treatments-xml/data/65/8E/D9/658ED9F6309A5704B883EA9244F5743E.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.891.38563" ID-GBIF-Dataset="45a6be79-8f9b-4fa6-9687-bb3d57a3d54a" ID-PMC="PMC6882923" ID-Pensoft-Pub="1313-2970-891-17" ID-Pensoft-UUID="E45641CF778A5EDE9155B5D9DD2027BD" ID-PubMed="31802970" ID-ZooBank="26724E9CABBB41E985A287021244E574" ModsDocID="1313-2970-891-17" checkinTime="1574358819888" checkinUser="pensoft" docAuthor="Maddison, Wayne P." docDate="2019" docId="658ED9F6309A5704B883EA9244F5743E" docLanguage="en" docName="ZooKeys 891: 17-29" docOrigin="ZooKeys 891" docSource="http://dx.doi.org/10.3897/zookeys.891.38563" docTitle="Lapsias cyrboides Simon 1900" docType="treatment" docVersion="4" id="E45641CF778A5EDE9155B5D9DD2027BD" lastPageNumber="17" masterDocId="E45641CF778A5EDE9155B5D9DD2027BD" masterDocTitle="A new lapsiine jumping spider from North America, with a review of Simon's Lapsias species (Araneae, Salticidae, Spartaeinae)" masterLastPageNumber="29" masterPageNumber="17" pageNumber="17" updateTime="1668168118219" updateUser="ExternalLinkService">
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<mods:title>A new lapsiine jumping spider from North America, with a review of Simon's Lapsias species (Araneae, Salticidae, Spartaeinae)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Maddison, Wayne P.</mods:namePart>
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<mods:title>ZooKeys</mods:title>
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<mods:date>2019</mods:date>
<mods:detail type="volume">
<mods:number>891</mods:number>
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<mods:start>17</mods:start>
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<mods:url>http://dx.doi.org/10.3897/zookeys.891.38563</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.891.38563</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-891-17</mods:identifier>
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<treatment ID-GBIF-Taxon="160511235" LSID="urn:lsid:plazi:treatment:658ED9F6309A5704B883EA9244F5743E" httpUri="http://treatment.plazi.org/id/658ED9F6309A5704B883EA9244F5743E" lastPageNumber="17" pageId="0" pageNumber="17">
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<paragraph pageId="0" pageNumber="17">
<taxonomicName LSID="658ED9F6-309A-5704-B883-EA9244F5743E" authority="Simon, 1900" authorityName="Simon" authorityYear="1900" class="Arachnida" family="Salticidae" genus="Lapsias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lapsias cyrboides" order="Araneae" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="cyrboides">Lapsias cyrboides Simon, 1900</taxonomicName>
<figureCitation captionStart="Figures 30–40" captionStartId="F4" captionText="Figures 30 - 40. Lapsias cyrboides lectotype male (30 - 34, 36), paralectotype male (35), and female tentatively considered of this species (37 - 40). 30 - 32 Palp 30 ventral view 31 retrolateral view of tibia 32 ventral view 33 body from above 34 body from below 35 paralectotype male body from above 36 lectotype male face 37 epigyne from below 38 vulva from above 39 female body from above 40 body from below. Scale bars: 0.1 mm (30); 1.0 mm (otherwise)." figureDoi="10.3897/zookeys.891.38563.figures30-40" httpUri="https://binary.pensoft.net/fig/357965" pageId="0" pageNumber="17" tableDoi="10.3897/zookeys.891.38563.figures30-40">Figs 30-40</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="type material">
<paragraph pageId="0" pageNumber="17">Type material.</paragraph>
<paragraph pageId="0" pageNumber="17">
In MNHN, 3 males, 4 females, 3 juveniles from Colonia Tovar, Aragua State, Venezuela, in a vial with label &quot;20924 Laps. 
<taxonomicName lsidName="cyrboides" pageId="0" pageNumber="17" rank="species" species="cyrboides">cyrboides</taxonomicName>
E.S., Tovar!&quot; and a more recent label &quot;det 
<normalizedToken originalValue="Szűts">Szűts</normalizedToken>
0014&quot;. 
<bibRefCitation author="Galiano, ME" journalOrPublisher="Physis, Revista de la Sociedad Argentina de Ciencias Naturales (C)" pageId="0" pageNumber="17" pagination="273 - 470" refId="B2" refString="Galiano, ME, 1963. Las especies americanas de aranas de la familia Salticidae descriptas por Eugene Simon: Redescripciones basadas en los ejemplares tipicos. Physis, Revista de la Sociedad Argentina de Ciencias Naturales (C) 23: 273 - 470" title="Las especies americanas de aranas de la familia Salticidae descriptas por Eugene Simon: Redescripciones basadas en los ejemplares tipicos." volume="23" year="1963">Galiano (1963)</bibRefCitation>
designated one male as a lectotype, in separate microvial with her label &quot;Typus? M.E. Galiano II 1959&quot;. She mentions one female designated also as lectotype, but no female is separated and labelled as such. Because Galiano (incorrectly) designated two lectotypes, the name is not yet fixed to a single specimen. This ambiguity is resolved by designating her male lectotype as the only lectotype.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="17" type="notes">
<paragraph pageId="0" pageNumber="17">Notes.</paragraph>
<paragraph pageId="0" pageNumber="17">
<bibRefCitation author="Simon, E" journalOrPublisher="Annales de la Societe Entomologique de Belgique" pageId="0" pageNumber="17" pagination="381 - 407" refId="B12" refString="Simon, E, 1900. Descriptions d'arachnides nouveaux de la famille des Attidae. Annales de la Societe Entomologique de Belgique 44: 381 - 407" title="Descriptions d'arachnides nouveaux de la famille des Attidae." volume="44" year="1900">Simon (1900)</bibRefCitation>
described a male and female. However, as noted by 
<bibRefCitation author="Galiano, ME" journalOrPublisher="Physis, Revista de la Sociedad Argentina de Ciencias Naturales (C)" pageId="0" pageNumber="17" pagination="273 - 470" refId="B2" refString="Galiano, ME, 1963. Las especies americanas de aranas de la familia Salticidae descriptas por Eugene Simon: Redescripciones basadas en los ejemplares tipicos. Physis, Revista de la Sociedad Argentina de Ciencias Naturales (C) 23: 273 - 470" title="Las especies americanas de aranas de la familia Salticidae descriptas por Eugene Simon: Redescripciones basadas en los ejemplares tipicos." volume="23" year="1963">Galiano (1963)</bibRefCitation>
, there are two species of female among the four females in the type vial, similar in body but easily distinguished by the epigyne. Two of the females (
<figureCitation captionStart="Figures 30–40" captionStartId="F4" captionText="Figures 30 - 40. Lapsias cyrboides lectotype male (30 - 34, 36), paralectotype male (35), and female tentatively considered of this species (37 - 40). 30 - 32 Palp 30 ventral view 31 retrolateral view of tibia 32 ventral view 33 body from above 34 body from below 35 paralectotype male body from above 36 lectotype male face 37 epigyne from below 38 vulva from above 39 female body from above 40 body from below. Scale bars: 0.1 mm (30); 1.0 mm (otherwise)." figureDoi="10.3897/zookeys.891.38563.figures30-40" httpUri="https://binary.pensoft.net/fig/357965" pageId="0" pageNumber="17" tableDoi="10.3897/zookeys.891.38563.figures30-40">Figs 37-40</figureCitation>
) have an anteriorly placed guide (
<figureCitation captionStart="Figures 30–40" captionStartId="F4" captionText="Figures 30 - 40. Lapsias cyrboides lectotype male (30 - 34, 36), paralectotype male (35), and female tentatively considered of this species (37 - 40). 30 - 32 Palp 30 ventral view 31 retrolateral view of tibia 32 ventral view 33 body from above 34 body from below 35 paralectotype male body from above 36 lectotype male face 37 epigyne from below 38 vulva from above 39 female body from above 40 body from below. Scale bars: 0.1 mm (30); 1.0 mm (otherwise)." figureDoi="10.3897/zookeys.891.38563.figures30-40" httpUri="https://binary.pensoft.net/fig/357965" pageId="0" pageNumber="17" tableDoi="10.3897/zookeys.891.38563.figures30-40">Fig. 37</figureCitation>
), while the other two females (
<figureCitation captionStart="Figures 41–51" captionStartId="F5" captionText="Figures 41 - 51. Lapsias tovarensis lectotype male (41 - 47) and female tentatively considered of this species (48 - 51). 41 - 43 Palp 41 ventral view 42 retrolateral view of tibia 43 ventral view 44 body from above 45 body from below 46 face 47 chelicerae from below 48 epigyne from below 49 vulva from above 50 female body from above 51 body from below. Abbreviations: c? sclerite homologous to that called the conductor in other lapsiines. Scale bars: 0.1 mm (41); 1.0 mm (otherwise)." figureDoi="10.3897/zookeys.891.38563.figures41-51" httpUri="https://binary.pensoft.net/fig/357966" pageId="0" pageNumber="17" tableDoi="10.3897/zookeys.891.38563.figures41-51">Figs 48-51</figureCitation>
) lack such a guide and instead show two wing-shaped atria extending laterally (
<figureCitation captionStart="Figures 41–51" captionStartId="F5" captionText="Figures 41 - 51. Lapsias tovarensis lectotype male (41 - 47) and female tentatively considered of this species (48 - 51). 41 - 43 Palp 41 ventral view 42 retrolateral view of tibia 43 ventral view 44 body from above 45 body from below 46 face 47 chelicerae from below 48 epigyne from below 49 vulva from above 50 female body from above 51 body from below. Abbreviations: c? sclerite homologous to that called the conductor in other lapsiines. Scale bars: 0.1 mm (41); 1.0 mm (otherwise)." figureDoi="10.3897/zookeys.891.38563.figures41-51" httpUri="https://binary.pensoft.net/fig/357966" pageId="0" pageNumber="17" tableDoi="10.3897/zookeys.891.38563.figures41-51">Fig. 48</figureCitation>
). It is reasonable to assume that these two kinds of female belong to the two smaller-bodied 
<taxonomicName class="Arachnida" family="Salticidae" genus="Lapsias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lapsias" order="Araneae" pageId="0" pageNumber="17" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="17">Lapsias</emphasis>
</taxonomicName>
at Colonia Tovar, 
<taxonomicName lsidName="L. cyrboides" pageId="0" pageNumber="17" rank="species" species="cyrboides">
<emphasis italics="true" pageId="0" pageNumber="17">L. cyrboides</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="L. tovarensis" pageId="0" pageNumber="17" rank="species" species="tovarensis">
<emphasis italics="true" pageId="0" pageNumber="17">L. tovarensis</emphasis>
</taxonomicName>
. Under 
<taxonomicName lsidName="L. cyrboides" pageId="0" pageNumber="17" rank="species" species="cyrboides">
<emphasis italics="true" pageId="0" pageNumber="17">L. cyrboides</emphasis>
</taxonomicName>
Simon described the female kind with anterior guide (&quot;Plaga genitalis...longior quam latior&quot;), but he did not justify this choice nor even mention the second form of female. Galiano followed 
<normalizedToken originalValue="Simon’s">Simon's</normalizedToken>
choice of matching female. The two forms of female are approximately the same size and carapace shape and are too faded to supply distinctive markings by which to match to the males. Nonetheless, I tentatively support 
<normalizedToken originalValue="Simon’s">Simon's</normalizedToken>
and 
<normalizedToken originalValue="Galiano’s">Galiano's</normalizedToken>
matching based on an expected correlation between the form of the 
<normalizedToken originalValue="female’s">female's</normalizedToken>
guide and that of the 
<normalizedToken originalValue="male’s">male's</normalizedToken>
RTA. An anterior guide is expected to be associated with an extraordinary RTA, for instance, as in 
<taxonomicName class="Arachnida" family="Salticidae" genus="Papuamyr" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Papuamyr omhifosa" order="Araneae" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="omhifosa">
<emphasis italics="true" pageId="0" pageNumber="17">Papuamyr omhifosa</emphasis>
</taxonomicName>
Maddison &amp; 
<normalizedToken originalValue="Szűts">Szűts</normalizedToken>
, 2019 (
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.842.32970" author="Maddison, WP" journalOrPublisher="ZooKeys" pageId="0" pageNumber="17" pagination="85 - 112" refId="B8" refString="Maddison, WP, Szűts, T, 2019. Myrmarachnine jumping spiders of the new subtribe Levieina from Papua New Guinea (Araneae, Salticidae, Myrmarachnini). ZooKeys 842: 85 - 112" title="Myrmarachnine jumping spiders of the new subtribe Levieina from Papua New Guinea (Araneae, Salticidae, Myrmarachnini)." url="https://doi.org/10.3897/zookeys.842.32970" volume="842" year="2019">
Maddison and 
<normalizedToken originalValue="Szűts">Szűts</normalizedToken>
2019
</bibRefCitation>
). Males of 
<taxonomicName lsidName="L. ciliatus" pageId="0" pageNumber="17" rank="species" species="ciliatus">
<emphasis italics="true" pageId="0" pageNumber="17">L. ciliatus</emphasis>
</taxonomicName>
and 
<taxonomicName lsidName="L. tovarensis" pageId="0" pageNumber="17" rank="species" species="tovarensis">
<emphasis italics="true" pageId="0" pageNumber="17">L. tovarensis</emphasis>
</taxonomicName>
have small RTAs, suggesting a simple or small guide along the epigastric furrow, as is common in salticids, while the male of 
<taxonomicName lsidName="L. cyrboides" pageId="0" pageNumber="17" rank="species" species="cyrboides">
<emphasis italics="true" pageId="0" pageNumber="17">L. cyrboides</emphasis>
</taxonomicName>
has an unusual dorsally projecting tibial apophysis, which predicts an unusually-placed female guide. Thus, the female with anterior guide is tentatively considered that of 
<taxonomicName lsidName="L. cyrboides" pageId="0" pageNumber="17" rank="species" species="cyrboides">
<emphasis italics="true" pageId="0" pageNumber="17">L. cyrboides</emphasis>
</taxonomicName>
, and the female with wing-shaped atria is considered that of 
<taxonomicName lsidName="L. tovarensis" pageId="0" pageNumber="17" rank="species" species="tovarensis">
<emphasis italics="true" pageId="0" pageNumber="17">L. tovarensis</emphasis>
</taxonomicName>
.
</paragraph>
<caption doi="10.3897/zookeys.891.38563.figures30-40" httpUri="https://binary.pensoft.net/fig/357965" pageId="0" pageNumber="17" start="Figures 30–40" startId="F4">
<paragraph pageId="0" pageNumber="17">
<emphasis bold="true" pageId="0" pageNumber="17">Figures 30-40.</emphasis>
<taxonomicName class="Arachnida" family="Salticidae" genus="Lapsias" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Lapsias cyrboides" order="Araneae" pageId="0" pageNumber="17" phylum="Arthropoda" rank="species" species="cyrboides">
<emphasis italics="true" pageId="0" pageNumber="17">Lapsias cyrboides</emphasis>
</taxonomicName>
lectotype male (
<emphasis bold="true" pageId="0" pageNumber="17">30-34, 36</emphasis>
), paralectotype male (
<emphasis bold="true" pageId="0" pageNumber="17">35</emphasis>
), and female tentatively considered of this species (
<emphasis bold="true" pageId="0" pageNumber="17">37-40</emphasis>
). 
<emphasis bold="true" pageId="0" pageNumber="17">30-32</emphasis>
Palp 
<emphasis bold="true" pageId="0" pageNumber="17">30</emphasis>
ventral view 
<emphasis bold="true" pageId="0" pageNumber="17">31</emphasis>
retrolateral view of tibia 
<emphasis bold="true" pageId="0" pageNumber="17">32</emphasis>
ventral view 
<emphasis bold="true" pageId="0" pageNumber="17">33</emphasis>
body from above 
<emphasis bold="true" pageId="0" pageNumber="17">34</emphasis>
body from below 
<emphasis bold="true" pageId="0" pageNumber="17">35</emphasis>
paralectotype male body from above 
<emphasis bold="true" pageId="0" pageNumber="17">36</emphasis>
lectotype male face 
<emphasis bold="true" pageId="0" pageNumber="17">37</emphasis>
epigyne from below 
<emphasis bold="true" pageId="0" pageNumber="17">38</emphasis>
vulva from above 
<emphasis bold="true" pageId="0" pageNumber="17">39</emphasis>
female body from above 
<emphasis bold="true" pageId="0" pageNumber="17">40</emphasis>
body from below. Scale bars: 0.1 mm (
<emphasis bold="true" pageId="0" pageNumber="17">30</emphasis>
); 1.0 mm (otherwise).
</paragraph>
</caption>
</subSubSection>
</treatment>
</document>