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<document id="EFBB2A371CFE5C29AD1B337061A3F1A9" ID-CLB-Dataset="3262" ID-DOI="10.5281/zenodo.5731880" ID-GBIF-Dataset="48249a3d-036b-4e20-896b-92042f714cff" ID-ISSN="2064-2474" ID-Zenodo-Dep="5731880" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638041255008" checkinUser="felipe" docAuthor="Papp, L" docDate="2008" docId="03E1F51CFFC21442FDB13FBFFD913B37" docLanguage="en" docName="ActaZoolAcadSciH.54.1.13-21.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 54 (1)" docStyle="DocumentStyle:4FD749C6DF7B87D8940F4D8C7B0A0514.3:ActaZoolAcadSciH.2002-2014.journal_article" docStyleId="4FD749C6DF7B87D8940F4D8C7B0A0514" docStyleName="ActaZoolAcadSciH.2002-2014.journal_article" docStyleVersion="3" docTitle="Micromacrocera L. PAPP 2008, gen. n." docType="treatment" docVersion="4" lastPageNumber="14" masterDocId="FFD88D64FFC31440FFFE3E08FF90386A" masterDocTitle="Micromacrocera Gen. N., The Smallest Macrocerine Fly (Diptera: Keroplatidae, Macrocerinae)" masterLastPageNumber="21" masterPageNumber="13" pageNumber="14" updateTime="1699470583137" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="9598A8CE96919498010651E888768839">Micromacrocera Gen. N., The Smallest Macrocerine Fly (Diptera: Keroplatidae, Macrocerinae)</mods:title>
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<mods:namePart id="3799117A38162CCA363FF7DAE7A44B53">Papp, L</mods:namePart>
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<mods:date id="30FEFC65F4AE295C8716EBB7662F6A56">2008</mods:date>
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<treatment id="03E1F51CFFC21442FDB13FBFFD913B37" ID-DOI="http://doi.org/10.5281/zenodo.5731838" ID-GBIF-Taxon="197857349" ID-Zenodo-Dep="5731838" LSID="urn:lsid:plazi:treatment:03E1F51CFFC21442FDB13FBFFD913B37" httpUri="http://treatment.plazi.org/id/03E1F51CFFC21442FDB13FBFFD913B37" lastPageId="2" lastPageNumber="14" pageId="1" pageNumber="14">
<subSubSection id="C3521781FFC21441FDB13FBFFC7439BB" box="[591,996,439,465]" pageId="1" pageNumber="14" type="nomenclature">
<paragraph id="8BF7440AFFC21441FDB13FBFFC7439BB" blockId="1.[591,996,439,465]" box="[591,996,439,465]" pageId="1" pageNumber="14">
<heading id="D0BFF366FFC21441FDB13FBFFC7439BB" box="[591,996,439,465]" centered="true" fontSize="10" level="2" pageId="1" pageNumber="14" reason="6">
<taxonomicName id="4C483F89FFC21441FDB13FBFFC1639BB" ID-CoL="5RYT" authority="L. PAPP" authorityName="L. PAPP" authorityYear="2008" box="[591,902,439,465]" class="Insecta" family="Keroplatidae" genus="Micromacrocera" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="14" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B93C9818FFC21441FDB13FBFFCB139BB" bold="true" box="[591,801,439,465]" pageId="1" pageNumber="14">Micromacrocera</emphasis>
L. PAPP
</taxonomicName>
,
<taxonomicNameLabel id="A20F2563FFC21441FC6B3FBFFC7439BB" box="[917,996,439,465]" pageId="1" pageNumber="14" rank="genus">gen. n.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3521781FFC21441FE803FF3FC3F3A7A" box="[382,943,507,528]" pageId="1" pageNumber="14" type="type_taxon">
<paragraph id="8BF7440AFFC21441FE803FF3FC3F3A7A" blockId="1.[382,1083,507,585]" box="[382,943,507,528]" pageId="1" pageNumber="14">
<typeStatus id="54F3FAA8FFC21441FE803FF3FE3F3A7A" box="[382,431,507,528]" pageId="1" pageNumber="14">Type</typeStatus>
species:
<taxonomicName id="4C483F89FFC21441FDF43FF3FCFC3A64" ID-CoL="42YKB" authority="L. PAPP" authorityName="L. PAPP" authorityYear="2008" box="[522,876,507,528]" class="Insecta" family="Keroplatidae" genus="Micromacrocera" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="14" phylum="Arthropoda" rank="species" species="stenobasis" status="sp. nov.">
<emphasis id="B93C9818FFC21441FDF43FF3FC883A7A" box="[522,792,507,528]" italics="true" pageId="1" pageNumber="14">Micromacrocera stenobasis</emphasis>
L. PAPP
</taxonomicName>
,
<taxonomicNameLabel id="A20F2563FFC21441FC873FF3FC3F3A7A" box="[889,943,507,528]" pageId="1" pageNumber="14" rank="species">sp. n.</taxonomicNameLabel>
</paragraph>
</subSubSection>
<subSubSection id="C3521781FFC21441FE803C10FDBF3A47" box="[382,559,536,557]" pageId="1" pageNumber="14" type="description">
<paragraph id="8BF7440AFFC21441FE803C10FDBF3A47" blockId="1.[382,1083,507,585]" box="[382,559,536,557]" pageId="1" pageNumber="14">Gender: feminine.</paragraph>
</subSubSection>
<subSubSection id="C3521781FFC21441FE803C3CFBAB3A23" box="[382,1083,564,585]" pageId="1" pageNumber="14" type="etymology">
<paragraph id="8BF7440AFFC21441FE803C3CFBAB3A23" blockId="1.[382,1083,507,585]" box="[382,1083,564,585]" pageId="1" pageNumber="14">
Etymology. This is the smallest species of the subfamily
<taxonomicName id="4C483F89FFC21441FC4E3C3CFBA73A23" authorityName="Papp" authorityYear="2008" box="[944,1079,564,585]" class="Insecta" family="Keroplatidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="14" phylum="Arthropoda" rank="subFamily" subFamily="Macrocerinae">Macrocerinae</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C3521781FFC21442FE803C66FD913B37" lastPageId="2" lastPageNumber="15" pageId="1" pageNumber="14" type="description">
<paragraph id="8BF7440AFFC21441FE803C66FD713B79" blockId="1.[317,1270,622,1824]" pageId="1" pageNumber="14">
Frons (postfrons) short oblique, cerebral sclerite large, occupying all dorsal part of head. Cerebral sclerite with some scattered setae. Occiput and postgena large. Eyes comparatively small, occupying less than half of the head in profile (
<figureCitation id="1373588FFFC21441FD303CA0FC953AD7" box="[718,773,680,701]" captionStart="Figs 14" captionStartId="2.[317,360,1743,1764]" captionTargetBox="[319,1265,973,1702]" captionTargetId="figure-230@2.[317,1269,972,1705]" captionTargetPageId="2" captionText="Figs 14. Micromacrocera stenobasis sp. n., holotype male. 1 = head laterally with scape and pedicel, 2 = mouth-parts in a ventral, sublateral view (stipes and cardo dark); 3 = first flagellomere, 4 = 10th flagellomere. Scale: 0.2 mm for all" figureDoi="http://doi.org/10.5281/zenodo.5731882" httpUri="https://zenodo.org/record/5731882/files/figure.png" pageId="1" pageNumber="14">Fig. 1</figureCitation>
). Three ocelli, anterior one forward, posterior ocelli dorsally directed. Pilosity of eyes distinct (
<figureCitation id="1373588FFFC21441FD333CCCFC933AB3" box="[717,771,708,729]" captionStart="Figs 14" captionStartId="2.[317,360,1743,1764]" captionTargetBox="[319,1265,973,1702]" captionTargetId="figure-230@2.[317,1269,972,1705]" captionTargetPageId="2" captionText="Figs 14. Micromacrocera stenobasis sp. n., holotype male. 1 = head laterally with scape and pedicel, 2 = mouth-parts in a ventral, sublateral view (stipes and cardo dark); 3 = first flagellomere, 4 = 10th flagellomere. Scale: 0.2 mm for all" figureDoi="http://doi.org/10.5281/zenodo.5731882" httpUri="https://zenodo.org/record/5731882/files/figure.png" pageId="1" pageNumber="14">Fig. 1</figureCitation>
). Face and clypeus separated by a rather broad membranous aperture. Face very short, in this respect it is similar to that of
<taxonomicName id="4C483F89FFC21441FC243CE9FBE23A9C" box="[986,1138,737,758]" class="Insecta" family="Keroplatidae" genus="Macrocera" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="14" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B93C9818FFC21441FC243CE9FBD53A9C" box="[986,1093,737,758]" italics="true" pageId="1" pageNumber="14">Macrocera</emphasis>
spp.
</taxonomicName>
Clypeus high and comparatively narrow. Foramen small.
</paragraph>
<paragraph id="8BF7440AFFC21441FE803D13FE1D3BC9" blockId="1.[317,1270,622,1824]" pageId="1" pageNumber="14">
Antenna in all probability with 14 flagellomeres, but only 10 can be seen on the specimen (left side, on the right side only 8), apical flagellomeres lost. Scape comparatively long, ventrally with longer setae, pedicel globose. Flagellomeres (
<figureCitation id="1373588FFFC21441FD213D5CFCA73B03" box="[735,823,852,873]" captionStart="Figs 14" captionStartId="2.[317,360,1743,1764]" captionTargetBox="[319,1265,973,1702]" captionTargetId="figure-230@2.[317,1269,972,1705]" captionTargetPageId="2" captionText="Figs 14. Micromacrocera stenobasis sp. n., holotype male. 1 = head laterally with scape and pedicel, 2 = mouth-parts in a ventral, sublateral view (stipes and cardo dark); 3 = first flagellomere, 4 = 10th flagellomere. Scale: 0.2 mm for all" figureDoi="http://doi.org/10.5281/zenodo.5731882" httpUri="https://zenodo.org/record/5731882/files/figure.png" pageId="1" pageNumber="14">Figs 34</figureCitation>
) simple, cylindrical. Hair-like setae (cilia) of flagellomeres rather long, on the 10th flagellomere they are longer than diameter of flagellomere (
<figureCitation id="1373588FFFC21441FEBA3D86FE113BC9" box="[324,385,910,931]" captionStart="Figs 14" captionStartId="2.[317,360,1743,1764]" captionTargetBox="[319,1265,973,1702]" captionTargetId="figure-230@2.[317,1269,972,1705]" captionTargetPageId="2" captionText="Figs 14. Micromacrocera stenobasis sp. n., holotype male. 1 = head laterally with scape and pedicel, 2 = mouth-parts in a ventral, sublateral view (stipes and cardo dark); 3 = first flagellomere, 4 = 10th flagellomere. Scale: 0.2 mm for all" figureDoi="http://doi.org/10.5281/zenodo.5731882" httpUri="https://zenodo.org/record/5731882/files/figure.png" pageId="1" pageNumber="14">Fig. 4</figureCitation>
).
</paragraph>
<paragraph id="8BF7440AFFC21441FE803DA3FC133BB7" blockId="1.[317,1270,622,1824]" pageId="1" pageNumber="14">
Mouth-parts rather small, labella small, fleshy. Maxillary palpus with four palpomeres but seems 3-partite, since palpomeres 12 partly fused (
<figureCitation id="1373588FFFC21441FCC53DC0FCE73BB7" box="[827,887,968,989]" captionStart="Figs 14" captionStartId="2.[317,360,1743,1764]" captionTargetBox="[319,1265,973,1702]" captionTargetId="figure-230@2.[317,1269,972,1705]" captionTargetPageId="2" captionText="Figs 14. Micromacrocera stenobasis sp. n., holotype male. 1 = head laterally with scape and pedicel, 2 = mouth-parts in a ventral, sublateral view (stipes and cardo dark); 3 = first flagellomere, 4 = 10th flagellomere. Scale: 0.2 mm for all" figureDoi="http://doi.org/10.5281/zenodo.5731882" httpUri="https://zenodo.org/record/5731882/files/figure.png" pageId="1" pageNumber="14">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BF7440AFFC21441FE803DECFCDF3C3A" blockId="1.[317,1270,622,1824]" pageId="1" pageNumber="14">
Thorax (
<figureCitation id="1373588FFFC21441FE2F3DECFD9B3B93" box="[465,523,996,1017]" captionStart="Figs 510" captionStartId="3.[317,360,1721,1742]" captionTargetBox="[320,1268,1052,1691]" captionTargetId="figure-230@3.[317,1269,1049,1691]" captionTargetPageId="3" captionText="Figs 510. Micromacrocera stenobasis sp. n., holotype male. 5 = thorax with coxae, lateral view, 6 = apex of right fore tibia, anterior view, 7 = contour of fore basitarsus, anterior view, 8 = apex of left hind tibia, anterior view, 9 = fore tarsomere 5, lateral view, 10 =apex of fore tarsus, dorsal view. Scales: 0.4 mm for Fig. 5, 0.2 mm for Fig. 7, 0.1 mm for Figs 6, 810" figureDoi="http://doi.org/10.5281/zenodo.5731884" httpUri="https://zenodo.org/record/5731884/files/figure.png" pageId="1" pageNumber="14">Fig. 5</figureCitation>
) small, height at middle (without coxae) only
<quantity id="4CB0E9EFFFC21441FC2E3DECFBB63B93" box="[976,1062,996,1017]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="14" unit="mm" value="0.5">0.50 mm</quantity>
(
<figureCitation id="1373588FFFC21441FBCA3DECFBE03B93" box="[1076,1136,996,1017]" captionStart="Figs 510" captionStartId="3.[317,360,1721,1742]" captionTargetBox="[320,1268,1052,1691]" captionTargetId="figure-230@3.[317,1269,1049,1691]" captionTargetPageId="3" captionText="Figs 510. Micromacrocera stenobasis sp. n., holotype male. 5 = thorax with coxae, lateral view, 6 = apex of right fore tibia, anterior view, 7 = contour of fore basitarsus, anterior view, 8 = apex of left hind tibia, anterior view, 9 = fore tarsomere 5, lateral view, 10 =apex of fore tarsus, dorsal view. Scales: 0.4 mm for Fig. 5, 0.2 mm for Fig. 7, 0.1 mm for Figs 6, 810" figureDoi="http://doi.org/10.5281/zenodo.5731884" httpUri="https://zenodo.org/record/5731884/files/figure.png" pageId="1" pageNumber="14">Fig. 5</figureCitation>
). Structure of thorax unusual: mediotergite largely dorsal, laterotergite not large but continued dorsally. Scutellum bulging, minute,
<quantity id="4CB0E9EFFFC21441FE1C3A16FDA83C59" box="[482,568,1054,1075]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="1" pageNumber="14" unit="mm" value="0.07">0.07 mm</quantity>
only. (I saw only 4 setal bases on scutellum, almost dorsal ones, i.e. not marginals; scutellum otherwise microtomentose only).
</paragraph>
<paragraph id="8BF7440AFFC21441FE803A50FB753D73" blockId="1.[317,1270,622,1824]" pageId="1" pageNumber="14">
Wing (
<figureCitation id="1373588FFFC21441FE3F3A50FD963C07" box="[449,518,1112,1133]" captionStart="Fig" captionStartId="6.[378,413,1804,1825]" captionTargetBox="[320,1269,957,1764]" captionTargetId="figure-216@6.[317,1269,957,1766]" captionTargetPageId="6" captionText="Fig. 15. Micromacrocera stenobasis sp. n., holotype male, photo of the wings dorsally" figureDoi="http://doi.org/10.5281/zenodo.5731894" httpUri="https://zenodo.org/record/5731894/files/figure.png" pageId="1" pageNumber="14">Fig. 15</figureCitation>
) membrane without macrotrichia and with microtrichia evenly distributed but wholly irregular. No strong costal fringe. Costa thick, reaches 3/4 of distance between R
<subScript id="17CC464FFFC21441FB623A89FB333CE7" attach="left" box="[1180,1187,1153,1165]" fontSize="5" pageId="1" pageNumber="14">5</subScript>
and M
<subScript id="17CC464FFFC21441FB173A89FB603CE7" attach="right" box="[1257,1264,1153,1165]" fontSize="5" pageId="1" pageNumber="14">1</subScript>
. Longitudinal veins (incl. Cu1) setose dorsally, medial stalk (M
<subScript id="17CC464FFFC21441FC6E3A96FC353CC0" attach="left" box="[912,933,1182,1194]" fontSize="5" pageId="1" pageNumber="14">1+2</subScript>
) free of setae. R
<subScript id="17CC464FFFC21441FBBE3A96FBD73CC0" attach="left" box="[1088,1095,1182,1194]" fontSize="5" pageId="1" pageNumber="14">4</subScript>
absent but a small thickening on R
<subScript id="17CC464FFFC21441FE2B3AB2FE4C3CAC" attach="left" box="[469,476,1210,1222]" fontSize="5" pageId="1" pageNumber="14">5</subScript>
represents base of R
<subScript id="17CC464FFFC21441FD5F3AB2FD383CAC" attach="left" box="[673,680,1210,1222]" fontSize="5" pageId="1" pageNumber="14">4</subScript>
. M
<subScript id="17CC464FFFC21441FD363AB2FD5F3CAC" attach="left" box="[712,719,1210,1222]" fontSize="5" pageId="1" pageNumber="14">1</subScript>
and M
<subScript id="17CC464FFFC21441FCEC3AB2FC893CAC" attach="none" box="[786,793,1210,1222]" fontSize="5" pageId="1" pageNumber="14">2</subScript>
not completely reaching wing margin. Base of M
<subScript id="17CC464FFFC21441FB113AB2FB663CAC" attach="right" box="[1263,1270,1210,1222]" fontSize="5" pageId="1" pageNumber="14">3</subScript>
not pigmented. Cu
<subScript id="17CC464FFFC21441FE0F3ADFFE683C89" attach="left" box="[497,504,1239,1251]" fontSize="5" pageId="1" pageNumber="14">2</subScript>
and A
<subScript id="17CC464FFFC21441FDC43ADFFDD13C89" attach="left" box="[570,577,1239,1251]" fontSize="5" pageId="1" pageNumber="14">1</subScript>
present as short indistinct parallel brownish folds on wing base (to the level of R-M fusion only). A
<subScript id="17CC464FFFC21441FDAB3AFCFDCC3D6A" attach="left" box="[597,604,1268,1280]" fontSize="5" pageId="1" pageNumber="14">2</subScript>
not discernible at all. No alula present between anal angle and wing scale, and even almost no alular region on wing. Prealular (basal) region with a straight margin (
<figureCitation id="1373588FFFC21441FB693B0CFB493D73" box="[1175,1241,1284,1305]" captionStart="Fig" captionStartId="6.[378,413,1804,1825]" captionTargetBox="[320,1269,957,1764]" captionTargetId="figure-216@6.[317,1269,957,1766]" captionTargetPageId="6" captionText="Fig. 15. Micromacrocera stenobasis sp. n., holotype male, photo of the wings dorsally" figureDoi="http://doi.org/10.5281/zenodo.5731894" httpUri="https://zenodo.org/record/5731894/files/figure.png" pageId="1" pageNumber="14">Fig. 15</figureCitation>
).
</paragraph>
<paragraph id="8BF7440AFFC21441FE803B29FD5A3D5C" blockId="1.[317,1270,622,1824]" box="[382,714,1313,1334]" pageId="1" pageNumber="14">Halter simple, knob not too large.</paragraph>
<paragraph id="8BF7440AFFC21441FE803B36FDE63D89" blockId="1.[317,1270,622,1824]" pageId="1" pageNumber="14">
Femoral and tibial setulae simple, fine and rather long, and not ordered into rows. Fore coxa medium-long, the middle coxa short, the hind coxa very short (
<figureCitation id="1373588FFFC21441FC473B53FC673D1A" box="[953,1015,1371,1392]" captionStart="Figs 510" captionStartId="3.[317,360,1721,1742]" captionTargetBox="[320,1268,1052,1691]" captionTargetId="figure-230@3.[317,1269,1049,1691]" captionTargetPageId="3" captionText="Figs 510. Micromacrocera stenobasis sp. n., holotype male. 5 = thorax with coxae, lateral view, 6 = apex of right fore tibia, anterior view, 7 = contour of fore basitarsus, anterior view, 8 = apex of left hind tibia, anterior view, 9 = fore tarsomere 5, lateral view, 10 =apex of fore tarsus, dorsal view. Scales: 0.4 mm for Fig. 5, 0.2 mm for Fig. 7, 0.1 mm for Figs 6, 810" figureDoi="http://doi.org/10.5281/zenodo.5731884" httpUri="https://zenodo.org/record/5731884/files/figure.png" pageId="1" pageNumber="14">Fig. 5</figureCitation>
). Tibial spurs 1+2+1, i.e. posteroventral spur on hind tibia missing (at least I was unable to find its base even under a higher magnification). Fore tibia antero-apically without a large depressed or even emarginated area (
<figureCitation id="1373588FFFC21441FB313B9CFEDB3DAC" captionStart="Figs 510" captionStartId="3.[317,360,1721,1742]" captionTargetBox="[320,1268,1052,1691]" captionTargetId="figure-230@3.[317,1269,1049,1691]" captionTargetPageId="3" captionText="Figs 510. Micromacrocera stenobasis sp. n., holotype male. 5 = thorax with coxae, lateral view, 6 = apex of right fore tibia, anterior view, 7 = contour of fore basitarsus, anterior view, 8 = apex of left hind tibia, anterior view, 9 = fore tarsomere 5, lateral view, 10 =apex of fore tarsus, dorsal view. Scales: 0.4 mm for Fig. 5, 0.2 mm for Fig. 7, 0.1 mm for Figs 6, 810" figureDoi="http://doi.org/10.5281/zenodo.5731884" httpUri="https://zenodo.org/record/5731884/files/figure.png" pageId="1" pageNumber="14">Fig. 6</figureCitation>
), and with a row of 5 medium-long pointed setae, similar to some
<taxonomicName id="4C483F89FFC21441FC3A3BB9FB8D3DAC" authorityName="Billberg" authorityYear="1820" box="[964,1053,1457,1478]" class="Insecta" family="Sciaridae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="14" phylum="Arthropoda" rank="family">Sciaridae</taxonomicName>
(e.g.
<taxonomicName id="4C483F89FFC21441FBAD3BB9FB3A3DAC" authorityName="Winnertz" authorityYear="1867" box="[1107,1194,1457,1478]" class="Insecta" family="Sciaridae" genus="Bradysia" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="14" phylum="Arthropoda" rank="genus">
<emphasis id="B93C9818FFC21441FBAD3BB9FB3A3DAC" box="[1107,1194,1457,1478]" italics="true" pageId="1" pageNumber="14">Bradysia</emphasis>
</taxonomicName>
), rather than to any other
<taxonomicName id="4C483F89FFC21441FE153BC6FDE23D89" authorityName="Papp" authorityYear="2008" box="[491,626,1486,1507]" class="Insecta" family="Keroplatidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="14" phylum="Arthropoda" rank="subFamily" subFamily="Macrocerinae">Macrocerinae</taxonomicName>
.
</paragraph>
<paragraph id="8BF7440AFFC21441FE803BE3FDB33E53" blockId="1.[317,1270,622,1824]" pageId="1" pageNumber="14">
No anterior or posterior combs of small thornlets on mid and hind tibiae (
<figureCitation id="1373588FFFC21441FBCD3BE3FBFC3E6A" box="[1075,1132,1515,1536]" captionStart="Figs 510" captionStartId="3.[317,360,1721,1742]" captionTargetBox="[320,1268,1052,1691]" captionTargetId="figure-230@3.[317,1269,1049,1691]" captionTargetPageId="3" captionText="Figs 510. Micromacrocera stenobasis sp. n., holotype male. 5 = thorax with coxae, lateral view, 6 = apex of right fore tibia, anterior view, 7 = contour of fore basitarsus, anterior view, 8 = apex of left hind tibia, anterior view, 9 = fore tarsomere 5, lateral view, 10 =apex of fore tarsus, dorsal view. Scales: 0.4 mm for Fig. 5, 0.2 mm for Fig. 7, 0.1 mm for Figs 6, 810" figureDoi="http://doi.org/10.5281/zenodo.5731884" httpUri="https://zenodo.org/record/5731884/files/figure.png" pageId="1" pageNumber="14">Fig. 8</figureCitation>
). Tarsal claws simple, very small, and straight, except for apical part. No developed pulvilli; they are as hairy as empodium (
<figureCitation id="1373588FFFC21441FE4C382CFD883E53" box="[434,536,1572,1593]" captionStart="Figs 510" captionStartId="3.[317,360,1721,1742]" captionTargetBox="[320,1268,1052,1691]" captionTargetId="figure-230@3.[317,1269,1049,1691]" captionTargetPageId="3" captionText="Figs 510. Micromacrocera stenobasis sp. n., holotype male. 5 = thorax with coxae, lateral view, 6 = apex of right fore tibia, anterior view, 7 = contour of fore basitarsus, anterior view, 8 = apex of left hind tibia, anterior view, 9 = fore tarsomere 5, lateral view, 10 =apex of fore tarsus, dorsal view. Scales: 0.4 mm for Fig. 5, 0.2 mm for Fig. 7, 0.1 mm for Figs 6, 810" figureDoi="http://doi.org/10.5281/zenodo.5731884" httpUri="https://zenodo.org/record/5731884/files/figure.png" pageId="1" pageNumber="14">Figs 910</figureCitation>
).
</paragraph>
<paragraph id="8BF7440AFFC21441FE803849FCBB3E8C" blockId="1.[317,1270,622,1824]" pageId="1" pageNumber="14">
Abdomen with 8 normal preabdominal (pregenital) segments. Both tergites and sternites broad, in each segment tergite and sternite almost meet laterally, i.e. intervening membranous area rather small. Spiracle pairs 17 are situated in membrane. Tergite 8 (
<figureCitation id="1373588FFFC21441FC253873FBB13EFA" box="[987,1057,1659,1680]" captionStart="Figs 1112" captionStartId="4.[317,360,1748,1769]" captionTargetBox="[321,1266,1083,1719]" captionTargetId="figure-402@4.[320,1267,1082,1719]" captionTargetPageId="4" captionText="Figs 1112. Micromacrocera stenobasis sp. n., holotype male, terminalia.11 = tergite and sternite 8, ventral view (tergal setae omitted), 12 = tergite 9 and proctiger, dorsal view (hypoproct covered). Scale: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.5731888" httpUri="https://zenodo.org/record/5731888/files/figure.png" pageId="1" pageNumber="14">Fig. 11</figureCitation>
) not much shortened, sternite 8 short, not much more than half as long as tergite (0.57x). Tergite 9 comparatively long, anterior edge broadly emarginated, caudally largely pentagonal (
<figureCitation id="1373588FFFC21441FC6338BCFC773EA3" box="[925,999,1716,1737]" captionStart="Figs 1112" captionStartId="4.[317,360,1748,1769]" captionTargetBox="[321,1266,1083,1719]" captionTargetId="figure-402@4.[320,1267,1082,1719]" captionTargetPageId="4" captionText="Figs 1112. Micromacrocera stenobasis sp. n., holotype male, terminalia.11 = tergite and sternite 8, ventral view (tergal setae omitted), 12 = tergite 9 and proctiger, dorsal view (hypoproct covered). Scale: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.5731888" httpUri="https://zenodo.org/record/5731888/files/figure.png" pageId="1" pageNumber="14">Fig. 12</figureCitation>
). Cerci comparatively large and broad with 2 pairs of rather short apical setae.
</paragraph>
<paragraph id="8BF7440AFFC21442FE8038E6FD913B37" blockId="1.[317,1270,622,1824]" lastBlockId="2.[317,1271,436,861]" lastPageId="2" lastPageNumber="15" pageId="1" pageNumber="14">
Male genitalia rather small. Left and right gonocoxites fused but on a short ventral section only (
<figureCitation id="1373588FFFC21441FE863903FE7B3F4A" box="[376,491,1803,1824]" captionStart="Figs 1314" captionStartId="5.[317,360,1773,1794]" captionTargetBox="[461,1131,424,1741]" captionTargetId="figure-10@5.[455,1131,424,1745]" captionTargetPageId="5" captionText="Figs 1314. Micromacrocera stenobasis sp. n., holotype male, genitalia. 13 = dorsal view, 14 = ventral view. Scale: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.5731890" httpUri="https://zenodo.org/record/5731890/files/figure.png" pageId="1" pageNumber="14">Figs 1314</figureCitation>
). Genital structures in three layers (from ventral to dorsal): gonocoxal fusion, parameral complex, and most dorsal, just under tergite 9, gonocoxal apodemes with their fine plate (see below). Medial wall of gonocoxites less long (high) than breadth of gonostylus. Parameres with long lateral arms joining long gonocoxal apodemes. Sclerite connecting parameres with basiphallus very short compared to that of
<taxonomicName id="4C483F89FFC11442FD8B3C03FC803A4A" box="[629,784,523,544]" class="Insecta" family="Keroplatidae" genus="Macrocera" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="15" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B93C9818FFC11442FD8B3C03FD703A4A" box="[629,736,523,544]" italics="true" pageId="2" pageNumber="15">Macrocera</emphasis>
spp.
</taxonomicName>
Phallus (distiphallus) membranous, disintegrated when treated with sodium hydroxide. The medial slightly asymmetrical sclerotized structure, ventral to the level of tip of parameres and gonocoxal apodemes, must be regarded as ejaculatory apodeme. There are 2 pairs of parameral lateral arms: one dorsal, connected to gonocoxal apodemes; the ventral pair as long as dorsal one, completely overlapping it, (
<figureCitation id="1373588FFFC11442FCBC3C76FC183AF9" box="[834,904,638,659]" captionStart="Figs 1314" captionStartId="5.[317,360,1773,1794]" captionTargetBox="[461,1131,424,1741]" captionTargetId="figure-10@5.[455,1131,424,1745]" captionTargetPageId="5" captionText="Figs 1314. Micromacrocera stenobasis sp. n., holotype male, genitalia. 13 = dorsal view, 14 = ventral view. Scale: 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.5731890" httpUri="https://zenodo.org/record/5731890/files/figure.png" pageId="2" pageNumber="15">Fig. 13</figureCitation>
), slightly ventrally curved pair of processes may support phallus. Gonocoxal apodemes connected by a lamina, which is distinct though not strongly sclerotized, and, that connects gonocoxal apodeme apices with basiphallus (not seen in
<taxonomicName id="4C483F89FFC11442FEC33CDDFE383A80" box="[317,424,725,746]" class="Insecta" family="Keroplatidae" genus="Macrocera" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B93C9818FFC11442FEC33CDDFE383A80" box="[317,424,725,746]" italics="true" pageId="2" pageNumber="15">Macrocera</emphasis>
</taxonomicName>
). Inner wall of gonocoxites almost complete, i.e. ends not far from base of gonostylus with a basal lateral process; lateral half of the apex of gonocoxites with a U-shaped emargination, which allow moving the apex of gonostylus laterally. That inner (medial) wall plain, i.e. not concave as in
<taxonomicName id="4C483F89FFC11442FE903D23FD823B2A" authorityName="Papp &amp; Sevcik" authorityYear="2005" box="[366,530,811,832]" class="Insecta" family="Keroplatidae" genus="Sciarokeroplatus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B93C9818FFC11442FE903D23FD823B2A" box="[366,530,811,832]" italics="true" pageId="2" pageNumber="15">Sciarokeroplatus</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFD939FBFFC11442FDBA3D23FC9E3B2A" author="PAPP, L. &amp; SEVCIK, J." box="[580,782,811,832]" pageId="2" pageNumber="15" pagination="113 - 123" refId="ref3391" refString="PAPP, L. &amp; SEVCIK, J. (2005) Sciarokeroplatinae, a new subfamily of Keroplatidae (Diptera). Acta Zoologica Academiae Scientiarum Hungaricae 51 (2): 113 - 123." type="journal article" year="2005">
<taxonomicName id="4C483F89FFC11442FDBA3D23FDE33B54" authority="Papp, 2008" authorityName="Papp" authorityYear="2008" box="[580,627,811,832]" pageId="2" pageNumber="16" rank="species" species="papp" status="sp. nov.">PAPP</taxonomicName>
&amp; ŠEVČÍK 2005
</bibRefCitation>
) and present not far from gonostylar base, contrary to
<taxonomicName id="4C483F89FFC11442FEA93D40FE6B3B37" authorityName="Papp &amp; Sevcik" authorityYear="2005" box="[343,507,840,861]" class="Insecta" family="Keroplatidae" genus="Sciarokeroplatus" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="15" phylum="Arthropoda" rank="genus">
<emphasis id="B93C9818FFC11442FEA93D40FE6B3B37" box="[343,507,840,861]" italics="true" pageId="2" pageNumber="15">Sciarokeroplatus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>