treatments-xml/data/03/ED/87/03ED879CFF88FF9717AEFD09FCEEFDB7.xml

<document id="395992A51F034E25BA41721A20106989" ID-DOI="10.11646/zootaxa.3937.2.6" ID-GBIF-Dataset="1c07ebfb-47ef-4aa4-aa8b-652fa505657c" ID-ISSN="1175-5326" ID-Zenodo-Dep="243845" ID-ZooBank="E654E1ED-708D-4537-A7EC-8FFF8EC2DCF4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461021025898" checkinUser="plazi" docAuthor="Soares, Karla D. A., Gadig, Otto F. B. &amp; Gomes, Ulisses L." docDate="2015" docId="03ED879CFF88FF9717AEFD09FCEEFDB7" docLanguage="en" docName="zt03937p361.pdf" docOrigin="Zootaxa 3937 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scyliorhinus ugoi Soares, Gadig &amp; Gomes, 2015, sp. nov." docType="treatment" docVersion="8" lastPageNumber="359" masterDocId="FFD4FFE4FF89FF9B1739FFDBFFC8FF8A" masterDocTitle="Scyliorhinus ugoi, a new species of catshark from Brazil (Chondrichthyes: Carcharhiniformes: Scyliorhinidae)" masterLastPageNumber="361" masterPageNumber="347" pageNumber="348" updateTime="1698570162864" updateUser="plazi">
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<mods:title id="2D580B2E480856BAE5B7225CA7E718A8">Scyliorhinus ugoi, a new species of catshark from Brazil (Chondrichthyes: Carcharhiniformes: Scyliorhinidae)</mods:title>
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<mods:namePart id="32CFEF0E05E255276DF2DF6DE7FCE430">Soares, Karla D. A.</mods:namePart>
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<mods:namePart id="A19F289C83F6DFF7D5747A336249AB29">Gadig, Otto F. B.</mods:namePart>
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<mods:namePart id="2F5ECD7736FA6C821AA8238C84F1AC5A">Gomes, Ulisses L.</mods:namePart>
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<paragraph id="8BFB368AFF88FF9A17AEFD09FE14FD66" blockId="1.[151,476,722,782]" box="[151,476,722,748]" pageId="1" pageNumber="348">
<heading id="D0B381E6FF88FF9A17AEFD09FE14FD66" bold="true" box="[151,476,722,748]" fontSize="11" level="1" pageId="1" pageNumber="348" reason="1">
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<taxonomicName id="4C444D09FF88FF9A17AEFD09FEA4FD66" ID-CoL="4W57X" box="[151,364,722,748]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="ugoi" status="sp. nov.">
<emphasis id="B930EA98FF88FF9A17AEFD09FEA4FD66" bold="true" box="[151,364,722,748]" italics="true" pageId="1" pageNumber="348">Scyliorhinus ugoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20357E3FF88FF9A164AFD09FE14FD66" box="[371,476,722,748]" pageId="1" pageNumber="348" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="8BFB368AFF88FF9A17AEFD2EFEDAFC84" blockId="1.[151,476,722,782]" box="[151,274,757,782]" pageId="1" pageNumber="348">
(
<figureCitation id="137F2A0FFF88FF9A17A6FD2EFEC1FC84" box="[159,265,757,782]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStart-4="FIGURE 5" captionStart-5="FIGURE 6" captionStart-6="FIGURE 7" captionStart-7="FIGURE 8" captionStart-8="FIGURE 9" captionStartId-0="4.[151,249,1130,1152]" captionStartId-1="6.[151,260,1897,1919]" captionStartId-2="7.[151,249,648,670]" captionStartId-3="7.[151,249,1454,1476]" captionStartId-4="8.[151,249,1047,1069]" captionStartId-5="8.[151,249,1888,1910]" captionStartId-6="9.[151,249,988,1010]" captionStartId-7="10.[151,249,1319,1341]" captionStartId-8="11.[151,249,987,1009]" captionTargetBox-0="[371,1214,204,1099]" captionTargetBox-1="[346,1240,665,1868]" captionTargetBox-2="[436,1163,193,607]" captionTargetBox-3="[495,1143,740,1431]" captionTargetBox-4="[341,1245,447,1021]" captionTargetBox-5="[477,1087,1188,1844]" captionTargetBox-6="[356,1216,199,957]" captionTargetBox-7="[356,1227,308,1298]" captionTargetBox-8="[340,1247,193,965]" captionTargetId-0="figure@4.[362,1226,195,1105]" captionTargetId-1="figure@6.[340,1248,661,1875]" captionTargetId-2="figure@7.[431,1163,193,621]" captionTargetId-3="figure@7.[433,1159,720,1432]" captionTargetId-4="figure@8.[340,1248,445,1025]" captionTargetId-5="figure@8.[469,1097,1180,1853]" captionTargetId-6="figure@9.[340,1247,193,967]" captionTargetId-7="figure@10.[348,1240,301,1298]" captionTargetId-8="figure@11.[340,1248,193,966]" captionTargetPageId-0="4" captionTargetPageId-1="6" captionTargetPageId-2="7" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="8" captionTargetPageId-6="9" captionTargetPageId-7="10" captionTargetPageId-8="11" captionText-0="FIGURE 1. Scyliorhinus ugoi sp. nov., holotype, MNRJ 42619, female, 500 mm TL. A) Dorsal view. B) Lateral view; paratype, UERJ 2179, male, 419 mm TL. C) Dorsal view. D) Lateral view. Scale bar: 5 cm." captionText-1="FIGURE 2. Scyliorhinus ugoi sp. nov. A) Dorsal view of the head of a female, MNRJ 42619, female, 500 mm TL. B) Dorsal view of the head of a male, UERJ 2179, 419 mm TL. C) Lateral view of head, UERJ 2179 male, 419 mm TL. D) First dorsal fin. E) Second dorsal fin. F) Anal fin. G) Caudal fin. H) Ventral view of trunk, MNRJ 42619, female, 500 mm TL. Scale bars: 2 cm." captionText-2="FIGURE 3. Eye of Scyliorhinus ugoi sp. nov., MNRJ 42619, female, 500 mm TL. Scale bar: 2 cm." captionText-3="FIGURE 4. Nostrils and mouth of Scyliorhinus ugoi sp. nov., MNRJ 42619, female, 500 mm TL. Scale bar: 2 cm." captionText-4="FIGURE 5. Lateral trunk denticles of Scyliorhinus ugoi sp. nov. A – C) UERJ 1726, female, 602 mm TL. D – F) UERJ 1725, adult male, 534 mm TL. A, D) Region above pectoral fin. B, E) Region below first dorsal fin. C, F) Region below second dorsal fin." captionText-5="FIGURE 6. Pectoral fin skeleton of Scyliorhinus ugoi sp. nov., UERJ 1722, female, 600 mm TL. Scale bar: 2 cm. DRA, distal segment; MES, mesopterygium; MET, metapterygium; MTS, metapterygial axis; PRA, proximal segment; PRO, propterygium." captionText-6="FIGURE 7. Clasper of Scyliorhinus ugoi sp. nov., UERJ 1725, male, 530 mm TL. A) Clasper gland. B) Skeleton. Scale bar = 5 mm. AP, apopyle; CRH, cover rhipidion; G, end-style; EN, envelope; ERH, exorhipidion; HP, hypopyle; RD, marginal dorsal cartilage; RD 2, accessory marginal dorsal cartilage; RH, rhipidion; RV, marginal ventral cartilage; TD, terminal dorsal cartilage; TDC, terminal dermal cover; TV, terminal ventral cartilage; TV 2, terminal ventral cartilage 2." captionText-7="FIGURE 8. Cranium of Scyliorhinus ugoi sp. nov., UERJ 1725, male, 530 mm TL. A) Dorsal view. B) Ventral view; UERJ 1726, female, 597 mm TL. C) Dorsal view. D) Ventral view. Scale bar = 1 cm. AF, anterior fontanelle; ASC, anterior semicircular canal; CRF, cranial roof; FOE, external foramen of preorbital canal; FPE, external profundus foramen; HF, hyomandibular facet; ICF, internal carotid foramen; IS, internasal septum; LR, lateral rostral cartilage; MR, medial rostral cartilage; NA, nasal aperture; NC, nasal capsule; NF, nasal fontanelle; OC, optic capsule; PEP, preorbital process; PRF, parietal fossa; PSC, posterior semicircular canal; PT, pterotic process; PTP, postorbital process; SC, supraorbital crest; SF, stapedial foramen; SG, supraorbital groove; SS, suborbital shelf." captionText-8="FIGURE 9. Distribution of Scyliorhinus ugoi sp. nov. Yellow diamonds represent records of type series, and white star locality of the holotype." httpUri-0="https://zenodo.org/record/243846/files/figure.png" httpUri-1="https://zenodo.org/record/243847/files/figure.png" httpUri-2="https://zenodo.org/record/243848/files/figure.png" httpUri-3="https://zenodo.org/record/243849/files/figure.png" httpUri-4="https://zenodo.org/record/243850/files/figure.png" httpUri-5="https://zenodo.org/record/243851/files/figure.png" httpUri-6="https://zenodo.org/record/243852/files/figure.png" httpUri-7="https://zenodo.org/record/243853/files/figure.png" httpUri-8="https://zenodo.org/record/243854/files/figure.png" pageId="1" pageNumber="348">Figs. 1–9</figureCitation>
)
</paragraph>
<paragraph id="8BFB368AFF88FF9A17AEFCE6FE84FC1A" blockId="1.[151,1436,829,974]" pageId="1" pageNumber="348">
Proposed common name: dark freckled catshark (English), cação-gato negrinho (Portuguese). 
<emphasis id="B930EA98FF88FF9A17AEFC87FEC3FCF8" box="[151,267,860,882]" italics="true" pageId="1" pageNumber="348">Scyliohinus</emphasis>
sp.: 
<bibRefCitation id="EFD54B7BFF88FF9A1603FC87FE71FCF8" author="Gadig" box="[314,441,860,882]" pageId="1" pageNumber="348" refString="Gadig, O. B. F. (2001) Tubaroes da costa brasileira. Unpubl. PhD thesis, Universidade Estadual Paulista ' Julio Mesquita Filho', Rio Claro, Sao Paulo, 343 pp." type="book" year="2001">Gadig, 2001</bibRefCitation>
, pp. 149–150, coast of Espírito Santo state; 
<bibRefCitation id="EFD54B7BFF88FF9A144DFC87FBE7FCF8" author="Gomes" box="[884,1071,860,882]" pageId="1" pageNumber="348" refString="Gomes, U. L., Signori, C. N., Gadig, O. B. F. &amp; Santos, H. R. S. (2010) Guia para Identificacao de Tubaroes e Raias do Rio de Janeiro. Technical Books, Rio de Janeiro, 234 pp." type="book" year="2010">
Gomes 
<emphasis id="B930EA98FF88FF9A14FBFC86FC26FCF8" box="[962,1006,860,882]" italics="true" pageId="1" pageNumber="348">et al</emphasis>
. 2010
</bibRefCitation>
, pp. 85–86, fig. 110, coast of Rio de Janeiro state.
</paragraph>
</subSubSection>
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<paragraph id="8BFB368AFF88FF9A17AEFC42FD1AFC44" blockId="1.[151,1436,829,974]" pageId="1" pageNumber="348">
<taxonomicName id="4C444D09FF88FF9A17AEFC42FDA7FC25" ID-CoL="4W57C" authority="Gadig &amp; Gomes, 2003" authorityName="Gadig &amp; Gomes" authorityYear="2003" box="[151,623,921,943]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="hesperius">
<emphasis id="B930EA98FF88FF9A17AEFC42FEB3FC25" box="[151,379,921,943]" italics="true" pageId="1" pageNumber="348">Scyliorhinus hesperius</emphasis>
: 
<bibRefCitation id="EFD54B7BFF88FF9A16B1FC42FDA7FC25" author="Gadig" box="[392,623,921,943]" pageId="1" pageNumber="348" refString="Gadig, O. B. F. &amp; Gomes, U. L. (2003) Classe Chondrichthyes. In: Menezes, N. A., Buckup, P. A., Figueiredo, J. L. &amp; Moura, R. L. (Eds.), Catalogo das especies de peixes marinhos do Brasil. Museu de Zoologia da Universidade de Sao Paulo, Sao Paulo, pp. 21 - 22." type="book chapter" year="2003">Gadig &amp; Gomes, 2003</bibRefCitation>
</taxonomicName>
, p. 22, coasts of Espírito Santo and Rio de Janeiro states. 
<taxonomicName id="4C444D09FF88FF9A17AEFC63FEDDFC44" ID-CoL="63SXP" box="[151,277,952,974]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="genus">
<emphasis id="B930EA98FF88FF9A17AEFC63FEDDFC44" box="[151,277,952,974]" italics="true" pageId="1" pageNumber="348">Scyliorhinus</emphasis>
</taxonomicName>
sp. 1: 
<bibRefCitation id="EFD54B7BFF88FF9A1661FC63FE16FC44" author="Soares" box="[344,478,952,974]" pageId="1" pageNumber="348" refString="Soares, K. D. A. (2014) Estudo taxonomico das especies de Scyliorhinus do grupo haeckelii / besnardi (Carcharhiniformes, Scyliorhinidae) das regioes Sudeste e Sul do Brasil (Oceano Atlantico Sul Ocidental). Unpubl. Master's thesis, Universidade do Estado do Rio de Janeiro, Rio de Janeiro, 208 pp." type="book" year="2014">Soares, 2014</bibRefCitation>
, pp. 80–84, figs. 17, 68.
</paragraph>
</subSubSection>
<subSubSection id="C35E6501FF88FF9A17AEFC21FCB7FB4D" pageId="1" pageNumber="348" type="materials_examined">
<paragraph id="8BFB368AFF88FF9A17AEFC21FB64FB99" blockId="1.[151,1437,1018,1979]" box="[151,1196,1018,1043]" pageId="1" pageNumber="348">
<typeStatus id="54FF8828FF88FF9A17AEFC21FECBFB99" box="[151,259,1018,1043]" pageId="1" pageNumber="348" type="holotype">
<emphasis id="B930EA98FF88FF9A17AEFC21FECBFB99" bold="true" box="[151,259,1018,1043]" pageId="1" pageNumber="348">Holotype</emphasis>
</typeStatus>
. 
<collectionCode id="ED55AE4FFF88FF9A1629FC20FE92FB98" box="[272,346,1019,1042]" httpUri="http://grbio.org/cool/zi1i-a0b5" name="Museu Nacional/Universidade Federal de Rio de Janeiro" pageId="1" pageNumber="348">MNRJ</collectionCode>
42619, adult female, 
<quantity id="4CBC9B6FFF88FF9A1577FC21FD63FB98" box="[590,683,1018,1043]" metricMagnitude="-1" metricUnit="m" metricValue="4.96" pageId="1" pageNumber="348" unit="mm" value="496.0">496 mm</quantity>
TL (southern Bahia, Central Brazilian coast).
</paragraph>
<paragraph id="8BFB368AFF88FF9A17FEFBC5FA4EFB29" blockId="1.[151,1437,1018,1979]" pageId="1" pageNumber="348">
<typeStatus id="54FF8828FF88FF9A17FEFBC5FEF7FBBD" box="[199,319,1054,1079]" pageId="1" pageNumber="348" type="paratype">
<emphasis id="B930EA98FF88FF9A17FEFBC5FEF7FBBD" bold="true" box="[199,319,1054,1079]" pageId="1" pageNumber="348">Paratypes</emphasis>
</typeStatus>
. 
<collectionCode id="ED55AE4FFF88FF9A166FFBC4FE7BFBBD" box="[342,435,1055,1079]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="348">MZUSP</collectionCode>
<date id="FFFA104AFF88FF9A16FDFBC4FDD2FBBD" box="[452,538,1055,1079]" pageId="1" pageNumber="348" value="48-11-04">110448</date>
, adult male, 
<quantity id="4CBC9B6FFF88FF9A15FDFBC5FCE3FBBC" box="[708,811,1054,1079]" metricMagnitude="-1" metricUnit="m" metricValue="4.65" pageId="1" pageNumber="348" unit="mm" value="465.0">465 mm</quantity>
TL (Alagoas, Northeastern 
<collectingCountry id="F353761AFF88FF9A13BCFBC5FB03FBBD" box="[1157,1227,1054,1079]" name="Brazil" pageId="1" pageNumber="348">Brazil</collectingCountry>
, 9° S 
<geoCoordinate id="EE70504DFF88FF9A1223FBC4FA47FBBD" box="[1306,1423,1054,1079]" direction="west" orientation="longitude" pageId="1" pageNumber="348" precision="925" value="-34.833332">34° 50’W</geoCoordinate>
); 
<collectionCode id="ED55AE4FFF88FF9A17AEFB98FF3CFBD1" box="[151,244,1091,1115]" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="1" pageNumber="348">MZUSP</collectionCode>
<date id="FFFA104AFF88FF9A17C4FB98FE9BFBD1" box="[253,339,1091,1115]" pageId="1" pageNumber="348" value="49-11-04">110449</date>
, adult male, 
<quantity id="4CBC9B6FFF88FF9A16D0FB99FD80FBD0" box="[489,584,1090,1115]" metricMagnitude="-1" metricUnit="m" metricValue="4.45" pageId="1" pageNumber="348" unit="mm" value="445.0">445 mm</quantity>
TL (Rio Grande do Norte, Northeastern 
<collectingCountry id="F353761AFF88FF9A131AFB99FBA2FBD1" box="[1059,1130,1090,1115]" name="Brazil" pageId="1" pageNumber="348">Brazil</collectingCountry>
, 
<geoCoordinate id="EE70504DFF88FF9A134CFB99FB01FBD1" box="[1141,1225,1090,1115]" direction="south" orientation="latitude" pageId="1" pageNumber="348" precision="925" value="-6.233333">6° 14’S</geoCoordinate>
<geoCoordinate id="EE70504DFF88FF9A13E8FB98FA89FBD1" box="[1233,1345,1090,1115]" direction="west" orientation="longitude" pageId="1" pageNumber="348" precision="925" value="-34.85">34° 51’W</geoCoordinate>
); 
<collectionCode id="ED55AE4FFF88FF9A1263FB98FA54FBD0" box="[1370,1436,1091,1114]" pageId="1" pageNumber="348">UERJ</collectionCode>
1725, adult male, 
<quantity id="4CBC9B6FFF88FF9A164EFBBDFE14FBF4" box="[375,476,1126,1151]" metricMagnitude="-1" metricUnit="m" metricValue="5.3" pageId="1" pageNumber="348" unit="mm" value="530.0">530 mm</quantity>
TL (Central Brazilian coast, between southern Bahia and northern Rio de Janeiro; cranium, jaws and claspers); 
<collectionCode id="ED55AE4FFF88FF9A16E6FB50FDE9FB28" box="[479,545,1163,1186]" pageId="1" pageNumber="348">UERJ</collectionCode>
2179, immature male, 
<quantity id="4CBC9B6FFF88FF9A1410FB51FC4EFB28" box="[809,902,1162,1187]" metricMagnitude="-1" metricUnit="m" metricValue="4.15" pageId="1" pageNumber="348" unit="mm" value="415.0">415 mm</quantity>
TL (southern Bahia, Central Brazilian coast).
</paragraph>
<paragraph id="8BFB368AFF88FF9A17FEFB75FCB7FB4D" blockId="1.[151,1437,1018,1979]" box="[199,895,1198,1223]" pageId="1" pageNumber="348">
<emphasis id="B930EA98FF88FF9A17FEFB75FDE4FB4D" bold="true" box="[199,556,1198,1223]" pageId="1" pageNumber="348">Additional material examined</emphasis>
. 
<specimenCount id="9D42FD03FF88FF9A1502FB75FD0CFB4D" box="[571,708,1198,1223]" pageId="1" pageNumber="348" type="generic">5 specimens</specimenCount>
(see Appendix).
</paragraph>
</subSubSection>
<subSubSection id="C35E6501FF88FF9A17FEFB09FDC8F969" pageId="1" pageNumber="348" type="diagnosis">
<paragraph id="8BFB368AFF88FF9A17FEFB09FDC8F969" blockId="1.[151,1437,1018,1979]" pageId="1" pageNumber="348">
<emphasis id="B930EA98FF88FF9A17FEFB09FE89FB61" bold="true" box="[199,321,1234,1259]" pageId="1" pageNumber="348">Diagnosis.</emphasis>
A southwestern Atlantic Ocean 
<taxonomicName id="4C444D09FF88FF9A15F5FB08FC9EFB61" box="[716,854,1235,1259]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="genus">
<emphasis id="B930EA98FF88FF9A15F5FB08FC9EFB61" box="[716,854,1235,1259]" italics="true" pageId="1" pageNumber="348">Scyliorhinus</emphasis>
</taxonomicName>
distinguished by a considerably broad head (
<emphasis id="B930EA98FF88FF9A1244FB08FA54FB60" box="[1405,1436,1235,1258]" italics="true" pageId="1" pageNumber="348">vs.</emphasis>
slender in the 
<emphasis id="B930EA98FF88FF9A1604FB2CFDE2FA84" box="[317,554,1270,1294]" italics="true" pageId="1" pageNumber="348">
<taxonomicName id="4C444D09FF88FF9A1604FB2CFE76FA84" box="[317,446,1270,1294]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="haeckelii">S. haeckelii</taxonomicName>
/ 
<taxonomicName id="4C444D09FF88FF9A16F2FB2DFDE2FA84" box="[459,554,1270,1294]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="besnardi">besnardi</taxonomicName>
</emphasis>
group and 
<taxonomicName id="4C444D09FF88FF9A158BFB2CFCF4FA84" box="[690,828,1270,1294]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="hesperius">
<emphasis id="B930EA98FF88FF9A158BFB2CFCF4FA84" box="[690,828,1270,1294]" italics="true" pageId="1" pageNumber="348">S. hesperius</emphasis>
</taxonomicName>
), and color pattern with brown saddles on a light brown background, predorsal saddles with anterior and posterior sharp median projections (
<emphasis id="B930EA98FF88FF9A13E1FAC0FB27FAB8" box="[1240,1263,1307,1330]" italics="true" pageId="1" pageNumber="348">vs</emphasis>
. no projections in the 
<emphasis id="B930EA98FF88FF9A17DDFAE4FE07FADC" box="[228,463,1342,1366]" italics="true" pageId="1" pageNumber="348">
<taxonomicName id="4C444D09FF88FF9A17DDFAE4FEADFADC" box="[228,357,1342,1366]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="haeckelii">S. haeckelii</taxonomicName>
/ 
<taxonomicName id="4C444D09FF88FF9A1648FAE5FE07FADC" box="[369,463,1342,1366]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="besnardi">besnardi</taxonomicName>
</emphasis>
group or 
<taxonomicName id="4C444D09FF88FF9A1571FAE4FD1AFADC" box="[584,722,1342,1366]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="hesperius">
<emphasis id="B930EA98FF88FF9A1571FAE4FD1AFADC" box="[584,722,1342,1366]" italics="true" pageId="1" pageNumber="348">S. hesperius</emphasis>
</taxonomicName>
); and light and dark, spiracle-sized or slightly smaller spots, lacking large white spots (
<emphasis id="B930EA98FF88FF9A16F4FAB8FE2CFAF0" box="[461,484,1379,1402]" italics="true" pageId="1" pageNumber="348">vs</emphasis>
. large white spots in 
<taxonomicName id="4C444D09FF88FF9A15DCFAB8FCA7FAF1" box="[741,879,1379,1403]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="hesperius">
<emphasis id="B930EA98FF88FF9A15DCFAB8FCA7FAF1" box="[741,879,1379,1403]" italics="true" pageId="1" pageNumber="348">S. hesperius</emphasis>
</taxonomicName>
). The following combination of characters also distinguishes these species: snout rounded and moderately short, preoral length 5% TL (
<emphasis id="B930EA98FF88FF9A1391FA5CFB0EFA14" box="[1192,1222,1415,1438]" italics="true" pageId="1" pageNumber="348">vs.</emphasis>
4.5% TL in the 
<emphasis id="B930EA98FF88FF9A12BEFA5CFEAAFA49" italics="true" pageId="1" pageNumber="348">
<taxonomicName id="4C444D09FF88FF9A12BEFA5CFF30FA49" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="haeckelii">S. haeckelii</taxonomicName>
/ 
<taxonomicName id="4C444D09FF88FF9A163DFA70FEAAFA49" box="[260,354,1451,1475]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="besnardi">besnardi</taxonomicName>
</emphasis>
group); preorbital length 6.4% TL and head length 20% TL (
<emphasis id="B930EA98FF88FF9A1306FA70FB9EFA48" box="[1087,1110,1451,1474]" italics="true" pageId="1" pageNumber="348">vs</emphasis>
. 5% TL and 18% TL in 
<taxonomicName id="4C444D09FF88FF9A12BEFA70FECAFA6C" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="hesperius">
<emphasis id="B930EA98FF88FF9A12BEFA70FECAFA6C" italics="true" pageId="1" pageNumber="348">S. hesperius</emphasis>
</taxonomicName>
); mouth width corresponding to 9% TL (
<emphasis id="B930EA98FF88FF9A15EDFA14FD23FA6C" box="[724,747,1487,1510]" italics="true" pageId="1" pageNumber="348">vs</emphasis>
. 8% TL in the 
<emphasis id="B930EA98FF88FF9A14ACFA14FBB7FA6C" box="[917,1151,1486,1510]" italics="true" pageId="1" pageNumber="348">
<taxonomicName id="4C444D09FF88FF9A14ACFA14FBDCFA6C" box="[917,1044,1486,1510]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="haeckelii">S. haeckelii</taxonomicName>
/ 
<taxonomicName id="4C444D09FF88FF9A1319FA15FBB7FA6C" box="[1056,1151,1486,1510]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="besnardi">besnardi</taxonomicName>
</emphasis>
group and 7.2% TL in 
<taxonomicName id="4C444D09FF88FF9A12BEFA14FECAF981" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="hesperius">
<emphasis id="B930EA98FF88FF9A12BEFA14FECAF981" italics="true" pageId="1" pageNumber="348">S. hesperius</emphasis>
</taxonomicName>
); pectoral-pelvic space 1.5 times pelvic-anal space (
<emphasis id="B930EA98FF88FF9A145DFA28FCB3F980" box="[868,891,1523,1546]" italics="true" pageId="1" pageNumber="348">vs</emphasis>
. equal in the 
<emphasis id="B930EA98FF88FF9A1319FA28FAC5F981" box="[1056,1293,1523,1547]" italics="true" pageId="1" pageNumber="348">
<taxonomicName id="4C444D09FF88FF9A1319FA28FB69F981" box="[1056,1185,1523,1547]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="haeckelii">S. haeckelii</taxonomicName>
/ 
<taxonomicName id="4C444D09FF88FF9A1397FA28FAC5F981" box="[1198,1293,1523,1547]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="besnardi">besnardi</taxonomicName>
</emphasis>
group), and interdorsal space 2.3 times dorsal-caudal space (
<emphasis id="B930EA98FF88FF9A1581F9CCFD1EF9A4" box="[696,726,1559,1582]" italics="true" pageId="1" pageNumber="348">vs.</emphasis>
1.6 times in 
<emphasis id="B930EA98FF88FF9A1450F9CCFB9BF9A4" box="[873,1107,1558,1582]" italics="true" pageId="1" pageNumber="348">
<taxonomicName id="4C444D09FF88FF9A1450F9CCFC20F9A4" box="[873,1000,1558,1582]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="haeckelii">S. haeckelii</taxonomicName>
/ 
<taxonomicName id="4C444D09FF88FF9A14CDF9CDFB9BF9A4" box="[1012,1107,1558,1582]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="besnardi">besnardi</taxonomicName>
</emphasis>
group); anal-fin base 8% TL (
<emphasis id="B930EA98FF88FF9A1799F9E0FF7FF9D8" box="[160,183,1595,1618]" italics="true" pageId="1" pageNumber="348">vs</emphasis>
. 6.7% TL in S. 
<taxonomicName id="4C444D09FF88FF9A1651F9E0FE1BF9D9" box="[360,467,1595,1619]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="hesperius">
<emphasis id="B930EA98FF88FF9A1651F9E0FE1BF9D9" box="[360,467,1595,1619]" italics="true" pageId="1" pageNumber="348">hesperius</emphasis>
</taxonomicName>
and 8.3% TL in the 
<emphasis id="B930EA98FF88FF9A1581F9E0FC57F9D9" box="[696,927,1595,1619]" italics="true" pageId="1" pageNumber="348">
<taxonomicName id="4C444D09FF88FF9A1581F9E0FCFDF9D9" box="[696,821,1595,1619]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="haeckelii">S. haeckelii</taxonomicName>
/ 
<taxonomicName id="4C444D09FF88FF9A1478F9E0FC57F9D9" box="[833,927,1595,1619]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="besnardi">besnardi</taxonomicName>
</emphasis>
group); cranium with flat basal plate, without keels; cranial width larger in females (42.7% NL) (
<emphasis id="B930EA98FF88FF9A15DBF984FCC8F9FC" box="[738,768,1631,1654]" italics="true" pageId="1" pageNumber="348">vs.</emphasis>
37.4% NL in the 
<emphasis id="B930EA98FF88FF9A14D9F984FB00F9FC" box="[992,1224,1630,1654]" italics="true" pageId="1" pageNumber="348">
<taxonomicName id="4C444D09FF88FF9A14D9F984FB95F9FC" box="[992,1117,1630,1654]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="haeckelii">S. haeckelii</taxonomicName>
/ 
<taxonomicName id="4C444D09FF88FF9A1353F985FB00F9FC" box="[1130,1224,1630,1654]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="besnardi">besnardi</taxonomicName>
</emphasis>
group); a terminal dermal cover present in hindmost portion of clasper, covering terminal cartilages (lacking in 
<taxonomicName id="4C444D09FF88FF9A123AF958FA42F911" box="[1283,1418,1667,1691]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="hesperius">
<emphasis id="B930EA98FF88FF9A123AF958FA42F911" box="[1283,1418,1667,1691]" italics="true" pageId="1" pageNumber="348">S. hesperius</emphasis>
</taxonomicName>
); medium-sized, adult male at 
<quantity id="4CBC9B6FFF88FF9A16E2F97DFDFFF934" box="[475,567,1702,1727]" metricMagnitude="-1" metricUnit="m" metricValue="4.45" pageId="1" pageNumber="348" unit="mm" value="445.0">445 mm</quantity>
TL and adult female 
<quantity id="4CBC9B6FFF88FF9A1413F97DFC4EF934" box="[810,902,1702,1727]" metricMagnitude="-1" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="348" unit="mm" value="500.0">500 mm</quantity>
TL (
<emphasis id="B930EA98FF88FF9A14F8F97CFC17F934" box="[961,991,1703,1726]" italics="true" pageId="1" pageNumber="348">vs.</emphasis>
<quantity id="4CBC9B6FFF88FF9A14DFF97DFB8CF934" box="[998,1092,1702,1727]" metricMagnitude="-1" metricUnit="m" metricValue="3.53" pageId="1" pageNumber="348" unit="mm" value="353.0">353 mm</quantity>
and 
<quantity id="4CBC9B6FFF88FF9A1342F97CFB13F934" box="[1147,1243,1703,1727]" metricMagnitude="-1" metricUnit="m" metricValue="4.1" pageId="1" pageNumber="348" unit="mm" value="410.0">410 mm</quantity>
, respectively, for the 
<emphasis id="B930EA98FF88FF9A17F9F910FE6FF969" box="[192,423,1739,1763]" italics="true" pageId="1" pageNumber="348">
<taxonomicName id="4C444D09FF88FF9A17F9F910FEF5F969" box="[192,317,1739,1763]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="haeckelii">S. haeckelii</taxonomicName>
/ 
<taxonomicName id="4C444D09FF88FF9A1670F910FE6FF969" box="[329,423,1739,1763]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="besnardi">besnardi</taxonomicName>
</emphasis>
group).
</paragraph>
</subSubSection>
<subSubSection id="C35E6501FF88FF9117FEF935FDA7FF7D" lastPageId="10" lastPageNumber="357" pageId="1" pageNumber="348" type="description">
<paragraph id="8BFB368AFF88FF9A17FEF935FC0CF8A1" blockId="1.[151,1437,1018,1979]" pageId="1" pageNumber="348">
<emphasis id="B930EA98FF88FF9A17FEF935FE91F88D" bold="true" box="[199,345,1774,1799]" pageId="1" pageNumber="348">Description.</emphasis>
Morphometric measurements are given in 
<tableCitation id="C6C60331FF88FF9A140EF935FC40F88D" box="[823,904,1774,1799]" captionStart="TABLE 1" captionStartId="2.[151,239,152,175]" captionTargetBox="[159,1430,276,1951]" captionTargetPageId="2" captionText="TABLE 1. Measurements in millimeters and proportions as percentage of total length for Scyliorhinus ugoi sp. nov. Mean (x), standard deviation (SD) and range of proportions (%)." httpUri="http://table.plazi.org/id/DF3B6602FF8BFF9917AEFF43FC4FFF45" pageId="1" pageNumber="348" tableUuid="DF3B6602FF8BFF9917AEFF43FC4FFF45">Table 1</tableCitation>
and cranial measurements in 
<tableCitation id="C6C60331FF88FF9A13EAF935FAE0F88D" box="[1235,1320,1774,1799]" captionStart="TABLE 2" captionStartId="5.[151,239,152,175]" captionTargetBox="[159,1335,317,628]" captionTargetPageId="5" captionText="TABLE 2. Measurements in millimeters and proportions as percentage of nasobasal length for Scyliorhinus ugoi sp. nov. Female (F), male (M), and range of proportions (% NL)." httpUri="http://table.plazi.org/id/DF3B6602FF8CFF9E17AEFF43FD20FF45" pageId="1" pageNumber="348" tableUuid="DF3B6602FF8CFF9E17AEFF43FD20FF45">Table 2</tableCitation>
. Modes of meristic counts are given as well their range in parentheses (if different).
</paragraph>
<paragraph id="8BFB368AFF88FF9A17FEF8EDFD63F831" blockId="1.[151,1437,1018,1979]" pageId="1" pageNumber="348">
Head (
<figureCitation id="137F2A0FFF88FF9A162CF8EDFE94F8C5" box="[277,348,1846,1871]" captionStart="FIGURE 2" captionStartId="6.[151,260,1897,1919]" captionTargetBox="[346,1240,665,1868]" captionTargetId="figure@6.[340,1248,661,1875]" captionTargetPageId="6" captionText="FIGURE 2. Scyliorhinus ugoi sp. nov. A) Dorsal view of the head of a female, MNRJ 42619, female, 500 mm TL. B) Dorsal view of the head of a male, UERJ 2179, 419 mm TL. C) Lateral view of head, UERJ 2179 male, 419 mm TL. D) First dorsal fin. E) Second dorsal fin. F) Anal fin. G) Caudal fin. H) Ventral view of trunk, MNRJ 42619, female, 500 mm TL. Scale bars: 2 cm." httpUri="https://zenodo.org/record/243847/files/figure.png" pageId="1" pageNumber="348">Fig. 2</figureCitation>
a–c) broad and depressed, greatest width 0.63 times of head length. Snout thick, somewhat flattened and relatively short, preoral length corresponding to 4.3–5.6% TL and 0.5 times the mouth width. Ampullae pores not greatly enlarged on snout. Preoral length 1.1–1.4 times in preorbital length (
<emphasis id="B930EA98FF88FF9A13E5F8A4FB3BF81C" box="[1244,1267,1919,1942]" italics="true" pageId="1" pageNumber="348">vs</emphasis>
. 
<quantity id="4CBC9B6FFF88FF9A123BF8A4FABBF81D" box="[1282,1395,1918,1943]" metricMagnitude="-2" metricUnit="m" metricValue="3.9369999999999994" metricValueMax="4.318" metricValueMin="3.5559999999999996" pageId="1" pageNumber="348" unit="in" value="1.5499999999999998" valueMax="1.7" valueMin="1.4">1.4–1.7 in</quantity>
the 
<emphasis id="B930EA98FF88FF9A17AEF878FE3BF831" box="[151,499,1955,1979]" italics="true" pageId="1" pageNumber="348">
<taxonomicName id="4C444D09FF88FF9A17AEF878FE41F831" box="[151,393,1955,1979]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="haeckelii">Scyliorhinus haeckelii</taxonomicName>
/ 
<taxonomicName id="4C444D09FF88FF9A16ACF878FE3BF831" box="[405,499,1955,1979]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="1" pageNumber="348" phylum="Chordata" rank="species" species="besnardi">besnardi</taxonomicName>
</emphasis>
group) (
<tableCitation id="C6C60331FF88FF9A156DF879FD51F831" box="[596,665,1954,1979]" captionStart="TABLE 1" captionStartId="2.[151,239,152,175]" captionTargetBox="[159,1430,276,1951]" captionTargetPageId="2" captionText="TABLE 1. Measurements in millimeters and proportions as percentage of total length for Scyliorhinus ugoi sp. nov. Mean (x), standard deviation (SD) and range of proportions (%)." httpUri="http://table.plazi.org/id/DF3B6602FF8BFF9917AEFF43FC4FFF45" pageId="1" pageNumber="348" tableUuid="DF3B6602FF8BFF9917AEFF43FC4FFF45">Tab. 1</tableCitation>
).
</paragraph>
<caption id="DF3B6602FF8BFF9917AEFF43FC4FFF45" ID-Table-UUID="DF3B6602FF8BFF9917AEFF43FC4FFF45" httpUri="http://table.plazi.org/id/DF3B6602FF8BFF9917AEFF43FC4FFF45" pageId="2" pageNumber="349" targetBox="[159,1430,276,1951]" targetIsTable="true" targetPageId="2">
<paragraph id="8BFB368AFF8BFF9917AEFF43FC4FFF45" blockId="2.[151,1436,151,207]" pageId="2" pageNumber="349">
<emphasis id="B930EA98FF8BFF9917AEFF43FECAFF25" bold="true" box="[151,258,152,175]" pageId="2" pageNumber="349">TABLE 1</emphasis>
. Measurements in millimeters and proportions as percentage of total length for 
<taxonomicName id="4C444D09FF8BFF99138DFF4CFABCFF25" box="[1204,1396,151,175]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="2" pageNumber="349" phylum="Chordata" rank="species" species="ugoi" status="sp. nov.">
<emphasis id="B930EA98FF8BFF99138DFF4CFABCFF25" box="[1204,1396,151,175]" italics="true" pageId="2" pageNumber="349">Scyliorhinus ugoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20357E3FF8BFF991245FF43FF0EFF45" pageId="2" pageNumber="349" rank="species">
<emphasis id="B930EA98FF8BFF991245FF43FF0EFF45" bold="true" pageId="2" pageNumber="349">sp. nov.</emphasis>
</taxonomicNameLabel>
Mean (
<emphasis id="B930EA98FF8BFF991623FF63FEEEFF45" box="[282,294,184,207]" italics="true" pageId="2" pageNumber="349">x</emphasis>
), standard deviation (SD) and range of proportions (%).
</paragraph>
</caption>
<paragraph id="8BFB368AFF8BFF9817A6FF30FE8AF84F" blockId="2.[159,1336,234,256]" lastBlockId="3.[151,1437,1567,1989]" lastPageId="3" lastPageNumber="350" pageId="2" pageNumber="349">
Measurements 
<taxonomicName id="4C444D09FF8BFF991505FF31FD25FE8A" box="[572,749,234,256]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="2" pageNumber="349" phylum="Chordata" rank="species" species="ugoi" status="sp. nov.">
<emphasis id="B930EA98FF8BFF991505FF31FD25FE8A" box="[572,749,234,256]" italics="true" pageId="2" pageNumber="349">Scyliorhinus ugoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20357E3FF8BFF9915CAFF31FC86FE8A" box="[755,846,234,256]" pageId="2" pageNumber="349" rank="species">
<emphasis id="B930EA98FF8BFF9915CAFF31FC86FE8A" bold="true" box="[755,846,234,256]" pageId="2" pageNumber="349">sp. nov.</emphasis>
</taxonomicNameLabel>
<emphasis id="B930EA98FF8BFF9914C0FF31FA5EF842" italics="true" pageId="2" pageNumber="349">
<taxonomicName id="4C444D09FF8BFF9914C0FF31FB1EFE8A" box="[1017,1238,234,256]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="2" pageNumber="349" phylum="Chordata" rank="species" species="haeckelii">Scyliorhinus haeckelii</taxonomicName>
/ 
<taxonomicName id="4C444D09FF8BFF9913DBFF31FAF0FE8A" box="[1250,1336,234,256]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="2" pageNumber="349" phylum="Chordata" rank="species" species="besnardi">besnardi</taxonomicName>
......continued on the next page
</emphasis>
Measurements 
<taxonomicName id="4C444D09FF8AFF981505FF12FD25FF55" box="[572,749,201,223]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="3" pageNumber="350" phylum="Chordata" rank="species" species="ugoi" status="sp. nov.">
<emphasis id="B930EA98FF8AFF981505FF12FD25FF55" box="[572,749,201,223]" italics="true" pageId="3" pageNumber="350">Scyliorhinus ugoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20357E3FF8AFF9815CAFF12FC86FF55" box="[755,846,201,223]" pageId="3" pageNumber="350" rank="species">
<emphasis id="B930EA98FF8AFF9815CAFF12FC86FF55" bold="true" box="[755,846,201,223]" pageId="3" pageNumber="350">sp. nov.</emphasis>
</taxonomicNameLabel>
<emphasis id="B930EA98FF8AFF9814C0FF12FAF0FF55" box="[1017,1336,201,223]" italics="true" pageId="3" pageNumber="350">
<taxonomicName id="4C444D09FF8AFF9814C0FF12FB1EFF55" box="[1017,1238,201,223]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="3" pageNumber="350" phylum="Chordata" rank="species" species="haeckelii">Scyliorhinus haeckelii</taxonomicName>
/ 
<taxonomicName id="4C444D09FF8AFF9813DBFF12FAF0FF55" box="[1250,1336,201,223]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="3" pageNumber="350" phylum="Chordata" rank="species" species="besnardi">besnardi</taxonomicName>
</emphasis>
Eyes (
<figureCitation id="137F2A0FFF8AFF981628F9C4FE9FF9B2" box="[273,343,1567,1592]" captionStart="FIGURE 3" captionStartId="7.[151,249,648,670]" captionTargetBox="[436,1163,193,607]" captionTargetId="figure@7.[431,1163,193,621]" captionTargetPageId="7" captionText="FIGURE 3. Eye of Scyliorhinus ugoi sp. nov., MNRJ 42619, female, 500 mm TL. Scale bar: 2 cm." httpUri="https://zenodo.org/record/243848/files/figure.png" pageId="3" pageNumber="350">Fig. 3</figureCitation>
) large and spindle-shaped; eye length 5.2 times in head length and 2.5–4.2 times eye height. Eyes dorsolateral on head, with lower edges well medial to horizontal head rim in dorsal view; subocular ridges strong. Nictitating lower eyelids of rudimentary 
<typeStatus id="54FF8828FF8AFF9815FBF9B3FD3AF90A" box="[706,754,1640,1664]" pageId="3" pageNumber="350">type</typeStatus>
(
<emphasis id="B930EA98FF8AFF98143FF9B2FC8CF90A" box="[774,836,1641,1664]" italics="true" pageId="3" pageNumber="350">sensu</emphasis>
<bibRefCitation id="EFD54B7BFF8AFF981477F9B3FBDEF90A" author="Compagno" box="[846,1046,1640,1664]" pageId="3" pageNumber="350" refString="Compagno, L. J. V. (1970) Systematics of the genus Hemitriakis (Selachii: Carcharhinidae), and related genera. Proceedings of California Academic Sciences, Series 4, 38, 63 - 98." type="journal article" year="1970">Compagno, 1970</bibRefCitation>
), with shallow subocular pouches and secondary lower eyelids free from upper eyelids. Prespiracular length corresponding to 10.5% TL (
<tableCitation id="C6C60331FF8AFF98127DF957FA43F92E" box="[1348,1419,1676,1701]" captionStart="TABLE 1" captionStartId="2.[151,239,152,175]" captionTargetBox="[159,1430,276,1951]" captionTargetPageId="2" captionText="TABLE 1. Measurements in millimeters and proportions as percentage of total length for Scyliorhinus ugoi sp. nov. Mean (x), standard deviation (SD) and range of proportions (%)." httpUri="http://table.plazi.org/id/DF3B6602FF8BFF9917AEFF43FC4FFF45" pageId="3" pageNumber="350" tableUuid="DF3B6602FF8BFF9917AEFF43FC4FFF45">Tab. 1</tableCitation>
). Nostrils (
<figureCitation id="137F2A0FFF8AFF98163BF974FE8FF942" box="[258,327,1711,1736]" captionStart="FIGURE 4" captionStartId="7.[151,249,1454,1476]" captionTargetBox="[495,1143,740,1431]" captionTargetId="figure@7.[433,1159,720,1432]" captionTargetPageId="7" captionText="FIGURE 4. Nostrils and mouth of Scyliorhinus ugoi sp. nov., MNRJ 42619, female, 500 mm TL. Scale bar: 2 cm." httpUri="https://zenodo.org/record/243849/files/figure.png" pageId="3" pageNumber="350">Fig. 4</figureCitation>
) with broad incurrent apertures, without nasoral grooves or nasal barbels, small oval excurrent apertures and short posterior nasal flaps. Anterior nasal flaps large, covering posterior nasal flaps and excurrent apertures somewhat anterior to mouth. Internarial space 2.4–4.6 times in interorbital space.Mouth (
<figureCitation id="137F2A0FFF8AFF981256F92CFF6EF8BE" captionStart="FIGURE 4" captionStartId="7.[151,249,1454,1476]" captionTargetBox="[495,1143,740,1431]" captionTargetId="figure@7.[433,1159,720,1432]" captionTargetPageId="7" captionText="FIGURE 4. Nostrils and mouth of Scyliorhinus ugoi sp. nov., MNRJ 42619, female, 500 mm TL. Scale bar: 2 cm." httpUri="https://zenodo.org/record/243849/files/figure.png" pageId="3" pageNumber="350">Fig. 4</figureCitation>
) broadly arched, moderately wide and short, mouth width 2.2 times in head length; mouth length 0.5 times in mouth width. Teeth prominently exposed in ventral view. Lower labial furrows short, 0.2 times in mouth width; upper labial furrows absent. Tongue moderate-sized flat and rounded, occupying most of mouth floor. Dorsal labial cartilages 1.5 times the ventral one; anterior tip of dorsal cartilages reaching orbital processes of palatoquadrate.
</paragraph>
<paragraph id="8BFB368AFF8BFF991505FECEFAEDF815" pageId="2" pageNumber="349">
<table id="F944C42AFF8B006417A6FECFFA5EF815" box="[159,1430,276,1951]" colsContinueIn="3.[159,1342,242,1504]" gridcols="5" gridrows="41" pageId="2" pageNumber="349">
<tr id="357434C8FF8B006417A6FECFFA5EFEA0" box="[159,1430,276,298]" gridrow="0" pageId="2" pageNumber="349" rowspan-0="1">
<th id="76A55DB4FF8B00641502FECFFD01FEA0" box="[571,713,276,298]" gridcol="1" gridrow="0" pageId="2" pageNumber="349">x SD</th>
<th id="76A55DB4FF8B00641420FECFFCB6FEA0" box="[793,894,276,298]" gridcol="2" gridrow="0" pageId="2" pageNumber="349">%</th>
<th id="76A55DB4FF8B006414C0FECFFB41FEA0" box="[1017,1161,276,298]" gridcol="3" gridrow="0" pageId="2" pageNumber="349">x SD</th>
<th id="76A55DB4FF8B006413E0FECFFA5EFEA0" box="[1241,1430,276,298]" gridcol="4" gridrow="0" pageId="2" pageNumber="349">%</th>
</tr>
<tr id="357434C8FF8B006417A6FEE6FA5EFED9" box="[159,1430,317,339]" gridrow="1" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FEE6FE66FED9" box="[159,430,317,339]" gridcol="0" gridrow="1" pageId="2" pageNumber="349">
Total length 
<emphasis id="B930EA98FF8BFF991627FEE6FE98FED9" bold="true" box="[286,336,317,339]" pageId="2" pageNumber="349">(TL)</emphasis>
</th>
<td id="76A55DB4FF8B00641502FEE6FD01FED9" box="[571,713,317,339]" gridcol="1" gridrow="1" pageId="2" pageNumber="349">516 ±68.1</td>
<td id="76A55DB4FF8B00641420FEE6FCB6FED9" box="[793,894,317,339]" gridcol="2" gridrow="1" pageId="2" pageNumber="349">100</td>
<td id="76A55DB4FF8B006414C0FEE6FB41FED9" box="[1017,1161,317,339]" gridcol="3" gridrow="1" pageId="2" pageNumber="349">354.4 ±78.7</td>
<td id="76A55DB4FF8B006413E0FEE6FA5EFED9" box="[1241,1430,317,339]" gridcol="4" gridrow="1" pageId="2" pageNumber="349">100</td>
</tr>
<tr id="357434C8FF8B006417A6FEBDFA5EFEF6" box="[159,1430,358,380]" gridrow="2" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FEBDFE66FEF6" box="[159,430,358,380]" gridcol="0" gridrow="2" pageId="2" pageNumber="349">Precaudal length</th>
<td id="76A55DB4FF8B00641502FEBDFD01FEF6" box="[571,713,358,380]" gridcol="1" gridrow="2" pageId="2" pageNumber="349">393.7± 55.7</td>
<td id="76A55DB4FF8B00641420FEBDFCB6FEF6" box="[793,894,358,380]" gridcol="2" gridrow="2" pageId="2" pageNumber="349">74.7–76.8</td>
<td id="76A55DB4FF8B006414C0FEBDFB41FEF6" box="[1017,1161,358,380]" gridcol="3" gridrow="2" pageId="2" pageNumber="349">275.6 ±63.5</td>
<td id="76A55DB4FF8B006413E0FEBDFA5EFEF6" box="[1241,1430,358,380]" gridcol="4" gridrow="2" pageId="2" pageNumber="349">76.5–77.9</td>
</tr>
<tr id="357434C8FF8B006417A6FE4BFA5EFE2C" box="[159,1430,400,422]" gridrow="3" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FE4BFE66FE2C" box="[159,430,400,422]" gridcol="0" gridrow="3" pageId="2" pageNumber="349">Eye-spiracle length</th>
<td id="76A55DB4FF8B00641502FE4BFD01FE2C" box="[571,713,400,422]" gridcol="1" gridrow="3" pageId="2" pageNumber="349">6.1 ± 1.3</td>
<td id="76A55DB4FF8B00641420FE4BFCB6FE2C" box="[793,894,400,422]" gridcol="2" gridrow="3" pageId="2" pageNumber="349">0.9–1.3</td>
<td id="76A55DB4FF8B006414C0FE4BFB41FE2C" box="[1017,1161,400,422]" gridcol="3" gridrow="3" pageId="2" pageNumber="349">3.3 ±1.2</td>
<td id="76A55DB4FF8B006413E0FE4BFA5EFE2C" box="[1241,1430,400,422]" gridcol="4" gridrow="3" pageId="2" pageNumber="349">0.6–1.3</td>
</tr>
<tr id="357434C8FF8B006417A6FE62FA5EFE45" box="[159,1430,441,463]" gridrow="4" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FE62FE66FE45" box="[159,430,441,463]" gridcol="0" gridrow="4" pageId="2" pageNumber="349">Preoral length</th>
<td id="76A55DB4FF8B00641502FE62FD01FE45" box="[571,713,441,463]" gridcol="1" gridrow="4" pageId="2" pageNumber="349">23.8 ±4.7</td>
<td id="76A55DB4FF8B00641420FE62FCB6FE45" box="[793,894,441,463]" gridcol="2" gridrow="4" pageId="2" pageNumber="349">4.3–5.6</td>
<td id="76A55DB4FF8B006414C0FE62FB41FE45" box="[1017,1161,441,463]" gridcol="3" gridrow="4" pageId="2" pageNumber="349">16.9 ±3.2</td>
<td id="76A55DB4FF8B006413E0FE62FA5EFE45" box="[1241,1430,441,463]" gridcol="4" gridrow="4" pageId="2" pageNumber="349">4.3–4.7</td>
</tr>
<tr id="357434C8FF8B006417A6FE39FA5EFE72" box="[159,1430,482,504]" gridrow="5" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FE39FE66FE72" box="[159,430,482,504]" gridcol="0" gridrow="5" pageId="2" pageNumber="349">Preorbital length</th>
<td id="76A55DB4FF8B00641502FE39FD01FE72" box="[571,713,482,504]" gridcol="1" gridrow="5" pageId="2" pageNumber="349">32.6 ±3.8</td>
<td id="76A55DB4FF8B00641420FE39FCB6FE72" box="[793,894,482,504]" gridcol="2" gridrow="5" pageId="2" pageNumber="349">6.4</td>
<td id="76A55DB4FF8B006414C0FE39FB41FE72" box="[1017,1161,482,504]" gridcol="3" gridrow="5" pageId="2" pageNumber="349">23.2 ±4.5</td>
<td id="76A55DB4FF8B006413E0FE39FA5EFE72" box="[1241,1430,482,504]" gridcol="4" gridrow="5" pageId="2" pageNumber="349">6.2–8.0</td>
</tr>
<tr id="357434C8FF8B006417A6FDD7FA5EFDA8" box="[159,1430,524,546]" gridrow="6" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FDD7FE66FDA8" box="[159,430,524,546]" gridcol="0" gridrow="6" pageId="2" pageNumber="349">Prespiracular length</th>
<td id="76A55DB4FF8B00641502FDD7FD01FDA8" box="[571,713,524,546]" gridcol="1" gridrow="6" pageId="2" pageNumber="349">53.1 ±5.9</td>
<td id="76A55DB4FF8B00641420FDD7FCB6FDA8" box="[793,894,524,546]" gridcol="2" gridrow="6" pageId="2" pageNumber="349">10.4–10.7</td>
<td id="76A55DB4FF8B006414C0FDD7FB41FDA8" box="[1017,1161,524,546]" gridcol="3" gridrow="6" pageId="2" pageNumber="349">37.6 ±7.7</td>
<td id="76A55DB4FF8B006413E0FDD7FA5EFDA8" box="[1241,1430,524,546]" gridcol="4" gridrow="6" pageId="2" pageNumber="349">10.6–10.7</td>
</tr>
<tr id="357434C8FF8B006417A6FDEEFA5EFDC1" box="[159,1430,565,587]" gridrow="7" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FDEEFE66FDC1" box="[159,430,565,587]" gridcol="0" gridrow="7" pageId="2" pageNumber="349">Prebranchial length</th>
<td id="76A55DB4FF8B00641502FDEEFD01FDC1" box="[571,713,565,587]" gridcol="1" gridrow="7" pageId="2" pageNumber="349">76.6 ±11.4</td>
<td id="76A55DB4FF8B00641420FDEEFCB6FDC1" box="[793,894,565,587]" gridcol="2" gridrow="7" pageId="2" pageNumber="349">14.5–15.2</td>
<td id="76A55DB4FF8B006414C0FDEEFB41FDC1" box="[1017,1161,565,587]" gridcol="3" gridrow="7" pageId="2" pageNumber="349">54.2±11.9</td>
<td id="76A55DB4FF8B006413E0FDEEFA5EFDC1" box="[1241,1430,565,587]" gridcol="4" gridrow="7" pageId="2" pageNumber="349">14.8–15.4</td>
</tr>
<tr id="357434C8FF8B006417A6FD85FA5EFDFE" box="[159,1430,606,628]" gridrow="8" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FD85FE66FDFE" box="[159,430,606,628]" gridcol="0" gridrow="8" pageId="2" pageNumber="349">Head length</th>
<td id="76A55DB4FF8B00641502FD85FD01FDFE" box="[571,713,606,628]" gridcol="1" gridrow="8" pageId="2" pageNumber="349">103.2 ±14.3</td>
<td id="76A55DB4FF8B00641420FD85FCB6FDFE" box="[793,894,606,628]" gridcol="2" gridrow="8" pageId="2" pageNumber="349">19.5–20.3</td>
<td id="76A55DB4FF8B006414C0FD85FB41FDFE" box="[1017,1161,606,628]" gridcol="3" gridrow="8" pageId="2" pageNumber="349">71 ±16.1</td>
<td id="76A55DB4FF8B006413E0FD85FA5EFDFE" box="[1241,1430,606,628]" gridcol="4" gridrow="8" pageId="2" pageNumber="349">18.7–19.2</td>
</tr>
<tr id="357434C8FF8B006417A6FD53FA5EFD14" box="[159,1430,648,670]" gridrow="9" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FD53FE66FD14" box="[159,430,648,670]" gridcol="0" gridrow="9" pageId="2" pageNumber="349">Prepectoral length</th>
<td id="76A55DB4FF8B00641502FD53FD01FD14" box="[571,713,648,670]" gridcol="1" gridrow="9" pageId="2" pageNumber="349">94.0 ±12.1</td>
<td id="76A55DB4FF8B00641420FD53FCB6FD14" box="[793,894,648,670]" gridcol="2" gridrow="9" pageId="2" pageNumber="349">17.9</td>
<td id="76A55DB4FF8B006414C0FD53FB41FD14" box="[1017,1161,648,670]" gridcol="3" gridrow="9" pageId="2" pageNumber="349">66 ±15.3</td>
<td id="76A55DB4FF8B006413E0FD53FA5EFD14" box="[1241,1430,648,670]" gridcol="4" gridrow="9" pageId="2" pageNumber="349">16.1–18.2</td>
</tr>
<tr id="357434C8FF8B006417A6FD6AFA5EFD4D" box="[159,1430,689,711]" gridrow="10" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FD6AFE66FD4D" box="[159,430,689,711]" gridcol="0" gridrow="10" pageId="2" pageNumber="349">Prepelvic length</th>
<td id="76A55DB4FF8B00641502FD6AFD01FD4D" box="[571,713,689,711]" gridcol="1" gridrow="10" pageId="2" pageNumber="349">217.5±33.8</td>
<td id="76A55DB4FF8B00641420FD6AFCB6FD4D" box="[793,894,689,711]" gridcol="2" gridrow="10" pageId="2" pageNumber="349">41.0–43.4</td>
<td id="76A55DB4FF8B006414C0FD6AFB41FD4D" box="[1017,1161,689,711]" gridcol="3" gridrow="10" pageId="2" pageNumber="349">148 ±33.5</td>
<td id="76A55DB4FF8B006413E0FD6AFA5EFD4D" box="[1241,1430,689,711]" gridcol="4" gridrow="10" pageId="2" pageNumber="349">38.2–43.7</td>
</tr>
<tr id="357434C8FF8B006417A6FD01FA5EFD7A" box="[159,1430,730,752]" gridrow="11" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FD01FE66FD7A" box="[159,430,730,752]" gridcol="0" gridrow="11" pageId="2" pageNumber="349">Snout-vent length</th>
<td id="76A55DB4FF8B00641502FD01FD01FD7A" box="[571,713,730,752]" gridcol="1" gridrow="11" pageId="2" pageNumber="349">232.2±34.9</td>
<td id="76A55DB4FF8B00641420FD01FCB6FD7A" box="[793,894,730,752]" gridcol="2" gridrow="11" pageId="2" pageNumber="349">43.6–45.4</td>
<td id="76A55DB4FF8B006414C0FD01FB41FD7A" box="[1017,1161,730,752]" gridcol="3" gridrow="11" pageId="2" pageNumber="349">152 ±33.9</td>
<td id="76A55DB4FF8B006413E0FD01FA5EFD7A" box="[1241,1430,730,752]" gridcol="4" gridrow="11" pageId="2" pageNumber="349">40.9–42.3</td>
</tr>
<tr id="357434C8FF8B006417A6FCDFFA5EFC90" box="[159,1430,772,794]" gridrow="12" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FCDFFE66FC90" box="[159,430,772,794]" gridcol="0" gridrow="12" pageId="2" pageNumber="349">Vent-caudal length</th>
<td id="76A55DB4FF8B00641502FCDFFD01FC90" box="[571,713,772,794]" gridcol="1" gridrow="12" pageId="2" pageNumber="349">262.5 ±43.4</td>
<td id="76A55DB4FF8B00641420FCDFFCB6FC90" box="[793,894,772,794]" gridcol="2" gridrow="12" pageId="2" pageNumber="349">53.7–55.3</td>
<td id="76A55DB4FF8B006414C0FCDFFB41FC90" box="[1017,1161,772,794]" gridcol="3" gridrow="12" pageId="2" pageNumber="349">189.6 ±40.9</td>
<td id="76A55DB4FF8B006413E0FCDFFA5EFC90" box="[1241,1430,772,794]" gridcol="4" gridrow="12" pageId="2" pageNumber="349">51.3–58.3</td>
</tr>
<tr id="357434C8FF8B006417A6FCF6FA5EFCC9" box="[159,1430,813,835]" gridrow="13" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FCF6FE66FCC9" box="[159,430,813,835]" gridcol="0" gridrow="13" pageId="2" pageNumber="349">Preanal length</th>
<td id="76A55DB4FF8B00641502FCF6FD01FCC9" box="[571,713,813,835]" gridcol="1" gridrow="13" pageId="2" pageNumber="349">314.1 ±50.3</td>
<td id="76A55DB4FF8B00641420FCF6FCB6FCC9" box="[793,894,813,835]" gridcol="2" gridrow="13" pageId="2" pageNumber="349">59.1–64.4</td>
<td id="76A55DB4FF8B006414C0FCF6FB41FCC9" box="[1017,1161,813,835]" gridcol="3" gridrow="13" pageId="2" pageNumber="349">213.2 ±51.4</td>
<td id="76A55DB4FF8B006413E0FCF6FA5EFCC9" box="[1241,1430,813,835]" gridcol="4" gridrow="13" pageId="2" pageNumber="349">55–62.7</td>
</tr>
<tr id="357434C8FF8B006417A6FC8DFA5EFCE6" box="[159,1430,854,876]" gridrow="14" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FC8DFE66FCE6" box="[159,430,854,876]" gridcol="0" gridrow="14" pageId="2" pageNumber="349">Pre-first dorsal length</th>
<td id="76A55DB4FF8B00641502FC8DFD01FCE6" box="[571,713,854,876]" gridcol="1" gridrow="14" pageId="2" pageNumber="349">257.5 ±38.2</td>
<td id="76A55DB4FF8B00641420FC8DFCB6FCE6" box="[793,894,854,876]" gridcol="2" gridrow="14" pageId="2" pageNumber="349">48.4–51.2</td>
<td id="76A55DB4FF8B006414C0FC8DFB41FCE6" box="[1017,1161,854,876]" gridcol="3" gridrow="14" pageId="2" pageNumber="349">177 ±41.3</td>
<td id="76A55DB4FF8B006413E0FC8DFA5EFCE6" box="[1241,1430,854,876]" gridcol="4" gridrow="14" pageId="2" pageNumber="349">44.9–51.3</td>
</tr>
<tr id="357434C8FF8B006417A6FC5BFA5EFC1C" box="[159,1430,896,918]" gridrow="15" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FC5BFE66FC1C" box="[159,430,896,918]" gridcol="0" gridrow="15" pageId="2" pageNumber="349">Interdorsal space</th>
<td id="76A55DB4FF8B00641502FC5BFD01FC1C" box="[571,713,896,918]" gridcol="1" gridrow="15" pageId="2" pageNumber="349">57.2 ±9.5</td>
<td id="76A55DB4FF8B00641420FC5BFCB6FC1C" box="[793,894,896,918]" gridcol="2" gridrow="15" pageId="2" pageNumber="349">10.9–11.0</td>
<td id="76A55DB4FF8B006414C0FC5BFB41FC1C" box="[1017,1161,896,918]" gridcol="3" gridrow="15" pageId="2" pageNumber="349">40.4 ±13.5</td>
<td id="76A55DB4FF8B006413E0FC5BFA5EFC1C" box="[1241,1430,896,918]" gridcol="4" gridrow="15" pageId="2" pageNumber="349">7.0–10.7</td>
</tr>
<tr id="357434C8FF8B006417A6FC72FA5EFC35" box="[159,1430,937,959]" gridrow="16" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FC72FE66FC35" box="[159,430,937,959]" gridcol="0" gridrow="16" pageId="2" pageNumber="349">Dorsal-caudal space</th>
<td id="76A55DB4FF8B00641502FC72FD01FC35" box="[571,713,937,959]" gridcol="1" gridrow="16" pageId="2" pageNumber="349">22.3 ±4.7</td>
<td id="76A55DB4FF8B00641420FC72FCB6FC35" box="[793,894,937,959]" gridcol="2" gridrow="16" pageId="2" pageNumber="349">4.2– 5.1</td>
<td id="76A55DB4FF8B006414C0FC72FB41FC35" box="[1017,1161,937,959]" gridcol="3" gridrow="16" pageId="2" pageNumber="349">18.2 ±3.9</td>
<td id="76A55DB4FF8B006413E0FC72FA5EFC35" box="[1241,1430,937,959]" gridcol="4" gridrow="16" pageId="2" pageNumber="349">5.8–6.7</td>
</tr>
<tr id="357434C8FF8B006417A6FC09FA5EFC62" box="[159,1430,978,1000]" gridrow="17" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FC09FE66FC62" box="[159,430,978,1000]" gridcol="0" gridrow="17" pageId="2" pageNumber="349">Pectoral-pelvic space</th>
<td id="76A55DB4FF8B00641502FC09FD01FC62" box="[571,713,978,1000]" gridcol="1" gridrow="17" pageId="2" pageNumber="349">96.3 ±16.4</td>
<td id="76A55DB4FF8B00641420FC09FCB6FC62" box="[793,894,978,1000]" gridcol="2" gridrow="17" pageId="2" pageNumber="349">17.6–20.1</td>
<td id="76A55DB4FF8B006414C0FC09FB41FC62" box="[1017,1161,978,1000]" gridcol="3" gridrow="17" pageId="2" pageNumber="349">63.5 ±15.9</td>
<td id="76A55DB4FF8B006413E0FC09FA5EFC62" box="[1241,1430,978,1000]" gridcol="4" gridrow="17" pageId="2" pageNumber="349">10–20.3</td>
</tr>
<tr id="357434C8FF8B006417A6FC27FA5EFB98" box="[159,1430,1020,1042]" gridrow="18" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FC27FE66FB98" box="[159,430,1020,1042]" gridcol="0" gridrow="18" pageId="2" pageNumber="349">Pelvic-anal space</th>
<td id="76A55DB4FF8B00641502FC27FD01FB98" box="[571,713,1020,1042]" gridcol="1" gridrow="18" pageId="2" pageNumber="349">59.7 ±13.3</td>
<td id="76A55DB4FF8B00641420FC27FCB6FB98" box="[793,894,1020,1042]" gridcol="2" gridrow="18" pageId="2" pageNumber="349">10.5–12.8</td>
<td id="76A55DB4FF8B006414C0FC27FB41FB98" box="[1017,1161,1020,1042]" gridcol="3" gridrow="18" pageId="2" pageNumber="349">44.1 ±12.3</td>
<td id="76A55DB4FF8B006413E0FC27FA5EFB98" box="[1241,1430,1020,1042]" gridcol="4" gridrow="18" pageId="2" pageNumber="349">12.7–14.6</td>
</tr>
<tr id="357434C8FF8B006417A6FBFEFA5EFBB1" box="[159,1430,1061,1083]" gridrow="19" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FBFEFE66FBB1" box="[159,430,1061,1083]" gridcol="0" gridrow="19" pageId="2" pageNumber="349">Anal-caudal space</th>
<td id="76A55DB4FF8B00641502FBFEFD01FBB1" box="[571,713,1061,1083]" gridcol="1" gridrow="19" pageId="2" pageNumber="349">39.2 ±5.6</td>
<td id="76A55DB4FF8B00641420FBFEFCB6FBB1" box="[793,894,1061,1083]" gridcol="2" gridrow="19" pageId="2" pageNumber="349">7.4–7.6</td>
<td id="76A55DB4FF8B006414C0FBFEFB41FBB1" box="[1017,1161,1061,1083]" gridcol="3" gridrow="19" pageId="2" pageNumber="349">28.5 ±6.5</td>
<td id="76A55DB4FF8B006413E0FBFEFA5EFBB1" box="[1241,1430,1061,1083]" gridcol="4" gridrow="19" pageId="2" pageNumber="349">8.7–8.9</td>
</tr>
<tr id="357434C8FF8B006417A6FB95FA5EFBEE" box="[159,1430,1102,1124]" gridrow="20" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FB95FE66FBEE" box="[159,430,1102,1124]" gridcol="0" gridrow="20" pageId="2" pageNumber="349">Eye length</th>
<td id="76A55DB4FF8B00641502FB95FD01FBEE" box="[571,713,1102,1124]" gridcol="1" gridrow="20" pageId="2" pageNumber="349">19.8 ±2.8</td>
<td id="76A55DB4FF8B00641420FB95FCB6FBEE" box="[793,894,1102,1124]" gridcol="2" gridrow="20" pageId="2" pageNumber="349">3.8–4.0</td>
<td id="76A55DB4FF8B006414C0FB95FB41FBEE" box="[1017,1161,1102,1124]" gridcol="3" gridrow="20" pageId="2" pageNumber="349">13.6 ±3.4</td>
<td id="76A55DB4FF8B006413E0FB95FA5EFBEE" box="[1241,1430,1102,1124]" gridcol="4" gridrow="20" pageId="2" pageNumber="349">3.3–4.3</td>
</tr>
<tr id="357434C8FF8B006417A6FBA3FA5EFB04" box="[159,1430,1144,1166]" gridrow="21" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FBA3FE66FB04" box="[159,430,1144,1166]" gridcol="0" gridrow="21" pageId="2" pageNumber="349">Eye height</th>
<td id="76A55DB4FF8B00641502FBA3FD01FB04" box="[571,713,1144,1166]" gridcol="1" gridrow="21" pageId="2" pageNumber="349">3.8 ±0.3</td>
<td id="76A55DB4FF8B00641420FBA3FCB6FB04" box="[793,894,1144,1166]" gridcol="2" gridrow="21" pageId="2" pageNumber="349">0.9–1.4</td>
<td id="76A55DB4FF8B006414C0FBA3FB41FB04" box="[1017,1161,1144,1166]" gridcol="3" gridrow="21" pageId="2" pageNumber="349">4.7 ±1.1</td>
<td id="76A55DB4FF8B006413E0FBA3FA5EFB04" box="[1241,1430,1144,1166]" gridcol="4" gridrow="21" pageId="2" pageNumber="349">1.3–1.5</td>
</tr>
<tr id="357434C8FF8B006417A6FB7AFA5EFB3D" box="[159,1430,1185,1207]" gridrow="22" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FB7AFE66FB3D" box="[159,430,1185,1207]" gridcol="0" gridrow="22" pageId="2" pageNumber="349">Internarial space</th>
<td id="76A55DB4FF8B00641502FB7AFD01FB3D" box="[571,713,1185,1207]" gridcol="1" gridrow="22" pageId="2" pageNumber="349">10.1 ±3.6</td>
<td id="76A55DB4FF8B00641420FB7AFCB6FB3D" box="[793,894,1185,1207]" gridcol="2" gridrow="22" pageId="2" pageNumber="349">1.4–2.4</td>
<td id="76A55DB4FF8B006414C0FB7AFB41FB3D" box="[1017,1161,1185,1207]" gridcol="3" gridrow="22" pageId="2" pageNumber="349">9 ±3.4</td>
<td id="76A55DB4FF8B006413E0FB7AFA5EFB3D" box="[1241,1430,1185,1207]" gridcol="4" gridrow="22" pageId="2" pageNumber="349">2.0–4.0</td>
</tr>
<tr id="357434C8FF8B006417A6FB11FA5EFB6A" box="[159,1430,1226,1248]" gridrow="23" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FB11FE66FB6A" box="[159,430,1226,1248]" gridcol="0" gridrow="23" pageId="2" pageNumber="349">Interorbital space</th>
<td id="76A55DB4FF8B00641502FB11FD01FB6A" box="[571,713,1226,1248]" gridcol="1" gridrow="23" pageId="2" pageNumber="349">31.1 ±3.1</td>
<td id="76A55DB4FF8B00641420FB11FCB6FB6A" box="[793,894,1226,1248]" gridcol="2" gridrow="23" pageId="2" pageNumber="349">6.1–6.6</td>
<td id="76A55DB4FF8B006414C0FB11FB41FB6A" box="[1017,1161,1226,1248]" gridcol="3" gridrow="23" pageId="2" pageNumber="349">24.3 ±4.8</td>
<td id="76A55DB4FF8B006413E0FB11FA5EFB6A" box="[1241,1430,1226,1248]" gridcol="4" gridrow="23" pageId="2" pageNumber="349">6.4–7.3</td>
</tr>
<tr id="357434C8FF8B006417A6FB2FFA5EFA80" box="[159,1430,1268,1290]" gridrow="24" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FB2FFE66FA80" box="[159,430,1268,1290]" gridcol="0" gridrow="24" pageId="2" pageNumber="349">Spiracle length</th>
<td id="76A55DB4FF8B00641502FB2FFD01FA80" box="[571,713,1268,1290]" gridcol="1" gridrow="24" pageId="2" pageNumber="349">4.3 ±1.1</td>
<td id="76A55DB4FF8B00641420FB2FFCB6FA80" box="[793,894,1268,1290]" gridcol="2" gridrow="24" pageId="2" pageNumber="349">0.4–0.9</td>
<td id="76A55DB4FF8B006414C0FB2FFB41FA80" box="[1017,1161,1268,1290]" gridcol="3" gridrow="24" pageId="2" pageNumber="349">2.6 ±1.1</td>
<td id="76A55DB4FF8B006413E0FB2FFA5EFA80" box="[1241,1430,1268,1290]" gridcol="4" gridrow="24" pageId="2" pageNumber="349">0.6–1.3</td>
</tr>
<tr id="357434C8FF8B006417A6FAC6FA5EFAB9" box="[159,1430,1309,1331]" gridrow="25" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FAC6FE66FAB9" box="[159,430,1309,1331]" gridcol="0" gridrow="25" pageId="2" pageNumber="349">Mouth length</th>
<td id="76A55DB4FF8B00641502FAC6FD01FAB9" box="[571,713,1309,1331]" gridcol="1" gridrow="25" pageId="2" pageNumber="349">23.5 ±5.0</td>
<td id="76A55DB4FF8B00641420FAC6FCB6FAB9" box="[793,894,1309,1331]" gridcol="2" gridrow="25" pageId="2" pageNumber="349">3.7–5. 1</td>
<td id="76A55DB4FF8B006414C0FAC6FB41FAB9" box="[1017,1161,1309,1331]" gridcol="3" gridrow="25" pageId="2" pageNumber="349">14.9 ±4.8</td>
<td id="76A55DB4FF8B006413E0FAC6FA5EFAB9" box="[1241,1430,1309,1331]" gridcol="4" gridrow="25" pageId="2" pageNumber="349">4.7–5.6</td>
</tr>
<tr id="357434C8FF8B006417A6FA9DFA5EFAD6" box="[159,1430,1350,1372]" gridrow="26" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FA9DFE66FAD6" box="[159,430,1350,1372]" gridcol="0" gridrow="26" pageId="2" pageNumber="349">Mouth width</th>
<td id="76A55DB4FF8B00641502FA9DFD01FAD6" box="[571,713,1350,1372]" gridcol="1" gridrow="26" pageId="2" pageNumber="349">46.6 ±7.4</td>
<td id="76A55DB4FF8B00641420FA9DFCB6FAD6" box="[793,894,1350,1372]" gridcol="2" gridrow="26" pageId="2" pageNumber="349">9.0–9.7</td>
<td id="76A55DB4FF8B006414C0FA9DFB41FAD6" box="[1017,1161,1350,1372]" gridcol="3" gridrow="26" pageId="2" pageNumber="349">27.8 ±6.4</td>
<td id="76A55DB4FF8B006413E0FA9DFA5EFAD6" box="[1241,1430,1350,1372]" gridcol="4" gridrow="26" pageId="2" pageNumber="349">8.0</td>
</tr>
<tr id="357434C8FF8B006417A6FAABFA5EFA0C" box="[159,1430,1392,1414]" gridrow="27" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FAABFE66FA0C" box="[159,430,1392,1414]" gridcol="0" gridrow="27" pageId="2" pageNumber="349">Lower labial furrow length</th>
<td id="76A55DB4FF8B00641502FAABFD01FA0C" box="[571,713,1392,1414]" gridcol="1" gridrow="27" pageId="2" pageNumber="349">10.6 ±1.9</td>
<td id="76A55DB4FF8B00641420FAABFCB6FA0C" box="[793,894,1392,1414]" gridcol="2" gridrow="27" pageId="2" pageNumber="349">1.9–2.1</td>
<td id="76A55DB4FF8B006414C0FAABFB41FA0C" box="[1017,1161,1392,1414]" gridcol="3" gridrow="27" pageId="2" pageNumber="349">6.6 ±1.8</td>
<td id="76A55DB4FF8B006413E0FAABFA5EFA0C" box="[1241,1430,1392,1414]" gridcol="4" gridrow="27" pageId="2" pageNumber="349">1.3–2.0</td>
</tr>
<tr id="357434C8FF8B006417A6FA42FA5EFA25" box="[159,1430,1433,1455]" gridrow="28" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FA42FE66FA25" box="[159,430,1433,1455]" gridcol="0" gridrow="28" pageId="2" pageNumber="349">First gill slit height</th>
<td id="76A55DB4FF8B00641502FA42FD01FA25" box="[571,713,1433,1455]" gridcol="1" gridrow="28" pageId="2" pageNumber="349">15.2 ±2.7</td>
<td id="76A55DB4FF8B00641420FA42FCB6FA25" box="[793,894,1433,1455]" gridcol="2" gridrow="28" pageId="2" pageNumber="349">2.8–3.1</td>
<td id="76A55DB4FF8B006414C0FA42FB41FA25" box="[1017,1161,1433,1455]" gridcol="3" gridrow="28" pageId="2" pageNumber="349">9 ±2.3</td>
<td id="76A55DB4FF8B006413E0FA42FA5EFA25" box="[1241,1430,1433,1455]" gridcol="4" gridrow="28" pageId="2" pageNumber="349">2.0–2.8</td>
</tr>
<tr id="357434C8FF8B006417A6FA19FA5EFA52" box="[159,1430,1474,1496]" gridrow="29" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FA19FE66FA52" box="[159,430,1474,1496]" gridcol="0" gridrow="29" pageId="2" pageNumber="349">Fifth gill slit height</th>
<td id="76A55DB4FF8B00641502FA19FD01FA52" box="[571,713,1474,1496]" gridcol="1" gridrow="29" pageId="2" pageNumber="349">8.6 ±2.3</td>
<td id="76A55DB4FF8B00641420FA19FCB6FA52" box="[793,894,1474,1496]" gridcol="2" gridrow="29" pageId="2" pageNumber="349">0.9–1.8</td>
<td id="76A55DB4FF8B006414C0FA19FB41FA52" box="[1017,1161,1474,1496]" gridcol="3" gridrow="29" pageId="2" pageNumber="349">4.7 ±1.3</td>
<td id="76A55DB4FF8B006413E0FA19FA5EFA52" box="[1241,1430,1474,1496]" gridcol="4" gridrow="29" pageId="2" pageNumber="349">1.3–1.9</td>
</tr>
<tr id="357434C8FF8B006417A6FA37FA5EF988" box="[159,1430,1516,1538]" gridrow="30" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6FA37FE66F988" box="[159,430,1516,1538]" gridcol="0" gridrow="30" pageId="2" pageNumber="349">Pectoral length</th>
<td id="76A55DB4FF8B00641502FA37FD01F988" box="[571,713,1516,1538]" gridcol="1" gridrow="30" pageId="2" pageNumber="349">68.5 ±7.4</td>
<td id="76A55DB4FF8B00641420FA37FCB6F988" box="[793,894,1516,1538]" gridcol="2" gridrow="30" pageId="2" pageNumber="349">12.8–13.6</td>
<td id="76A55DB4FF8B006414C0FA37FB41F988" box="[1017,1161,1516,1538]" gridcol="3" gridrow="30" pageId="2" pageNumber="349">41 ±11.4</td>
<td id="76A55DB4FF8B006413E0FA37FA5EF988" box="[1241,1430,1516,1538]" gridcol="4" gridrow="30" pageId="2" pageNumber="349">10.7–13.8</td>
</tr>
<tr id="357434C8FF8B006417A6F9CEFA5EF9A1" box="[159,1430,1557,1579]" gridrow="31" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6F9CEFE66F9A1" box="[159,430,1557,1579]" gridcol="0" gridrow="31" pageId="2" pageNumber="349">Pectoral anterior margin</th>
<td id="76A55DB4FF8B00641502F9CEFD01F9A1" box="[571,713,1557,1579]" gridcol="1" gridrow="31" pageId="2" pageNumber="349">74.6 ±9.0</td>
<td id="76A55DB4FF8B00641420F9CEFCB6F9A1" box="[793,894,1557,1579]" gridcol="2" gridrow="31" pageId="2" pageNumber="349">14.5–15.0</td>
<td id="76A55DB4FF8B006414C0F9CEFB41F9A1" box="[1017,1161,1557,1579]" gridcol="3" gridrow="31" pageId="2" pageNumber="349">47.5 ±11.9</td>
<td id="76A55DB4FF8B006413E0F9CEFA5EF9A1" box="[1241,1430,1557,1579]" gridcol="4" gridrow="31" pageId="2" pageNumber="349">12.7–15.6</td>
</tr>
<tr id="357434C8FF8B006417A6F9E5FA5EF9DE" box="[159,1430,1598,1620]" gridrow="32" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6F9E5FE66F9DE" box="[159,430,1598,1620]" gridcol="0" gridrow="32" pageId="2" pageNumber="349">Pectoral base</th>
<td id="76A55DB4FF8B00641502F9E5FD01F9DE" box="[571,713,1598,1620]" gridcol="1" gridrow="32" pageId="2" pageNumber="349">34.1 ±10.3</td>
<td id="76A55DB4FF8B00641420F9E5FCB6F9DE" box="[793,894,1598,1620]" gridcol="2" gridrow="32" pageId="2" pageNumber="349">3.1–7.2</td>
<td id="76A55DB4FF8B006414C0F9E5FB41F9DE" box="[1017,1161,1598,1620]" gridcol="3" gridrow="32" pageId="2" pageNumber="349">24.3 ±7.1</td>
<td id="76A55DB4FF8B006413E0F9E5FA5EF9DE" box="[1241,1430,1598,1620]" gridcol="4" gridrow="32" pageId="2" pageNumber="349">6.7–9.7</td>
</tr>
<tr id="357434C8FF8B006417A6F9B3FA5EF9F4" box="[159,1430,1640,1662]" gridrow="33" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6F9B3FE66F9F4" box="[159,430,1640,1662]" gridcol="0" gridrow="33" pageId="2" pageNumber="349">Pectoral posterior margin</th>
<td id="76A55DB4FF8B00641502F9B3FD01F9F4" box="[571,713,1640,1662]" gridcol="1" gridrow="33" pageId="2" pageNumber="349">45.6 ±6.3</td>
<td id="76A55DB4FF8B00641420F9B3FCB6F9F4" box="[793,894,1640,1662]" gridcol="2" gridrow="33" pageId="2" pageNumber="349">8.7–9.0</td>
<td id="76A55DB4FF8B006414C0F9B3FB41F9F4" box="[1017,1161,1640,1662]" gridcol="3" gridrow="33" pageId="2" pageNumber="349">30 ±8.8</td>
<td id="76A55DB4FF8B006413E0F9B3FA5EF9F4" box="[1241,1430,1640,1662]" gridcol="4" gridrow="33" pageId="2" pageNumber="349">5.3–8.9</td>
</tr>
<tr id="357434C8FF8B006417A6F94AFA5EF92D" box="[159,1430,1681,1703]" gridrow="34" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6F94AFE66F92D" box="[159,430,1681,1703]" gridcol="0" gridrow="34" pageId="2" pageNumber="349">Pectoral inner margin</th>
<td id="76A55DB4FF8B00641502F94AFD01F92D" box="[571,713,1681,1703]" gridcol="1" gridrow="34" pageId="2" pageNumber="349">32.2 ±2.3</td>
<td id="76A55DB4FF8B00641420F94AFCB6F92D" box="[793,894,1681,1703]" gridcol="2" gridrow="34" pageId="2" pageNumber="349">5.9–7.1</td>
<td id="76A55DB4FF8B006414C0F94AFB41F92D" box="[1017,1161,1681,1703]" gridcol="3" gridrow="34" pageId="2" pageNumber="349">21.9 ±5.8</td>
<td id="76A55DB4FF8B006413E0F94AFA5EF92D" box="[1241,1430,1681,1703]" gridcol="4" gridrow="34" pageId="2" pageNumber="349">5.3–7.6</td>
</tr>
<tr id="357434C8FF8B006417A6F961FA5EF95A" box="[159,1430,1722,1744]" gridrow="35" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6F961FE66F95A" box="[159,430,1722,1744]" gridcol="0" gridrow="35" pageId="2" pageNumber="349">Pelvic length</th>
<td id="76A55DB4FF8B00641502F961FD01F95A" box="[571,713,1722,1744]" gridcol="1" gridrow="35" pageId="2" pageNumber="349">54.3 ±10.1</td>
<td id="76A55DB4FF8B00641420F961FCB6F95A" box="[793,894,1722,1744]" gridcol="2" gridrow="35" pageId="2" pageNumber="349">9.0–10.5</td>
<td id="76A55DB4FF8B006414C0F961FB41F95A" box="[1017,1161,1722,1744]" gridcol="3" gridrow="35" pageId="2" pageNumber="349">37.58 ±11.4</td>
<td id="76A55DB4FF8B006413E0F961FA5EF95A" box="[1241,1430,1722,1744]" gridcol="4" gridrow="35" pageId="2" pageNumber="349">8.7–12.5</td>
</tr>
<tr id="357434C8FF8B006417A6F93FFA5EF970" box="[159,1430,1764,1786]" gridrow="36" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6F93FFE66F970" box="[159,430,1764,1786]" gridcol="0" gridrow="36" pageId="2" pageNumber="349">Pelvic inner margin</th>
<td id="76A55DB4FF8B00641502F93FFD01F970" box="[571,713,1764,1786]" gridcol="1" gridrow="36" pageId="2" pageNumber="349">18.0 ±3.1</td>
<td id="76A55DB4FF8B00641420F93FFCB6F970" box="[793,894,1764,1786]" gridcol="2" gridrow="36" pageId="2" pageNumber="349">3.1–3.8</td>
<td id="76A55DB4FF8B006414C0F93FFB41F970" box="[1017,1161,1764,1786]" gridcol="3" gridrow="36" pageId="2" pageNumber="349">16.3 ±6.7</td>
<td id="76A55DB4FF8B006413E0F93FFA5EF970" box="[1241,1430,1764,1786]" gridcol="4" gridrow="36" pageId="2" pageNumber="349">4–6.8</td>
</tr>
<tr id="357434C8FF8B006417A6F8D6FA5EF8A9" box="[159,1430,1805,1827]" gridrow="37" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6F8D6FE66F8A9" box="[159,430,1805,1827]" gridcol="0" gridrow="37" pageId="2" pageNumber="349">Pelvic base</th>
<td id="76A55DB4FF8B00641502F8D6FD01F8A9" box="[571,713,1805,1827]" gridcol="1" gridrow="37" pageId="2" pageNumber="349">38.0 ±11.0</td>
<td id="76A55DB4FF8B00641420F8D6FCB6F8A9" box="[793,894,1805,1827]" gridcol="2" gridrow="37" pageId="2" pageNumber="349">6.4–8.9</td>
<td id="76A55DB4FF8B006414C0F8D6FB41F8A9" box="[1017,1161,1805,1827]" gridcol="3" gridrow="37" pageId="2" pageNumber="349">24.4 ±6.6</td>
<td id="76A55DB4FF8B006413E0F8D6FA5EF8A9" box="[1241,1430,1805,1827]" gridcol="4" gridrow="37" pageId="2" pageNumber="349">4.7–8.0</td>
</tr>
<tr id="357434C8FF8B006417A6F8EDFA5EF8C6" box="[159,1430,1846,1868]" gridrow="38" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6F8EDFE66F8C6" box="[159,430,1846,1868]" gridcol="0" gridrow="38" pageId="2" pageNumber="349">Pelvic anterior margin</th>
<td id="76A55DB4FF8B00641502F8EDFD01F8C6" box="[571,713,1846,1868]" gridcol="1" gridrow="38" pageId="2" pageNumber="349">36.0 ±5.8</td>
<td id="76A55DB4FF8B00641420F8EDFCB6F8C6" box="[793,894,1846,1868]" gridcol="2" gridrow="38" pageId="2" pageNumber="349">6.6–7.4</td>
<td id="76A55DB4FF8B006414C0F8EDFB41F8C6" box="[1017,1161,1846,1868]" gridcol="3" gridrow="38" pageId="2" pageNumber="349">23 ±6.1</td>
<td id="76A55DB4FF8B006413E0F8EDFA5EF8C6" box="[1241,1430,1846,1868]" gridcol="4" gridrow="38" pageId="2" pageNumber="349">5.3–7.0</td>
</tr>
<tr id="357434C8FF8B006417A6F8BBFA5EF8FC" box="[159,1430,1888,1910]" gridrow="39" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6F8BBFE66F8FC" box="[159,430,1888,1910]" gridcol="0" gridrow="39" pageId="2" pageNumber="349">Pelvic posterior margin</th>
<td id="76A55DB4FF8B00641502F8BBFD01F8FC" box="[571,713,1888,1910]" gridcol="1" gridrow="39" pageId="2" pageNumber="349">33.6 ±4.8</td>
<td id="76A55DB4FF8B00641420F8BBFCB6F8FC" box="[793,894,1888,1910]" gridcol="2" gridrow="39" pageId="2" pageNumber="349">6.4–6.6</td>
<td id="76A55DB4FF8B006414C0F8BBFB41F8FC" box="[1017,1161,1888,1910]" gridcol="3" gridrow="39" pageId="2" pageNumber="349">22.7 ±7.2</td>
<td id="76A55DB4FF8B006413E0F8BBFA5EF8FC" box="[1241,1430,1888,1910]" gridcol="4" gridrow="39" pageId="2" pageNumber="349">3.2–7.4</td>
</tr>
<tr id="357434C8FF8B006417A6F852FA5EF815" box="[159,1430,1929,1951]" gridrow="40" pageId="2" pageNumber="349">
<th id="76A55DB4FF8B006417A6F852FE66F815" box="[159,430,1929,1951]" gridcol="0" gridrow="40" pageId="2" pageNumber="349">Clasper outer length</th>
<td id="76A55DB4FF8B00641502F852FD01F815" box="[571,713,1929,1951]" gridcol="1" gridrow="40" pageId="2" pageNumber="349">20.3 ±4.5</td>
<td id="76A55DB4FF8B00641420F852FCB6F815" box="[793,894,1929,1951]" gridcol="2" gridrow="40" pageId="2" pageNumber="349">3.8–4.1</td>
<td id="76A55DB4FF8B006414C0F852FB41F815" box="[1017,1161,1929,1951]" gridcol="3" gridrow="40" pageId="2" pageNumber="349">16.8 ±8.1</td>
<td id="76A55DB4FF8B006413E0F852FA5EF815" box="[1241,1430,1929,1951]" gridcol="4" gridrow="40" pageId="2" pageNumber="349">2.0–6.2</td>
</tr>
</table>
</paragraph>
<caption id="DF3B6602FF8AFF9817A6FF7BFE44FF3C" box="[159,396,160,182]" pageId="3" pageNumber="350" targetBox="[159,1342,242,1504]" targetIsTable="true" targetPageId="3">
<paragraph id="8BFB368AFF8AFF9817A6FF7BFE44FF3C" blockId="3.[159,1336,160,223]" box="[159,396,160,182]" pageId="3" pageNumber="350">
<emphasis id="B930EA98FF8AFF9817A6FF7BFEC4FF3C" bold="true" box="[159,268,160,182]" pageId="3" pageNumber="350">TABLE 1.</emphasis>
(Continued)
</paragraph>
</caption>
<paragraph id="8BFB368AFF8AFF981505FF28FAEEFA6A" pageId="3" pageNumber="350">
<table id="F944C42AFF8A006417A6FF29FAF6FA6A" box="[159,1342,242,1504]" colsContinueFrom="2.[159,1430,276,1951]" gridcols="5" gridrows="31" pageId="3" pageNumber="350">
<tr id="357434C8FF8A006417A6FF29FAF6FE82" box="[159,1342,242,264]" gridrow="0" pageId="3" pageNumber="350" rowspan-0="1">
<th id="76A55DB4FF8A00641502FF29FD01FE82" box="[571,713,242,264]" gridcol="1" gridrow="0" pageId="3" pageNumber="350">x SD</th>
<th id="76A55DB4FF8A00641420FF29FC4CFE82" box="[793,900,242,264]" gridcol="2" gridrow="0" pageId="3" pageNumber="350">%</th>
<th id="76A55DB4FF8A006414C0FF29FB41FE82" box="[1017,1161,242,264]" gridcol="3" gridrow="0" pageId="3" pageNumber="350">x SD</th>
<th id="76A55DB4FF8A006413E0FF29FAF6FE82" box="[1241,1342,242,264]" gridcol="4" gridrow="0" pageId="3" pageNumber="350">%</th>
</tr>
<tr id="357434C8FF8A006417A6FEC7FAF6FEB8" box="[159,1342,284,306]" gridrow="1" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FEC7FE05FEB8" box="[159,461,284,306]" gridcol="0" gridrow="1" pageId="3" pageNumber="350">Clasper inner length</th>
<td id="76A55DB4FF8A00641502FEC7FD01FEB8" box="[571,713,284,306]" gridcol="1" gridrow="1" pageId="3" pageNumber="350">44.5 ±14.8</td>
<td id="76A55DB4FF8A00641420FEC7FC4CFEB8" box="[793,900,284,306]" gridcol="2" gridrow="1" pageId="3" pageNumber="350">8.1–9.0</td>
<td id="76A55DB4FF8A006414C0FEC7FB41FEB8" box="[1017,1161,284,306]" gridcol="3" gridrow="1" pageId="3" pageNumber="350">35.7 ±12.6</td>
<td id="76A55DB4FF8A006413E0FEC7FAF6FEB8" box="[1241,1342,284,306]" gridcol="4" gridrow="1" pageId="3" pageNumber="350">6.0–10.4</td>
</tr>
<tr id="357434C8FF8A006417A6FE9EFAF6FED1" box="[159,1342,325,347]" gridrow="2" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FE9EFE05FED1" box="[159,461,325,347]" gridcol="0" gridrow="2" pageId="3" pageNumber="350">Clasper base width</th>
<td id="76A55DB4FF8A00641502FE9EFD01FED1" box="[571,713,325,347]" gridcol="1" gridrow="2" pageId="3" pageNumber="350">3.5 ±2.1</td>
<td id="76A55DB4FF8A00641420FE9EFC4CFED1" box="[793,900,325,347]" gridcol="2" gridrow="2" pageId="3" pageNumber="350">0.4–0.8</td>
<td id="76A55DB4FF8A006414C0FE9EFB41FED1" box="[1017,1161,325,347]" gridcol="3" gridrow="2" pageId="3" pageNumber="350">4 ±1.4</td>
<td id="76A55DB4FF8A006413E0FE9EFAF6FED1" box="[1241,1342,325,347]" gridcol="4" gridrow="2" pageId="3" pageNumber="350">0.6–1.1</td>
</tr>
<tr id="357434C8FF8A006417A6FEB5FAF6FE0E" box="[159,1342,366,388]" gridrow="3" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FEB5FE05FE0E" box="[159,461,366,388]" gridcol="0" gridrow="3" pageId="3" pageNumber="350">Anal base</th>
<td id="76A55DB4FF8A00641502FEB5FD01FE0E" box="[571,713,366,388]" gridcol="1" gridrow="3" pageId="3" pageNumber="350">39.8 ±8.0</td>
<td id="76A55DB4FF8A00641420FEB5FC4CFE0E" box="[793,900,366,388]" gridcol="2" gridrow="3" pageId="3" pageNumber="350">5.7–7.7</td>
<td id="76A55DB4FF8A006414C0FEB5FB41FE0E" box="[1017,1161,366,388]" gridcol="3" gridrow="3" pageId="3" pageNumber="350">30.2 ±6.3</td>
<td id="76A55DB4FF8A006413E0FEB5FAF6FE0E" box="[1241,1342,366,388]" gridcol="4" gridrow="3" pageId="3" pageNumber="350">8.0–8.7</td>
</tr>
<tr id="357434C8FF8A006417A6FE43FAF6FE24" box="[159,1342,408,430]" gridrow="4" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FE43FE05FE24" box="[159,461,408,430]" gridcol="0" gridrow="4" pageId="3" pageNumber="350">Anal anterior margin</th>
<td id="76A55DB4FF8A00641502FE43FD01FE24" box="[571,713,408,430]" gridcol="1" gridrow="4" pageId="3" pageNumber="350">44.1±6.3</td>
<td id="76A55DB4FF8A00641420FE43FC4CFE24" box="[793,900,408,430]" gridcol="2" gridrow="4" pageId="3" pageNumber="350">8.3–8.7</td>
<td id="76A55DB4FF8A006414C0FE43FB41FE24" box="[1017,1161,408,430]" gridcol="3" gridrow="4" pageId="3" pageNumber="350">28.1 ±6.3</td>
<td id="76A55DB4FF8A006413E0FE43FAF6FE24" box="[1241,1342,408,430]" gridcol="4" gridrow="4" pageId="3" pageNumber="350">8.0–8.7</td>
</tr>
<tr id="357434C8FF8A006417A6FE1AFAF6FE5D" box="[159,1342,449,471]" gridrow="5" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FE1AFE05FE5D" box="[159,461,449,471]" gridcol="0" gridrow="5" pageId="3" pageNumber="350">Anal posterior margin</th>
<td id="76A55DB4FF8A00641502FE1AFD01FE5D" box="[571,713,449,471]" gridcol="1" gridrow="5" pageId="3" pageNumber="350">26.8 ±3.1</td>
<td id="76A55DB4FF8A00641420FE1AFC4CFE5D" box="[793,900,449,471]" gridcol="2" gridrow="5" pageId="3" pageNumber="350">5.1–5.4</td>
<td id="76A55DB4FF8A006414C0FE1AFB41FE5D" box="[1017,1161,449,471]" gridcol="3" gridrow="5" pageId="3" pageNumber="350">17.6 ±4.9</td>
<td id="76A55DB4FF8A006413E0FE1AFAF6FE5D" box="[1241,1342,449,471]" gridcol="4" gridrow="5" pageId="3" pageNumber="350">4.7–6.0</td>
</tr>
<tr id="357434C8FF8A006417A6FE31FAF6FD8A" box="[159,1342,490,512]" gridrow="6" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FE31FE05FD8A" box="[159,461,490,512]" gridcol="0" gridrow="6" pageId="3" pageNumber="350">Anal inner margin</th>
<td id="76A55DB4FF8A00641502FE31FD01FD8A" box="[571,713,490,512]" gridcol="1" gridrow="6" pageId="3" pageNumber="350">13.5 ±2.3</td>
<td id="76A55DB4FF8A00641420FE31FC4CFD8A" box="[793,900,490,512]" gridcol="2" gridrow="6" pageId="3" pageNumber="350">2.6–2.9</td>
<td id="76A55DB4FF8A006414C0FE31FB41FD8A" box="[1017,1161,490,512]" gridcol="3" gridrow="6" pageId="3" pageNumber="350">9 ±3.7</td>
<td id="76A55DB4FF8A006413E0FE31FAF6FD8A" box="[1241,1342,490,512]" gridcol="4" gridrow="6" pageId="3" pageNumber="350">2.0–4.9</td>
</tr>
<tr id="357434C8FF8A006417A6FDCFFAF6FDA0" box="[159,1342,532,554]" gridrow="7" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FDCFFE05FDA0" box="[159,461,532,554]" gridcol="0" gridrow="7" pageId="3" pageNumber="350">Anal height</th>
<td id="76A55DB4FF8A00641502FDCFFD01FDA0" box="[571,713,532,554]" gridcol="1" gridrow="7" pageId="3" pageNumber="350">22.3 ±3.8</td>
<td id="76A55DB4FF8A00641420FDCFFC4CFDA0" box="[793,900,532,554]" gridcol="2" gridrow="7" pageId="3" pageNumber="350">4.2–4.6</td>
<td id="76A55DB4FF8A006414C0FDCFFB41FDA0" box="[1017,1161,532,554]" gridcol="3" gridrow="7" pageId="3" pageNumber="350">13 ±3.6</td>
<td id="76A55DB4FF8A006413E0FDCFFAF6FDA0" box="[1241,1342,532,554]" gridcol="4" gridrow="7" pageId="3" pageNumber="350">3.3–5.1</td>
</tr>
<tr id="357434C8FF8A006417A6FDE6FAF6FDD9" box="[159,1342,573,595]" gridrow="8" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FDE6FE05FDD9" box="[159,461,573,595]" gridcol="0" gridrow="8" pageId="3" pageNumber="350">First dorsal anterior margin</th>
<td id="76A55DB4FF8A00641502FDE6FD01FDD9" box="[571,713,573,595]" gridcol="1" gridrow="8" pageId="3" pageNumber="350">51.1 ±6.3</td>
<td id="76A55DB4FF8A00641420FDE6FC4CFDD9" box="[793,900,573,595]" gridcol="2" gridrow="8" pageId="3" pageNumber="350">9.7–10.0</td>
<td id="76A55DB4FF8A006414C0FDE6FB41FDD9" box="[1017,1161,573,595]" gridcol="3" gridrow="8" pageId="3" pageNumber="350">34.9 ±8.5</td>
<td id="76A55DB4FF8A006413E0FDE6FAF6FDD9" box="[1241,1342,573,595]" gridcol="4" gridrow="8" pageId="3" pageNumber="350">8.7–10.6</td>
</tr>
<tr id="357434C8FF8A006417A6FDBDFAF6FDF6" box="[159,1342,614,636]" gridrow="9" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FDBDFE05FDF6" box="[159,461,614,636]" gridcol="0" gridrow="9" pageId="3" pageNumber="350">First dorsal base</th>
<td id="76A55DB4FF8A00641502FDBDFD01FDF6" box="[571,713,614,636]" gridcol="1" gridrow="9" pageId="3" pageNumber="350">36.1 ±5.7</td>
<td id="76A55DB4FF8A00641420FDBDFC4CFDF6" box="[793,900,614,636]" gridcol="2" gridrow="9" pageId="3" pageNumber="350">6.6–7.1</td>
<td id="76A55DB4FF8A006414C0FDBDFB41FDF6" box="[1017,1161,614,636]" gridcol="3" gridrow="9" pageId="3" pageNumber="350">24.1 ±6.4</td>
<td id="76A55DB4FF8A006413E0FDBDFAF6FDF6" box="[1241,1342,614,636]" gridcol="4" gridrow="9" pageId="3" pageNumber="350">6.7–8.0</td>
</tr>
<tr id="357434C8FF8A006417A6FD4BFAF6FD2C" box="[159,1342,656,678]" gridrow="10" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FD4BFE05FD2C" box="[159,461,656,678]" gridcol="0" gridrow="10" pageId="3" pageNumber="350">First dorsal height</th>
<td id="76A55DB4FF8A00641502FD4BFD01FD2C" box="[571,713,656,678]" gridcol="1" gridrow="10" pageId="3" pageNumber="350">30.2 ±4.9</td>
<td id="76A55DB4FF8A00641420FD4BFC4CFD2C" box="[793,900,656,678]" gridcol="2" gridrow="10" pageId="3" pageNumber="350">5.7 –5.9</td>
<td id="76A55DB4FF8A006414C0FD4BFB41FD2C" box="[1017,1161,656,678]" gridcol="3" gridrow="10" pageId="3" pageNumber="350">19.1 ±5.7</td>
<td id="76A55DB4FF8A006413E0FD4BFAF6FD2C" box="[1241,1342,656,678]" gridcol="4" gridrow="10" pageId="3" pageNumber="350">2.7–6.0</td>
</tr>
<tr id="357434C8FF8A006417A6FD62FAF6FD45" box="[159,1342,697,719]" gridrow="11" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FD62FE05FD45" box="[159,461,697,719]" gridcol="0" gridrow="11" pageId="3" pageNumber="350">First dorsal inner margin</th>
<td id="76A55DB4FF8A00641502FD62FD01FD45" box="[571,713,697,719]" gridcol="1" gridrow="11" pageId="3" pageNumber="350">15.7 ±2.7</td>
<td id="76A55DB4FF8A00641420FD62FC4CFD45" box="[793,900,697,719]" gridcol="2" gridrow="11" pageId="3" pageNumber="350">3.1–3.4</td>
<td id="76A55DB4FF8A006414C0FD62FB41FD45" box="[1017,1161,697,719]" gridcol="3" gridrow="11" pageId="3" pageNumber="350">10.2 ±2.8</td>
<td id="76A55DB4FF8A006413E0FD62FAF6FD45" box="[1241,1342,697,719]" gridcol="4" gridrow="11" pageId="3" pageNumber="350">2.0–3.6</td>
</tr>
<tr id="357434C8FF8A006417A6FD39FAF6FD72" box="[159,1342,738,760]" gridrow="12" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FD39FE05FD72" box="[159,461,738,760]" gridcol="0" gridrow="12" pageId="3" pageNumber="350">Second dorsal anterior margin</th>
<td id="76A55DB4FF8A00641502FD39FD01FD72" box="[571,713,738,760]" gridcol="1" gridrow="12" pageId="3" pageNumber="350">36.6 ±7.9</td>
<td id="76A55DB4FF8A00641420FD39FC4CFD72" box="[793,900,738,760]" gridcol="2" gridrow="12" pageId="3" pageNumber="350">5.0–7.6</td>
<td id="76A55DB4FF8A006414C0FD39FB41FD72" box="[1017,1161,738,760]" gridcol="3" gridrow="12" pageId="3" pageNumber="350">26.5 ±6.1</td>
<td id="76A55DB4FF8A006413E0FD39FAF6FD72" box="[1241,1342,738,760]" gridcol="4" gridrow="12" pageId="3" pageNumber="350">5.3–7.6</td>
</tr>
<tr id="357434C8FF8A006417A6FCD7FAF6FCA8" box="[159,1342,780,802]" gridrow="13" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FCD7FE05FCA8" box="[159,461,780,802]" gridcol="0" gridrow="13" pageId="3" pageNumber="350">Second dorsal base</th>
<td id="76A55DB4FF8A00641502FCD7FD01FCA8" box="[571,713,780,802]" gridcol="1" gridrow="13" pageId="3" pageNumber="350">30.1 ±4.8</td>
<td id="76A55DB4FF8A00641420FCD7FC4CFCA8" box="[793,900,780,802]" gridcol="2" gridrow="13" pageId="3" pageNumber="350">5.5–6.1</td>
<td id="76A55DB4FF8A006414C0FCD7FB41FCA8" box="[1017,1161,780,802]" gridcol="3" gridrow="13" pageId="3" pageNumber="350">19.5 ±4.9</td>
<td id="76A55DB4FF8A006413E0FCD7FAF6FCA8" box="[1241,1342,780,802]" gridcol="4" gridrow="13" pageId="3" pageNumber="350">5.3–6.6</td>
</tr>
<tr id="357434C8FF8A006417A6FCEEFAF6FCC1" box="[159,1342,821,843]" gridrow="14" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FCEEFE05FCC1" box="[159,461,821,843]" gridcol="0" gridrow="14" pageId="3" pageNumber="350">Second dorsal height</th>
<td id="76A55DB4FF8A00641502FCEEFD01FCC1" box="[571,713,821,843]" gridcol="1" gridrow="14" pageId="3" pageNumber="350">21.1 ±4.6</td>
<td id="76A55DB4FF8A00641420FCEEFC4CFCC1" box="[793,900,821,843]" gridcol="2" gridrow="14" pageId="3" pageNumber="350">3.8–4.7</td>
<td id="76A55DB4FF8A006414C0FCEEFB41FCC1" box="[1017,1161,821,843]" gridcol="3" gridrow="14" pageId="3" pageNumber="350">12.6 ±3.4</td>
<td id="76A55DB4FF8A006413E0FCEEFAF6FCC1" box="[1241,1342,821,843]" gridcol="4" gridrow="14" pageId="3" pageNumber="350">3.3–4.7</td>
</tr>
<tr id="357434C8FF8A006417A6FC85FAF6FCFE" box="[159,1342,862,884]" gridrow="15" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FC85FE05FCFE" box="[159,461,862,884]" gridcol="0" gridrow="15" pageId="3" pageNumber="350">Second dorsal inner margin</th>
<td id="76A55DB4FF8A00641502FC85FD01FCFE" box="[571,713,862,884]" gridcol="1" gridrow="15" pageId="3" pageNumber="350">13.7 ±2.5</td>
<td id="76A55DB4FF8A00641420FC85FC4CFCFE" box="[793,900,862,884]" gridcol="2" gridrow="15" pageId="3" pageNumber="350">2.6–2.9</td>
<td id="76A55DB4FF8A006414C0FC85FB41FCFE" box="[1017,1161,862,884]" gridcol="3" gridrow="15" pageId="3" pageNumber="350">9 ±2.4</td>
<td id="76A55DB4FF8A006413E0FC85FAF6FCFE" box="[1241,1342,862,884]" gridcol="4" gridrow="15" pageId="3" pageNumber="350">1.3–3.0</td>
</tr>
<tr id="357434C8FF8A006417A6FC53FAF6FC14" box="[159,1342,904,926]" gridrow="16" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FC53FE05FC14" box="[159,461,904,926]" gridcol="0" gridrow="16" pageId="3" pageNumber="350">Dorsal caudal margin</th>
<td id="76A55DB4FF8A00641502FC53FD01FC14" box="[571,713,904,926]" gridcol="1" gridrow="16" pageId="3" pageNumber="350">122.5 ±13.4</td>
<td id="76A55DB4FF8A00641420FC53FC4CFC14" box="[793,900,904,926]" gridcol="2" gridrow="16" pageId="3" pageNumber="350">23.4–25.3</td>
<td id="76A55DB4FF8A006414C0FC53FB41FC14" box="[1017,1161,904,926]" gridcol="3" gridrow="16" pageId="3" pageNumber="350">78.7 ±16</td>
<td id="76A55DB4FF8A006413E0FC53FAF6FC14" box="[1241,1342,904,926]" gridcol="4" gridrow="16" pageId="3" pageNumber="350">23.2–23.5</td>
</tr>
<tr id="357434C8FF8A006417A6FC6AFAF6FC4D" box="[159,1342,945,967]" gridrow="17" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FC6AFE05FC4D" box="[159,461,945,967]" gridcol="0" gridrow="17" pageId="3" pageNumber="350">Postventral caudal margin</th>
<td id="76A55DB4FF8A00641502FC6AFD01FC4D" box="[571,713,945,967]" gridcol="1" gridrow="17" pageId="3" pageNumber="350">50.7 ±4.0</td>
<td id="76A55DB4FF8A00641420FC6AFC4CFC4D" box="[793,900,945,967]" gridcol="2" gridrow="17" pageId="3" pageNumber="350">9.4–10.7</td>
<td id="76A55DB4FF8A006414C0FC6AFB41FC4D" box="[1017,1161,945,967]" gridcol="3" gridrow="17" pageId="3" pageNumber="350">35.4 ±7.5</td>
<td id="76A55DB4FF8A006413E0FC6AFAF6FC4D" box="[1241,1342,945,967]" gridcol="4" gridrow="17" pageId="3" pageNumber="350">8.7–10.0</td>
</tr>
<tr id="357434C8FF8A006417A6FC01FAF6FC7A" box="[159,1342,986,1008]" gridrow="18" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FC01FE05FC7A" box="[159,461,986,1008]" gridcol="0" gridrow="18" pageId="3" pageNumber="350">Subterminal caudal margin</th>
<td id="76A55DB4FF8A00641502FC01FD01FC7A" box="[571,713,986,1008]" gridcol="1" gridrow="18" pageId="3" pageNumber="350">48.5 ±9.0</td>
<td id="76A55DB4FF8A00641420FC01FC4CFC7A" box="[793,900,986,1008]" gridcol="2" gridrow="18" pageId="3" pageNumber="350">8.8–10.7</td>
<td id="76A55DB4FF8A006414C0FC01FB41FC7A" box="[1017,1161,986,1008]" gridcol="3" gridrow="18" pageId="3" pageNumber="350">30.9 ±6.8</td>
<td id="76A55DB4FF8A006413E0FC01FAF6FC7A" box="[1241,1342,986,1008]" gridcol="4" gridrow="18" pageId="3" pageNumber="350">8.0–8.5</td>
</tr>
<tr id="357434C8FF8A006417A6FBDFFAF6FB90" box="[159,1342,1028,1050]" gridrow="19" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FBDFFE05FB90" box="[159,461,1028,1050]" gridcol="0" gridrow="19" pageId="3" pageNumber="350">Terminal caudal margin</th>
<td id="76A55DB4FF8A00641502FBDFFD01FB90" box="[571,713,1028,1050]" gridcol="1" gridrow="19" pageId="3" pageNumber="350">25.8 ±4.0</td>
<td id="76A55DB4FF8A00641420FBDFFC4CFB90" box="[793,900,1028,1050]" gridcol="2" gridrow="19" pageId="3" pageNumber="350">5.2–5.4</td>
<td id="76A55DB4FF8A006414C0FBDFFB41FB90" box="[1017,1161,1028,1050]" gridcol="3" gridrow="19" pageId="3" pageNumber="350">18.4 ±5.1</td>
<td id="76A55DB4FF8A006413E0FBDFFAF6FB90" box="[1241,1342,1028,1050]" gridcol="4" gridrow="19" pageId="3" pageNumber="350">0.6–7.0</td>
</tr>
<tr id="357434C8FF8A006417A6FBF6FAF6FBC9" box="[159,1342,1069,1091]" gridrow="20" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FBF6FE05FBC9" box="[159,461,1069,1091]" gridcol="0" gridrow="20" pageId="3" pageNumber="350">Terminal caudal lobe</th>
<td id="76A55DB4FF8A00641502FBF6FD01FBC9" box="[571,713,1069,1091]" gridcol="1" gridrow="20" pageId="3" pageNumber="350">29.0 ±4.0</td>
<td id="76A55DB4FF8A00641420FBF6FC4CFBC9" box="[793,900,1069,1091]" gridcol="2" gridrow="20" pageId="3" pageNumber="350">5.4–5.6</td>
<td id="76A55DB4FF8A006414C0FBF6FB41FBC9" box="[1017,1161,1069,1091]" gridcol="3" gridrow="20" pageId="3" pageNumber="350">18.6 ±5.3</td>
<td id="76A55DB4FF8A006413E0FBF6FAF6FBC9" box="[1241,1342,1069,1091]" gridcol="4" gridrow="20" pageId="3" pageNumber="350">4.0–7.2</td>
</tr>
<tr id="357434C8FF8A006417A6FB8DFAF6FBE6" box="[159,1342,1110,1132]" gridrow="21" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FB8DFE05FBE6" box="[159,461,1110,1132]" gridcol="0" gridrow="21" pageId="3" pageNumber="350">Head height</th>
<td id="76A55DB4FF8A00641502FB8DFD01FBE6" box="[571,713,1110,1132]" gridcol="1" gridrow="21" pageId="3" pageNumber="350">32.6 ±7.4</td>
<td id="76A55DB4FF8A00641420FB8DFC4CFBE6" box="[793,900,1110,1132]" gridcol="2" gridrow="21" pageId="3" pageNumber="350">5.4–6.6</td>
<td id="76A55DB4FF8A006414C0FB8DFB41FBE6" box="[1017,1161,1110,1132]" gridcol="3" gridrow="21" pageId="3" pageNumber="350">22.7 ±5.8</td>
<td id="76A55DB4FF8A006413E0FB8DFAF6FBE6" box="[1241,1342,1110,1132]" gridcol="4" gridrow="21" pageId="3" pageNumber="350">4.7–7.0</td>
</tr>
<tr id="357434C8FF8A006417A6FB5BFAF6FB1C" box="[159,1342,1152,1174]" gridrow="22" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FB5BFE05FB1C" box="[159,461,1152,1174]" gridcol="0" gridrow="22" pageId="3" pageNumber="350">Trunk height</th>
<td id="76A55DB4FF8A00641502FB5BFD01FB1C" box="[571,713,1152,1174]" gridcol="1" gridrow="22" pageId="3" pageNumber="350">53.8 ±11.4</td>
<td id="76A55DB4FF8A00641420FB5BFC4CFB1C" box="[793,900,1152,1174]" gridcol="2" gridrow="22" pageId="3" pageNumber="350">9.5–12.2</td>
<td id="76A55DB4FF8A006414C0FB5BFB41FB1C" box="[1017,1161,1152,1174]" gridcol="3" gridrow="22" pageId="3" pageNumber="350">42.2 ±8.9</td>
<td id="76A55DB4FF8A006413E0FB5BFAF6FB1C" box="[1241,1342,1152,1174]" gridcol="4" gridrow="22" pageId="3" pageNumber="350">10.7–11.6</td>
</tr>
<tr id="357434C8FF8A006417A6FB72FAF6FB35" box="[159,1342,1193,1215]" gridrow="23" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FB72FE05FB35" box="[159,461,1193,1215]" gridcol="0" gridrow="23" pageId="3" pageNumber="350">Abdomen height</th>
<td id="76A55DB4FF8A00641502FB72FD01FB35" box="[571,713,1193,1215]" gridcol="1" gridrow="23" pageId="3" pageNumber="350">37.3 ±7.0</td>
<td id="76A55DB4FF8A00641420FB72FC4CFB35" box="[793,900,1193,1215]" gridcol="2" gridrow="23" pageId="3" pageNumber="350">7.1–8.2</td>
<td id="76A55DB4FF8A006414C0FB72FB41FB35" box="[1017,1161,1193,1215]" gridcol="3" gridrow="23" pageId="3" pageNumber="350">28.5 ±8</td>
<td id="76A55DB4FF8A006413E0FB72FAF6FB35" box="[1241,1342,1193,1215]" gridcol="4" gridrow="23" pageId="3" pageNumber="350">6.0–8.9</td>
</tr>
<tr id="357434C8FF8A006417A6FB09FAF6FB62" box="[159,1342,1234,1256]" gridrow="24" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FB09FE05FB62" box="[159,461,1234,1256]" gridcol="0" gridrow="24" pageId="3" pageNumber="350">Caudal height</th>
<td id="76A55DB4FF8A00641502FB09FD01FB62" box="[571,713,1234,1256]" gridcol="1" gridrow="24" pageId="3" pageNumber="350">28.8 ±8.3</td>
<td id="76A55DB4FF8A00641420FB09FC4CFB62" box="[793,900,1234,1256]" gridcol="2" gridrow="24" pageId="3" pageNumber="350">5.0–7.6</td>
<td id="76A55DB4FF8A006414C0FB09FB41FB62" box="[1017,1161,1234,1256]" gridcol="3" gridrow="24" pageId="3" pageNumber="350">20.9 ±6</td>
<td id="76A55DB4FF8A006413E0FB09FAF6FB62" box="[1241,1342,1234,1256]" gridcol="4" gridrow="24" pageId="3" pageNumber="350">5.3–7.4</td>
</tr>
<tr id="357434C8FF8A006417A6FB27FAF6FA98" box="[159,1342,1276,1298]" gridrow="25" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FB27FE05FA98" box="[159,461,1276,1298]" gridcol="0" gridrow="25" pageId="3" pageNumber="350">Caudal peduncle height</th>
<td id="76A55DB4FF8A00641502FB27FD01FA98" box="[571,713,1276,1298]" gridcol="1" gridrow="25" pageId="3" pageNumber="350">15.8 ±2.5</td>
<td id="76A55DB4FF8A00641420FB27FC4CFA98" box="[793,900,1276,1298]" gridcol="2" gridrow="25" pageId="3" pageNumber="350">3.1–3.3</td>
<td id="76A55DB4FF8A006414C0FB27FB41FA98" box="[1017,1161,1276,1298]" gridcol="3" gridrow="25" pageId="3" pageNumber="350">12.6 ±3.8</td>
<td id="76A55DB4FF8A006413E0FB27FAF6FA98" box="[1241,1342,1276,1298]" gridcol="4" gridrow="25" pageId="3" pageNumber="350">3.3–4.9</td>
</tr>
<tr id="357434C8FF8A006417A6FAFEFAF6FAB1" box="[159,1342,1317,1339]" gridrow="26" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FAFEFE05FAB1" box="[159,461,1317,1339]" gridcol="0" gridrow="26" pageId="3" pageNumber="350">Head width</th>
<td id="76A55DB4FF8A00641502FAFEFD01FAB1" box="[571,713,1317,1339]" gridcol="1" gridrow="26" pageId="3" pageNumber="350">65.0 ±8.6</td>
<td id="76A55DB4FF8A00641420FAFEFC4CFAB1" box="[793,900,1317,1339]" gridcol="2" gridrow="26" pageId="3" pageNumber="350">13. 0–13.3</td>
<td id="76A55DB4FF8A006414C0FAFEFB41FAB1" box="[1017,1161,1317,1339]" gridcol="3" gridrow="26" pageId="3" pageNumber="350">41.4 ±13.5</td>
<td id="76A55DB4FF8A006413E0FAFEFAF6FAB1" box="[1241,1342,1317,1339]" gridcol="4" gridrow="26" pageId="3" pageNumber="350">3.3–11.7</td>
</tr>
<tr id="357434C8FF8A006417A6FA95FAF6FAEE" box="[159,1342,1358,1380]" gridrow="27" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FA95FE05FAEE" box="[159,461,1358,1380]" gridcol="0" gridrow="27" pageId="3" pageNumber="350">Trunk width</th>
<td id="76A55DB4FF8A00641502FA95FD01FAEE" box="[571,713,1358,1380]" gridcol="1" gridrow="27" pageId="3" pageNumber="350">63.8 ±13.7</td>
<td id="76A55DB4FF8A00641420FA95FC4CFAEE" box="[793,900,1358,1380]" gridcol="2" gridrow="27" pageId="3" pageNumber="350">10.9–14.5</td>
<td id="76A55DB4FF8A006414C0FA95FB41FAEE" box="[1017,1161,1358,1380]" gridcol="3" gridrow="27" pageId="3" pageNumber="350">43.7 ±10.3</td>
<td id="76A55DB4FF8A006413E0FA95FAF6FAEE" box="[1241,1342,1358,1380]" gridcol="4" gridrow="27" pageId="3" pageNumber="350">11.4–12.7</td>
</tr>
<tr id="357434C8FF8A006417A6FAA3FAF6FA04" box="[159,1342,1400,1422]" gridrow="28" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FAA3FE05FA04" box="[159,461,1400,1422]" gridcol="0" gridrow="28" pageId="3" pageNumber="350">Abdomen width</th>
<td id="76A55DB4FF8A00641502FAA3FD01FA04" box="[571,713,1400,1422]" gridcol="1" gridrow="28" pageId="3" pageNumber="350">42.3 ±9.2</td>
<td id="76A55DB4FF8A00641420FAA3FC4CFA04" box="[793,900,1400,1422]" gridcol="2" gridrow="28" pageId="3" pageNumber="350">6.6–8.7</td>
<td id="76A55DB4FF8A006414C0FAA3FB41FA04" box="[1017,1161,1400,1422]" gridcol="3" gridrow="28" pageId="3" pageNumber="350">26.7 ±7.2</td>
<td id="76A55DB4FF8A006413E0FAA3FAF6FA04" box="[1241,1342,1400,1422]" gridcol="4" gridrow="28" pageId="3" pageNumber="350">6.7–8.9</td>
</tr>
<tr id="357434C8FF8A006417A6FA7AFAF6FA3D" box="[159,1342,1441,1463]" gridrow="29" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FA7AFE05FA3D" box="[159,461,1441,1463]" gridcol="0" gridrow="29" pageId="3" pageNumber="350">Caudal width</th>
<td id="76A55DB4FF8A00641502FA7AFD01FA3D" box="[571,713,1441,1463]" gridcol="1" gridrow="29" pageId="3" pageNumber="350">22.0 ±5.0</td>
<td id="76A55DB4FF8A00641420FA7AFC4CFA3D" box="[793,900,1441,1463]" gridcol="2" gridrow="29" pageId="3" pageNumber="350">3.8–4.9</td>
<td id="76A55DB4FF8A006414C0FA7AFB41FA3D" box="[1017,1161,1441,1463]" gridcol="3" gridrow="29" pageId="3" pageNumber="350">16 ±4.2</td>
<td id="76A55DB4FF8A006413E0FA7AFAF6FA3D" box="[1241,1342,1441,1463]" gridcol="4" gridrow="29" pageId="3" pageNumber="350">4.0–4.9</td>
</tr>
<tr id="357434C8FF8A006417A6FA11FAF6FA6A" box="[159,1342,1482,1504]" gridrow="30" pageId="3" pageNumber="350">
<th id="76A55DB4FF8A006417A6FA11FE05FA6A" box="[159,461,1482,1504]" gridcol="0" gridrow="30" pageId="3" pageNumber="350">Caudal peduncle width</th>
<td id="76A55DB4FF8A00641502FA11FD01FA6A" box="[571,713,1482,1504]" gridcol="1" gridrow="30" pageId="3" pageNumber="350">12.0 ±2.2</td>
<td id="76A55DB4FF8A00641420FA11FC4CFA6A" box="[793,900,1482,1504]" gridcol="2" gridrow="30" pageId="3" pageNumber="350">1.9–2.3</td>
<td id="76A55DB4FF8A006414C0FA11FB41FA6A" box="[1017,1161,1482,1504]" gridcol="3" gridrow="30" pageId="3" pageNumber="350">10.8 ±3.6</td>
<td id="76A55DB4FF8A006413E0FA11FAF6FA6A" box="[1241,1342,1482,1504]" gridcol="4" gridrow="30" pageId="3" pageNumber="350">2.7–4.1</td>
</tr>
</table>
</paragraph>
<caption id="DF3B6602FF8DFF9F17AEFBB1FBF2FB14" httpUri="https://zenodo.org/record/243846/files/figure.png" pageId="4" pageNumber="351" targetBox="[371,1214,204,1099]" targetPageId="4">
<paragraph id="8BFB368AFF8DFF9F17AEFBB1FBF2FB14" blockId="4.[151,1436,1130,1182]" pageId="4" pageNumber="351">
<emphasis id="B930EA98FF8DFF9F17AEFBB1FEDDFB0A" bold="true" box="[151,277,1130,1152]" pageId="4" pageNumber="351">FIGURE 1.</emphasis>
<taxonomicName id="4C444D09FF8DFF9F1627FBB1FE1AFB0A" box="[286,466,1130,1152]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="4" pageNumber="351" phylum="Chordata" rank="species" species="ugoi" status="sp. nov.">
<emphasis id="B930EA98FF8DFF9F1627FBB1FE1AFB0A" box="[286,466,1130,1152]" italics="true" pageId="4" pageNumber="351">Scyliorhinus ugoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20357E3FF8DFF9F16E5FBB1FDF3FB0A" box="[476,571,1130,1152]" pageId="4" pageNumber="351" rank="species">
<emphasis id="B930EA98FF8DFF9F16E5FBB1FDF3FB0A" bold="true" box="[476,571,1130,1152]" pageId="4" pageNumber="351">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype, MNRJ 42619, female, 500 mm TL. A) Dorsal view. B) Lateral view; paratype, UERJ 2179, male, 419 mm TL. C) Dorsal view. D) Lateral view. Scale bar: 5 cm.
</paragraph>
</caption>
<paragraph id="8BFB368AFF8DFF9F17FEFB1EFAD3FA89" blockId="4.[151,1437,1221,1643]" pageId="4" pageNumber="351">
First two gill openings about equally long; first gill opening twice as large as fifth (
<figureCitation id="137F2A0FFF8DFF9F1348FB1EFB70FB54" box="[1137,1208,1221,1246]" captionStart="FIGURE 2" captionStartId="6.[151,260,1897,1919]" captionTargetBox="[346,1240,665,1868]" captionTargetId="figure@6.[340,1248,661,1875]" captionTargetPageId="6" captionText="FIGURE 2. Scyliorhinus ugoi sp. nov. A) Dorsal view of the head of a female, MNRJ 42619, female, 500 mm TL. B) Dorsal view of the head of a male, UERJ 2179, 419 mm TL. C) Lateral view of head, UERJ 2179 male, 419 mm TL. D) First dorsal fin. E) Second dorsal fin. F) Anal fin. G) Caudal fin. H) Ventral view of trunk, MNRJ 42619, female, 500 mm TL. Scale bars: 2 cm." httpUri="https://zenodo.org/record/243847/files/figure.png" pageId="4" pageNumber="351">Fig. 2</figureCitation>
c). All gill openings slightly concave and not elevated on dorsolateral surface of head, gill filaments not visible externally.
</paragraph>
<paragraph id="8BFB368AFF8DFF9F17FEFAD6FC12F9A9" blockId="4.[151,1437,1221,1643]" pageId="4" pageNumber="351">Sexual heterodonty weak; adult males presenting teeth longer with undeveloped lateral cusps on central portion of lower jaw. Upper teeth slightly higher-crowned than lower teeth, with longer, stronger transverse ridges, otherwise very similar. Medial teeth (M) and anteroposterior teeth (AN) poorly differentiated; symphysial teeth (S) with higher-crown and smaller than anteroposteriors, with erect or semi-erect cusps and one weak cusplet on either side of the cusp. Anteroposteriors in both jaws larger than medials and symphysials, with semi oblique cusps, usually one strong high cusplet on either side, longitudinal ridges confined to the basal ledges, and low, flat roots. Gradient monognathic heterodonty well-developed in anteroposterior teeth; anteroposteriors smaller distally, with thicker and more oblique cusps, and lower cusplets.</paragraph>
<paragraph id="8BFB368AFF8DFF9F17FEF9F6FD5CF9E1" blockId="4.[151,1437,1221,1643]" pageId="4" pageNumber="351">Teeth in 52/49 (47–56/45–53) rows, 3/2 (2–3/2–3) functional series. Teeth not highly differentiated in upper and lower jaws or along jaws. Tooth formula:</paragraph>
<paragraph id="8BFB368AFF8DFF9F17AEF941FC8EF939" blockId="4.[151,838,1690,1750]" box="[151,838,1690,1715]" pageId="4" pageNumber="351">Left AN16–19 M6–8 S1 S1 M6–8 AN16–19 Right</paragraph>
<paragraph id="8BFB368AFF8DFF9F17E5F966FD31F95C" blockId="4.[151,838,1690,1750]" box="[220,761,1725,1750]" pageId="4" pageNumber="351">AN15–17 M6–9 M7–10 AN16–17</paragraph>
<paragraph id="8BFB368AFF8DFF9F17FEF8DEFBD2F824" blockId="4.[151,1437,1797,1966]" pageId="4" pageNumber="351">
Lateral trunk denticles (
<figureCitation id="137F2A0FFF8DFF9F16E1F8DEFDD4F894" box="[472,540,1797,1822]" captionStart="FIGURE 5" captionStartId="8.[151,249,1047,1069]" captionTargetBox="[341,1245,447,1021]" captionTargetId="figure@8.[340,1248,445,1025]" captionTargetPageId="8" captionText="FIGURE 5. Lateral trunk denticles of Scyliorhinus ugoi sp. nov. A – C) UERJ 1726, female, 602 mm TL. D – F) UERJ 1725, adult male, 534 mm TL. A, D) Region above pectoral fin. B, E) Region below first dorsal fin. C, F) Region below second dorsal fin." httpUri="https://zenodo.org/record/243850/files/figure.png" pageId="4" pageNumber="351">Fig. 5</figureCitation>
) with flat, elongated teardrop-shaped crowns 1.4–2.1 times as long as wide, anterior part covered with ectodermal pits (
<emphasis id="B930EA98FF8DFF9F159BF8F0FD28F8C8" box="[674,736,1835,1858]" italics="true" pageId="4" pageNumber="351">sensu</emphasis>
Muñoz- Chápuli, 1985). Crown with a strong medial ridge that extends entire length of crown on the long medial cusp. Lateral cusps not well developed, 0.3 times in medial cusp; lateral ridges anterior to them very short or absent. Males with dermal denticles longer than females. Density of dermal denticles on posterior regions lower than on anterior regions.
</paragraph>
<caption id="DF3B6602FF8CFF9E17AEFF43FD20FF45" ID-Table-UUID="DF3B6602FF8CFF9E17AEFF43FD20FF45" httpUri="http://table.plazi.org/id/DF3B6602FF8CFF9E17AEFF43FD20FF45" pageId="5" pageNumber="352" targetBox="[159,1335,317,628]" targetIsTable="true" targetPageId="5">
<paragraph id="8BFB368AFF8CFF9E17AEFF43FD20FF45" blockId="5.[151,1436,151,207]" pageId="5" pageNumber="352">
<emphasis id="B930EA98FF8CFF9E17AEFF43FEC9FF25" bold="true" box="[151,257,152,175]" pageId="5" pageNumber="352">TABLE 2</emphasis>
. Measurements in millimeters and proportions as percentage of nasobasal length for 
<taxonomicName id="4C444D09FF8CFF9E1346FF4CFAFFFF25" box="[1151,1335,151,175]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="5" pageNumber="352" phylum="Chordata" rank="species" species="ugoi" status="sp. nov.">
<emphasis id="B930EA98FF8CFF9E1346FF4CFAFFFF25" box="[1151,1335,151,175]" italics="true" pageId="5" pageNumber="352">Scyliorhinus ugoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20357E3FF8CFF9E1204FF43FA54FF25" box="[1341,1436,152,175]" pageId="5" pageNumber="352" rank="species">
<emphasis id="B930EA98FF8CFF9E1204FF43FA54FF25" bold="true" box="[1341,1436,152,175]" pageId="5" pageNumber="352">sp. nov.</emphasis>
</taxonomicNameLabel>
Female (F), male (M), and range of proportions (%NL).
</paragraph>
</caption>
<paragraph id="8BFB368AFF8CFF9E17A6FF30FB38FE8A" blockId="5.[159,1295,234,298]" box="[159,1264,234,256]" pageId="5" pageNumber="352">
Measurements 
<taxonomicName id="4C444D09FF8CFF9E14E6FF31FB47FE8A" box="[991,1167,234,256]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="5" pageNumber="352" phylum="Chordata" rank="species" species="ugoi" status="sp. nov.">
<emphasis id="B930EA98FF8CFF9E14E6FF31FB47FE8A" box="[991,1167,234,256]" italics="true" pageId="5" pageNumber="352">Scyliorhinus ugoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20357E3FF8CFF9E13AFFF31FB38FE8A" box="[1174,1264,234,256]" pageId="5" pageNumber="352" rank="species">
<emphasis id="B930EA98FF8CFF9E13AFFF31FB38FE8A" bold="true" box="[1174,1264,234,256]" pageId="5" pageNumber="352">sp. nov.</emphasis>
</taxonomicNameLabel>
</paragraph>
<paragraph id="8BFB368AFF8CFF9E1402FECEFECCFD36" blockId="5.[159,1295,234,298]" lastBlockId="5.[159,1335,648,700]" pageId="5" pageNumber="352">F % NL M % NL Distance between rostral base and anterior edge of anterior 21 39.5 19.5 36.7–38.8 fontanelle</paragraph>
<paragraph id="8BFB368AFF8CFF9E17A6FEE6FAECFDFE" pageId="5" pageNumber="352">
<table id="F944C42AFF8C006417A6FEE6FAFFFDFE" box="[159,1335,317,628]" gridcols="5" gridrows="3" pageId="5" pageNumber="352">
<tr id="357434C8FF8C006417A6FEE6FAFFFEF6" box="[159,1335,317,380]" gridrow="0" pageId="5" pageNumber="352">
<th id="76A55DB4FF8C006417A6FEE6FDDBFEF6" box="[159,531,317,380]" gridcol="0" gridrow="0" pageId="5" pageNumber="352">
Nasobasal length (NL
<emphasis id="B930EA98FF8CFF9E1645FEE6FE4CFED9" bold="true" box="[380,388,317,339]" pageId="5" pageNumber="352">)</emphasis>
Rostral length
</th>
<th id="76A55DB4FF8C00641402FEE6FCAFFEF6" box="[827,871,317,380]" gridcol="1" gridrow="0" pageId="5" pageNumber="352">54.3 14.6</th>
<th id="76A55DB4FF8C00641490FEE6FBDCFEF6" box="[937,1044,317,380]" gridcol="2" gridrow="0" pageId="5" pageNumber="352">100 20.8–29.3</th>
<th id="76A55DB4FF8C00641349FEE6FB54FEF6" box="[1136,1180,317,380]" gridcol="3" gridrow="0" pageId="5" pageNumber="352">51.5 12</th>
<th id="76A55DB4FF8C006413EBFEE6FAFFFEF6" box="[1234,1335,317,380]" gridcol="4" gridrow="0" pageId="5" pageNumber="352">100 22.4–24</th>
</tr>
<tr id="357434C8FF8C006417A6FE4BFAFFFE72" box="[159,1335,400,504]" gridrow="1" pageId="5" pageNumber="352">
<th id="76A55DB4FF8C006417A6FE4BFDDBFE72" box="[159,531,400,504]" gridcol="0" gridrow="1" pageId="5" pageNumber="352">Width across lateral rostral cartilages Width across nasal capsules Nasal capsule width</th>
<td id="76A55DB4FF8C00641402FE4BFCAFFE72" box="[827,871,400,504]" gridcol="1" gridrow="1" pageId="5" pageNumber="352">7.6 43.6 21.6</td>
<td id="76A55DB4FF8C00641490FE4BFBDCFE72" box="[937,1044,400,504]" gridcol="2" gridrow="1" pageId="5" pageNumber="352">13.8–14.5 77.5– 85.4 37.9–43.7</td>
<td id="76A55DB4FF8C00641349FE4BFB54FE72" box="[1136,1180,400,504]" gridcol="3" gridrow="1" pageId="5" pageNumber="352">6.5 40 21</td>
<td id="76A55DB4FF8C006413EBFE4BFAFFFE72" box="[1234,1335,400,504]" gridcol="4" gridrow="1" pageId="5" pageNumber="352">12.2–13 76–79.6 40.8</td>
</tr>
<tr id="357434C8FF8C006417A6FDD7FAFFFDFE" box="[159,1335,524,628]" gridrow="2" pageId="5" pageNumber="352">
<th id="76A55DB4FF8C006417A6FDD7FDDBFDFE" box="[159,531,524,628]" gridcol="0" gridrow="2" pageId="5" pageNumber="352">Nasal capsule length Nasal aperture width Distance between nasal apertures</th>
<td id="76A55DB4FF8C00641402FDD7FCAFFDFE" box="[827,871,524,628]" gridcol="1" gridrow="2" pageId="5" pageNumber="352">20.6 13.5 6.6</td>
<td id="76A55DB4FF8C00641490FDD7FBDCFDFE" box="[937,1044,524,628]" gridcol="2" gridrow="2" pageId="5" pageNumber="352">37.5–39.6 25–25.8 10.4–13.8</td>
<td id="76A55DB4FF8C00641349FDD7FB54FDFE" box="[1136,1180,524,628]" gridcol="3" gridrow="2" pageId="5" pageNumber="352">19 14 6.5</td>
<td id="76A55DB4FF8C006413EBFDD7FAFFFDFE" box="[1234,1335,524,628]" gridcol="4" gridrow="2" pageId="5" pageNumber="352">35.1–38.7 26.5–27.7 12.2–13</td>
</tr>
</table>
</paragraph>
<paragraph id="8BFB368AFF8CFF9E17A6FD0BFAFFFBD0" pageId="5" pageNumber="352">
<table id="F944C42AFF8C006417A6FD0BFAFFFBD0" box="[159,1335,720,1114]" gridcols="5" gridrows="4" pageId="5" pageNumber="352">
<tr id="357434C8FF8C006417A6FD0BFAFFFCB2" box="[159,1335,720,824]" gridrow="0" pageId="5" pageNumber="352">
<th id="76A55DB4FF8C006417A6FD0BFE31FCB2" box="[159,505,720,824]" gridcol="0" gridrow="0" pageId="5" pageNumber="352">Anterior fontanelle width Basal plate width Orbit length</th>
<th id="76A55DB4FF8C00641402FD0BFCAFFCB2" box="[827,871,720,824]" gridcol="1" gridrow="0" pageId="5" pageNumber="352">13.3 22.6 24.3</th>
<th id="76A55DB4FF8C00641490FD0BFBC6FCB2" box="[937,1038,720,824]" gridcol="2" gridrow="0" pageId="5" pageNumber="352">24.1–25 39.6–45.8 44.8–47.9</th>
<th id="76A55DB4FF8C00641349FD0BFB54FCB2" box="[1136,1180,720,824]" gridcol="3" gridrow="0" pageId="5" pageNumber="352">12.5 19.5 24</th>
<th id="76A55DB4FF8C006413EAFD0BFAFFFCB2" box="[1235,1335,720,824]" gridcol="4" gridrow="0" pageId="5" pageNumber="352">22.4–26 37–38.7 46.3–47</th>
</tr>
<tr id="357434C8FF8C006417A6FC97FAFFFC3E" box="[159,1335,844,948]" gridrow="1" pageId="5" pageNumber="352">
<th id="76A55DB4FF8C006417A6FC97FE31FC3E" box="[159,505,844,948]" gridcol="0" gridrow="1" pageId="5" pageNumber="352">Preorbital process length Postorbital process length Nasal capsules length</th>
<td id="76A55DB4FF8C00641402FC97FCAFFC3E" box="[827,871,844,948]" gridcol="1" gridrow="1" pageId="5" pageNumber="352">3.3 4.5 11</td>
<td id="76A55DB4FF8C00641490FC97FBC6FC3E" box="[937,1038,844,948]" gridcol="2" gridrow="1" pageId="5" pageNumber="352">6.2–7.4 8.3–8.6 18.7–20.7</td>
<td id="76A55DB4FF8C00641349FC97FB54FC3E" box="[1136,1180,844,948]" gridcol="3" gridrow="1" pageId="5" pageNumber="352">3.5 4.5 10.5</td>
<td id="76A55DB4FF8C006413EAFC97FAFFFC3E" box="[1235,1335,844,948]" gridcol="4" gridrow="1" pageId="5" pageNumber="352">6.1–7.4 8.1–9.2 20.4</td>
</tr>
<tr id="357434C8FF8C006417A6FC13FAFFFBBA" box="[159,1335,968,1072]" gridrow="2" pageId="5" pageNumber="352">
<th id="76A55DB4FF8C006417A6FC13FE31FBBA" box="[159,505,968,1072]" gridcol="0" gridrow="2" pageId="5" pageNumber="352">Suborbital shelf width Otic capsules width Width across preorbital processes</th>
<td id="76A55DB4FF8C00641402FC13FCAFFBBA" box="[827,871,968,1072]" gridcol="1" gridrow="2" pageId="5" pageNumber="352">36 28 31.6</td>
<td id="76A55DB4FF8C00641490FC13FBC6FBBA" box="[937,1038,968,1072]" gridcol="2" gridrow="2" pageId="5" pageNumber="352">65.5–73 54.1–57.4 56.9–62.5</td>
<td id="76A55DB4FF8C00641349FC13FB54FBBA" box="[1136,1180,968,1072]" gridcol="3" gridrow="2" pageId="5" pageNumber="352">32.5 28.5 29.5</td>
<td id="76A55DB4FF8C006413EAFC13FAFFFBBA" box="[1235,1335,968,1072]" gridcol="4" gridrow="2" pageId="5" pageNumber="352">62.8–63.2 55.1–55.5 57.1–57.5</td>
</tr>
<tr id="357434C8FF8C006417A6FB9FFAFFFBD0" box="[159,1335,1092,1114]" gridrow="3" pageId="5" pageNumber="352">
<th id="76A55DB4FF8C006417A6FB9FFE31FBD0" box="[159,505,1092,1114]" gridcol="0" gridrow="3" pageId="5" pageNumber="352">Width across postorbital processes</th>
<td id="76A55DB4FF8C00641402FB9FFCAFFBD0" box="[827,871,1092,1114]" gridcol="1" gridrow="3" pageId="5" pageNumber="352">39.5</td>
<td id="76A55DB4FF8C00641490FB9FFBC6FBD0" box="[937,1038,1092,1114]" gridcol="2" gridrow="3" pageId="5" pageNumber="352">72.4–77</td>
<td id="76A55DB4FF8C00641349FB9FFB54FBD0" box="[1136,1180,1092,1114]" gridcol="3" gridrow="3" pageId="5" pageNumber="352">36</td>
<td id="76A55DB4FF8C006413EAFB9FFAFFFBD0" box="[1235,1335,1092,1114]" gridcol="4" gridrow="3" pageId="5" pageNumber="352">69.4–70.3</td>
</tr>
</table>
</paragraph>
<paragraph id="8BFB368AFF8CFF9E17FEFB45FB49FAA9" blockId="5.[151,1437,1182,2035]" pageId="5" pageNumber="352">
Body (
<figureCitation id="137F2A0FFF8CFF9E162FFB45FE93FB3D" box="[278,347,1182,1207]" captionStart="FIGURE 1" captionStartId="4.[151,249,1130,1152]" captionTargetBox="[371,1214,204,1099]" captionTargetId="figure@4.[362,1226,195,1105]" captionTargetPageId="4" captionText="FIGURE 1. Scyliorhinus ugoi sp. nov., holotype, MNRJ 42619, female, 500 mm TL. A) Dorsal view. B) Lateral view; paratype, UERJ 2179, male, 419 mm TL. C) Dorsal view. D) Lateral view. Scale bar: 5 cm." httpUri="https://zenodo.org/record/243846/files/figure.png" pageId="5" pageNumber="352">Fig. 1</figureCitation>
) stout and depressed on head region, tapering considerably to caudal fin. Prepectoral length corresponding to 17.9% of total length (TL) and prepelvic length 41– 43.4% TL. Snout– vent length 0.8 times in vent-caudal length. Pectoral-pelvic space 1.5–1.6 times pelvic–anal space (
<emphasis id="B930EA98FF8CFF9E130DFB3CFB9AFB74" box="[1076,1106,1255,1278]" italics="true" pageId="5" pageNumber="352">vs.</emphasis>
<quantity id="4CBC9B6FFF8CFF9E1362FB3CFB1FFB75" box="[1115,1239,1254,1279]" metricMagnitude="-2" metricUnit="m" metricValue="2.794" metricValueMax="3.5559999999999996" metricValueMin="2.032" pageId="5" pageNumber="352" unit="in" value="1.1" valueMax="1.4" valueMin="0.8">0.8–1.4 in</quantity>
the 
<emphasis id="B930EA98FF8CFF9E122BFB3DFEAAFAA9" italics="true" pageId="5" pageNumber="352">
<taxonomicName id="4C444D09FF8CFF9E122BFB3DFF30FAA9" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="5" pageNumber="352" phylum="Chordata" rank="species" species="haeckelii">Scyliorhinus haeckelii</taxonomicName>
/ 
<taxonomicName id="4C444D09FF8CFF9E163DFAD0FEAAFAA9" box="[260,354,1291,1315]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="5" pageNumber="352" phylum="Chordata" rank="species" species="besnardi">besnardi</taxonomicName>
</emphasis>
group). Interdorsal space 2.1–2.5 times dorsal-caudal space (
<tableCitation id="C6C60331FF8CFF9E1313FAD1FBB8FAA9" box="[1066,1136,1290,1315]" captionStart="TABLE 1" captionStartId="2.[151,239,152,175]" captionTargetBox="[159,1430,276,1951]" captionTargetPageId="2" captionText="TABLE 1. Measurements in millimeters and proportions as percentage of total length for Scyliorhinus ugoi sp. nov. Mean (x), standard deviation (SD) and range of proportions (%)." httpUri="http://table.plazi.org/id/DF3B6602FF8BFF9917AEFF43FC4FFF45" pageId="5" pageNumber="352" tableUuid="DF3B6602FF8BFF9917AEFF43FC4FFF45">Tab. 1</tableCitation>
).
</paragraph>
<paragraph id="8BFB368AFF8CFF9E17FEFAF5FAABFA39" blockId="5.[151,1437,1182,2035]" pageId="5" pageNumber="352">
Pectoral fins (
<figureCitation id="137F2A0FFF8CFF9E165BFAF5FE67FACD" box="[354,431,1326,1351]" captionStart="FIGURE 1" captionStartId="4.[151,249,1130,1152]" captionTargetBox="[371,1214,204,1099]" captionTargetId="figure@4.[362,1226,195,1105]" captionTargetPageId="4" captionText="FIGURE 1. Scyliorhinus ugoi sp. nov., holotype, MNRJ 42619, female, 500 mm TL. A) Dorsal view. B) Lateral view; paratype, UERJ 2179, male, 419 mm TL. C) Dorsal view. D) Lateral view. Scale bar: 5 cm." httpUri="https://zenodo.org/record/243846/files/figure.png" pageId="5" pageNumber="352">Figs. 1</figureCitation>
and 
<figureCitation id="137F2A0FFF8CFF9E16DCFAF4FE3CFACD" box="[485,500,1327,1351]" captionStart="FIGURE 2" captionStartId="6.[151,260,1897,1919]" captionTargetBox="[346,1240,665,1868]" captionTargetId="figure@6.[340,1248,661,1875]" captionTargetPageId="6" captionText="FIGURE 2. Scyliorhinus ugoi sp. nov. A) Dorsal view of the head of a female, MNRJ 42619, female, 500 mm TL. B) Dorsal view of the head of a male, UERJ 2179, 419 mm TL. C) Lateral view of head, UERJ 2179 male, 419 mm TL. D) First dorsal fin. E) Second dorsal fin. F) Anal fin. G) Caudal fin. H) Ventral view of trunk, MNRJ 42619, female, 500 mm TL. Scale bars: 2 cm." httpUri="https://zenodo.org/record/243847/files/figure.png" pageId="5" pageNumber="352">2</figureCitation>
) large and rounded-triangular, not falcate, with narrowly rounded apexes and broad bases. Origin of pectorals under interspace between third and fourth gill openings. Pectoral base 0.3– 0.7 times mouth width (
<emphasis id="B930EA98FF8CFF9E1603FAACFE91FA04" box="[314,345,1399,1422]" italics="true" pageId="5" pageNumber="352">vs.</emphasis>
0.8 times in the 
<emphasis id="B930EA98FF8CFF9E1520FAADFCB0FA04" box="[537,888,1398,1422]" italics="true" pageId="5" pageNumber="352">
<taxonomicName id="4C444D09FF8CFF9E1520FAADFCC5FA04" box="[537,781,1398,1422]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="5" pageNumber="352" phylum="Chordata" rank="species" species="haeckelii">Scyliorhinus haeckelii</taxonomicName>
/ 
<taxonomicName id="4C444D09FF8CFF9E1423FAADFCB0FA04" box="[794,888,1398,1422]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="5" pageNumber="352" phylum="Chordata" rank="species" species="besnardi">besnardi</taxonomicName>
</emphasis>
group). Pectoral anterior margin 2–4.8 times its base (
<emphasis id="B930EA98FF8CFF9E17E5FA40FF32FA38" box="[220,250,1435,1458]" italics="true" pageId="5" pageNumber="352">vs.</emphasis>
<quantity id="4CBC9B6FFF8CFF9E1638FA40FEB1FA39" box="[257,377,1434,1459]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" metricValueMax="5.842" metricValueMin="4.318" pageId="5" pageNumber="352" unit="in" value="2.0" valueMax="2.3" valueMin="1.7">1.7– 2.3 in</quantity>
the 
<emphasis id="B930EA98FF8CFF9E1694FA40FCC5FA39" box="[429,781,1435,1459]" italics="true" pageId="5" pageNumber="352">
<taxonomicName id="4C444D09FF8CFF9E1694FA40FD6BFA39" box="[429,675,1435,1459]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="5" pageNumber="352" phylum="Chordata" rank="species" species="haeckelii">Scyliorhinus haeckelii</taxonomicName>
/ 
<taxonomicName id="4C444D09FF8CFF9E1596FA40FCC5FA39" box="[687,781,1435,1459]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="5" pageNumber="352" phylum="Chordata" rank="species" species="besnardi">besnardi</taxonomicName>
</emphasis>
group) and 1.6 times the posterior margin (
<tableCitation id="C6C60331FF8CFF9E1235FA41FA9AFA39" box="[1292,1362,1434,1459]" captionStart="TABLE 1" captionStartId="2.[151,239,152,175]" captionTargetBox="[159,1430,276,1951]" captionTargetPageId="2" captionText="TABLE 1. Measurements in millimeters and proportions as percentage of total length for Scyliorhinus ugoi sp. nov. Mean (x), standard deviation (SD) and range of proportions (%)." httpUri="http://table.plazi.org/id/DF3B6602FF8BFF9917AEFF43FC4FFF45" pageId="5" pageNumber="352" tableUuid="DF3B6602FF8BFF9917AEFF43FC4FFF45">Tab. 1</tableCitation>
).
</paragraph>
<paragraph id="8BFB368AFF8CFF9E17FEFA65FA50F97D" blockId="5.[151,1437,1182,2035]" pageId="5" pageNumber="352">
Pectoral fin skeleton aplesodic (
<figureCitation id="137F2A0FFF8CFF9E1577FA65FD51FA5D" box="[590,665,1470,1495]" captionStart="FIGURE 6" captionStartId="8.[151,249,1888,1910]" captionTargetBox="[477,1087,1188,1844]" captionTargetId="figure@8.[469,1097,1180,1853]" captionTargetPageId="8" captionText="FIGURE 6. Pectoral fin skeleton of Scyliorhinus ugoi sp. nov., UERJ 1722, female, 600 mm TL. Scale bar: 2 cm. DRA, distal segment; MES, mesopterygium; MET, metapterygium; MTS, metapterygial axis; PRA, proximal segment; PRO, propterygium." httpUri="https://zenodo.org/record/243851/files/figure.png" pageId="5" pageNumber="352">Fig. 6</figureCitation>
) with radials mostly divided into three segments, longest distal radial segment (DRA) 1.8 times length of its proximal radial segment (PRA). Pectoral skeleton tribasal, propterygium (
<collectingCountry id="F353761AFF8CFF9E1607F9DCFEB0F995" box="[318,376,1543,1567]" name="Palau" pageId="5" pageNumber="352">PRO</collectingCountry>
) with a single radial, mesopterygium (MES) with 3 radials, and metapterygium (MET) with 8 (7–9) radials on basal segment and 1 (1–2) on metapterygial axis (MTS); total radial count 13 (12–15). Propterygium small, short and rectangular or trapezoidal. Mesopterygium short, diamond-shaped and slightly elongated and distally expanded in the axes of its radials. A fused plate formed by the proximal segment of the second, third and fourth mesopterygial radials. Metapterygium basal segment triangular, elongated slightly diagonal to the axes of its radials; metapterygial axis quadrangular, trisegmental, and with length about 0.4 times of metapterygial basal segment. This segment presents a projection directed anteriorly in its inner margin.
</paragraph>
<paragraph id="8BFB368AFF8CFF9E17FEF8D9FBA8F8B5" blockId="5.[151,1437,1182,2035]" pageId="5" pageNumber="352">
Pelvic fins (
<figureCitation id="137F2A0FFF8CFF9E1668F8D9FE5EF891" box="[337,406,1794,1819]" captionStart="FIGURE 2" captionStartId="6.[151,260,1897,1919]" captionTargetBox="[346,1240,665,1868]" captionTargetId="figure@6.[340,1248,661,1875]" captionTargetPageId="6" captionText="FIGURE 2. Scyliorhinus ugoi sp. nov. A) Dorsal view of the head of a female, MNRJ 42619, female, 500 mm TL. B) Dorsal view of the head of a male, UERJ 2179, 419 mm TL. C) Lateral view of head, UERJ 2179 male, 419 mm TL. D) First dorsal fin. E) Second dorsal fin. F) Anal fin. G) Caudal fin. H) Ventral view of trunk, MNRJ 42619, female, 500 mm TL. Scale bars: 2 cm." httpUri="https://zenodo.org/record/243847/files/figure.png" pageId="5" pageNumber="352">Fig. 2</figureCitation>
) broadly triangular; pelvic anterior margins 0.5 times of pectoral anterior margins. Pelvic anterior margin 0.9–1.1 times posterior margin (
<tableCitation id="C6C60331FF8CFF9E158EF8FDFD32F8B5" box="[695,762,1830,1855]" captionStart="TABLE 1" captionStartId="2.[151,239,152,175]" captionTargetBox="[159,1430,276,1951]" captionTargetPageId="2" captionText="TABLE 1. Measurements in millimeters and proportions as percentage of total length for Scyliorhinus ugoi sp. nov. Mean (x), standard deviation (SD) and range of proportions (%)." httpUri="http://table.plazi.org/id/DF3B6602FF8BFF9917AEFF43FC4FFF45" pageId="5" pageNumber="352" tableUuid="DF3B6602FF8BFF9917AEFF43FC4FFF45">Tab.1</tableCitation>
). Total radial count 13 (12–15).
</paragraph>
<paragraph id="8BFB368AFF8CFF9E17FEF891FA55F879" blockId="5.[151,1437,1182,2035]" pageId="5" pageNumber="352">
Claspers (
<figureCitation id="137F2A0FFF8CFF9E1601F891FEB4F8E9" box="[312,380,1866,1891]" captionStart="FIGURE 7" captionStartId="9.[151,249,988,1010]" captionTargetBox="[356,1216,199,957]" captionTargetId="figure@9.[340,1247,193,967]" captionTargetPageId="9" captionText="FIGURE 7. Clasper of Scyliorhinus ugoi sp. nov., UERJ 1725, male, 530 mm TL. A) Clasper gland. B) Skeleton. Scale bar = 5 mm. AP, apopyle; CRH, cover rhipidion; G, end-style; EN, envelope; ERH, exorhipidion; HP, hypopyle; RD, marginal dorsal cartilage; RD 2, accessory marginal dorsal cartilage; RH, rhipidion; RV, marginal ventral cartilage; TD, terminal dorsal cartilage; TDC, terminal dermal cover; TV, terminal ventral cartilage; TV 2, terminal ventral cartilage 2." httpUri="https://zenodo.org/record/243852/files/figure.png" pageId="5" pageNumber="352">Fig. 7</figureCitation>
) moderately long and slender, cylindrical and blunt-tipped, extending free rear tips of pelvic fins, by a distance about 0.2 times pelvic inner margin. Most of clasper except dorsomedial and posteromedial surface of glans (including rhipidion) covered by large denticles with anteriorly directed cusps. An envelope (EN) delimits anteriorly an exorhipidion (ERH) not differentiated from lateral edge of glans, specialized clasper hooks absent. Rhipidion (RH) well developed medially covered by a poorly developed cover rhipidion (CRH).
</paragraph>
<paragraph id="8BFB368AFF8FFF9D17AEFF4CFECAFDEA" blockId="6.[151,1436,151,608]" pageId="6" pageNumber="353">
Pseudosiphon poorly developed and visualized only internally; pseudopera absent. Apopyle (AP) and hypopyle (HP) connected by a long clasper groove, with its dorsal margins fused over the clasper groove. Clasper skeleton (
<figureCitation id="137F2A0FFF8FFF9D17A6FF04FF2FFF72" box="[159,231,223,248]" captionStart="FIGURE 7" captionStartId="9.[151,249,988,1010]" captionTargetBox="[356,1216,199,957]" captionTargetId="figure@9.[340,1247,193,967]" captionTargetPageId="9" captionText="FIGURE 7. Clasper of Scyliorhinus ugoi sp. nov., UERJ 1725, male, 530 mm TL. A) Clasper gland. B) Skeleton. Scale bar = 5 mm. AP, apopyle; CRH, cover rhipidion; G, end-style; EN, envelope; ERH, exorhipidion; HP, hypopyle; RD, marginal dorsal cartilage; RD 2, accessory marginal dorsal cartilage; RH, rhipidion; RV, marginal ventral cartilage; TD, terminal dorsal cartilage; TDC, terminal dermal cover; TV, terminal ventral cartilage; TV 2, terminal ventral cartilage 2." httpUri="https://zenodo.org/record/243852/files/figure.png" pageId="6" pageNumber="353">Fig. 7</figureCitation>
b) relatively simple. The axial cartilage connected proximally by a single basal segment and a dorsal beta cartilage to the pelvic basipterygium. Clasper shaft slender and formed from the axial cartilage and tightly rolled dorsal and ventral marginal (RD and RV) cartilages. A poorly developed acessory dorsal marginal cartilage (RD2) present between the terminal portion of the dorsal marginal cartilage and anterior portion of the dorsal terminal (TD); this cartilage supports the rhipidion. The skeleton of the clasper glans comprises a large, curved, wedge-shaped dorsal terminal and a similarly shaped ventral terminal (TV), articulating with and separated by a terminal extension of the axial cartilage, the narrow end-style (G); ventral terminal longer than dorsal terminal cartilage. There is no accessory terminal cartilage or dorsal terminal 2 cartilage, but a slender ventral terminal cartilage (TV2) is positioned on the ventral terminal cartilage. A terminal dermal cover (TDC – new term) present in the hindmost portion of the clasper, covering the terminal cartilages and presenting no denticles.
</paragraph>
<caption id="DF3B6602FF8FFF9D17AEF8B2FEA7F856" httpUri="https://zenodo.org/record/243847/files/figure.png" pageId="6" pageNumber="353" targetBox="[346,1240,665,1868]" targetPageId="6">
<paragraph id="8BFB368AFF8FFF9D17AEF8B2FEA7F856" blockId="6.[151,1436,1897,2012]" pageId="6" pageNumber="353">
<emphasis id="B930EA98FF8FFF9D17AEF8B2FEECF8F5" bold="true" box="[151,292,1897,1919]" pageId="6" pageNumber="353">FIGURE 2.</emphasis>
<taxonomicName id="4C444D09FF8FFF9D1609F8B2FE2CF8F5" box="[304,484,1897,1919]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="6" pageNumber="353" phylum="Chordata" rank="species" species="ugoi" status="sp. nov.">
<emphasis id="B930EA98FF8FFF9D1609F8B2FE2CF8F5" box="[304,484,1897,1919]" italics="true" pageId="6" pageNumber="353">Scyliorhinus ugoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20357E3FF8FFF9D16D4F8B2FD82F8F5" box="[493,586,1897,1919]" pageId="6" pageNumber="353" rank="species">
<emphasis id="B930EA98FF8FFF9D16D4F8B2FD82F8F5" bold="true" box="[493,586,1897,1919]" pageId="6" pageNumber="353">sp. nov.</emphasis>
</taxonomicNameLabel>
A) Dorsal view of the head of a female, MNRJ 42619, female, 500 mm TL. B) Dorsal view of the head of a male, UERJ 2179, 419 mm TL. C) Lateral view of head, UERJ 2179 male, 419 mm TL. D) First dorsal fin. E) Second dorsal fin. F) Anal fin. G) Caudal fin. H) Ventral view of trunk, MNRJ 42619, female, 500 mm TL. Scale bars: 2 cm.
</paragraph>
</caption>
<caption id="DF3B6602FF8EFF9C17AEFD53FB6EFD14" box="[151,1190,648,670]" httpUri="https://zenodo.org/record/243848/files/figure.png" pageId="7" pageNumber="354" targetBox="[436,1163,193,607]" targetPageId="7">
<paragraph id="8BFB368AFF8EFF9C17AEFD53FB6EFD14" blockId="7.[151,1190,648,670]" box="[151,1190,648,670]" pageId="7" pageNumber="354">
<emphasis id="B930EA98FF8EFF9C17AEFD53FEDAFD14" bold="true" box="[151,274,648,670]" pageId="7" pageNumber="354">FIGURE 3.</emphasis>
Eye of 
<taxonomicName id="4C444D09FF8EFF9C165DFD53FDD1FD14" box="[356,537,648,670]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="7" pageNumber="354" phylum="Chordata" rank="species" species="ugoi" status="sp. nov.">
<emphasis id="B930EA98FF8EFF9C165DFD53FDD1FD14" box="[356,537,648,670]" italics="true" pageId="7" pageNumber="354">Scyliorhinus ugoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20357E3FF8EFF9C1519FD53FDB3FD14" box="[544,635,648,670]" pageId="7" pageNumber="354" rank="species">
<emphasis id="B930EA98FF8EFF9C1519FD53FDB3FD14" bold="true" box="[544,635,648,670]" pageId="7" pageNumber="354">sp. nov.</emphasis>
</taxonomicNameLabel>
, MNRJ 42619, female, 500 mm TL. Scale bar: 2 cm.
</paragraph>
</caption>
<caption id="DF3B6602FF8EFF9C17AEFA75FAF2FA4E" box="[151,1338,1454,1476]" httpUri="https://zenodo.org/record/243849/files/figure.png" pageId="7" pageNumber="354" targetBox="[495,1143,740,1431]" targetPageId="7">
<paragraph id="8BFB368AFF8EFF9C17AEFA75FAF2FA4E" blockId="7.[151,1338,1454,1476]" box="[151,1338,1454,1476]" pageId="7" pageNumber="354">
<emphasis id="B930EA98FF8EFF9C17AEFA75FEDAFA4E" bold="true" box="[151,274,1454,1476]" pageId="7" pageNumber="354">FIGURE 4.</emphasis>
Nostrils and mouth of 
<taxonomicName id="4C444D09FF8EFF9C16C7FA75FD79FA4E" box="[510,689,1454,1476]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="7" pageNumber="354" phylum="Chordata" rank="species" species="ugoi" status="sp. nov.">
<emphasis id="B930EA98FF8EFF9C16C7FA75FD79FA4E" box="[510,689,1454,1476]" italics="true" pageId="7" pageNumber="354">Scyliorhinus ugoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20357E3FF8EFF9C158EFA75FCD9FA4E" box="[695,785,1454,1476]" pageId="7" pageNumber="354" rank="species">
<emphasis id="B930EA98FF8EFF9C158EFA75FCD9FA4E" bold="true" box="[695,785,1454,1476]" pageId="7" pageNumber="354">sp. nov.</emphasis>
</taxonomicNameLabel>
, MNRJ 42619, female, 500 mm TL. Scale bar: 2 cm.
</paragraph>
</caption>
<paragraph id="8BFB368AFF8EFF9C17FEFA2AFC29F9FF" blockId="7.[151,1437,1521,2014]" pageId="7" pageNumber="354">
First dorsal fin (
<figureCitation id="137F2A0FFF8EFF9C16BBFA2AFE1AF983" box="[386,466,1521,1546]" captionStart="FIGURE 1" captionStartId="4.[151,249,1130,1152]" captionTargetBox="[371,1214,204,1099]" captionTargetId="figure@4.[362,1226,195,1105]" captionTargetPageId="4" captionText="FIGURE 1. Scyliorhinus ugoi sp. nov., holotype, MNRJ 42619, female, 500 mm TL. A) Dorsal view. B) Lateral view; paratype, UERJ 2179, male, 419 mm TL. C) Dorsal view. D) Lateral view. Scale bar: 5 cm." httpUri="https://zenodo.org/record/243846/files/figure.png" pageId="7" pageNumber="354">Figs. 1</figureCitation>
, 
<figureCitation id="137F2A0FFF8EFF9C16DAFA2AFE3AF983" box="[483,498,1521,1545]" captionStart="FIGURE 2" captionStartId="6.[151,260,1897,1919]" captionTargetBox="[346,1240,665,1868]" captionTargetId="figure@6.[340,1248,661,1875]" captionTargetPageId="6" captionText="FIGURE 2. Scyliorhinus ugoi sp. nov. A) Dorsal view of the head of a female, MNRJ 42619, female, 500 mm TL. B) Dorsal view of the head of a male, UERJ 2179, 419 mm TL. C) Lateral view of head, UERJ 2179 male, 419 mm TL. D) First dorsal fin. E) Second dorsal fin. F) Anal fin. G) Caudal fin. H) Ventral view of trunk, MNRJ 42619, female, 500 mm TL. Scale bars: 2 cm." httpUri="https://zenodo.org/record/243847/files/figure.png" pageId="7" pageNumber="354">2</figureCitation>
) triangular, apically narrow and not falcate, with nearly straight anterior margin, rounded apex and angular free rear tip. First dorsal origin above to insertions of pelvic fins. First dorsal base 1.6 times in interdorsal space, 3.4 times in dorsal caudal margin. First dorsal anterior margin 1.4 times its base; first dorsal height 0.8 times first dorsal base. Total radial count 13 (12–14).
</paragraph>
<paragraph id="8BFB368AFF8EFF9C17FEF95AFBEFF8A3" blockId="7.[151,1437,1521,2014]" pageId="7" pageNumber="354">
Second dorsal fin (
<figureCitation id="137F2A0FFF8EFF9C1699F95AFE26F913" box="[416,494,1665,1690]" captionStart="FIGURE 1" captionStartId="4.[151,249,1130,1152]" captionTargetBox="[371,1214,204,1099]" captionTargetId="figure@4.[362,1226,195,1105]" captionTargetPageId="4" captionText="FIGURE 1. Scyliorhinus ugoi sp. nov., holotype, MNRJ 42619, female, 500 mm TL. A) Dorsal view. B) Lateral view; paratype, UERJ 2179, male, 419 mm TL. C) Dorsal view. D) Lateral view. Scale bar: 5 cm." httpUri="https://zenodo.org/record/243846/files/figure.png" pageId="7" pageNumber="354">Figs. 1</figureCitation>
, 
<figureCitation id="137F2A0FFF8EFF9C16C7F95AFDC5F913" box="[510,525,1665,1689]" captionStart="FIGURE 2" captionStartId="6.[151,260,1897,1919]" captionTargetBox="[346,1240,665,1868]" captionTargetId="figure@6.[340,1248,661,1875]" captionTargetPageId="6" captionText="FIGURE 2. Scyliorhinus ugoi sp. nov. A) Dorsal view of the head of a female, MNRJ 42619, female, 500 mm TL. B) Dorsal view of the head of a male, UERJ 2179, 419 mm TL. C) Lateral view of head, UERJ 2179 male, 419 mm TL. D) First dorsal fin. E) Second dorsal fin. F) Anal fin. G) Caudal fin. H) Ventral view of trunk, MNRJ 42619, female, 500 mm TL. Scale bars: 2 cm." httpUri="https://zenodo.org/record/243847/files/figure.png" pageId="7" pageNumber="354">2</figureCitation>
) low, triangular, apically narrow and not falcate; second dorsal fin height 0.7 times first dorsal fin height, base 0.8 times first dorsal fin base. Second dorsal fin with nearly straight anterior margin, bluntly rounded apex, narrowly rounded free rear tip, and straight inner margin. Second dorsal fin origin slightly behind anal midbase. Second dorsal fin base 0.7 times in dorsocaudal space, second dorsal fin height 1.4 times in second dorsal fin base. Total radial count 10 (9–12).
</paragraph>
<paragraph id="8BFB368AFF8EFF9317FEF8EEFE96FF5E" blockId="7.[151,1437,1521,2014]" lastBlockId="8.[151,1437,151,357]" lastPageId="8" lastPageNumber="355" pageId="7" pageNumber="354">
Anal fin low (
<figureCitation id="137F2A0FFF8EFF9C1649F8EEFE09F8C7" box="[368,449,1845,1870]" captionStart="FIGURE 1" captionStartId="4.[151,249,1130,1152]" captionTargetBox="[371,1214,204,1099]" captionTargetId="figure@4.[362,1226,195,1105]" captionTargetPageId="4" captionText="FIGURE 1. Scyliorhinus ugoi sp. nov., holotype, MNRJ 42619, female, 500 mm TL. A) Dorsal view. B) Lateral view; paratype, UERJ 2179, male, 419 mm TL. C) Dorsal view. D) Lateral view. Scale bar: 5 cm." httpUri="https://zenodo.org/record/243846/files/figure.png" pageId="7" pageNumber="354">Figs. 1</figureCitation>
, 
<figureCitation id="137F2A0FFF8EFF9C16EAF8EEFE2BF8C7" box="[467,483,1845,1869]" captionStart="FIGURE 2" captionStartId="6.[151,260,1897,1919]" captionTargetBox="[346,1240,665,1868]" captionTargetId="figure@6.[340,1248,661,1875]" captionTargetPageId="6" captionText="FIGURE 2. Scyliorhinus ugoi sp. nov. A) Dorsal view of the head of a female, MNRJ 42619, female, 500 mm TL. B) Dorsal view of the head of a male, UERJ 2179, 419 mm TL. C) Lateral view of head, UERJ 2179 male, 419 mm TL. D) First dorsal fin. E) Second dorsal fin. F) Anal fin. G) Caudal fin. H) Ventral view of trunk, MNRJ 42619, female, 500 mm TL. Scale bars: 2 cm." httpUri="https://zenodo.org/record/243847/files/figure.png" pageId="7" pageNumber="354">2</figureCitation>
), apically narrow, not falcate and somewhat larger than second dorsal fin, anal fin height 1.1 times second dorsal fin height and base 1.3 times second dorsal fin base. Anal fin anterior margin nearly straight, apex narrowly rounded, free rear tip acutely pointed, and inner margin straight. Anal fin base without preanal ridges, anal fin origin and anal fin base length behind pelvic fins insertions. Anal fin base 0.5 times interdorsal space (
<emphasis id="B930EA98FF8EFF9C1683F81DFE10F857" box="[442,472,1990,2013]" italics="true" pageId="7" pageNumber="354">vs.</emphasis>
<quantity id="4CBC9B6FFF8EFF9C16D8F81EFD9BF854" box="[481,595,1989,2014]" metricMagnitude="-2" metricUnit="m" metricValue="1.7779999999999998" metricValueMax="2.032" metricValueMin="1.524" pageId="7" pageNumber="354" unit="in" value="0.7" valueMax="0.8" valueMin="0.6">0.6–0.8 in</quantity>
the 
<emphasis id="B930EA98FF8EFF9C15BFF81EFC2DF857" box="[646,997,1989,2013]" italics="true" pageId="7" pageNumber="354">
<taxonomicName id="4C444D09FF8EFF9C15BFF81EFCB2F857" box="[646,890,1989,2013]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="7" pageNumber="354" phylum="Chordata" rank="species" species="haeckelii">Scyliorhinus haeckelii</taxonomicName>
/ 
<taxonomicName id="4C444D09FF8EFF9C14BEF81EFC2DF857" box="[903,997,1989,2013]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="7" pageNumber="354" phylum="Chordata" rank="species" species="besnardi">besnardi</taxonomicName>
</emphasis>
group) and 1.3–1.5 times dorsocaudal space. Anal anterior margin 1.5–1.7 times posterior one, anal fin height 1.8 times in anal fin base. Total radial count 20 (18–22).
</paragraph>
<paragraph id="8BFB368AFF81FF9317FEFF04FF30FEEF" blockId="8.[151,1437,151,357]" pageId="8" pageNumber="355">
Caudal fin (
<figureCitation id="137F2A0FFF81FF931660FF04FE65FF72" box="[345,429,223,248]" captionStart="FIGURE 1" captionStartId="4.[151,249,1130,1152]" captionTargetBox="[371,1214,204,1099]" captionTargetId="figure@4.[362,1226,195,1105]" captionTargetPageId="4" captionText="FIGURE 1. Scyliorhinus ugoi sp. nov., holotype, MNRJ 42619, female, 500 mm TL. A) Dorsal view. B) Lateral view; paratype, UERJ 2179, male, 419 mm TL. C) Dorsal view. D) Lateral view. Scale bar: 5 cm." httpUri="https://zenodo.org/record/243846/files/figure.png" pageId="8" pageNumber="355">Figs. 1</figureCitation>
, 
<figureCitation id="137F2A0FFF81FF9316FBFF3BFE19FF72" box="[450,465,224,248]" captionStart="FIGURE 2" captionStartId="6.[151,260,1897,1919]" captionTargetBox="[346,1240,665,1868]" captionTargetId="figure@6.[340,1248,661,1875]" captionTargetPageId="6" captionText="FIGURE 2. Scyliorhinus ugoi sp. nov. A) Dorsal view of the head of a female, MNRJ 42619, female, 500 mm TL. B) Dorsal view of the head of a male, UERJ 2179, 419 mm TL. C) Lateral view of head, UERJ 2179 male, 419 mm TL. D) First dorsal fin. E) Second dorsal fin. F) Anal fin. G) Caudal fin. H) Ventral view of trunk, MNRJ 42619, female, 500 mm TL. Scale bars: 2 cm." httpUri="https://zenodo.org/record/243847/files/figure.png" pageId="8" pageNumber="355">2</figureCitation>
) narrow-lobed and asymmetrical, with large and hardly developed terminal lobe. Caudal fin short, dorsal margin 3.2 times in precaudal length; subterminal margin 0.9–1 time in terminal margin. Dorsal caudal fin margin slightly convex, without lateral undulations. No crests of denticles on the caudal fin margins.
</paragraph>
<caption id="DF3B6602FF81FF9317AEFBCCFE91FBE0" httpUri="https://zenodo.org/record/243850/files/figure.png" pageId="8" pageNumber="355" targetBox="[341,1245,447,1021]" targetPageId="8">
<paragraph id="8BFB368AFF81FF9317AEFBCCFE91FBE0" blockId="8.[151,1436,1047,1130]" pageId="8" pageNumber="355">
<emphasis id="B930EA98FF81FF9317AEFBCCFEDAFBA7" bold="true" box="[151,274,1047,1069]" pageId="8" pageNumber="355">FIGURE 5.</emphasis>
Lateral trunk denticles of 
<taxonomicName id="4C444D09FF81FF931526FBCCFD19FBA7" box="[543,721,1047,1069]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="8" pageNumber="355" phylum="Chordata" rank="species" species="ugoi" status="sp. nov.">
<emphasis id="B930EA98FF81FF931526FBCCFD19FBA7" box="[543,721,1047,1069]" italics="true" pageId="8" pageNumber="355">Scyliorhinus ugoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20357E3FF81FF9315E1FBCCFCF9FBA7" box="[728,817,1047,1069]" pageId="8" pageNumber="355" rank="species">
<emphasis id="B930EA98FF81FF9315E1FBCCFCF9FBA7" bold="true" box="[728,817,1047,1069]" pageId="8" pageNumber="355">sp. nov.</emphasis>
</taxonomicNameLabel>
A–C) UERJ 1726, female, 602 mm TL. D–F) UERJ 1725, adult male, 534 mm TL. A, D) Region above pectoral fin. B, E) Region below first dorsal fin. C, F) Region below second dorsal fin.
</paragraph>
</caption>
<caption id="DF3B6602FF81FF9317AEF8BBFEEFF83E" httpUri="https://zenodo.org/record/243851/files/figure.png" pageId="8" pageNumber="355" targetBox="[477,1087,1188,1844]" targetPageId="8">
<paragraph id="8BFB368AFF81FF9317AEF8BBFEEFF83E" blockId="8.[151,1436,1888,1972]" pageId="8" pageNumber="355">
<emphasis id="B930EA98FF81FF9317AEF8BBFEDCF8FC" bold="true" box="[151,276,1888,1910]" pageId="8" pageNumber="355">FIGURE 6.</emphasis>
Pectoral fin skeleton of 
<taxonomicName id="4C444D09FF81FF931521F8BBFD03F8FC" box="[536,715,1888,1910]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="8" pageNumber="355" phylum="Chordata" rank="species" species="ugoi" status="sp. nov.">
<emphasis id="B930EA98FF81FF931521F8BBFD03F8FC" box="[536,715,1888,1910]" italics="true" pageId="8" pageNumber="355">Scyliorhinus ugoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20357E3FF81FF9315EDF8BBFCE7F8FC" box="[724,815,1888,1910]" pageId="8" pageNumber="355" rank="species">
<emphasis id="B930EA98FF81FF9315EDF8BBFCE7F8FC" bold="true" box="[724,815,1888,1910]" pageId="8" pageNumber="355">sp. nov.</emphasis>
</taxonomicNameLabel>
, UERJ 1722, female, 600 mm TL. Scale bar: 2 cm. DRA, distal segment; MES, mesopterygium; MET, metapterygium; MTS, metapterygial axis; PRA, proximal segment; PRO, propterygium.
</paragraph>
</caption>
<caption id="DF3B6602FF80FF9217AEFC07FB76FBC4" httpUri="https://zenodo.org/record/243852/files/figure.png" pageId="9" pageNumber="356" targetBox="[356,1216,199,957]" targetPageId="9">
<paragraph id="8BFB368AFF80FF9217AEFC07FB76FBC4" blockId="9.[151,1436,988,1102]" pageId="9" pageNumber="356">
<emphasis id="B930EA98FF80FF9217AEFC07FED9FC78" bold="true" box="[151,273,988,1010]" pageId="9" pageNumber="356">FIGURE 7.</emphasis>
Clasper of 
<taxonomicName id="4C444D09FF80FF9216BFFC07FDFFFC78" box="[390,567,988,1010]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="9" pageNumber="356" phylum="Chordata" rank="species" species="ugoi" status="sp. nov.">
<emphasis id="B930EA98FF80FF9216BFFC07FDFFFC78" box="[390,567,988,1010]" italics="true" pageId="9" pageNumber="356">Scyliorhinus ugoi</emphasis>
</taxonomicName>
<emphasis id="B930EA98FF80FF921504FC07FD5EFC78" bold="true" box="[573,662,988,1010]" pageId="9" pageNumber="356">
<taxonomicNameLabel id="A20357E3FF80FF921504FC07FD5EFC78" box="[573,662,988,1010]" pageId="9" pageNumber="356" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, UERJ 1725, male, 530 mm TL. A) Clasper gland. B) Skeleton. Scale bar = 5 mm. AP, apopyle; CRH, cover rhipidion; G, end-style; EN, envelope; ERH, exorhipidion; HP, hypopyle; RD, marginal dorsal cartilage; RD2, accessory marginal dorsal cartilage; RH, rhipidion; RV, marginal ventral cartilage; TD, terminal dorsal cartilage; TDC, terminal dermal cover; TV, terminal ventral cartilage; TV2, terminal ventral cartilage 2.
</paragraph>
</caption>
<paragraph id="8BFB368AFF80FF9217FEFB4FFD03FA92" blockId="9.[151,1437,1172,2025]" pageId="9" pageNumber="356">Total vertebral counts 127 (119–135), monospondylous precaudal centra (MP) 39 (38–39). Transition between monospondylous and diplospondylous (DP) centra behind pelvic fin bases and over clasper shafts. Last MP centra before MP–DP transition smaller than the anterior and larger than the first DP, not forming a 'stutter- zone' of alternating long and short centra.</paragraph>
<paragraph id="8BFB368AFF80FF9217FEFAFFFCFDFAB6" blockId="9.[151,1437,1172,2025]" box="[199,821,1316,1341]" pageId="9" pageNumber="356">
Intestinal valve of conicospiral 
<typeStatus id="54FF8828FF80FF92151EFAFEFD92FAB7" box="[551,602,1317,1341]" pageId="9" pageNumber="356">type</typeStatus>
, with 6 (6–8) turns.
</paragraph>
<paragraph id="8BFB368AFF80FF9217FEFA93FC5DF84F" blockId="9.[151,1437,1172,2025]" pageId="9" pageNumber="356">
Cranium (
<figureCitation id="137F2A0FFF80FF921607FA93FE40FAEB" box="[318,392,1352,1377]" captionStart="FIGURE 8" captionStartId="10.[151,249,1319,1341]" captionTargetBox="[356,1227,308,1298]" captionTargetId="figure@10.[348,1240,301,1298]" captionTargetPageId="10" captionText="FIGURE 8. Cranium of Scyliorhinus ugoi sp. nov., UERJ 1725, male, 530 mm TL. A) Dorsal view. B) Ventral view; UERJ 1726, female, 597 mm TL. C) Dorsal view. D) Ventral view. Scale bar = 1 cm. AF, anterior fontanelle; ASC, anterior semicircular canal; CRF, cranial roof; FOE, external foramen of preorbital canal; FPE, external profundus foramen; HF, hyomandibular facet; ICF, internal carotid foramen; IS, internasal septum; LR, lateral rostral cartilage; MR, medial rostral cartilage; NA, nasal aperture; NC, nasal capsule; NF, nasal fontanelle; OC, optic capsule; PEP, preorbital process; PRF, parietal fossa; PSC, posterior semicircular canal; PT, pterotic process; PTP, postorbital process; SC, supraorbital crest; SF, stapedial foramen; SG, supraorbital groove; SS, suborbital shelf." httpUri="https://zenodo.org/record/243853/files/figure.png" pageId="9" pageNumber="356">Fig. 8</figureCitation>
) broad and flattened, corresponding to 10% TL (
<tableCitation id="C6C60331FF80FF9214DFFA93FBF2FAEB" box="[998,1082,1352,1377]" captionStart="TABLE 2" captionStartId="5.[151,239,152,175]" captionTargetBox="[159,1335,317,628]" captionTargetPageId="5" captionText="TABLE 2. Measurements in millimeters and proportions as percentage of nasobasal length for Scyliorhinus ugoi sp. nov. Female (F), male (M), and range of proportions (% NL)." httpUri="http://table.plazi.org/id/DF3B6602FF8CFF9E17AEFF43FD20FF45" pageId="9" pageNumber="356" tableUuid="DF3B6602FF8CFF9E17AEFF43FD20FF45">Tab. 2</tableCitation>
). Rostral cartilages short and slender; rostrum length 20.8–29.3% of nasobasal length (NL). Medial rostral cartilage (MR) originates from anteroventral tip of internasal septum and extends anterodorsally. Distance between lateral rostral cartilages (LR) proportionately larger on females (13.8–14.5% NL) than males (12.2–13% NL). Nasal capsules (NC) large, transversely oval, its width larger than length. Width across capsules 76– 85.4% NL. Nasal apertures (NA) broadly circular and laterally positioned on capsules; two thirds of its area covered by nasal fontanelle (NF). Males with nasal apertures proportionately larger (26.5– 27.7% NL) than females (25– 25.8% NL). Internasal septum (IS) high and compressed, not hypercalcified in adults. Distance between rostral base and anterior edge of anterior fontanelle (AF) proportionately larger in females (39.5% NL) than males (36.7– 38.8% NL). Cranial roof (CRF) broadly arched between supraorbital crests (SC) in frontal view, with a definite saddle-like medial depression in lateral view. Anterior fontanelle broad and subquadrate on males and heart-shaped on females, length 1.3 times width, width 4 times in nasobasal length. Basal plate flat, without keels; width larger in females (39.6–45.8% NL) (
<emphasis id="B930EA98FF80FF9215ACF922FD7BF89A" box="[661,691,1785,1808]" italics="true" pageId="9" pageNumber="356">vs.</emphasis>
33.3–40.8% NL in the 
<emphasis id="B930EA98FF80FF9214EEF923FAF3F89A" box="[983,1339,1784,1808]" italics="true" pageId="9" pageNumber="356">
<taxonomicName id="4C444D09FF80FF9214EEF923FB19F89A" box="[983,1233,1784,1808]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="9" pageNumber="356" phylum="Chordata" rank="species" species="haeckelii">Scyliorhinus haeckelii</taxonomicName>
/ 
<taxonomicName id="4C444D09FF80FF9213E4F923FAF3F89A" box="[1245,1339,1784,1808]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="9" pageNumber="356" phylum="Chordata" rank="species" species="besnardi">besnardi</taxonomicName>
</emphasis>
group). Orbits oval-subquadrate in shape; its length 2.2 times in nasobasal length. Supraorbital crests low, strong and deeply arched. Postorbital processes (PTP) angular, situated on anterior ends of optic capsules; width across postorbital processes 1.2 times preorbital processes (PEP). Optic capsules short, length 4.7 times in nasobasal length; its width 2.5–2.7 times optic capsules length. Hyomandibular facets (HF) large, extending across the entire lateral face of each optic capsule, with slightly exserted ends.
</paragraph>
<paragraph id="8BFB368AFF80FF9117FEF80BFDA7FF7D" blockId="9.[151,1437,1172,2025]" lastBlockId="10.[151,1436,151,248]" lastPageId="10" lastPageNumber="357" pageId="9" pageNumber="356">
Color pattern with brown saddles on a light brown background, predorsal saddles with anterior and posterior sharp median projections; light and dark spots spiracle-sized or slightly smaller, no large white spots. Light brown on ventral region; some specimens with dark points on body and fins. Remarkably darker than the 
<emphasis id="B930EA98FF83FF91122BFF67FEAAFF72" italics="true" pageId="10" pageNumber="357">
<taxonomicName id="4C444D09FF83FF91122BFF67FF30FF72" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="10" pageNumber="357" phylum="Chordata" rank="species" species="haeckelii">Scyliorhinus haeckelii</taxonomicName>
/ 
<taxonomicName id="4C444D09FF83FF91163DFF3BFEAAFF72" box="[260,354,224,248]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="10" pageNumber="357" phylum="Chordata" rank="species" species="besnardi">besnardi</taxonomicName>
</emphasis>
group and S. 
<taxonomicName id="4C444D09FF83FF9116C4FF3BFDAFFF72" box="[509,615,224,248]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="10" pageNumber="357" phylum="Chordata" rank="species" species="hesperius">
<emphasis id="B930EA98FF83FF9116C4FF3BFDAFFF72" box="[509,615,224,248]" italics="true" pageId="10" pageNumber="357">hesperius</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<caption id="DF3B6602FF83FF9117AEFAFCFC9AFA7F" httpUri="https://zenodo.org/record/243853/files/figure.png" pageId="10" pageNumber="357" targetBox="[356,1227,308,1298]" targetPageId="10">
<paragraph id="8BFB368AFF83FF9117AEFAFCFC9AFA7F" blockId="10.[151,1436,1319,1525]" pageId="10" pageNumber="357">
<emphasis id="B930EA98FF83FF9117AEFAFCFEDBFAB7" bold="true" box="[151,275,1319,1341]" pageId="10" pageNumber="357">FIGURE 8.</emphasis>
Cranium of 
<taxonomicName id="4C444D09FF83FF9116A1FAFCFD81FAB4" box="[408,585,1319,1342]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="10" pageNumber="357" phylum="Chordata" rank="species" species="ugoi" status="sp. nov.">
<emphasis id="B930EA98FF83FF9116A1FAFCFD81FAB4" box="[408,585,1319,1342]" italics="true" pageId="10" pageNumber="357">Scyliorhinus ugoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20357E3FF83FF911576FAFCFD60FAB7" box="[591,680,1319,1341]" pageId="10" pageNumber="357" rank="species">
<emphasis id="B930EA98FF83FF911576FAFCFD60FAB7" bold="true" box="[591,680,1319,1341]" pageId="10" pageNumber="357">sp. nov.</emphasis>
</taxonomicNameLabel>
, UERJ 1725, male, 530 mm TL. A) Dorsal view. B) Ventral view; UERJ 1726, female, 597 mm TL. C) Dorsal view. D) Ventral view. Scale bar = 1 cm. AF, anterior fontanelle; ASC, anterior semicircular canal; CRF, cranial roof; FOE, external foramen of preorbital canal; FPE, external profundus foramen; HF, hyomandibular facet; ICF, internal carotid foramen; IS, internasal septum; LR, lateral rostral cartilage; MR, medial rostral cartilage; NA, nasal aperture; NC, nasal capsule; NF, nasal fontanelle; OC, optic capsule; PEP, preorbital process; PRF, parietal fossa; PSC, posterior semicircular canal; PT, pterotic process; PTP, postorbital process; SC, supraorbital crest; SF, stapedial foramen; SG, supraorbital groove; SS, suborbital shelf.
</paragraph>
</caption>
<subSubSection id="C35E6501FF83FF9117FEF9F9FD9DF829" pageId="10" pageNumber="357" type="distribution">
<paragraph id="8BFB368AFF83FF9117FEF9F9FD9DF829" blockId="10.[151,1437,1570,1955]" pageId="10" pageNumber="357">
<emphasis id="B930EA98FF83FF9117FEF9F9FD84F9B1" bold="true" box="[199,588,1570,1595]" pageId="10" pageNumber="357">Distribution and biological data.</emphasis>
This species (
<figureCitation id="137F2A0FFF83FF9115D4F9F9FCF9F9B1" box="[749,817,1570,1595]" captionStart="FIGURE 9" captionStartId="11.[151,249,987,1009]" captionTargetBox="[340,1247,193,965]" captionTargetId="figure@11.[340,1248,193,966]" captionTargetPageId="11" captionText="FIGURE 9. Distribution of Scyliorhinus ugoi sp. nov. Yellow diamonds represent records of type series, and white star locality of the holotype." httpUri="https://zenodo.org/record/243854/files/figure.png" pageId="10" pageNumber="357">Fig. 9</figureCitation>
) is recorded from the coast of the states of Rio Grande do Norte, Pernambuco, Alagoas, Bahia (northeastern 
<collectingCountry id="F353761AFF83FF911430F99DFC99F9D5" box="[777,849,1606,1631]" name="Brazil" pageId="10" pageNumber="357">Brazil</collectingCountry>
), Espírito Santo, and northern Rio de Janeiro (southeastern 
<collectingCountry id="F353761AFF83FF91160AF9B1FEB4F909" box="[307,380,1642,1667]" name="Brazil" pageId="10" pageNumber="357">Brazil</collectingCountry>
). Accurate occurrence records are known only for a few specimens, but this species has been collected for several years between Rio Grande do Norte and Rio de Janeiro states. Most specimens recorded are demersal, about 
<quantity id="4CBC9B6FFF83FF91166BF969FE1AF940" box="[338,466,1714,1739]" metricMagnitude="2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="10" pageNumber="357" unit="m" value="450.0" valueMax="500.0" valueMin="400.0">400–500 m</quantity>
deep, associated with coral formations, where it possibly lays its egg-cases. Stomach contents include small invertebrates (cephalopods), small bony fishes, and one hagfish (Class Myxini) egg-case was found in the stomach of an adult specimen. Maximum total length about 
<quantity id="4CBC9B6FFF83FF9114CEF921FB9EF898" box="[1015,1110,1786,1811]" metricMagnitude="-1" metricUnit="m" metricValue="6.3" pageId="10" pageNumber="357" unit="mm" value="630.0">630 mm</quantity>
. Size of the first maturity not well defined, but 
<quantity id="4CBC9B6FFF83FF911664F8C5FE73F8BC" box="[349,443,1822,1847]" metricMagnitude="-1" metricUnit="m" metricValue="4.5" pageId="10" pageNumber="357" unit="mm" value="450.0">450 mm</quantity>
TL males and 
<quantity id="4CBC9B6FFF83FF91155CF8C4FD0BF8BC" box="[613,707,1823,1847]" metricMagnitude="-1" metricUnit="m" metricValue="4.7" pageId="10" pageNumber="357" unit="mm" value="470.0">470 mm</quantity>
TL females are adults. Males ranging in size from 
<quantity id="4CBC9B6FFF83FF91123EF8C5FA55F8BC" box="[1287,1437,1822,1847]" metricMagnitude="-1" metricUnit="m" metricValue="4.765" metricValueMax="5.34" metricValueMin="4.1899999999999995" pageId="10" pageNumber="357" unit="mm" value="476.5" valueMax="534.0" valueMin="419.0">419–534 mm</quantity>
and females from 
<quantity id="4CBC9B6FFF83FF91165CF899FE35F8D0" box="[357,509,1858,1883]" metricMagnitude="-1" metricUnit="m" metricValue="5.305" metricValueMax="6.3" metricValueMin="4.31" pageId="10" pageNumber="357" unit="mm" value="530.5" valueMax="630.0" valueMin="431.0">431–630 mm</quantity>
. Reproduction is by oviparity, producing at least two amber colored egg capsules, one for each uterus, which are smooth, lacking longitudinal grooves and measuring about 
<quantity id="4CBC9B6FFF83FF9113BDF8BDFB28F8F4" box="[1156,1248,1894,1919]" metricMagnitude="-1" metricUnit="m" metricValue="1.52" pageId="10" pageNumber="357" unit="mm" value="152.0">152 mm</quantity>
long and 
<quantity id="4CBC9B6FFF83FF911277F8BDFA55F8F4" box="[1358,1437,1894,1919]" metricMagnitude="-2" metricUnit="m" metricValue="5.8" pageId="10" pageNumber="357" unit="mm" value="58.0">58 mm</quantity>
wide (
<bibRefCitation id="EFD54B7BFF83FF9117E5F851FEA3F829" author="Gadig" box="[220,363,1930,1955]" pageId="10" pageNumber="357" refString="Gadig, O. B. F. (2001) Tubaroes da costa brasileira. Unpubl. PhD thesis, Universidade Estadual Paulista ' Julio Mesquita Filho', Rio Claro, Sao Paulo, 343 pp." type="book" year="2001">Gadig, 2001</bibRefCitation>
; 
<bibRefCitation id="EFD54B7BFF83FF91164EF850FD81F829" author="Gomes" box="[375,585,1930,1955]" pageId="10" pageNumber="357" refString="Gomes, U. L., Signori, C. N., Gadig, O. B. F. &amp; Santos, H. R. S. (2010) Guia para Identificacao de Tubaroes e Raias do Rio de Janeiro. Technical Books, Rio de Janeiro, 234 pp." type="book" year="2010">
Gomes 
<emphasis id="B930EA98FF83FF9116F4F850FE36F828" box="[461,510,1930,1954]" italics="true" pageId="10" pageNumber="357">et al</emphasis>
, 2010
</bibRefCitation>
).
</paragraph>
</subSubSection>
<caption id="DF3B6602FF82FF9017AEFC00FE42FB9A" httpUri="https://zenodo.org/record/243854/files/figure.png" pageId="11" pageNumber="358" targetBox="[340,1247,193,965]" targetPageId="11">
<paragraph id="8BFB368AFF82FF9017AEFC00FE42FB9A" blockId="11.[151,1435,987,1040]" pageId="11" pageNumber="358">
<emphasis id="B930EA98FF82FF9017AEFC00FEDDFC7B" bold="true" box="[151,277,987,1009]" pageId="11" pageNumber="358">FIGURE 9.</emphasis>
Distribution of 
<taxonomicName id="4C444D09FF82FF9016FBFC00FDBDFC7B" box="[450,629,987,1009]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="ugoi" status="sp. nov.">
<emphasis id="B930EA98FF82FF9016FBFC00FDBDFC7B" box="[450,629,987,1009]" italics="true" pageId="11" pageNumber="358">Scyliorhinus ugoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20357E3FF82FF901546FC00FD2BFC7B" box="[639,739,987,1009]" pageId="11" pageNumber="358" rank="species">
<emphasis id="B930EA98FF82FF901546FC00FD2BFC7B" bold="true" box="[639,739,987,1009]" pageId="11" pageNumber="358">sp. nov.</emphasis>
</taxonomicNameLabel>
Yellow diamonds represent records of type series, and white star locality of the holotype.
</paragraph>
</caption>
<subSubSection id="C35E6501FF82FF9017FEFBE7FB25FAFF" pageId="11" pageNumber="358" type="etymology">
<paragraph id="8BFB368AFF82FF9017FEFBE7FAD2FBDF" blockId="11.[151,1437,1084,2009]" box="[199,1306,1084,1109]" pageId="11" pageNumber="358">
<emphasis id="B930EA98FF82FF9017FEFBE7FE86FBDF" bold="true" box="[199,334,1084,1109]" pageId="11" pageNumber="358">Etymology.</emphasis>
The specific name 
<taxonomicName id="4C444D09FF82FF901511FBE6FD92FBDE" box="[552,602,1085,1108]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="ugoi">
<emphasis id="B930EA98FF82FF901511FBE6FD92FBDE" box="[552,602,1085,1108]" italics="true" pageId="11" pageNumber="358">ugoi</emphasis>
</taxonomicName>
is dedicated to Ugo de Luna Gomes, son of Ulisses L. Gomes.
</paragraph>
<paragraph id="8BFB368AFF82FF9017FEFBBBFB25FAFF" blockId="11.[151,1437,1084,2009]" pageId="11" pageNumber="358">
<emphasis id="B930EA98FF82FF9017FEFBBBFCCEFBF3" bold="true" box="[199,774,1120,1145]" pageId="11" pageNumber="358">
Comparisons with other species of 
<taxonomicName id="4C444D09FF82FF90154CFBBBFCCEFBF3" box="[629,774,1120,1145]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="genus">
<emphasis id="B930EA98FF82FF90154CFBBBFCCEFBF3" bold="true" box="[629,774,1120,1145]" italics="true" pageId="11" pageNumber="358">Scyliorhinus</emphasis>
</taxonomicName>
</emphasis>
. Seven species of 
<taxonomicName id="4C444D09FF82FF9014DEFBBAFBBBFBF3" box="[999,1139,1121,1145]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="genus">
<emphasis id="B930EA98FF82FF9014DEFBBAFBBBFBF3" box="[999,1139,1121,1145]" italics="true" pageId="11" pageNumber="358">Scyliorhinus</emphasis>
</taxonomicName>
are readily distinguished from 
<taxonomicName id="4C444D09FF82FF9017E1FB5EFE57FB16" box="[216,415,1157,1181]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="ugoi">
<emphasis id="B930EA98FF82FF9017E1FB5EFE57FB16" box="[216,415,1157,1181]" italics="true" pageId="11" pageNumber="358">Scyliorhinus ugoi</emphasis>
</taxonomicName>
by color pattern. 
<taxonomicName id="4C444D09FF82FF90154BFB5EFBC0FB17" authority="Garman, 1881" authorityName="Garman" authorityYear="1881" box="[626,1032,1157,1181]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="retifer">
<emphasis id="B930EA98FF82FF90154BFB5EFC85FB17" box="[626,845,1157,1181]" italics="true" pageId="11" pageNumber="358">Scyliorhinus retifer</emphasis>
(
<collectingCountry id="F353761AFF82FF901464FB5EFC73FB17" box="[861,955,1157,1181]" name="Germany" pageId="11" pageNumber="358">Garman</collectingCountry>
, 1881)
</taxonomicName>
has a unique reticulated pattern of dark lines; 
<taxonomicName id="4C444D09FF82FF901626FB72FDA2FB4B" authority="Goode &amp; Bean, 1896" authorityName="Goode &amp; Bean" authorityYear="1896" box="[287,618,1192,1217]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="boa">
<emphasis id="B930EA98FF82FF901626FB72FEA1FB4B" box="[287,361,1193,1217]" italics="true" pageId="11" pageNumber="358">S. boa</emphasis>
Goode &amp; Bean, 1896
</taxonomicName>
, 
<taxonomicName id="4C444D09FF82FF901544FB72FC70FB4B" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[637,952,1192,1217]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="canicula">
<emphasis id="B930EA98FF82FF901544FB72FD35FB4B" box="[637,765,1193,1217]" italics="true" pageId="11" pageNumber="358">S. canicula</emphasis>
Linnaeus, 1758
</taxonomicName>
, 
<taxonomicName id="4C444D09FF82FF9014F3FB72FABDFB4B" authority="Maurin &amp; Bonnet, 1970" authorityName="Maurin &amp; Bonnet" authorityYear="1970" box="[970,1397,1192,1217]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="cervigoni">
<emphasis id="B930EA98FF82FF9014F3FB72FB9DFB4A" box="[970,1109,1193,1216]" italics="true" pageId="11" pageNumber="358">S. cervigoni</emphasis>
Maurin &amp; Bonnet, 1970
</taxonomicName>
, 
<taxonomicName id="4C444D09FF82FF9012BEFB72FE6EFB6F" authority="Fowler, 1934" authorityName="Fowler" authorityYear="1934" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="garmani">
<emphasis id="B930EA98FF82FF9012BEFB72FF3EFB6E" italics="true" pageId="11" pageNumber="358">S. garmani</emphasis>
(Fowler, 1934)
</taxonomicName>
, and 
<taxonomicName id="4C444D09FF82FF9016D1FB16FCDBFB6F" authority="Linnaeus, 1758" authorityName="Linnaeus" authorityYear="1758" box="[488,787,1228,1253]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="stellaris">
<emphasis id="B930EA98FF82FF9016D1FB16FD97FB6F" box="[488,607,1229,1253]" italics="true" pageId="11" pageNumber="358">S. stellaris</emphasis>
Linnaeus, 1758
</taxonomicName>
have conspicuous, small to medium-sized, rounded dark spots and indistinct saddle blotches (
<bibRefCitation id="EFD54B7BFF82FF901509FB2BFD10FA83" author="Springer" box="[560,728,1264,1289]" pageId="11" pageNumber="358" refString="Springer, S. (1979) A revision of the catsharks, Family Scyliorhinidae. NOAA Technical Report NMFS, 422, 1 - 97. Springer, S. &amp; Sadowsky, V. (1970) Subspecies of the western Atlantic catshark, Scyliorhinus retifer. Proceedings of the Biological Society of Washington, 83 (7), 83 - 98." type="journal article" year="1979">Springer, 1979</bibRefCitation>
; 
<bibRefCitation id="EFD54B7BFF82FF9015DAFB2AFC6DFA83" author="Compagno" box="[739,933,1265,1289]" pageId="11" pageNumber="358" refString="Compagno, L. J. V. (1984) FAO species catalogue. Vol. 4. Sharks of the world: An annotated and illustrated catalogue of shark species known to date. Part 2. Carcharhiniformes. FAO Fisheries Synopsis, no. 125, 4, 251 - 655." type="journal article" year="1984">Compagno, 1984</bibRefCitation>
; 
<bibRefCitation id="EFD54B7BFF82FF901489FB2AFB78FA83" author="Compagno" box="[944,1200,1264,1289]" pageId="11" pageNumber="358" refString="Compagno, L. J. V., Dando, M. &amp; Fowler, S. (2005) Sharks of the world. Princeton University Press, Princeton, 368 pp." type="book" year="2005">
Compagno 
<emphasis id="B930EA98FF82FF901309FB2AFBA8FA83" box="[1072,1120,1265,1289]" italics="true" pageId="11" pageNumber="358">et al</emphasis>
., 2005
</bibRefCitation>
). In the 
<emphasis id="B930EA98FF82FF90122BFB2AFEAAFAA7" italics="true" pageId="11" pageNumber="358">
<taxonomicName id="4C444D09FF82FF90122BFB2AFF30FAA7" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="haeckelii">Scyliorhinus haeckelii</taxonomicName>
/ 
<taxonomicName id="4C444D09FF82FF90163DFACEFEAAFAA7" box="[260,354,1301,1325]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="besnardi">besnardi</taxonomicName>
</emphasis>
group, when contrasting saddles are present, specimens lack pre-dorsal saddles with anterior and posterior sharp median projections (
<bibRefCitation id="EFD54B7BFF82FF90155CFAE2FD36FADB" author="Soares" box="[613,766,1337,1361]" pageId="11" pageNumber="358" refString="Soares, K. D. A. (2014) Estudo taxonomico das especies de Scyliorhinus do grupo haeckelii / besnardi (Carcharhiniformes, Scyliorhinidae) das regioes Sudeste e Sul do Brasil (Oceano Atlantico Sul Ocidental). Unpubl. Master's thesis, Universidade do Estado do Rio de Janeiro, Rio de Janeiro, 208 pp." type="book" year="2014">Soares, 2014</bibRefCitation>
). 
<taxonomicName id="4C444D09FF82FF90142AFAE2FB54FADB" authority="Springer, 1966" authorityName="Springer" authorityYear="1966" box="[787,1180,1336,1361]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="meadi">
<emphasis id="B930EA98FF82FF90142AFAE2FC24FADB" box="[787,1004,1337,1361]" italics="true" pageId="11" pageNumber="358">Scyliorhinus meadi</emphasis>
<bibRefCitation id="EFD54B7BFF82FF9014CCFAE3FB54FADB" author="Springer" box="[1013,1180,1336,1361]" pageId="11" pageNumber="358" refString="Springer, S. (1966) A review of Western Atlantic catsharks, Scyliorhinidae, with descriptions of a new genus and five new species. Fishery Bulletin, United States Fish and Wildlife Service, 65 (3), 581 - 624." type="journal article" year="1966">Springer, 1966</bibRefCitation>
</taxonomicName>
has a color pattern of dark saddles on a light background, without light or dark spots (
<bibRefCitation id="EFD54B7BFF82FF901450FA87FBDAFAFF" author="Springer" box="[873,1042,1372,1397]" pageId="11" pageNumber="358" refString="Springer, S. (1979) A revision of the catsharks, Family Scyliorhinidae. NOAA Technical Report NMFS, 422, 1 - 97. Springer, S. &amp; Sadowsky, V. (1970) Subspecies of the western Atlantic catshark, Scyliorhinus retifer. Proceedings of the Biological Society of Washington, 83 (7), 83 - 98." type="journal article" year="1979">Springer, 1979</bibRefCitation>
; 
<bibRefCitation id="EFD54B7BFF82FF901324FA86FB29FAFF" author="Compagno" box="[1053,1249,1373,1397]" pageId="11" pageNumber="358" refString="Compagno, L. J. V. (1984) FAO species catalogue. Vol. 4. Sharks of the world: An annotated and illustrated catalogue of shark species known to date. Part 2. Carcharhiniformes. FAO Fisheries Synopsis, no. 125, 4, 251 - 655." type="journal article" year="1984">Compagno, 1984</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35E6501FF82FF9717FEFA5AFCEEFDB7" lastPageId="12" lastPageNumber="359" pageId="11" pageNumber="358" type="reference_group">
<paragraph id="8BFB368AFF82FF9017FEFA5AFAB4F9A3" blockId="11.[151,1437,1084,2009]" pageId="11" pageNumber="358">
<taxonomicName id="4C444D09FF82FF9017FEFA5AFE46FA12" ID-CoL="4W57X" box="[199,398,1409,1433]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="ugoi">
<emphasis id="B930EA98FF82FF9017FEFA5AFE46FA12" box="[199,398,1409,1433]" italics="true" pageId="11" pageNumber="358">Scyliorhinus ugoi</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C444D09FF82FF901699FA5AFCDCFA13" ID-CoL="4W576" authority="Compagno, 1988" authorityName="Compagno" authorityYear="1988" box="[416,788,1409,1433]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="comoroensis">
<emphasis id="B930EA98FF82FF901699FA5AFD8EFA12" box="[416,582,1409,1432]" italics="true" pageId="11" pageNumber="358">S comoroensis</emphasis>
Compagno, 1988
</taxonomicName>
, 
<taxonomicName id="4C444D09FF82FF90141FFA5AFB0DFA13" ID-CoL="4W574" authority="Muller &amp; Henle, 1838" authorityName="Muller &amp; Henle" authorityYear="1838" box="[806,1221,1408,1433]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="capensis">
<emphasis id="B930EA98FF82FF90141FFA5AFC6FFA12" box="[806,935,1409,1432]" italics="true" pageId="11" pageNumber="358">S. capensis</emphasis>
(Müller &amp; Henle, 1838)
</taxonomicName>
, 
<taxonomicName id="4C444D09FF82FF9013EEFA5AFAFEFA12" ID-CoL="4W57V" box="[1239,1334,1409,1432]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="torrei">
<emphasis id="B930EA98FF82FF9013EEFA5AFAFEFA12" box="[1239,1334,1409,1432]" italics="true" pageId="11" pageNumber="358">S. torrei</emphasis>
</taxonomicName>
Howell- Rivero, 1936, 
<taxonomicName id="4C444D09FF82FF90160FFA7EFDAEFA37" ID-CoL="4W57C" authority="Springer, 1966" authorityName="Springer" authorityYear="1966" box="[310,614,1444,1469]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="hesperius">
<emphasis id="B930EA98FF82FF90160FFA7EFE75FA37" box="[310,445,1445,1469]" italics="true" pageId="11" pageNumber="358">S. hesperius</emphasis>
<bibRefCitation id="EFD54B7BFF82FF9016FAFA7FFDAEFA37" author="Springer" box="[451,614,1444,1469]" pageId="11" pageNumber="358" refString="Springer, S. (1966) A review of Western Atlantic catsharks, Scyliorhinidae, with descriptions of a new genus and five new species. Fishery Bulletin, United States Fish and Wildlife Service, 65 (3), 581 - 624." type="journal article" year="1966">Springer, 1966</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C444D09FF82FF90154DFA7EFBA6FA37" ID-CoL="4W57T" authority="Shirai, Hagiwara &amp; Nakaya, 1992" authorityName="Shirai, Hagiwara &amp; Nakaya" authorityYear="1992" box="[628,1134,1444,1469]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="tokubee">
<emphasis id="B930EA98FF82FF90154DFA7EFD22FA37" box="[628,746,1445,1469]" italics="true" pageId="11" pageNumber="358">S. tokubee</emphasis>
<bibRefCitation id="EFD54B7BFF82FF9015C9FA7FFBA6FA37" author="Shirai" box="[752,1134,1444,1469]" pageId="11" pageNumber="358" refString="Shirai, S., Hagiwara, S. &amp; Nakaya, K. (1992) Scyliorhinus tokubee sp. nov. from Izu Peninsula, Southern Japan (Scyliorhinidae, Elasmobranchii). Japanese Journal of Ichthyology, 39 (1), 9 - 16." type="journal article" year="1992">Shirai, Hagiwara &amp; Nakaya, 1992</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C444D09FF82FF901397FA7EFF10FA6B" ID-CoL="79ZWW" authority="Tanaka, 1908" authorityName="Tanaka" authorityYear="1908" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="torazame">
<emphasis id="B930EA98FF82FF901397FA7EFAFCFA36" box="[1198,1332,1445,1468]" italics="true" pageId="11" pageNumber="358">S. torazame</emphasis>
(Tanaka, 1908)
</taxonomicName>
are similar in having a color pattern of light spots on a darker background, combined with dark saddles, but only 
<taxonomicName id="4C444D09FF82FF9017EDFA36FE53F98E" ID-CoL="4W57X" box="[212,411,1517,1541]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="ugoi">
<emphasis id="B930EA98FF82FF9017EDFA36FE53F98E" box="[212,411,1517,1541]" italics="true" pageId="11" pageNumber="358">Scyliorhinus ugoi</emphasis>
</taxonomicName>
presents small dark spots and pre-dorsal saddles with anterior and posterior sharp median projections (
<bibRefCitation id="EFD54B7BFF82FF901643F9CBFDEBF9A3" author="Springer" box="[378,547,1552,1577]" pageId="11" pageNumber="358" refString="Springer, S. (1979) A revision of the catsharks, Family Scyliorhinidae. NOAA Technical Report NMFS, 422, 1 - 97. Springer, S. &amp; Sadowsky, V. (1970) Subspecies of the western Atlantic catshark, Scyliorhinus retifer. Proceedings of the Biological Society of Washington, 83 (7), 83 - 98." type="journal article" year="1979">Springer, 1979</bibRefCitation>
; Compagno, 1988; 
<bibRefCitation id="EFD54B7BFF82FF9015C5F9CBFC0DF9A3" author="Shirai" box="[764,965,1552,1577]" pageId="11" pageNumber="358" refString="Shirai, S., Hagiwara, S. &amp; Nakaya, K. (1992) Scyliorhinus tokubee sp. nov. from Izu Peninsula, Southern Japan (Scyliorhinidae, Elasmobranchii). Japanese Journal of Ichthyology, 39 (1), 9 - 16." type="journal article" year="1992">
Shirai 
<emphasis id="B930EA98FF82FF90147CF9CAFCBDF9A3" box="[837,885,1553,1577]" italics="true" pageId="11" pageNumber="358">et al</emphasis>
., 1992
</bibRefCitation>
; 
<bibRefCitation id="EFD54B7BFF82FF9014E9F9CAFB19F9A3" author="Compagno" box="[976,1233,1552,1577]" pageId="11" pageNumber="358" refString="Compagno, L. J. V., Dando, M. &amp; Fowler, S. (2005) Sharks of the world. Princeton University Press, Princeton, 368 pp." type="book" year="2005">
Compagno 
<emphasis id="B930EA98FF82FF901368F9CAFB49F9A3" box="[1105,1153,1553,1577]" italics="true" pageId="11" pageNumber="358">et al</emphasis>
., 2005
</bibRefCitation>
; 
<bibRefCitation id="EFD54B7BFF82FF9013E5F9CAFAB8F9A3" author="Soares" box="[1244,1392,1553,1577]" pageId="11" pageNumber="358" refString="Soares, K. D. A. (2014) Estudo taxonomico das especies de Scyliorhinus do grupo haeckelii / besnardi (Carcharhiniformes, Scyliorhinidae) das regioes Sudeste e Sul do Brasil (Oceano Atlantico Sul Ocidental). Unpubl. Master's thesis, Universidade do Estado do Rio de Janeiro, Rio de Janeiro, 208 pp." type="book" year="2014">Soares, 2014</bibRefCitation>
).
</paragraph>
<paragraph id="8BFB368AFF82FF9017FEF9EEFCD8F91F" blockId="11.[151,1437,1084,2009]" pageId="11" pageNumber="358">
<taxonomicName id="4C444D09FF82FF9017FEF9EEFE75F9C7" ID-CoL="9G677" box="[199,445,1589,1613]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="canicula">
<emphasis id="B930EA98FF82FF9017FEF9EEFE75F9C7" box="[199,445,1589,1613]" italics="true" pageId="11" pageNumber="358">Scyliorhinus canicula</emphasis>
</taxonomicName>
has relatively larger, broader anterior nasal flaps that reach the mouth, covering the shallow nasoral grooves that are lacking in 
<taxonomicName id="4C444D09FF82FF9015ECF982FCEEF9FA" ID-CoL="4W57X" box="[725,806,1625,1648]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="ugoi">
<emphasis id="B930EA98FF82FF9015ECF982FCEEF9FA" box="[725,806,1625,1648]" italics="true" pageId="11" pageNumber="358">S. ugoi</emphasis>
</taxonomicName>
and in all other species of the genus (
<bibRefCitation id="EFD54B7BFF82FF9013D2F983FA50F9FB" author="Springer" box="[1259,1432,1624,1649]" pageId="11" pageNumber="358" refString="Springer, S. (1979) A revision of the catsharks, Family Scyliorhinidae. NOAA Technical Report NMFS, 422, 1 - 97. Springer, S. &amp; Sadowsky, V. (1970) Subspecies of the western Atlantic catshark, Scyliorhinus retifer. Proceedings of the Biological Society of Washington, 83 (7), 83 - 98." type="journal article" year="1979">Springer, 1979</bibRefCitation>
; 
<bibRefCitation id="EFD54B7BFF82FF9017AEF9A6FE91F91F" author="Compagno" box="[151,345,1661,1685]" pageId="11" pageNumber="358" refString="Compagno, L. J. V. (1984) FAO species catalogue. Vol. 4. Sharks of the world: An annotated and illustrated catalogue of shark species known to date. Part 2. Carcharhiniformes. FAO Fisheries Synopsis, no. 125, 4, 251 - 655." type="journal article" year="1984">Compagno, 1984</bibRefCitation>
; 
<bibRefCitation id="EFD54B7BFF82FF90165CF9A6FDAEF91F" author="Compagno" box="[357,614,1660,1685]" pageId="11" pageNumber="358" refString="Compagno, L. J. V., Dando, M. &amp; Fowler, S. (2005) Sharks of the world. Princeton University Press, Princeton, 368 pp." type="book" year="2005">
Compagno 
<emphasis id="B930EA98FF82FF9016DCF9A6FDDDF91F" box="[485,533,1661,1685]" italics="true" pageId="11" pageNumber="358">et al</emphasis>
., 2005
</bibRefCitation>
; 
<bibRefCitation id="EFD54B7BFF82FF901548F9A6FCCCF91F" author="Soares" box="[625,772,1661,1685]" pageId="11" pageNumber="358" refString="Soares, K. D. A. (2014) Estudo taxonomico das especies de Scyliorhinus do grupo haeckelii / besnardi (Carcharhiniformes, Scyliorhinidae) das regioes Sudeste e Sul do Brasil (Oceano Atlantico Sul Ocidental). Unpubl. Master's thesis, Universidade do Estado do Rio de Janeiro, Rio de Janeiro, 208 pp." type="book" year="2014">Soares, 2014</bibRefCitation>
).
</paragraph>
<paragraph id="8BFB368AFF82FF9017FEF97AFA89F8C3" blockId="11.[151,1437,1084,2009]" pageId="11" pageNumber="358">
<emphasis id="B930EA98FF82FF9017FEF97AFDECF933" box="[199,548,1697,1721]" italics="true" pageId="11" pageNumber="358">
<taxonomicName id="4C444D09FF82FF9017FEF97AFE72F933" ID-CoL="6XZWW" box="[199,442,1697,1721]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="haeckelii">Scyliorhinus haeckelii</taxonomicName>
/ 
<taxonomicName id="4C444D09FF82FF9016FFF97AFDECF933" ID-CoL="79ZWZ" box="[454,548,1697,1721]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="besnardi">besnardi</taxonomicName>
</emphasis>
from the Southwestern Atlantic Ocean differs from 
<taxonomicName id="4C444D09FF82FF90134CF97AFB09F932" ID-CoL="4W57X" box="[1141,1217,1697,1720]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="ugoi">
<emphasis id="B930EA98FF82FF90134CF97AFB09F932" box="[1141,1217,1697,1720]" italics="true" pageId="11" pageNumber="358">S. ugoi</emphasis>
</taxonomicName>
in its smaller size, with adult males known at about 
<quantity id="4CBC9B6FFF82FF901524F91FFD7FF956" box="[541,695,1732,1757]" metricMagnitude="-1" metricUnit="m" metricValue="3.6" metricValueMax="3.7" metricValueMin="3.5" pageId="11" pageNumber="358" unit="mm" value="360.0" valueMax="370.0" valueMin="350.0">350–370 mm</quantity>
, while in 
<taxonomicName id="4C444D09FF82FF901414F91EFCB5F956" ID-CoL="4W57X" box="[813,893,1733,1756]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="ugoi">
<emphasis id="B930EA98FF82FF901414F91EFCB5F956" box="[813,893,1733,1756]" italics="true" pageId="11" pageNumber="358">S. ugoi</emphasis>
</taxonomicName>
males at about 
<quantity id="4CBC9B6FFF82FF901300F91EFB51F956" box="[1081,1177,1733,1757]" metricMagnitude="-1" metricUnit="m" metricValue="4.2" pageId="11" pageNumber="358" unit="mm" value="420.0">420 mm</quantity>
are still immature. Its snout is more slender, usually more pointed and longer than that of 
<taxonomicName id="4C444D09FF82FF9014A7F932FC24F88A" ID-CoL="4W57X" box="[926,1004,1769,1792]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="ugoi">
<emphasis id="B930EA98FF82FF9014A7F932FC24F88A" box="[926,1004,1769,1792]" italics="true" pageId="11" pageNumber="358">S. ugoi</emphasis>
</taxonomicName>
, with preoral length 1.4–1.7 times in preorbital length 
<emphasis id="B930EA98FF82FF90165FF8D6FE4DF8AE" box="[358,389,1805,1828]" italics="true" pageId="11" pageNumber="358">vs.</emphasis>
1.1– 1. 4 times in 
<taxonomicName id="4C444D09FF82FF901557F8D6FCBDF8AF" ID-CoL="4W57X" authority="Soares, 2014" authorityName="Soares" authorityYear="2014" box="[622,885,1805,1829]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="ugoi">
<emphasis id="B930EA98FF82FF901557F8D6FD77F8AE" box="[622,703,1805,1828]" italics="true" pageId="11" pageNumber="358">S. ugoi</emphasis>
(
<bibRefCitation id="EFD54B7BFF82FF9015EBF8D6FCA6F8AF" author="Soares" box="[722,878,1805,1829]" pageId="11" pageNumber="358" refString="Soares, K. D. A. (2014) Estudo taxonomico das especies de Scyliorhinus do grupo haeckelii / besnardi (Carcharhiniformes, Scyliorhinidae) das regioes Sudeste e Sul do Brasil (Oceano Atlantico Sul Ocidental). Unpubl. Master's thesis, Universidade do Estado do Rio de Janeiro, Rio de Janeiro, 208 pp." type="book" year="2014">Soares, 2014</bibRefCitation>
)
</taxonomicName>
. The wider and stout head of 
<taxonomicName id="4C444D09FF82FF9013E6F8D6FAF9F8AE" ID-CoL="4W57X" box="[1247,1329,1805,1828]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="ugoi">
<emphasis id="B930EA98FF82FF9013E6F8D6FAF9F8AE" box="[1247,1329,1805,1828]" italics="true" pageId="11" pageNumber="358">S. ugoi</emphasis>
</taxonomicName>
is easily observable in all specimens and, together with its characteristic color pattern, permits a precise distinction.
</paragraph>
<paragraph id="8BFB368AFF82FF9017FEF88FFD23F83F" blockId="11.[151,1437,1084,2009]" pageId="11" pageNumber="358">
The abdomen of 
<taxonomicName id="4C444D09FF82FF9016B2F88EFE11F8E6" ID-CoL="4W57X" box="[395,473,1877,1900]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="ugoi">
<emphasis id="B930EA98FF82FF9016B2F88EFE11F8E6" box="[395,473,1877,1900]" italics="true" pageId="11" pageNumber="358">S. ugoi</emphasis>
</taxonomicName>
is relatively longer than in 
<emphasis id="B930EA98FF82FF901427F88EFBC3F8E7" box="[798,1035,1877,1901]" italics="true" pageId="11" pageNumber="358">
<taxonomicName id="4C444D09FF82FF901427F88EFC68F8E7" ID-CoL="6XZWW" box="[798,928,1877,1901]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="haeckelii">S. haeckelii</taxonomicName>
/ 
<taxonomicName id="4C444D09FF82FF901494F88EFBC3F8E7" ID-CoL="79ZWZ" box="[941,1035,1877,1901]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="besnardi">besnardi</taxonomicName>
</emphasis>
, with pectoral–pelvic space 1.5–1.6 times pelvic–anal space 
<emphasis id="B930EA98FF82FF9016FAF8A2FE12F81A" box="[451,474,1913,1936]" italics="true" pageId="11" pageNumber="358">vs</emphasis>
0.8–1.4. Interdorsal space 2.1–2.5 times dorsal–caudal space 
<emphasis id="B930EA98FF82FF9013ECF8A2FB24F81A" box="[1237,1260,1913,1936]" italics="true" pageId="11" pageNumber="358">vs</emphasis>
. 
<quantity id="4CBC9B6FFF82FF90123DF8A2FAA2F81B" box="[1284,1386,1912,1937]" metricMagnitude="-2" metricUnit="m" metricValue="4.064" metricValueMax="5.08" metricValueMin="3.048" pageId="11" pageNumber="358" unit="in" value="1.6" valueMax="2.0" valueMin="1.2">1.2–2 in</quantity>
the 
<emphasis id="B930EA98FF82FF9017AEF846FE3BF83F" box="[151,499,1949,1973]" italics="true" pageId="11" pageNumber="358">
<taxonomicName id="4C444D09FF82FF9017AEF846FE41F83F" ID-CoL="6XZWW" box="[151,393,1949,1973]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="haeckelii">Scyliorhinus haeckelii</taxonomicName>
/ 
<taxonomicName id="4C444D09FF82FF9016ACF846FE3BF83F" ID-CoL="79ZWZ" box="[405,499,1949,1973]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="besnardi">besnardi</taxonomicName>
</emphasis>
group (
<bibRefCitation id="EFD54B7BFF82FF901573F846FD17F83F" author="Soares" box="[586,735,1949,1973]" pageId="11" pageNumber="358" refString="Soares, K. D. A. (2014) Estudo taxonomico das especies de Scyliorhinus do grupo haeckelii / besnardi (Carcharhiniformes, Scyliorhinidae) das regioes Sudeste e Sul do Brasil (Oceano Atlantico Sul Ocidental). Unpubl. Master's thesis, Universidade do Estado do Rio de Janeiro, Rio de Janeiro, 208 pp." type="book" year="2014">Soares, 2014</bibRefCitation>
).
</paragraph>
<paragraph id="8BFB368AFF82FF9717FEF81BFEFFFF72" blockId="11.[151,1437,1084,2009]" lastBlockId="12.[151,1436,151,573]" lastPageId="12" lastPageNumber="359" pageId="11" pageNumber="358">
Monospondylous vertebral counts of 
<taxonomicName id="4C444D09FF82FF9015B3F81AFD11F852" ID-CoL="4W57X" box="[650,729,1985,2008]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="ugoi">
<emphasis id="B930EA98FF82FF9015B3F81AFD11F852" box="[650,729,1985,2008]" italics="true" pageId="11" pageNumber="358">S. ugoi</emphasis>
</taxonomicName>
(38–39) differ from counts for 
<taxonomicName id="4C444D09FF82FF90135AF81AFB01F853" ID-CoL="6XZX8" box="[1123,1225,1985,2009]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="meadi">
<emphasis id="B930EA98FF82FF90135AF81AFB01F853" box="[1123,1225,1985,2009]" italics="true" pageId="11" pageNumber="358">S. meadi</emphasis>
</taxonomicName>
(43–49), 
<taxonomicName id="4C444D09FF82FF901204F81AFA54F852" ID-CoL="4W57V" box="[1341,1436,1985,2008]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="11" pageNumber="358" phylum="Chordata" rank="species" species="torrei">
<emphasis id="B930EA98FF82FF901204F81AFA54F852" box="[1341,1436,1985,2008]" italics="true" pageId="11" pageNumber="358">S. torrei</emphasis>
</taxonomicName>
(30–34), 
<taxonomicName id="4C444D09FF85FF9717C6FF43FE61FF25" ID-CoL="4W576" box="[255,425,152,175]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="12" pageNumber="359" phylum="Chordata" rank="species" species="comoroensis">
<emphasis id="B930EA98FF85FF9717C6FF43FE61FF25" box="[255,425,152,175]" italics="true" pageId="12" pageNumber="359">S. comoroensis</emphasis>
</taxonomicName>
(45) and 
<taxonomicName id="4C444D09FF85FF971525FF43FD53FF25" ID-CoL="4W574" box="[540,667,152,175]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="12" pageNumber="359" phylum="Chordata" rank="species" species="capensis">
<emphasis id="B930EA98FF85FF971525FF43FD53FF25" box="[540,667,152,175]" italics="true" pageId="12" pageNumber="359">S. capensis</emphasis>
</taxonomicName>
(48), but do not differ appreciably from counts for 
<taxonomicName id="4C444D09FF85FF971232FF43FABAFF3A" ID-CoL="4W57K" box="[1291,1394,152,176]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="12" pageNumber="359" phylum="Chordata" rank="species" species="retifer">
<emphasis id="B930EA98FF85FF971232FF43FABAFF3A" box="[1291,1394,152,176]" italics="true" pageId="12" pageNumber="359">S. retifer</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C444D09FF85FF9712BEFF43FF09FF5E" ID-CoL="4W56X" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="12" pageNumber="359" phylum="Chordata" rank="species" species="boa">
<emphasis id="B930EA98FF85FF9712BEFF43FF09FF5E" italics="true" pageId="12" pageNumber="359">S. boa</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C444D09FF85FF9717E0FF67FEA2FF5E" ID-CoL="4W57C" box="[217,362,188,212]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="12" pageNumber="359" phylum="Chordata" rank="species" species="hesperius">
<emphasis id="B930EA98FF85FF9717E0FF67FEA2FF5E" box="[217,362,188,212]" italics="true" pageId="12" pageNumber="359">S. hesperius</emphasis>
</taxonomicName>
and 
<emphasis id="B930EA98FF85FF971696FF67FD69FF5E" box="[431,673,188,212]" italics="true" pageId="12" pageNumber="359">
<taxonomicName id="4C444D09FF85FF971696FF67FDFEFF5E" ID-CoL="6XZWW" box="[431,566,188,212]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="12" pageNumber="359" phylum="Chordata" rank="species" species="haeckelii">S. haeckelii</taxonomicName>
/ 
<taxonomicName id="4C444D09FF85FF97157AFF67FD69FF5E" ID-CoL="79ZWZ" box="[579,673,188,212]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="12" pageNumber="359" phylum="Chordata" rank="species" species="besnardi">besnardi</taxonomicName>
</emphasis>
(Springer &amp; Sadowsky, 1970; 
<bibRefCitation id="EFD54B7BFF85FF971333FF67FB7DFF5E" author="Springer" box="[1034,1205,188,213]" pageId="12" pageNumber="359" refString="Springer, S. (1979) A revision of the catsharks, Family Scyliorhinidae. NOAA Technical Report NMFS, 422, 1 - 97. Springer, S. &amp; Sadowsky, V. (1970) Subspecies of the western Atlantic catshark, Scyliorhinus retifer. Proceedings of the Biological Society of Washington, 83 (7), 83 - 98." type="journal article" year="1979">Springer, 1979</bibRefCitation>
; 
<bibRefCitation id="EFD54B7BFF85FF9713FDFF67FA50FF5F" author="Compagno" box="[1220,1432,188,213]" pageId="12" pageNumber="359" refString="Compagno, L. J. V. (1988 b) Scyliorhinus comoroensis sp. n., a new catshark from the Comoro Islands, western Indian Ocean (Carcharhiniformes, Scyliorhinidae). Bulletin Museum National d'Histoire Naturelle, 10 (3), 603 - 625." type="journal article" year="1988" yearSuffix="b">Compagno, 1988b</bibRefCitation>
; 
<bibRefCitation id="EFD54B7BFF85FF9717AEFF3BFEE3FF72" author="Soares" box="[151,299,224,248]" pageId="12" pageNumber="359" refString="Soares, K. D. A. (2014) Estudo taxonomico das especies de Scyliorhinus do grupo haeckelii / besnardi (Carcharhiniformes, Scyliorhinidae) das regioes Sudeste e Sul do Brasil (Oceano Atlantico Sul Ocidental). Unpubl. Master's thesis, Universidade do Estado do Rio de Janeiro, Rio de Janeiro, 208 pp." type="book" year="2014">Soares, 2014</bibRefCitation>
).
</paragraph>
<paragraph id="8BFB368AFF85FF9717FEFEDFFCF4FE5A" blockId="12.[151,1436,151,573]" pageId="12" pageNumber="359">
The occurrence of 
<taxonomicName id="4C444D09FF85FF97169CFEDFFDE7FE96" ID-CoL="4W57C" box="[421,559,260,284]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="12" pageNumber="359" phylum="Chordata" rank="species" species="hesperius">
<emphasis id="B930EA98FF85FF97169CFEDFFDE7FE96" box="[421,559,260,284]" italics="true" pageId="12" pageNumber="359">S. hesperius</emphasis>
</taxonomicName>
in the uppermost slope (
<quantity id="4CBC9B6FFF85FF971462FEDFFC17FE96" box="[859,991,260,285]" metricMagnitude="2" metricUnit="m" metricValue="3.655" metricValueMax="4.57" metricValueMin="2.7399999999999998" pageId="12" pageNumber="359" unit="m" value="365.5" valueMax="457.0" valueMin="274.0">274–457 m</quantity>
in depth) of the Atlantic from Central 
<collectingCountry id="F353761AFF85FF9717AEFEFCFF3FFECA" box="[151,247,295,320]" name="United States of America" pageId="12" pageNumber="359">America</collectingCountry>
to 
<collectingCountry id="F353761AFF85FF971627FEFCFEAAFECA" box="[286,354,295,320]" name="Brazil" pageId="12" pageNumber="359">Brazil</collectingCountry>
(
<bibRefCitation id="EFD54B7BFF85FF97164BFEFCFD42FECA" author="Gadig" box="[370,650,295,320]" pageId="12" pageNumber="359" refString="Gadig, O. B. F. &amp; Gomes, U. L. (2003) Classe Chondrichthyes. In: Menezes, N. A., Buckup, P. A., Figueiredo, J. L. &amp; Moura, R. L. (Eds.), Catalogo das especies de peixes marinhos do Brasil. Museu de Zoologia da Universidade de Sao Paulo, Sao Paulo, pp. 21 - 22." type="book chapter" year="2003">Gadig &amp; Gomes, 2003</bibRefCitation>
) is an error of identification and refers, in fact, to 
<taxonomicName id="4C444D09FF85FF9713D1FEF3FF14FEEE" ID-CoL="4W57X" authority="Soares, 2014" authorityName="Soares" authorityYear="2014" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="12" pageNumber="359" phylum="Chordata" rank="species" species="ugoi">
<emphasis id="B930EA98FF85FF9713D1FEF3FAF0FEB5" box="[1256,1336,296,319]" italics="true" pageId="12" pageNumber="359">S. ugoi</emphasis>
(
<bibRefCitation id="EFD54B7BFF85FF971270FEF3FF1CFEEE" author="Soares" pageId="12" pageNumber="359" refString="Soares, K. D. A. (2014) Estudo taxonomico das especies de Scyliorhinus do grupo haeckelii / besnardi (Carcharhiniformes, Scyliorhinidae) das regioes Sudeste e Sul do Brasil (Oceano Atlantico Sul Ocidental). Unpubl. Master's thesis, Universidade do Estado do Rio de Janeiro, Rio de Janeiro, 208 pp." type="book" year="2014">Soares, 2014</bibRefCitation>
)
</taxonomicName>
. Beyond the differences in color pattern, 
<taxonomicName id="4C444D09FF85FF9715F9FE97FCC7FEEE" ID-CoL="4W57X" box="[704,783,332,356]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="12" pageNumber="359" phylum="Chordata" rank="species" species="ugoi">
<emphasis id="B930EA98FF85FF9715F9FE97FCC7FEEE" box="[704,783,332,356]" italics="true" pageId="12" pageNumber="359">S. ugoi</emphasis>
</taxonomicName>
has a larger head, with preorbital length 6.4% TL and head length 19.5–20.3% TL 
<emphasis id="B930EA98FF85FF9716C9FEAAFDCFFE02" box="[496,519,369,392]" italics="true" pageId="12" pageNumber="359">vs</emphasis>
. 4.9–5.3% TL and 18.9% TL in 
<emphasis id="B930EA98FF85FF971498FEABFBFCFE02" box="[929,1076,368,392]" italics="true" pageId="12" pageNumber="359">
<taxonomicName id="4C444D09FF85FF971498FEABFBF8FE02" ID-CoL="4W57C" box="[929,1072,368,392]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="12" pageNumber="359" phylum="Chordata" rank="species" species="hesperius">S. hesperius</taxonomicName>
,
</emphasis>
respectively. The mouth is also larger in 
<taxonomicName id="4C444D09FF85FF9717C4FE4FFE83FE26" ID-CoL="4W57X" box="[253,331,404,428]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="12" pageNumber="359" phylum="Chordata" rank="species" species="ugoi">
<emphasis id="B930EA98FF85FF9717C4FE4FFE83FE26" box="[253,331,404,428]" italics="true" pageId="12" pageNumber="359">S. ugoi</emphasis>
</taxonomicName>
, its width 9.0–9.7% TL 
<emphasis id="B930EA98FF85FF97155EFE4EFD4DFE26" box="[615,645,405,428]" italics="true" pageId="12" pageNumber="359">vs.</emphasis>
7.0– 
<quantity id="4CBC9B6FFF85FF9715F9FE4FFCCAFE27" box="[704,770,404,429]" metricMagnitude="-1" metricUnit="m" metricValue="1.905" pageId="12" pageNumber="359" unit="in" value="7.5">7.5 in</quantity>
<taxonomicName id="4C444D09FF85FF971432FE4FFC5CFE26" ID-CoL="4W57C" box="[779,916,404,428]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="12" pageNumber="359" phylum="Chordata" rank="species" species="hesperius">
<emphasis id="B930EA98FF85FF971432FE4FFC5CFE26" box="[779,916,404,428]" italics="true" pageId="12" pageNumber="359">S. hesperius</emphasis>
</taxonomicName>
, as is its anal base length at 7.7–8.2% TL 
<emphasis id="B930EA98FF85FF971244FE4EFA5CFE26" box="[1405,1428,405,428]" italics="true" pageId="12" pageNumber="359">vs</emphasis>
. 5.7–7.7% TL in 
<taxonomicName id="4C444D09FF85FF971676FE63FE1FFE5A" ID-CoL="4W57C" box="[335,471,440,464]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="12" pageNumber="359" phylum="Chordata" rank="species" species="hesperius">
<emphasis id="B930EA98FF85FF971676FE63FE1FFE5A" box="[335,471,440,464]" italics="true" pageId="12" pageNumber="359">S. hesperius</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD54B7BFF85FF9716DCFE6CFD47FE5A" author="Springer" box="[485,655,439,464]" pageId="12" pageNumber="359" refString="Springer, S. (1979) A revision of the catsharks, Family Scyliorhinidae. NOAA Technical Report NMFS, 422, 1 - 97. Springer, S. &amp; Sadowsky, V. (1970) Subspecies of the western Atlantic catshark, Scyliorhinus retifer. Proceedings of the Biological Society of Washington, 83 (7), 83 - 98." type="journal article" year="1979">Springer, 1979</bibRefCitation>
; 
<bibRefCitation id="EFD54B7BFF85FF9715A2FE63FCF8FE5A" author="Soares" box="[667,816,440,464]" pageId="12" pageNumber="359" refString="Soares, K. D. A. (2014) Estudo taxonomico das especies de Scyliorhinus do grupo haeckelii / besnardi (Carcharhiniformes, Scyliorhinidae) das regioes Sudeste e Sul do Brasil (Oceano Atlantico Sul Ocidental). Unpubl. Master's thesis, Universidade do Estado do Rio de Janeiro, Rio de Janeiro, 208 pp." type="book" year="2014">Soares, 2014</bibRefCitation>
).
</paragraph>
<paragraph id="8BFB368AFF85FF9717FEFE06FCEEFDB7" blockId="12.[151,1436,151,573]" pageId="12" pageNumber="359">
It is apparent from the comparisons presented above that 
<taxonomicName id="4C444D09FF85FF971476FE07FC55FE7E" ID-CoL="4W57X" box="[847,925,476,500]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="12" pageNumber="359" phylum="Chordata" rank="species" species="ugoi">
<emphasis id="B930EA98FF85FF971476FE07FC55FE7E" box="[847,925,476,500]" italics="true" pageId="12" pageNumber="359">S. ugoi</emphasis>
</taxonomicName>
is not particularly close to its geographically nearest congener, 
<emphasis id="B930EA98FF85FF971658FDDBFD83FD92" box="[353,587,512,536]" italics="true" pageId="12" pageNumber="359">
<taxonomicName id="4C444D09FF85FF971658FDDBFE28FD92" ID-CoL="6XZWW" box="[353,480,512,536]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="12" pageNumber="359" phylum="Chordata" rank="species" species="haeckelii">S. haeckelii</taxonomicName>
/ 
<taxonomicName id="4C444D09FF85FF9716D5FDDBFD83FD92" ID-CoL="79ZWZ" box="[492,587,512,536]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="12" pageNumber="359" phylum="Chordata" rank="species" species="besnardi">besnardi</taxonomicName>
</emphasis>
of the southwestern Atlantic Ocean, and is differentiated from 
<taxonomicName id="4C444D09FF85FF97122DFDDBFA54FD92" ID-CoL="4W57C" box="[1300,1436,512,536]" class="Elasmobranchii" family="Scyliorhinidae" genus="Scyliorhinus" kingdom="Animalia" order="Carcharhiniformes" pageId="12" pageNumber="359" phylum="Chordata" rank="species" species="hesperius">
<emphasis id="B930EA98FF85FF97122DFDDBFA54FD92" box="[1300,1436,512,536]" italics="true" pageId="12" pageNumber="359">S. hesperius</emphasis>
</taxonomicName>
of Central 
<collectingCountry id="F353761AFF85FF971637FDFFFEA7FDB7" box="[270,367,548,573]" name="United States of America" pageId="12" pageNumber="359">America</collectingCountry>
by color pattern and body proportions.
</paragraph>
</subSubSection>
</treatment>
</document>