178 lines
22 KiB
XML
178 lines
22 KiB
XML
<document id="E9DDC90B85F517146F0453FC97661604" ID-DOI="10.11646/zootaxa.5047.1.6" ID-ISSN="1175-5326" ID-Zenodo-Dep="5533748" ID-ZooBank="E48D4D6E-F5C9-4DC1-9257-E46677019AE2" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1632814976319" checkinUser="plazi" docAuthor="Muadsub, Sopita & Pinkaew, Nantasak" docDate="2021" docId="657687AC3E664239FF74FC7AFF5CF831" docLanguage="en" docName="zootaxa.5047.1.6.pdf" docOrigin="Zootaxa 5047 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Loboschiza sictriquetra Muadsub & Pinkaew 2021, sp. nov." docType="treatment" docVersion="3" lastPageNumber="78" masterDocId="994FFFD43E6C4233FFE3FFBDFF87FFC7" masterDocTitle="Three new species of Loboschiza Diakonoff, 1968 (Lepidoptera: Tortricidae Olethreutinae) with redescriptions of L. mediana (Walker, 1864) and L. martia (Meyrick, 1911), two new records from Thailand" masterLastPageNumber="80" masterPageNumber="68" pageNumber="78" updateTime="1699416056941" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="E0237A3A5E199CF1C3EB96EAA981FEFA">Three new species of Loboschiza Diakonoff, 1968 (Lepidoptera: Tortricidae Olethreutinae) with redescriptions of L. mediana (Walker, 1864) and L. martia (Meyrick, 1911), two new records from Thailand</mods:title>
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<mods:name id="94A903EE70B60A54916B8DFB4CBFA594" type="personal">
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<mods:role id="EFBEB7A088E44C82F02FFCA309EF8F3F">
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<mods:roleTerm id="CC943B254DC2E266A2C578178722395A">Author</mods:roleTerm>
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<mods:namePart id="ED89E45D4E2086ACD60C6A976ED5823C">Muadsub, Sopita</mods:namePart>
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<mods:affiliation id="9C2848139F63CFF609259689AA87A67A">Department of Entomology, Faculty of Agriculture at Kamphaeng Saen, Kasetsart University, Kamphaeng Saen Campus, Nakhon Pathom, 73140, Thailand</mods:affiliation>
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<mods:nameIdentifier id="846CC28F2F86F6510B5C4D140A026815" type="email">muadsub_s@hotmail.com</mods:nameIdentifier>
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<mods:roleTerm id="8EDCAD6592A2F0A9AB51F7640F3F4DE2">Author</mods:roleTerm>
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<mods:namePart id="234FD135FE1126A75C2F7888D8FEB968">Pinkaew, Nantasak</mods:namePart>
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<mods:affiliation id="949A2B104F3C2F2DDF1C974C2015C82F">Department of Entomology, Faculty of Agriculture at Kamphaeng Saen, Kasetsart University, Kamphaeng Saen Campus, Nakhon Pathom, 73140, Thailand & Center for Advanced Studies in Tropical Natural Resources, NRU-KU, Kasetsart University, Chatuchak, Bangkok, 10900, Thailand. muadsub _ s @ hotmail. com; https: // orcid. org / 0000 - 0002 - 9672 - 1878</mods:affiliation>
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<mods:title id="4683DD1C1974B60F0CFB33BABBEA76E7">Zootaxa</mods:title>
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<mods:date id="D180168DE1147675A0990FA3C53507D5">2021</mods:date>
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<mods:number id="25205DA54B80C9E69F4FBF12BE6D2C0A">2021-09-28</mods:number>
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<mods:number id="600597B99C199756CF4A3A534ED412BF">5047</mods:number>
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<mods:identifier id="891AC4E8F25DE851F7F125643BC4EC21" type="DOI">10.11646/zootaxa.5047.1.6</mods:identifier>
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<taxonomicName id="2ADF4D393E664239FF74FC7AFE35FC25" authority="Muadsub & Pinkaew, 2021" authorityName="Muadsub & Pinkaew" authorityYear="2021" box="[151,434,967,994]" class="Insecta" family="Tortricidae" genus="Loboschiza" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="78" phylum="Arthropoda" rank="species" species="sictriquetra" status="sp. nov.">
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<emphasis id="DFABEAA83E664239FF74FC7AFE35FC25" bold="true" box="[151,434,967,994]" italics="true" pageId="10" pageNumber="78">Loboschiza sictriquetra</emphasis>
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<taxonomicNameLabel id="C49857D33E664239FE5AFC75FD90FC25" box="[441,535,968,994]" pageId="10" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="A5C565313E664239FF74FBADFE19FBED" box="[151,414,1040,1067]" pageId="10" pageNumber="78" type="description">
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<paragraph id="ED6036BA3E664239FF74FBADFE19FBED" blockId="10.[151,954,967,1067]" box="[151,414,1040,1067]" pageId="10" pageNumber="78">
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(
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<figureCitation id="75E42A3F3E664239FF7CFBADFF68FBEC" box="[159,239,1040,1067]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1921,1945]" captionTargetBox="[168,1418,748,1890]" captionTargetId="figure-255@2.[160,1428,741,1898]" captionTargetPageId="2" captionText="FIGURES 1–5. Wing pattern of Loboschiza spp. 1. L. mediana (male, np12141). 2. L. martia (male, np8493). 3. L. ellipsa sp. nov. (holotype, male). 4. L. namphromensis sp. nov. (holotype, male). 5. L. sictriquetra sp. nov. (holotype, male), scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5533750" httpUri="https://zenodo.org/record/5533750/files/figure.png" pageId="10" pageNumber="78">Figs. 5</figureCitation>
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,
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<figureCitation id="75E42A3F3E664239FF18FBADFE9EFBED" box="[251,281,1040,1066]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1951,1975]" captionTargetBox="[214,1373,181,1927]" captionTargetId="figure-27@3.[214,1373,181,1927]" captionTargetPageId="3" captionText="FIGURES 6–10. Labial palpi of Loboschiza spp. 6. L. mediana (male, np2536). 7. L. martia (male, np8493). 8. L. ellipsa sp. nov. (holotype, male). 9. L. namphromensis sp. nov. (holotype, male). 10. L. sictriquetra sp. nov. (holotype, male)." figureDoi="http://doi.org/10.5281/zenodo.5533752" httpUri="https://zenodo.org/record/5533752/files/figure.png" pageId="10" pageNumber="78">10</figureCitation>
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,
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<figureCitation id="75E42A3F3E664239FEC6FBADFEC3FBED" box="[293,324,1040,1066]" captionStart="FIGURES 17–20" captionStartId="8.[151,264,1919,1943]" captionTargetBox="[285,1298,196,1880]" captionTargetId="figure-17@8.[278,1308,181,1895]" captionTargetPageId="8" captionText="FIGURES 17–20. Abdominal segments of male of Loboschiza spp. 17. L. mediana (NP3958). 18. L. ellipsa sp. nov. (paratype, NP3699). 19. L. namphromensis sp. nov. (paratype, NP3701). 20. L. sictriquetra sp. nov. (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5533762" httpUri="https://zenodo.org/record/5533762/files/figure.png" pageId="10" pageNumber="78">20</figureCitation>
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,
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<figureCitation id="75E42A3F3E664239FEACFBADFE11FBED" box="[335,406,1040,1066]" captionStart="FIGURES 21–28" captionStartId="9.[151,264,1788,1812]" captionTargetBox="[161,1425,181,1765]" captionTargetId="figure-27@9.[161,1425,181,1765]" captionTargetPageId="9" captionText="FIGURES 21–28. Male genitalia of Loboschiza spp. 21. L. mediana (NP3957). 22. L. martia (NP3078). 23–24. L. ellipsa sp. nov. (holotype). 25–26. L. namphromensis sp. nov. (holotype). 27–28. L. sictriquetra sp. nov. (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5533764" httpUri="https://zenodo.org/record/5533764/files/figure.png" pageId="10" pageNumber="78">27–28</figureCitation>
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)
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</paragraph>
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<subSubSection id="A5C565313E664239FF74FBE5FA0FFB51" pageId="10" pageNumber="78" type="materials_examined">
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<paragraph id="ED6036BA3E664239FF74FBE5FA0FFB51" blockId="10.[151,1437,1112,2038]" pageId="10" pageNumber="78">
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<emphasis id="DFABEAA83E664239FF74FBE5FEC9FBB5" bold="true" box="[151,334,1112,1138]" pageId="10" pageNumber="78">Type Material.</emphasis>
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<materialsCitation id="5DB73CE73E664239FEB8FBE5FA0FFB51" collectingDate="2020-12-16" collectionCode="KKIC" collectorName="Huai Kha Khaeng W. S." country="Thailand" elevation="550" latitude="15.490833" location="Uthai Thani Prov." longLatPrecision="21" longitude="99.300835" pageId="10" pageNumber="78" specimenCount="1" specimenCount-male="1" stateProvince="Uthai Thani" typeStatus="holotype">
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<typeStatus id="326488183E664239FEB8FBE5FE4FFBB5" box="[347,456,1112,1138]" pageId="10" pageNumber="78" type="holotype">Holotype</typeStatus>
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:
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<specimenCount id="FBD9FD333E664239FE3BFBE4FE77FBB4" box="[472,496,1113,1139]" pageId="10" pageNumber="78" type="male">♂</specimenCount>
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;
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<collectingCountry id="95C8762A3E664239FDE2FBE5FD12FBB5" box="[513,661,1112,1138]" name="Thailand" pageId="10" pageNumber="78">THAILAND</collectingCountry>
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.
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<collectingRegion id="2F1BF8583E664239FD45FBE5FCFBFBB4" box="[678,892,1112,1139]" country="Thailand" name="Uthai Thani" pageId="10" pageNumber="78">Uthai Thani Prov.</collectingRegion>
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,
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<collectorName id="402A536C3E664239FC6EFBE5FB20FBB5" box="[909,1191,1112,1138]" pageId="10" pageNumber="78">Huai Kha Khaeng W.S.</collectorName>
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, (Khao Nang Rum),
|
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<geoCoordinate id="88EB507D3E664239FF74FBC1FE9CFB5F" box="[151,283,1148,1176]" degrees="15" direction="north" minutes="29" orientation="latitude" pageId="10" pageNumber="78" precision="15" seconds="27" value="15.490833">
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15
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<superScript id="1AAA9BF23E664239FF50FBC1FF3CFB4D" attach="none" box="[179,187,1148,1162]" fontSize="6" pageId="10" pageNumber="78">o</superScript>
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29ʹ27ʺN
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</geoCoordinate>
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,
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<geoCoordinate id="88EB507D3E664239FEC5FBC1FE21FB5F" box="[294,422,1148,1176]" degrees="99" direction="east" minutes="18" orientation="longitude" pageId="10" pageNumber="78" precision="15" seconds="03" value="99.300835">
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99
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18ʹ03ʺE
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</geoCoordinate>
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, alt.
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<quantity id="2A279B5F3E664239FE3FFBC1FDA1FB50" box="[476,550,1148,1175]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="10" pageNumber="78" unit="m" value="550.0">
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<elevation id="66F2D1893E664239FE3FFBC1FDA1FB50" box="[476,550,1148,1175]" metricMagnitude="2" metricUnit="m" metricValue="5.5" pageId="10" pageNumber="78" unit="m" value="550.0">550 m</elevation>
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</quantity>
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;
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<date id="9961107A3E664239FDD2FBC1FD4CFB51" box="[561,715,1148,1175]" pageId="10" pageNumber="78" value="2020-12-16">
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<collectingDate id="8925E9923E664239FDD2FBC1FD4CFB51" box="[561,715,1148,1175]" pageId="10" pageNumber="78" value="2020-12-16">16 Dec. 2020</collectingDate>
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</date>
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; Pinkaew et al. leg.; np13625 (genitalia slide NP3965);
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<collectionCode id="8BCEAE7F3E664239FADFFBC1FA03FB51" box="[1340,1412,1148,1174]" pageId="10" pageNumber="78">KKIC</collectionCode>
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.
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="A5C565313E664239FF24FB1DFC1DFB19" pageId="10" pageNumber="78" type="diagnosis">
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<paragraph id="ED6036BA3E664239FF24FB1DFC1DFB19" blockId="10.[151,1437,1112,2038]" pageId="10" pageNumber="78">
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<emphasis id="DFABEAA83E664239FF24FB1DFEC5FB7D" bold="true" box="[199,322,1184,1210]" pageId="10" pageNumber="78">Diagnosis.</emphasis>
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<taxonomicName id="2ADF4D393E664239FEA9FB1DFDD1FB7D" authority="Muadsub & Pinkaew, 2021" authorityName="Muadsub & Pinkaew" authorityYear="2021" box="[330,598,1184,1210]" class="Insecta" family="Tortricidae" genus="Loboschiza" kingdom="Animalia" order="Lepidoptera" pageId="10" pageNumber="78" phylum="Arthropoda" rank="species" species="sictriquetra" status="sp. nov.">
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<emphasis id="DFABEAA83E664239FEA9FB1DFDD1FB7D" box="[330,598,1184,1210]" italics="true" pageId="10" pageNumber="78">Loboschiza sictriquetra</emphasis>
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</taxonomicName>
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<emphasis id="DFABEAA83E664239FDBCFB1DFD3CFB7D" bold="true" box="[607,699,1184,1210]" pageId="10" pageNumber="78">
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<taxonomicNameLabel id="C49857D33E664239FDBCFB1DFD3CFB7D" box="[607,699,1184,1210]" pageId="10" pageNumber="78" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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differs from all other species in the genus by the conspicuously large socii and the elongate subtriangular valva in the male genitalia.
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</paragraph>
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</subSubSection>
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<subSubSection id="A5C565313E664239FF24FB55FCA1F84D" pageId="10" pageNumber="78" type="description">
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<paragraph id="ED6036BA3E664239FF24FB55FE01FA55" blockId="10.[151,1437,1112,2038]" pageId="10" pageNumber="78">
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<emphasis id="DFABEAA83E664239FF24FB55FEDEFAC5" bold="true" box="[199,345,1256,1282]" pageId="10" pageNumber="78">Description.</emphasis>
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Head (
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<figureCitation id="75E42A3F3E664239FE4EFB55FD85FAC5" box="[429,514,1256,1282]" captionStart="FIGURES 6–10" captionStartId="3.[151,264,1951,1975]" captionTargetBox="[214,1373,181,1927]" captionTargetId="figure-27@3.[214,1373,181,1927]" captionTargetPageId="3" captionText="FIGURES 6–10. Labial palpi of Loboschiza spp. 6. L. mediana (male, np2536). 7. L. martia (male, np8493). 8. L. ellipsa sp. nov. (holotype, male). 9. L. namphromensis sp. nov. (holotype, male). 10. L. sictriquetra sp. nov. (holotype, male)." figureDoi="http://doi.org/10.5281/zenodo.5533752" httpUri="https://zenodo.org/record/5533752/files/figure.png" pageId="10" pageNumber="78">Fig. 10</figureCitation>
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): Lower frons orange white, upper frons and vertex pale yellowish orange; labial palpi porrect, first segment orange white, second segment broadened apically, orange white, gradually darkening to pale yellowish orange apically, with a small brown mark dorsomedially and a larger brown mark apically, third segment pale yellowish orange, slightly paler toward apex; antenna blackish, basal segments mixed with pale yellowish orange scales.
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</paragraph>
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<paragraph id="ED6036BA3E664239FF24FA21FAB2F949" blockId="10.[151,1437,1112,2038]" pageId="10" pageNumber="78">
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Thorax: Pronotal collar greyish orange; tegulae light grey with orange patch outer 1/2, mesonotum dark greyish brown, anterior 1/3 and posterior 1/3 mixed with orange scales; posterior crest absent. Forewing subrectangular, wingspan 10.4 mm in male (
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<figureCitation id="75E42A3F3E664239FE34FA59FD9EFA38" box="[471,537,1508,1535]" captionStart="FIGURES 1–5" captionStartId="2.[151,264,1921,1945]" captionTargetBox="[168,1418,748,1890]" captionTargetId="figure-255@2.[160,1428,741,1898]" captionTargetPageId="2" captionText="FIGURES 1–5. Wing pattern of Loboschiza spp. 1. L. mediana (male, np12141). 2. L. martia (male, np8493). 3. L. ellipsa sp. nov. (holotype, male). 4. L. namphromensis sp. nov. (holotype, male). 5. L. sictriquetra sp. nov. (holotype, male), scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5533750" httpUri="https://zenodo.org/record/5533750/files/figure.png" pageId="10" pageNumber="78">Fig. 5</figureCitation>
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) (n = 1); costal margin evenly curved; termen rather straight and oblique; costal strigulae well developed, yellowish orange, separated by blackish streaks; wing pattern two-toned, basal 2/3 with light greyish brown ground color, with variable well-defined orange spots and bands, apical 1/3 with transverse, irregular sinuous striae on blackish ground, fringe scales blackish; underside light greyish brown with white spots along costa and termen. Hindwing light brown, slightly paler toward base; underside light greyish brown.
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</paragraph>
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<paragraph id="ED6036BA3E664239FF24F925FCA1F84D" blockId="10.[151,1437,1112,2038]" pageId="10" pageNumber="78">
|
||
Abdomen: Posterior margin of sternum VIII bilobed (
|
||
<figureCitation id="75E42A3F3E664239FCCCF925FC02F975" box="[815,901,1688,1714]" captionStart="FIGURES 17–20" captionStartId="8.[151,264,1919,1943]" captionTargetBox="[285,1298,196,1880]" captionTargetId="figure-17@8.[278,1308,181,1895]" captionTargetPageId="8" captionText="FIGURES 17–20. Abdominal segments of male of Loboschiza spp. 17. L. mediana (NP3958). 18. L. ellipsa sp. nov. (paratype, NP3699). 19. L. namphromensis sp. nov. (paratype, NP3701). 20. L. sictriquetra sp. nov. (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5533762" httpUri="https://zenodo.org/record/5533762/files/figure.png" pageId="10" pageNumber="78">Fig. 20</figureCitation>
|
||
). Male genitalia (
|
||
<figureCitation id="75E42A3F3E664239FBACF925FB24F975" box="[1103,1187,1688,1714]" captionStart="FIGURES 21–28" captionStartId="9.[151,264,1788,1812]" captionTargetBox="[161,1425,181,1765]" captionTargetId="figure-27@9.[161,1425,181,1765]" captionTargetPageId="9" captionText="FIGURES 21–28. Male genitalia of Loboschiza spp. 21. L. mediana (NP3957). 22. L. martia (NP3078). 23–24. L. ellipsa sp. nov. (holotype). 25–26. L. namphromensis sp. nov. (holotype). 27–28. L. sictriquetra sp. nov. (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5533764" httpUri="https://zenodo.org/record/5533764/files/figure.png" pageId="10" pageNumber="78">Fig. 27</figureCitation>
|
||
) with tegumen ovate, apodeme for muscle m4 enlarged, truncate and oblique, extended inwardly near other; uncus absent; socii large, sinuate, 1/2 length of tegumen, densely setose; gnathos indistinct; vinculum small, rather narrow; juxta small, subtriangular; caulis short; anellus surrounding base of phallus; phallus subcylindrical, moderately long and wide, slightly tapered apically, with dense cornuti; valva elongate-subtriangular, densely setose, widest medially, slightly narrow to rounded apex, inner margin slightly curved, median of outer margin extending outward forming a pointed apex, with a small, short, upwardly curved spine (
|
||
<figureCitation id="75E42A3F3E664239FD26F8CDFC9DF84D" box="[709,794,1904,1930]" captionStart="FIGURES 21–28" captionStartId="9.[151,264,1788,1812]" captionTargetBox="[161,1425,181,1765]" captionTargetId="figure-27@9.[161,1425,181,1765]" captionTargetPageId="9" captionText="FIGURES 21–28. Male genitalia of Loboschiza spp. 21. L. mediana (NP3957). 22. L. martia (NP3078). 23–24. L. ellipsa sp. nov. (holotype). 25–26. L. namphromensis sp. nov. (holotype). 27–28. L. sictriquetra sp. nov. (holotype)." figureDoi="http://doi.org/10.5281/zenodo.5533764" httpUri="https://zenodo.org/record/5533764/files/figure.png" pageId="10" pageNumber="78">Fig. 28</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A5C565313E664239FF24F829FD54F868" box="[199,723,1940,1967]" pageId="10" pageNumber="78" type="distribution">
|
||
<paragraph id="ED6036BA3E664239FF24F829FD54F868" blockId="10.[151,1437,1112,2038]" box="[199,723,1940,1967]" pageId="10" pageNumber="78">
|
||
<emphasis id="DFABEAA83E664239FF24F829FEDEF869" bold="true" box="[199,345,1940,1966]" pageId="10" pageNumber="78">Distribution</emphasis>
|
||
.
|
||
<collectingCountry id="95C8762A3E664239FE84F829FE4CF869" box="[359,459,1940,1966]" name="Thailand" pageId="10" pageNumber="78">Thailand</collectingCountry>
|
||
(
|
||
<collectingRegion id="2F1BF8583E664239FE3AF829FD4BF868" box="[473,716,1940,1967]" country="Thailand" name="Uthai Thani" pageId="10" pageNumber="78">Uthai Thani province</collectingRegion>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="A5C565313E664239FF24F805FF5CF831" pageId="10" pageNumber="78" type="etymology">
|
||
<paragraph id="ED6036BA3E664239FF24F805FF5CF831" blockId="10.[151,1437,1112,2038]" pageId="10" pageNumber="78">
|
||
<emphasis id="DFABEAA83E664239FF24F805FECFF815" bold="true" box="[199,328,1976,2002]" pageId="10" pageNumber="78">Etymology</emphasis>
|
||
. The specific epithet (
|
||
<emphasis id="DFABEAA83E664239FDAFF804FD00F815" box="[588,647,1977,2002]" italics="true" pageId="10" pageNumber="78">sic =</emphasis>
|
||
like/
|
||
<emphasis id="DFABEAA83E664239FD28F804FD62F815" box="[715,741,1977,2002]" italics="true" pageId="10" pageNumber="78">tri</emphasis>
|
||
= three/
|
||
<emphasis id="DFABEAA83E664239FCB3F804FC1EF815" box="[848,921,1977,2002]" italics="true" pageId="10" pageNumber="78">quetra</emphasis>
|
||
= angled or sided) refers to the shape of the valva.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |