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<document id="29043C9A7CDFCD110262E73ADAC29974" ID-CLB-Dataset="21117" ID-DOI="10.1080/00222933.2015.1042411" ID-GBIF-Dataset="fcb40b61-77e8-4d89-bb8a-ad06b7fe4b46" ID-ISSN="1464-5262" ID-Zenodo-Dep="4000854" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1598383427830" checkinUser="carolina" docAuthor="Buffington, M. L. &amp; Copeland, R. S." docDate="2015" docId="03ED87BDFF8CFFC820D2DFD5FE96FAA5" docLanguage="en" docName="JNATHIST.49.43-44.2597-2607.pdf" docOrigin="Journal of Natural History 49 (43)" docStyle="DocumentStyle:210803DF447D5DCB84349979EDD3507F.6:JNatHist.2013-2015.journal_article.2cover" docStyleId="210803DF447D5DCB84349979EDD3507F" docStyleName="JNatHist.2013-2015.journal_article.2cover" docStyleVersion="6" docTitle="Muhaka Buffington &amp; Copeland 2015, new genus" docType="treatment" docUuid="CB5647B6-47A8-48A7-B063-262FCAF8A4BC" docUuidSource="ZooBank" docVersion="9" lastPageNumber="2602" masterDocId="FFD4FFC5FF8EFFCD2165DB3AFFBAFF86" masterDocTitle="Muhaka icipe, an enigmatic new genus and species of Kleidotomini (Hymenoptera: Figitidae: Eucoilinae) from an East African coastal forest" masterLastPageNumber="2607" masterPageNumber="2597" pageNumber="2599" updateTime="1698845466776" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="CDEBF7C5F30E51B61E5065DFB7C14618">Muhaka icipe, an enigmatic new genus and species of Kleidotomini (Hymenoptera: Figitidae: Eucoilinae) from an East African coastal forest</mods:title>
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<mods:namePart id="0CA409490B6F48097B3EB5E5CF7E53DF">Buffington, M. L.</mods:namePart>
<mods:affiliation id="F411E7FD6CD100DF2184CFE4D916B6B9">Systematic Entomology Laboratory, ARS / USDA, c / o Smithsonian Institution, National Museum of Natural History, Washington, DC, USA;</mods:affiliation>
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<mods:namePart id="04E2E045DA814997ECD6D14F025B4F1E">Copeland, R. S.</mods:namePart>
<mods:affiliation id="4D746AD11BBFECDA875A5461D1EA08D5">International Centre of Insect Physiology and Ecology (ICIPE), Nairobi, Kenya; &amp; Division of Invertebrate Zoology, National Museums of Kenya, Nairobi, Kenya</mods:affiliation>
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<mods:date id="92A0CCEE4240AA922F20A05E6B3021D4">2015</mods:date>
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<mods:number id="D56F92AAFBE6389156D14EE2D5B01E06">49</mods:number>
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<treatment id="03ED87BDFF8CFFC820D2DFD5FE96FAA5" ID-DOI="http://doi.org/10.5281/zenodo.4324247" ID-GBIF-Taxon="167308393" ID-Zenodo-Dep="4324247" LSID="urn:lsid:zoobank.org:act:CB5647B6-47A8-48A7-B063-262FCAF8A4BC" httpUri="http://treatment.plazi.org/id/03ED87BDFF8CFFC820D2DFD5FE96FAA5" lastPageId="5" lastPageNumber="2602" pageId="2" pageNumber="2599">
<subSubSection id="C35E6520FF8CFFCF20D2DFD5FC0AFA8E" box="[439,944,1263,1288]" pageId="2" pageNumber="2599" type="nomenclature">
<paragraph id="8BFB36ABFF8CFFCF20D2DFD5FC0AFA8E" blockId="2.[439,944,1263,1319]" box="[439,944,1263,1288]" pageId="2" pageNumber="2599">
<heading id="D0B381C7FF8CFFCF20D2DFD5FC0AFA8E" box="[439,944,1263,1288]" centered="true" fontSize="10" level="2" pageId="2" pageNumber="2599" reason="2">
<taxonomicName id="4C444D28FF8CFFCF20D2DFD5FC94FA81" authority="Buffington &amp; Copeland" authorityName="Buffington &amp; Copeland" authorityYear="2015" box="[439,814,1263,1287]" class="Insecta" family="Figitidae" genus="Muhaka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="2599" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B930EAB9FF8CFFCF20D2DFD5FDAFFA81" bold="true" box="[439,533,1263,1287]" italics="true" pageId="2" pageNumber="2599">Muhaka</emphasis>
Buffington &amp; Copeland
</taxonomicName>
,
<taxonomicNameLabel id="A20357C2FF8CFFCF225EDFCAFC0AFA8E" box="[827,944,1264,1288]" pageId="2" pageNumber="2599" rank="genus">new genus</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35E6520FF8CFFCF2314DE35FD4CFAA1" box="[625,758,1295,1319]" pageId="2" pageNumber="2599" type="description">
<paragraph id="8BFB36ABFF8CFFCF2314DE35FD4CFAA1" blockId="2.[439,944,1263,1319]" box="[625,758,1295,1319]" pageId="2" pageNumber="2599">
<figureCitation id="137F2A2EFF8CFFCF2314DE35FD4CFAA1" box="[625,758,1295,1319]" captionStart-0="Figure 1" captionStart-1="Figure 2" captionStart-2="Figure 3" captionStartId-0="2.[212,279,978,1000]" captionStartId-1="3.[144,211,921,943]" captionStartId-2="4.[212,279,902,924]" captionTargetBox-0="[212,1172,168,946]" captionTargetBox-1="[144,1104,168,889]" captionTargetBox-2="[213,1172,168,870]" captionTargetId-0="figure@2.[212,1173,168,946]" captionTargetId-1="figure@3.[144,1104,168,889]" captionTargetId-2="figure@4.[212,1172,168,870]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="Figure 1. Anterolateral habitus of Muhaka icipe Buffington and Copeland, n. gen., n. sp." captionText-1="Figure 2. Scanning electron micrographs of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (holotype specimen). (A) Head and mesosoma, dorsal view; (B) head, anteroventral view; (C) head and mesosoma, dorsolateral view; (D) head and pronotum, anterolateral view. Abbreviations: ank, ankos; kem, kemnina; sct, bifurcate posterior margin of scutellum." captionText-2="Figure 3. Habitus and habitat images of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (A) Dorsal habitus, with inset of scutellar and propodeal morphology; (B) lateral habitus (holotype specimen); (C) fore wing, bright-field illumination; (D) Muhaka Forest 6 m Malaise trap." figureDoi-0="http://doi.org/10.5281/zenodo.4000856" figureDoi-1="http://doi.org/10.5281/zenodo.4000858" figureDoi-2="http://doi.org/10.5281/zenodo.4000860" httpUri-0="https://zenodo.org/record/4000856/files/figure.png" httpUri-1="https://zenodo.org/record/4000858/files/figure.png" httpUri-2="https://zenodo.org/record/4000860/files/figure.png" pageId="2" pageNumber="2599">Figures 13</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="C35E6520FF8CFFCF21B1DE7FFC78FADA" box="[212,962,1348,1372]" pageId="2" pageNumber="2599" type="type_taxon">
<paragraph id="8BFB36ABFF8CFFCF21B1DE7FFC78FADA" blockId="2.[212,962,1348,1372]" box="[212,962,1348,1372]" pageId="2" pageNumber="2599">
<heading id="D0B381C7FF8CFFCF21B1DE7FFC78FADA" box="[212,962,1348,1372]" fontSize="10" level="3" pageId="2" pageNumber="2599" reason="5">
<typeStatus id="54FF8809FF8CFFCF21B1DE7FFEB7FADA" box="[212,269,1349,1372]" pageId="2" pageNumber="2599">Type</typeStatus>
species:
<taxonomicName id="4C444D28FF8CFFCF2017DE7EFC91FADA" authority="Buffington &amp; Copeland" authorityName=", Buffington &amp; Copeland" authorityYear="2015" box="[370,811,1348,1372]" class="Insecta" family="Figitidae" genus="Muhaka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="2599" phylum="Arthropoda" rank="species" species="icipe" status="sp. nov.">
<emphasis id="B930EAB9FF8CFFCF2017DE7EFDB1FADA" box="[370,523,1348,1372]" italics="true" pageId="2" pageNumber="2599">Muhaka icipe</emphasis>
, Buffington &amp; Copeland
</taxonomicName>
,
<taxonomicNameLabel id="A20357C2FF8CFFCF225DDE7FFC04FADA" box="[824,958,1348,1372]" pageId="2" pageNumber="2599" rank="species">new species</taxonomicNameLabel>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C35E6520FF8CFFCE21B1DEA7FC32FA9E" lastPageId="3" lastPageNumber="2600" pageId="2" pageNumber="2599" type="diagnosis">
<paragraph id="8BFB36ABFF8CFFCF21B1DEA7FEFAFA32" blockId="2.[212,1172,1437,1691]" box="[212,320,1437,1460]" pageId="2" pageNumber="2599">
<heading id="D0B381C7FF8CFFCF21B1DEA7FEFAFA32" box="[212,320,1437,1460]" fontSize="10" level="3" pageId="2" pageNumber="2599" reason="5">
<emphasis id="B930EAB9FF8CFFCF21B1DEA7FEFAFA32" box="[212,320,1437,1460]" italics="true" pageId="2" pageNumber="2599">Diagnosis</emphasis>
</heading>
</paragraph>
<paragraph id="8BFB36ABFF8CFFCE21B1DEFEFC32FA9E" blockId="2.[212,1172,1437,1691]" lastBlockId="3.[144,1104,1089,1304]" lastPageId="3" lastPageNumber="2600" pageId="2" pageNumber="2599">
Unique within Eucoilinae by the possession of a distinct valley-like depression on the vertex (ankos), encompassing the lateral ocelli, and whose anterior ridge directs the anterior ocellus in an anterior orientation; also unique within Eucoilinae is the possession of overhanging ridges over the toruli (kemnina). Superficially,
<taxonomicName id="4C444D28FF8CFFCF221CDD1EFC6FF9BA" authorityName="Buffington &amp; Copeland" authorityYear="2015" box="[889,981,1572,1596]" class="Insecta" family="Figitidae" genus="Muhaka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="2599" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAB9FF8CFFCF221CDD1EFC6FF9BA" box="[889,981,1572,1596]" italics="true" pageId="2" pageNumber="2599">Muhaka</emphasis>
</taxonomicName>
may be mistaken for
<taxonomicName id="4C444D28FF8CFFCF2199DD7EFEC3F9DA" authorityName="Quinlan" authorityYear="1984" box="[252,377,1604,1628]" class="Hexapoda" family="Figitidae" genus="Stentorceps" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="2599" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAB9FF8CFFCF2199DD7EFEC3F9DA" box="[252,377,1604,1628]" italics="true" pageId="2" pageNumber="2599">Stentorceps</emphasis>
</taxonomicName>
or
<taxonomicName id="4C444D28FF8CFFCF20C7DD7EFD96F9DA" authorityName="Buffington (Buffington" authorityYear="2012" box="[418,556,1604,1628]" class="Hexapoda" family="Figitidae" genus="Nanocthulhu" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="2599" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAB9FF8CFFCF20C7DD7EFD96F9DA" box="[418,556,1604,1628]" italics="true" pageId="2" pageNumber="2599">Nanocthulhu</emphasis>
</taxonomicName>
in that these three genera contain species with unusual head ornamentation, as well as relatively large, paddle-like mandibles. However, careful examination of the shape and position of these head characters readily separates
<taxonomicName id="4C444D28FF8CFFCF255DDDB9FB2EF91D" authorityName="Buffington &amp; Copeland" authorityYear="2015" box="[1080,1172,1667,1691]" class="Insecta" family="Figitidae" genus="Muhaka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="2" pageNumber="2599" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAB9FF8CFFCF255DDDB9FB2EF91D" box="[1080,1172,1667,1691]" italics="true" pageId="2" pageNumber="2599">Muhaka</emphasis>
</taxonomicName>
from the other two genera.
<taxonomicName id="4C444D28FF8DFFCE20D1DF7BFDAAFBDF" authorityName="Buffington &amp; Copeland" authorityYear="2015" box="[436,528,1089,1113]" class="Insecta" family="Figitidae" genus="Muhaka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="2600" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAB9FF8DFFCE20D1DF7BFDAAFBDF" box="[436,528,1089,1113]" italics="true" pageId="3" pageNumber="2600">Muhaka</emphasis>
</taxonomicName>
clearly belongs in the Kleidotomini, and its fore wing venation is characteristic of the tribe (
<figureCitation id="137F2A2EFF8DFFCE2349DF5BFD29FBFF" box="[556,659,1121,1145]" captionStart="Figure 1" captionStartId="2.[212,279,978,1000]" captionTargetBox="[212,1172,168,946]" captionTargetId="figure@2.[212,1173,168,946]" captionTargetPageId="2" captionText="Figure 1. Anterolateral habitus of Muhaka icipe Buffington and Copeland, n. gen., n. sp." figureDoi="http://doi.org/10.5281/zenodo.4000856" httpUri="https://zenodo.org/record/4000856/files/figure.png" pageId="3" pageNumber="2600">Figures 1</figureCitation>
and
<figureCitation id="137F2A2EFF8DFFCE23ACDF5BFD53FBFF" box="[713,745,1121,1145]" captionStart="Figure 3" captionStartId="4.[212,279,902,924]" captionTargetBox="[213,1172,168,870]" captionTargetId="figure@4.[212,1172,168,870]" captionTargetPageId="4" captionText="Figure 3. Habitus and habitat images of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (A) Dorsal habitus, with inset of scutellar and propodeal morphology; (B) lateral habitus (holotype specimen); (C) fore wing, bright-field illumination; (D) Muhaka Forest 6 m Malaise trap." figureDoi="http://doi.org/10.5281/zenodo.4000860" httpUri="https://zenodo.org/record/4000860/files/figure.png" pageId="3" pageNumber="2600">3C</figureCitation>
). Within Kleidotomini,
<taxonomicName id="4C444D28FF8DFFCE2291DF5BFBEAFBFF" authorityName="Buffington &amp; Copeland" authorityYear="2015" box="[1012,1104,1121,1145]" class="Insecta" family="Figitidae" genus="Muhaka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="2600" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAB9FF8DFFCE2291DF5BFBEAFBFF" box="[1012,1104,1121,1145]" italics="true" pageId="3" pageNumber="2600">Muhaka</emphasis>
</taxonomicName>
is most similar in appearance to
<taxonomicName id="4C444D28FF8DFFCE2366DFBBFDDCFB1F" authorityName="Kieffer" authorityYear="1901" box="[515,614,1153,1177]" class="Hexapoda" family="Figitidae" genus="Triplasta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="2600" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAB9FF8DFFCE2366DFBBFDDCFB1F" box="[515,614,1153,1177]" italics="true" pageId="3" pageNumber="2600">Triplasta</emphasis>
</taxonomicName>
species. Both taxa have weak striae on the lateral aspects of the pronotum and along the base of the syntergum of the metasoma. Additionally, in both genera the posterior margin of the metapleuron is distinct and the posterolateral face on the ventral corner of the metapleuron is glabrous. However, in
<taxonomicName id="4C444D28FF8DFFCE21F5DE3AFF49FA9E" authorityName="Kieffer" authorityYear="1901" box="[144,243,1280,1304]" class="Hexapoda" family="Figitidae" genus="Triplasta" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="2600" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAB9FF8DFFCE21F5DE3AFF49FA9E" box="[144,243,1280,1304]" italics="true" pageId="3" pageNumber="2600">Triplasta</emphasis>
</taxonomicName>
species, the metasomal base is glabrous (setose in
<taxonomicName id="4C444D28FF8DFFCE2279DE3AFCC2FA9E" authorityName="Buffington &amp; Copeland" authorityYear="2015" box="[796,888,1280,1304]" class="Insecta" family="Figitidae" genus="Muhaka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="2600" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAB9FF8DFFCE2279DE3AFCC2FA9E" box="[796,888,1280,1304]" italics="true" pageId="3" pageNumber="2600">Muhaka</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<caption id="DF3B6623FF8DFFCE21F5D8A3FBB6FB81" ID-DOI="http://doi.org/10.5281/zenodo.4000858" ID-Zenodo-Dep="4000858" httpUri="https://zenodo.org/record/4000858/files/figure.png" pageId="3" pageNumber="2600" startId="3.[144,211,921,943]" targetBox="[144,1104,168,889]" targetPageId="3">
<paragraph id="8BFB36ABFF8DFFCE21F5D8A3FBB6FB81" blockId="3.[144,1105,921,1031]" pageId="3" pageNumber="2600">
Figure 2. Scanning electron micrographs of
<taxonomicName id="4C444D28FF8DFFCE2335D8A3FC5BFC29" authority="Buffington and Copeland" authorityName="Buffington and Copeland" authorityYear="2015" box="[592,993,921,943]" class="Insecta" family="Figitidae" genus="Muhaka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="3" pageNumber="2600" phylum="Arthropoda" rank="species" species="icipe" status="n. gen., n. sp.">
<emphasis id="B930EAB9FF8DFFCE2335D8A3FD62FC28" box="[592,728,921,942]" italics="true" pageId="3" pageNumber="2600">Muhaka icipe</emphasis>
Buffington and Copeland
</taxonomicName>
,
<taxonomicNameLabel id="A20357C2FF8DFFCE2289D8A0FF17FC4A" pageId="3" pageNumber="2600" rank="species">n. gen., n. sp.</taxonomicNameLabel>
(holotype specimen). (A) Head and mesosoma, dorsal view; (B) head, anteroventral view; (C) head and mesosoma, dorsolateral view; (D) head and pronotum, anterolateral view. Abbreviations: ank, ankos; kem, kemnina; sct, bifurcate posterior margin of scutellum.
</paragraph>
</caption>
<subSubSection id="C35E6520FF8DFFC821F5DE67FCE0FB84" lastPageId="5" lastPageNumber="2602" pageId="3" pageNumber="2600" type="description">
<paragraph id="8BFB36ABFF8DFFCE21F5DE67FEB5FAF2" blockId="3.[144,1104,1373,1691]" box="[144,271,1373,1396]" pageId="3" pageNumber="2600">
<heading id="D0B381C7FF8DFFCE21F5DE67FEB5FAF2" box="[144,271,1373,1396]" fontSize="10" level="3" pageId="3" pageNumber="2600" reason="5">
<emphasis id="B930EAB9FF8DFFCE21F5DE67FEB5FAF2" box="[144,271,1373,1396]" italics="true" pageId="3" pageNumber="2600">Description</emphasis>
</heading>
</paragraph>
<paragraph id="8BFB36ABFF8DFFC921F5DEBEFC4EFBF9" blockId="3.[144,1104,1373,1691]" lastBlockId="4.[212,1171,1095,1151]" lastPageId="4" lastPageNumber="2601" pageId="3" pageNumber="2600">
<emphasis id="B930EAB9FF8DFFCE21F5DEBEFF6EFA1A" box="[144,212,1412,1436]" italics="true" pageId="3" pageNumber="2600">Head.</emphasis>
Malar sulcus simple, sinuate, converging towards anterior margin of clypeus, deeply impressed in ventral half immediately posterior to clypeus (
<figureCitation id="137F2A2EFF8DFFCE2232DE9EFC72FA3A" box="[855,968,1444,1468]" captionStart="Figure 2" captionStartId="3.[144,211,921,943]" captionTargetBox="[144,1104,168,889]" captionTargetId="figure@3.[144,1104,168,889]" captionTargetPageId="3" captionText="Figure 2. Scanning electron micrographs of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (holotype specimen). (A) Head and mesosoma, dorsal view; (B) head, anteroventral view; (C) head and mesosoma, dorsolateral view; (D) head and pronotum, anterolateral view. Abbreviations: ank, ankos; kem, kemnina; sct, bifurcate posterior margin of scutellum." figureDoi="http://doi.org/10.5281/zenodo.4000858" httpUri="https://zenodo.org/record/4000858/files/figure.png" pageId="3" pageNumber="2600">Figure 2D</figureCitation>
); anterior of clypeus pointed, protruding above base of mandibles (
<figureCitation id="137F2A2EFF8DFFCE23BDDEFEFCF3FA5A" box="[728,841,1476,1500]" captionStart="Figure 2" captionStartId="3.[144,211,921,943]" captionTargetBox="[144,1104,168,889]" captionTargetId="figure@3.[144,1104,168,889]" captionTargetPageId="3" captionText="Figure 2. Scanning electron micrographs of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (holotype specimen). (A) Head and mesosoma, dorsal view; (B) head, anteroventral view; (C) head and mesosoma, dorsolateral view; (D) head and pronotum, anterolateral view. Abbreviations: ank, ankos; kem, kemnina; sct, bifurcate posterior margin of scutellum." figureDoi="http://doi.org/10.5281/zenodo.4000858" httpUri="https://zenodo.org/record/4000858/files/figure.png" pageId="3" pageNumber="2600">Figure 2D</figureCitation>
); resulting area between malar sulci keel-like (
<figureCitation id="137F2A2EFF8DFFCE2013DEDEFE5DFA7A" box="[374,487,1508,1532]" captionStart="Figure 2" captionStartId="3.[144,211,921,943]" captionTargetBox="[144,1104,168,889]" captionTargetId="figure@3.[144,1104,168,889]" captionTargetPageId="3" captionText="Figure 2. Scanning electron micrographs of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (holotype specimen). (A) Head and mesosoma, dorsal view; (B) head, anteroventral view; (C) head and mesosoma, dorsolateral view; (D) head and pronotum, anterolateral view. Abbreviations: ank, ankos; kem, kemnina; sct, bifurcate posterior margin of scutellum." figureDoi="http://doi.org/10.5281/zenodo.4000858" httpUri="https://zenodo.org/record/4000858/files/figure.png" pageId="3" pageNumber="2600">Figure 2D</figureCitation>
). Orbital furrows absent. Malar space smooth. Kemnina present posterior to toruli, distinctly overhanging toruli when observed dorsally (
<figureCitation id="137F2A2EFF8DFFCE21FDDD19FE93F9BD" box="[152,297,1571,1595]" captionStart="Figure 2" captionStartId="3.[144,211,921,943]" captionTargetBox="[144,1104,168,889]" captionTargetId="figure@3.[144,1104,168,889]" captionTargetPageId="3" captionText="Figure 2. Scanning electron micrographs of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (holotype specimen). (A) Head and mesosoma, dorsal view; (B) head, anteroventral view; (C) head and mesosoma, dorsolateral view; (D) head and pronotum, anterolateral view. Abbreviations: ank, ankos; kem, kemnina; sct, bifurcate posterior margin of scutellum." figureDoi="http://doi.org/10.5281/zenodo.4000858" httpUri="https://zenodo.org/record/4000858/files/figure.png" pageId="3" pageNumber="2600">Figure 2BD</figureCitation>
). Ankos present posterior to base of kemnina, resulting in anterior ocellus facing anteriorly (versus dorsally in other eucoilines); lateral ocelli nested within depression of ankos, with short scattered setae present (
<figureCitation id="137F2A2EFF8DFFCE23C6DD59FCABF9FD" box="[675,785,1635,1659]" captionStart="Figure 2" captionStartId="3.[144,211,921,943]" captionTargetBox="[144,1104,168,889]" captionTargetId="figure@3.[144,1104,168,889]" captionTargetPageId="3" captionText="Figure 2. Scanning electron micrographs of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (holotype specimen). (A) Head and mesosoma, dorsal view; (B) head, anteroventral view; (C) head and mesosoma, dorsolateral view; (D) head and pronotum, anterolateral view. Abbreviations: ank, ankos; kem, kemnina; sct, bifurcate posterior margin of scutellum." figureDoi="http://doi.org/10.5281/zenodo.4000858" httpUri="https://zenodo.org/record/4000858/files/figure.png" pageId="3" pageNumber="2600">Figure 2C</figureCitation>
). Mandibles extremely large, paddle-shaped, roughly 1/3 total length of head (
<figureCitation id="137F2A2EFF8DFFCE23D2DDB9FCA5F91D" box="[695,799,1667,1691]" captionStart="Figure 1" captionStartId="2.[212,279,978,1000]" captionTargetBox="[212,1172,168,946]" captionTargetId="figure@2.[212,1173,168,946]" captionTargetPageId="2" captionText="Figure 1. Anterolateral habitus of Muhaka icipe Buffington and Copeland, n. gen., n. sp." figureDoi="http://doi.org/10.5281/zenodo.4000856" httpUri="https://zenodo.org/record/4000856/files/figure.png" pageId="3" pageNumber="2600">Figures 1</figureCitation>
and
<figureCitation id="137F2A2EFF8DFFCE223EDDB9FC1AF91D" box="[859,928,1667,1691]" captionStart="Figure 2" captionStartId="3.[144,211,921,943]" captionTargetBox="[144,1104,168,889]" captionTargetId="figure@3.[144,1104,168,889]" captionTargetPageId="3" captionText="Figure 2. Scanning electron micrographs of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (holotype specimen). (A) Head and mesosoma, dorsal view; (B) head, anteroventral view; (C) head and mesosoma, dorsolateral view; (D) head and pronotum, anterolateral view. Abbreviations: ank, ankos; kem, kemnina; sct, bifurcate posterior margin of scutellum." figureDoi="http://doi.org/10.5281/zenodo.4000858" httpUri="https://zenodo.org/record/4000858/files/figure.png" pageId="3" pageNumber="2600">2B, D</figureCitation>
), sub-quadrate, spatulate in anterior view (
<figureCitation id="137F2A2EFF8AFFC9209EDF7DFDD4FBD9" box="[507,622,1095,1119]" captionStart="Figure 2" captionStartId="3.[144,211,921,943]" captionTargetBox="[144,1104,168,889]" captionTargetId="figure@3.[144,1104,168,889]" captionTargetPageId="3" captionText="Figure 2. Scanning electron micrographs of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (holotype specimen). (A) Head and mesosoma, dorsal view; (B) head, anteroventral view; (C) head and mesosoma, dorsolateral view; (D) head and pronotum, anterolateral view. Abbreviations: ank, ankos; kem, kemnina; sct, bifurcate posterior margin of scutellum." figureDoi="http://doi.org/10.5281/zenodo.4000858" httpUri="https://zenodo.org/record/4000858/files/figure.png" pageId="4" pageNumber="2601">Figure 2D</figureCitation>
); basal mandibular impression present, indicating mandible articulates in longitudinal plane; basal mandibular keel absent.
</paragraph>
<caption id="DF3B6623FF8AFFC921B1D8BCFEE6FC75" ID-DOI="http://doi.org/10.5281/zenodo.4000860" ID-Zenodo-Dep="4000860" httpUri="https://zenodo.org/record/4000860/files/figure.png" pageId="4" pageNumber="2601" startId="4.[212,279,902,924]" targetBox="[213,1172,168,870]" targetPageId="4">
<paragraph id="8BFB36ABFF8AFFC921B1D8BCFEE6FC75" blockId="4.[212,1172,902,1011]" pageId="4" pageNumber="2601">
Figure 3. Habitus and habitat images of
<taxonomicName id="4C444D28FF8AFFC92317D8BCFBB9FC1A" authority="Buffington and Copeland" authorityName="Buffington and Copeland" authorityYear="2015" box="[626,1027,902,924]" class="Insecta" family="Figitidae" genus="Muhaka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="2601" phylum="Arthropoda" rank="species" species="icipe" status="n. gen., n. sp.">
<emphasis id="B930EAB9FF8AFFC92317D8BCFD40FC1D" box="[626,762,902,923]" italics="true" pageId="4" pageNumber="2601">Muhaka icipe</emphasis>
Buffington and Copeland
</taxonomicName>
,
<taxonomicNameLabel id="A20357C2FF8AFFC9256BD8BCFB2EFC1D" box="[1038,1172,902,923]" pageId="4" pageNumber="2601" rank="species">n. gen., n. sp.</taxonomicNameLabel>
(A) Dorsal habitus, with inset of scutellar and propodeal morphology; (B) lateral habitus (holotype specimen); (C) fore wing, bright-field illumination; (D)
<taxonomicName id="4C444D28FF8AFFC922CAD8FAFBEFFC50" authority="Forest" authorityName="Forest" box="[943,1109,960,982]" class="Insecta" family="Figitidae" genus="Muhaka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="2601" phylum="Arthropoda" rank="genus">Muhaka Forest</taxonomicName>
6 m Malaise trap.
</paragraph>
</caption>
<paragraph id="8BFB36ABFF8AFFC921B1DF86FE1BFAB5" blockId="4.[212,1172,1212,1331]" pageId="4" pageNumber="2601">
<emphasis id="B930EAB9FF8AFFC921B1DF86FEFEFB52" box="[212,324,1212,1236]" italics="true" pageId="4" pageNumber="2601">Antennae.</emphasis>
Male: 13 flagellomeres, sub equal in length; multiporous plate sensilla on all flagellomeres; single campaniform sensillum present on distal margin of flagellomeres 413; flagellomere 1 distinctly modified, slightly elongate, excavated laterally. Female unknown.
</paragraph>
<paragraph id="8BFB36ABFF8AFFC921B1DE48FC94F98E" blockId="4.[212,1171,1392,1544]" pageId="4" pageNumber="2601">
<emphasis id="B930EAB9FF8AFFC921B1DE48FEF0FA0F" box="[212,330,1394,1417]" italics="true" pageId="4" pageNumber="2601">Pronotum.</emphasis>
Pronotal plate narrow, with setae present along anterior aspect; dorsal margin rounded; pronotal fovea open laterally; ventral half of pronotal plate extended anteriorly (
<figureCitation id="137F2A2EFF8AFFC920A5DE8AFD8EFA4E" box="[448,564,1456,1480]" captionStart="Figure 2" captionStartId="3.[144,211,921,943]" captionTargetBox="[144,1104,168,889]" captionTargetId="figure@3.[144,1104,168,889]" captionTargetPageId="3" captionText="Figure 2. Scanning electron micrographs of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (holotype specimen). (A) Head and mesosoma, dorsal view; (B) head, anteroventral view; (C) head and mesosoma, dorsolateral view; (D) head and pronotum, anterolateral view. Abbreviations: ank, ankos; kem, kemnina; sct, bifurcate posterior margin of scutellum." figureDoi="http://doi.org/10.5281/zenodo.4000858" httpUri="https://zenodo.org/record/4000858/files/figure.png" pageId="4" pageNumber="2601">Figure 2C</figureCitation>
). Pronotal trough present, ventrad of pronotal plate, deep with broad, confluent setae (
<figureCitation id="137F2A2EFF8AFFC92317DEEAFD51FA6E" box="[626,747,1488,1512]" captionStart="Figure 2" captionStartId="3.[144,211,921,943]" captionTargetBox="[144,1104,168,889]" captionTargetId="figure@3.[144,1104,168,889]" captionTargetPageId="3" captionText="Figure 2. Scanning electron micrographs of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (holotype specimen). (A) Head and mesosoma, dorsal view; (B) head, anteroventral view; (C) head and mesosoma, dorsolateral view; (D) head and pronotum, anterolateral view. Abbreviations: ank, ankos; kem, kemnina; sct, bifurcate posterior margin of scutellum." figureDoi="http://doi.org/10.5281/zenodo.4000858" httpUri="https://zenodo.org/record/4000858/files/figure.png" pageId="4" pageNumber="2601">Figure 2C</figureCitation>
). Lateral pronotal carina, pronotal triangle and pronotal impression absent (
<figureCitation id="137F2A2EFF8AFFC923CFDECAFCA4F98E" box="[682,798,1520,1544]" captionStart="Figure 2" captionStartId="3.[144,211,921,943]" captionTargetBox="[144,1104,168,889]" captionTargetId="figure@3.[144,1104,168,889]" captionTargetPageId="3" captionText="Figure 2. Scanning electron micrographs of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (holotype specimen). (A) Head and mesosoma, dorsal view; (B) head, anteroventral view; (C) head and mesosoma, dorsolateral view; (D) head and pronotum, anterolateral view. Abbreviations: ank, ankos; kem, kemnina; sct, bifurcate posterior margin of scutellum." figureDoi="http://doi.org/10.5281/zenodo.4000858" httpUri="https://zenodo.org/record/4000858/files/figure.png" pageId="4" pageNumber="2601">Figure 2C</figureCitation>
).
</paragraph>
<paragraph id="8BFB36ABFF8AFFC921B1DD7CFBAFF9FB" blockId="4.[212,1171,1605,1661]" pageId="4" pageNumber="2601">
<emphasis id="B930EAB9FF8AFFC921B1DD7CFEDCF9DB" box="[212,358,1606,1629]" italics="true" pageId="4" pageNumber="2601">Mesoscutum.</emphasis>
Parascutal impression present, incomplete (2C). Notauli, mesoscutal keel, parapsidal ridges, parapsidal hair-lines absent (
<figureCitation id="137F2A2EFF8AFFC9224FDD5FFC12F9FB" box="[810,936,1637,1661]" captionStart="Figure 2" captionStartId="3.[144,211,921,943]" captionTargetBox="[144,1104,168,889]" captionTargetId="figure@3.[144,1104,168,889]" captionTargetPageId="3" captionText="Figure 2. Scanning electron micrographs of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (holotype specimen). (A) Head and mesosoma, dorsal view; (B) head, anteroventral view; (C) head and mesosoma, dorsolateral view; (D) head and pronotum, anterolateral view. Abbreviations: ank, ankos; kem, kemnina; sct, bifurcate posterior margin of scutellum." figureDoi="http://doi.org/10.5281/zenodo.4000858" httpUri="https://zenodo.org/record/4000858/files/figure.png" pageId="4" pageNumber="2601">Figures 2C</figureCitation>
and
<figureCitation id="137F2A2EFF8AFFC92286DD5FFBBCF9FB" box="[995,1030,1637,1661]" captionStart="Figure 3" captionStartId="4.[212,279,902,924]" captionTargetBox="[213,1172,168,870]" captionTargetId="figure@4.[212,1172,168,870]" captionTargetPageId="4" captionText="Figure 3. Habitus and habitat images of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (A) Dorsal habitus, with inset of scutellar and propodeal morphology; (B) lateral habitus (holotype specimen); (C) fore wing, bright-field illumination; (D) Muhaka Forest 6 m Malaise trap." figureDoi="http://doi.org/10.5281/zenodo.4000860" httpUri="https://zenodo.org/record/4000860/files/figure.png" pageId="4" pageNumber="2601">3A</figureCitation>
).
</paragraph>
<paragraph id="8BFB36ABFF8BFFC821F5DB93FEE9FE86" blockId="5.[144,1104,168,256]" pageId="5" pageNumber="2602">
<emphasis id="B930EAB9FF8BFFC821F5DB93FEA3FF46" box="[144,281,169,192]" italics="true" pageId="5" pageNumber="2602">Mesopectus.</emphasis>
Mesopleural carina simple, distinctly raised (
<figureCitation id="137F2A2EFF8BFFC8227DDB92FC30FF46" box="[792,906,168,192]" captionStart="Figure 2" captionStartId="3.[144,211,921,943]" captionTargetBox="[144,1104,168,889]" captionTargetId="figure@3.[144,1104,168,889]" captionTargetPageId="3" captionText="Figure 2. Scanning electron micrographs of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (holotype specimen). (A) Head and mesosoma, dorsal view; (B) head, anteroventral view; (C) head and mesosoma, dorsolateral view; (D) head and pronotum, anterolateral view. Abbreviations: ank, ankos; kem, kemnina; sct, bifurcate posterior margin of scutellum." figureDoi="http://doi.org/10.5281/zenodo.4000858" httpUri="https://zenodo.org/record/4000858/files/figure.png" pageId="5" pageNumber="2602">Figure 2C</figureCitation>
). Precoxal carina of lower part of mesopleuron present, complete (
<figureCitation id="137F2A2EFF8BFFC823B2DBF2FCF5FF66" box="[727,847,200,224]" captionStart="Figure 2" captionStartId="3.[144,211,921,943]" captionTargetBox="[144,1104,168,889]" captionTargetId="figure@3.[144,1104,168,889]" captionTargetPageId="3" captionText="Figure 2. Scanning electron micrographs of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (holotype specimen). (A) Head and mesosoma, dorsal view; (B) head, anteroventral view; (C) head and mesosoma, dorsolateral view; (D) head and pronotum, anterolateral view. Abbreviations: ank, ankos; kem, kemnina; sct, bifurcate posterior margin of scutellum." figureDoi="http://doi.org/10.5281/zenodo.4000858" httpUri="https://zenodo.org/record/4000858/files/figure.png" pageId="5" pageNumber="2602">Figure 2C</figureCitation>
). Surcoxal depression reduced, smooth.
</paragraph>
<paragraph id="8BFB36ABFF8BFFC821F5DA13FC5FFEE6" blockId="5.[144,1104,297,352]" pageId="5" pageNumber="2602">
<emphasis id="B930EAB9FF8BFFC821F5DA13FEBFFEC7" box="[144,261,297,321]" italics="true" pageId="5" pageNumber="2602">Scutellum.</emphasis>
Scutellar plate small, narrow; glandular release pit positioned posteriorly. Dorsal surface of the scutellum longitudinally striate (
<figureCitation id="137F2A2EFF8BFFC8239CDA72FCC2FEE6" box="[761,888,328,352]" captionStart="Figure 2" captionStartId="3.[144,211,921,943]" captionTargetBox="[144,1104,168,889]" captionTargetId="figure@3.[144,1104,168,889]" captionTargetPageId="3" captionText="Figure 2. Scanning electron micrographs of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (holotype specimen). (A) Head and mesosoma, dorsal view; (B) head, anteroventral view; (C) head and mesosoma, dorsolateral view; (D) head and pronotum, anterolateral view. Abbreviations: ank, ankos; kem, kemnina; sct, bifurcate posterior margin of scutellum." figureDoi="http://doi.org/10.5281/zenodo.4000858" httpUri="https://zenodo.org/record/4000858/files/figure.png" pageId="5" pageNumber="2602">Figures 2A</figureCitation>
and
<figureCitation id="137F2A2EFF8BFFC822D6DA72FC6FFEE6" box="[947,981,328,352]" captionStart="Figure 3" captionStartId="4.[212,279,902,924]" captionTargetBox="[213,1172,168,870]" captionTargetId="figure@4.[212,1172,168,870]" captionTargetPageId="4" captionText="Figure 3. Habitus and habitat images of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (A) Dorsal habitus, with inset of scutellar and propodeal morphology; (B) lateral habitus (holotype specimen); (C) fore wing, bright-field illumination; (D) Muhaka Forest 6 m Malaise trap." figureDoi="http://doi.org/10.5281/zenodo.4000860" httpUri="https://zenodo.org/record/4000860/files/figure.png" pageId="5" pageNumber="2602">3A</figureCitation>
).
</paragraph>
<paragraph id="8BFB36ABFF8BFFC821F5DAB3FCBAFDC7" blockId="5.[144,1104,393,577]" pageId="5" pageNumber="2602">
<emphasis id="B930EAB9FF8BFFC821F5DAB3FE43FE27" box="[144,505,393,417]" italics="true" pageId="5" pageNumber="2602">Metapectalpropodeal complex.</emphasis>
Spiracular groove poorly defined, ventral margin absent (
<figureCitation id="137F2A2EFF8BFFC82197DA93FED4FE47" box="[242,366,425,449]" captionStart="Figure 2" captionStartId="3.[144,211,921,943]" captionTargetBox="[144,1104,168,889]" captionTargetId="figure@3.[144,1104,168,889]" captionTargetPageId="3" captionText="Figure 2. Scanning electron micrographs of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (holotype specimen). (A) Head and mesosoma, dorsal view; (B) head, anteroventral view; (C) head and mesosoma, dorsolateral view; (D) head and pronotum, anterolateral view. Abbreviations: ank, ankos; kem, kemnina; sct, bifurcate posterior margin of scutellum." figureDoi="http://doi.org/10.5281/zenodo.4000858" httpUri="https://zenodo.org/record/4000858/files/figure.png" pageId="5" pageNumber="2602">Figure 2C</figureCitation>
). Posterior margin of metapectus gently sculptured, ridged. Metapleural ridge, submetapleural ridge absent; cavity absent along posterior margin of metapleuron, ventral to submetapleural ridge; posteroventral margin slightly drawn out, glabrous, with distinctly flat posterior aspect (
<figureCitation id="137F2A2EFF8BFFC82233D933FC6AFDA7" box="[854,976,521,545]" captionStart="Figure 2" captionStartId="3.[144,211,921,943]" captionTargetBox="[144,1104,168,889]" captionTargetId="figure@3.[144,1104,168,889]" captionTargetPageId="3" captionText="Figure 2. Scanning electron micrographs of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (holotype specimen). (A) Head and mesosoma, dorsal view; (B) head, anteroventral view; (C) head and mesosoma, dorsolateral view; (D) head and pronotum, anterolateral view. Abbreviations: ank, ankos; kem, kemnina; sct, bifurcate posterior margin of scutellum." figureDoi="http://doi.org/10.5281/zenodo.4000858" httpUri="https://zenodo.org/record/4000858/files/figure.png" pageId="5" pageNumber="2602">Figure 2C</figureCitation>
). Anterior impression of metepimeron and metepisternum absent.
</paragraph>
<paragraph id="8BFB36ABFF8BFFC821F5D950FC26FD27" blockId="5.[144,1104,617,673]" pageId="5" pageNumber="2602">
<emphasis id="B930EAB9FF8BFFC821F5D950FF67FD07" box="[144,221,618,641]" italics="true" pageId="5" pageNumber="2602">Wings.</emphasis>
Hyaline; setose (
<figureCitation id="137F2A2EFF8BFFC820C6D953FDB7FD07" box="[419,525,617,641]" captionStart="Figure 1" captionStartId="2.[212,279,978,1000]" captionTargetBox="[212,1172,168,946]" captionTargetId="figure@2.[212,1173,168,946]" captionTargetPageId="2" captionText="Figure 1. Anterolateral habitus of Muhaka icipe Buffington and Copeland, n. gen., n. sp." figureDoi="http://doi.org/10.5281/zenodo.4000856" httpUri="https://zenodo.org/record/4000856/files/figure.png" pageId="5" pageNumber="2602">Figures 1</figureCitation>
and
<figureCitation id="137F2A2EFF8BFFC8232FD953FD35FD07" box="[586,655,617,641]" captionStart="Figure 3" captionStartId="4.[212,279,902,924]" captionTargetBox="[213,1172,168,870]" captionTargetId="figure@4.[212,1172,168,870]" captionTargetPageId="4" captionText="Figure 3. Habitus and habitat images of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (A) Dorsal habitus, with inset of scutellar and propodeal morphology; (B) lateral habitus (holotype specimen); (C) fore wing, bright-field illumination; (D) Muhaka Forest 6 m Malaise trap." figureDoi="http://doi.org/10.5281/zenodo.4000860" httpUri="https://zenodo.org/record/4000860/files/figure.png" pageId="5" pageNumber="2602">3A, C</figureCitation>
). Apical margin complete (not emarginate). Overall wing shape pernaform (
<bibRefCitation id="EFD54B5AFF8BFFC82327D9B3FC36FD27" author="Buffington ML &amp; Sandler R" box="[578,908,649,673]" pageId="5" pageNumber="2602" pagination="586 - 597" refId="ref4192" refString="Buffington ML, Sandler R. 2012. The occurrence and phylogenetic implications of wing interference patterns in Cynipoidea (Insecta: Hymenoptera). Invertebr Syst. 6: 586 - 597." type="journal article" year="2012">Buffington and Sandler 2012</bibRefCitation>
).
</paragraph>
<paragraph id="8BFB36ABFF8BFFC821F5D9F1FEF6FCE7" blockId="5.[144,1104,714,865]" pageId="5" pageNumber="2602">
<emphasis id="B930EAB9FF8BFFC821F5D9F1FF76FD64" box="[144,204,715,738]" italics="true" pageId="5" pageNumber="2602">Legs.</emphasis>
Fore- and mid-coxae sub-equal in size, hind coxa twice as long as other coxae; all coxae glabrous; metacoxa without posterior dorsoventral hair line. Femora with sparse setal lines; tibiae and tarsomeres with dense, adpressed setae. Length of metatarsomere 1 slightly less than combined length of remaining metatarsomeres.
</paragraph>
<paragraph id="8BFB36ABFF8BFFC821F5D8B1FCE0FB84" blockId="5.[144,1104,906,1027]" pageId="5" pageNumber="2602">
<emphasis id="B930EAB9FF8BFFC821F5D8B1FEB6FC24" box="[144,268,907,930]" italics="true" pageId="5" pageNumber="2602">Metasoma.</emphasis>
Metasoma subequal in size to head and mesosoma (
<figureCitation id="137F2A2EFF8BFFC82208D8B0FC58FC24" box="[877,994,906,930]" captionStart="Figure 3" captionStartId="4.[212,279,902,924]" captionTargetBox="[213,1172,168,870]" captionTargetId="figure@4.[212,1172,168,870]" captionTargetPageId="4" captionText="Figure 3. Habitus and habitat images of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (A) Dorsal habitus, with inset of scutellar and propodeal morphology; (B) lateral habitus (holotype specimen); (C) fore wing, bright-field illumination; (D) Muhaka Forest 6 m Malaise trap." figureDoi="http://doi.org/10.5281/zenodo.4000860" httpUri="https://zenodo.org/record/4000860/files/figure.png" pageId="5" pageNumber="2602">Figure 3A</figureCitation>
). Base of syntergum with hairy ring, comprised of dense adpressed setae, incomplete dorsally (
<figureCitation id="137F2A2EFF8BFFC821FDD8F0FEACFC64" box="[152,278,970,994]" captionStart="Figure 2" captionStartId="3.[144,211,921,943]" captionTargetBox="[144,1104,168,889]" captionTargetId="figure@3.[144,1104,168,889]" captionTargetPageId="3" captionText="Figure 2. Scanning electron micrographs of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (holotype specimen). (A) Head and mesosoma, dorsal view; (B) head, anteroventral view; (C) head and mesosoma, dorsolateral view; (D) head and pronotum, anterolateral view. Abbreviations: ank, ankos; kem, kemnina; sct, bifurcate posterior margin of scutellum." figureDoi="http://doi.org/10.5281/zenodo.4000858" httpUri="https://zenodo.org/record/4000858/files/figure.png" pageId="5" pageNumber="2602">Figures 2C</figureCitation>
and
<figureCitation id="137F2A2EFF8BFFC82034D8F0FEC9FC64" box="[337,371,970,994]" captionStart="Figure 3" captionStartId="4.[212,279,902,924]" captionTargetBox="[213,1172,168,870]" captionTargetId="figure@4.[212,1172,168,870]" captionTargetPageId="4" captionText="Figure 3. Habitus and habitat images of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (A) Dorsal habitus, with inset of scutellar and propodeal morphology; (B) lateral habitus (holotype specimen); (C) fore wing, bright-field illumination; (D) Muhaka Forest 6 m Malaise trap." figureDoi="http://doi.org/10.5281/zenodo.4000860" httpUri="https://zenodo.org/record/4000860/files/figure.png" pageId="5" pageNumber="2602">3A</figureCitation>
); remainder of metasoma glabrous (
<figureCitation id="137F2A2EFF8BFFC82276D8F0FC32FC64" box="[787,904,970,994]" captionStart="Figure 3" captionStartId="4.[212,279,902,924]" captionTargetBox="[213,1172,168,870]" captionTargetId="figure@4.[212,1172,168,870]" captionTargetPageId="4" captionText="Figure 3. Habitus and habitat images of Muhaka icipe Buffington and Copeland, n. gen., n. sp. (A) Dorsal habitus, with inset of scutellar and propodeal morphology; (B) lateral habitus (holotype specimen); (C) fore wing, bright-field illumination; (D) Muhaka Forest 6 m Malaise trap." figureDoi="http://doi.org/10.5281/zenodo.4000860" httpUri="https://zenodo.org/record/4000860/files/figure.png" pageId="5" pageNumber="2602">Figure 3A</figureCitation>
). Terga posterior to syntergum gradually directed posteriorly. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C35E6520FF8BFFC821F5DF79FE62FB05" box="[144,472,1091,1155]" pageId="5" pageNumber="2602" type="distribution">
<paragraph id="8BFB36ABFF8BFFC821F5DF79FEAEFBDD" blockId="5.[144,472,1091,1155]" box="[144,276,1091,1115]" pageId="5" pageNumber="2602">
<heading id="D0B381C7FF8BFFC821F5DF79FEAEFBDD" box="[144,276,1091,1115]" fontSize="10" level="3" pageId="5" pageNumber="2602" reason="5">
<emphasis id="B930EAB9FF8BFFC821F5DF79FEAEFBDD" box="[144,276,1091,1115]" italics="true" pageId="5" pageNumber="2602">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8BFB36ABFF8BFFC821F5DF50FE62FB05" blockId="5.[144,472,1091,1155]" box="[144,472,1130,1155]" pageId="5" pageNumber="2602">
Afrotropical Region:
<collectingCountry id="F353763BFF8BFFC820E3DF51FE6EFB05" box="[390,468,1131,1155]" name="Kenya" pageId="5" pageNumber="2602">Kenya</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C35E6520FF8BFFC821F5DFF9FE96FAA5" pageId="5" pageNumber="2602" type="etymology">
<paragraph id="8BFB36ABFF8BFFC821F5DFF9FEB3FB5D" blockId="5.[144,1103,1219,1315]" box="[144,265,1219,1243]" pageId="5" pageNumber="2602">
<heading id="D0B381C7FF8BFFC821F5DFF9FEB3FB5D" box="[144,265,1219,1243]" fontSize="10" level="3" pageId="5" pageNumber="2602" reason="5">
<emphasis id="B930EAB9FF8BFFC821F5DFF9FEB3FB5D" box="[144,265,1219,1243]" italics="true" pageId="5" pageNumber="2602">Etymology</emphasis>
</heading>
</paragraph>
<paragraph id="8BFB36ABFF8BFFC821F5DFD1FE96FAA5" blockId="5.[144,1103,1219,1315]" pageId="5" pageNumber="2602">
Genus named in honour of
<taxonomicName id="4C444D28FF8BFFC820ADDFD1FDCBFA85" authority="forest" authorityName="Forest" box="[456,625,1259,1283]" class="Insecta" family="Figitidae" genus="Muhaka" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="2605" phylum="Arthropoda" rank="genus">Muhaka forest</taxonomicName>
, the type-locality of the genus; it is a noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>