Pachelpheus" kingdom="Animalia" order="Decapoda" pageId="6" pageNumber="81" phylum="Arthropoda" rank="species" species="pachyacanthus" status="gen. et sp. nov.">
<emphasis id="B930EAE9FFC29D7A8E38F9DAFDB9F9EF" box="[279,565,1634,1658]" italics="true" pageId="6" pageNumber="81">Pachelpheus pachyacanthus</emphasis>
</taxonomicName>
<emphasis id="B930EAE9FFC29D7A8D14F9DAFD58F9EF" bold="true" box="[571,724,1634,1658]" pageId="6" pageNumber="81">
<taxonomicNameLabel id="A2035792FFC29D7A8D14F9DAFD58F9EF" box="[571,724,1634,1658]" pageId="6" pageNumber="81" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918): A—mandible, mesial view; B—maxillule, lateral view; C—maxilla, lateral view; D—first maxilliped, lateral view; E—same, endopod, mesial view; F—second maxilliped, lateral view; G—third maxilliped, lateral view; H—same, apex of ultimate article, lateral view.
</paragraph>
</caption>
<paragraph id="8BFB36FBFFC29D7A8FE8F8ADFD77F80E" blockId="6.[151,1436,1813,2019]" pageId="6" pageNumber="81">
Second pereiopod not particularly slender or elongate; ischium long, unarmed; merus much shorter than ischium; carpus with five subarticles, first longest but not longer than sum of others, ratio of subarticles approximately equal to 3.5/1.0/1.0/1.1/1.8; chela longer than distal-most carpal subarticle, simple, cutting edges of fingers furnished with rows of microscopic setae (
<figureCitation id="137F2A7EFFC29D7A8D5BF839FD61F80E" box="[628,749,1920,1947]" captionStart="FIGURE 5" captionStartId="8.[151,250,1846,1871]" captionTargetBox="[163,1420,190,1804]" captionTargetId="figure@8.[151,1436,181,1821]" captionTargetPageId="8" captionText="FIGURE 5. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918): A—second pereiopod, lateral view; B—same, chela, mesial view; C—third pereiopod, lateral view; D—same, distal portion of carpus, propodus and dactylus, lateral view; E—fourth pereiopod, lateral view; F—fifth pereiopod, lateral view; G—same, ischium and merus, dorsomesial view; H—same, ischium and merus, lateral view; I—same, distal portion of carpus, propodus and dactylus, mesial view." figureDoi="http://doi.org/10.5281/zenodo.3635662" httpUri="https://zenodo.org/record/3635662/files/figure.png" pageId="6" pageNumber="81">Fig. 5A, B</figureCitation>
).
</paragraph>
<paragraph id="8BFB36FBFFC29D7B8FE8F81DFCA5FDF6" blockId="6.[151,1436,1813,2019]" lastBlockId="7.[151,1437,153,611]" lastPageId="7" lastPageNumber="82" pageId="6" pageNumber="81">
Third pereiopod robust; ischium armed with one very stout, thick cuspidate seta on ventrolateral surface; merus about four times as long as maximal width, armed with two stout, thick cuspidate setae on ventrolateral surface; carpus slenderer than merus, about 0.6 times as long as merus, distoventral margin armed with one slender spiniform seta; propodus noticeably longer than carpus, ventral margin with two slender spiniform setae on proximal half and pair of spiniform setae near dactylar base; dactylus moderately slender, somewhat elongate, conical, simple, slightly curving distally, about 0.7 length of propodus (
<figureCitation id="137F2A7EFFC39D7B8C3BFEBDFC18FE8A" box="[788,916,260,287]" captionStart="FIGURE 5" captionStartId="8.[151,250,1846,1871]" captionTargetBox="[163,1420,190,1804]" captionTargetId="figure@8.[151,1436,181,1821]" captionTargetPageId="8" captionText="FIGURE 5. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918): A—second pereiopod, lateral view; B—same, chela, mesial view; C—third pereiopod, lateral view; D—same, distal portion of carpus, propodus and dactylus, lateral view; E—fourth pereiopod, lateral view; F—fifth pereiopod, lateral view; G—same, ischium and merus, dorsomesial view; H—same, ischium and merus, lateral view; I—same, distal portion of carpus, propodus and dactylus, mesial view." figureDoi="http://doi.org/10.5281/zenodo.3635662" httpUri="https://zenodo.org/record/3635662/files/figure.png" pageId="7" pageNumber="82">Fig. 5C, D</figureCitation>
). Fourth pereiopod generally similar to third, except for armature of merus, latter armed with three stout, thick cuspidate setae on ventrolateral surface (
<figureCitation id="137F2A7EFFC39D7B8A17FE91FA03FED6" box="[1336,1423,296,323]" captionStart="FIGURE 5" captionStartId="8.[151,250,1846,1871]" captionTargetBox="[163,1420,190,1804]" captionTargetId="figure@8.[151,1436,181,1821]" captionTargetPageId="8" captionText="FIGURE 5. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918): A—second pereiopod, lateral view; B—same, chela, mesial view; C—third pereiopod, lateral view; D—same, distal portion of carpus, propodus and dactylus, lateral view; E—fourth pereiopod, lateral view; F—fifth pereiopod, lateral view; G—same, ischium and merus, dorsomesial view; H—same, ischium and merus, lateral view; I—same, distal portion of carpus, propodus and dactylus, mesial view." figureDoi="http://doi.org/10.5281/zenodo.3635662" httpUri="https://zenodo.org/record/3635662/files/figure.png" pageId="7" pageNumber="82">Fig. 5E</figureCitation>
). Fifth pereiopod somewhat slenderer than fourth pereiopod; ischium armed with one very stout, thick, flattened, distally truncate (possibly abraded) cuspidate seta on ventrolateral surface; merus about five times as long as maximal width, armed with four stout, thick, flattened, distally truncate (possibly abraded) cuspidate setae on ventrolateral surface; carpus slenderer than merus, about half-length of merus, distoventral margin armed with one spiniform seta; propodus much longer than carpus, ventral margin unarmed, one stout spiniform seta present on mesial side near dactylar base, distal half of propodus with rows of progressively longer serrulate setae on ventrolateral surface, forming grooming brush; dactylus moderately slender, somewhat elongate, conical, simple, more conspicuously curving distally, about half-length of propodus (
<figureCitation id="137F2A7EFFC39D7B8D80FDF1FD7DFDF6" box="[687,753,584,611]" captionStart="FIGURE 5" captionStartId="8.[151,250,1846,1871]" captionTargetBox="[163,1420,190,1804]" captionTargetId="figure@8.[151,1436,181,1821]" captionTargetPageId="8" captionText="FIGURE 5. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918): A—second pereiopod, lateral view; B—same, chela, mesial view; C—third pereiopod, lateral view; D—same, distal portion of carpus, propodus and dactylus, lateral view; E—fourth pereiopod, lateral view; F—fifth pereiopod, lateral view; G—same, ischium and merus, dorsomesial view; H—same, ischium and merus, lateral view; I—same, distal portion of carpus, propodus and dactylus, mesial view." figureDoi="http://doi.org/10.5281/zenodo.3635662" httpUri="https://zenodo.org/record/3635662/files/figure.png" pageId="7" pageNumber="82">Fig. 5</figureCitation>
F–I).
</paragraph>
<caption id="DF3B6673FFC39D7B8FB8F9BEFC69F9F2" ID-DOI="http://doi.org/10.5281/zenodo.3635660" ID-Zenodo-Dep="3635660" httpUri="https://zenodo.org/record/3635660/files/figure.png" pageId="7" pageNumber="82" startId="7.[151,250,1542,1567]" targetBox="[156,1427,643,1510]" targetPageId="7">
<paragraph id="8BFB36FBFFC39D7B8FB8F9BEFC69F9F2" blockId="7.[151,1437,1542,1639]" pageId="7" pageNumber="82">
<emphasis id="B930EAE9FFC39D7B8FB8F9BEFE9FF98A" bold="true" box="[151,275,1542,1567]" pageId="7" pageNumber="82">FIGURE 4.</emphasis>
<taxonomicName id="4C444D78FFC39D7B8E36F9BEFDB5F98B" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[281,569,1542,1566]" class="Malacostraca" family="Alpheidae" genus="Pachelpheus" kingdom="Animalia" order="Decapoda" pageId="7" pageNumber="82" phylum="Arthropoda" rank="species" species="pachyacanthus" status="gen. et sp. nov.">
<emphasis id="B930EAE9FFC39D7B8E36F9BEFDB5F98B" box="[281,569,1542,1566]" italics="true" pageId="7" pageNumber="82">Pachelpheus pachyacanthus</emphasis>
</taxonomicName>
<emphasis id="B930EAE9FFC39D7B8D10F9BEFD50F98B" bold="true" box="[575,732,1542,1566]" pageId="7" pageNumber="82">
<taxonomicNameLabel id="A2035792FFC39D7B8D10F9BEFD50F98B" box="[575,732,1542,1566]" pageId="7" pageNumber="82" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, male (cl 3.5 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51917):A—right cheliped, lateral view; B—same, distal portion of merus, carpus and chela, mesial view; C—same, merus and carpus, mesial view; D—same, carpus and chela, lateral view.
</paragraph>
</caption>
<paragraph id="8BFB36FBFFC39D7B8FE8F92CFA91F963" blockId="7.[151,1436,1684,1999]" pageId="7" pageNumber="82">
First male pleopod with small endopod furnished with some setae (
<figureCitation id="137F2A7EFFC39D7B8C9CF92CFB81F93A" box="[947,1037,1684,1711]" captionStart="FIGURE 2" captionStartId="5.[151,250,1789,1814]" captionTargetBox="[185,1402,194,1754]" captionTargetId="figure@5.[164,1423,181,1764]" captionTargetPageId="5" captionText="FIGURE 2. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918) [A–G, K, L]; holotype, male (cl 3.5 mm) from the same locality (FLMNH UF 51917) [H–J]: A—frontal margin, dorsal view; B—same, lateral view; C—fifth and sixth pleonites, lateral view; D—telson, dorsal view; E—same, detail of posterior margin, dorsal view; F—ventromesial carina on first article of antennular peduncle, lateral view; G—antennal scaphocerite, dorsal view; H—male first pleopod, mesial (posterior) view; I—male second pleopod, mesial (posterior) view; J—same, appendix masculina and appendix interna, lateral (anterior) view; K—uropod, dorsal view; L—same, detail of distolateral angle of exopod and spiniform setae, dorsal and slightly posterior view." figureDoi="http://doi.org/10.5281/zenodo.3635656" httpUri="https://zenodo.org/record/3635656/files/figure.png" pageId="7" pageNumber="82">Fig. 2H</figureCitation>
). Second male pleopod with appendix masculina much longer than appendix interna, furnished with numerous stiff setae apically and subapically, as illustrated (
<figureCitation id="137F2A7EFFC39D7B8E38F964FEF1F962" box="[279,381,1756,1783]" captionStart="FIGURE 2" captionStartId="5.[151,250,1789,1814]" captionTargetBox="[185,1402,194,1754]" captionTargetId="figure@5.[164,1423,181,1764]" captionTargetPageId="5" captionText="FIGURE 2. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918) [A–G, K, L]; holotype, male (cl 3.5 mm) from the same locality (FLMNH UF 51917) [H–J]: A—frontal margin, dorsal view; B—same, lateral view; C—fifth and sixth pleonites, lateral view; D—telson, dorsal view; E—same, detail of posterior margin, dorsal view; F—ventromesial carina on first article of antennular peduncle, lateral view; G—antennal scaphocerite, dorsal view; H—male first pleopod, mesial (posterior) view; I—male second pleopod, mesial (posterior) view; J—same, appendix masculina and appendix interna, lateral (anterior) view; K—uropod, dorsal view; L—same, detail of distolateral angle of exopod and spiniform setae, dorsal and slightly posterior view." figureDoi="http://doi.org/10.5281/zenodo.3635656" httpUri="https://zenodo.org/record/3635656/files/figure.png" pageId="7" pageNumber="82">Fig. 2I, J</figureCitation>
). Second female pleopod without appendix masculina, with appendix interna only.
</paragraph>
<paragraph id="8BFB36FBFFC39D7B8FE8F8B8FB1EF83E" blockId="7.[151,1436,1684,1999]" pageId="7" pageNumber="82">
Uropod with lateral lobe of protopod very broad, rounded; exopod narrowly ovoid, slightly truncate distally; distolateral tooth absent; diaeresis (transverse suture) straight, lateral section conspicuously thickened, elevated, armed with two very stout, thick spiniform setae; distal and distomesial margin with row of slender spiniform setae above marginal plumose setae; endopod more narrowly ovoid, distal and distomesial margin with row of slender spiniform setae above marginal plumose setae, less numerous than on exopod (
<figureCitation id="137F2A7EFFC39D7B8B23F828FB0AF83E" box="[1036,1158,1936,1963]" captionStart="FIGURE 2" captionStartId="5.[151,250,1789,1814]" captionTargetBox="[185,1402,194,1754]" captionTargetId="figure@5.[164,1423,181,1764]" captionTargetPageId="5" captionText="FIGURE 2. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918) [A–G, K, L]; holotype, male (cl 3.5 mm) from the same locality (FLMNH UF 51917) [H–J]: A—frontal margin, dorsal view; B—same, lateral view; C—fifth and sixth pleonites, lateral view; D—telson, dorsal view; E—same, detail of posterior margin, dorsal view; F—ventromesial carina on first article of antennular peduncle, lateral view; G—antennal scaphocerite, dorsal view; H—male first pleopod, mesial (posterior) view; I—male second pleopod, mesial (posterior) view; J—same, appendix masculina and appendix interna, lateral (anterior) view; K—uropod, dorsal view; L—same, detail of distolateral angle of exopod and spiniform setae, dorsal and slightly posterior view." figureDoi="http://doi.org/10.5281/zenodo.3635656" httpUri="https://zenodo.org/record/3635656/files/figure.png" pageId="7" pageNumber="82">Fig. 2K, L</figureCitation>
).
</paragraph>
<paragraph id="8BFB36FBFFC39D7B8FE8F80CFD39F85A" blockId="7.[151,1436,1684,1999]" box="[199,693,1972,1999]" pageId="7" pageNumber="82">Gill formula as given for genus (see above).</paragraph>
<caption id="DF3B6673FFCC9D748FB8F88EFD72F84A" ID-DOI="http://doi.org/10.5281/zenodo.3635662" ID-Zenodo-Dep="3635662" httpUri="https://zenodo.org/record/3635662/files/figure.png" pageId="8" pageNumber="83" startId="8.[151,250,1846,1871]" targetBox="[163,1420,190,1804]" targetPageId="8">
<paragraph id="8BFB36FBFFCC9D748FB8F88EFD72F84A" blockId="8.[151,1437,1846,2015]" pageId="8" pageNumber="83">
<emphasis id="B930EAE9FFCC9D748FB8F88EFE9EF8DA" bold="true" box="[151,274,1846,1871]" pageId="8" pageNumber="83">FIGURE 5.</emphasis>
<taxonomicName id="4C444D78FFCC9D748E38F88FFDB9F8DA" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[279,565,1847,1871]" class="Malacostraca" family="Alpheidae" genus="Pachelpheus" kingdom="Animalia" order="Decapoda" pageId="8" pageNumber="83" phylum="Arthropoda" rank="species" species="pachyacanthus" status="gen. et sp. nov.">
<emphasis id="B930EAE9FFCC9D748E38F88FFDB9F8DA" box="[279,565,1847,1871]" italics="true" pageId="8" pageNumber="83">Pachelpheus pachyacanthus</emphasis>
</taxonomicName>
<emphasis id="B930EAE9FFCC9D748D14F88FFD58F8DA" bold="true" box="[571,724,1847,1871]" pageId="8" pageNumber="83">
<taxonomicNameLabel id="A2035792FFCC9D748D14F88FFD58F8DA" box="[571,724,1847,1871]" pageId="8" pageNumber="83" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918): A—second pereiopod, lateral view; B—same, chela, mesial view; C—third pereiopod, lateral view; D—same, distal portion of carpus, propodus and dactylus, lateral view; E—fourth pereiopod, lateral view; F—fifth pereiopod, lateral view; G—same, ischium and merus, dorsomesial view; H—same, ischium and merus, lateral view; I—same, distal portion of carpus, propodus and dactylus, mesial view.
</paragraph>
</caption>
<caption id="DF3B6673FFCD9D758FB8F8A7FB7CF8C9" ID-DOI="http://doi.org/10.5281/zenodo.3635664" ID-Zenodo-Dep="3635664" httpUri="https://zenodo.org/record/3635664/files/figure.png" pageId="9" pageNumber="84" startId="9.[151,250,1823,1848]" targetBox="[151,1436,181,1798]" targetPageId="9">
<paragraph id="8BFB36FBFFCD9D758FB8F8A7FB7CF8C9" blockId="9.[151,1437,1823,1884]" pageId="9" pageNumber="84">
<emphasis id="B930EAE9FFCD9D758FB8F8A7FE9FF8AD" bold="true" box="[151,275,1823,1848]" pageId="9" pageNumber="84">FIGURE 6.</emphasis>
<taxonomicName id="4C444D78FFCD9D758E36F898FDB5F8AD" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[281,569,1824,1848]" class="Malacostraca" family="Alpheidae" genus="Pachelpheus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="84" phylum="Arthropoda" rank="species" species="pachyacanthus" status="gen. et sp. nov.">
<emphasis id="B930EAE9FFCD9D758E36F898FDB5F8AD" box="[281,569,1824,1848]" italics="true" pageId="9" pageNumber="84">Pachelpheus pachyacanthus</emphasis>
</taxonomicName>
<emphasis id="B930EAE9FFCD9D758D10F898FD50F8AD" bold="true" box="[575,732,1824,1848]" pageId="9" pageNumber="84">
<taxonomicNameLabel id="A2035792FFCD9D758D10F898FD50F8AD" box="[575,732,1824,1848]" pageId="9" pageNumber="84" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype, male (cl 3.5 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51917), living individual:A—dorsal view; B—lateral view. Photographs by the author.
</paragraph>
</caption>
<paragraph id="8BFB36FBFFCD9D758FE8F832FABDF85C" blockId="9.[151,1436,1930,2029]" pageId="9" pageNumber="84">
<emphasis id="B930EAE9FFCD9D758FE8F832FED4F830" bold="true" box="[199,344,1930,1957]" pageId="9" pageNumber="84">Colouration</emphasis>
. Body semi-translucent, more or less evenly covered with large reddish spots (= groups of red chromatophores); yellow ovary visible through partial translucence of carapace in the female (
<figureCitation id="137F2A7EFFCD9D758B97F816FA8BF85D" box="[1208,1287,1966,1992]" captionStart="FIGURE 6" captionStartId="9.[151,250,1823,1848]" captionTargetBox="[151,1436,181,1798]" captionTargetId="figure@9.[151,1436,181,1798]" captionTargetPageId="9" captionText="FIGURE 6. Pachelpheus pachyacanthus gen. et sp. nov., holotype, male (cl 3.5 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51917), living individual:A—dorsal view; B—lateral view. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.3635664" httpUri="https://zenodo.org/record/3635664/files/figure.png" pageId="9" pageNumber="84">Figs. 6</figureCitation>
,
<figureCitation id="137F2A7EFFCD9D758A3CF817FAAFF85C" box="[1299,1315,1967,1993]" captionStart="FIGURE 7" captionStartId="10.[151,250,1637,1662]" captionTargetBox="[152,1436,921,1612]" captionTargetId="figure@10.[152,1436,921,1612]" captionTargetPageId="10" captionText="FIGURE 7. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918), living individual (chelipeds missing), lateral view. Photograph by the author." figureDoi="http://doi.org/10.5281/zenodo.3635666" httpUri="https://zenodo.org/record/3635666/files/figure.png" pageId="9" pageNumber="84">7</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35E6570FFCD9D768FE8F86BFCCAFF6E" lastPageId="10" lastPageNumber="85" pageId="9" pageNumber="84" type="etymology">
<paragraph id="8BFB36FBFFCD9D768FE8F86BFCCAFF6E" blockId="9.[151,1436,1930,2029]" lastBlockId="10.[151,1437,153,899]" lastPageId="10" lastPageNumber="85" pageId="9" pageNumber="84">
<emphasis id="B930EAE9FFCD9D758FE8F86BFEC4F878" bold="true" box="[199,328,2003,2029]" pageId="9" pageNumber="84">Etymology</emphasis>
. The new species’ name is a combination of two Latinised Greek words
<emphasis id="B930EAE9FFCD9D758BA3F86AFB43F879" box="[1164,1231,2002,2028]" italics="true" pageId="9" pageNumber="84">pachy</emphasis>
(from the ancient Greek
<emphasis id="B930EAE9FFCE9D768FCDFF21FEBDFF26" box="[226,305,153,179]" italics="true" pageId="10" pageNumber="85">pakhús</emphasis>
= thick) and
<emphasis id="B930EAE9FFCE9D768EEAFF21FDA6FF26" box="[453,554,153,179]" italics="true" pageId="10" pageNumber="85">acanthus</emphasis>
(from the ancient Greek
<emphasis id="B930EAE9FFCE9D768C6CFF21FC24FF26" box="[835,936,153,179]" italics="true" pageId="10" pageNumber="85">akanthos</emphasis>
—spine) and refers to the presence of unusually thick cuspidate setae on the third, fourth and fifth pereiopods, as well as two very thick spiniform setae on the lateral portion of the uropodal diaeresis; used as an adjective.
</paragraph>
</subSubSection>
<subSubSection id="C35E6570FFCE9D768FE8FEBDFE9AFE3A" pageId="10" pageNumber="85" type="biology_ecology">
<paragraph id="8BFB36FBFFCE9D768FE8FEBDFE9AFE3A" blockId="10.[151,1437,153,899]" pageId="10" pageNumber="85">
<emphasis id="B930EAE9FFCE9D768FE8FEBDFEAAFE8A" bold="true" box="[199,294,261,287]" pageId="10" pageNumber="85">Ecology</emphasis>
. Both specimens were collected from burrows of unknown hosts, on a shallow subtidal sand flat, about
<quantity id="4CBC9B1EFFCE9D768FB8FE91FF5CFED6" box="[151,208,297,323]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="10" pageNumber="85" unit="m" value="30.0">30 m</quantity>
from the sandy shore and
<quantity id="4CBC9B1EFFCE9D768ED9FE90FDA3FED6" box="[502,559,296,323]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="10" pageNumber="85" unit="m" value="50.0">50 m</quantity>
from a large rocky outcrop, at a depth of about
<quantity id="4CBC9B1EFFCE9D768B6CFE90FB13FED6" box="[1091,1183,296,323]" metricMagnitude="0" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="10" pageNumber="85" unit="m" value="1.25" valueMax="1.5" valueMin="1.0">1–1.5 m</quantity>
at incoming tight. The sand flat is characterised by relatively fine, more or less compacted sand, with an abundance of fragmented coral rubble, and some muddier patches, larger pieces of rubble, and small living colonies of the coral
<taxonomicName id="4C444D78FFCE9D768BE1FEC9FF51FE3A" baseAuthorityName="Linnaeus" baseAuthorityYear="1758" class="Anthozoa" family="Pocilloporidae" genus="Pocillopora" kingdom="Animalia" order="Scleractinia" pageId="10" pageNumber="85" phylum="Cnidaria" rank="species" species="damicornis">
<emphasis id="B930EAE9FFCE9D768BE1FEC9FF51FE3A" italics="true" pageId="10" pageNumber="85">Pocillopora damicornis</emphasis>
</taxonomicName>
(L.).
</paragraph>
</subSubSection>
<subSubSection id="C35E6570FFCE9D768FE8FE01FD1EF874" pageId="10" pageNumber="85" type="discussion">
<paragraph id="8BFB36FBFFCE9D768FE8FE01FB74FC16" blockId="10.[151,1437,153,899]" pageId="10" pageNumber="85">
<emphasis id="B930EAE9FFCE9D768FE8FE01FEB8FE46" bold="true" box="[199,308,441,467]" pageId="10" pageNumber="85">Remarks</emphasis>
. The presence of cuspidate setae on the meri of the third to fifth pereiopods is not uncommon in alpheid shrimps, being also known in several so-called “lower” alpheid genera, such as
<taxonomicName id="4C444D78FFCE9D768B46FE65FA8FFE62" authorityName="Anker &amp; Dworschak" authorityYear="2007" box="[1129,1283,477,503]" class="Malacostraca" family="Alpheidae" genus="Jengalpheops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAE9FFCE9D768B46FE65FA8FFE62" box="[1129,1283,477,503]" italics="true" pageId="10" pageNumber="85">Jengalpheops</emphasis>
</taxonomicName>
(monotypic),
<taxonomicName id="4C444D78FFCE9D768FB8FDB9FEC9FD8E" authorityName="Powell" authorityYear="1979" box="[151,325,513,539]" class="Malacostraca" family="Alpheidae" genus="Potamalpheops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAE9FFCE9D768FB8FDB9FEC9FD8E" box="[151,325,513,539]" italics="true" pageId="10" pageNumber="85">Potamalpheops</emphasis>
</taxonomicName>
(most if not all species) and
<taxonomicName id="4C444D78FFCE9D768D8FFDB9FC5DFD8E" authority="Coutiere, 1897" authorityName="Coutiere" authorityYear="1897" box="[672,977,513,539]" class="Malacostraca" family="Alpheidae" genus="Athanopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAE9FFCE9D768D8FFDB9FC90FD8E" box="[672,796,513,539]" italics="true" pageId="10" pageNumber="85">Athanopsis</emphasis>
<bibRefCitation id="EFD54B0AFFCE9D768C08FDB9FC5DFD8E" author="Coutiere, H." box="[807,977,513,539]" pageId="10" pageNumber="85" pagination="380 - 386" refId="ref7859" refString="Coutiere, H. (1897) Note sur quelques genres nouveaux ou peu connus d'Alpheides, formant la sousfamille des Alpheopsides. Bulletin du Museum d'Histoire Naturelle, 2, 380 - 386." type="journal article" year="1897">Coutiere, 1897</bibRefCitation>
</taxonomicName>
(some species), but also in “higher” or more derived genera, such as
<taxonomicName id="4C444D78FFCE9D768ECCFD9DFC91FDAA" authority="Coutiere, 1897" authorityName="Coutiere" authorityYear="1897" box="[483,797,549,575]" class="Malacostraca" family="Alpheidae" genus="Parabetaeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAE9FFCE9D768ECCFD9DFDFDFDAA" box="[483,625,549,575]" italics="true" pageId="10" pageNumber="85">Parabetaeus</emphasis>
<bibRefCitation id="EFD54B0AFFCE9D768D58FD9DFC91FDAA" author="Coutiere, H." box="[631,797,549,575]" pageId="10" pageNumber="85" pagination="380 - 386" refId="ref7859" refString="Coutiere, H. (1897) Note sur quelques genres nouveaux ou peu connus d'Alpheides, formant la sousfamille des Alpheopsides. Bulletin du Museum d'Histoire Naturelle, 2, 380 - 386." type="journal article" year="1897">Coutière, 1897</bibRefCitation>
</taxonomicName>
(all species),
<taxonomicName id="4C444D78FFCE9D768C97FD9EFB9DFDAA" authorityName="Dana" authorityYear="1852" box="[952,1041,550,575]" class="Malacostraca" family="Alpheidae" genus="Betaeus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAE9FFCE9D768C97FD9EFB9DFDAA" box="[952,1041,550,575]" italics="true" pageId="10" pageNumber="85">Betaeus</emphasis>
</taxonomicName>
(most if not all species) and
<taxonomicName id="4C444D78FFCE9D768A7CFD9EFF5DFDF6" authorityName="Yaldwyn" authorityYear="1971" class="Malacostraca" family="Alpheidae" genus="Betaeopsis" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAE9FFCE9D768A7CFD9EFF5DFDF6" italics="true" pageId="10" pageNumber="85">Betaeopsis</emphasis>
</taxonomicName>
(both species) (e.g.,
<bibRefCitation id="EFD54B0AFFCE9D768EEEFDF1FDDFFDF6" author="Powell, C. B." box="[449,595,585,611]" pageId="10" pageNumber="85" pagination="116 - 150" refId="ref8556" refString="Powell, C. B. (1979) Three alpheid shrimps of a new genus from West African fresh and brackish waters: taxonomy and ecologial zonation (Crustacea Decapoda Natantia). Revue de Zoologie Africaine, 93, 116 - 150." type="journal article" year="1979">Powell 1979</bibRefCitation>
;
<bibRefCitation id="EFD54B0AFFCE9D768D4FFDF1FCE7FDF6" author="Nomura, K. &amp; Anker, A." box="[608,875,585,611]" pageId="10" pageNumber="85" pagination="43 - 54" refId="ref8414" refString="Nomura, K. &amp; Anker, A. (2001) Redescription of Parabetaeus culliereti Coutiere and status of the genus Neoalpheopsis Banner (Decapoda: Caridea: Alpheidae). Crustacean Research, 30, 43 - 54. https: // doi. org / 10.18353 / crustacea. 30.0 _ 43" type="journal article" year="2001">Nomura &amp; Anker 2001</bibRefCitation>
;
<bibRefCitation id="EFD54B0AFFCE9D768C58FDF1FBD0FDF6" author="Anker, A. &amp; Jeng, M. S." box="[887,1116,585,611]" pageId="10" pageNumber="85" pagination="570 - 584" refId="ref7191" refString="Anker, A. &amp; Jeng, M. S. (2002) Redefinition of Betaeopsis Yaldwyn, 1971, with invalidation of generic status of Hamalpheus Bruce &amp; Iliffe, 1991 (Crustacea: Decapoda: Alpheidae). Proceedings of the Biological Society of Washington, 115, 570 - 584." type="journal article" year="2002">Anker &amp; Jeng 2002</bibRefCitation>
;
<bibRefCitation id="EFD54B0AFFCE9D768B48FDF1FA1BFDF6" author="Anker, A. &amp; Dworschak, P. C." box="[1127,1431,585,611]" pageId="10" pageNumber="85" pagination="290 - 302" refId="ref7012" refString="Anker, A. &amp; Dworschak, P. C. (2007) Jengalpheops rufus, gen. nov., sp. nov., a new commensal alpheid shrimp from the Philippines (Crustacea: Decapoda). Zoological Studies, 46, 290 - 302." type="journal article" year="2007">Anker &amp; Dworschak 2007</bibRefCitation>
; Anker 2012, 2015). However, in
<taxonomicName id="4C444D78FFCE9D768D33FDD5FCD2FD12" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[540,862,621,647]" class="Malacostraca" family="Alpheidae" genus="Pachelpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="85" phylum="Arthropoda" rank="species" species="pachyacanthus" status="sp. nov.">
<emphasis id="B930EAE9FFCE9D768D33FDD5FCD2FD12" box="[540,862,621,647]" italics="true" pageId="10" pageNumber="85">Pachelpheus pachyacanthus</emphasis>
</taxonomicName>
<emphasis id="B930EAE9FFCE9D768C46FDD5FC4AFD12" bold="true" box="[873,966,621,647]" pageId="10" pageNumber="85">
<taxonomicNameLabel id="A2035792FFCE9D768C46FDD5FC4AFD12" box="[873,966,621,647]" pageId="10" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the cuspidate setae of walking legs are conspicuously thickened and attain truly unusual dimensions; in fact they are readily visible in the general view of the shrimps, especially on the merus of the fifth pereiopod (
<figureCitation id="137F2A7EFFCE9D768C17FD0DFC09FD5A" box="[824,901,693,719]" captionStart="FIGURE 1" captionStartId="4.[151,250,1177,1202]" captionTargetBox="[178,1411,246,1150]" captionTargetId="figure@4.[164,1423,234,1150]" captionTargetPageId="4" captionText="FIGURE 1. Pachelpheus pachyacanthus gen. et sp. nov., holotype, male (cl 3.5 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51917): habitus, lateral view." figureDoi="http://doi.org/10.5281/zenodo.3635654" httpUri="https://zenodo.org/record/3635654/files/figure.png" pageId="10" pageNumber="85">Figs. 1</figureCitation>
,
<figureCitation id="137F2A7EFFCE9D768CBBFD0DFC28FD5A" box="[916,932,693,719]" captionStart="FIGURE 6" captionStartId="9.[151,250,1823,1848]" captionTargetBox="[151,1436,181,1798]" captionTargetId="figure@9.[151,1436,181,1798]" captionTargetPageId="9" captionText="FIGURE 6. Pachelpheus pachyacanthus gen. et sp. nov., holotype, male (cl 3.5 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51917), living individual:A—dorsal view; B—lateral view. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.3635664" httpUri="https://zenodo.org/record/3635664/files/figure.png" pageId="10" pageNumber="85">6</figureCitation>
,
<figureCitation id="137F2A7EFFCE9D768C9FFD0DFC4CFD5A" box="[944,960,693,719]" captionStart="FIGURE 7" captionStartId="10.[151,250,1637,1662]" captionTargetBox="[152,1436,921,1612]" captionTargetId="figure@10.[152,1436,921,1612]" captionTargetPageId="10" captionText="FIGURE 7. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918), living individual (chelipeds missing), lateral view. Photograph by the author." figureDoi="http://doi.org/10.5281/zenodo.3635666" httpUri="https://zenodo.org/record/3635666/files/figure.png" pageId="10" pageNumber="85">7</figureCitation>
). The most robust of these cuspidate setae seem to be somewhat flattened and their sockets are relatively shallow, i.e. not accommodating them completely; in addition, those of the fifth pereiopod seem to be somewhat abraded (
<figureCitation id="137F2A7EFFCE9D768CBCFD45FB82FC82" box="[915,1038,764,791]" captionStart="FIGURE 5" captionStartId="8.[151,250,1846,1871]" captionTargetBox="[163,1420,190,1804]" captionTargetId="figure@8.[151,1436,181,1821]" captionTargetPageId="8" captionText="FIGURE 5. Pachelpheus pachyacanthus gen. et sp. nov., paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918): A—second pereiopod, lateral view; B—same, chela, mesial view; C—third pereiopod, lateral view; D—same, distal portion of carpus, propodus and dactylus, lateral view; E—fourth pereiopod, lateral view; F—fifth pereiopod, lateral view; G—same, ischium and merus, dorsomesial view; H—same, ischium and merus, lateral view; I—same, distal portion of carpus, propodus and dactylus, mesial view." figureDoi="http://doi.org/10.5281/zenodo.3635662" httpUri="https://zenodo.org/record/3635662/files/figure.png" pageId="10" pageNumber="85">Fig. 5G, H</figureCitation>
). Their function remains enigmatic, but it is possible that they are somehow involved in the shrimp’s clinging either to the host or to the burrow walls, and perhaps also in digging. It also remains unknown whether these setae are actually homologous to the more conventional cuspidate setae, which are conical, not flattened, never abraded, and lodged in deeper pits.
</paragraph>
<caption id="DF3B6673FFCE9D768FB8F9DDFB56F937" ID-DOI="http://doi.org/10.5281/zenodo.3635666" ID-Zenodo-Dep="3635666" httpUri="https://zenodo.org/record/3635666/files/figure.png" pageId="10" pageNumber="85" startId="10.[151,250,1637,1662]" targetBox="[152,1436,921,1612]" targetPageId="10">
<paragraph id="8BFB36FBFFCE9D768FB8F9DDFB56F937" blockId="10.[151,1437,1637,1698]" pageId="10" pageNumber="85">
<emphasis id="B930EAE9FFCE9D768FB8F9DDFE9EF9EB" bold="true" box="[151,274,1637,1662]" pageId="10" pageNumber="85">FIGURE 7.</emphasis>
<taxonomicName id="4C444D78FFCE9D768E38F9DDFDB9F9E8" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[279,565,1637,1661]" class="Malacostraca" family="Alpheidae" genus="Pachelpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="85" phylum="Arthropoda" rank="species" species="pachyacanthus" status="gen. et sp. nov.">
<emphasis id="B930EAE9FFCE9D768E38F9DDFDB9F9E8" box="[279,565,1637,1661]" italics="true" pageId="10" pageNumber="85">Pachelpheus pachyacanthus</emphasis>
</taxonomicName>
<emphasis id="B930EAE9FFCE9D768D14F9DEFD58F9EB" bold="true" box="[571,724,1638,1662]" pageId="10" pageNumber="85">
<taxonomicNameLabel id="A2035792FFCE9D768D14F9DEFD58F9EB" box="[571,724,1638,1662]" pageId="10" pageNumber="85" rank="species">gen. et sp. nov.</taxonomicNameLabel>
</emphasis>
, paratype, female (cl 3.6 mm) from Isla Rancheria, Coiba Archipelago, Panama (FLMNH UF 51918), living individual (chelipeds missing), lateral view. Photograph by the author.
</paragraph>
</caption>
<paragraph id="8BFB36FBFFCE9D768FE8F972FD1EF874" blockId="10.[151,1437,1738,2017]" pageId="10" pageNumber="85">
<taxonomicName id="4C444D78FFCE9D768FE8F972FD89F971" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[199,517,1738,1764]" class="Malacostraca" family="Alpheidae" genus="Pachelpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="85" phylum="Arthropoda" rank="species" species="pachyacanthus" status="sp. nov.">
<emphasis id="B930EAE9FFCE9D768FE8F972FD89F971" box="[199,517,1738,1764]" italics="true" pageId="10" pageNumber="85">Pachelpheus pachyacanthus</emphasis>
</taxonomicName>
<emphasis id="B930EAE9FFCE9D768D23F973FDE9F970" bold="true" box="[524,613,1739,1765]" pageId="10" pageNumber="85">
<taxonomicNameLabel id="A2035792FFCE9D768D23F973FDE9F970" box="[524,613,1739,1765]" pageId="10" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
is presumably an infaunal symbiont (“commensal”), dwelling in burrows of yet unknown hosts. Other infaunal animals collected at the
<typeStatus id="54FF8859FFCE9D768C66F957FCF5F89C" box="[841,889,1775,1801]" pageId="10" pageNumber="85">type</typeStatus>
locality of
<taxonomicName id="4C444D78FFCE9D768CD7F957FB37F89D" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[1016,1211,1774,1800]" class="Malacostraca" family="Alpheidae" genus="Pachelpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="85" phylum="Arthropoda" rank="species" species="pachyacanthus" status="sp. nov.">
<emphasis id="B930EAE9FFCE9D768CD7F957FB37F89D" box="[1016,1211,1774,1800]" italics="true" pageId="10" pageNumber="85">P. pachyacanthus</emphasis>
</taxonomicName>
<emphasis id="B930EAE9FFCE9D768BEDF957FA97F89C" bold="true" box="[1218,1307,1775,1801]" pageId="10" pageNumber="85">
<taxonomicNameLabel id="A2035792FFCE9D768BEDF957FA97F89C" box="[1218,1307,1775,1801]" pageId="10" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
include the snapping shrimps
<taxonomicName id="4C444D78FFCE9D768E4BF8AAFE32F8B9" authorityName="Fabricius" authorityYear="1798" box="[356,446,1810,1836]" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAE9FFCE9D768E4BF8AAFE32F8B9" box="[356,446,1810,1836]" italics="true" pageId="10" pageNumber="85">Alpheus</emphasis>
</taxonomicName>
cf.
<emphasis id="B930EAE9FFCE9D768EC4F8ABFDACF8B9" box="[491,544,1811,1836]" italics="true" pageId="10" pageNumber="85">naos</emphasis>
<bibRefCitation id="EFD54B0AFFCE9D768D09F8AAFC04F8B8" author="Anker, A. &amp; Hurt, C. &amp; Knowlton, N." box="[550,904,1810,1837]" pageId="10" pageNumber="85" pagination="1 - 23" refId="ref7128" refString="Anker, A., Hurt, C. &amp; Knowlton, N. (2007) Three transisthmian snapping shrimps (Crustacea: Decapoda: Alpheidae: Alpheus) associated with innkeeper worms (Echiura: Thalassematidae). Zootaxa, 1626 (1), 1 - 23. https: // doi. org / 10.11646 / zootaxa. 1626.1.1" type="journal article" year="2007">Anker, Hurt &amp; Knowlton, 2007</bibRefCitation>
and
<taxonomicName id="4C444D78FFCE9D768CEFF8ABFF5EF8C4" authority="Bracken-Grissom &amp; Felder, 2014" authorityName="Bracken-Grissom &amp; Felder" authorityYear="2014" class="Malacostraca" family="Alpheidae" genus="Alpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="85" phylum="Arthropoda" rank="species" species="hephaestus">
<emphasis id="B930EAE9FFCE9D768CEFF8ABFBD0F8B9" box="[960,1116,1810,1837]" italics="true" pageId="10" pageNumber="85">A. hephaestus</emphasis>
<bibRefCitation id="EFD54B0AFFCE9D768B4BF8AAFF5EF8C4" author="Bracken-Grissom, H. D. &amp; Felder, D. L." pageId="10" pageNumber="85" pagination="451 - 491" refId="ref7792" refString="Bracken-Grissom, H. D. &amp; Felder, D. L. (2014) Provisional revision of American snapping shrimp allied to Alpheus floridanus Kingsley, 1878 (Crustacea: Decapoda: Alpheidae) with notes on A. floridanus africanus. Zootaxa, 3895 (4), 451 - 491. https: // doi. org / 10.11646 / zootaxa. 3895.4.1" type="journal article" year="2014">Bracken-Grissom &amp; Felder, 2014</bibRefCitation>
</taxonomicName>
, mud and ghost shrimps (
<taxonomicName id="4C444D78FFCE9D768EDEF88EFD09F8C5" authorityName="Borradaile" authorityYear="1903" box="[497,645,1846,1872]" class="Malacostraca" family="Upogebiidae" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="85" phylum="Arthropoda" rank="family">Upogebiidae</taxonomicName>
,
<taxonomicName id="4C444D78FFCE9D768DA1F88EFD67F8C4" box="[654,747,1846,1873]" class="Insecta" family="Axiidae" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="85" phylum="Arthropoda" rank="family">Axiidae</taxonomicName>
), pea crabs (
<taxonomicName id="4C444D78FFCE9D768C59F88EFB94F8C4" authorityName="De Haan" authorityYear="1833" box="[886,1048,1846,1873]" class="Malacostraca" family="Pinnotheridae" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="85" phylum="Arthropoda" rank="family">Pinnotheridae</taxonomicName>
), stomatopods (
<taxonomicName id="4C444D78FFCE9D768BE8F88EFAB2F8C4" box="[1223,1342,1846,1873]" class="Malacostraca" family="Squillidae" kingdom="Animalia" order="Stomatopoda" pageId="10" pageNumber="85" phylum="Arthropoda" rank="family">Squillidae</taxonomicName>
), thalassematid echiurans, sipunculans, and a single galeommatid bivalve, indicating a highly diversified burrowing fauna (material under study). Interestingly, the single species of the genus
<taxonomicName id="4C444D78FFCE9D768CA2F8C6FBABF80D" authorityName="Anker &amp; Dworschak" authorityYear="2007" box="[909,1063,1918,1944]" class="Malacostraca" family="Alpheidae" genus="Jengalpheops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="85" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAE9FFCE9D768CA2F8C6FBABF80D" box="[909,1063,1918,1944]" italics="true" pageId="10" pageNumber="85">Jengalpheops</emphasis>
</taxonomicName>
, which may be a close relative of
<taxonomicName id="4C444D78FFCE9D768FB8F81AFEAAF829" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[151,294,1954,1980]" class="Malacostraca" family="Alpheidae" genus="Pachelpheus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="85" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B930EAE9FFCE9D768FB8F81AFEAAF829" box="[151,294,1954,1980]" italics="true" pageId="10" pageNumber="85">Pachelpheus</emphasis>
</taxonomicName>
<emphasis id="B930EAE9FFCE9D768E01F81BFE15F828" bold="true" box="[302,409,1955,1981]" pageId="10" pageNumber="85">
<taxonomicNameLabel id="A2035792FFCE9D768E01F81BFE15F828" box="[302,409,1955,1981]" pageId="10" pageNumber="85" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, is an obligate symbiont of burrows of large ghost shrimps,
<taxonomicName id="4C444D78FFCE9D768B68F81AFA97F829" authorityName="A.Milne-Edwards" authorityYear="1870" box="[1095,1307,1954,1980]" class="Malacostraca" family="Callianassidae" genus="Glypturus" kingdom="Animalia" order="Decapoda" pageId="10" pageNumber="85" phylum="Arthropoda" rank="species" species="armatus">
<emphasis id="B930EAE9FFCE9D768B68F81AFA97F829" box="[1095,1307,1954,1980]" italics="true" pageId="10" pageNumber="85">Glypturus armatus</emphasis>
</taxonomicName>
(A. Milne- Edwards, 1870) (
<bibRefCitation id="EFD54B0AFFCE9D768E78F87EFD0AF874" author="Anker, A. &amp; Dworschak, P. C." box="[343,646,1990,2017]" pageId="10" pageNumber="85" pagination="290 - 302" refId="ref7012" refString="Anker, A. &amp; Dworschak, P. C. (2007) Jengalpheops rufus, gen. nov., sp. nov., a new commensal alpheid shrimp from the Philippines (Crustacea: Decapoda). Zoological Studies, 46, 290 - 302." type="journal article" year="2007">Anker &amp; Dworschak 2007</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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