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<mods:title id="C025EEE53F427BF8B7E41B26A96F6293">Updated catalogue and taxonomic notes on the Old-World scorpion genus Buthus Leach, 1815 (Scorpiones, Buthidae)</mods:title>
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<treatment id="6553F2398FA7F2C374E7163BE0665CE3" ID-GBIF-Taxon="132648865" LSID="urn:lsid:plazi:treatment:6553F2398FA7F2C374E7163BE0665CE3" httpUri="http://treatment.plazi.org/id/6553F2398FA7F2C374E7163BE0665CE3" lastPageNumber="36" pageId="21" pageNumber="36">
<subSubSection id="3587AC6752004C5F29AAC6C6110CF269" pageId="21" pageNumber="36" type="nomenclature">
<paragraph id="782144A3E69B6B6BB059DC6949E83A31" pageId="21" pageNumber="36">
49.
<taxonomicName id="0C1E563593CD18E59FA233DC45C87115" ID-CoL="NZ28" LSID="https://science.mnhn.fr/institution/mnhn/collection/rs/item/rs8501" authority="Lourenco, 2002" authorityName="Lourenco" authorityYear="2002" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buthus tassili" order="Scorpiones" pageId="21" pageNumber="36" phylum="Arthropoda" rank="species" species="tassili">
Buthus tassili
<normalizedToken id="7CB7E34773CA3F55D57EFE42DCFD2D1D" originalValue="Lourenço">Lourenco</normalizedToken>
, 2002
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection id="5BD3FEC2F154909C7EFA3900FB1E9A80" pageId="21" pageNumber="36" type="reference_group">
<paragraph id="4F1BD7E3E397986DDCD884E0EEDAC259" pageId="21" pageNumber="36">
<taxonomicName id="BE947EE6BE363CEF16B95B742910114A" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buthus tassili" order="Scorpiones" pageId="21" pageNumber="36" phylum="Arthropoda" rank="species" species="tassili">Buthus tassili</taxonomicName>
:
<bibRefCitation id="5B9E842993559AA66D789612B9AAF3A6" author="Lourenco, WR" journalOrPublisher="Biogeographica" pageId="30" pageNumber="45" pagination="109 - 127" title="Considerations sur les modeles de distribution et differentiation du genre Buthus Leach, 1815, avec la description d'une nouvelle espece des montagnes du Tassili des Ajjer, Algerie (Scorpiones, Buthidae)." volume="78" year="2002">
<normalizedToken id="EA5355E33093C96FCC9DC80991719C5F" originalValue="Lourenço">Lourenco</normalizedToken>
2002
</bibRefCitation>
: 113-115, fig. 10, 12, 14;
<bibRefCitation id="4FFEBFB0A8144E5ED0EE561ED4287FE7" author="Lourenco, WR" journalOrPublisher="Revue Suisse de Zoologie" pageId="30" pageNumber="45" pagination="875 - 912" title="Complements a la faune de scorpions (Arachnida) de l'Afrique du Nord, avec des considerations sur le genre Buthus Leach, 1815." volume="110" year="2003">
<normalizedToken id="EAE62E37243498E2461CD3468947C5AF" originalValue="Lourenço">Lourenco</normalizedToken>
2003
</bibRefCitation>
: 906-909, fig. 80-86; Rossi, Tropea and
<normalizedToken id="F54FF4A2A6481D8E791D40FF01D3F84A" originalValue="Yağmur">Yagmur</normalizedToken>
2013: 3-5, 7.
</paragraph>
<paragraph id="A869D4093F62345705C853294DC6C9A4" pageId="21" pageNumber="36">
<taxonomicName id="B0885BABF80C6490B9A861DA79B5A902" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buthus occitanus" order="Scorpiones" pageId="21" pageNumber="36" phylum="Arthropoda" rank="species" species="occitanus">Buthus occitanus</taxonomicName>
(MIS):
<bibRefCitation id="4A6B4A560215654E23ECCDE6838D469D" author="Pallary, P" journalOrPublisher="Bulletin de la Societe d'Histoire Naturelle de l'Afrique du Nord" pageId="33" pageNumber="48" pagination="133 - 141" title="Les scorpions du Sahara central." volume="20" year="1929">Pallary 1929</bibRefCitation>
: 134, 140;
<bibRefCitation id="4391728A79C412DE98B5978FADA5944A" author="Pallary, P" journalOrPublisher="Memoires de la Societe d'Histoire Naturelle d'Afrique du Nord" pageId="33" pageNumber="48" pagination="90 - 100" title="Scorpions du Sahara central. Mission du Hoggar III (Fevrier a Mai 1928)." volume="4" year="1934">Pallary 1934</bibRefCitation>
: 98-99.
</paragraph>
<paragraph id="D94252519683CDA2D7914BA512E34417" pageId="21" pageNumber="36">
<taxonomicName id="85672E1F0F6A939DAB828067F74D69F7" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buthus occitanus var. neeli" order="Scorpiones" pageId="21" pageNumber="36" phylum="Arthropoda" rank="variety" species="occitanus" subspecies="tunetanus" variety="neeli">Buthus occitanus tunetanus neeli</taxonomicName>
:
<bibRefCitation id="90630337A3BB2DA4526F86B38EF19C9B" author="Gysin, J" journalOrPublisher="Archives de l'Institut Pasteur d'Algerie" pageId="27" pageNumber="42" pagination="65 - 72" title="Une nouvelle variete de scorpion au Hoggar Buthusoccitanus (Am.) ssp tunetanus (Herbst) var neeli var. nov." volume="47" year="1969">Gysin 1969</bibRefCitation>
: 65-71, fig. 1-5;
<normalizedToken id="620F4A6CA746EAC5D19F03428F20D18B" originalValue="Peréz">Perez</normalizedToken>
1974: 22;
<bibRefCitation id="4EF3981CA64E1D4F8361BAB3F4E4CBDC" pageId="21" pageNumber="36">Touloun 2012</bibRefCitation>
: 37, 40.
</paragraph>
<paragraph id="C138591BD48DD7E398419BD878BCA793" pageId="21" pageNumber="36">
<taxonomicName id="B21E81C84D7AE8370A7819C4AA17ECAA" class="Arachnida" family="Buthidae" genus="Buthus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Buthus occitanus" order="Scorpiones" pageId="21" pageNumber="36" phylum="Arthropoda" rank="subspecies" species="occitanus" subspecies="tunetatus">Buthus occitanus tunetatus</taxonomicName>
,
<normalizedToken id="2931F21C820718AC53D86F0007E0D178" originalValue="“Spécimens">&quot;Specimens</normalizedToken>
des
<normalizedToken id="1692B8ABBFF35A714D7616C31BE5CED7" originalValue="régions">regions</normalizedToken>
montagneuses centrales du Sahara&quot;:
<bibRefCitation id="639BCBBE938DB9A9B53CA2371C879612" author="Vachon, M" journalOrPublisher="Institut Pasteur d'Algerie, Alger" pageId="35" pageNumber="50" title="Etudes sur les scorpions. (published 1948 - 1951 in Archives de l'Institut Pasteur d'Algerie, 1948, 26: 25 - 90, 162 - 208, 288 - 316, 441 - 481; 1949, 27: 66 - 100, 134 - 169, 281 - 288, 334 - 396; 1950, 28: 152 - 216, 382 - 413; 1951, 29: 46 - 104)." url="https://www.ntnu.no/ub/scorpion-files/vachon.php" year="1952 a">Vachon 1952a</bibRefCitation>
: 279.
</paragraph>
</subSubSection>
<subSubSection id="850D4901EBD139A022F167D7036B2447" pageId="21" pageNumber="36" type="type material">
<paragraph id="5A7735E7EDB4FB3795DC15C6C118E99D" pageId="21" pageNumber="36">Type material.</paragraph>
<paragraph id="166FDAD341C4D236953D8AC980705D79" pageId="21" pageNumber="36">1 M holotype (MNHN N° RS8501), Tin Tazarif (approx. 24.466°, 10.466°), Illizi, Algeria. Paratype: 1 F (MNHN N° RS8622), same locality.</paragraph>
</subSubSection>
<subSubSection id="904E4DE0850690834C12393EB083C564" pageId="21" pageNumber="36" type="distribution">
<paragraph id="758938D6A431D45443A5FC7CB76DF347" pageId="21" pageNumber="36">Distribution.</paragraph>
<paragraph id="0C08848B7049E731AB9D73B725118E78" pageId="21" pageNumber="36">
This species is known from a wide area around the Hoggar and Tassili
<normalizedToken id="A2BBCAD42999B0DD4968D9F31B41126A" originalValue="NAjjer">N'Ajjer</normalizedToken>
Mountains, including at least one locality in Libya.
</paragraph>
</subSubSection>
<subSubSection id="BEB52491832945ADA9F1E7E170D96CC9" pageId="21" pageNumber="36" type="remarks">
<paragraph id="7FB285542DB20B91DFE214248BA213E5" pageId="21" pageNumber="36">Remarks.</paragraph>
<paragraph id="0A3D85B298ED09167D7AC84F1993E391" pageId="21" pageNumber="36">
The Tin Tazarif coordinates given here, standing at 880 m a.s.l., do not match the altitude given for the point by
<bibRefCitation id="A7FA3C8CBCDCE06A60DE53DA121E8CAE" author="Lourenco, WR" journalOrPublisher="Biogeographica" pageId="30" pageNumber="45" pagination="109 - 127" title="Considerations sur les modeles de distribution et differentiation du genre Buthus Leach, 1815, avec la description d'une nouvelle espece des montagnes du Tassili des Ajjer, Algerie (Scorpiones, Buthidae)." volume="78" year="2002">
<normalizedToken id="A8295D038D7F80E588655E1FEDBD6556" originalValue="Lourenço">Lourenco</normalizedToken>
(2002)
</bibRefCitation>
, of 1.800 m a.s.l., but correspond well to the map location given by the author in Figure 6 of the same article. Nevertheless there are several locations in the Tassili
<normalizedToken id="A3DDD52F08E36D91CF6F0902A8A670AD" originalValue="NAjjer">N'Ajjer</normalizedToken>
Mountains at or above 1.800 m a.s.l., located closer to Jebel Azao, its highest peak. The species ranges from the Hoggar Mountains to the Tassili
<normalizedToken id="1947FB2B05715210E6599249B4DEDFD2" originalValue="NAjjer">N'Ajjer</normalizedToken>
Mountains, including Ghat in Libya.
<normalizedToken id="F5503AB6FB12B19E0F4801C42B62DD51" originalValue="Gysins">Gysin's</normalizedToken>
name, B. o. t.
<taxonomicName id="1A9F9365DDE4C4710296E258971A8E02" genus="Buthoidea" lsidName="Buthoidea neeli var. neeli" pageId="21" pageNumber="36" rank="variety" species="neeli" variety="neeli">neeli</taxonomicName>
, is not available under the ICZN article 10.2 and 45.5, as already stated by
<bibRefCitation id="9EA5DF0B0DD03B30D2A650BAF691B6CF" pageId="21" pageNumber="36">Fet and Lowe (2000)</bibRefCitation>
. Therefore, it cannot enter formal synonymy. Nevertheless, because
<normalizedToken id="1909C405132AEEF22981E3493FB1F1D5" originalValue="Gysins">Gysin's</normalizedToken>
description (1969) brings relevant taxonomic information (figures and new localities) for
<taxonomicName id="145E4DB77D4541E54C943E1090DEC8F4" lsidName="B. tassili" pageId="21" pageNumber="36" rank="species" species="tassili">B. tassili</taxonomicName>
, we have decided to clearly state this new informal synonymy. The specimens studied of both species come from the Algerian Hoggar Mountains and share a typical darkened fifth segment of the metasoma (see
<taxonomicName id="4585B30BC2556D71EB31D81D1F8B3640" lsidName="B. nigrovesiculosus" pageId="21" pageNumber="36" rank="species" species="nigrovesiculosus">B. nigrovesiculosus</taxonomicName>
above).
</paragraph>
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