treatments-xml/data/2B/25/82/2B25821F771753A3A0ACDB210B907CAF.xml
2024-06-21 12:32:19 +02:00

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<document ID-DOI="http://dx.doi.org/10.3897/aiep.51.63469" ID-GBIF-Dataset="ff4ff95f-52c8-4c59-b52d-d7a654405d54" ID-GBIF-Taxon="191980407" ID-Pensoft-Pub="1734-1515-1-77" ID-Pensoft-UUID="BA60DFBF2C245293A2B5BC8A9877F1F3" ID-ZooBank="84149491A8AD433B9E356F876B68FDF3" ModsDocID="1734-1515-51-1-77" checkinTime="1641859653430" checkinUser="pensoft" docAuthor="Teena, Jayakumar T. K., Murugan, A., Kumar, Ajith T. T. &amp; Lal, Kuldeep K." docDate="2021" docId="2B25821F771753A3A0ACDB210B907CAF" docLanguage="en" docName="ActIchthPisc 51(1): 77-83" docOrigin="Acta Ichthyologica et Piscatoria 51 (1)" docPubDate="2021-03-31" docSource="http://dx.doi.org/10.3897/aiep.51.63469" docTitle="Pycnocraspedum squamipinne Alcock 1889" docType="treatment" docVersion="4" id="BA60DFBF2C245293A2B5BC8A9877F1F3" lastPageNumber="77" masterDocId="BA60DFBF2C245293A2B5BC8A9877F1F3" masterDocTitle="Redescription of a rare cusk eel, Pycnocraspedum squamipinne (Actinopterygii, Ophidiiformes, Ophidiidae), from Bay of Bengal" masterLastPageNumber="83" masterPageNumber="77" pageId="0" pageNumber="77" updateTime="1641865355756" updateUser="ExternalLinkService">
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<mods:title>Redescription of a rare cusk eel, Pycnocraspedum squamipinne (Actinopterygii, Ophidiiformes, Ophidiidae), from Bay of Bengal</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Teena, Jayakumar T. K.</mods:namePart>
<mods:affiliation>ICAR - National Bureau of Fish Genetic Resources, Lucknow 226002, Uttar Pradesh, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Murugan, A.</mods:namePart>
<mods:affiliation>Vivekananda College, Kanyakumari 629701, Tamil Nadu, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Kumar, Ajith T. T.</mods:namePart>
<mods:affiliation>ICAR - National Bureau of Fish Genetic Resources, Lucknow 226002, Uttar Pradesh, India</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lal, Kuldeep K.</mods:namePart>
<mods:affiliation>ICAR - National Bureau of Fish Genetic Resources, Lucknow 226002, Uttar Pradesh, India</mods:affiliation>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>Acta Ichthyologica et Piscatoria</mods:title>
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<mods:part>
<mods:date>2021</mods:date>
<mods:detail type="pubDate">
<mods:number>2021-03-31</mods:number>
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<mods:detail type="volume">
<mods:number>51</mods:number>
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<mods:detail type="issue">
<mods:number>1</mods:number>
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<mods:start>77</mods:start>
<mods:end>83</mods:end>
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<mods:url>http://dx.doi.org/10.3897/aiep.51.63469</mods:url>
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<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/aiep.51.63469</mods:identifier>
<mods:identifier type="Pensoft-Pub">1734-1515-1-77</mods:identifier>
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<treatment ID-GBIF-Taxon="191980407" LSID="urn:lsid:plazi:treatment:2B25821F771753A3A0ACDB210B907CAF" httpUri="http://treatment.plazi.org/id/2B25821F771753A3A0ACDB210B907CAF" lastPageNumber="77" pageId="0" pageNumber="77">
<subSubSection pageId="0" pageNumber="77" type="nomenclature">
<paragraph pageId="0" pageNumber="77">
<taxonomicName LSID="2B25821F-7717-53A3-A0AC-DB210B907CAF" authority="Alcock, 1889" authorityName="Alcock" authorityYear="1889" class="Actinopterygii" family="Ophidiidae" genus="Pycnocraspedum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pycnocraspedum squamipinne" order="Ophidiiformes" pageId="0" pageNumber="77" phylum="Chordata" rank="species" species="squamipinne">Pycnocraspedum squamipinne Alcock, 1889</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="77" type="description">
<paragraph pageId="0" pageNumber="77">
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. (A) Pycnocraspedum squamipinne, 40.3 cm TL, off Chennai coast, Bay of Bengal; (B) Pycnocraspedum squamipinne syntype 29.9 cm TL, ZSI F 11703, Bay of Bengal." figureDoi="10.3897/aiep.51.63469.figure1" httpUri="https://binary.pensoft.net/fig/526609" pageId="0" pageNumber="77">Figs 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. (A) Basibranchial tooth patches of Pycnocraspedum squamipinne, 29.8 cm SL, scale bar = 1 cm; (B) Dentition in jaws premaxillary, vomerine, palatine, dentary, and basibranchial tooth patches, scale bar = 1 cm." figureDoi="10.3897/aiep.51.63469.figure2" httpUri="https://binary.pensoft.net/fig/526610" pageId="0" pageNumber="77">, 2</figureCitation>
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pycnocraspedum squamipinne, 20 cm SL (A) left sagitta, (B) right sagitta; (C) First gill raker of Pycnocraspedum squamipinne, 29.8 cm SL, scale bar = 1 cm." figureDoi="10.3897/aiep.51.63469.figure3" httpUri="https://binary.pensoft.net/fig/526611" pageId="0" pageNumber="77">, 3</figureCitation>
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Radiograph of Pycnocraspedum squamipinne, 38 cm SL." figureDoi="10.3897/aiep.51.63469.figure4" httpUri="https://binary.pensoft.net/fig/526612" pageId="0" pageNumber="77">, 4</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="77" type="reference_group">
<paragraph pageId="0" pageNumber="77">
<taxonomicName authorityName="Alcock" authorityYear="1889" class="Actinopterygii" family="Ophidiidae" genus="Pycnocraspedum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pycnocraspedum squamipinne" order="Ophidiiformes" pageId="0" pageNumber="77" phylum="Chordata" rank="species" species="squamipinne">Pycnocraspedum squamipinne</taxonomicName>
Alcock, 1889 (Type locality: Bay of Bengal,
<geoCoordinate degrees="20" direction="north" minutes="17" orientation="latitude" precision="15" seconds="30" value="20.291666">20°17'30&quot;N</geoCoordinate>
,
<geoCoordinate degrees="88" direction="east" minutes="50" orientation="longitude" precision="925" value="88.833336">88°50'E</geoCoordinate>
)
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="77" type="materials_examined">
<paragraph pageId="0" pageNumber="77">Materials examined.</paragraph>
<paragraph pageId="0" pageNumber="77">
<materialsCitation accessionNumber="F11700" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="M. S." country="INDIA" latitude="20.291666" location="Bay of Bengal" longLatPrecision="868" longitude="88.833336" specimenCount="1" typeStatus="Syntype">
<collectingCountry name="India">INDIA</collectingCountry>
1;
<typeStatus>Syntype</typeStatus>
of
<taxonomicName genus="P." lsidName="P. squamipinne" pageId="0" pageNumber="77" rank="species" species="squamipinne">
<emphasis italics="true" pageId="0" pageNumber="77">P. squamipinne</emphasis>
</taxonomicName>
(TL =
<quantity metricMagnitude="-1" metricUnit="m" metricValue="2.91" unit="mm" value="291.0">291 mm</quantity>
);
<location LSID="urn:lsid:plazi:treatment:2B25821F771753A3A0ACDB210B907CAF:80D72E14DADF3A4435C297B6EC3BEA47" country="INDIA" latitude="20.291666" longLatPrecision="868" longitude="88.833336" name="Bay of Bengal">Bay of Bengal</location>
;
<geoCoordinate degrees="20" direction="north" minutes="17" orientation="latitude" precision="15" seconds="30" value="20.291666">20°17'30&quot;N</geoCoordinate>
,
<geoCoordinate degrees="88" direction="east" minutes="50" orientation="longitude" precision="925" value="88.833336">88°50'E</geoCoordinate>
R.I.
<collectorName name="M Sudhakar">M.S.</collectorName>
<taxonomicName class="Actinopterygii" genus="Investigator" kingdom="Animalia" lsidName="Investigator" order="Ophidiiformes" pageId="0" pageNumber="77" phylum="Chordata" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="77">Investigator</emphasis>
</taxonomicName>
coll. leg.; ZSI
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/F11700">F11700</accessionNumber>
1
</materialsCitation>
;
<materialsCitation accessionNumber="F 11703" collectingDate="2021-01-01" collectingDateMax="2021-12-31" collectingDateMin="2021-01-01" collectorName="M Sudhakar" country="India" latitude="20.291666" location="Bay of Bengal" longLatPrecision="868" longitude="88.833336" specimenCount="1" typeStatus="Syntype">
<typeStatus>Syntype</typeStatus>
of
<taxonomicName genus="P." lsidName="P. squamipinne" pageId="0" pageNumber="77" rank="species" species="squamipinne">
<emphasis italics="true" pageId="0" pageNumber="77">P. squamipinne</emphasis>
</taxonomicName>
(TL =
<quantity metricMagnitude="-1" metricUnit="m" metricValue="2.9899999999999998" unit="mm" value="299.0">299 mm</quantity>
) same collection data as preceding; ZSI
<accessionNumber httpUri="http://www.ncbi.nlm.nih.gov/nucleotide/F11703">F 11703</accessionNumber>
(
<quantity metricMagnitude="0" metricUnit="m" metricValue="1.299" unit="mm" value="1299.0">1, 299 mm</quantity>
TL)
</materialsCitation>
.
</paragraph>
<paragraph pageId="0" pageNumber="77">
<materialsCitation collectingDate="2018-03" collectorName="Teena Jayakumar T. K, T. K and T. T Ajith Kumar" country="INDIA" location="Chennai" specimenCount="1" stateProvince="Tamil Nadu">
<collectingCountry name="India">INDIA</collectingCountry>
5;
<location LSID="urn:lsid:plazi:treatment:2B25821F771753A3A0ACDB210B907CAF:CBB2BD80842E03A5127F2776C9BE3A3C" country="INDIA" name="Chennai" stateProvince="Tamil Nadu">Chennai</location>
coast,
<collectingRegion country="India" name="Tamil Nadu">Tamil Nadu</collectingRegion>
;
<collectingDate value="2018-03">March 2018</collectingDate>
;
<collectorName>Teena Jayakumar
<collectorName>T.K and T.T Ajith Kumar</collectorName>
</collectorName>
leg.; NBFGR / OPHPSQU1 to 5 • 10; same collection data as for preceding; NBFGR / CE1 to 10
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="77" type="diagnosis">
<paragraph pageId="0" pageNumber="77">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="77">
Deep bodied cusk eel with head 3.0-3.9 times in SL; eye diameter 5.5-6.8 in HL; interorbital 4.8-6.3 in HL; snout 3.8-5.2 in HL; depth 4.3-6.8 in SL; predorsal length 3.3-4.7 in SL, distance between pelvic origin to anal origin 2.5-4.4 in SL; no pseudobranchiae and 12-13 pyloric caeca. Two median basibranchial tooth patches. Single opercular spine and 3 short blunt spines at lower angle of preopercle.
<taxonomicName authorityName="Alcock" authorityYear="1889" class="Actinopterygii" family="Ophidiidae" genus="Pycnocraspedum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pycnocraspedum squamipinne" order="Ophidiiformes" pageId="0" pageNumber="77" phylum="Chordata" rank="species" species="squamipinne">
<emphasis italics="true" pageId="0" pageNumber="77">Pycnocraspedum squamipinne</emphasis>
</taxonomicName>
differs from its congeners in origin of dorsal fin above preopercle, absence of pseudobranchial filaments, pectoral fins and pelvic fins short and not reaching anus.
</paragraph>
<caption doi="10.3897/aiep.51.63469.figure1" httpUri="https://binary.pensoft.net/fig/526609" pageId="0" pageNumber="77" start="Figure 1" startId="F1">
<paragraph pageId="0" pageNumber="77">
<emphasis bold="true" pageId="0" pageNumber="77">Figure 1.</emphasis>
<emphasis bold="true" pageId="0" pageNumber="77">(A</emphasis>
)
<taxonomicName authorityName="Alcock" authorityYear="1889" class="Actinopterygii" family="Ophidiidae" genus="Pycnocraspedum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pycnocraspedum squamipinne" order="Ophidiiformes" pageId="0" pageNumber="77" phylum="Chordata" rank="species" species="squamipinne">
<emphasis italics="true" pageId="0" pageNumber="77">Pycnocraspedum squamipinne</emphasis>
</taxonomicName>
, 40.3 cm TL, off Chennai coast, Bay of Bengal; (
<emphasis bold="true" pageId="0" pageNumber="77">B)</emphasis>
<taxonomicName authorityName="Alcock" authorityYear="1889" class="Actinopterygii" family="Ophidiidae" genus="Pycnocraspedum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pycnocraspedum squamipinne" order="Ophidiiformes" pageId="0" pageNumber="77" phylum="Chordata" rank="species" species="squamipinne">
<emphasis italics="true" pageId="0" pageNumber="77">Pycnocraspedum squamipinne</emphasis>
</taxonomicName>
syntype 29.9 cm TL, ZSI F11703, Bay of Bengal.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="77" type="description">
<paragraph pageId="0" pageNumber="77">Description.</paragraph>
<paragraph pageId="0" pageNumber="77">
Body compressed, moderately deep and tapers caudally, its depth 4.3-6.8 in SL (Fig.
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. (A) Pycnocraspedum squamipinne, 40.3 cm TL, off Chennai coast, Bay of Bengal; (B) Pycnocraspedum squamipinne syntype 29.9 cm TL, ZSI F 11703, Bay of Bengal." figureDoi="10.3897/aiep.51.63469.figure1" httpUri="https://binary.pensoft.net/fig/526609" pageId="0" pageNumber="77">1</figureCitation>
). Body covered by small cycloid scales. Head large, compressed, fully scaled except lips, head length 3.0-3.9 in SL. Eye diameter 15.2%-18.4% HL; preorbital length 19.2%-23.6% HL and postorbital length 61.9%-72.7% HL. Head length in preanal distance 1.6-1.9. Anterior nostril tubular, behind snout rim with small skin flap, smaller than posterior nostril being simple pore in front of eye. Eyes moderately large, elliptical, snout length greater than diameter of eye, interorbital space wide equal to or slightly greater than eye diameter. Mouth large and oblique, snout blunt, maxilla extending beyond posterior margin of eye, wide posteriorly. Granular teeth in bands in jaws, vomer and palatines, vomerine tooth-patch narrow, v-shaped, tongue pointed, reaching below vomer. Median basibranchial with elongate tooth patch from first gill arch to third and second small tooth patch near fourth gill arch followed by pair of teardrop-shaped (rounded anteriorly and pointed posteriorly) lower pharyngeal tooth patches (Fig.
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. (A) Basibranchial tooth patches of Pycnocraspedum squamipinne, 29.8 cm SL, scale bar = 1 cm; (B) Dentition in jaws premaxillary, vomerine, palatine, dentary, and basibranchial tooth patches, scale bar = 1 cm." figureDoi="10.3897/aiep.51.63469.figure2" httpUri="https://binary.pensoft.net/fig/526610" pageId="0" pageNumber="77">2</figureCitation>
). Sagittal otolith long and oval (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pycnocraspedum squamipinne, 20 cm SL (A) left sagitta, (B) right sagitta; (C) First gill raker of Pycnocraspedum squamipinne, 29.8 cm SL, scale bar = 1 cm." figureDoi="10.3897/aiep.51.63469.figure3" httpUri="https://binary.pensoft.net/fig/526611" pageId="0" pageNumber="77">3A, B</figureCitation>
). Ratio of length to height ranging from 2.4 to 2.9 times; length to thickness from 5.9 to 7.4 times. Pseudobranchial filaments absent. Developed gill rakers 4 on first gill arch, having small granular teeth (Fig.
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Pycnocraspedum squamipinne, 20 cm SL (A) left sagitta, (B) right sagitta; (C) First gill raker of Pycnocraspedum squamipinne, 29.8 cm SL, scale bar = 1 cm." figureDoi="10.3897/aiep.51.63469.figure3" httpUri="https://binary.pensoft.net/fig/526611" pageId="0" pageNumber="77">3C</figureCitation>
). Branchiostegal rays 8; 6 in ceratohyal and 2 on epihyal. Opercle thin, soft with weak posterodorsal spine, three broad and blunt spines at lower angle of preopercle. Origin of dorsal fin above hind margin of preopercle. Pectoral fins elongate with prominent skin flap above its base but not reaching anus. Pelvic fins united basally and separated distally. Caudal distinguishable from dorsal and anal by closer grouping of last 10 rays. Lateral line well developed, terminating at point about length of postorbit from caudal fin base. Stomach black with 13 long and finger-like pyloric caeca. Total of 12 precaudal vertebrae and 47-49 total vertebrae number (Fig.
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Radiograph of Pycnocraspedum squamipinne, 38 cm SL." figureDoi="10.3897/aiep.51.63469.figure4" httpUri="https://binary.pensoft.net/fig/526612" pageId="0" pageNumber="77">4</figureCitation>
). Meristic and morphometric characters of the specimen are presented in Tables
<tableCitation captionStart="Table 1" captionStartId="T1" captionText="Table 1. Comparison of morphometric and meristic characters of Pycnocraspedum squamipinne (n = 15) and syntypes (ZSI F 11700 and ZSI F 11703)." httpUri="http://table.plazi.org/id/BE3A931F473D9032CB19CCA02F023013" pageId="0" pageNumber="77" tableUuid="BE3A931F473D9032CB19CCA02F023013">1</tableCitation>
,
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Comparison of morphometric and meristic characters among Pycnocraspedum species." httpUri="http://table.plazi.org/id/A3633DC7AE3D82C969E321C42240882A" pageId="0" pageNumber="77" tableUuid="A3633DC7AE3D82C969E321C42240882A">2</tableCitation>
.
</paragraph>
<caption doi="10.3897/aiep.51.63469.figure2" httpUri="https://binary.pensoft.net/fig/526610" pageId="0" pageNumber="77" start="Figure 2" startId="F2">
<paragraph pageId="0" pageNumber="77">
<emphasis bold="true" pageId="0" pageNumber="77">Figure 2.</emphasis>
(
<emphasis bold="true" pageId="0" pageNumber="77">A</emphasis>
) Basibranchial tooth patches of
<taxonomicName authorityName="Alcock" authorityYear="1889" class="Actinopterygii" family="Ophidiidae" genus="Pycnocraspedum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pycnocraspedum squamipinne" order="Ophidiiformes" pageId="0" pageNumber="77" phylum="Chordata" rank="species" species="squamipinne">
<emphasis italics="true" pageId="0" pageNumber="77">Pycnocraspedum squamipinne</emphasis>
</taxonomicName>
, 29.8 cm SL, scale bar = 1 cm; (
<emphasis bold="true" pageId="0" pageNumber="77">B)</emphasis>
Dentition in jaws premaxillary, vomerine, palatine, dentary, and basibranchial tooth patches, scale bar = 1 cm.
</paragraph>
</caption>
<caption doi="10.3897/aiep.51.63469.figure3" httpUri="https://binary.pensoft.net/fig/526611" pageId="0" pageNumber="77" start="Figure 3" startId="F3">
<paragraph pageId="0" pageNumber="77">
<emphasis bold="true" pageId="0" pageNumber="77">Figure 3.</emphasis>
<taxonomicName authorityName="Alcock" authorityYear="1889" class="Actinopterygii" family="Ophidiidae" genus="Pycnocraspedum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pycnocraspedum squamipinne" order="Ophidiiformes" pageId="0" pageNumber="77" phylum="Chordata" rank="species" species="squamipinne">
<emphasis italics="true" pageId="0" pageNumber="77">Pycnocraspedum squamipinne</emphasis>
</taxonomicName>
, 20 cm SL (
<emphasis bold="true" pageId="0" pageNumber="77">A</emphasis>
) left sagitta, (
<emphasis bold="true" pageId="0" pageNumber="77">B</emphasis>
) right sagitta; (
<emphasis bold="true" pageId="0" pageNumber="77">C</emphasis>
) First gill raker of
<taxonomicName authorityName="Alcock" authorityYear="1889" class="Actinopterygii" family="Ophidiidae" genus="Pycnocraspedum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pycnocraspedum squamipinne" order="Ophidiiformes" pageId="0" pageNumber="77" phylum="Chordata" rank="species" species="squamipinne">
<emphasis italics="true" pageId="0" pageNumber="77">Pycnocraspedum squamipinne</emphasis>
</taxonomicName>
, 29.8 cm SL, scale bar = 1 cm.
</paragraph>
</caption>
<caption doi="10.3897/aiep.51.63469.figure4" httpUri="https://binary.pensoft.net/fig/526612" pageId="0" pageNumber="77" start="Figure 4" startId="F4">
<paragraph pageId="0" pageNumber="77">
<emphasis bold="true" pageId="0" pageNumber="77">Figure 4.</emphasis>
Radiograph of
<taxonomicName authorityName="Alcock" authorityYear="1889" class="Actinopterygii" family="Ophidiidae" genus="Pycnocraspedum" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pycnocraspedum squamipinne" order="Ophidiiformes" pageId="0" pageNumber="77" phylum="Chordata" rank="species" species="squamipinne">
<emphasis italics="true" pageId="0" pageNumber="77">Pycnocraspedum squamipinne</emphasis>
</taxonomicName>
, 38 cm SL.
</paragraph>
</caption>
</subSubSection>
<subSubSection pageId="0" pageNumber="77" type="color">
<paragraph pageId="0" pageNumber="77">Color.</paragraph>
<paragraph pageId="0" pageNumber="77">Body greyish brown but on ventral side dull silvery brown. Body without markings or spots. Distal margins of dorsal and anal fins dark black, ground color of dorsal and anal fin pale brown. Pectoral fins thick black but base of the pectoral light grey. Pelvic fin rays white with blackish tinge. Lateral line with blackish tinge. Orobranchial cavity greyish white, peritoneum black.</paragraph>
</subSubSection>
</treatment>
</document>