152 lines
15 KiB
XML
152 lines
15 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.131.1921" ID-GBIF-Dataset="72a81528-63ef-4531-9f9c-ade551a651d8" ID-PMC="PMC3208436" ID-Pensoft-Pub="1313-2970-131-1" ID-PubMed="22140329" ModsDocAuthor="" ModsDocDate="2011" ModsDocID="1313-2970-131-1" ModsDocOrigin="ZooKeys 131" ModsDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" checkinTime="1451249839150" checkinUser="pensoft" docAuthor="Likhitrakarn, Natdanai, Golovatch, Sergei I. & Panha, Somsak" docDate="2011" docId="8C125C108C56957E2370A7DA1C1F197D" docLanguage="en" docName="ZooKeys 131: 1-161" docOrigin="ZooKeys 131" docSource="http://dx.doi.org/10.3897/zookeys.131.1921" docTitle="Orthomorpha Bollman 1893" docType="treatment" docVersion="6" lastPageNumber="6" masterDocId="FFFF0F54FF88BD1C371C965D22453928" masterDocTitle="Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)" masterLastPageNumber="161" masterPageNumber="1" pageNumber="5" updateTime="1668152297771" updateUser="ExternalLinkService">
|
|
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
|
<mods:titleInfo>
|
|
<mods:title>Revision of the Southeast Asian millipede genus Orthomorpha Bollman, 1893, with the proposal of a new genus (Diplopoda, Polydesmida, Paradoxosomatidae)</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Likhitrakarn, Natdanai</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Golovatch, Sergei I.</mods:namePart>
|
|
</mods:name>
|
|
<mods:name type="personal">
|
|
<mods:role>
|
|
<mods:roleTerm>Author</mods:roleTerm>
|
|
</mods:role>
|
|
<mods:namePart>Panha, Somsak</mods:namePart>
|
|
</mods:name>
|
|
<mods:typeOfResource>text</mods:typeOfResource>
|
|
<mods:relatedItem type="host">
|
|
<mods:titleInfo>
|
|
<mods:title>ZooKeys</mods:title>
|
|
</mods:titleInfo>
|
|
<mods:part>
|
|
<mods:date>2011</mods:date>
|
|
<mods:detail type="volume">
|
|
<mods:number>131</mods:number>
|
|
</mods:detail>
|
|
<mods:extent unit="page">
|
|
<mods:start>1</mods:start>
|
|
<mods:end>161</mods:end>
|
|
</mods:extent>
|
|
</mods:part>
|
|
</mods:relatedItem>
|
|
<mods:location>
|
|
<mods:url>http://dx.doi.org/10.3897/zookeys.131.1921</mods:url>
|
|
</mods:location>
|
|
<mods:classification>journal article</mods:classification>
|
|
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.131.1921</mods:identifier>
|
|
<mods:identifier type="Pensoft-Pub">1313-2970-131-1</mods:identifier>
|
|
</mods:mods>
|
|
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3506926" ID-GBIF-Taxon="152031094" ID-Zenodo-Dep="3506926" LSID="urn:lsid:plazi:treatment:8C125C108C56957E2370A7DA1C1F197D" httpUri="http://treatment.plazi.org/id/8C125C108C56957E2370A7DA1C1F197D" lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
|
|
<subSubSection pageId="4" pageNumber="5" type="nomenclature">
|
|
<paragraph pageId="4" pageNumber="5">
|
|
<taxonomicName LSID="http://species-id.net/wiki/Orthomorpha" authority="Bollman, 1893" authorityName="Bollman" authorityYear="1893" class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Orthomorpha Bollman, 1893</taxonomicName>
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="4" pageNumber="5" type="reference_group">
|
|
<paragraph pageId="4" pageNumber="5">
|
|
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Orthomorpha</taxonomicName>
|
|
<bibRefCitation author="Bollman, CH" journalOrPublisher="Bulletin of the United States National Museum" pageId="114" pageNumber="115" pagination="150 - 152" title=": Notes upon the North American myriapods described by C. L. Koch." volume="46" year="1893">Bollman 1893</bibRefCitation>
|
|
: 159 (N).
|
|
</paragraph>
|
|
<paragraph pageId="4" pageNumber="5">
|
|
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="genus">Orthomorpha</taxonomicName>
|
|
-
|
|
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="205 - 283" title="Paradoxosomatidae from Borneo (Diplopoda, Polydesmida)." volume="106" year="1963">Jeekel 1963</bibRefCitation>
|
|
: 261 (D).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="4" pageNumber="5" type="diagnosis">
|
|
<paragraph pageId="4" pageNumber="5">Diagnosis.</paragraph>
|
|
<paragraph pageId="4" pageNumber="5">
|
|
A genus of
|
|
<taxonomicName lsidName="" pageId="4" pageNumber="5" rank="tribe" tribe="Orthomorphini">Orthomorphini</taxonomicName>
|
|
with 20 segments. Body medium- to large-sized, adults ca 15-50 mm long, ca 1.1-3.1 and 1.5-6.7 mm wide on midbody pro- and metazona, respectively. Paraterga invariably well-developed, metazonite to prozonite width ratio being ca 1.6-1.7. Adenostyles on ♂ legs 1 missing. Sternal lobe or cone(s) between ♂ coxae 4 present or absent.
|
|
</paragraph>
|
|
<paragraph pageId="4" pageNumber="5">
|
|
Gonopod with a long, subcylindrical, distodorsally usually setose coxite and a normal, cylindrical cannula. Telopodite mostly very slender and long, modestly curved. Prefemoral portion densely setose, about as long as (rarely) to ca 2-3 times (usually) shorter than femorite (measured together with
|
|
<normalizedToken originalValue="“postfemoral”">"postfemoral"</normalizedToken>
|
|
part lying distal to lateral sulcus). Femorite without evidence of torsion (= seminal groove running only mesally), often slightly enlarged distally, mostly with a clear-cut, oblique, distolateral sulcus demarcating a
|
|
<normalizedToken originalValue="“postfemoral”">"postfemoral"</normalizedToken>
|
|
part. Solenophore only moderately strongly curved mesad or caudomesad, consisting of modestly developed laminae lateralis and medialis, yet with lamina lateralis somewhat larger than lamina medialis, both sheathing a similarly long, simple, flagelliform solenomere with a barely exposed tip; tip of solenophore never deeply split, normally poorly bi- or trifid, some of apical prongs being either minute denticles or lappets, or small teeth, or completely reduced.
|
|
</paragraph>
|
|
<paragraph pageId="4" pageNumber="5">
|
|
Type-species:
|
|
<taxonomicName class="Diplopoda" family="Polydesmidae" genus="Polydesmus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Polydesmus beaumontii" order="Polydesmida" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="beaumontii">Polydesmus beaumontii</taxonomicName>
|
|
Le Guillou, 1841, by subsequent designation by
|
|
<bibRefCitation pageId="4" pageNumber="5">Pocock (1909)</bibRefCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5" type="remarks">
|
|
<paragraph pageId="4" pageNumber="5">Remarks.</paragraph>
|
|
<paragraph lastPageId="5" lastPageNumber="6" pageId="4" pageNumber="5">
|
|
The
|
|
<taxonomicName lsidName="" pageId="4" pageNumber="5" rank="tribe" tribe="Orthomorphini">Orthomorphini</taxonomicName>
|
|
is certainly the most diverse tribe of Oriental
|
|
<taxonomicName lsidName="" pageId="4" pageNumber="5" rank="subFamily" subFamily="Paradoxosomatinae">Paradoxosomatinae</taxonomicName>
|
|
both at the generic and species levels. It is generally characterized by the gonopod showing a simple, usually subcylindrical (= normally not excavate mesa
|
|
<pageBreakToken pageId="5" pageNumber="6" start="start">lly</pageBreakToken>
|
|
) and elongate femorite devoid both of torsion (= the seminal groove running entirely or nearly entirely on the mesal side) and processes/outgrowths. A distolateral sulcus demarcating a
|
|
<normalizedToken originalValue="“postfemoral”">"postfemoral"</normalizedToken>
|
|
part is usually, but not always, present. A solenomere is always long and flagelliform, starting on top of the femorite (+
|
|
<normalizedToken originalValue="“postfemoral”">"postfemoral"</normalizedToken>
|
|
part, if any) at the base of a more or less elaborate solenophore, the latter being demarcated by an evident cingulum or mesal sulcus. The solenophore always consists of well- to moderately well developed, often subequal, lamellar lamina lateralis and lamina medialis, both only modestly curved mesad or caudomesad and both supporting and sheathing at least most of the solenomere. It is the solenophore that provides most of the generic characters in
|
|
<taxonomicName lsidName="" pageId="5" pageNumber="6" rank="tribe" tribe="Orthomorphini">Orthomorphini</taxonomicName>
|
|
, such as the presence or absence of additional structures (processes or lobes) at its base, near midway and/or at its tip (
|
|
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Academisch Proefschrift, Rotterdam" pageId="114" pageNumber="115" title="On the classification and geographical distribution of the family Paradoxosomatidae (Diplopoda, Polydesmida)." year="1968">Jeekel 1968</bibRefCitation>
|
|
;
|
|
<bibRefCitation pageId="5" pageNumber="6">Golovatch 1997a</bibRefCitation>
|
|
,
|
|
<bibRefCitation pageId="5" pageNumber="6">2000</bibRefCitation>
|
|
,
|
|
<bibRefCitation author="Golovatch, SI" journalOrPublisher="Arthropoda Selecta" pageId="114" pageNumber="115" pagination="119 - 124" title="On several new or poorly-known Oriental Paradoxosomatidae (Diplopoda Polydesmida), IX." volume="18" year="2009">2009</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph pageId="5" pageNumber="6">
|
|
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Orthomorpha</taxonomicName>
|
|
is basically characterized by very broad paraterga, coupled with the gonopod showing mostly an elongate, slender femorite (+
|
|
<normalizedToken originalValue="“postfemoral”">"postfemoral"</normalizedToken>
|
|
part, if any) and a long, modestly curved solenophore bearing additional structures neither at its midway nor near its base; the tip of the solenophore is never deeply split, normally poorly bi- or trifid, some of the apical prongs being either minute denticles or lappets, or small teeth, or completely reduced.
|
|
</paragraph>
|
|
<paragraph pageId="5" pageNumber="6">
|
|
Based on the broad paraterga and the conformation of the solenophore,
|
|
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Orthomorpha</taxonomicName>
|
|
comes closest to the continental Southeast Asian
|
|
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Antheromorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Antheromorpha" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Antheromorpha</taxonomicName>
|
|
Jeekel, 1968 (see above), the Philippine
|
|
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Luzonomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Luzonomorpha" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Luzonomorpha</taxonomicName>
|
|
Hoffman, 1973 and the basically Bornean
|
|
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Gigantomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gigantomorpha" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Gigantomorpha</taxonomicName>
|
|
Jeekel, 1963 (
|
|
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="205 - 283" title="Paradoxosomatidae from Borneo (Diplopoda, Polydesmida)." volume="106" year="1963">Jeekel 1963</bibRefCitation>
|
|
,
|
|
<bibRefCitation author="Jeekel, CAW" journalOrPublisher="Tijdschrift voor Entomologie" pageId="114" pageNumber="115" pagination="71 - 88" title="A revision of the Burmese Paradoxosomatidae (Diplopoda, Polydesmida) in the Museo Civico di Storia Naturale at Genoa (Part III)." volume="123" year="1980 a">1980a</bibRefCitation>
|
|
;
|
|
<bibRefCitation author="Hoffman, RL" journalOrPublisher="Journal of Natural History" pageId="114" pageNumber="115" pagination="361 - 389" title="Descriptions and allocation of new or poorly known genera and species of Paradoxosomatidae from south-eastern Asia (Diplopoda: Polydesmida)." url="doi: 10.1080/00222937300770281" volume="7" year="1973">Hoffman 1973</bibRefCitation>
|
|
;
|
|
<bibRefCitation author="Golovatch, SI" journalOrPublisher="Revue suisse de Zoologie" pageId="114" pageNumber="115" pagination="151 - 193" title="The millipede family Paradoxosomatidae on Borneo, with contributions to the faunas of some other islands of the Sunda area (Diplopooda, Polydesmida)." volume="103" year="1996">Golovatch 1996</bibRefCitation>
|
|
,
|
|
<bibRefCitation pageId="5" pageNumber="6">1997a</bibRefCitation>
|
|
). However,
|
|
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Antheromorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Antheromorpha" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Antheromorpha</taxonomicName>
|
|
species show a very deeply split tip of the solenophore,
|
|
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Gigantomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Gigantomorpha" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Gigantomorpha</taxonomicName>
|
|
species demonstrate a somewhat flattened and bisinuate gonofemorite, usually also a more elaborate, often deeply split solenophore tip, whereas in
|
|
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Luzonomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Luzonomorpha" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Luzonomorpha</taxonomicName>
|
|
the tip is deeply bispinose. A similarly poorly bi- or trifid gonopod is only observed in
|
|
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorphoides" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorphoides" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Orthomorphoides</taxonomicName>
|
|
gen. n., but its species differ from
|
|
<taxonomicName class="Diplopoda" family="Paradoxosomatidae" genus="Orthomorpha" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Orthomorpha" order="Polydesmida" pageId="5" pageNumber="6" phylum="Arthropoda" rank="genus">Orthomorpha</taxonomicName>
|
|
in much smaller bodies, in the rather poorly developed paraterga and at least in some of the apical prongs of the solenophore being short spines (see below).
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |